174 lines
20 KiB
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174 lines
20 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1119.85400" ID-GBIF-Dataset="418bd521-e4b9-4ee6-981f-a6f471eab287" ID-Pensoft-Pub="1313-2970-1119-1" ID-Pensoft-UUID="F158A7C5261D52B69288A62F3C777CAF" ID-ZooBank="A3BE91C6B79344E1A886A803BF104D8B" ModsDocID="1313-2970-1119-1" checkinTime="1661532673136" checkinUser="pensoft" docAuthor="Jirapatrasilp, Parin, Sutcharit, Chirasak & Panha, Somsak" docDate="2022" docId="27DC3A3214135276945122EA2D7FA9AD" docLanguage="en" docName="ZooKeys 1119: 1-115" docOrigin="ZooKeys 1119" docPubDate="2022-08-25" docSource="http://dx.doi.org/10.3897/zookeys.1119.85400" docTitle="Pupina godwinausteni Jirapatrasilp 2022, sp. nov." docType="treatment" docUuid="EF5C2A8A-36DE-4C06-896A-CE55F9520C60" docUuidSource="ZooBank" docVersion="4" id="F158A7C5261D52B69288A62F3C777CAF" lastPageNumber="1" masterDocId="F158A7C5261D52B69288A62F3C777CAF" masterDocTitle="Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia" masterLastPageNumber="115" masterPageNumber="1" pageNumber="1" updateTime="1661537468408" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jirapatrasilp, Parin</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5591-6724</mods:nameIdentifier>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier type="email">jirasak4@yahoo.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Panha, Somsak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand & Academy of Science, The Royal Society of Thailand, Bangkok 10300, Thailand</mods:affiliation>
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<mods:nameIdentifier type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-08-25</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1119</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>115</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1119.85400</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1119.85400</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1119-1</mods:identifier>
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<mods:identifier type="ZooBank">A3BE91C6B79344E1A886A803BF104D8B</mods:identifier>
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<mods:identifier type="Pensoft-UUID">F158A7C5261D52B69288A62F3C777CAF</mods:identifier>
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<treatment ID-GBIF-Taxon="199839041" LSID="urn:lsid:zoobank.org:act:EF5C2A8A-36DE-4C06-896A-CE55F9520C60" httpUri="http://treatment.plazi.org/id/27DC3A3214135276945122EA2D7FA9AD" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="https://zoobank.org/EF5C2A8A-36DE-4C06-896A-CE55F9520C60" authority="Jirapatrasilp" authorityName="Jirapatrasilp" authorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina godwinausteni" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="godwinausteni" status="sp. nov.">Pupina godwinausteni Jirapatrasilp</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Umbilical, columellar and parietal views of A Pupinella mansuyi, specimen CUMZ 12148 from Pha Chu, Nan B Pupina artata from the Pupina artata species group, specimen CUMZ 12003 from Ban Ping Khong, Chiang Mai C Pupina godwinausteni sp. nov. from the Pupina arula species group, holotype CUMZ 12090 / 1 D Pupina aureola from the Pupina aureola species group, specimen CUMZ 12130 from Sra Morakot, Krabi." figureDoi="10.3897/zookeys.1119.85400.figure10" httpUri="https://binary.pensoft.net/fig/735371" pageId="0" pageNumber="1">Figs 10C</figureCitation>
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<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Shells of Pupina arula species group from mainland Southeast Asia A-G Pupina peguensis A syntype of Pupina blanfordi NHMUK 1888.12.4.100 and specimens B CUMZ 12105 C CUMZ 12050 D CUMZ 12103 E NHMUK ex. Cuming coll. F CUMZ 12094, and G CUMZ 12108 H-J Pupina crosseana H lectotype MNHN-IM- 2000 - 35834 I paralectotype RBINS MT 966 / 10591, and J specimen CUMZ 12049 K, L Pupina siamensis K lectotype SMF 109948 and L specimen CUMZ 12052 M-P Pupina bilabiata sp. nov. M holotype CUMZ 12073 / 1 and specimens N CUMZ 12079 O CUMZ 12081, and P CUMZ 12086 Q, R Pupina godwinausteni sp. nov. Q holotype CUMZ 12090 / 1 and R paratype CUMZ 12090 / 2. Photo: P. Maestrati, MNHN (H), F. Trus, RBINS (I)." figureDoi="10.3897/zookeys.1119.85400.figure28" httpUri="https://binary.pensoft.net/fig/735389" pageId="0" pageNumber="1">, 28Q, R</figureCitation>
