480 lines
60 KiB
XML
480 lines
60 KiB
XML
<document id="02280294F4F9CC5C200122F612EA3A89" ID-CLB-Dataset="202083" ID-DOI="10.11646/zootaxa.5211.1.1" ID-GBIF-Dataset="bb97b9dd-2acc-4244-8f30-72bef07b9ed9" ID-ISSN="1175-5326" ID-Zenodo-Dep="7345007" ID-ZooBank="5754769C-B747-4714-BDD9-7D5509D48BEB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1669101340847" checkinUser="plazi" docAuthor="López-García, Margarita M. & Deloya, Cuauhtémoc" docDate="2022" docId="A209493BFD72FFAFAFA619C1FF55BDB0" docLanguage="en" docName="zootaxa.5211.1.1.pdf" docOrigin="Zootaxa 5211 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5211.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Ligyrus (Ligyrus) nasutus" docType="treatment" docVersion="9" lastPageNumber="47" masterDocId="5E303143FD5FFF81AF311956FFBEBC61" masterDocTitle="Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision" masterLastPageNumber="119" masterPageNumber="1" pageNumber="46" updateTime="1699396337306" updateUser="plazi">
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<mods:title id="B25FEDC7AB5BDDBE91D459A5FC4A6A8D">Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision</mods:title>
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<mods:namePart id="F2A7FF80B554093D3862F0A9389630BE">López-García, Margarita M.</mods:namePart>
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<mods:nameIdentifier id="9EA63A530DC35B8DF9BA26E017EC6030" type="email">margaralopezg@gmail.com</mods:nameIdentifier>
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<mods:namePart id="C9A25B9F3D00E8BBB7D8409D8DCF386A">Deloya, Cuauhtémoc</mods:namePart>
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<mods:nameIdentifier id="C8DE5C8A1E46C249536B77D441957C85" type="email">cuauhtemoc.deloya@inecol.mx</mods:nameIdentifier>
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<mods:title id="86A121CF7B2454E8B86B5838AD513609">Zootaxa</mods:title>
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<mods:part id="A4A09B42606E4D5220B16B8DA57C60F8">
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<mods:date id="65E84C7AED1749A03E8BA560D797F61D">2022</mods:date>
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<mods:number id="73488C66EA339AF00FAB673F7F464B32">2022-11-22</mods:number>
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<treatment id="A209493BFD72FFAFAFA619C1FF55BDB0" ID-DOI="http://doi.org/10.5281/zenodo.7383763" ID-GBIF-Taxon="205794482" ID-Zenodo-Dep="7383763" LSID="urn:lsid:plazi:treatment:A209493BFD72FFAFAFA619C1FF55BDB0" httpUri="http://treatment.plazi.org/id/A209493BFD72FFAFAFA619C1FF55BDB0" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46">
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<subSubSection id="62BAABA6FD72FFACAFA619C1FCCCBCD0" box="[151,882,151,177]" pageId="45" pageNumber="46" type="nomenclature">
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<paragraph id="2A1FF82DFD72FFACAFA619C1FCCCBCD0" blockId="45.[151,882,151,214]" box="[151,882,151,177]" pageId="45" pageNumber="46">
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<heading id="71574F41FD72FFACAFA619C1FCCCBCD0" bold="true" box="[151,882,151,177]" fontSize="11" level="2" pageId="45" pageNumber="46" reason="4">
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<emphasis id="18D4243FFD72FFACAFA619C1FCCCBCD0" bold="true" box="[151,882,151,177]" pageId="45" pageNumber="46">
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<taxonomicName id="EDA083AEFD72FFACAFA619C1FD7FBCD0" authority="(Burmeister, 1847)" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[151,705,151,177]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="nasutus" subGenus="Ligyrus">
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<emphasis id="18D4243FFD72FFACAFA619C1FF4ABCD0" bold="true" box="[151,244,151,177]" italics="true" pageId="45" pageNumber="46">Ligyrus</emphasis>
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(
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<emphasis id="18D4243FFD72FFACAE3419C1FEDCBCD0" bold="true" box="[261,354,151,177]" italics="true" pageId="45" pageNumber="46">Ligyrus</emphasis>
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)
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<emphasis id="18D4243FFD72FFACAE4219C1FE6CBCD0" bold="true" box="[371,466,151,177]" italics="true" pageId="45" pageNumber="46">nasutus</emphasis>
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||
(
|
||
<bibRefCitation id="4E3185DCFD72FFACAED219C1FD07BCD0" author="Burmeister, H." box="[483,697,151,177]" pageId="45" pageNumber="46" refId="ref88626" refString="Burmeister, H. (1847) Handbuch der Entomologie. Coleoptera Lamellicornia, Xylophila et Pectinicornia. Band 5. Enslin, Berlin, viii + 584 pp." type="book" year="1847">Burmeister, 1847</bibRefCitation>
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||
)
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||
</taxonomicName>
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||
revised status
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="62BAABA6FD72FFACAFA619EDFE5EBCB4" box="[151,480,187,214]" pageId="45" pageNumber="46" type="description">
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<paragraph id="2A1FF82DFD72FFACAFA619EDFE5EBCB4" blockId="45.[151,882,151,214]" box="[151,480,187,214]" pageId="45" pageNumber="46">
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(
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<figureCitation id="B29BE4A8FD72FFACAFAE19EDFF42BCB4" box="[159,252,187,213]" captionStart="FIGURE 1" captionStartId="81.[151,250,1535,1559]" captionTargetBox="[151,1436,181,1511]" captionTargetId="figure-17@81.[151,1436,181,1511]" captionTargetPageId="81" captionText="FIGURE 1. Head. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) Ligyrus (Ligyrodes) peruvianus, D) L. (Anagrylius) moroni, E) L. (Ligyrus) ruginasus, F) L. (Ligyrus) nasutus, G) L. (Ligyrus) bidentulus, H) Tomarus adoceteus, I) T. maimon, J) T. pullus, K) T. pumilus." figureDoi="http://doi.org/10.5281/zenodo.7345011" httpUri="https://zenodo.org/record/7345011/files/figure.png" pageId="45" pageNumber="46">Figs. 1F</figureCitation>
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,
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<figureCitation id="B29BE4A8FD72FFACAE3919EDFE85BCB7" box="[264,315,187,214]" captionStart="FIGURE 15" captionStartId="90.[151,250,1535,1559]" captionTargetBox="[154,1430,183,1509]" captionTargetId="figure-24@90.[151,1436,181,1512]" captionTargetPageId="90" captionText="FIGURE 15. Parameres in frontal view. A) Euligyrus ebenus, B) E. similis, C) Proculigyrus cicatricosus, D) Ligyrus (Anagrylius) cuniculus, E) L. (Anagrylius) moroni, F) L. (Ligyrodes) peruvianus, G) L. (Ligyrodes) relictus, H) L. (Ligyrodes) sallaei, I) L. (Ligyrus) allonasutus, J) L. (Ligyrus) bidentulus, K) L. (Ligyrus) burmeisteri, L) Ligyrus (Ligyrus) fossor, M) L. (Ligyrus) gibbosus, N) L. (Ligyrus) nasutus, O) L. (Ligyrus) neglectus, P) L. (Ligyrus) paranaensis, Q) L. (Ligyrus) peninsularis, R) L. (Ligyrus) rubripes, S) L. (Ligyrus) ruginasus, T) L. (Ligyrus) villosus." figureDoi="http://doi.org/10.5281/zenodo.7345063" httpUri="https://zenodo.org/record/7345063/files/figure.png" pageId="45" pageNumber="46">15N</figureCitation>
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,
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||
<figureCitation id="B29BE4A8FD72FFACAE7719EDFECCBCB4" box="[326,370,187,213]" captionStart="FIGURE 18" captionStartId="93.[151,250,1597,1621]" captionTargetBox="[152,1436,184,1567]" captionTargetId="figure-17@93.[151,1436,181,1573]" captionTargetPageId="93" captionText="FIGURE 18. Parameres in lateral view.A) L. (Ligyrus.) allonasutus, B) L. (Ligyrus) bidentulus, C) L. (Ligyrus) burmeisteri, D) Ligyrus (Ligyrus) fossor, E) L. (Ligyrus) gibbosus, F) L. (Ligyrus) nasutus, G) L. (Ligyrus) neglectus, H) L. (Ligyrus) paranaensis, I) L. (Ligyrus) peninsularis, J) L. (Ligyrus) rubripes, K) L. (Ligyrus) ruginasus, L) L. (Ligyrus) villosus." figureDoi="http://doi.org/10.5281/zenodo.7345077" httpUri="https://zenodo.org/record/7345077/files/figure.png" pageId="45" pageNumber="46">18F</figureCitation>
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,
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||
