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<document ID-DOI="http://dx.doi.org/10.3897/dez.65.21000" ID-GBIF-Dataset="03cce405-5887-4474-b040-35de18e8d020" ID-GBIF-Taxon="182223662" ID-Pensoft-Pub="1860-1324-1-1" ID-Pensoft-UUID="77D7AEC9294554E5BBDCABA37901022A" ID-Zenodo-Dep="1166397" ID-ZooBank="73DEE0F32BB04A21B4455E168FE50F54" LSID="urn:lsid:zoobank.org:act:469A110B-37CD-48A9-B118-E78BED4D6367" ModsDocID="1860-1324-65-1-1" checkinTime="1621372489409" checkinUser="pensoft" docAuthor="Liebherr, James K." docDate="2018" docId="EEC343DBD53D0DC5F2D9C350DB3B9533" docLanguage="en" docName="DeutEntomolZeit 65(1): 1-63" docOrigin="Deutsche Entomologische Zeitschrift 65 (1)" docPubDate="2018-01-18" docSource="http://dx.doi.org/10.3897/dez.65.21000" docTitle="Mecyclothorax picdupinsensis Liebherr 2018, sp. n." docType="treatment" docUuid="469A110B-37CD-48A9-B118-E78BED4D6367" docUuidSource="ZooBank" docVersion="3" id="77D7AEC9294554E5BBDCABA37901022A" lastPageId="43" lastPageNumber="44" masterDocId="77D7AEC9294554E5BBDCABA37901022A" masterDocTitle="Cladistic classification of Mecyclothorax Sharp (Coleoptera, Carabidae, Moriomorphini) and taxonomic revision of the New Caledonian subgenus Phacothorax Jeannel" masterLastPageNumber="63" masterPageNumber="1" pageId="42" pageNumber="43" updateTime="1643465225333" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Cladistic classification of Mecyclothorax Sharp (Coleoptera, Carabidae, Moriomorphini) and taxonomic revision of the New Caledonian subgenus Phacothorax Jeannel</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Liebherr, James K.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-9831-884X</mods:nameIdentifier>
<mods:affiliation>Cornell University Insect Collection, John H. and Anna B. Comstock Hall, Cornell University, Ithaca, NY 14853 - 2601, USA</mods:affiliation>
<mods:nameIdentifier type="email">jkl5@cornell.edu</mods:nameIdentifier>
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<mods:title>Deutsche Entomologische Zeitschrift</mods:title>
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<mods:date>2018</mods:date>
<mods:detail type="pubDate">
<mods:number>2018-01-18</mods:number>
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<mods:detail type="volume">
<mods:number>65</mods:number>
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<mods:detail type="issue">
<mods:number>1</mods:number>
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<mods:end>63</mods:end>
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<mods:url>http://dx.doi.org/10.3897/dez.65.21000</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/dez.65.21000</mods:identifier>
<mods:identifier type="Pensoft-Pub">1860-1324-1-1</mods:identifier>
<mods:identifier type="ZooBank">73DEE0F32BB04A21B4455E168FE50F54</mods:identifier>
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<mods:identifier type="Zenodo-Dep">1166397</mods:identifier>
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<subSection lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43" type="results">
<treatment ID-GBIF-Taxon="182223662" LSID="urn:lsid:zoobank.org:act:469A110B-37CD-48A9-B118-E78BED4D6367" httpUri="http://treatment.plazi.org/id/EEC343DBD53D0DC5F2D9C350DB3B9533" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43">
<subSubSection pageId="42" pageNumber="43" type="nomenclature">
<paragraph pageId="42" pageNumber="43">
15.
