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<paragraph id="00469A65FFB5B859FF6B40F67C7A0899" blockId="47.[151,1437,1967,2031]" box="[151,1396,1967,1995]" pageId="47" pageNumber="48">
<taxonomicName id="C7F9E1E6FFB5B859FF6B40F67D3F0899" ID-CoL="42XB2" ID-ENA="1330989" authority="Hasan, Islam, Kuramoto, Kurabayashi &amp; Sumida, 2014" authorityName="Hasan, Islam, Kuramoto, Kurabayashi &amp; Sumida" authorityYear="2014" box="[151,1073,1967,1995]" class="Amphibia" family="Microhylidae" genus="Microhyla" isUncertain="true" kingdom="Animalia" order="Anura" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="mukhlesuri">
<emphasis id="328D4677FFB5B859FF6B40F678B00899" bold="true" box="[151,446,1967,1994]" pageId="47" pageNumber="48">
<emphasis id="328D4677FFB5B859FF6B40F678010899" bold="true" box="[151,271,1967,1994]" italics="true" pageId="47" pageNumber="48">Microhyla</emphasis>
cf.
<emphasis id="328D4677FFB5B859FEC640F678B00899" bold="true" box="[314,446,1967,1994]" italics="true" pageId="47" pageNumber="48">mukhlesuri</emphasis>
</emphasis>
<materialsCitation id="B0919038FFB5B859FE3940E87D3F0899" ID-GBIF-Occurrence="4120689394" box="[453,1073,1968,1995]" collectionCode="IABHU" pageId="47" pageNumber="48" specimenCount="1" typeStatus="holotype">Hasan, Islam, Kuramoto, Kurabayashi &amp; Sumida, 2014</materialsCitation>
</taxonomicName>
<emphasis id="328D4677FFB5B859FBB240E97C7A0899" bold="true" box="[1102,1396,1968,1994]" pageId="47" pageNumber="48">Non-native; Established.</emphasis>
</paragraph>
</subSubSection>
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<taxonomicName id="C7F9E1E6FFB5B859FF0B408D7DB508BD" ID-CoL="42XB2" authority="Hasan, Islam, Kuramoto, Kurabayashi &amp; Sumida, 2014: 408" authorityName="Hasan, Islam, Kuramoto, Kurabayashi &amp; Sumida" authorityPageNumber="408" authorityYear="2014" box="[247,1211,2004,2031]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="mukhlesuri">
<emphasis id="328D4677FFB5B859FF0B408D78FF08BD" box="[247,497,2004,2030]" italics="true" pageId="47" pageNumber="48">Microhyla mukhlesuri</emphasis>
Hasan, Islam, Kuramoto, Kurabayashi &amp; Sumida, 2014: 408
</taxonomicName>
.
<typeStatus id="DF4224C7FFB5B859FB35408D7C3808BD" box="[1225,1334,2004,2030]" pageId="47" pageNumber="48" type="holotype">Holotype</typeStatus>
: IABHU
<materialsCitation id="B0919038FFAAB846FEAB47CF78E40F86" ID-GBIF-Occurrence="4120689380" country="Bangladesh" elevation="9" latitude="22.583334" location="Bangladesh" longLatPrecision="1259" longitude="91.916664" pageId="48" pageNumber="49" specimenCount="1">
3956, by original designation.
<typeStatus id="DF4224C7FFAAB846FD5547CE7BEF0FE2" box="[681,737,151,177]" pageId="48" pageNumber="49">Type</typeStatus>
locality: “Raozan, Chittagong (
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,
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,&gt;
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asl
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.),
<collectingCountry id="78EEDAF5FFAAB846FEAB47E278EE0F86" box="[343,480,187,213]" name="Bangladesh" pageId="48" pageNumber="49">Bangladesh</collectingCountry>
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.
</paragraph>
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<subSubSection id="48E3C9EEFFAAB846FF6B478678B60FAA" box="[151,440,223,249]" pageId="48" pageNumber="49" type="vernacular_names">
<paragraph id="00469A65FFAAB846FF6B478678B60FAA" blockId="48.[151,1437,150,2041]" box="[151,440,223,249]" pageId="48" pageNumber="49">Mukhlesurs Chorus Frog</paragraph>
</subSubSection>
<subSubSection id="48E3C9EEFFAAB846FF6B465A78120E4E" box="[151,284,259,285]" pageId="48" pageNumber="49" type="description">
<paragraph id="00469A65FFAAB846FF6B465A78120E4E" blockId="48.[151,1437,150,2041]" box="[151,284,259,285]" pageId="48" pageNumber="49">
(
<figureCitation id="98C286E0FFAAB846FF63465A781A0E4E" box="[159,276,259,285]" captionStart="FIGURE 6" captionStartId="350.[152,255,1943,1967]" captionTargetBox="[154,1433,181,1920]" captionTargetId="figure-34@350.[154,1433,181,1920]" captionTargetPageId="350" captionText="FIGURE 6. A) Microhyla cf. butleri, B) Microhyla cf. heymonsi, C) Microhyla mantheyi, D) Microhyla cf. mukhlesuri, E) Micryletta subaraji, F) Chalcorana labialis, G) Hylarana erythraea, and H) Pulchrana baramica. Photographs by: Kelvin K.P. Lim [AB, G], Alex Figueroa [CD, F], and Law Ing Thong [E, H]." figureDoi="http://doi.org/10.5281/zenodo.7960331" httpUri="https://zenodo.org/record/7960331/files/figure.png" pageId="48" pageNumber="49">Figure 6D</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="48E3C9EEFFAAB846FF6B467F7D060D32" pageId="48" pageNumber="49" type="reference_group">
<paragraph id="00469A65FFAAB846FF6B467F78760E12" blockId="48.[151,1437,150,2041]" box="[151,376,294,321]" pageId="48" pageNumber="49">
<emphasis id="328D4677FFAAB846FF6B467F78760E12" bold="true" box="[151,376,294,321]" pageId="48" pageNumber="49">
<collectingCountry id="78EEDAF5FFAAB846FF6B467F781E0E13" box="[151,272,294,320]" name="Singapore" pageId="48" pageNumber="49">Singapore</collectingCountry>
records.
</emphasis>
</paragraph>
<paragraph id="00469A65FFAAB846FF0B46127D1E0D6E" blockId="48.[151,1437,150,2041]" pageId="48" pageNumber="49">
<taxonomicName id="C7F9E1E6FFAAB846FF0B461278C80E36" ID-CoL="73GRF" authorityName="Boulenger" authorityYear="1884" box="[247,454,331,357]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="fissipes">
<emphasis id="328D4677FFAAB846FF0B461278C80E36" box="[247,454,331,357]" italics="true" pageId="48" pageNumber="49">Microhyla fissipes</emphasis>
</taxonomicName>
(non Boulenger, 1884)—N.
<bibRefCitation id="6468E794FFAAB846FCEC46127AAB0E36" author="Lim, K. K. P. &amp; Lim, N. T. - L." box="[784,933,331,357]" pageId="48" pageNumber="49" pagination="107" refId="ref326554" refString="Lim, K. K. P. &amp; Lim, N. T. - L. (2006) Geographic distribution: Luperosaurus browni. Herpetological Review, 37 (1), 107." type="journal article" year="2006">Lim, 2006: 6</bibRefCitation>
(Pulau Tekong).—N.T.-L.
<bibRefCitation id="6468E794FFAAB846FB274612788F0EDA" author="Lim, K. K. P. &amp; Lim, N. T. - L." pageId="48" pageNumber="49" pagination="107" refId="ref326554" refString="Lim, K. K. P. &amp; Lim, N. T. - L. (2006) Geographic distribution: Luperosaurus browni. Herpetological Review, 37 (1), 107." type="journal article" year="2006">
Lim
<emphasis id="328D4677FFAAB846FAEF46127C430E36" box="[1299,1357,331,357]" italics="true" pageId="48" pageNumber="49">et al.</emphasis>
, 2006: 101
</bibRefCitation>
(Pulau Tekong).—N. Baker &amp; K.P. Lim, 2008: 65, 165.—D.C.J. Yeo &amp; C.S.W. Chia, 2010: 32.—T.M. Leong &amp; K.K.P.
<bibRefCitation id="6468E794FFAAB846FD6F46CA7A4C0EFE" author="Lim, K. K. P. &amp; Leong, T. M. &amp; Lim, F. L. K." box="[659,834,403,429]" pageId="48" pageNumber="49" pagination="143 - 156" refId="ref326919" refString="Lim, K. K. P., Leong, T. M. &amp; Lim, F. L. K. (2011) The king cobra, Ophiophagus hannah (Cantor) in Singapore (Reptilia: Squamata: Elapidae). Nature in Singapore, 4, 143 - 156." type="journal article" year="2011">Lim, 2011: 140</bibRefCitation>
.—N. Baker &amp; K.P. Lim, 2012: 65, 165.—T.H. Ng &amp; D.C.J. Yeo, 2012: 95, 9697.—
<bibRefCitation id="6468E794FFAAB846FD4846EE7AF40E82" author="Manthey, U. &amp; Denzer, W." box="[692,1018,439,465]" pageId="48" pageNumber="49" pagination="3 - 21" refId="ref329158" refString="Manthey, U. &amp; Denzer, W. (2014) S ¸ dostasiatische anuren im fokus: spezies der gattung Microhyla Tschudi, 1838 (Anura: Microhylidae: Microhylinae). Sauria, 36 (3), 3 - 21." type="journal article" year="2014">Manthey &amp; Denzer, 2014: 11</bibRefCitation>
.—
<bibRefCitation id="6468E794FFAAB846FBE046EE7C170E82" author="Hasan, M. &amp; Sarker, M. A. R. &amp; Kurabayashi, A. &amp; Kuramoto, M. &amp; Sumida, M." box="[1052,1305,438,465]" pageId="48" pageNumber="49" pagination="63 - 80" refId="ref312913" refString="Hasan, M., Sarker, M. A. R., Kurabayashi, A., Kuramoto, M. &amp; Sumida, M. (2015) Genetic variation, advertisement call, and morphometry of Microhyla nilphamariensis from Bangladesh. Philippine Journal of Systematic Biology, 9, 63 - 80." type="journal article" year="2015">
Hasan
<emphasis id="328D4677FFAAB846FB9046EE7DAB0E82" box="[1132,1189,439,465]" italics="true" pageId="48" pageNumber="49">et al.</emphasis>
, 2015: 72
</bibRefCitation>
.—I.S. Law, 2015: 30 (Nanyang Technological University).—I.S. Law &amp;
<bibRefCitation id="6468E794FFAAB846FC0446827C0A0EA6" author="Groenewoud, D." box="[1016,1284,474,501]" pageId="48" pageNumber="49" pagination="42" refId="ref311492" refString="Groenewoud, D. (2015 b) Singapore bent-toed gecko swimming underwater. Singapore Biodiversity Records, 2015, 42." type="journal article" year="2015">Groenewoud, 2015b: 63</bibRefCitation>
.—R. Subaraj
<emphasis id="328D4677FFAAB846FEAB46A678810D4A" box="[343,399,511,537]" italics="true" pageId="48" pageNumber="49">et al.</emphasis>
, 2016: 30 (Lorong Halus Wetland).—K.K.P.
