139 lines
17 KiB
XML
139 lines
17 KiB
XML
<document id="0ED2694FC6FC5E88D6FBAAFB0A9DEF83" ID-DOI="10.11646/zootaxa.2306.1.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="5304869" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629654615282" checkinUser="felipe" docAuthor="Malohlava, Vladimír & Bocak, Ladislav" docDate="2009" docId="FE0687C3FFAB4E1EE5E2FD3CFC90F837" docLanguage="en" docName="zt02306p050.pdf" docOrigin="Zootaxa 2306 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2306.1.4" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Paratelius Kazantsev 1992" docType="treatment" docVersion="4" lastPageNumber="45" masterDocId="023FFFBBFFAA4E1FE575FFC2FFDBFFF4" masterDocTitle="A revision of Paratelius Kazantsev, 1992 (Coleoptera: Lycidae)" masterLastPageNumber="50" masterPageNumber="44" pageNumber="45" updateTime="1699089800590" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="C963E647F5CD0D6EB75938A46F2C58F0">A revision of Paratelius Kazantsev, 1992 (Coleoptera: Lycidae)</mods:title>
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<mods:namePart id="9B78440DDEFB4C4898567E5283DB5A55">Malohlava, Vladimír</mods:namePart>
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<mods:namePart id="5CBAE88804B2708703B66B0A331F5239">Bocak, Ladislav</mods:namePart>
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<mods:title id="75236BD5317779E22E051D50CCF58B86">Zootaxa</mods:title>
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<mods:date id="AE43E5DA0D54CD6DDF3086232E2D9AEA">2009</mods:date>
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<mods:number id="DB92A1F63E0CC64C43F9F4CFD28D0C74">2306</mods:number>
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<mods:url id="9E99A9E4A4BF2E3C3BB1AA84D0183705">https://biotaxa.org/Zootaxa/article/view/zootaxa.2306.1.4</mods:url>
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<mods:classification id="BD1DFB63BDE34379633871DE2D7E6CBB">journal article</mods:classification>
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<mods:identifier id="FB447C350C9279039F72879092229229" type="DOI">10.11646/zootaxa.2306.1.4</mods:identifier>
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<treatment id="FE0687C3FFAB4E1EE5E2FD3CFC90F837" ID-DOI="http://doi.org/10.5281/zenodo.5312440" ID-Zenodo-Dep="5312440" LSID="urn:lsid:plazi:treatment:FE0687C3FFAB4E1EE5E2FD3CFC90F837" httpUri="http://treatment.plazi.org/id/FE0687C3FFAB4E1EE5E2FD3CFC90F837" lastPageNumber="45" pageId="1" pageNumber="45">
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<subSubSection id="3EB5655EFFAB4E1EE5E2FD3CFDE1FCEC" box="[151,570,766,792]" pageId="1" pageNumber="45" type="nomenclature">
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<paragraph id="761036D5FFAB4E1EE5E2FD3CFDE1FCEC" blockId="1.[151,570,766,792]" box="[151,570,766,792]" pageId="1" pageNumber="45">
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<heading id="2D5881B9FFAB4E1EE5E2FD3CFDE1FCEC" bold="true" box="[151,570,766,792]" fontSize="11" level="1" pageId="1" pageNumber="45" reason="1">
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<emphasis id="44DBEAC7FFAB4E1EE5E2FD3CFDE1FCEC" bold="true" box="[151,570,766,792]" pageId="1" pageNumber="45">
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Genus
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<taxonomicName id="B1AF4D56FFAB4E1EE59BFD3CFDE1FCEC" authority="Kazantsev, 1992" authorityName="Kazantsev" authorityYear="1992" box="[238,570,766,792]" class="Insecta" family="Lycidae" genus="Paratelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="45" phylum="Arthropoda" rank="genus">
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<emphasis id="44DBEAC7FFAB4E1EE59BFD3CFEB1FCEC" bold="true" box="[238,362,766,792]" italics="true" pageId="1" pageNumber="45">Paratelius</emphasis>
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<bibRefCitation id="123E4B24FFAB4E1EE404FD3CFDE1FCEC" author="Kazantsev, S. V." box="[369,570,766,792]" pageId="1" pageNumber="45" pagination="97 - 104" refId="ref3983" refString="Kazantsev, S. V. (1992) Contribution a l'etude des lycides orientaux. Les Ateliini (Coleoptera). Revue Francaise d'Entomologie (Nouvelle serie), 14, 97 - 104." type="journal article" year="1992">Kazantsev, 1992</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="3EB5655EFFAB4E1EE5E2FC85FB95FC5B" pageId="1" pageNumber="45" type="diagnosis">
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<paragraph id="761036D5FFAB4E1EE5E2FC85FB95FC5B" blockId="1.[151,1436,839,1987]" pageId="1" pageNumber="45">
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<emphasis id="44DBEAC7FFAB4E1EE5E2FC85FEC0FC95" bold="true" box="[151,283,839,865]" pageId="1" pageNumber="45">Diagnosis:</emphasis>