<taxonomicName LSID="http://zoobank.org/469A110B-37CD-48A9-B118-E78BED4D6367" authority="Liebherr" authorityName="Liebherr" authorityYear="2018" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax picdupinsensis" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="picdupinsensis" status="sp. n.">Mecyclothorax picdupinsensis Liebherr</taxonomicName>
<taxonomicNameLabel pageId="42" pageNumber="43">sp. n.</taxonomicNameLabel>
<figureCitation captionStart="Figure 20" captionStartId="F21" captionText="Figure 20. New Caledonian Mecyclothorax (Phacothorax) spp., dorsal view: A, M. manautei; B, M. paniensis; C, M. mouensis; D, M. kanak (Mt. Humboldt, 580 m); E, M. kanak (Mt. Humboldt, 630 m); F, M. kanak (Mt. Dzumac); G, M. kanak (Col de Yate); H, M. picdupinsensis." figureDoi="10.3897/dez.65.21000.figure20" httpUri="https://binary.pensoft.net/fig/179976" pageId="42" pageNumber="43">Figures 20H</figureCitation>
<figureCitation captionStart="Figure 21" captionStartId="F22" captionText="Figure 21. Male aedeagal median lobe and parameres, and ring sclerite-mediotergite plus antecostal margin, tergite IX-of Mecyclothorax (Phacothorax) spp.: A-B, M. kanak, right view, dorsal view in situ (Mt. Humboldt, 630 m); C-D, M. kanak, right view, dorsal view in situ (Mt. Humboldt, 1400 m); E, M. kanak, form Q, right view (Mt. Humboldt, 1300 m); F-G, M. kanak, right view, dorsal view in situ (Mt. Dzumac); H-I, M. kanak, right view, dorsal view in situ (Riviere Bleue); J-K, M. kanak, right view, dorsal view in situ (Col de Yate); L-N, M. picdupinsensis, right views, dorsal view of ring sclerite. See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure21" httpUri="https://binary.pensoft.net/fig/179977" pageId="42" pageNumber="43">, 21L-N</figureCitation>
<figureCitation captionStart="Figure 22" captionStartId="F23" captionText="Figure 22. Paired left (above) and right (below) parameres of Mecyclothorax (Phacothorax) spp. (ectal view): A, M. kanak; B, M. picdupinsensis." figureDoi="10.3897/dez.65.21000.figure22" httpUri="https://binary.pensoft.net/fig/179979" pageId="42" pageNumber="43">, 22B</figureCitation>
<figureCitation captionStart="Figure 23" captionStartId="F24" captionText="Figure 23. Female reproductive tract, gonocoxites and associated laterotergites, Mecyclothorax (Phacothorax) spp: A, M. manautei; B, M. paniensis; C, M. mouensis; D, M. kanak (Mt. Dzumac; common oviduct artificially darkened to show configuration); E, M. kanak (Pic du Grand Kaori); F, M. picdupinsensis. See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure23" httpUri="https://binary.pensoft.net/fig/179980" pageId="42" pageNumber="43">, 23F</figureCitation>
<figureCitation captionStart="Figure 24" captionStartId="F25" captionText="Figure 24. Left female gonocoxa of Mecyclothorax (Phacothorax) spp., ventral view: A, M. manautei; B, M. paniensis; C, M. mouensis; D, M. kanak; E, M. picdupinsensis. See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure24" httpUri="https://binary.pensoft.net/fig/179981" pageId="42" pageNumber="43">, 24E</figureCitation>
<figureCitation captionStart="Figure 25" captionStartId="F26" captionText="Figure 25. Geographical distributions of Mecyclothorax (Phacothorax) spp. (asterisked locality attributed to M. mouensis based on non-type female specimen)." figureDoi="10.3897/dez.65.21000.figure25" httpUri="https://binary.pensoft.net/fig/179982" pageId="42" pageNumber="43">, 25</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="42" pageNumber="43" type="diagnosis">
<paragraph pageId="42" pageNumber="43">Diagnosis.</paragraph>
<paragraph pageId="42" pageNumber="43">
This final member of the species triplet also including
<taxonomicName genus="M." lsidName="M. mouensis" pageId="42" pageNumber="43" rank="species" species="mouensis">
<emphasis italics="true" pageId="42" pageNumber="43">M. mouensis</emphasis>
</taxonomicName>
and
<taxonomicName genus="M." lsidName="M. kanak" pageId="42" pageNumber="43" rank="species" species="kanak">
<emphasis italics="true" pageId="42" pageNumber="43">M. kanak</emphasis>
</taxonomicName>
can be diagnosed from the former by characters listed in the
<taxonomicName genus="M." lsidName="M. kanak" pageId="42" pageNumber="43" rank="species" species="kanak">
<emphasis italics="true" pageId="42" pageNumber="43">M. kanak</emphasis>
</taxonomicName>
species diagnosis. As stated above, this species cannot be diagnosed from
<taxonomicName genus="M." lsidName="M. kanak" pageId="42" pageNumber="43" rank="species" species="kanak">
<emphasis italics="true" pageId="42" pageNumber="43">M. kanak</emphasis>
</taxonomicName>
based on external characters. The basis for species recognition lies both in the form of the male aedeagal median lobe apex-i.e., broadly rounded with a minute hitch present (Fig.
