114 lines
16 KiB
XML
114 lines
16 KiB
XML
<document id="CE4CFD4C1C5D252C65C14C543D3065C4" ID-CLB-Dataset="301492" ID-DOI="10.23885/1814-3326-2017-13-1-41-49" ID-GBIF-Dataset="576c34d2-bc69-4740-9026-54ab6e0bdf97" ID-ISSN="2713-1785" ID-Zenodo-Dep="13273923" IM.bibliography_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.taxonomicNames_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1723143279861" checkinUser="felipe" docAuthor="Nabozhenko, M. V. & Keskin, B." docDate="2017" docId="E063791BFFC9F32463F0E9DA4EC1F81C" docLanguage="en" docName="CaucEntomolBull.13.1.41-49.pdf" docOrigin="Caucasian Entomological Bulletin (Caucas. entomol. bull.) 13 (1)" docSource="http://dx.doi.org/10.23885/1814-3326-2017-13-1-41-49" docStyle="DocumentStyle:6A2F040C6DCB19A13F917346BF1BCB54.2:CaucEntomolBull.2017-.journal_article" docStyleId="6A2F040C6DCB19A13F917346BF1BCB54" docStyleName="CaucEntomolBull.2017-.journal_article" docStyleVersion="2" docTitle="Helops Fabricius 1775, s. str." docType="treatment" docVersion="2" lastPageNumber="42" masterDocId="1C5A0163FFCBF3266052EB7D4D70FFF7" masterDocTitle="Taxonomic review of the genus Helops Fabricius, 1775 (Coleoptera: Tenebrionidae) of Turkey" masterLastPageNumber="49" masterPageNumber="41" pageNumber="42" updateTime="1723642514758" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="2BD6954E37DF6A9E36D1BF9C338084AF">Taxonomic review of the genus Helops Fabricius, 1775 (Coleoptera: Tenebrionidae) of Turkey</mods:title>
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<mods:namePart id="CD6CBB58B3C110523EC190FFB090A661">Nabozhenko, M. V.</mods:namePart>
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<mods:namePart id="B39836ABD8B93A672643CD74ADFA5746">Keskin, B.</mods:namePart>
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<mods:title id="8ADD2C5A3FBDC4BD5B50C11710586DE7">Caucasian Entomological Bulletin</mods:title>
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<mods:date id="E3C9DBB347F3A9EE5EC8237D5492FF09">2017</mods:date>
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<mods:title id="666CDC58E6BDD13EF751525E0AA42B49">Caucas. entomol. bull.</mods:title>
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<treatment id="E063791BFFC9F32463F0E9DA4EC1F81C" ID-DOI="http://doi.org/10.5281/zenodo.13321436" ID-Zenodo-Dep="13321436" LSID="urn:lsid:plazi:treatment:E063791BFFC9F32463F0E9DA4EC1F81C" httpUri="http://treatment.plazi.org/id/E063791BFFC9F32463F0E9DA4EC1F81C" lastPageNumber="42" pageId="2" pageNumber="42">
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<subSubSection id="20D09B86FFC9F32463F0E9DA4871FD4A" box="[930,1281,679,701]" pageId="2" pageNumber="42" type="nomenclature">
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<paragraph id="6875C80DFFC9F32463F0E9DA4871FD4A" blockId="2.[930,1281,623,701]" box="[930,1281,679,701]" pageId="2" pageNumber="42">
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<heading id="333D7F61FFC9F32463F0E9DA4871FD4A" box="[930,1281,679,701]" centered="true" fontSize="9" level="2" pageId="2" pageNumber="42" reason="3">
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Genus
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<taxonomicName id="AFCAB38EFFC9F32463BAE9DA495BFD4B" authority="Fabricius, 1775" authorityName="Fabricius" authorityYear="1775" box="[1000,1067,679,700]" class="Insecta" family="Leiodidae" genus="Helops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="42" phylum="Arthropoda" rank="genus" sensu="stricto">
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<emphasis id="5ABE141FFFC9F32463BAE9DA495BFD4B" box="[1000,1067,679,700]" italics="true" pageId="2" pageNumber="42">Helops</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="418DA964FFC9F3246463E9D54915FD4A" box="[1073,1125,680,701]" pageId="2" pageNumber="42" sensu="stricto">s. str.</taxonomicNameLabel>