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<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Live specimens of A Pupina siamensis, specimen CUMZ 12071 from Tham Khao Loi Temple, Rayong B-D Pupina bilabiata sp. nov. B paratype CUMZ 12073 / 2 from Banpot Pisai Temple, Chumphon and specimens C CUMZ 12082 from Pha Jor Cave, Nong Bua Lam Phu and D CUMZ 12087 from Ban Yai, Surat Thani E, F Pupina godwinausteni sp. nov.: paratypes E CUMZ 12090 / 26 and F CUMZ 12091 from Khao Wong Cave, Uthai Thani G, H Pupina aureola: specimens G CUMZ 12117 from Lod Cave, Nakhon Sri Thammarat and H CUMZ 12121 from Tham Thong Panara Temple, Nakhon Sri Thammarat, showing its microhabitat in rotten log. All not to scale." figureDoi="10.3897/zookeys.1119.85400.figure32" httpUri="https://binary.pensoft.net/fig/735393" pageId="0" pageNumber="1">, 32E, F</figureCitation>
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<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. A Pupina bilabiata sp. nov., specimen CUMZ 12086 from Na San Temple, Surat Thani B, C Pupina godwinausteni sp. nov. B holotype CUMZ 12090 / 1 and C paratype CUMZ 12090 / 2 from Khao Wong Cave, Uthai Thani D Pupina arula, syntype UMZC I. 103025 " Ind " E Pupina mouhoti, possible syntype NHMUK ex. Cuming coll. from Camboja F Pupina vescoi, syntype NHMUK 1893.2.4.767 from Cochin China. Photo: J. Ablett, H. Taylor, NHM (D)." figureDoi="10.3897/zookeys.1119.85400.figure33" httpUri="https://binary.pensoft.net/fig/735394" pageId="0" pageNumber="1">, 33B, C</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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CUMZ 12090/1 (Figs
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Umbilical, columellar and parietal views of A Pupinella mansuyi, specimen CUMZ 12148 from Pha Chu, Nan B Pupina artata from the Pupina artata species group, specimen CUMZ 12003 from Ban Ping Khong, Chiang Mai C Pupina godwinausteni sp. nov. from the Pupina arula species group, holotype CUMZ 12090 / 1 D Pupina aureola from the Pupina aureola species group, specimen CUMZ 12130 from Sra Morakot, Krabi." figureDoi="10.3897/zookeys.1119.85400.figure10" httpUri="https://binary.pensoft.net/fig/735371" pageId="0" pageNumber="1">10C</figureCitation>
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,
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<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Shells of Pupina arula species group from mainland Southeast Asia A-G Pupina peguensis A syntype of Pupina blanfordi NHMUK 1888.12.4.100 and specimens B CUMZ 12105 C CUMZ 12050 D CUMZ 12103 E NHMUK ex. Cuming coll. F CUMZ 12094, and G CUMZ 12108 H-J Pupina crosseana H lectotype MNHN-IM- 2000 - 35834 I paralectotype RBINS MT 966 / 10591, and J specimen CUMZ 12049 K, L Pupina siamensis K lectotype SMF 109948 and L specimen CUMZ 12052 M-P Pupina bilabiata sp. nov. M holotype CUMZ 12073 / 1 and specimens N CUMZ 12079 O CUMZ 12081, and P CUMZ 12086 Q, R Pupina godwinausteni sp. nov. Q holotype CUMZ 12090 / 1 and R paratype CUMZ 12090 / 2. Photo: P. Maestrati, MNHN (H), F. Trus, RBINS (I)." figureDoi="10.3897/zookeys.1119.85400.figure28" httpUri="https://binary.pensoft.net/fig/735389" pageId="0" pageNumber="1">28Q</figureCitation>
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,
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<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. A Pupina bilabiata sp. nov., specimen CUMZ 12086 from Na San Temple, Surat Thani B, C Pupina godwinausteni sp. nov. B holotype CUMZ 12090 / 1 and C paratype CUMZ 12090 / 2 from Khao Wong Cave, Uthai Thani D Pupina arula, syntype UMZC I. 103025 " Ind " E Pupina mouhoti, possible syntype NHMUK ex. Cuming coll. from Camboja F Pupina vescoi, syntype NHMUK 1893.2.4.767 from Cochin China. Photo: J. Ablett, H. Taylor, NHM (D)." figureDoi="10.3897/zookeys.1119.85400.figure33" httpUri="https://binary.pensoft.net/fig/735394" pageId="0" pageNumber="1">33B</figureCitation>
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), 5 June 2017, coll. C. Sutcharit, R. Srisonchai, A. Pholyotha. Measurement: shell height 8.8 mm, shell width 6.8 mm and 5 whorls.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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CUMZ 12090/2-26 (24 shells and 1 specimen in ethanol; Figs
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<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Shells of Pupina arula species group from mainland Southeast Asia A-G Pupina peguensis A syntype of Pupina blanfordi NHMUK 1888.12.4.100 and specimens B CUMZ 12105 C CUMZ 12050 D CUMZ 12103 E NHMUK ex. Cuming coll. F CUMZ 12094, and G CUMZ 12108 H-J Pupina crosseana H lectotype MNHN-IM- 2000 - 35834 I paralectotype RBINS MT 966 / 10591, and J specimen CUMZ 12049 K, L Pupina siamensis K lectotype SMF 109948 and L specimen CUMZ 12052 M-P Pupina bilabiata sp. nov. M holotype CUMZ 12073 / 1 and specimens N CUMZ 12079 O CUMZ 12081, and P CUMZ 12086 Q, R Pupina godwinausteni sp. nov. Q holotype CUMZ 12090 / 1 and R paratype CUMZ 12090 / 2. Photo: P. Maestrati, MNHN (H), F. Trus, RBINS (I)." figureDoi="10.3897/zookeys.1119.85400.figure28" httpUri="https://binary.pensoft.net/fig/735389" pageId="0" pageNumber="1">28R</figureCitation>
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,
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<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Live specimens of A Pupina siamensis, specimen CUMZ 12071 from Tham Khao Loi Temple, Rayong B-D Pupina bilabiata sp. nov. B paratype CUMZ 12073 / 2 from Banpot Pisai Temple, Chumphon and specimens C CUMZ 12082 from Pha Jor Cave, Nong Bua Lam Phu and D CUMZ 12087 from Ban Yai, Surat Thani E, F Pupina godwinausteni sp. nov.: paratypes E CUMZ 12090 / 26 and F CUMZ 12091 from Khao Wong Cave, Uthai Thani G, H Pupina aureola: specimens G CUMZ 12117 from Lod Cave, Nakhon Sri Thammarat and H CUMZ 12121 from Tham Thong Panara Temple, Nakhon Sri Thammarat, showing its microhabitat in rotten log. All not to scale." figureDoi="10.3897/zookeys.1119.85400.figure32" httpUri="https://binary.pensoft.net/fig/735393" pageId="0" pageNumber="1">32E</figureCitation>
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,
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<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. A Pupina bilabiata sp. nov., specimen CUMZ 12086 from Na San Temple, Surat Thani B, C Pupina godwinausteni sp. nov. B holotype CUMZ 12090 / 1 and C paratype CUMZ 12090 / 2 from Khao Wong Cave, Uthai Thani D Pupina arula, syntype UMZC I. 103025 " Ind " E Pupina mouhoti, possible syntype NHMUK ex. Cuming coll. from Camboja F Pupina vescoi, syntype NHMUK 1893.2.4.767 from Cochin China. Photo: J. Ablett, H. Taylor, NHM (D)." figureDoi="10.3897/zookeys.1119.85400.figure33" httpUri="https://binary.pensoft.net/fig/735394" pageId="0" pageNumber="1">33C</figureCitation>
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) and NHMUK 20210335 (3 shells), same data as holotype; CUMZ 12091 (20 shells and 24 specimens in ethanol; Fig.
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<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Live specimens of A Pupina siamensis, specimen CUMZ 12071 from Tham Khao Loi Temple, Rayong B-D Pupina bilabiata sp. nov. B paratype CUMZ 12073 / 2 from Banpot Pisai Temple, Chumphon and specimens C CUMZ 12082 from Pha Jor Cave, Nong Bua Lam Phu and D CUMZ 12087 from Ban Yai, Surat Thani E, F Pupina godwinausteni sp. nov.: paratypes E CUMZ 12090 / 26 and F CUMZ 12091 from Khao Wong Cave, Uthai Thani G, H Pupina aureola: specimens G CUMZ 12117 from Lod Cave, Nakhon Sri Thammarat and H CUMZ 12121 from Tham Thong Panara Temple, Nakhon Sri Thammarat, showing its microhabitat in rotten log. All not to scale." figureDoi="10.3897/zookeys.1119.85400.figure32" httpUri="https://binary.pensoft.net/fig/735393" pageId="0" pageNumber="1">32F</figureCitation>
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) from the type locality, 5 Dec. 2020, coll. P. Jirapatrasilp, C. Sutcharit, A. Pholyotha.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type locality">
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<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Khao Wong Cave, Ban Rai District, Uthai Thani Province, Thailand,
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<geoCoordinate degrees="15" direction="north" minutes="01" orientation="latitude" precision="1" seconds="52.6" value="15.031278">15°01'52.6"N</geoCoordinate>
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,
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<geoCoordinate degrees="99" direction="east" minutes="27" orientation="longitude" precision="1" seconds="23.3" value="99.45647">99°27'23.3"E</geoCoordinate>
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, 246 m amsl.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Other material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<materialsCitation collectingDate="2016-07-28" county="Uthai Thani Province" location="Tham Namthip Bureau of Monks" municipality="Lan Sak District" specimenCount="1">
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CUMZ 12092 (2 shells) from
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<location LSID="urn:lsid:plazi:treatment:27DC3A3214135276945122EA2D7FA9AD:4162C86070AC64B33C863B2CE1FCD725" county="Uthai Thani Province" municipality="Lan Sak District" name="Tham Namthip Bureau of Monks">Tham Namthip Bureau of Monks</location>
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,
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<collectingMunicipality>Lan Sak District</collectingMunicipality>
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,
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<collectingCounty>Uthai Thani Province</collectingCounty>
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,
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<collectingDate value="2016-07-28">28 July 2016</collectingDate>
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</materialsCitation>
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.