<figureCitation captionStart="Figure 21" captionStartId="F22" captionText="Figure 21. Male aedeagal median lobe and parameres, and ring sclerite-mediotergite plus antecostal margin, tergite IX-of Mecyclothorax (Phacothorax) spp.: A-B, M. kanak, right view, dorsal view in situ (Mt. Humboldt, 630 m); C-D, M. kanak, right view, dorsal view in situ (Mt. Humboldt, 1400 m); E, M. kanak, form Q, right view (Mt. Humboldt, 1300 m); F-G, M. kanak, right view, dorsal view in situ (Mt. Dzumac); H-I, M. kanak, right view, dorsal view in situ (Riviere Bleue); J-K, M. kanak, right view, dorsal view in situ (Col de Yate); L-N, M. picdupinsensis, right views, dorsal view of ring sclerite. See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure21" httpUri="https://binary.pensoft.net/fig/179977" pageId="42" pageNumber="43">21L</figureCitation>
)-and female gonocoxae with a broad mediobasal patch covered with quadrate cuticular scales (Fig.
<figureCitation captionStart="Figure 24" captionStartId="F25" captionText="Figure 24. Left female gonocoxa of Mecyclothorax (Phacothorax) spp., ventral view: A, M. manautei; B, M. paniensis; C, M. mouensis; D, M. kanak; E, M. picdupinsensis. See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure24" httpUri="https://binary.pensoft.net/fig/179981" pageId="42" pageNumber="43">24E</figureCitation>
). Also, based on specimens at hand, the female bursa copulatrix is shorter and broader in this species, and the helminthoid sclerite is more narrowly extended as a laminar structure that is heavily melanized (Fig.
<figureCitation captionStart="Figure 23" captionStartId="F24" captionText="Figure 23. Female reproductive tract, gonocoxites and associated laterotergites, Mecyclothorax (Phacothorax) spp: A, M. manautei; B, M. paniensis; C, M. mouensis; D, M. kanak (Mt. Dzumac; common oviduct artificially darkened to show configuration); E, M. kanak (Pic du Grand Kaori); F, M. picdupinsensis. See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure23" httpUri="https://binary.pensoft.net/fig/179980" pageId="42" pageNumber="43">23F</figureCitation>
). Standardized body length 3.03.4 mm. Chaetotaxy +/+//+/-//+/2/+/+.
</paragraph>
</subSubSection>
<subSubSection lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43" type="description">
<paragraph pageId="42" pageNumber="43">Description</paragraph>
<paragraph pageId="42" pageNumber="43">
(n = 5). The description of
<taxonomicName genus="M." lsidName="M. kanak" pageId="42" pageNumber="43" rank="species" species="kanak">
<emphasis italics="true" pageId="42" pageNumber="43">M. kanak</emphasis>
</taxonomicName>
can serve to describe this species, with substitution of the following ranges of ratios of the measured specimens: ocular ratio 1.44-1.52; ocular lobe ratio 0.71-0.80; EyL/EyD = 2.08-2.38; MPW/BPW = 2.30-2.44; MPW/PL = 1.33-1.38; APW/BPW = 1.32-1.36; MEW/EL = 0.85-0.89.
</paragraph>
<paragraph pageId="42" pageNumber="43">
<emphasis italics="true" pageId="42" pageNumber="43">Male genitalia</emphasis>
(n = 2). Antecostal margin of abdominal mediotergite IX angulate, not extended (Fig.
<figureCitation captionStart="Figure 21" captionStartId="F22" captionText="Figure 21. Male aedeagal median lobe and parameres, and ring sclerite-mediotergite plus antecostal margin, tergite IX-of Mecyclothorax (Phacothorax) spp.: A-B, M. kanak, right view, dorsal view in situ (Mt. Humboldt, 630 m); C-D, M. kanak, right view, dorsal view in situ (Mt. Humboldt, 1400 m); E, M. kanak, form Q, right view (Mt. Humboldt, 1300 m); F-G, M. kanak, right view, dorsal view in situ (Mt. Dzumac); H-I, M. kanak, right view, dorsal view in situ (Riviere Bleue); J-K, M. kanak, right view, dorsal view in situ (Col de Yate); L-N, M. picdupinsensis, right views, dorsal view of ring sclerite. See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure21" httpUri="https://binary.pensoft.net/fig/179977" pageId="42" pageNumber="43">21M</figureCitation>
); right paramere narrow, evenly narrowed from narrow base to whiplike apex, with one seta near midlength on ventral margin complementing two long apical setae (Fig.
<figureCitation captionStart="Figure 22" captionStartId="F23" captionText="Figure 22. Paired left (above) and right (below) parameres of Mecyclothorax (Phacothorax) spp. (ectal view): A, M. kanak; B, M. picdupinsensis." figureDoi="10.3897/dez.65.21000.figure22" httpUri="https://binary.pensoft.net/fig/179979" pageId="42" pageNumber="43">22B</figureCitation>
); left paramere narrow basally, evenly narrowed to whiplike apex, two longer apical setae present; aedeagal median lobe robust, broad dorsoventrally, evenly curved dorsally and ventrally to a broadly downturned apex which bears a variously developed apical invagination, present as a small tubercle or change of angle of the apical margin (Fig.
<figureCitation captionStart="Figure 21" captionStartId="F22" captionText="Figure 21. Male aedeagal median lobe and parameres, and ring sclerite-mediotergite plus antecostal margin, tergite IX-of Mecyclothorax (Phacothorax) spp.: A-B, M. kanak, right view, dorsal view in situ (Mt. Humboldt, 630 m); C-D, M. kanak, right view, dorsal view in situ (Mt. Humboldt, 1400 m); E, M. kanak, form Q, right view (Mt. Humboldt, 1300 m); F-G, M. kanak, right view, dorsal view in situ (Mt. Dzumac); H-I, M. kanak, right view, dorsal view in situ (Riviere Bleue); J-K, M. kanak, right view, dorsal view in situ (Col de Yate); L-N, M. picdupinsensis, right views, dorsal view of ring sclerite. See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure21" httpUri="https://binary.pensoft.net/fig/179977" pageId="42" pageNumber="43">21L</figureCitation>
); aedeagal median lobe internal sac with sclerotized region in position of dorsal plate, internal sac folded longitudinally toward apex (Fig.
<figureCitation captionStart="Figure 21" captionStartId="F22" captionText="Figure 21. Male aedeagal median lobe and parameres, and ring sclerite-mediotergite plus antecostal margin, tergite IX-of Mecyclothorax (Phacothorax) spp.: A-B, M. kanak, right view, dorsal view in situ (Mt. Humboldt, 630 m); C-D, M. kanak, right view, dorsal view in situ (Mt. Humboldt, 1400 m); E, M. kanak, form Q, right view (Mt. Humboldt, 1300 m); F-G, M. kanak, right view, dorsal view in situ (Mt. Dzumac); H-I, M. kanak, right view, dorsal view in situ (Riviere Bleue); J-K, M. kanak, right view, dorsal view in situ (Col de Yate); L-N, M. picdupinsensis, right views, dorsal view of ring sclerite. See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure21" httpUri="https://binary.pensoft.net/fig/179977" pageId="42" pageNumber="43">21L, N</figureCitation>
).
</paragraph>
<paragraph lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43">
<emphasis italics="true" pageId="42" pageNumber="43">Female reproductive tract</emphasis>
(n = 1). Bursa copulatrix length subequal to circumference, its surface thin, translucent (Fig.
<figureCitation captionStart="Figure 23" captionStartId="F24" captionText="Figure 23. Female reproductive tract, gonocoxites and associated laterotergites, Mecyclothorax (Phacothorax) spp: A, M. manautei; B, M. paniensis; C, M. mouensis; D, M. kanak (Mt. Dzumac; common oviduct artificially darkened to show configuration); E, M. kanak (Pic du Grand Kaori); F, M. picdupinsensis. See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure23" httpUri="https://binary.pensoft.net/fig/179980" pageId="42" pageNumber="43">23F</figureCitation>
); spermathecal duct entering at apex of bursa separate from bursa-common oviduct juncture; rounded laminar helminthoid sclerite extended from bursal wall between, and separate from both, bursal-common oviduct juncture and spermatheca; spermatheca of only slightly greater circumference than spermathecal duct, reservoir about 1/2 length of spermathecal duct, spermathecal gland duct entering at base of spermathecal reservoir (Fig.
<figureCitation captionStart="Figure 23" captionStartId="F24" captionText="Figure 23. Female reproductive tract, gonocoxites and associated laterotergites, Mecyclothorax (Phacothorax) spp: A, M. manautei; B, M. paniensis; C, M. mouensis; D, M. kanak (Mt. Dzumac; common oviduct artificially darkened to show configuration); E, M. kanak (Pic du Grand Kaori); F, M. picdupinsensis. See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure23" httpUri="https://binary.pensoft.net/fig/179980" pageId="42" pageNumber="43">23F</figureCitation>
); ligular apophysis present on common oviduct; basal gonocoxite 1 with apical fringe of two setae laterally, a very small setae may be present immediately mesad, and several smaller setae occur near apex of medial margin (Fig.
<figureCitation captionStart="Figure 24" captionStartId="F25" captionText="Figure 24. Left female gonocoxa of Mecyclothorax (Phacothorax) spp., ventral view: A, M. manautei; B, M. paniensis; C, M. mouensis; D, M. kanak; E, M. picdupinsensis. See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure24" httpUri="https://binary.pensoft.net/fig/179981" pageId="42" pageNumber="43">24E</figureCitation>
); medial margin bearing a dense field of mi
<pageBreakToken pageId="43" pageNumber="44" start="start">crotrichia</pageBreakToken>
basally, this field about 8 microtrichia maximal breadth, with the macrotrichia appearing like flattened scales; gonocoxite 2 moderately narrow basally, basal width 0.4
<normalizedToken originalValue="×">x</normalizedToken>
medial length; two gracile lateral ensiform setae of moderate length present, the apical seta broader.
</paragraph>
<paragraph pageId="43" pageNumber="44">
<emphasis italics="true" pageId="43" pageNumber="44">Types</emphasis>
- Holotype male (MNHN): NEW CALEDONIA 11787 / 22°15
<normalizedToken originalValue="Sx166°49">'Sx
<geoCoordinate degrees="166" direction="east" minutes="49" orientation="longitude" precision="925" value="166.81667">166°49'E</geoCoordinate>
</normalizedToken>
, 280m / Pic du Pin, site 1,rainfor. / 26Nov2004, berlesate / G.Monteith,P.Grimbacher // QUEENSLAND / MUSEUM LOAN / DATE: July 2005 / No. LE 05.24 //
<emphasis italics="true" pageId="43" pageNumber="44">
New Caledonia
<taxonomicName authorityName="Sharp" authorityYear="1903" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">Mecyclothorax</taxonomicName>
</emphasis>
revision / measured specimen 1 // HOLOTYPE /
<taxonomicName authorityName="Sharp" authorityYear="1903" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">Mecyclothorax</taxonomicName>
/
<taxonomicName authorityName="Liebherr" authorityYear="2018" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax picdupinsensis" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="picdupinsensis">Mecyclothorax picdupinsensis</taxonomicName>
/ J.K.Liebherr 2017 (black-bordered red label) /
</paragraph>
<paragraph pageId="43" pageNumber="44">
Paratypes (11 specimens). NEW CALEDONIA: Pic du Pin[code PA 48],
<geoCoordinate degrees="22" direction="south" minutes="15" orientation="latitude" precision="925" value="-22.25">22°15'S</geoCoordinate>
<geoCoordinate degrees="166" direction="east" minutes="49" orientation="longitude" precision="925" value="166.81667">166°49'E</geoCoordinate>
, 28-vi-1970, Franz (NHMW, 3), base, 280 m el.,
<geoCoordinate degrees="22" direction="south" minutes="14.9" orientation="latitude" precision="92" value="-22.248333">22°14.9'S</geoCoordinate>
<geoCoordinate degrees="166" direction="east" minutes="49.7" orientation="longitude" precision="92" value="166.82834">166°49.7'E</geoCoordinate>
, sifted rainforest, litter, 26-xii-2006, Wanat &amp; Dobosz (MNHW, 1),
<geoCoordinate degrees="22" direction="south" minutes="15" orientation="latitude" precision="925" value="-22.25">22°15'S</geoCoordinate>
<geoCoordinate degrees="166" direction="east" minutes="49" orientation="longitude" precision="925" value="166.81667">166°49'E</geoCoordinate>
, sifted litter, 22-x-2008, Wanat (MNHW, 1), site 1, rainforest, 280 m el.,
<geoCoordinate degrees="22" direction="south" minutes="15" orientation="latitude" precision="925" value="-22.25">22°15'S</geoCoordinate>
<geoCoordinate degrees="166" direction="east" minutes="49" orientation="longitude" precision="925" value="166.81667">166°49'E</geoCoordinate>
, berlesate, 26-xi-2004, lot 11787, Monteith &amp; Grimbacher (QMB, 5), 21-xii-2004, lot 12045, Monteith (QMB, 1).
</paragraph>
</subSubSection>
<subSubSection pageId="43" pageNumber="44" type="etymology">
<paragraph pageId="43" pageNumber="44">Etymology.</paragraph>
<paragraph pageId="43" pageNumber="44">
This species name represents the genitive adjectival form of the type locality, Pic du Pin;
<taxonomicName authorityName="Liebherr" authorityYear="2018" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax picdupinsensis" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="picdupinsensis">Mecyclothorax picdupinsensis</taxonomicName>
. This final species name includes reference to the New Caledonian Pine,
<taxonomicName class="Coniferopsida" family="Araucariaceae" genus="Araucaria" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Araucaria columnaris" order="Pinales" pageId="43" pageNumber="44" phylum="Gymnospermae" rank="species" species="columnaris">
<emphasis italics="true" pageId="43" pageNumber="44">Araucaria columnaris</emphasis>
</taxonomicName>
J. R. Forster and W. J. Hooker, and partially recapitulates the final species named in
<bibRefCitation DOI="https://doi.org/10.3897/dez.63.10241" author="Liebherr, JK" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="51" pageNumber="52" pagination="211 - 270" refId="B61" refString="Liebherr, JK, 2016. Cyphocoleus Chaudoir (Coleoptera, Carabidae, Odacanthini): descriptive taxonomy, phylogenetic relationships, and the Cenozoic history of New Caledonia. Deutsche Entomologische Zeitschrift 63: 211 - 270, DOI: https://doi.org/10.3897/dez.63.10241" title="Cyphocoleus Chaudoir (Coleoptera, Carabidae, Odacanthini): descriptive taxonomy, phylogenetic relationships, and the Cenozoic history of New Caledonia." url="https://doi.org/10.3897/dez.63.10241" volume="63" year="2016">Liebherr (2016)</bibRefCitation>
;
<taxonomicName authorityName="Liebherr" authorityYear="2016" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus iledespinsensis" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="iledespinsensis">
<emphasis italics="true" pageId="43" pageNumber="44">Cyphocoleus iledespinsensis</emphasis>
</taxonomicName>
Liebherr.
</paragraph>
</subSubSection>
<subSubSection pageId="43" pageNumber="44" type="distribution">
<paragraph pageId="43" pageNumber="44">Distribution and habitat.</paragraph>
<paragraph pageId="43" pageNumber="44">
This species is restricted to the vicinity of Pic du Pin, southern Grande Terre (Fig.
<figureCitation captionStart="Figure 25" captionStartId="F26" captionText="Figure 25. Geographical distributions of Mecyclothorax (Phacothorax) spp. (asterisked locality attributed to M. mouensis based on non-type female specimen)." figureDoi="10.3897/dez.65.21000.figure25" httpUri="https://binary.pensoft.net/fig/179982" pageId="43" pageNumber="44">25</figureCitation>
), with all collecting localities recorded at 280 m elevation. Nine of the 12 specimens were collected from sieved litter, whereas three specimens (NHMW) are associated with the collecting note: &quot;Total stehendes Stammen von
<taxonomicName genus="Aruacaria" lsidName="Aruacaria biramulata" pageId="43" pageNumber="44" rank="species" species="biramulata">
<emphasis italics="true" pageId="43" pageNumber="44">Aruacaria biramulata</emphasis>
</taxonomicName>
(Herbert Franz field notebook:
<normalizedToken originalValue="Jäch">Jaech</normalizedToken>
, pers. comm.)&quot;: Thus this species may climb the lower reaches and boles of larger trees in addition to occupying the ground-level litter layer.
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>