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<taxonomicName id="B1AF4D56FFAB4E1EE451FC8AFE7BFC95" authorityName="Kazantsev" authorityYear="1992" box="[292,416,840,865]" class="Insecta" family="Lycidae" genus="Paratelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="45" phylum="Arthropoda" rank="genus">
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<emphasis id="44DBEAC7FFAB4E1EE451FC8AFE7BFC95" box="[292,416,840,865]" italics="true" pageId="1" pageNumber="45">Paratelius</emphasis>
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</taxonomicName>
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resembles
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<taxonomicName id="B1AF4D56FFAB4E1EE75DFC8AFD55FC95" authorityName="Waterhouse" authorityYear="1879" box="[552,654,840,865]" class="Insecta" family="Lycidae" genus="Scarelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="45" phylum="Arthropoda" rank="genus">
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<emphasis id="44DBEAC7FFAB4E1EE75DFC8AFD55FC95" box="[552,654,840,865]" italics="true" pageId="1" pageNumber="45">Scarelus</emphasis>
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</taxonomicName>
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in general appearance and has similar long setae between dense, short pubescence on elytra.
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<taxonomicName id="B1AF4D56FFAB4E1EE49BFCADFDB6FC7C" authorityName="Kazantsev" authorityYear="1992" box="[494,621,879,904]" class="Insecta" family="Lycidae" genus="Paratelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="45" phylum="Arthropoda" rank="genus">
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<emphasis id="44DBEAC7FFAB4E1EE49BFCADFDB6FC7C" box="[494,621,879,904]" italics="true" pageId="1" pageNumber="45">Paratelius</emphasis>
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</taxonomicName>
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, like
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<emphasis id="44DBEAC7FFAB4E1EE7C1FCADFCCBFC7C" box="[692,784,879,904]" italics="true" pageId="1" pageNumber="45">
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<taxonomicName id="B1AF4D56FFAB4E1EE7C1FCADFCD7FC7C" baseAuthorityName="Kazantsev" baseAuthorityYear="1992" box="[692,780,879,904]" class="Insecta" family="Lycidae" genus="Atelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="45" phylum="Arthropoda" rank="genus">Atelius</taxonomicName>
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,
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</emphasis>
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bears four longitudinal costae on each elytron. In all known species the phallus is very long, slender and slightly curved (
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<figureCitation id="EE942A50FFAB4E1EE6C3FC57FB9AFC5B" box="[950,1089,917,943]" captionStart="FIGURES 1" captionStartId="2.[151,269,1666,1690]" captionTargetBox="[198,1406,197,1640]" captionTargetId="figure-77@2.[175,1411,197,1641]" captionTargetPageId="2" captionText="FIGURES 1A–J. Male genitalia in the ventral and lateral views. A, B Paratelius snizeki n. sp.; C, D P. emasensis n. sp.; E, F P. nigricornis n. sp.; G, H P. diversicornis (Pic).; I, J. P. juvencus (Kleine). Scales 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5304871" httpUri="https://zenodo.org/record/5304871/files/figure.png" pageId="1" pageNumber="45">Figs 1A–1J</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="3EB5655EFFAB4E1EE5B0FC79FDA0F97A" pageId="1" pageNumber="45" type="description">
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<paragraph id="761036D5FFAB4E1EE5B0FC79FDA0F97A" blockId="1.[151,1436,839,1987]" pageId="1" pageNumber="45">
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<emphasis id="44DBEAC7FFAB4E1EE5B0FC79FEA6FC21" bold="true" box="[197,381,955,981]" pageId="1" pageNumber="45">Redescription:</emphasis>
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Similar to
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<taxonomicName id="B1AF4D56FFAB4E1EE775FC7EFDBFFC21" authorityName="Waterhouse" authorityYear="1879" box="[512,612,956,981]" class="Insecta" family="Lycidae" genus="Scarelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="45" phylum="Arthropoda" rank="genus">
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<emphasis id="44DBEAC7FFAB4E1EE775FC7EFDBFFC21" box="[512,612,956,981]" italics="true" pageId="1" pageNumber="45">Scarelus</emphasis>
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</taxonomicName>
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in general appearance. Small-bodied, slender, 5.8–7.0 mm long. Most species brown to reddish brown colored, some uniformly testaceous; when dark colored, humeral part of elytra lighter and apical half of elytra dark. Head small, free, hypognathous, covered with short pubescence, surface mat, with fine structure. Frons convex; eyes small; antennal tubercles inconspicuous, divided by shallow depression; antennal cavities very close each to other, separated by narrow cranial bridge. Antennae 11-segmented, compressed, slightly serrate, long, surpassing elytral apex by 3 to 5 antennomeres, covered with dense, recumbent pubescence; apical antennomere longest, narrowed to the apex, pointed. Mandibles stout, quite long, slightly curved, shining, with smooth internal margin. Other mouthparts tiny. Labium and maxillae clearly visible without dissecting, reduced. Labium reaching from gular margin of mouth cavity to one third of its diameter: maxillae reaching to a half of diameter. Labial palpi 2-segmented, maxillary palpi 4- segmented, apical palpomere stoutest in both palpi. Labrum projected forwards, triangular. Pronotum trapezoidal, flat, posterior margin wider than frontal one, frontal margin slightly rounded, lateral margins convex, posterior angles acutely projected, posterior margin almost straight. Pronotal disc with single median longitudinal carina with very deep and narrow depression in middle part. Scutellum flat, trapezoidal, shiny, sparsely pubescent, usually emarginate at apex. Elytra flat, with four longitudinal costae. Costae connected by regular transverse ridges, whole elytra densely and shortly pubescent, also bearing sparse, stout, long, erect setae. Legs slender, compressed, densely pubescent. Tarsus with 5 tarsomeres. Male genitalia trilobate, uniform in general shape (
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<figureCitation id="EE942A50FFAB4E1EE4A3F98FFDC5F993" box="[470,542,1613,1639]" captionStart="FIGURES 1" captionStartId="2.[151,269,1666,1690]" captionTargetBox="[198,1406,197,1640]" captionTargetId="figure-77@2.[175,1411,197,1641]" captionTargetPageId="2" captionText="FIGURES 1A–J. Male genitalia in the ventral and lateral views. A, B Paratelius snizeki n. sp.; C, D P. emasensis n. sp.; E, F P. nigricornis n. sp.; G, H P. diversicornis (Pic).; I, J. P. juvencus (Kleine). Scales 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5304871" httpUri="https://zenodo.org/record/5304871/files/figure.png" pageId="1" pageNumber="45">Figs 1</figureCitation>
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A-J). Phallus very long, widest at base, paramerae very short, globular in shape, phallobase longer than paramerae.
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</paragraph>
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</subSubSection>
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<subSubSection id="3EB5655EFFAB4E1EE5B0F958FC90F837" pageId="1" pageNumber="45" type="distribution">
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<paragraph id="761036D5FFAB4E1EE5B0F958FC90F837" blockId="1.[151,1436,839,1987]" pageId="1" pageNumber="45">
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<emphasis id="44DBEAC7FFAB4E1EE5B0F958FEB2F940" bold="true" box="[197,361,1690,1716]" pageId="1" pageNumber="45">Distribution:</emphasis>
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<collectingCountry id="0EB87645FFAB4E1EE404F958FE39F940" box="[369,482,1690,1716]" name="Malaysia" pageId="1" pageNumber="45">Malaysia</collectingCountry>
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:
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<collectingRegion id="B46BF837FFAB4E1EE49BF958FDA7F940" box="[494,636,1690,1716]" country="Malaysia" name="Sabah" pageId="1" pageNumber="45">Sabah State</collectingRegion>
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. Distribution is limited to mountain ranges in northeastern Borneo. These mountains were produced by a counterclockwise rotation of Borneo since the Late Miocene (
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<bibRefCitation id="123E4B24FFAB4E1EE03DF903FE90F8F6" author="Hall, R. & Blundell, D." pageId="1" pageNumber="45" refId="ref3955" refString="Hall, R. & Blundell, D. (1996) Tectonic Evolution of Southeast Asia. Geological Society Special Publications No. 106. London." type="book" year="1996">Hall & Blundell, 1996</bibRefCitation>
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). Based on the endemic occurence in a single mountain range, we may suppose speciation of all
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<taxonomicName id="B1AF4D56FFAB4E1EE5C8F8CDFE43F8DC" box="[189,408,1806,1832]" class="Insecta" family="Lycidae" genus="Paratelius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="45" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis id="44DBEAC7FFAB4E1EE5C8F8CDFEE2F8DC" box="[189,313,1807,1832]" italics="true" pageId="1" pageNumber="45">Paratelius</emphasis>
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species
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</taxonomicName>
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occurred in this region.