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<materialsCitation collectingDate="2018-10-01" county="Uthai Thani Province" location="Hup Pa Tat" municipality="Lan Sak District" specimenCount="1">
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CUMZ 12093 (
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<specimenCount type="generic">1 specimen</specimenCount>
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in ethanol) from
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<location LSID="urn:lsid:plazi:treatment:27DC3A3214135276945122EA2D7FA9AD:D3BDCF0E671BE4644A7BB1BA45858AA6" county="Uthai Thani Province" municipality="Lan Sak District" name="Hup Pa Tat">Hup Pa Tat</location>
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,
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<collectingMunicipality>Lan Sak District</collectingMunicipality>
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,
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<collectingCounty>Uthai Thani Province</collectingCounty>
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,
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<collectingDate value="2018-10-01">1 Oct. 2018</collectingDate>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">Shell globose; last whorl ca. 80% of shell height. Apertural lip very thickened and slightly expanded; with a furrow between inner and outer peristomes; inner peristome thickened, cord-like; apertural lip angled when observed from lateral view. Columellar tooth curved downward like an earlobe.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
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<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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The globose shell shape of
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<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. godwinausteni" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="godwinausteni">
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<emphasis italics="true" pageId="0" pageNumber="1">P. godwinausteni</emphasis>
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</taxonomicName>
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sp. nov. is most similar to
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<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. siamensis" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="siamensis">
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<emphasis italics="true" pageId="0" pageNumber="1">P. siamensis</emphasis>
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</taxonomicName>
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, but
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<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. godwinausteni" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="godwinausteni">
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<emphasis italics="true" pageId="0" pageNumber="1">P. godwinausteni</emphasis>
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</taxonomicName>
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sp. nov. differs from
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<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. siamensis" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="siamensis">
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<emphasis italics="true" pageId="0" pageNumber="1">P. siamensis</emphasis>
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</taxonomicName>
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in having a larger shell, a more prominent parietal callus, a thicker apertural lip with a furrow between inner and outer peristomes, with an inner peristome thickened and cord-like, a longer posterior canal, a wider and more curved columellar tooth, and a more angled apertural lip when observed from lateral view.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">Shell height 7.7-9.5 mm; shell width 5.5-7.0 mm. Shell globose, solid, semi-transparent, brown, devoid of prominent sculpture on glazed smooth surface. Apex obtuse. Growth lines on shell surface inconspicuous. Whorls 5, last whorl large (ca. 80% of shell height) and bulging. Spire angle ca. 90°, somewhat extended. Sutures slightly impressed, but shallow. Aperture circular; lip highly thickened (ca. 0.4-0.5 mm wide and 0.5-0.6 mm thick) with darker colour, slightly expanded; apertural lip when observed from lateral view angled. Apertural lip with a furrow between inner and outer peristomes, with inner peristome thickened and cord-like. Parietal callus thickened with darker colour. Peristome interrupted by two canals; posterior canal ca. 1.0-1.2 mm long, 0.6 mm at its widest, curved outward and bulging at the outer margin; anterior canal slit-like, as long as apertural lip width. Parietal tooth thick; columellar tooth curved downward like an earlobe (ca. 2.2 mm long, 1.2 mm wide and 0.5 mm thick), covering anterior canal. Umbilicus closed. Operculum round, yellowish, and transparent corneous with uneven edge.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="etymology">
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<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="1">The specific epithet is dedicated to H.H. Godwin-Austen, a British malacologist, who prominently contributed to malacological studies in South and Southeast Asia.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="1">This new species is found in Uthai Thani Province, Thailand.</paragraph>
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</subSubSection>
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</treatment>
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</document> |