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Fabricius, 1775
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="20D09B86FFC9F324630EE99D484BFD01" box="[860,1339,736,758]" pageId="2" pageNumber="42" type="type_taxon">
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<paragraph id="6875C80DFFC9F324630EE99D484BFD01" blockId="2.[860,1339,736,758]" box="[860,1339,736,758]" pageId="2" pageNumber="42">
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<typeStatus id="B77176AFFFC9F324630EE99D4EFEFD02" box="[860,910,736,757]" pageId="2" pageNumber="42">Type</typeStatus>
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species
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<taxonomicName id="AFCAB38EFFC9F32463B3E99D4847FD01" authority="(Linnaeus, 1875)" baseAuthorityName="Linnaeus" baseAuthorityYear="1875" box="[993,1335,736,758]" class="Insecta" family="Leiodidae" genus="Helops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="42" phylum="Arthropoda" rank="species" species="caeruleus">
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<emphasis id="5ABE141FFFC9F32463B3E99D49F8FD02" box="[993,1160,736,757]" italics="true" pageId="2" pageNumber="42">Helops caeruleus</emphasis>
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(Linnaeus, 1875)
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="20D09B86FFC9F324630EE9814EC1F81C" pageId="2" pageNumber="42" type="description">
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<paragraph id="6875C80DFFC9F324630EE9814808FA93" blockId="2.[813,1400,764,2027]" pageId="2" pageNumber="42">
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<emphasis id="5ABE141FFFC9F324630EE98149E5FCE6" bold="true" box="[860,1173,764,786]" pageId="2" pageNumber="42">General morphology of adults.</emphasis>
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Body large (length
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<quantity id="AF3265E8FFC9F3246503E9814E05FCD9" metricMagnitude="-2" metricUnit="m" metricValue="2.05" metricValueMax="2.7" metricValueMin="1.4" pageId="2" pageNumber="42" unit="mm" value="20.5" valueMax="27.0" valueMin="14.0">14– 27 mm</quantity>
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), slender, weakly or moderately convex, completely or partly with blue, purple or green-blue shade (Color plate 1: 1–6; Color plate 2: 7–9). Eyes large, strongly transverse. Punctation of head often dense and very coarse. Mentum with strongly protruded elevation at middle.Antennae long, reaching elytral third or quarter. Male antennomere 11 larger than 10
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<superScript id="9FBF6545FFC9F3246393E8BC4EBDFC3A" attach="left" box="[961,973,961,973]" fontSize="5" pageId="2" pageNumber="42">th</superScript>
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one, female antennomere 11 smaller or subequal to 10
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<superScript id="9FBF6545FFC9F3246397E8A34EA1FC1D" attach="left" box="[965,977,990,1002]" fontSize="5" pageId="2" pageNumber="42">th</superScript>
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one. Pronotum cordiform, margins with thickened bead, disc with coarse and dense punctation. Prothoracic hypomera with coarse large punctures, with punctures and wrinkles or with only coarse short irregular wrinkles. Elytra with protruded humoral angles, distinct striae. Epipleura not reaching thickened (not flattened) apex. Flightless, wings strongly reduced, very small, without costal and rarely radial veins. Mesocoxal cavities closed externally by mesepimera and metaventrite, trochantin presented. Abdominal ventrites with coarse and dense punctation, without hair brush; ventrite 5 beaded apically, without long and strong suberected setae near apex. Legs slender. Pro- and mesotibiae of male with very dense line of erected brush. Male protarsomeres 1–3 weakly widened.