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<taxonomicName id="B1AF4D56FFAB4E1EE7B4F8CDFCE6F8DC" authorityName="Kazantsev" authorityYear="1992" box="[705,829,1807,1832]" class="Insecta" family="Lycidae" genus="Paratelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="45" phylum="Arthropoda" rank="genus">
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<emphasis id="44DBEAC7FFAB4E1EE7B4F8CDFCE6F8DC" box="[705,829,1807,1832]" italics="true" pageId="1" pageNumber="45">Paratelius</emphasis>
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</taxonomicName>
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has not been recently collected in the lowlands of Borneo, and only the
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<typeStatus id="A9148877FFAB4E1EE4E3F8F7FE13F8BB" box="[406,456,1845,1871]" pageId="1" pageNumber="45">type</typeStatus>
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specimen of
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<taxonomicName id="B1AF4D56FFAB4E1EE712F8F7FCC6F8BA" box="[615,797,1845,1870]" class="Insecta" family="Lycidae" genus="Paratelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="45" phylum="Arthropoda" rank="species" species="diversicornis">
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<emphasis id="44DBEAC7FFAB4E1EE712F8F7FCC6F8BA" box="[615,797,1845,1870]" italics="true" pageId="1" pageNumber="45">P. diversicornis</emphasis>
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</taxonomicName>
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is recorded from
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<collectingCountry id="0EB87645FFAB4E1EE685F8F7FB9BF8BB" box="[1008,1088,1845,1871]" name="Brunei" pageId="1" pageNumber="45">Brunei</collectingCountry>
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without precise locality data. Although Borneo was connected during the last glacial maximum to Indochina (
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<bibRefCitation id="123E4B24FFAB4E1EE139F89EFB0CF882" author="Voris, H. K." box="[1100,1239,1884,1910]" pageId="1" pageNumber="45" pagination="1153 - 1167" refId="ref4192" refString="Voris, H. K. (2000) Maps of Pleistocene sea levels in south-east Asia: shorelines, river systems and time duration. Journal of Biogeography, 27, 1153 - 1167." type="journal article" year="2000">Voris, 2000</bibRefCitation>
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) and the generic structure of both faunas is similar, neither
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<taxonomicName id="B1AF4D56FFAB4E1EE7F0F841FD24F868" authorityName="Kazantsev" authorityYear="1992" box="[645,767,1923,1948]" class="Insecta" family="Lycidae" genus="Paratelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="45" phylum="Arthropoda" rank="genus">
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<emphasis id="44DBEAC7FFAB4E1EE7F0F841FD24F868" box="[645,767,1923,1948]" italics="true" pageId="1" pageNumber="45">Paratelius</emphasis>
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</taxonomicName>
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from Borneo nor related
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<taxonomicName id="B1AF4D56FFAB4E1EE15CF841FBA1F868" baseAuthorityName="Kazantsev" baseAuthorityYear="1992" box="[1065,1146,1923,1948]" class="Insecta" family="Lycidae" genus="Atelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="45" phylum="Arthropoda" rank="genus">
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<emphasis id="44DBEAC7FFAB4E1EE15CF841FBA1F868" box="[1065,1146,1923,1948]" italics="true" pageId="1" pageNumber="45">Atelius</emphasis>
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</taxonomicName>
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from
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<collectingCountry id="0EB87645FFAB4E1EE1B4F841FAFFF869" box="[1217,1316,1923,1949]" name="Vietnam" pageId="1" pageNumber="45">Vietnam</collectingCountry>
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expanded their ranges over the subaerial shelf of the Asian continent.
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</paragraph>
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</subSubSection>
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</treatment>
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