<bibRefCitation id="6468E794FFAAB846FC7B46A67D7C0D4A" author="Lim, K. K. P. &amp; Chua, M. A. H. &amp; Lim, N. T. - L." box="[903,1138,511,537]" pageId="48" pageNumber="49" pagination="165 - 198" refId="ref327018" refString="Lim, K. K. P., Chua, M. A. H. &amp; Lim, N. T. - L. (2016) Freshwater fishes, terrestrial herpetofauna and mammals of Pulau Tekong, Singapore. Nature in Singapore, 9, 165 - 198." type="journal article" year="2016">
Lim
<emphasis id="328D4677FFAAB846FC4146A67AFA0D4A" box="[957,1012,511,537]" italics="true" pageId="48" pageNumber="49">et al.</emphasis>
, 2016: 174
</bibRefCitation>
(Pulau Tekong).—D.T.
<bibRefCitation id="6468E794FFAAB846FA8346A67B1C0D6E" author="Le, D. T. &amp; Nguyen, L. P. &amp; Phan, C. T. &amp; Nguyen, T. Q. &amp; Phan, T. Q. &amp; Dang, P. H. &amp; Ziegler, T." pageId="48" pageNumber="49" pagination="317 - 324" refId="ref322324" refString="Le, D. T., Nguyen, L. P., Phan, C. T., Nguyen, T. Q., Phan, T. Q., Dang, P. H. &amp; Ziegler, T. (2021) New records of frogs from Quang Nam Province, central Vietnam. Herpetology Notes, 14, 317 - 324." type="journal article" year="2021">
Le
<emphasis id="328D4677FFAAB846FEAB457A78810D6E" box="[343,399,547,573]" italics="true" pageId="48" pageNumber="49">et al.</emphasis>
, 2021: 319
</bibRefCitation>
.—B.C. Ng, 2021: 136 (Pang Sua Woodland).
</paragraph>
<paragraph id="00469A65FFAAB846FF0B451E7D060D32" blockId="48.[151,1437,150,2041]" box="[247,1032,582,609]" pageId="48" pageNumber="49">
<taxonomicName id="C7F9E1E6FFAAB846FF0B451E7B1C0D32" ID-CoL="42XB2" authorityName="Hasan, Islam, Kuramoto, Kurabayashi &amp; Sumida" authorityYear="2014" box="[247,530,583,609]" class="Amphibia" family="Microhylidae" genus="Microhyla" isUncertain="true" kingdom="Animalia" order="Anura" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="mukhlesuri">
<emphasis id="328D4677FFAAB846FF0B451E78650D32" box="[247,363,583,609]" italics="true" pageId="48" pageNumber="49">Microhyla</emphasis>
cf.
<emphasis id="328D4677FFAAB846FE6A451E7B1C0D32" box="[406,530,583,609]" italics="true" pageId="48" pageNumber="49">mukhlesuri</emphasis>
</taxonomicName>
—Sankar
<emphasis id="328D4677FFAAB846FD7F451E7BB50D32" box="[643,699,583,609]" italics="true" pageId="48" pageNumber="49">et al.</emphasis>
, 2022: 465 (Kranji Marshes).
</paragraph>
</subSubSection>
<subSubSection id="48E3C9EEFFAAB846FF6B453378DB09C2" pageId="48" pageNumber="49" type="discussion">
<paragraph id="00469A65FFAAB846FF6B45337BB20AE9" blockId="48.[151,1437,150,2041]" pageId="48" pageNumber="49">
<emphasis id="328D4677FFAAB846FF6B453378050DD7" bold="true" box="[151,267,618,644]" pageId="48" pageNumber="49">Remarks.</emphasis>
In a fairly recent publication, Yuan
<emphasis id="328D4677FFAAB846FD5D45327BD40DD7" box="[673,730,618,644]" italics="true" pageId="48" pageNumber="49">et al.</emphasis>
(2016) demonstrated that the Red River in
<collectingCountry id="78EEDAF5FFAAB846FB3845337C090DD7" box="[1220,1287,618,644]" name="China" pageId="48" pageNumber="49">China</collectingCountry>
and
<collectingCountry id="78EEDAF5FFAAB846FAC145337C920DD7" box="[1341,1436,618,644]" name="Vietnam" pageId="48" pageNumber="49">Vietnam</collectingCountry>
acts as a biogeographic barrier for
<taxonomicName id="C7F9E1E6FFAAB846FD7845D67BF30DFB" authorityName="Boulenger" authorityYear="1884" box="[644,765,654,680]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="fissipes">
<emphasis id="328D4677FFAAB846FD7845D67BF30DFB" box="[644,765,654,680]" italics="true" pageId="48" pageNumber="49">M. fissipes</emphasis>
</taxonomicName>
, and redescribed populations south of the Red River as
<taxonomicName id="C7F9E1E6FFAAB846FA8245D6787D0D9F" authorityName="Hasan, Islam, Kuramoto, Kurabayashi &amp; Sumida" authorityYear="2014" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="mukhlesuri">
<emphasis id="328D4677FFAAB846FA8245D6787D0D9F" italics="true" pageId="48" pageNumber="49">M. mukhlesuri</emphasis>
</taxonomicName>
. Prior to this,
<bibRefCitation id="6468E794FFAAB846FDDE45EB7A050D9E" author="Matsui, M. &amp; Ito, H. &amp; Shimada, T. &amp; Ota, H. &amp; Saidapur, S. K. &amp; Khonsue, W. &amp; Tanaka-Ueno, T. &amp; Wu, G. F." box="[546,779,690,717]" pageId="48" pageNumber="49" pagination="489 - 496" refId="ref329723" refString="Matsui, M., Ito, H., Shimada, T., Ota, H., Saidapur, S. K., Khonsue, W., Tanaka-Ueno, T. &amp; Wu, G. F. (2005) Taxonomic relationships within the Pan-Oriental narrow-mouth toad Microhyla ornata as revealed by mtDNA analysis (Amphibia, Anura, Microhylidae). Zoological Science, 22 (4), 489 - 496. https: // doi. org / 10.2108 / zsj. 22.489" type="journal article" year="2005">
Matsui
<emphasis id="328D4677FFAAB846FD8645EA7BB80D9F" box="[634,694,690,716]" italics="true" pageId="48" pageNumber="49">et al.</emphasis>
(2005)
</bibRefCitation>
redescribed
<taxonomicName id="C7F9E1E6FFAAB846FC5D45EA7D1A0D9F" baseAuthorityName="Dumeril &amp; Bibron" baseAuthorityYear="1841" box="[929,1044,691,716]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="ornata">
<emphasis id="328D4677FFAAB846FC5D45EA7D1A0D9F" box="[929,1044,691,716]" italics="true" pageId="48" pageNumber="49">M. ornata</emphasis>
</taxonomicName>
from
<collectingCountry id="78EEDAF5FFAAB846FBA345EB7DBF0D9F" box="[1119,1201,690,716]" name="Taiwan" pageId="48" pageNumber="49">Taiwan</collectingCountry>
and southern
<collectingCountry id="78EEDAF5FFAAB846FAA545EB7C920D9F" box="[1369,1436,690,716]" name="China" pageId="48" pageNumber="49">China</collectingCountry>
south to northern Peninsular
<collectingCountry id="78EEDAF5FFAAB846FDC0458C7BAD0DBC" box="[572,675,725,751]" name="Malaysia" pageId="48" pageNumber="49">Malaysia</collectingCountry>
as
<taxonomicName id="C7F9E1E6FFAAB846FD35458F7A4E0DA3" authorityName="Boulenger" authorityYear="1884" box="[713,832,726,752]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="fissipes">
<emphasis id="328D4677FFAAB846FD35458F7A4E0DA3" box="[713,832,726,752]" italics="true" pageId="48" pageNumber="49">M. fissipes</emphasis>
</taxonomicName>
. Thus,
<taxonomicName id="C7F9E1E6FFAAB846FC69458F7D020DA3" authorityName="Boulenger" authorityYear="1884" box="[917,1036,726,752]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="fissipes">
<emphasis id="328D4677FFAAB846FC69458F7D020DA3" box="[917,1036,726,752]" italics="true" pageId="48" pageNumber="49">M. fissipes</emphasis>
</taxonomicName>
from northern Peninsular
<collectingCountry id="78EEDAF5FFAAB846FACA458C7C930DBC" box="[1334,1437,725,751]" name="Malaysia" pageId="48" pageNumber="49">Malaysia</collectingCountry>
correspond to
<emphasis id="328D4677FFAAB846FE6B45A37B4E0C40" box="[407,576,761,787]" italics="true" pageId="48" pageNumber="49">
<taxonomicName id="C7F9E1E6FFAAB846FE6B45A37B320C40" authorityName="Hasan, Islam, Kuramoto, Kurabayashi &amp; Sumida" authorityYear="2014" box="[407,572,761,787]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="mukhlesuri">M. mukhlesuri</taxonomicName>
.
</emphasis>
In 2004, frogs identified as
<taxonomicName id="C7F9E1E6FFAAB846FC8045A37AFD0C40" authorityName="Boulenger" authorityYear="1884" box="[892,1011,761,787]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="fissipes">
<emphasis id="328D4677FFAAB846FC8045A37AFD0C40" box="[892,1011,761,787]" italics="true" pageId="48" pageNumber="49">M. fissipes</emphasis>
</taxonomicName>
were first recorded in Endau-Rompin National Park and then again in 2005 (Wood
<emphasis id="328D4677FFAAB846FCF544477A4B0C64" box="[777,837,797,823]" italics="true" pageId="48" pageNumber="49">et al.</emphasis>
2008). Unknown in the southern part of Peninsular
<collectingCountry id="78EEDAF5FFAAB846FF0B4418786E0C08" box="[247,352,833,859]" name="Malaysia" pageId="48" pageNumber="49">Malaysia</collectingCountry>
,
<taxonomicName id="C7F9E1E6FFAAB846FE91441B78E80C08" authorityName="Boulenger" authorityYear="1884" box="[365,486,833,859]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="fissipes">
<emphasis id="328D4677FFAAB846FE91441B78E80C08" box="[365,486,833,859]" italics="true" pageId="48" pageNumber="49">M. fissipes</emphasis>
</taxonomicName>
was regarded as introduced (
<bibRefCitation id="6468E794FFAAB846FCC944187D460C08" author="Daicus, B. &amp; Hashim, R." box="[821,1096,833,859]" pageId="48" pageNumber="49" pagination="65 - 72" refId="ref300547" refString="Daicus, B. &amp; Hashim, R. (2004) Herpetofauna of the western region of Endau-Rompin, Johore, Peninsular Malaysia. Malaysian Journal of Science, 23 (1), 65 - 72." type="journal article" year="2004">Daicus &amp; Hashim 2004</bibRefCitation>
). Likewise, on
<date id="7447BCA5FFAAB846FB0A44187C920C0F" box="[1270,1436,833,860]" pageId="48" pageNumber="49" value="2005-08-03">3 August 2005</date>
<specimenCount id="16FF51ECFFAAB846FF0B443D786A0C2D" box="[247,356,868,894]" pageId="48" pageNumber="49" type="adult">five adult</specimenCount>
males and seven tadpoles of
<taxonomicName id="C7F9E1E6FFAAB846FD41443C7A390C2C" authorityName="Boulenger" authorityYear="1884" box="[701,823,869,895]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="fissipes">
<emphasis id="328D4677FFAAB846FD41443C7A390C2C" box="[701,823,869,895]" italics="true" pageId="48" pageNumber="49">M. fissipes</emphasis>
</taxonomicName>
were collected from Pulau Tekong (
<bibRefCitation id="6468E794FFAAB846FB1C443D7C960C2D" author="Lim, K. K. P. &amp; Lim, N. T. - L." box="[1248,1432,868,894]" pageId="48" pageNumber="49" pagination="107" refId="ref326554" refString="Lim, K. K. P. &amp; Lim, N. T. - L. (2006) Geographic distribution: Luperosaurus browni. Herpetological Review, 37 (1), 107." type="journal article" year="2006">
Lim
<emphasis id="328D4677FFAAB846FAE4443C7C5D0C2D" box="[1304,1363,868,894]" italics="true" pageId="48" pageNumber="49">et al.</emphasis>
2006
</bibRefCitation>
;
<bibRefCitation id="6468E794FFAAB846FF0B44D178BF0CF1" author="Lim, K. K. P. &amp; Chua, M. A. H. &amp; Lim, N. T. - L." box="[247,433,904,930]" pageId="48" pageNumber="49" pagination="165 - 198" refId="ref327018" refString="Lim, K. K. P., Chua, M. A. H. &amp; Lim, N. T. - L. (2016) Freshwater fishes, terrestrial herpetofauna and mammals of Pulau Tekong, Singapore. Nature in Singapore, 9, 165 - 198." type="journal article" year="2016">
Lim
<emphasis id="328D4677FFAAB846FECC44D078650CF1" box="[304,363,904,930]" italics="true" pageId="48" pageNumber="49">et al.</emphasis>
2016
</bibRefCitation>
). Ten years later, on
<date id="7447BCA5FFAAB846FD5944D17A7A0CF0" box="[677,884,904,931]" pageId="48" pageNumber="49" value="2015-02-14">14 February 2015</date>
, the first record for
<collectingCountry id="78EEDAF5FFAAB846FB9D44D17C2E0CF1" box="[1121,1312,904,930]" name="Singapore Island" pageId="48" pageNumber="49">Singapore Island</collectingCountry>
was made at NTU (Law 2015). Nowadays, this species is well-established and is found throughout
<collectingCountry id="78EEDAF5FFAAB846FB0044F57C600C95" box="[1276,1390,940,966]" name="Singapore" pageId="48" pageNumber="49">Singapore</collectingCountry>
(A. Figueroa, unpubl. data). The extent of its distribution in southern Peninsular
<collectingCountry id="78EEDAF5FFAAB846FBA244897DCB0CB9" box="[1118,1221,976,1002]" name="Malaysia" pageId="48" pageNumber="49">Malaysia</collectingCountry>
remains unknown, and as of yet, the populations in southern Peninsular
<collectingCountry id="78EEDAF5FFAAB846FCAE44AA7AB70B5E" box="[850,953,1011,1037]" name="Malaysia" pageId="48" pageNumber="49">Malaysia</collectingCountry>
and
<collectingCountry id="78EEDAF5FFAAB846FC0E44AA7D6A0B5E" box="[1010,1124,1011,1037]" name="Singapore" pageId="48" pageNumber="49">Singapore</collectingCountry>
have not been examined to determine if they represent
<taxonomicName id="C7F9E1E6FFAAB846FDD243417BA80B62" authorityName="Boulenger" authorityYear="1884" box="[558,678,1047,1073]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="fissipes">
<emphasis id="328D4677FFAAB846FDD243417BA80B62" box="[558,678,1047,1073]" italics="true" pageId="48" pageNumber="49">M. fissipes</emphasis>
</taxonomicName>
,
<taxonomicName id="C7F9E1E6FFAAB846FD4943417A590B62" authorityName="Hasan, Islam, Kuramoto, Kurabayashi &amp; Sumida" authorityYear="2014" box="[693,855,1047,1073]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="mukhlesuri">
<emphasis id="328D4677FFAAB846FD4943417A590B62" box="[693,855,1047,1073]" italics="true" pageId="48" pageNumber="49">M. mukhlesuri</emphasis>
</taxonomicName>
, or some other member of the
<taxonomicName id="C7F9E1E6FFAAB846FB4943417C230B62" authorityName="Boulenger" authorityYear="1884" box="[1205,1325,1047,1073]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="fissipes">
<emphasis id="328D4677FFAAB846FB4943417C230B62" box="[1205,1325,1047,1073]" italics="true" pageId="48" pageNumber="49">M. fissipes</emphasis>
</taxonomicName>
complex. However, preliminary field identification of specimens in
<collectingCountry id="78EEDAF5FFAAB846FC8743627AE30B06" box="[891,1005,1083,1109]" name="Singapore" pageId="48" pageNumber="49">Singapore</collectingCountry>
key out to
<taxonomicName id="C7F9E1E6FFAAB846FB9B43657C060B06" authorityName="Hasan, Islam, Kuramoto, Kurabayashi &amp; Sumida" authorityYear="2014" box="[1127,1288,1083,1109]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="mukhlesuri">
<emphasis id="328D4677FFAAB846FB9B43657C060B06" box="[1127,1288,1083,1109]" italics="true" pageId="48" pageNumber="49">M. mukhlesuri</emphasis>
</taxonomicName>
(A. Figueroa pers. obs.). Until the taxonomic status of these populations is established, it is difficult to trace the species introduction pathway. However, we envision that
<taxonomicName id="C7F9E1E6FFAAB846FCC343DA7AEB0BCF" authorityName="Hasan, Islam, Kuramoto, Kurabayashi &amp; Sumida" authorityYear="2014" box="[831,997,1154,1180]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="mukhlesuri">
<emphasis id="328D4677FFAAB846FCC343DA7AEB0BCF" box="[831,997,1154,1180]" italics="true" pageId="48" pageNumber="49">M. mukhlesuri</emphasis>
</taxonomicName>
made its way to southern Peninsular
<collectingCountry id="78EEDAF5FFAAB846FF0B43FF78500B93" box="[247,350,1190,1216]" name="Malaysia" pageId="48" pageNumber="49">Malaysia</collectingCountry>
from the north, then to Pulau Tekong where it may have reached as a stowaway via ornamental plants or cargo (Ng &amp; Yeo 2012). Since Pulau Tekong currently serves as a military base,
<taxonomicName id="C7F9E1E6FFAAB846FB3B43927C680BB7" authorityName="Hasan, Islam, Kuramoto, Kurabayashi &amp; Sumida" authorityYear="2014" box="[1223,1382,1226,1252]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="mukhlesuri">
<emphasis id="328D4677FFAAB846FB3B43927C680BB7" box="[1223,1382,1226,1252]" italics="true" pageId="48" pageNumber="49">M. mukhlesuri</emphasis>
</taxonomicName>
may have reached
<collectingCountry id="78EEDAF5FFAAB846FE7343B47B450A54" box="[399,587,1261,1287]" name="Singapore Island" pageId="48" pageNumber="49">Singapore Island</collectingCountry>
by hitchhiking on military personnel frequenting the island or paraphernalia being transported back and forth. As such, we conditionally recognise the populations in
<collectingCountry id="78EEDAF5FFAAB846FB1942487C590A78" box="[1253,1367,1297,1323]" name="Singapore" pageId="48" pageNumber="49">Singapore</collectingCountry>
as
<taxonomicName id="C7F9E1E6FFAAB846FA82424B78940A1C" authorityName="Hasan, Islam, Kuramoto, Kurabayashi &amp; Sumida" authorityYear="2014" class="Amphibia" family="Microhylidae" genus="Microhyla" isUncertain="true" kingdom="Animalia" order="Anura" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="mukhlesuri">
<emphasis id="328D4677FFAAB846FA82424B7C930A78" box="[1406,1437,1298,1323]" italics="true" pageId="48" pageNumber="49">M.</emphasis>
cf.
<emphasis id="328D4677FFAAB846FEE2426C78940A1C" box="[286,410,1333,1359]" italics="true" pageId="48" pageNumber="49">mukhlesuri</emphasis>
</taxonomicName>
. Similar to the other three species of
<taxonomicName id="C7F9E1E6FFAAB846FCAA426C7AC40A1C" box="[854,970,1333,1359]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="48" pageNumber="49" phylum="Chordata" rank="genus">
<emphasis id="328D4677FFAAB846FCAA426C7AC40A1C" box="[854,970,1333,1359]" italics="true" pageId="48" pageNumber="49">Microhyla</emphasis>
</taxonomicName>
in
<collectingCountry id="78EEDAF5FFAAB846FC0F426C7D690A1C" box="[1011,1127,1333,1359]" name="Singapore" pageId="48" pageNumber="49">Singapore</collectingCountry>
, cryptic diversity was also uncovered in
<taxonomicName id="C7F9E1E6FFAAB846FE6D42037A380A20" authority="(Gorin et al. 2020)" baseAuthorityName="Gorin" baseAuthorityYear="2020" box="[401,822,1369,1395]" class="Amphibia" family="Microhylidae" genus="Microhyla" isUncertain="true" kingdom="Animalia" order="Anura" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="mukhlesuri">
<emphasis id="328D4677FFAAB846FE6D420378BE0A20" box="[401,432,1370,1395]" italics="true" pageId="48" pageNumber="49">M.</emphasis>
cf.
<emphasis id="328D4677FFAAB846FE2742007B590A20" box="[475,599,1369,1395]" italics="true" pageId="48" pageNumber="49">mukhlesuri</emphasis>
(
<bibRefCitation id="6468E794FFAAB846FD9442007A200A20" author="Gorin, V. A. &amp; Solovyeva, E. N. &amp; Hasan, M. &amp; Okamiya, H. &amp; Karunarathna, D. M. S. S. &amp; Pawangkhanant, P. &amp; de Silva, A. &amp; Juthong, W. &amp; Milto, K. D. &amp; Nguyen, L. T. &amp; Suwannapoom, C. &amp; Haas, A. &amp; Bickford, D. P. &amp; Das, I. &amp; Poyarkov, N. A." box="[616,814,1369,1395]" pageId="48" pageNumber="49" pagination="9411" refId="ref308218" refString="Gorin, V. A., Solovyeva, E. N., Hasan, M., Okamiya, H., Karunarathna, D. M. S. S., Pawangkhanant, P., de Silva, A., Juthong, W., Milto, K. D., Nguyen, L. T., Suwannapoom, C., Haas, A., Bickford, D. P., Das, I. &amp; Poyarkov, N. A. (2020) A little frog leaps a long way: compounded colonizations of the Indian Subcontinent discovered in the tiny Oriental frog genus Microhyla (Amphibia: Microhylidae). PeerJ, 8, e 9411. https: // doi. org / 10.7717 / peerj. 9411" type="journal article" year="2020">
Gorin
<emphasis id="328D4677FFAAB846FD4D42037BE40A20" box="[689,746,1369,1395]" italics="true" pageId="48" pageNumber="49">et al.</emphasis>
2020
</bibRefCitation>
)
</taxonomicName>
. Thus, if the populations in
<collectingCountry id="78EEDAF5FFAAB846FB8C42007DEC0A20" box="[1136,1250,1369,1395]" name="Singapore" pageId="48" pageNumber="49">Singapore</collectingCountry>
do belong to
<taxonomicName id="C7F9E1E6FFAAB846FA824203787D0AC5" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="mukhlesuri">
<emphasis id="328D4677FFAAB846FA824203787D0AC5" italics="true" pageId="48" pageNumber="49">M. mukhlesuri</emphasis>
</taxonomicName>
, then future investigations into
<taxonomicName id="C7F9E1E6FFAAB846FD2442247A770AC5" box="[728,889,1404,1430]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="mukhlesuri">
<emphasis id="328D4677FFAAB846FD2442247A770AC5" box="[728,889,1404,1430]" italics="true" pageId="48" pageNumber="49">M. mukhlesuri</emphasis>
</taxonomicName>
may show yet a different taxonomic designation for the species that colonised
<collectingCountry id="78EEDAF5FFAAB846FDBF42F97BB60AE9" box="[579,696,1440,1466]" name="Singapore" pageId="48" pageNumber="49">Singapore</collectingCountry>
.
</paragraph>
<paragraph id="00469A65FFAAB846FEAB429D78DB09C2" blockId="48.[151,1437,150,2041]" pageId="48" pageNumber="49">
Given its small size, secretive lifestyle within leaf-litter, and superficial resemblance to native
<taxonomicName id="C7F9E1E6FFAAB846FA82429C78690951" class="Amphibia" family="Microhylidae" genus="Microhyla" isUncertain="true" kingdom="Animalia" order="Anura" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="butleri">
<emphasis id="328D4677FFAAB846FA82429C7C930A8D" box="[1406,1437,1477,1502]" italics="true" pageId="48" pageNumber="49">M.</emphasis>
cf.
<emphasis id="328D4677FFAAB846FEE042B178690951" box="[284,359,1512,1538]" italics="true" pageId="48" pageNumber="49">butleri</emphasis>
</taxonomicName>
,
<taxonomicName id="C7F9E1E6FFAAB846FE8942B17B350951" box="[373,571,1512,1538]" class="Amphibia" family="Microhylidae" genus="Microhyla" isUncertain="true" kingdom="Animalia" order="Anura" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="mukhlesuri">
<emphasis id="328D4677FFAAB846FE8942B1789A0952" box="[373,404,1512,1537]" italics="true" pageId="48" pageNumber="49">M.</emphasis>
cf.
<emphasis id="328D4677FFAAB846FE4342B17B350951" box="[447,571,1512,1538]" italics="true" pageId="48" pageNumber="49">mukhlesuri</emphasis>
</taxonomicName>
may have gone undetected for some time. At the time of Ng &amp; Yeos (2012) publication,
<taxonomicName id="C7F9E1E6FFAAB846FE7941557B270976" box="[389,553,1547,1573]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="mukhlesuri">
<emphasis id="328D4677FFAAB846FE7941557B270976" box="[389,553,1547,1573]" italics="true" pageId="48" pageNumber="49">M. mukhlesuri</emphasis>
</taxonomicName>
was only known from Pulau Tekong; thus, the authors believed it would not compete with native microhylids. However, given that
<taxonomicName id="C7F9E1E6FFAAB846FC9F41697D27091A" box="[867,1065,1583,1609]" class="Amphibia" family="Microhylidae" genus="Microhyla" isUncertain="true" kingdom="Animalia" order="Anura" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="mukhlesuri">
<emphasis id="328D4677FFAAB846FC9F41697A8C091A" box="[867,898,1584,1609]" italics="true" pageId="48" pageNumber="49">M.</emphasis>
cf.
<emphasis id="328D4677FFAAB846FC5141767D27091A" box="[941,1065,1583,1609]" italics="true" pageId="48" pageNumber="49">mukhlesuri</emphasis>
</taxonomicName>
is now widespread and common throughout
<collectingCountry id="78EEDAF5FFAAB846FE8B410A78E5093E" box="[375,491,1619,1645]" name="Singapore" pageId="48" pageNumber="49">Singapore</collectingCountry>
, and that at many locales it is the dominant microhylid, we designate
<taxonomicName id="C7F9E1E6FFAAB846FB27410D7C92093E" box="[1243,1436,1619,1645]" class="Amphibia" family="Microhylidae" genus="Microhyla" isUncertain="true" kingdom="Animalia" order="Anura" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="mukhlesuri">
<emphasis id="328D4677FFAAB846FB27410D7DF4093E" box="[1243,1274,1620,1645]" italics="true" pageId="48" pageNumber="49">M.</emphasis>
cf.
<emphasis id="328D4677FFAAB846FADC410A7C92093E" box="[1312,1436,1619,1645]" italics="true" pageId="48" pageNumber="49">mukhlesuri</emphasis>
</taxonomicName>
an invasive species.
</paragraph>
</subSubSection>
<subSubSection id="48E3C9EEFFAAB846FF6B41C37B2509E6" box="[151,555,1690,1717]" pageId="48" pageNumber="49" type="distribution">
<paragraph id="00469A65FFAAB846FF6B41C37B2509E6" blockId="48.[151,1437,150,2041]" box="[151,555,1690,1717]" pageId="48" pageNumber="49">
<emphasis id="328D4677FFAAB846FF6B41C3782409E7" bold="true" box="[151,298,1690,1716]" pageId="48" pageNumber="49">Occurrence.</emphasis>
Ubiquitous. Common.
</paragraph>
</subSubSection>
<subSubSection id="48E3C9EEFFAAB846FF6B41E77B8C0872" pageId="48" pageNumber="49" type="conservation">
<paragraph id="00469A65FFAAB846FF6B41E77BB3098A" blockId="48.[151,1437,150,2041]" box="[151,701,1726,1753]" pageId="48" pageNumber="49">
<emphasis id="328D4677FFAAB846FF6B41E77B0A098A" bold="true" box="[151,516,1726,1753]" pageId="48" pageNumber="49">
<collectingCountry id="78EEDAF5FFAAB846FF6B41E7781E098B" box="[151,272,1726,1752]" name="Singapore" pageId="48" pageNumber="49">Singapore</collectingCountry>
conservation status.
</emphasis>
Not Applicable.
</paragraph>
<paragraph id="00469A65FFAAB846FF6B41BB7BC409AE" blockId="48.[151,1437,150,2041]" box="[151,714,1762,1789]" pageId="48" pageNumber="49">
<emphasis id="328D4677FFAAB846FF6B41BB78AF09AF" bold="true" box="[151,417,1762,1788]" pageId="48" pageNumber="49">Conservation priority.</emphasis>
None, non-native species.
</paragraph>
<paragraph id="00469A65FFAAB846FF6B405F7B8C0872" blockId="48.[151,1437,150,2041]" box="[151,642,1798,1825]" pageId="48" pageNumber="49">
<emphasis id="328D4677FFAAB846FF6B405F78DC0872" bold="true" box="[151,466,1798,1825]" pageId="48" pageNumber="49">IUCN conservation status.</emphasis>
<collectingRegion id="C23D5487FFAAB846FE25405E7B0D0872" box="[473,515,1799,1825]" country="Maldives" name="Uthuru" pageId="48" pageNumber="49">Not</collectingRegion>
Evaluated.
</paragraph>
</subSubSection>
<subSubSection id="48E3C9EEFFAAB846FF6B40737D1908AA" pageId="48" pageNumber="49" type="materials_examined">
<paragraph id="00469A65FFAAB846FF6B4073789108DE" blockId="48.[151,1437,150,2041]" pageId="48" pageNumber="49">
<emphasis id="328D4677FFAAB846FF6B40737A5D0816" bold="true" box="[151,851,1834,1861]" pageId="48" pageNumber="49">
<collectionCode id="66E802A0FFAAB846FF6B407378160817" box="[151,280,1834,1860]" pageId="48" pageNumber="49">LKCNHM</collectionCode>
&amp;
<collectionCode id="66E802A0FFAAB846FEC0407278A40816" box="[316,426,1835,1861]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="48" pageNumber="49" type="Museum">NHMUK</collectionCode>
Museum specimens. Pulau Tekong:
</emphasis>
<collectionCode id="66E802A0FFAAB846FCA540737A9F0816" box="[857,913,1834,1861]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="48" pageNumber="49">ZRC</collectionCode>
.1.11543
<collectionCode id="66E802A0FFAAB846FC0340737D390816" box="[1023,1079,1834,1861]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="48" pageNumber="49">ZRC</collectionCode>
.
<materialsCitation id="B0919038FFAAB846FBC640737895083A" ID-GBIF-Occurrence="4120689350" collectingDate="2005-08-03" collectionCode="WNP" pageId="48" pageNumber="49" specimenCount="1">
1.11554 (
<date id="7447BCA5FFAAB846FB5940737C4C0816" box="[1189,1346,1834,1861]" pageId="48" pageNumber="49" value="2005-08-03">
<collectingDate id="6403454DFFAAB846FB5940737C4C0816" box="[1189,1346,1834,1861]" pageId="48" pageNumber="49" value="2005-08-03">03-Aug-2005</collectingDate>
</date>
);
<emphasis id="328D4677FFAAB846FAA840727895083A" bold="true" pageId="48" pageNumber="49">
Venus Drive [
<collectionCode id="66E802A0FFAAB846FEB14016789F083A" box="[333,401,1871,1897]" pageId="48" pageNumber="49">WNP</collectionCode>
]
</emphasis>
</materialsCitation>
<emphasis id="328D4677FFAAB846FE67401678AD083A" bold="true" box="[411,419,1871,1897]" pageId="48" pageNumber="49">:</emphasis>
<collectionCode id="66E802A0FFAAB846FE52401678E9083A" box="[430,487,1871,1897]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="48" pageNumber="49">ZRC</collectionCode>
.1.13132
<collectionCode id="66E802A0FFAAB846FDAA40167B81083A" box="[598,655,1871,1897]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="48" pageNumber="49">ZRC</collectionCode>
.1.13133 (2017);
<emphasis id="328D4677FFAAB846FCAB40167D5C083A" bold="true" box="[855,1106,1871,1897]" pageId="48" pageNumber="49">
Venus Loop [
<collectionCode id="66E802A0FFAAB846FC0740167D31083A" box="[1019,1087,1871,1897]" pageId="48" pageNumber="49">WNP</collectionCode>
]:
</emphasis>
<materialsCitation id="B0919038FFAAB846FBA14017789508DE" ID-GBIF-Occurrence="4120689272" collectingDate="2016-06-17" collectionCode="ZRC" pageId="48" pageNumber="49" specimenCount="1">
<collectionCode id="66E802A0FFAAB846FBA140177D98083A" box="[1117,1174,1870,1897]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="48" pageNumber="49">ZRC</collectionCode>
.1.12531
<collectionCode id="66E802A0FFAAB846FAFF40177C32083A" box="[1283,1340,1870,1897]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="48" pageNumber="49">ZRC</collectionCode>
.1.12532 (
<date id="7447BCA5FFAAB846FF03402A789C08DE" box="[255,402,1907,1933]" pageId="48" pageNumber="49" value="2016-06-17">
<collectingDate id="6403454DFFAAB846FF03402A789C08DE" box="[255,402,1907,1933]" pageId="48" pageNumber="49" value="2016-06-17">17-Jun-2016</collectingDate>
</date>
)
</materialsCitation>
.
</paragraph>
<paragraph id="00469A65FFAAB846FF6B40CF7A3908E2" blockId="48.[151,1437,150,2041]" box="[151,823,1942,1969]" pageId="48" pageNumber="49">
<emphasis id="328D4677FFAAB846FF6B40CF7B8408E3" bold="true" box="[151,650,1942,1969]" pageId="48" pageNumber="49">
Additional
<collectingCountry id="78EEDAF5FFAAB846FEE240CF789908E3" box="[286,407,1942,1968]" name="Singapore" pageId="48" pageNumber="49">Singapore</collectingCountry>
museum specimens.
</emphasis>
No specimens.
</paragraph>
<paragraph id="00469A65FFAAB846FF6B40E37D1908AA" blockId="48.[151,1437,150,2041]" pageId="48" pageNumber="49">
<emphasis id="328D4677FFAAB846FF6B40E3789F0887" bold="true" box="[151,401,1978,2004]" pageId="48" pageNumber="49">
<collectingCountry id="78EEDAF5FFAAB846FF6B40E3781E0887" box="[151,272,1978,2004]" name="Singapore" pageId="48" pageNumber="49">Singapore</collectingCountry>
localities.
</emphasis>
Kranji Marshes—Lorong Halus Wetland—Nanyang Technological University Jurong Campus—Pang Sua Woodland—Pulau Tekong—Windsor Nature Park.
</paragraph>
</subSubSection>
</treatment>
</document>