<figureCitation id="B29BE4A8FD72FFACAE4F19EDFE0EBCB4" box="[382,432,187,213]" captionStart="FIGURE 26" captionStartId="101.[151,250,1827,1851]" captionTargetBox="[159,1430,189,1791]" captionTargetId="figure-17@101.[151,1436,181,1803]" captionTargetPageId="101" captionText="FIGURE 26. Habitus of the species of Ligyrus. Subgenus Ligyrus: A) Ligyrus fossor, B) L. gibbosus, C) L. nasutus, D) L. neglectus, E) L. paranaensis, F) L. peninsularis, G) L. rubripes, H) L. ruginasus, I) L. villosus. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7345101" httpUri="https://zenodo.org/record/7345101/files/figure.png" pageId="45" pageNumber="46">26C</figureCitation>
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||
;
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||
<figureCitation id="B29BE4A8FD72FFACAE8A19EDFE66BCB4" box="[443,472,187,213]" captionStart="FIGURE 34" captionStartId="107.[151,250,2007,2031]" captionTargetBox="[246,1346,1181,1977]" captionTargetId="figure-32@107.[242,1346,1181,1983]" captionTargetPageId="107" captionText="FIGURE 34. Distributional records of Ligyrus (Ligyrus) allonasutus and L. nasutus." figureDoi="http://doi.org/10.5281/zenodo.7345125" httpUri="https://zenodo.org/record/7345125/files/figure.png" pageId="45" pageNumber="46">34</figureCitation>
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||
)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="62BAABA6FD72FFACAFA61856FC98BD79" box="[151,806,256,280]" pageId="45" pageNumber="46" type="reference_group">
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<paragraph id="2A1FF82DFD72FFACAFA61856FC98BD79" blockId="45.[151,1437,255,586]" box="[151,806,256,280]" pageId="45" pageNumber="46">
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<treatmentCitationGroup id="0AB0DF03FD72FFACAFA61856FC98BD79" box="[151,806,256,280]" pageId="45" pageNumber="46">
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<treatmentCitation id="AB01DE3CFD72FFACAFA61856FD84BD79" author="Burmeister, H." box="[151,570,256,280]" page="120" pageId="45" pageNumber="46" year="1847">
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<taxonomicName id="EDA083AEFD72FFACAFA61856FD84BD79" authority="Burmeister, 1847: 120" authorityName="Burmeister" authorityPageNumber="120" authorityYear="1847" box="[151,570,256,280]" class="Insecta" family="Dynastidae" genus="Podalgus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="nasutus">
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||
<emphasis id="18D4243FFD72FFACAFA61856FEF5BD76" box="[151,331,256,280]" italics="true" pageId="45" pageNumber="46">Podalgus nasutus</emphasis>
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<bibRefCitation id="4E3185DCFD72FFACAE621856FD84BD79" author="Burmeister, H." box="[339,570,256,280]" pageId="45" pageNumber="46" refId="ref88626" refString="Burmeister, H. (1847) Handbuch der Entomologie. Coleoptera Lamellicornia, Xylophila et Pectinicornia. Band 5. Enslin, Berlin, viii + 584 pp." type="book" year="1847">Burmeister, 1847: 120</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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. Original combination.
|
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="62BAABA6FD72FFACAC1D1856FBCDBE2B" pageId="45" pageNumber="46" type="materials_examined">
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<paragraph id="2A1FF82DFD72FFACAC1D1856FB0FBD34" blockId="45.[151,1437,255,586]" pageId="45" pageNumber="46">
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||
Female
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||
<typeStatus id="F51B468FFD72FFACAC4F1856FC65BD79" box="[894,987,256,280]" pageId="45" pageNumber="46" type="lectotype">lectotype</typeStatus>
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||
designated by
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||
<treatmentCitation id="AB01DE3CFD72FFACAB471856FA87BD76" author="Endrodi, S." box="[1142,1337,255,280]" page="59" pageId="45" pageNumber="46" year="1969">
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<bibRefCitation id="4E3185DCFD72FFACAB471856FA87BD76" author="Endrodi, S." box="[1142,1337,255,280]" pageId="45" pageNumber="46" pagination="1 - 145" refId="ref89768" refString="Endrodi, S. (1969) Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). Entomologische Abhandlungen, 37, 1 - 145." type="journal article" year="1969">Endrödi (1969: 59)</bibRefCitation>
|
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</treatmentCitation>
|
||
(MLUH) “nasutus* / Yucat. / Berg //
|
||
<typeStatus id="F51B468FFD72FFACAED81849FDE4BD56" box="[489,602,287,311]" pageId="45" pageNumber="46" type="lectotype">Lectotypus</typeStatus>
|
||
/
|
||
<taxonomicName id="EDA083AEFD72FFACAD401848FD7EBD57" authorityName="Casey" authorityYear="1915" box="[625,704,286,310]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">Ligyrus</taxonomicName>
|
||
/ (
|
||
<taxonomicName id="EDA083AEFD72FFACADEF1848FC80BD57" authorityName="Burmeister" authorityYear="1847" box="[734,830,286,310]" class="Insecta" family="Dynastidae" genus="Podalgus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">Podalgus</taxonomicName>
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||
) /
|
||
<taxonomicName id="EDA083AEFD72FFACAC6D1849FC54BD57" authority="Burm" authorityName="Burmeister" authorityYear="1847" box="[860,1002,287,310]" class="Insecta" family="Dynastidae" genus="Podalgus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="nasutus">nasutus Burm</taxonomicName>
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||
/ Endrody” Type locality:
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||
<collectingCountry id="52B7B8BDFD72FFACAA331848FACEBD57" box="[1282,1392,286,310]" name="Guatemala" pageId="45" pageNumber="46">Guatemala</collectingCountry>
|
||
and Yucatan. Although
|
||
<collectingCountry id="52B7B8BDFD72FFACAEBB186BFE46BD34" box="[394,504,317,341]" name="Guatemala" pageId="45" pageNumber="46">Guatemala</collectingCountry>
|
||
is mentioned in the original description no
|
||
<typeStatus id="F51B468FFD72FFACAC83186BFB84BD34" box="[946,1082,317,341]" pageId="45" pageNumber="46" type="paralectotype">paralectotype</typeStatus>
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||
was found.
|
||
</paragraph>
|
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<paragraph id="2A1FF82DFD72FFACAFA6180AFBCDBE2B" blockId="45.[151,1437,255,586]" pageId="45" pageNumber="46">
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<treatmentCitationGroup id="0AB0DF03FD72FFACAFA6180AFBCDBE2B" pageId="45" pageNumber="46">
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<treatmentCitation id="AB01DE3CFD72FFACAFA6180AFDB8BD15" author="Bates, H. W." box="[151,518,348,372]" page="317" pageId="45" pageNumber="46" year="1888">
|
||
<taxonomicName id="EDA083AEFD72FFACAFA6180AFDB8BD15" authority="Bates, 1888: 317" authorityName="Bates" authorityPageNumber="317" authorityYear="1888" box="[151,518,348,372]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="pygidialis">
|
||
<emphasis id="18D4243FFD72FFACAFA6180AFEEEBD15" box="[151,336,348,372]" italics="true" pageId="45" pageNumber="46">Ligyrus pygidialis</emphasis>
|
||
<materialsCitation id="9AC8F270FD72FFACAE66180AFDB8BD15" ID-GBIF-Occurrence="3986696855" box="[343,518,348,372]" location="Male" municipality="Original" pageId="45" pageNumber="46" specimenCount="1" typeStatus="lectotype">
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<bibRefCitation id="4E3185DCFD72FFACAE66180AFDB8BD15" author="Bates, H. W." box="[343,518,348,372]" pageId="45" pageNumber="46" pagination="296 - 342" refId="ref88329" refString="Bates, H. W. (1888) Pectinicornia and Lamellicornia, Family Dynastinae. In: Godman, F. D. & Salvin, O. (Eds.), Biologia Centrali-Americana, Insecta, Coleoptera, 2 (Part 2), pp. 296 - 342." type="journal article" year="1888">Bates, 1888: 317</bibRefCitation>
|
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</materialsCitation>
|
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</taxonomicName>
|
||
</treatmentCitation>
|
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.
|
||
<collectingMunicipality id="CA7B6257FD72FFACAD21180AFDDABD15" box="[528,612,348,372]" pageId="45" pageNumber="46">Original</collectingMunicipality>
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||
combination.
|
||
<location id="2F7FAEF6FD72FFACADC9180AFC92BD15" LSID="urn:lsid:plazi:treatment:A209493BFD72FFAFAFA619C1FF55BDB0:2F7FAEF6FD72FFACADC9180AFC92BD15" box="[760,812,348,372]" municipality="Original" name="Male" pageId="45" pageNumber="46">Male</location>
|
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<typeStatus id="F51B468FFD72FFACAC02180AFC2EBD15" box="[819,912,348,372]" pageId="45" pageNumber="46" type="lectotype">lectotype</typeStatus>
|
||
<materialsCitation id="9AC8F270FD72FFACACA6180AFB90BDD0" ID-GBIF-Occurrence="3986696494" collectionCode="BMNH" location="Yucatan" pageId="45" pageNumber="46" specimenCount="1" specimenCount-male="1" stateProvince="Yucatan" typeStatus="lectotype">
|
||
designated by
|
||
<treatmentCitation id="AB01DE3CFD72FFACAB1B180AFB53BD12" author="Endrodi, S." box="[1066,1261,347,372]" page="59" pageId="45" pageNumber="46" year="1969">
|
||
<bibRefCitation id="4E3185DCFD72FFACAB1B180AFB53BD12" author="Endrodi, S." box="[1066,1261,347,372]" pageId="45" pageNumber="46" pagination="1 - 145" refId="ref89768" refString="Endrodi, S. (1969) Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). Entomologische Abhandlungen, 37, 1 - 145." type="journal article" year="1969">Endrödi (1969: 59)</bibRefCitation>
|
||
</treatmentCitation>
|
||
(
|
||
<collectionCode id="4CB160E8FD72FFACABCD180AFAF5BD15" box="[1276,1355,348,372]" pageId="45" pageNumber="46">BMNH</collectionCode>
|
||
) “LECTO- / TYPE // Type // B.C.A., Col.,II(2) /
|
||
<taxonomicName id="EDA083AEFD72FFACADB1182CFD71BDF3" authorityName="Casey" authorityYear="1915" box="[640,719,378,402]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">Ligyrus</taxonomicName>
|
||
/ pygidialis. // N.
|
||
<collectingRegion id="E86436CFFD72FFACACBC182DFC5CBDF3" box="[909,994,379,402]" country="Mexico" name="Yucatan" pageId="45" pageNumber="46">Yucatan</collectingRegion>
|
||
. / Gaumer. //
|
||
<taxonomicName id="EDA083AEFD72FFACAB40182CFB7EBDF3" authorityName="Casey" authorityYear="1915" box="[1137,1216,378,402]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">Ligyrus</taxonomicName>
|
||
/ pygidialis / Bates
|
||
<specimenCount id="3CA633A4FD72FFACAABA182DFA22BDF3" box="[1419,1436,379,402]" pageId="45" pageNumber="46" type="male">♂</specimenCount>
|
||
//
|
||
<typeStatus id="F51B468FFD72FFACAFEC18CFFEF0BDD0" box="[221,334,409,433]" pageId="45" pageNumber="46" type="lectotype">Lectotypus</typeStatus>
|
||
/
|
||
<taxonomicName id="EDA083AEFD72FFACAE5218CFFE0CBDD0" authorityName="Casey" authorityYear="1915" box="[355,434,409,433]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">Ligyrus</taxonomicName>
|
||
/ pygidialis / Bat. / Endrody”. Female
|
||
<typeStatus id="F51B468FFD72FFACAC7018CFFC77BDD0" box="[833,969,409,433]" pageId="45" pageNumber="46" type="paralectotype">paralectotype</typeStatus>
|
||
(
|
||
<collectionCode id="4CB160E8FD72FFACACE918CFFB99BDD0" box="[984,1063,409,433]" pageId="45" pageNumber="46">BMNH</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
<materialsCitation id="9AC8F270FD72FFACAB0418CFFE9ABD8F" ID-GBIF-Occurrence="3986696459" collectionCode="BMNH" collectorName="Male" location="Yucatan" pageId="45" pageNumber="46" specimenCount="1" specimenCount-female="1" stateProvince="Yucatan" typeStatus="paralectotype">
|
||
“PARA- / LECTO- / TYPE // Type // B.C.A., Col.,II(2) /
|
||
<taxonomicName id="EDA083AEFD72FFACAE9918EEFE49BDB1" authorityName="Casey" authorityYear="1915" box="[424,503,440,464]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">Ligyrus</taxonomicName>
|
||
/ pygidialis. // N.
|
||
<collectingRegion id="E86436CFFD72FFACAD8218EEFCB6BDAE" box="[691,776,440,463]" country="Mexico" name="Yucatan" pageId="45" pageNumber="46">Yucatan</collectingRegion>
|
||
. / Gaumer. //
|
||
<taxonomicName id="EDA083AEFD72FFACACA418EEFC5ABDB1" authorityName="Casey" authorityYear="1915" box="[917,996,440,464]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">Ligyrus</taxonomicName>
|
||
/ pygidialis / Bates
|
||
<specimenCount id="3CA633A4FD72FFACAB8418EEFB7CBDB1" box="[1205,1218,440,464]" pageId="45" pageNumber="46" type="female">♀</specimenCount>
|
||
”.
|
||
<collectorName id="87559DFBFD72FFACABE918EEFAB2BDB1" box="[1240,1292,440,464]" pageId="45" pageNumber="46">Male</collectorName>
|
||
<typeStatus id="F51B468FFD72FFACAA2518EEFA22BDB1" box="[1300,1436,440,464]" pageId="45" pageNumber="46" type="paralectotype">paralectotype</typeStatus>
|
||
(
|
||
<collectionCode id="4CB160E8FD72FFACAFFF1880FEA3BD8F" box="[206,285,470,494]" pageId="45" pageNumber="46">BMNH</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
<materialsCitation id="9AC8F270FD72FFACAE181880FD10BE6C" ID-GBIF-Occurrence="3986697002" location="There" pageId="45" pageNumber="46" specimenCount="13" stateProvince="Yucatan" typeStatus="paralectotype">
|
||
“PARA- / LECTO- / TYPE // B.C.A., Col.,II(2) /
|
||
<taxonomicName id="EDA083AEFD72FFACAC261880FCD8BD8F" authorityName="Casey" authorityYear="1915" box="[791,870,470,494]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">Ligyrus</taxonomicName>
|
||
/ pygidialis. // N.
|
||
<collectingRegion id="E86436CFFD72FFACAB261881FBD2BD8F" box="[1047,1132,471,494]" country="Mexico" name="Yucatan" pageId="45" pageNumber="46">Yucatan</collectingRegion>
|
||
. / Gaumer.”.
|
||
<location id="2F7FAEF6FD72FFACABDC1880FA96BD8F" LSID="urn:lsid:plazi:treatment:A209493BFD72FFAFAFA619C1FF55BDB0:2F7FAEF6FD72FFACABDC1880FA96BD8F" box="[1261,1320,470,494]" name="There" pageId="45" pageNumber="46" stateProvince="Yucatan">There</location>
|
||
are
|
||
<specimenCount id="3CA633A4FD72FFACAA631880FF42BE6C" pageId="45" pageNumber="46" type="generic" typeStatus="syntypes">13 syntypes</specimenCount>
|
||
which should be labeled as
|
||
<typeStatus id="F51B468FFD72FFACAD2D18A3FD10BE6C" box="[540,686,501,525]" pageId="45" pageNumber="46" type="paralectotype">paralectotypes</typeStatus>
|
||
</materialsCitation>
|
||
<materialsCitation id="9AC8F270FD72FFACAD8418A3FD47BE4D" ID-GBIF-Occurrence="3986696663" collectionCode="MNHN" location="The" pageId="45" pageNumber="46" specimenCode="EC7079-7091" specimenCount="1" stateProvince="Yucatan" typeStatus="paratype">
|
||
at
|
||
<collectionCode id="4CB160E8FD72FFACADFF18A3FCA5BE6D" box="[718,795,501,524]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="45" pageNumber="46">MNHN</collectionCode>
|
||
(
|
||
<specimenCode id="7A065056FD72FFACAC1818A3FC02BE6C" box="[809,956,501,525]" collectionCode="EC" pageId="45" pageNumber="46">EC7079-7091</specimenCode>
|
||
).
|
||
<location id="2F7FAEF6FD72FFACACFC18A3FC4ABE6C" LSID="urn:lsid:plazi:treatment:A209493BFD72FFAFAFA619C1FF55BDB0:2F7FAEF6FD72FFACACFC18A3FC4ABE6C" box="[973,1012,501,525]" name="The" pageId="45" pageNumber="46" stateProvince="Yucatan">The</location>
|
||
following are considered invalid types as the species was not originally described with
|
||
<typeStatus id="F51B468FFD72FFACADA51B42FD47BE4D" box="[660,761,532,556]" pageId="45" pageNumber="46" type="paratype">paratypes</typeStatus>
|
||
</materialsCitation>
|
||
: Two male
|
||
<typeStatus id="F51B468FFD72FFACAC5C1B42FC70BE4D" box="[877,974,532,556]" pageId="45" pageNumber="46" type="paratype">paratypes</typeStatus>
|
||
<materialsCitation id="9AC8F270FD72FFACACE41B42FA87BE4D" ID-GBIF-Occurrence="3986696451" box="[981,1337,532,556]" collectionCode="USNM" pageId="45" pageNumber="46" specimenCount="1" specimenCount-female="1" stateProvince="Yucatan" typeStatus="paratype">
|
||
and
|
||
<specimenCount id="3CA633A4FD72FFACAB311B42FBCFBE4D" box="[1024,1137,532,556]" pageId="45" pageNumber="46" type="female">two female</specimenCount>
|
||
<typeStatus id="F51B468FFD72FFACAB461B42FB66BE4D" box="[1143,1240,532,556]" pageId="45" pageNumber="46" type="paratype">paratypes</typeStatus>
|
||
(
|
||
<collectionCode id="4CB160E8FD72FFACABD71B42FA8CBE4D" box="[1254,1330,532,556]" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="45" pageNumber="46" type="Museum">USNM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
<materialsCitation id="9AC8F270FD72FFACAA0E1B42FBD0BE2B" ID-GBIF-Occurrence="3986696515" collectionCode="USNM" location="Bates" pageId="45" pageNumber="46" specimenCount="2" stateProvince="Yucatan" typeStatus="paratype">
|
||
“
|
||
<collectingRegion id="E86436CFFD72FFACAA791B42FA22BE4D" box="[1352,1436,532,556]" country="Mexico" name="Yucatan" pageId="45" pageNumber="46">Yucatan</collectingRegion>
|
||
/ Temax / Gaumer //
|
||
<collectionCode id="4CB160E8FD72FFACAEA81B64FE5CBE2B" box="[409,482,562,586]" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="45" pageNumber="46" type="Museum">USNM</collectionCode>
|
||
/
|
||
<typeStatus id="F51B468FFD72FFACAEC71B64FDCABE2B" box="[502,628,562,586]" pageId="45" pageNumber="46" type="paratype">PARATYPE</typeStatus>
|
||
/ 49710 //
|
||
<taxonomicName id="EDA083AEFD72FFACADD01B64FC8EBE2B" authorityName="Casey" authorityYear="1915" box="[737,816,562,586]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">Ligyrus</taxonomicName>
|
||
/ pygidialis /
|
||
<location id="2F7FAEF6FD72FFACAC8A1B65FC4DBE2B" LSID="urn:lsid:plazi:treatment:A209493BFD72FFAFAFA619C1FF55BDB0:2F7FAEF6FD72FFACAC8A1B65FC4DBE2B" box="[955,1011,563,586]" name="Bates" pageId="45" pageNumber="46" stateProvince="Yucatan">Bates</location>
|
||
/
|
||
<typeStatus id="F51B468FFD72FFACAB361B64FBDBBE2B" box="[1031,1125,562,586]" pageId="45" pageNumber="46" type="paratype">Paratype</typeStatus>
|
||
”
|
||
</materialsCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="62BAABA6FD72FFACAFA61B2EFDF8B897" pageId="45" pageNumber="46" type="description">
|
||
<paragraph id="2A1FF82DFD72FFACAFA61B2EFDF8B897" blockId="45.[151,1437,632,2027]" pageId="45" pageNumber="46">
|
||
<emphasis id="18D4243FFD72FFACAFA61B2EFE97BEF3" bold="true" box="[151,297,632,658]" pageId="45" pageNumber="46">Description.</emphasis>
|
||
Habitus as in
|
||
<figureCitation id="B29BE4A8FD72FFACAEE81B2EFDFDBEF3" box="[473,579,632,658]" captionStart="FIGURE 26" captionStartId="101.[151,250,1827,1851]" captionTargetBox="[159,1430,189,1791]" captionTargetId="figure-17@101.[151,1436,181,1803]" captionTargetPageId="101" captionText="FIGURE 26. Habitus of the species of Ligyrus. Subgenus Ligyrus: A) Ligyrus fossor, B) L. gibbosus, C) L. nasutus, D) L. neglectus, E) L. paranaensis, F) L. peninsularis, G) L. rubripes, H) L. ruginasus, I) L. villosus. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7345101" httpUri="https://zenodo.org/record/7345101/files/figure.png" pageId="45" pageNumber="46">Fig. 26C</figureCitation>
|
||
. Length
|
||
<quantity id="ED5855C8FD72FFACAD9D1B2FFCE8BEF2" box="[684,854,633,659]" metricMagnitude="-2" metricUnit="m" metricValue="1.61" metricValueMax="1.7399999999999998" metricValueMin="1.48" pageId="45" pageNumber="46" unit="mm" value="16.1" valueMax="17.4" valueMin="14.8">14.8–17.4 mm</quantity>
|
||
; humeral width
|
||
<quantity id="ED5855C8FD72FFACAB261B2FFB0DBEF2" box="[1047,1203,633,659]" metricMagnitude="-3" metricUnit="m" metricValue="8.754999999999999" metricValueMax="9.2" metricValueMin="8.31" pageId="45" pageNumber="46" unit="mm" value="8.754999999999999" valueMax="9.2" valueMin="8.31">8.31–9.2 mm</quantity>
|
||
. Color dark reddish brown.
|
||
<emphasis id="18D4243FFD72FFACAFC11BCBFE87BED6" bold="true" box="[240,313,669,695]" pageId="45" pageNumber="46">Head:</emphasis>
|
||
Frons deeply rugopunctate, with large and sparse punctures, nearly smooth on vertex, with only sparse, small punctures. Frontoclypeal region with a bilobed, complete carina (
|
||
<figureCitation id="B29BE4A8FD72FFACAB381B96FBE3BEBA" box="[1033,1117,704,731]" captionStart="FIGURE 1" captionStartId="81.[151,250,1535,1559]" captionTargetBox="[151,1436,181,1511]" captionTargetId="figure-17@81.[151,1436,181,1511]" captionTargetPageId="81" captionText="FIGURE 1. Head. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) Ligyrus (Ligyrodes) peruvianus, D) L. (Anagrylius) moroni, E) L. (Ligyrus) ruginasus, F) L. (Ligyrus) nasutus, G) L. (Ligyrus) bidentulus, H) Tomarus adoceteus, I) T. maimon, J) T. pullus, K) T. pumilus." figureDoi="http://doi.org/10.5281/zenodo.7345011" httpUri="https://zenodo.org/record/7345011/files/figure.png" pageId="45" pageNumber="46">Fig. 1F</figureCitation>
|
||
). Ocular canthus acute, with 10 ventral setae. Clypeal surface rugopunctate to transversely rugose, striae denser than those on frons. Clypeus triangular, base 3.5 times wider than apex (
|
||
<figureCitation id="B29BE4A8FD72FFACADB41A5EFD65BF42" box="[645,731,776,803]" captionStart="FIGURE 1" captionStartId="81.[151,250,1535,1559]" captionTargetBox="[151,1436,181,1511]" captionTargetId="figure-17@81.[151,1436,181,1511]" captionTargetPageId="81" captionText="FIGURE 1. Head. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) Ligyrus (Ligyrodes) peruvianus, D) L. (Anagrylius) moroni, E) L. (Ligyrus) ruginasus, F) L. (Ligyrus) nasutus, G) L. (Ligyrus) bidentulus, H) Tomarus adoceteus, I) T. maimon, J) T. pullus, K) T. pumilus." figureDoi="http://doi.org/10.5281/zenodo.7345011" httpUri="https://zenodo.org/record/7345011/files/figure.png" pageId="45" pageNumber="46">Fig. 1F</figureCitation>
|
||
). Clypeal teeth triangular, separated by 1 diameter tooth (
|
||
<figureCitation id="B29BE4A8FD72FFACAA5E1A5EFF09BF26" captionStart="FIGURE 1" captionStartId="81.[151,250,1535,1559]" captionTargetBox="[151,1436,181,1511]" captionTargetId="figure-17@81.[151,1436,181,1511]" captionTargetPageId="81" captionText="FIGURE 1. Head. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) Ligyrus (Ligyrodes) peruvianus, D) L. (Anagrylius) moroni, E) L. (Ligyrus) ruginasus, F) L. (Ligyrus) nasutus, G) L. (Ligyrus) bidentulus, H) Tomarus adoceteus, I) T. maimon, J) T. pullus, K) T. pumilus." figureDoi="http://doi.org/10.5281/zenodo.7345011" httpUri="https://zenodo.org/record/7345011/files/figure.png" pageId="45" pageNumber="46">Fig. 1F</figureCitation>
|
||
). Mandible with 2 apical and 1 lateral, widely rounded tooth, tooth 1 longer and narrower than tooth 2. Maxilla rectangular; galea with 3 dorsal teeth. Apex of labrum straight. Interocular distance 3.5 times an eye width. Antennal club very long.
|
||
<emphasis id="18D4243FFD72FFACAE7B1A23FE72BFEE" bold="true" box="[330,460,885,911]" pageId="45" pageNumber="46">Pronotum:</emphasis>
|
||
Surface with large, dense punctures. Apex without tubercle (with a small swelling) or fovea.
|
||
<emphasis id="18D4243FFD72FFACAFD41ACFFE84BFD2" bold="true" box="[229,314,921,947]" pageId="45" pageNumber="46">Elytra:</emphasis>
|
||
First interval punctate, punctures similar to other intervals. Inner surface of apex with small transverse tubercles forming about 70 parallel lines.
|
||
<emphasis id="18D4243FFD72FFACAD9A1AEAFC97BFB7" bold="true" box="[683,809,956,982]" pageId="45" pageNumber="46">Abdomen:</emphasis>
|
||
Apex of tergite IV with a diagonal line of 11 transverse, parallel, short carinae. Pygidial surface with small punctures, denser on base and anterior corners. Surface in lateral view strongly convex (male) to flat (female).
|
||
<emphasis id="18D4243FFD72FFACAD9D1D53FD53B87E" bold="true" box="[684,749,1029,1055]" pageId="45" pageNumber="46">Legs:</emphasis>
|
||
Protibia tridentate, basal tooth slightly distant from others. Protibial surface sparsely punctate. Protarsus of male simple, inner claw simple. Metatibia strongly contracted at apex. Apex of metatibia entire, with 15–20 spinules.
|
||
<emphasis id="18D4243FFD72FFACADC61D1BFC79B807" bold="true" box="[759,967,1100,1127]" pageId="45" pageNumber="46">Female genitalia:</emphasis>
|
||
Subcoxite subrectangular (2 times wider than long). Coxite subrectangular (wider than long); surface strongly concave. Subcoxite wider and shorter than coxite.
|
||
<emphasis id="18D4243FFD72FFACAFDA1DC3FE21B8CF" bold="true" box="[235,415,1172,1199]" pageId="45" pageNumber="46">Male genitalia:</emphasis>
|
||
Spiculum gastrale thin, apex truncate. Parameres with sides nearly parallel, lateral ventral teeth long (
|
||
<figureCitation id="B29BE4A8FD72FFACAE281DEEFE33B8B2" box="[281,397,1208,1235]" captionStart="FIGURE 15" captionStartId="90.[151,250,1535,1559]" captionTargetBox="[154,1430,183,1509]" captionTargetId="figure-24@90.[151,1436,181,1512]" captionTargetPageId="90" captionText="FIGURE 15. Parameres in frontal view. A) Euligyrus ebenus, B) E. similis, C) Proculigyrus cicatricosus, D) Ligyrus (Anagrylius) cuniculus, E) L. (Anagrylius) moroni, F) L. (Ligyrodes) peruvianus, G) L. (Ligyrodes) relictus, H) L. (Ligyrodes) sallaei, I) L. (Ligyrus) allonasutus, J) L. (Ligyrus) bidentulus, K) L. (Ligyrus) burmeisteri, L) Ligyrus (Ligyrus) fossor, M) L. (Ligyrus) gibbosus, N) L. (Ligyrus) nasutus, O) L. (Ligyrus) neglectus, P) L. (Ligyrus) paranaensis, Q) L. (Ligyrus) peninsularis, R) L. (Ligyrus) rubripes, S) L. (Ligyrus) ruginasus, T) L. (Ligyrus) villosus." figureDoi="http://doi.org/10.5281/zenodo.7345063" httpUri="https://zenodo.org/record/7345063/files/figure.png" pageId="45" pageNumber="46">Figs. 15N</figureCitation>
|
||
,
|
||
<figureCitation id="B29BE4A8FD72FFACAEAB1DEFFE77B8B2" box="[410,457,1209,1235]" captionStart="FIGURE 18" captionStartId="93.[151,250,1597,1621]" captionTargetBox="[152,1436,184,1567]" captionTargetId="figure-17@93.[151,1436,181,1573]" captionTargetPageId="93" captionText="FIGURE 18. Parameres in lateral view.A) L. (Ligyrus.) allonasutus, B) L. (Ligyrus) bidentulus, C) L. (Ligyrus) burmeisteri, D) Ligyrus (Ligyrus) fossor, E) L. (Ligyrus) gibbosus, F) L. (Ligyrus) nasutus, G) L. (Ligyrus) neglectus, H) L. (Ligyrus) paranaensis, I) L. (Ligyrus) peninsularis, J) L. (Ligyrus) rubripes, K) L. (Ligyrus) ruginasus, L) L. (Ligyrus) villosus." figureDoi="http://doi.org/10.5281/zenodo.7345077" httpUri="https://zenodo.org/record/7345077/files/figure.png" pageId="45" pageNumber="46">18F</figureCitation>
|
||
). Internal sac with lamellar spiny belt long (5 times longer than copulatory lamellae); accessory lamella thickened, irregular.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="62BAABA6FD72FFAFAFF61C56FE4ABCB4" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46" type="diagnosis">
|
||
<paragraph id="2A1FF82DFD72FFACAFF61C56FEE3B9CA" blockId="45.[151,1437,632,2027]" pageId="45" pageNumber="46">
|
||
<emphasis id="18D4243FFD72FFACAFF61C56FEFCB97B" bold="true" box="[199,322,1280,1306]" pageId="45" pageNumber="46">Diagnosis.</emphasis>
|
||
<taxonomicName id="EDA083AEFD72FFACAE761C57FE47B97B" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[327,505,1281,1306]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="nasutus">
|
||
<emphasis id="18D4243FFD72FFACAE761C57FE47B97B" box="[327,505,1281,1306]" italics="true" pageId="45" pageNumber="46">Ligyrus nasutus</emphasis>
|
||
</taxonomicName>
|
||
can be recognized by the frontal carina bilobed, not interrupted (
|
||
<figureCitation id="B29BE4A8FD72FFACABF61C56FAA4B97A" box="[1223,1306,1280,1307]" captionStart="FIGURE 1" captionStartId="81.[151,250,1535,1559]" captionTargetBox="[151,1436,181,1511]" captionTargetId="figure-17@81.[151,1436,181,1511]" captionTargetPageId="81" captionText="FIGURE 1. Head. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) Ligyrus (Ligyrodes) peruvianus, D) L. (Anagrylius) moroni, E) L. (Ligyrus) ruginasus, F) L. (Ligyrus) nasutus, G) L. (Ligyrus) bidentulus, H) Tomarus adoceteus, I) T. maimon, J) T. pullus, K) T. pumilus." figureDoi="http://doi.org/10.5281/zenodo.7345011" httpUri="https://zenodo.org/record/7345011/files/figure.png" pageId="45" pageNumber="46">Fig. 1F</figureCitation>
|
||
); pronotum without tubercle or fovea, with a basal small swelling (
|
||
<figureCitation id="B29BE4A8FD72FFACAC251C72FCC0B95F" box="[788,894,1316,1342]" captionStart="FIGURE 26" captionStartId="101.[151,250,1827,1851]" captionTargetBox="[159,1430,189,1791]" captionTargetId="figure-17@101.[151,1436,181,1803]" captionTargetPageId="101" captionText="FIGURE 26. Habitus of the species of Ligyrus. Subgenus Ligyrus: A) Ligyrus fossor, B) L. gibbosus, C) L. nasutus, D) L. neglectus, E) L. paranaensis, F) L. peninsularis, G) L. rubripes, H) L. ruginasus, I) L. villosus. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7345101" httpUri="https://zenodo.org/record/7345101/files/figure.png" pageId="45" pageNumber="46">Fig. 26C</figureCitation>
|
||
); pronotal punctation deep and large; metatibia elongate with apex strongly narrowed; parameres with sides nearly parallel; lateral ventral teeth long (
|
||
<figureCitation id="B29BE4A8FD72FFACAA151C1EFA29B902" box="[1316,1431,1352,1379]" captionStart="FIGURE 15" captionStartId="90.[151,250,1535,1559]" captionTargetBox="[154,1430,183,1509]" captionTargetId="figure-24@90.[151,1436,181,1512]" captionTargetPageId="90" captionText="FIGURE 15. Parameres in frontal view. A) Euligyrus ebenus, B) E. similis, C) Proculigyrus cicatricosus, D) Ligyrus (Anagrylius) cuniculus, E) L. (Anagrylius) moroni, F) L. (Ligyrodes) peruvianus, G) L. (Ligyrodes) relictus, H) L. (Ligyrodes) sallaei, I) L. (Ligyrus) allonasutus, J) L. (Ligyrus) bidentulus, K) L. (Ligyrus) burmeisteri, L) Ligyrus (Ligyrus) fossor, M) L. (Ligyrus) gibbosus, N) L. (Ligyrus) nasutus, O) L. (Ligyrus) neglectus, P) L. (Ligyrus) paranaensis, Q) L. (Ligyrus) peninsularis, R) L. (Ligyrus) rubripes, S) L. (Ligyrus) ruginasus, T) L. (Ligyrus) villosus." figureDoi="http://doi.org/10.5281/zenodo.7345063" httpUri="https://zenodo.org/record/7345063/files/figure.png" pageId="45" pageNumber="46">Figs. 15N</figureCitation>
|
||
,
|
||
<figureCitation id="B29BE4A8FD72FFACAFA61C3BFF78B9E6" box="[151,198,1389,1415]" captionStart="FIGURE 18" captionStartId="93.[151,250,1597,1621]" captionTargetBox="[152,1436,184,1567]" captionTargetId="figure-17@93.[151,1436,181,1573]" captionTargetPageId="93" captionText="FIGURE 18. Parameres in lateral view.A) L. (Ligyrus.) allonasutus, B) L. (Ligyrus) bidentulus, C) L. (Ligyrus) burmeisteri, D) Ligyrus (Ligyrus) fossor, E) L. (Ligyrus) gibbosus, F) L. (Ligyrus) nasutus, G) L. (Ligyrus) neglectus, H) L. (Ligyrus) paranaensis, I) L. (Ligyrus) peninsularis, J) L. (Ligyrus) rubripes, K) L. (Ligyrus) ruginasus, L) L. (Ligyrus) villosus." figureDoi="http://doi.org/10.5281/zenodo.7345077" httpUri="https://zenodo.org/record/7345077/files/figure.png" pageId="45" pageNumber="46">18F</figureCitation>
|
||
); internal sac with accessory lamella thick and irregular (as in
|
||
<taxonomicName id="EDA083AEFD72FFACAC9A1C3BFBF3B9E7" authorityName="Lopez-Garcia & Deloya" authorityYear="2022" box="[939,1101,1388,1414]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="allonasutus">
|
||
<emphasis id="18D4243FFD72FFACAC9A1C3BFBF3B9E7" box="[939,1101,1388,1414]" italics="true" pageId="45" pageNumber="46">L. allonasutus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="B29BE4A8FD72FFACAB6E1C3AFB79B9E6" box="[1119,1223,1388,1415]" captionStart="FIGURE 21" captionStartId="96.[151,250,1739,1763]" captionTargetBox="[161,1418,194,1706]" captionTargetId="figure-24@96.[151,1435,181,1715]" captionTargetPageId="96" captionText="FIGURE 21. Internal sac. A) Euligyrus ebenus, B) Proculigyrus cicatricosus, C) Ligyrus (Anagrylius) moroni, D) L. (Ligyrodes) sallaei, E) L. (Ligyrus) allonasutus. Abbreviations:AL:Accessory lamellae, Br: Brush of setae, CL: Copulatory lamella, LSB: Lamellar spiny belt, Ra: Raspula." figureDoi="http://doi.org/10.5281/zenodo.7345085" httpUri="https://zenodo.org/record/7345085/files/figure.png" pageId="45" pageNumber="46">Fig. 21E</figureCitation>
|
||
); coxite of female strongly concave.
|
||
</paragraph>
|
||
<paragraph id="2A1FF82DFD72FFAFAFF61CE2FE4ABCB4" blockId="45.[151,1437,632,2027]" lastBlockId="46.[151,1437,151,465]" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46">
|
||
<emphasis id="18D4243FFD72FFACAFF61CE2FE00B9AE" bold="true" box="[199,446,1460,1487]" pageId="45" pageNumber="46">Taxonomic remarks.</emphasis>
|
||
<taxonomicName id="EDA083AEFD72FFACAEF71CE3FDC7B9AF" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[454,633,1461,1486]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="nasutus">
|
||
<emphasis id="18D4243FFD72FFACAEF71CE3FDC7B9AF" box="[454,633,1461,1486]" italics="true" pageId="45" pageNumber="46">Ligyrus nasutus</emphasis>
|
||
</taxonomicName>
|
||
was described with
|
||
<specimenCount id="3CA633A4FD72FFACAC531CE3FC50B9AF" box="[866,1006,1460,1487]" pageId="45" pageNumber="46" type="female">two females</specimenCount>
|
||
, one from Yucatán and the other from
|
||
<collectingCountry id="52B7B8BDFD72FFACAFA61C8EFEAAB993" box="[151,276,1496,1522]" name="Guatemala" pageId="45" pageNumber="46">Guatemala</collectingCountry>
|
||
.
|
||
<taxonomicName id="EDA083AEFD72FFACAE111C8FFE53B993" authorityName="Bates" authorityYear="1888" box="[288,493,1496,1522]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="pygidialis">
|
||
<emphasis id="18D4243FFD72FFACAE111C8FFE53B993" box="[288,493,1496,1522]" italics="true" pageId="45" pageNumber="46">Ligyrus pygidialis</emphasis>
|
||
</taxonomicName>
|
||
was described by
|
||
<bibRefCitation id="4E3185DCFD72FFACADF51C8FFCEBB992" author="Bates, H. W." box="[708,853,1497,1523]" pageId="45" pageNumber="46" pagination="296 - 342" refId="ref88329" refString="Bates, H. W. (1888) Pectinicornia and Lamellicornia, Family Dynastinae. In: Godman, F. D. & Salvin, O. (Eds.), Biologia Centrali-Americana, Insecta, Coleoptera, 2 (Part 2), pp. 296 - 342." type="journal article" year="1888">Bates (1888)</bibRefCitation>
|
||
based on a large series of males and females from
|
||
<collectingRegion id="E86436CFFD72FFACAFA61CABFF4FBA76" box="[151,241,1533,1559]" country="Mexico" name="Yucatan" pageId="45" pageNumber="46">Yucatán</collectingRegion>
|
||
(
|
||
<collectingCountry id="52B7B8BDFD72FFACAE311CAAFEE5BA77" box="[256,347,1532,1558]" name="Mexico" pageId="45" pageNumber="46">Mexico</collectingCountry>
|
||
), which according to his description were different to
|
||
<taxonomicName id="EDA083AEFD72FFACAC841CABFB96BA77" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[949,1064,1533,1558]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="nasutus">
|
||
<emphasis id="18D4243FFD72FFACAC841CABFB96BA77" box="[949,1064,1533,1558]" italics="true" pageId="45" pageNumber="46">L. nasutus</emphasis>
|
||
</taxonomicName>
|
||
by the shape of the frontoclypeal carina divided at middle. It is not clear if
|
||
<bibRefCitation id="4E3185DCFD72FFACAD541F77FD4BBA5A" author="Bates, H. W." box="[613,757,1569,1595]" pageId="45" pageNumber="46" pagination="296 - 342" refId="ref88329" refString="Bates, H. W. (1888) Pectinicornia and Lamellicornia, Family Dynastinae. In: Godman, F. D. & Salvin, O. (Eds.), Biologia Centrali-Americana, Insecta, Coleoptera, 2 (Part 2), pp. 296 - 342." type="journal article" year="1888">Bates (1888)</bibRefCitation>
|
||
reviewed the type specimen of
|
||
<taxonomicName id="EDA083AEFD72FFACAB691F77FB72BA5B" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[1112,1228,1569,1594]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="nasutus">
|
||
<emphasis id="18D4243FFD72FFACAB691F77FB72BA5B" box="[1112,1228,1569,1594]" italics="true" pageId="45" pageNumber="46">L. nasutus</emphasis>
|
||
</taxonomicName>
|
||
of
|
||
<collectingRegion id="E86436CFFD72FFACABDE1F77FAF2BA5A" box="[1263,1356,1569,1595]" country="Mexico" name="Yucatan" pageId="45" pageNumber="46">Yucatán</collectingRegion>
|
||
, but he cited additional specimens from
|
||
<collectingRegion id="E86436CFFD72FFACAD381F13FDD0BA3E" box="[521,622,1605,1631]" country="Mexico" name="Guerrero" pageId="45" pageNumber="46">Guerrero</collectingRegion>
|
||
(
|
||
<collectingCountry id="52B7B8BDFD72FFACAD4F1F12FD69BA3F" box="[638,727,1604,1630]" name="Mexico" pageId="45" pageNumber="46">Mexico</collectingCountry>
|
||
) and San Jerónimo (
|
||
<collectingCountry id="52B7B8BDFD72FFACACF31F12FBFCBA3F" box="[962,1090,1604,1630]" name="Guatemala" pageId="45" pageNumber="46">Guatemala</collectingCountry>
|
||
).
|
||
<bibRefCitation id="4E3185DCFD72FFACAB641F12FABCBA3F" author="Endrodi, S." box="[1109,1282,1604,1630]" pageId="45" pageNumber="46" pagination="1 - 145" refId="ref89768" refString="Endrodi, S. (1969) Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). Entomologische Abhandlungen, 37, 1 - 145." type="journal article" year="1969">Endrödi (1969)</bibRefCitation>
|
||
chose the female from
|
||
<collectingRegion id="E86436CFFD72FFACAE3F1F3FFED6BAE2" box="[270,360,1641,1667]" country="Mexico" name="Yucatan" pageId="45" pageNumber="46">Yucatán</collectingRegion>
|
||
(in poor condition) as the
|
||
<typeStatus id="F51B468FFD72FFACADB91F3EFD51BAE3" box="[648,751,1640,1666]" pageId="45" pageNumber="46" type="lectotype">lectotype</typeStatus>
|
||
for
|
||
<taxonomicName id="EDA083AEFD72FFACAC2B1F3FFC32BAE3" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[794,908,1641,1666]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="nasutus">
|
||
<emphasis id="18D4243FFD72FFACAC2B1F3FFC32BAE3" box="[794,908,1641,1666]" italics="true" pageId="45" pageNumber="46">L. nasutus</emphasis>
|
||
</taxonomicName>
|
||
and considered
|
||
<taxonomicName id="EDA083AEFD72FFACAB0F1F3FFB76BAE3" authorityName="Bates" authorityYear="1888" box="[1086,1224,1640,1666]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="pygidialis">
|
||
<emphasis id="18D4243FFD72FFACAB0F1F3FFB76BAE3" box="[1086,1224,1640,1666]" italics="true" pageId="45" pageNumber="46">L. pygidialis</emphasis>
|
||
</taxonomicName>
|
||
as an “aberration”. He only described and illustrated the parameres from the populations from western
|
||
<collectingCountry id="52B7B8BDFD72FFACAB071FDAFB32BAC7" box="[1078,1164,1676,1702]" name="Mexico" pageId="45" pageNumber="46">Mexico</collectingCountry>
|
||
and
|
||
<collectingCountry id="52B7B8BDFD72FFACABF01FDAFA84BAC7" box="[1217,1338,1676,1702]" name="Guatemala" pageId="45" pageNumber="46">Guatemala</collectingCountry>
|
||
to
|
||
<collectingCountry id="52B7B8BDFD72FFACAA6D1FDBFF74BAAB" name="Costa Rica" pageId="45" pageNumber="46">Costa Rica</collectingCountry>
|
||
but did not illustrate the parameres of the
|
||
<collectingRegion id="E86436CFFD72FFACAD831FE7FCB2BAAA" box="[690,780,1713,1739]" country="Mexico" name="Yucatan" pageId="45" pageNumber="46">Yucatán</collectingRegion>
|
||
populations, even though the
|
||
<typeStatus id="F51B468FFD72FFACAB541FE6FB72BAAB" box="[1125,1228,1712,1738]" pageId="45" pageNumber="46" type="lectotype">lectotype</typeStatus>
|
||
of
|
||
<taxonomicName id="EDA083AEFD72FFACABC51FE7FAD7BAAB" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[1268,1385,1713,1738]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="nasutus">
|
||
<emphasis id="18D4243FFD72FFACABC51FE7FAD7BAAB" box="[1268,1385,1713,1738]" italics="true" pageId="45" pageNumber="46">L. nasutus</emphasis>
|
||
</taxonomicName>
|
||
was from this region. After his work, the name
|
||
<taxonomicName id="EDA083AEFD72FFACAD4F1F83FD4CBA8F" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[638,754,1749,1774]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="nasutus">
|
||
<emphasis id="18D4243FFD72FFACAD4F1F83FD4CBA8F" box="[638,754,1749,1774]" italics="true" pageId="45" pageNumber="46">L. nasutus</emphasis>
|
||
</taxonomicName>
|
||
was widely used for supposedly a unique species with wide distribution, but there was confusion since the original description of Burmeister included
|
||
<specimenCount id="3CA633A4FD72FFACAB911FAFFAA1BB73" box="[1184,1311,1784,1811]" pageId="45" pageNumber="46" type="female">two female</specimenCount>
|
||
specimens from different species as types for
|
||
<taxonomicName id="EDA083AEFD72FFACAD161E4BFD22BB57" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[551,668,1821,1846]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="nasutus">
|
||
<emphasis id="18D4243FFD72FFACAD161E4BFD22BB57" box="[551,668,1821,1846]" italics="true" pageId="45" pageNumber="46">L. nasutus</emphasis>
|
||
</taxonomicName>
|
||
. In the present work, the
|
||
<typeStatus id="F51B468FFD72FFACACF41E4AFB89BB57" box="[965,1079,1820,1846]" pageId="45" pageNumber="46" type="lectotype">lectotypes</typeStatus>
|
||
of
|
||
<taxonomicName id="EDA083AEFD72FFACAB6E1E4BFB6BBB57" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[1119,1237,1821,1846]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="nasutus">
|
||
<emphasis id="18D4243FFD72FFACAB6E1E4BFB6BBB57" box="[1119,1237,1821,1846]" italics="true" pageId="45" pageNumber="46">L. nasutus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="EDA083AEFD72FFACAA3F1E4BFA22BB57" authorityName="Bates" authorityYear="1888" box="[1294,1436,1820,1846]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="pygidialis">
|
||
<emphasis id="18D4243FFD72FFACAA3F1E4BFA22BB57" box="[1294,1436,1820,1846]" italics="true" pageId="45" pageNumber="46">L. pygidialis</emphasis>
|
||
</taxonomicName>
|
||
were analyzed in comparison with additional specimens from the region of
|
||
<collectingRegion id="E86436CFFD72FFACACD61E17FBFFBB3A" box="[999,1089,1857,1883]" country="Mexico" name="Yucatan" pageId="45" pageNumber="46">Yucatán</collectingRegion>
|
||
and compared with specimens from other regions of
|
||
<collectingCountry id="52B7B8BDFD72FFACAEBB1E32FE5EBB1F" box="[394,480,1892,1918]" name="Mexico" pageId="45" pageNumber="46">Mexico</collectingCountry>
|
||
and
|
||
<collectingCountry id="52B7B8BDFD72FFACAD251E32FD33BB1F" box="[532,653,1892,1918]" name="Guatemala" pageId="45" pageNumber="46">Guatemala</collectingCountry>
|
||
to
|
||
<collectingCountry id="52B7B8BDFD72FFACAD811E33FCB2BB1E" box="[688,780,1893,1919]" name="Panama" pageId="45" pageNumber="46">Panama</collectingCountry>
|
||
. Clear differences were found between the specimens from
|
||
<collectingRegion id="E86436CFFD72FFACAFA61EDFFE81BBC3" box="[151,319,1928,1955]" country="Mexico" name="Yucatan" pageId="45" pageNumber="46">Yucatán region</collectingRegion>
|
||
and the other populations, mainly in the shape of the frontoclypeal carina, pronotum, and parameres, which is supported by the biogeographic identity of the region of
|
||
<collectingRegion id="E86436CFFD72FFACAC5E1EFBFC73BBA6" box="[879,973,1965,1991]" country="Mexico" name="Yucatan" pageId="45" pageNumber="46">Yucatán</collectingRegion>
|
||
. Following the
|
||
<bibRefCitation id="4E3185DCFD72FFACAB481EFAFD8DBB8A" author="International Commission on Zoological Nomenclature" pageId="45" pageNumber="46" refId="ref90721" refString="International Commission on Zoological Nomenclature (1999) International Code of Zoological Nomenclature. 4 th Edition. International Trust for Zoological Nomenclature, London, 306 pp." type="book" year="1999">International Commission on Zoological Nomenclature (1999)</bibRefCitation>
|
||
, as the
|
||
<typeStatus id="F51B468FFD72FFACADBC1E86FD4ABB8B" box="[653,756,2000,2026]" pageId="45" pageNumber="46" type="lectotype">lectotype</typeStatus>
|
||
of
|
||
<taxonomicName id="EDA083AEFD72FFACAC2F1E87FC2DBB8B" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[798,915,2001,2026]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="nasutus">
|
||
<emphasis id="18D4243FFD72FFACAC2F1E87FC2DBB8B" box="[798,915,2001,2026]" italics="true" pageId="45" pageNumber="46">L. nasutus</emphasis>
|
||
</taxonomicName>
|
||
is from
|
||
<collectingRegion id="E86436CFFD72FFACACC61E87FBEABB8A" box="[1015,1108,2001,2027]" country="Mexico" name="Yucatan" pageId="45" pageNumber="46">Yucatán</collectingRegion>
|
||
, only the populations of this region correspond to this species, and the remaining populations from western
|
||
<collectingCountry id="52B7B8BDFD71FFAFAB3B19C1FBDEBCD0" box="[1034,1120,151,177]" name="Mexico" pageId="46" pageNumber="47">Mexico</collectingCountry>
|
||
to
|
||
<collectingCountry id="52B7B8BDFD71FFAFABB219C1FB62BCD0" box="[1155,1244,151,177]" name="Panama" pageId="46" pageNumber="47">Panama</collectingCountry>
|
||
are named in the present work as
|
||
<taxonomicName id="EDA083AEFD71FFAFAE7F19EDFE52BCB4" authorityName="Lopez-Garcia & Deloya" authorityYear="2022" box="[334,492,187,213]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="allonasutus">
|
||
<emphasis id="18D4243FFD71FFAFAE7F19EDFE52BCB4" box="[334,492,187,213]" italics="true" pageId="46" pageNumber="47">L. allonasutus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="62BAABA6FD71FFAFAFF61988FF55BDB0" pageId="46" pageNumber="47" type="distribution">
|
||
<paragraph id="2A1FF82DFD71FFAFAFF61988FCF6BD7C" blockId="46.[151,1437,151,465]" pageId="46" pageNumber="47">
|
||
<emphasis id="18D4243FFD71FFAFAFF61988FEDFBC99" bold="true" box="[199,353,222,248]" pageId="46" pageNumber="47">Distribution.</emphasis>
|
||
<taxonomicName id="EDA083AEFD71FFAFAE571989FDA6BC99" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[358,536,223,248]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="nasutus">
|
||
<emphasis id="18D4243FFD71FFAFAE571989FDA6BC99" box="[358,536,223,248]" italics="true" pageId="46" pageNumber="47">Ligyrus nasutus</emphasis>
|
||
</taxonomicName>
|
||
is restricted to the
|
||
<collectingRegion id="E86436CFFD71FFAFADD61989FC31BC98" box="[743,911,223,249]" country="Mexico" name="Yucatan" pageId="46" pageNumber="47">Yucatán region</collectingRegion>
|
||
of
|
||
<collectingCountry id="52B7B8BDFD71FFAFAC801989FBB7BC98" box="[945,1033,223,249]" name="Mexico" pageId="46" pageNumber="47">Mexico</collectingCountry>
|
||
, in the states of
|
||
<collectingRegion id="E86436CFFD71FFAFAB881989FA8CBC98" box="[1209,1330,223,249]" country="Mexico" name="Campeche" pageId="46" pageNumber="47">Campeche</collectingRegion>
|
||
,
|
||
<collectingRegion id="E86436CFFD71FFAFAA0A1989FA26BC98" box="[1339,1432,223,249]" country="Mexico" name="Yucatan" pageId="46" pageNumber="47">Yucatán</collectingRegion>
|
||
,
|
||
<collectingRegion id="E86436CFFD71FFAFAFA61855FE88BD7C" box="[151,310,259,285]" country="Mexico" name="Quintana Roo" pageId="46" pageNumber="47">Quintana Roo</collectingRegion>
|
||
, and eastern
|
||
<collectingRegion id="E86436CFFD71FFAFAEF61855FD9FBD7C" box="[455,545,259,285]" country="Mexico" name="Chiapas" pageId="46" pageNumber="47">Chiapas</collectingRegion>
|
||
at elevations below
|
||
<specimenCount id="3CA633A4FD71FFAFAC391854FCF6BD7C" box="[776,840,258,285]" pageId="46" pageNumber="47" type="generic">
|
||
<quantity id="ED5855C8FD71FFAFAC391854FCFABD7C" box="[776,836,258,285]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="46" pageNumber="47" unit="m" value="50.0">50 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
</paragraph>
|
||
<paragraph id="2A1FF82DFD71FFAFAFF61870FF55BDB0" blockId="46.[151,1437,151,465]" pageId="46" pageNumber="47">
|
||
<emphasis id="18D4243FFD71FFAFAFF61870FDB8BD21" bold="true" box="[199,518,294,321]" pageId="46" pageNumber="47">
|
||
Locality records (
|
||
<figureCitation id="B29BE4A8FD71FFAFAE911870FE47BD21" box="[416,505,294,320]" captionStart="FIGURE 34" captionStartId="107.[151,250,2007,2031]" captionTargetBox="[246,1346,1181,1977]" captionTargetId="figure-32@107.[242,1346,1181,1983]" captionTargetPageId="107" captionText="FIGURE 34. Distributional records of Ligyrus (Ligyrus) allonasutus and L. nasutus." figureDoi="http://doi.org/10.5281/zenodo.7345125" httpUri="https://zenodo.org/record/7345125/files/figure.png" pageId="46" pageNumber="47">Fig. 34</figureCitation>
|
||
).
|
||
</emphasis>
|
||
62 examined specimens from BMNH, CNIN, EMAC, IEXA, UNSM, USNM. Some data from
|
||
<bibRefCitation id="4E3185DCFD71FFAFAE7E181DFDFCBD04" author="Ratcliffe, B. C. & Cave, R. D. & Cano, E. B." box="[335,578,331,357]" pageId="46" pageNumber="47" pagination="1 - 666" refId="ref93157" refString="Ratcliffe, B. C., Cave, R. D. & Cano, E. B. (2013) The dynastine scarab beetles of Mexico, Guatemala, and Belize (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum, 27, 1 - 666." type="journal article" year="2013">
|
||
Ratcliffe
|
||
<emphasis id="18D4243FFD71FFAFAE86181DFE51BD04" box="[439,495,331,357]" italics="true" pageId="46" pageNumber="47">et al.</emphasis>
|
||
(2013)
|
||
</bibRefCitation>
|
||
.
|
||
<emphasis id="18D4243FFD71FFAFAD7C181CFC7CBD05" bold="true" box="[589,962,330,356]" pageId="46" pageNumber="47">
|
||
<collectingCountry id="52B7B8BDFD71FFAFAD7C181CFD7DBD05" box="[589,707,330,356]" name="Mexico" pageId="46" pageNumber="47">MEXICO</collectingCountry>
|
||
(81).
|
||
<collectingRegion id="E86436CFFD71FFAFAC34181CFC3DBD05" box="[773,899,330,356]" country="Mexico" name="Campeche" pageId="46" pageNumber="47">Campeche</collectingRegion>
|
||
(16):
|
||
</emphasis>
|
||
Calkini; Campeche (
|
||
<quantity id="ED5855C8FD71FFAFAB9F181CFB56BD04" box="[1198,1256,330,357]" metricMagnitude="3" metricUnit="m" metricValue="8.04672" pageId="46" pageNumber="47" unit="mi" value="5.0">5 mi.</quantity>
|
||
N); Grutas Xtacambilxunán.
|
||
<emphasis id="18D4243FFD71FFAFAE061838FE72BDE9" bold="true" box="[311,460,366,392]" pageId="46" pageNumber="47">
|
||
<collectingRegion id="E86436CFFD71FFAFAE061838FE27BDE9" box="[311,409,366,392]" country="Mexico" name="Chiapas" pageId="46" pageNumber="47">Chiapas</collectingRegion>
|
||
(1):
|
||
</emphasis>
|
||
Palenque.
|
||
<emphasis id="18D4243FFD71FFAFAD7B1838FC8BBDE9" bold="true" box="[586,821,366,393]" pageId="46" pageNumber="47">
|
||
<collectingRegion id="E86436CFFD71FFAFAD7B1838FD4ABDE8" box="[586,756,366,393]" country="Mexico" name="Quintana Roo" pageId="46" pageNumber="47">Quintana Roo</collectingRegion>
|
||
(12):
|
||
</emphasis>
|
||
Cancún; Carrillo Puerto (
|
||
<quantity id="ED5855C8FD71FFAFAB6D1839FB6EBDE8" box="[1116,1232,367,393]" metricMagnitude="4" metricUnit="m" metricValue="2.2" metricValueMax="2.4" metricValueMin="2.0" pageId="46" pageNumber="47" unit="km" value="22.0" valueMax="24.0" valueMin="20.0">20–24 km</quantity>
|
||
N); Cobá; Nuevo X-Can; Playa del Carmen (
|
||
<quantity id="ED5855C8FD71FFAFAEF018C5FDA3BDCC" box="[449,541,403,429]" metricMagnitude="4" metricUnit="m" metricValue="1.09" pageId="46" pageNumber="47" unit="km" value="10.9">10.9 km</quantity>
|
||
S).
|
||
<emphasis id="18D4243FFD71FFAFAD7818C4FD57BDCD" bold="true" box="[585,745,402,429]" pageId="46" pageNumber="47">
|
||
<collectingRegion id="E86436CFFD71FFAFAD7818C4FD12BDCC" box="[585,684,402,429]" country="Mexico" name="Yucatan" pageId="46" pageNumber="47">Yucatán</collectingRegion>
|
||
(51):
|
||
</emphasis>
|
||
Chichén Itzá; Conkal; Mérida; Piste (
|
||
<quantity id="ED5855C8FD71FFAFABBB18C5FB6FBDCC" box="[1162,1233,403,429]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="46" pageNumber="47" unit="km" value="12.0">12 km</quantity>
|
||
N); Santa Helena; Temax.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |