240 lines
22 KiB
XML
240 lines
22 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1117.86984" ID-Pensoft-Pub="1313-2970-1117-95" ID-Pensoft-UUID="54DB7D7318935384856B59FD7D22DFF4" ID-ZooBank="C583CC7AB2AD42048FA083C49BB088EA" ModsDocID="1313-2970-1117-95" checkinTime="1660292277777" checkinUser="pensoft" docAuthor="Holzenthal, Ralph W., Blahnik, Roger J. & Rios-Touma, Blanca" docDate="2022" docId="69786D3A558B5E689389141F23A11488" docLanguage="en" docName="ZooKeys 1117: 95-122" docOrigin="ZooKeys 1117" docPubDate="2022-08-11" docSource="http://dx.doi.org/10.3897/zookeys.1117.86984" docTitle="Chimarra (Chimarra) asterae Holzenthal & Blahnik & Ríos-Touma 2022, sp. nov." docType="treatment" docUuid="F1432B21-5E24-4894-93D6-6F4546BB04BB" docUuidSource="ZooBank" docVersion="1" id="54DB7D7318935384856B59FD7D22DFF4" lastPageNumber="95" masterDocId="54DB7D7318935384856B59FD7D22DFF4" masterDocTitle="A new genus and new species of Ecuadorian Philopotamidae (Trichoptera)" masterLastPageNumber="122" masterPageNumber="95" pageNumber="95" updateTime="1660292277777" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A new genus and new species of Ecuadorian Philopotamidae (Trichoptera)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Holzenthal, Ralph W.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1853-6340</mods:nameIdentifier>
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<mods:affiliation>Department of Entomology, University of Minnesota, 1980 Folwell Avenue, 219 Hodson Hall, St. Paul, Minnesota 55108 USA</mods:affiliation>
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<mods:nameIdentifier type="email">holze001@umn.edu</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Blahnik, Roger J.</mods:namePart>
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<mods:affiliation>Department of Entomology, University of Minnesota, 1980 Folwell Avenue, 219 Hodson Hall, St. Paul, Minnesota 55108 USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Rios-Touma, Blanca</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3921-0908</mods:nameIdentifier>
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<mods:affiliation>Grupo de Investigacion en Biodiversidad, Medio Ambiente y Salud (BIOMAS), Facultad de Ingenierias y Ciencias Aplicadas, Via Nayon S / N, Campus UDLAPARK, CP 170503, Universidad de Las Americas, Quito, Pichincha, Ecuador</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-08-11</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1117</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>95</mods:start>
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<mods:end>122</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1117.86984</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1117.86984</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1117-95</mods:identifier>
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<mods:identifier type="ZooBank">C583CC7AB2AD42048FA083C49BB088EA</mods:identifier>
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<mods:identifier type="Pensoft-UUID">54DB7D7318935384856B59FD7D22DFF4</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:zoobank.org:act:F1432B21-5E24-4894-93D6-6F4546BB04BB" httpUri="http://treatment.plazi.org/id/69786D3A558B5E689389141F23A11488" lastPageNumber="95" pageId="0" pageNumber="95">
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<subSubSection pageId="0" pageNumber="95" type="nomenclature">
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<paragraph pageId="0" pageNumber="95">
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<taxonomicName LSID="https://zoobank.org/F1432B21-5E24-4894-93D6-6F4546BB04BB" authority="Holzenthal & Blahnik & Ríos-Touma, 2022" authorityName="Holzenthal & Blahnik & Ríos-Touma" authorityYear="2022" class="Insecta" family="Philopotamidae" genus="Chimarra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chimarra (Chimarra) asterae" order="Trichoptera" pageId="0" pageNumber="95" phylum="Arthropoda" rank="species" species="asterae" status="sp. nov." subGenus="Chimarra">Chimarra (Chimarra) asterae</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="95">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="95" type="description">
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<paragraph pageId="0" pageNumber="95">
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<figureCitation captionStart="Figure 6" captionStartId="F5" captionText="Figure 6. Chimarra (Chimarra) asterae sp. nov. Male genitalia A segments IX, X, lateral B segments IX, X, dorsal C inferior appendages, oblique lateral D inferior appendage, ventral E inferior appendage, dorsal F phallus, lateral G inferior appendage, C. duckworthi, lateral (for comparison) H inferior appendage, C. duckworthi, ventral (for comparison)." figureDoi="10.3897/zookeys.1117.86984.figure6" httpUri="https://binary.pensoft.net/fig/726434" pageId="0" pageNumber="95">Figs 6</figureCitation>
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<figureCitation captionStart="Figures 7, 8" captionStartId="F6" captionText="Figures 7, 8. Chimarra (Chimarra) asterae sp. nov. 7 female genitalia, ventral 8 Chimarra (Chimarra) duckworthi, female genitalia, ventral (for comparison). Abbreviations: vag. app. = vaginal apparatus." figureDoi="10.3897/zookeys.1117.86984.figures7-8" httpUri="https://binary.pensoft.net/fig/726435" pageId="0" pageNumber="95">, 7</figureCitation>
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<figureCitation captionStart="Figures 7, 8" captionStartId="F6" captionText="Figures 7, 8. Chimarra (Chimarra) asterae sp. nov. 7 female genitalia, ventral 8 Chimarra (Chimarra) duckworthi, female genitalia, ventral (for comparison). Abbreviations: vag. app. = vaginal apparatus." figureDoi="10.3897/zookeys.1117.86984.figures7-8" httpUri="https://binary.pensoft.net/fig/726435" pageId="0" pageNumber="95">, 8</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="95" type="type material">
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<paragraph pageId="0" pageNumber="95">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="95">
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<emphasis bold="true" pageId="0" pageNumber="95">
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<emphasis italics="true" pageId="0" pageNumber="95">Holotype</emphasis>
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.
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</emphasis>
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Male (pinned). Ecuador: Morona-Santiago: Macas, small gravel stream (Wallace/Real property),
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<geoCoordinate degrees="2.20299" direction="south" orientation="latitude" precision="1" value="-2.20299">2.20299°S</geoCoordinate>
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,
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<geoCoordinate degrees="78.08539" direction="west" orientation="longitude" precision="1" value="-78.08539">78.08539°W</geoCoordinate>
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, 1076 m a.s.l., 14.xi.2015,
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<normalizedToken originalValue="Ríos-Touma">Rios-Touma</normalizedToken>
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, Thomson, Amigo, Real-Wallace, UMSP000357522 (UMSP).
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<emphasis bold="true" pageId="0" pageNumber="95">
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<emphasis italics="true" pageId="0" pageNumber="95">Paratypes</emphasis>
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.
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</emphasis>
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Ecuador: same data as holotype 28 males, 39 females (pinned) (UMSP); same locality as holotype, except 27.i.2015, Holzenthal, Huisman,
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<normalizedToken originalValue="Ríos-Touma">Rios-Touma</normalizedToken>
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, Amigo, 4 males, 11 females (pinned), 3 males (in alcohol) (MECN).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="95" type="diagnosis">
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<paragraph pageId="0" pageNumber="95">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="95">
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<taxonomicName authorityName="Holzenthal & Blahnik & Ríos-Touma" authorityYear="2022" class="Insecta" family="Philopotamidae" genus="Chimarra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chimarra asterae" order="Trichoptera" pageId="0" pageNumber="95" phylum="Arthropoda" rank="species" species="asterae">
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<emphasis italics="true" pageId="0" pageNumber="95">Chimarra asterae</emphasis>
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</taxonomicName>
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is a member of the
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<taxonomicName authorityName="Ulmer" authorityYear="1909" class="Insecta" family="Philopotamidae" genus="Chimarra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chimarra (Chimarra) bidens" order="Trichoptera" pageId="0" pageNumber="95" phylum="Arthropoda" rank="species" species="bidens" subGenus="Chimarra">Chimarra (Chimarra) bidens</taxonomicName>
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group of
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<bibRefCitation DOI="https://doi.org/10.3897/zookeys.184.2911" author="Blahnik, RJ" journalOrPublisher="Memoirs of the American Entomological Institute" pageId="0" pageNumber="95" pagination="1 - 318" refId="B1" refString="Blahnik, RJ, 1998. Revision of the Neotropical species of the genus Chimarra, subgenus Chimarra (Trichoptera: Philopotamidae). Memoirs of the American Entomological Institute 59: 1 - 318, DOI: https://doi.org/10.3897/zookeys.184.2911" title="Revision of the Neotropical species of the genus Chimarra, subgenus Chimarra (Trichoptera: Philopotamidae)." url="https://doi.org/10.3897/zookeys.184.2911" volume="59" year="1998">Blahnik (1998)</bibRefCitation>
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, very similar to
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<taxonomicName genus="C." lsidName="C. duckworthi" pageId="0" pageNumber="95" rank="species" species="duckworthi">
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<emphasis italics="true" pageId="0" pageNumber="95">C. duckworthi</emphasis>
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</taxonomicName>
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Flint, 1967, particularly because of the general shape and length of the inferior appendages. The distinctly different sclerotization of the female genitalia provides the best evidence that the two forms are distinct species. The most distinctive differences in the male genitalia are in the structure of the inferior appendages, which have the basal part more broadly rounded, in lateral view, and the apex slightly more rounded, with a small notch or tooth-like projection pre-apically on the mesal surface (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F5" captionText="Figure 6. Chimarra (Chimarra) asterae sp. nov. Male genitalia A segments IX, X, lateral B segments IX, X, dorsal C inferior appendages, oblique lateral D inferior appendage, ventral E inferior appendage, dorsal F phallus, lateral G inferior appendage, C. duckworthi, lateral (for comparison) H inferior appendage, C. duckworthi, ventral (for comparison)." figureDoi="10.3897/zookeys.1117.86984.figure6" httpUri="https://binary.pensoft.net/fig/726434" pageId="0" pageNumber="95">6C-E</figureCitation>
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), absent in
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<taxonomicName genus="C." lsidName="C. duckworthi" pageId="0" pageNumber="95" rank="species" species="duckworthi">
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<emphasis italics="true" pageId="0" pageNumber="95">C. duckworthi</emphasis>
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</taxonomicName>
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(Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F5" captionText="Figure 6. Chimarra (Chimarra) asterae sp. nov. Male genitalia A segments IX, X, lateral B segments IX, X, dorsal C inferior appendages, oblique lateral D inferior appendage, ventral E inferior appendage, dorsal F phallus, lateral G inferior appendage, C. duckworthi, lateral (for comparison) H inferior appendage, C. duckworthi, ventral (for comparison)." figureDoi="10.3897/zookeys.1117.86984.figure6" httpUri="https://binary.pensoft.net/fig/726434" pageId="0" pageNumber="95">6G, H</figureCitation>
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).
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<taxonomicName authorityName="Holzenthal & Blahnik & Ríos-Touma" authorityYear="2022" class="Insecta" family="Philopotamidae" genus="Chimarra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chimarra asterae" order="Trichoptera" pageId="0" pageNumber="95" phylum="Arthropoda" rank="species" species="asterae">
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<emphasis italics="true" pageId="0" pageNumber="95">Chimarra asterae</emphasis>
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</taxonomicName>
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could also be confused with
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<taxonomicName genus="C." lsidName="C. caribea" pageId="0" pageNumber="95" rank="species" species="caribea">
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<emphasis italics="true" pageId="0" pageNumber="95">C. caribea</emphasis>
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</taxonomicName>
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Flint, 1968, which also has a small tooth-like projection near the apex of the inferior appendage. However, the overall length of the inferior appendage is longer in
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<taxonomicName genus="C." lsidName="C. caribea" pageId="0" pageNumber="95" rank="species" species="caribea">
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<emphasis italics="true" pageId="0" pageNumber="95">C. caribea</emphasis>
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</taxonomicName>
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and the tooth-like projection is somewhat more removed from the apex (
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<bibRefCitation DOI="https://doi.org/10.3897/zookeys.184.2911" author="Blahnik, RJ" journalOrPublisher="Memoirs of the American Entomological Institute" pageId="0" pageNumber="95" pagination="1 - 318" refId="B1" refString="Blahnik, RJ, 1998. Revision of the Neotropical species of the genus Chimarra, subgenus Chimarra (Trichoptera: Philopotamidae). Memoirs of the American Entomological Institute 59: 1 - 318, DOI: https://doi.org/10.3897/zookeys.184.2911" title="Revision of the Neotropical species of the genus Chimarra, subgenus Chimarra (Trichoptera: Philopotamidae)." url="https://doi.org/10.3897/zookeys.184.2911" volume="59" year="1998">Blahnik 1998</bibRefCitation>
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: fig. 54C, D,F).
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</paragraph>
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<caption doi="10.3897/zookeys.1117.86984.figure6" httpUri="https://binary.pensoft.net/fig/726434" pageId="0" pageNumber="95" start="Figure 6" startId="F5">
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<paragraph pageId="0" pageNumber="95">
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<emphasis bold="true" pageId="0" pageNumber="95">Figure 6.</emphasis>
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<taxonomicName authorityName="Holzenthal & Blahnik & Ríos-Touma" authorityYear="2022" class="Insecta" family="Philopotamidae" genus="Chimarra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chimarra (Chimarra) asterae" order="Trichoptera" pageId="0" pageNumber="95" phylum="Arthropoda" rank="species" species="asterae" subGenus="Chimarra">Chimarra (Chimarra) asterae</taxonomicName>
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sp. nov. Male genitalia
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<emphasis bold="true" pageId="0" pageNumber="95">A</emphasis>
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segments IX, X, lateral
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<emphasis bold="true" pageId="0" pageNumber="95">B</emphasis>
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segments IX, X, dorsal
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<emphasis bold="true" pageId="0" pageNumber="95">C</emphasis>
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inferior appendages, oblique lateral
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<emphasis bold="true" pageId="0" pageNumber="95">D</emphasis>
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inferior appendage, ventral
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<emphasis bold="true" pageId="0" pageNumber="95">E</emphasis>
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inferior appendage, dorsal
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<emphasis bold="true" pageId="0" pageNumber="95">F</emphasis>
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phallus, lateral
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<emphasis bold="true" pageId="0" pageNumber="95">G</emphasis>
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inferior appendage,
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<taxonomicName genus="C." lsidName="C. duckworthi" pageId="0" pageNumber="95" rank="species" species="duckworthi">
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<emphasis italics="true" pageId="0" pageNumber="95">C. duckworthi</emphasis>
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</taxonomicName>
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, lateral (for comparison)
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<emphasis bold="true" pageId="0" pageNumber="95">H</emphasis>
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inferior appendage,
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<taxonomicName genus="C." lsidName="C. duckworthi" pageId="0" pageNumber="95" rank="species" species="duckworthi">
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<emphasis italics="true" pageId="0" pageNumber="95">C. duckworthi</emphasis>
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</taxonomicName>
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, ventral (for comparison).
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="95">
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The female genitalia of
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<taxonomicName genus="C." lsidName="C. asterae" pageId="0" pageNumber="95" rank="species" species="asterae">
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<emphasis italics="true" pageId="0" pageNumber="95">C. asterae</emphasis>
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</taxonomicName>
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resemble
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<taxonomicName genus="C." lsidName="C. duckworthi" pageId="0" pageNumber="95" rank="species" species="duckworthi">
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<emphasis italics="true" pageId="0" pageNumber="95">C. duckworthi</emphasis>
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</taxonomicName>
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in having both a dorsal and paired ventral sclerites near the posterior opening of the vaginal apparatus, as well as distinct, membranous pocket-like lobes (probably receptacles for the inferior appendages of the male) associated with the ventral sclerites of segment IX (Fig.
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<figureCitation captionStart="Figures 7, 8" captionStartId="F6" captionText="Figures 7, 8. Chimarra (Chimarra) asterae sp. nov. 7 female genitalia, ventral 8 Chimarra (Chimarra) duckworthi, female genitalia, ventral (for comparison). Abbreviations: vag. app. = vaginal apparatus." figureDoi="10.3897/zookeys.1117.86984.figures7-8" httpUri="https://binary.pensoft.net/fig/726435" pageId="0" pageNumber="95">8</figureCitation>
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). The genitalia of
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<taxonomicName genus="C." lsidName="C. asterae" pageId="0" pageNumber="95" rank="species" species="asterae">
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<emphasis italics="true" pageId="0" pageNumber="95">C. asterae</emphasis>
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</taxonomicName>
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differs in that the sclerotized ventral furrows of the vaginal apparatus are short, and the lateral margins of the vaginal tract have distinct sclerites (Fig.
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<figureCitation captionStart="Figures 7, 8" captionStartId="F6" captionText="Figures 7, 8. Chimarra (Chimarra) asterae sp. nov. 7 female genitalia, ventral 8 Chimarra (Chimarra) duckworthi, female genitalia, ventral (for comparison). Abbreviations: vag. app. = vaginal apparatus." figureDoi="10.3897/zookeys.1117.86984.figures7-8" httpUri="https://binary.pensoft.net/fig/726435" pageId="0" pageNumber="95">7</figureCitation>
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). Also, it lacks the paired posteroventral sclerites, posterior to the sclerotized ventral furrows, which form an element of the vaginal apparatus in
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<taxonomicName genus="C." lsidName="C. duckworthi" pageId="0" pageNumber="95" rank="species" species="duckworthi">
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<emphasis italics="true" pageId="0" pageNumber="95">C. duckworthi</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<caption doi="10.3897/zookeys.1117.86984.figures7-8" httpUri="https://binary.pensoft.net/fig/726435" pageId="0" pageNumber="95" start="Figures 7, 8" startId="F6">
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<paragraph pageId="0" pageNumber="95">
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<emphasis bold="true" pageId="0" pageNumber="95">Figures 7, 8.</emphasis>
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<taxonomicName authorityName="Holzenthal & Blahnik & Ríos-Touma" authorityYear="2022" class="Insecta" family="Philopotamidae" genus="Chimarra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chimarra (Chimarra) asterae" order="Trichoptera" pageId="0" pageNumber="95" phylum="Arthropoda" rank="species" species="asterae" subGenus="Chimarra">Chimarra (Chimarra) asterae</taxonomicName>
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sp. nov.
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<emphasis bold="true" pageId="0" pageNumber="95">7</emphasis>
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female genitalia, ventral
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<emphasis bold="true" pageId="0" pageNumber="95">8</emphasis>
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<taxonomicName authorityName="Flint" authorityYear="1967" class="Insecta" family="Philopotamidae" genus="Chimarra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chimarra (Chimarra) duckworthi" order="Trichoptera" pageId="0" pageNumber="95" phylum="Arthropoda" rank="species" species="duckworthi" subGenus="Chimarra">Chimarra (Chimarra) duckworthi</taxonomicName>
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, female genitalia, ventral (for comparison). Abbreviations: vag. app. = vaginal apparatus.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="95" type="description">
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<paragraph pageId="0" pageNumber="95">Description.</paragraph>
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<paragraph pageId="0" pageNumber="95">
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<emphasis bold="true" pageId="0" pageNumber="95">Adult.</emphasis>
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Forewing length male 4.8-5.4 mm (
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<emphasis italics="true" pageId="0" pageNumber="95">n</emphasis>
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= 5); female 5.4-6.0 mm (
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<emphasis italics="true" pageId="0" pageNumber="95">n</emphasis>
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= 5). Color nearly uniformly brownish black (fuscous), except femora slightly paler. Head relatively short and rounded, (postocular parietal sclerite ca. half diameter of eye). Third segment of maxillary much longer than second, subequal to 5th. Male protarsal claws enlarged, asymmetrical in size and shape, outer claw longer and twisted.
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</paragraph>
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<paragraph pageId="0" pageNumber="95">
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<emphasis bold="true" pageId="0" pageNumber="95">Male.</emphasis>
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Abdominal segment IX, in lateral view, with very pronounced sinuous extension of anteroventral margin and small apodemes from anterodorsal margin; Posteroventral process very narrow, length greater than width, subacute apically. Tergum X membranous mesally, with sclerotized lateral lobes, each bearing pair of sensilla on short, rounded protuberance near dorsal margin in basal half, apex of lobe somewhat mesally cupped and bluntly rounded. Preanal appendage short, rounded, knob-like. Inferior appendage, in lateral view, relatively elongate (similar in length to
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<taxonomicName genus="C." lsidName="C. duckworthi" pageId="0" pageNumber="95" rank="species" species="duckworthi">
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<emphasis italics="true" pageId="0" pageNumber="95">C. duckworthi</emphasis>
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</taxonomicName>
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, shorter than
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<taxonomicName genus="C." lsidName="C. caribea" pageId="0" pageNumber="95" rank="species" species="caribea">
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<emphasis italics="true" pageId="0" pageNumber="95">C. caribea</emphasis>
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</taxonomicName>
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), mesally curved, with apical rotation, apices apposed, chisel-like, basoventral margin of appendage more broadly rounded than in
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<taxonomicName genus="C." lsidName="C. duckworthi" pageId="0" pageNumber="95" rank="species" species="duckworthi">
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<emphasis italics="true" pageId="0" pageNumber="95">C. duckworthi</emphasis>
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</taxonomicName>
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, apex of inferior appendage somewhat enlarged, with small tooth-like projection near apex on ventromesal margin. Phallobase with very distinct, acute apicoventral projection, two phallic spines, moderately elongate, differing slightly in length, endotheca tubular, more-or-less covered with small echinate spines, apicoventrally with curled sclerite and associated tract of very small spines. Phallotremal sclerite complex composed of elongate rod and ring structure and membranous structure with pair of associated wishbone-like sclerites apically.
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</paragraph>
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<paragraph pageId="0" pageNumber="95">
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<emphasis bold="true" pageId="0" pageNumber="95">Female.</emphasis>
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Ventral sclerites of sternum IX with prominent membranous lateral pouches (probably "clasper receptacles" of Blahnik, 1998). Vaginal apparatus moderately elongated with distinct rounded dorsal and paired ventral sclerites apically, ventrally with paired, narrow, furrow-like sclerites, beginning at mid-length, proximate posteriorly and diverging anteriorly; lateral margins of vaginal tract with elongate, narrow sclerites; vaginal tract narrowed anteriorly, with declivous, cup-like sclerite.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="95" type="etymology">
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<paragraph pageId="0" pageNumber="95">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="95">
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This new species is named in honor of Aster Real-Wallace, a young nature enthusiast and member of the Real-Wallace family, owners and protectors of a beautiful patch of remnant Amazonian pre-montane riverine forest on a tributary of the
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<normalizedToken originalValue="Río">Rio</normalizedToken>
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Upano, where this species was discovered.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |