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<document id="3FCACD37A371BB8733B4210BD48BC563" ID-CLB-Dataset="5965" ID-DOI="10.11646/zootaxa.3758.1.1" ID-GBIF-Dataset="903b427b-6b4f-4f0b-9270-1459a7e5be1e" ID-ISSN="1175-5326" ID-Zenodo-Dep="5037597" ID-ZooBank="63F695AA-891F-48D2-AE96-7492016B9590" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623093915434" checkinUser="felipe" docAuthor="PERKINS, PHILIP D." docDate="2014" docId="E654878A670B0417F2EEB157FB1CF88A" docLanguage="en" docName="zt03758p092.pdf.imf" docOrigin="Zootaxa 3758 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.3758.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydraena capensis PERKINS, 2014, new species" docType="treatment" docVersion="5" lastPageNumber="14" masterDocId="1A6DFFF26706041AF279B33BFF8FFFE5" masterDocTitle="A revision of the water beetle genus Hydraena Kugelann for southern Africa (Coleoptera: Hydraenidae)" masterLastPageNumber="92" masterPageNumber="1" pageNumber="14" updateTime="1699001492311" updateUser="ExternalLinkService">
<mods:mods id="C943AF2AAA381282E4E64590B0EC5877" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="ADA46E2F35E7C2BB0DF02872AE8AC02D">
<mods:title id="C2F216D5CA90DF5E747972C7DFE382AD">A revision of the water beetle genus Hydraena Kugelann for southern Africa (Coleoptera: Hydraenidae)</mods:title>
</mods:titleInfo>
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<mods:namePart id="690561A137B8AC8C59F7A024B8D69819">PERKINS, PHILIP D.</mods:namePart>
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<mods:title id="62DD8280D6549AA9D344D9EDF50618C9">Zootaxa</mods:title>
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<mods:part id="3B9B76E88A0B970AAEC588A5FED6FB20">
<mods:date id="5BAFB61FE58F8BB516DBE3F6872E4867">2014</mods:date>
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<mods:number id="FC7E83786E3E62466D0ABB6D36604397">2014-01-29</mods:number>
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<mods:number id="543BBB82B8AC1941ADE2B7559F27A328">3758</mods:number>
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<mods:classification id="EC9B7EE96934DB6332EC99D057AC1CA9">journal article</mods:classification>
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<mods:identifier id="72CAB2D43F178A8AE019E939A56E838F" type="DOI">10.11646/zootaxa.3758.1.1</mods:identifier>
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<treatment id="E654878A670B0417F2EEB157FB1CF88A" ID-DOI="http://doi.org/10.5281/zenodo.5037621" ID-GBIF-Taxon="183196961" ID-Zenodo-Dep="5037621" LSID="urn:lsid:plazi:treatment:E654878A670B0417F2EEB157FB1CF88A" httpUri="http://treatment.plazi.org/id/E654878A670B0417F2EEB157FB1CF88A" lastPageNumber="14" pageId="13" pageNumber="14">
<subSubSection id="26E76517670B0417F2EEB157FD94FD63" box="[151,539,620,646]" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="6E42369C670B0417F2EEB157FD94FD63" blockId="13.[151,606,620,679]" box="[151,539,620,646]" pageId="13" pageNumber="14">
<heading id="350A81F0670B0417F2EEB157FD94FD63" bold="true" box="[151,539,620,646]" fontSize="11" level="2" pageId="13" pageNumber="14" reason="4">
<emphasis id="5C89EA8E670B0417F2EEB157FD94FD63" bold="true" box="[151,539,620,646]" pageId="13" pageNumber="14">
<taxonomicName id="A9FD4D1F670B0417F2EEB157FEF3FD63" ID-CoL="3N3R8" authorityName="Perkins" authorityYear="2014" box="[151,380,620,646]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="capensis" status="sp. nov.">
<emphasis id="5C89EA8E670B0417F2EEB157FEF3FD63" bold="true" box="[151,380,620,646]" italics="true" pageId="13" pageNumber="14">Hydraena capensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="47BA57F5670B0417F3F5B157FD94FD63" box="[396,539,620,646]" pageId="13" pageNumber="14" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="26E76517670B0417F2EEB1B5FDD1FD42" box="[151,606,654,679]" pageId="13" pageNumber="14" type="description">
<paragraph id="6E42369C670B0417F2EEB1B5FDD1FD42" blockId="13.[151,606,620,679]" box="[151,606,654,679]" pageId="13" pageNumber="14">
<figureCitation id="F6C62A19670B0417F2EEB1B5FF6CFD42" box="[151,227,654,679]" captionStart="FIGURE 6" captionStartId="14.[151,249,1701,1723]" captionTargetBox="[216,1371,179,1679]" captionTargetId="figure-109@14.[216,1371,167,1679]" captionTargetPageId="14" captionText="FIGURE 6. Hydraena capensis, holotype, dorsal and lateral habitus." figureDoi="http://doi.org/10.5281/zenodo.5037613" httpUri="https://zenodo.org/record/5037613/files/figure.png" pageId="13" pageNumber="14">Figs. 6</figureCitation>
(habitus), 8 (aedeagus), 78 (map)
</paragraph>
</subSubSection>
<subSubSection id="26E76517670B0417F2EEB1EDFA88FCD2" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="6E42369C670B0417F2EEB1EDFA88FCD2" blockId="13.[151,1437,726,1903]" pageId="13" pageNumber="14">
<emphasis id="5C89EA8E670B0417F2EEB1EDFEC8FD0A" bold="true" box="[151,327,726,751]" pageId="13" pageNumber="14">Type material.</emphasis>
<materialsCitation id="DE953CC1670B0417F329B1EDFEB7FCD2" ID-GBIF-Occurrence="3311567315" collectingDate="1903-07-08" collectionCode="ZMHB" collectorName="Labels" country="South Africa" latitude="-34.016666" location="Chapman's Bay" longLatPrecision="1202" longitude="18.333334" pageId="13" pageNumber="14" specimenCount="1" stateProvince="Eastern Cape" typeStatus="holotype">
<typeStatus id="B146883E670B0417F329B1EDFE37FD0A" box="[336,440,726,751]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
(male):
<emphasis id="5C89EA8E670B0417F065B1EDFC6FFD0A" bold="true" box="[540,992,726,751]" pageId="13" pageNumber="14">
<collectingCountry id="16EA760C670B0417F065B1EDFD34FD0A" box="[540,699,726,751]" name="South Africa" pageId="13" pageNumber="14">South Africa</collectingCountry>
:
<collectingRegion id="AC39F87E670B0417F0B3B1EDFC6FFD0A" box="[714,992,726,751]" country="South Africa" name="Eastern Cape" pageId="13" pageNumber="14">Eastern Cape Province</collectingRegion>
</emphasis>
,
<location id="6B226047670B0417F188B1EDFB29FD0A" LSID="urn:lsid:plazi:treatment:E654878A670B0417F2EEB157FB1CF88A:6B226047670B0417F188B1EDFB29FD0A" box="[1009,1190,726,751]" country="South Africa" latitude="-34.016666" longLatPrecision="1202" longitude="18.333334" name="Chapman's Bay" pageId="13" pageNumber="14" stateProvince="Eastern Cape">Chapmans Bay</location>
, little pond,
<geoCoordinate id="0BC9505B670B0417F740B1EDFA18FD0A" box="[1337,1431,726,751]" degrees="34" direction="south" minutes="1" orientation="latitude" pageId="13" pageNumber="14" precision="925" value="-34.016666">34° 1' S</geoCoordinate>
,
<geoCoordinate id="0BC9505B670B0417F2EEB1C1FE8DFCF7" box="[151,258,762,787]" degrees="18" direction="east" minutes="20" orientation="longitude" pageId="13" pageNumber="14" precision="925" value="18.333334">18° 20' E</geoCoordinate>
,
<date id="1A43105C670B0417F376B1C1FE0AFCF6" box="[271,389,762,787]" pageId="13" pageNumber="14" value="1903-07-08">
<collectingDate id="0A07E9B4670B0417F376B1C1FE0AFCF6" box="[271,389,762,787]" pageId="13" pageNumber="14" value="1903-07-08">8 vii 1903</collectingDate>
</date>
, Vanhoffen, Ernst (
<collectionCode id="08ECAE59670B0417F026B1C1FD39FCF6" box="[607,694,762,787]" pageId="13" pageNumber="14">ZMHB</collectionCode>
).
<collectorName id="C308534A670B0417F0B0B1C1FC97FCF6" box="[713,792,762,787]" pageId="13" pageNumber="14">Labels</collectorName>
: &quot; Capland Süsswassertümpel bei Chapmansbay
<date id="1A43105C670B0417F743B1C1FA13FCF6" box="[1338,1436,762,787]" pageId="13" pageNumber="14" value="1903-07-08">
<collectingDate id="0A07E9B4670B0417F743B1C1FA13FCF6" box="[1338,1436,762,787]" pageId="13" pageNumber="14" value="1903-07-08">8.7.1903</collectingDate>
</date>
Vanhöffen S.
<emphasis id="5C89EA8E670B0417F351B025FEB7FCD2" bold="true" box="[296,312,798,823]" pageId="13" pageNumber="14">&quot;</emphasis>
</materialsCitation>
<materialsCitation id="DE953CC1670B0417F346B025FA8CFCD2" ID-GBIF-Occurrence="3311567368" box="[319,1283,798,823]" collectingDate="1903-07-08" collectionCode="ZMHB" collectorName="Labels" country="South Africa" latitude="-34.016666" location="Same" longLatPrecision="1202" longitude="18.333334" pageId="13" pageNumber="14" specimenCount="1" stateProvince="Eastern Cape" typeStatus="paratype">
<emphasis id="5C89EA8E670B0417F346B025FE23FCD2" bold="true" box="[319,428,798,823]" pageId="13" pageNumber="14">
<typeStatus id="B146883E670B0417F346B025FE23FCD2" box="[319,428,798,823]" pageId="13" pageNumber="14" type="paratype">Paratype</typeStatus>
</emphasis>
:
<emphasis id="5C89EA8E670B0417F3C2B025FCF8FCD2" bold="true" box="[443,887,798,823]" pageId="13" pageNumber="14">
<collectingCountry id="16EA760C670B0417F3C2B025FDD7FCD2" box="[443,600,798,823]" name="South Africa" pageId="13" pageNumber="14">South Africa</collectingCountry>
:
<collectingRegion id="AC39F87E670B0417F01DB025FCF8FCD2" box="[612,887,798,823]" country="South Africa" name="Eastern Cape" pageId="13" pageNumber="14">Eastern Cape Province</collectingRegion>
</emphasis>
,
<location id="6B226047670B0417F1FDB025FC4DFCD2" LSID="urn:lsid:plazi:treatment:E654878A670B0417F2EEB157FB1CF88A:6B226047670B0417F1FDB025FC4DFCD2" box="[900,962,798,823]" country="South Africa" name="Same" pageId="13" pageNumber="14" stateProvince="Eastern Cape">Same</location>
data as holotype (1
<collectionCode id="08ECAE59670B0417F6DCB025FB74FCD2" box="[1189,1275,798,823]" pageId="13" pageNumber="14">ZMHB</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="26E76517670B0417F2BEB079FE57FC46" pageId="13" pageNumber="14" type="diagnosis">
<paragraph id="6E42369C670B0417F2BEB079FE57FC46" blockId="13.[151,1437,726,1903]" pageId="13" pageNumber="14">
<emphasis id="5C89EA8E670B0417F2BEB079FE42FCBE" bold="true" box="[199,461,834,859]" pageId="13" pageNumber="14">Differential diagnosis.</emphasis>
This species is slightly more coarsely punctate dorsally, and has slightly more rounded elytra than
<taxonomicName id="A9FD4D1F670B0417F36FB05CFE01FC9B" box="[278,398,871,894]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="cooperi">
<emphasis id="5C89EA8E670B0417F36FB05CFE01FC9B" box="[278,398,871,894]" italics="true" pageId="13" pageNumber="14">H. cooperi</emphasis>
</taxonomicName>
and
<taxonomicName id="A9FD4D1F670B0417F3BFB05CFDFEFC9A" authorityName="Perkins" authorityYear="2014" box="[454,625,870,895]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="cooperoides">
<emphasis id="5C89EA8E670B0417F3BFB05CFDFEFC9A" box="[454,625,870,895]" italics="true" pageId="13" pageNumber="14">H. cooperoides</emphasis>
</taxonomicName>
(
<figureCitation id="F6C62A19670B0417F0F9B05DFD5FFC9A" box="[640,720,870,895]" captionStart="FIGURE 2" captionStartId="9.[151,249,1978,2000]" captionTargetBox="[277,1309,445,1957]" captionTargetId="figure-116@9.[277,1309,445,1957]" captionTargetPageId="9" captionText="FIGURE 2. Hydraena cooperi, holotype, dorsal and lateral habitus." figureDoi="http://doi.org/10.5281/zenodo.5037603" httpUri="https://zenodo.org/record/5037603/files/figure.png" pageId="13" pageNumber="14">Figs. 2</figureCitation>
,
<figureCitation id="F6C62A19670B0417F0A4B05DFD63FC9A" box="[733,748,870,895]" captionStart="FIGURE 3" captionStartId="11.[151,249,1726,1748]" captionTargetBox="[192,1395,193,1705]" captionTargetId="figure-110@11.[192,1395,193,1705]" captionTargetPageId="11" captionText="FIGURE 3. Hydraena cooperoides, holotype, dorsal and lateral habitus." figureDoi="http://doi.org/10.5281/zenodo.5037605" httpUri="https://zenodo.org/record/5037605/files/figure.png" pageId="13" pageNumber="14">3</figureCitation>
,
<figureCitation id="F6C62A19670B0417F080B05DFC86FC9A" box="[761,777,870,895]" captionStart="FIGURE 6" captionStartId="14.[151,249,1701,1723]" captionTargetBox="[216,1371,179,1679]" captionTargetId="figure-109@14.[216,1371,167,1679]" captionTargetPageId="14" captionText="FIGURE 6. Hydraena capensis, holotype, dorsal and lateral habitus." figureDoi="http://doi.org/10.5281/zenodo.5037613" httpUri="https://zenodo.org/record/5037613/files/figure.png" pageId="13" pageNumber="14">6</figureCitation>
). Reliable determinations will require careful examination of the aedeagi (
<figureCitation id="F6C62A19670B0417F33AB0B1FE21FC46" box="[323,430,906,931]" captionStart="FIGURES 45" captionStartId="12.[151,264,1996,2019]" captionTargetBox="[332,1258,500,1974]" captionTargetId="figure-143@12.[320,1266,481,1976]" captionTargetPageId="12" captionText="FIGURES 45. Aedeagi of holotypes of H. cooperi and H. cooperoides, and aedeagal variants of H. cooperi." figureDoi="http://doi.org/10.5281/zenodo.5037609" httpUri="https://zenodo.org/record/5037609/files/figure.png" pageId="13" pageNumber="14">Figs. 4, 5</figureCitation>
,
<figureCitation id="F6C62A19670B0417F3C3B0B1FE45FC46" box="[442,458,906,931]" captionStart="FIGURES 89" captionStartId="16.[151,264,1629,1652]" captionTargetId="figure-148@16.[271,1315,193,1608]" captionTargetPageId="16" captionText="FIGURES 89. Aedeagi of holotypes." figureDoi="http://doi.org/10.5281/zenodo.5037619" httpUri="https://zenodo.org/record/5037619/files/figure.png" pageId="13" pageNumber="14">8</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="26E76517670B0417F2BEB095FAAEF8E6" pageId="13" pageNumber="14" type="description">
<paragraph id="6E42369C670B0417F2BEB095FAA7FBEB" blockId="13.[151,1437,726,1903]" pageId="13" pageNumber="14">
<emphasis id="5C89EA8E670B0417F2BEB095FEDEFC22" bold="true" box="[199,337,942,967]" pageId="13" pageNumber="14">Description</emphasis>
. Size:
<typeStatus id="B146883E670B0417F3DFB095FD87FC22" box="[422,520,942,967]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.41/0.64; head 0.20/0.36; pronotum 0.34/0.48, PA 0.40, PB 0.47; elytra 0.90/0.64. Dorsum of head dark brown to piceous; pronotum light brown around dark brown to piceous macula; elytra brown; legs light brown; maxillary palpi testaceous.
</paragraph>
<paragraph id="6E42369C670B0417F2BEB720FC9BFB7A" blockId="13.[151,1437,726,1903]" pageId="13" pageNumber="14">Frons punctures ca. 1xef, larger and denser near eyes than medially; interstices weakly shining, 0.51xpd laterally, 0.52xpd medially. Clypeus mcr laterally, very finely sparsely punctulate medially. Mentum weakly concave medially, densely finely punctulate, interstices shining; postmentum punctate, with deep depression on each side. Genae raised, shining, without posterior ridge.</paragraph>
<paragraph id="6E42369C670B0417F2BEB790FAC8FAEE" blockId="13.[151,1437,726,1903]" pageId="13" pageNumber="14">Pronotum laterally slightly emarginate behind middle; anterior margin straight behind eyes, emarginate behind frons, scintilla absent; punctures on disc ca. 12xpd those of frons, interstices shining, 0.52xpd, punctures slightly denser at anterior and posterior; PF1 absent; PF2 very shallow; PF3 shallow; PF4 very shallow or obsolete.</paragraph>
<paragraph id="6E42369C670B0417F2BEB62DFE12FA5A" blockId="13.[151,1437,726,1903]" pageId="13" pageNumber="14">Elytra weakly arcuate laterally; summit of posterior declivity at ca. posterior 1/3; lateral explanate margin moderately wide; on basal 1/3 punctures ca. 1xpd largest pronotal punctures, punctures becoming finer and more widely spaced toward posterior. Intervals not raised, shining, on basal 1/3 ca. 2xpd, as are interstices between punctures of a row. Apices in dorsal aspect weakly separately rounded, in posterior aspect margins forming shallow angle with one another.</paragraph>
<paragraph id="6E42369C670B0417F2BEB6F1FAAEF8E6" blockId="13.[151,1437,726,1903]" pageId="13" pageNumber="14">
Ratios of P2 width and plaque shape (P2/w/l/s) ca. 2/1/1/5. P1 laminate; median carina sinuate in profile. P2 l/ w ca. 2/1, sides becoming parallel toward blunt apex, apex raised slightly above mesoventral intercoxal process. Plaques very small, indistinct, oval, margins not strongly demarcated, located at sides of moderately deep median depression. Metaventrite without midlongitudinal carina between intercoxal process and median depression. AIS width at slightly arcuate posterior margin ca. 2x P2. All tibiae slender. Protibiae very slightly arcuate, meso- and metatibiae straight. Abdominal apex symmetrical; last tergite with apicomedian notch. Female (microslide mount, n=1): last tergite moderately broadly rounded, without apicomedian notch, ca. 36 hooked setae in irregular row, many contiguous, no median gap; gonocoxite not midlongitudinally divided, apical margin moderately broadly rounded, low transverse ridge evident at border of microreticulation; spermatheca simple,
<typeStatus id="B146883E670B0417F6F0B5D0FB36F8E6" box="[1161,1209,1771,1795]" pageId="13" pageNumber="14">type</typeStatus>
C shape.
</paragraph>
</subSubSection>
<subSubSection id="26E76517670B0417F2BEB435FC11F8C2" box="[199,926,1806,1831]" pageId="13" pageNumber="14" type="etymology">
<paragraph id="6E42369C670B0417F2BEB435FC11F8C2" blockId="13.[151,1437,726,1903]" box="[199,926,1806,1831]" pageId="13" pageNumber="14">
<emphasis id="5C89EA8E670B0417F2BEB435FEC1F8C2" bold="true" box="[199,334,1806,1831]" pageId="13" pageNumber="14">Etymology.</emphasis>
Named in reference to the geographical distribution.
</paragraph>
</subSubSection>
<subSubSection id="26E76517670B0417F2BEB409FB1CF88A" pageId="13" pageNumber="14" type="distribution">
<paragraph id="6E42369C670B0417F2BEB409FC48F8AE" blockId="13.[151,1437,726,1903]" box="[199,967,1842,1867]" pageId="13" pageNumber="14">
<emphasis id="5C89EA8E670B0417F2BEB409FEEFF8AE" bold="true" box="[199,352,1842,1867]" pageId="13" pageNumber="14">Distribution.</emphasis>
Currently known only from the
<typeStatus id="B146883E670B0417F0B5B408FD73F8AE" box="[716,764,1843,1867]" pageId="13" pageNumber="14">type</typeStatus>
locality (
<figureCitation id="F6C62A19670B0417F11FB409FC34F8AE" box="[870,955,1842,1867]" captionStart="FIGURES 7778" captionStartId="87.[151,264,1980,2003]" captionTargetBox="[157,1436,193,1900]" captionTargetId="figure-22@87.[157,1442,193,1905]" captionTargetPageId="87" captionText="FIGURES 7778. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5037785" httpUri="https://zenodo.org/record/5037785/files/figure.png" pageId="13" pageNumber="14">Fig. 78</figureCitation>
).
</paragraph>
<paragraph id="6E42369C670B0417F2BEB46DFB1CF88A" blockId="13.[151,1437,726,1903]" box="[199,1171,1878,1903]" pageId="13" pageNumber="14">
<emphasis id="5C89EA8E670B0417F2BEB46DFE89F88A" bold="true" box="[199,262,1878,1903]" pageId="13" pageNumber="14">Note.</emphasis>
I am grateful to Manfred Uhlig for interpreting the locality label of the
<typeStatus id="B146883E670B0417F650B46DFB00F88A" box="[1065,1167,1878,1903]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
</treatment>
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