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</paragraph>
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<paragraph id="6875C80DFFC9F324630EEE174995F8FE" blockId="2.[813,1400,764,2027]" pageId="2" pageNumber="42">
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Male genitalia and terminalia (
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<figureCitation id="F0F1D488FFC9F32464DFEE17498CFA88" box="[1165,1276,1386,1408]" captionStart="Figs 19–23" captionStartId="5.[237,268,1546,1563]" captionTargetBox="[206,1394,290,1516]" captionTargetId="figure-184@5.[189,1398,272,1525]" captionTargetPageId="5" captionText="Figs 19–23. Helops caeruleus sevenii, male genitalia and terminalia. 19 – aedeagus, ventral view; 20 – median piece (penis); 21 – aedeagus, lateral view; 22 – lobe of gastral spicula, lateral view; 23 – male inner sternite VIII, ventrally. Рис. 19–23. Helops caeruleus sevenii, генитаΛии и терминаΛии самца. 19 – эΑеагус вентраΛьно; 20 – пенис; 21 – эΑеагус ΛатераΛьно; 22 – Λопасть гастраΛьной спикуΛы, ΛатераΛьно; 23 – VIII внутренний стернит самца, вентраΛьно." figureDoi="http://doi.org/10.5281/zenodo.13273935" httpUri="https://zenodo.org/record/13273935/files/figure.png" pageId="2" pageNumber="42">Figs 19–23</figureCitation>
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,
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<figureCitation id="F0F1D488FFC9F3246554EE174836FA88" box="[1286,1350,1386,1407]" captionStart="Figs 24–28" captionStartId="6.[236,267,1568,1585]" captionTargetBox="[206,1393,287,1542]" captionTargetId="figure-309@6.[189,1398,272,1551]" captionTargetPageId="6" captionText="Figs 24–28. Helops rossii, male genitalia and terminalia. 24 – aedeagus, ventral view;25 – median piece (penis);26 – aedeagus,lateral view;27 – lobe of gastral spicula,lateral view;28 – male inner sternite VIII, dorsally. Рис. 24–28. Helops rossii, генитаΛии и терминаΛии самца. 24 – эΑеагус вентраΛьно; 25 – пенис; 26 – эΑеагус ΛатераΛьно; 27 – Λопасть гастраΛьной спикуΛы, ΛатераΛьно; 28 – VIII внутренний стернит самца, ΑорсаΛьно." figureDoi="http://doi.org/10.5281/zenodo.13273937" httpUri="https://zenodo.org/record/13273937/files/figure.png" pageId="2" pageNumber="42">24–28</figureCitation>
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,
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<figureCitation id="F0F1D488FFC9F3246502EE174E37FA6C" captionStart="Figs 29–33" captionStartId="7.[237,268,1519,1536]" captionTargetBox="[200,1385,283,1495]" captionTargetId="figure-288@7.[189,1398,272,1501]" captionTargetPageId="7" captionText="Figs 29–33. Helops glabriventris glabriventris, male genitalia and terminalia. 29 – aedeagus, ventral view; 30 – median piece (penis); 31 – aedeagus, lateral view; 32 – lobe of gastral spicula, lateral view; 33 – male inner sternite VIII, ventrally. Рис. 29–33. Helops glabriventris glabriventris, генитаΛии и терминаΛии самца. 29 – эΑеагус вентраΛьно; 30 – пенис; 31 – эΑеагус ΛатераΛьно; 32 – Λопасть гастраΛьной спикуΛы, ΛатераΛьно; 33 – VIII внутренний стернит самца, вентраΛьно." figureDoi="http://doi.org/10.5281/zenodo.13273939" httpUri="https://zenodo.org/record/13273939/files/figure.png" pageId="2" pageNumber="42">29– 33</figureCitation>
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,
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<figureCitation id="F0F1D488FFC9F3246303EEFB4EE2FA6C" box="[849,914,1414,1435]" captionStart="Figs 34–38" captionStartId="8.[236,267,1542,1559]" captionTargetBox="[190,1396,280,1519]" captionTargetId="figure-284@8.[189,1398,272,1525]" captionTargetPageId="8" captionText="Figs 34–38. Helops cyanipes, male genitalia and terminalia. 34 – aedeagus,ventral view;35 – median piece (penis);36 – aedeagus,lateral view;37 – lobe of gastral spicula,lateral view;38 – male inner sternite VIII, ventrally. Рис. 34–38. Helops cyanipes, генитаΛии и терминаΛии самца. 34 – эΑеагус вентраΛьно; 35 – пенис; 36 – эΑеагус ΛатераΛьно; 37 – Λопасть гастраΛьной спикуΛы, ΛатераΛьно; 38 – VIII внутренний стернит самца, вентраΛьно." figureDoi="http://doi.org/10.5281/zenodo.13273943" httpUri="https://zenodo.org/record/13273943/files/figure.png" pageId="2" pageNumber="42">34–38</figureCitation>
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,
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<figureCitation id="F0F1D488FFC9F32463CEEEFB4EAEFA6B" box="[924,990,1414,1436]" captionStart="Figs 39–43" captionStartId="9.[212,243,1685,1702]" captionTargetBox="[199,1386,276,1661]" captionTargetId="figure-155@9.[189,1398,272,1670]" captionTargetPageId="9" captionText="Figs 39–43.Helops punctatissimus sp. n., male genitalia and terminalia. 39 – aedeagus, ventral view; 40 – median piece (penis); 41 – aedeagus, lateral view; 42 – male inner sternite VIII, ventrally; 43 – gastral spicula. Рис.39–43. Helops punctatissimus sp. n., генитаΛии и терминаΛии самца. 39 – эΑеагус вентраΛьно; 40 – пенис; 41 – эΑеагус ΛатераΛьно; 42 – VIII внутренний стернит самца, вентраΛьно; 43 – гастраΛьная спикуΛа." figureDoi="http://doi.org/10.5281/zenodo.13273945" httpUri="https://zenodo.org/record/13273945/files/figure.png" pageId="2" pageNumber="42">39–43</figureCitation>
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). Aedeagus long; apical piece lanceolate, without longitudinal impression dorsally, covered with short spines only at widest apical part. Ventral alae of apical piece merged and completely conceal penis. Basal piece near 2 times longer than apical piece. Penis with two narrow baculi merged and broadened at basal third and with pair sclerotized part at middle. Apical lobes of gastral spicula large, pubescent apically; rods of gastral spicula straight, not merged apically and with long, often involute processes near base of lobes. Inner sternite VIII densely pubescent with long strong setae marginally; lobes with dense fine punctation and structures for rigidity of construction: pair sclerite at middle and additional sclerotized parts at middle, coming from lateral sides to middle.
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</paragraph>
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<paragraph id="6875C80DFFC9F324630EEC72496AF861" blockId="2.[813,1400,764,2027]" pageId="2" pageNumber="42">Female genital tubes. Spermatheca long, strongly branched, main duct with 3 additional long branched ducts and 2–3 short not branched ducts; basal duct short. Accessory gland long, with one-way valve, which has very short branch externally.</paragraph>
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<paragraph id="6875C80DFFC9F324630EECE14EC1F81C" blockId="2.[813,1400,764,2027]" pageId="2" pageNumber="42">
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<emphasis id="5ABE141FFFC9F324630EECE14EE7F846" bold="true" box="[860,919,1948,1969]" pageId="2" pageNumber="42">Note.</emphasis>
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Male genitalia of
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<taxonomicName id="AFCAB38EFFC9F3246406ECE149E7F846" authorityName="Fabricius" authorityYear="1775" box="[1108,1175,1948,1969]" class="Insecta" family="Leiodidae" genus="Helops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="42" phylum="Arthropoda" rank="genus">
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<emphasis id="5ABE141FFFC9F3246406ECE149E7F846" box="[1108,1175,1948,1969]" italics="true" pageId="2" pageNumber="42">Helops</emphasis>
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</taxonomicName>
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poorly differ because species of the genus
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<taxonomicName id="AFCAB38EFFC9F3246457ECC44938F839" authorityName="Fabricius" authorityYear="1775" box="[1029,1096,1977,1998]" class="Insecta" family="Leiodidae" genus="Helops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="42" phylum="Arthropoda" rank="genus">
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<emphasis id="5ABE141FFFC9F3246457ECC44938F839" box="[1029,1096,1977,1998]" italics="true" pageId="2" pageNumber="42">Helops</emphasis>
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</taxonomicName>
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are allopatric (at least in the Middle East).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |