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<document id="97BB3BED47887EE075102755FE518306" ID-DOI="10.5281/zenodo.193523" ID-GBIF-Dataset="d1368e6e-3dea-4d66-bacd-94c02d3805fa" ID-ISSN="1175-5326" ID-Zenodo-Dep="193523" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460152417717" checkinUser="plazi" docAuthor="Köhler, Gunther" docDate="2010" docId="602D87848342EC6AFF41F89BD985FE82" docLanguage="en" docName="zt02354p018.pdf" docOrigin="Zootaxa 2354" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Anolis beckeri Boulenger 1881" docType="treatment" docVersion="8" lastPageNumber="12" masterDocId="9C14FFFC834AEC61FFD6FF8FDD27FFA0" masterDocTitle="A revision of the Central American species related to Anolis pentaprion with the resurrection of A. beckeri and the description of a new species (Squamata: Polychrotidae)" masterLastPageNumber="18" masterPageNumber="1" pageNumber="9" updateTime="1698235538294" updateUser="plazi">
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<mods:title id="ED721637EF00AAB3C8386F0DB6D9D530">A revision of the Central American species related to Anolis pentaprion with the resurrection of A. beckeri and the description of a new species (Squamata: Polychrotidae)</mods:title>
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<mods:namePart id="8C4A313EFA950562FDCA3FA7CF2ECC3C">Köhler, Gunther</mods:namePart>
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<mods:date id="317B3B2423C8D023264559D962F415CB">2010</mods:date>
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<treatment id="602D87848342EC6AFF41F89BD985FE82" ID-DOI="http://doi.org/10.5281/zenodo.6201489" ID-GBIF-Taxon="119383283" ID-Zenodo-Dep="6201489" LSID="urn:lsid:plazi:treatment:602D87848342EC6AFF41F89BD985FE82" httpUri="http://treatment.plazi.org/id/602D87848342EC6AFF41F89BD985FE82" lastPageId="11" lastPageNumber="12" pageId="8" pageNumber="9">
<subSubSection id="A09E65198342EC69FF41F89BDF29F88D" box="[151,526,1811,1838]" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="E83B36928342EC69FF41F89BDF29F88D" blockId="8.[151,526,1811,1838]" box="[151,526,1811,1838]" pageId="8" pageNumber="9">
<heading id="B37381FE8342EC69FF41F89BDF29F88D" bold="true" box="[151,526,1811,1838]" fontSize="11" level="1" pageId="8" pageNumber="9" reason="1">
<taxonomicName id="2F844D118342EC69FF41F89BDF29F88D" authority="Boulenger 1881" authorityName="Boulenger" authorityYear="1881" box="[151,526,1811,1838]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="beckeri">
<emphasis id="DAF0EA808342EC69FF41F89BDF29F88D" bold="true" box="[151,526,1811,1838]" pageId="8" pageNumber="9">
<emphasis id="DAF0EA808342EC69FF41F89BDC63F88E" bold="true" box="[151,324,1812,1838]" italics="true" pageId="8" pageNumber="9">Anolis beckeri</emphasis>
<bibRefCitation id="8C154B638342EC69FE9AF89CDF29F88D" author="Boulenger" box="[332,526,1811,1837]" pageId="8" pageNumber="16" refString="Boulenger, G. A. (1881) Description of a new species of Anolis from Yucatan. Proceedings of the Zoological Society London, 1881, 921 - 922." type="proceedings paper" year="1881">Boulenger 1881</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A09E65198342EC69FF41F8D3DE92F853" pageId="8" pageNumber="9" type="reference_group">
<paragraph id="E83B36928342EC69FF41F8D3D9DFF8D3" blockId="8.[151,1436,1884,2035]" box="[151,1272,1884,1907]" pageId="8" pageNumber="9">
<treatmentCitationGroup id="C89411BC8342EC69FF41F8D3D9DFF8D3" box="[151,1272,1884,1907]" pageId="8" pageNumber="9">
<taxonomicName id="2F844D118342EC69FF41F8D3DF3EF8D3" authority="Boulenger, 1881:921" authorityName="Boulenger" authorityPageNumber="921" authorityYear="1881" box="[151,537,1884,1907]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="beckeri">
<emphasis id="DAF0EA808342EC69FF41F8D3DC17F8D3" box="[151,304,1884,1907]" italics="true" pageId="8" pageNumber="9">Anolis beckeri</emphasis>
<treatmentCitation id="692510838342EC69FEE1F8D3DF3EF8D3" author="Boulenger" box="[311,537,1884,1907]" page="921" pageId="8" pageNumber="9" year="1881">
<bibRefCitation id="8C154B638342EC69FEE1F8D3DCCCF8D3" author="Boulenger" box="[311,491,1884,1907]" pageId="8" pageNumber="16" refString="Boulenger, G. A. (1881) Description of a new species of Anolis from Yucatan. Proceedings of the Zoological Society London, 1881, 921 - 922." type="proceedings paper" year="1881">Boulenger, 1881</bibRefCitation>
:921
</treatmentCitation>
</taxonomicName>
; two
<typeStatus id="373F88308342EC69FD85F8D3DF94F8D3" box="[595,691,1884,1907]" pageId="8" pageNumber="9" type="syntype">syntypes</typeStatus>
, IRSNB 2010 (12), from “Yucatán.”
<bibRefCitation id="8C154B638342EC69FB9AF8D3D9D3F8D3" author="Barbour" box="[1100,1268,1884,1907]" pageId="8" pageNumber="16" refString="Barbour, T. (1934) The anoles. II. The mainland species from Mexico southward. Bulletin of the Museum of Comparative Zoology, Harvard University, 77, 121 - 155." type="journal article" year="1934">Barbour (1934)</bibRefCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="E83B36928342EC69FF41F8F3DE5CF813" blockId="8.[151,1436,1884,2035]" pageId="8" pageNumber="9">
<taxonomicName id="2F844D118342EC69FF41F8F3DC84F833" authority="Stuart" authorityName="Stuart" box="[151,419,1916,1939]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="pentaprion">
<emphasis id="DAF0EA808342EC69FF41F8F3DC71F833" box="[151,342,1916,1939]" italics="true" pageId="8" pageNumber="9">Anolis pentaprion</emphasis>
: Stuart
</taxonomicName>
(1937; in part),
<bibRefCitation id="8C154B638342EC69FD87F8F3DE15F833" author="Stuart" box="[593,818,1916,1939]" pageId="8" pageNumber="17" refString="Stuart, L. C. (1948) The amphibians and reptiles of Alta Verapaz, Guatemala. Miscellaneous Publications Museum of Zoology University of Michigan, 69, 1 - 109." type="journal article" year="1948">Stuart (1948; in part)</bibRefCitation>
,
<bibRefCitation id="8C154B638342EC69FCE8F8F3D902F833" author="Myers" box="[830,1061,1916,1939]" lastPageId="8" lastPageNumber="17" pageId="8" pageNumber="16" refString="Myers, C. W. (1971) Central American lizards related to Anolis pentaprion: Two new species from the Cordillera de Talamanca. American Museum Novitates, 2471, 1 - 40." type="journal article" year="1971">Myers (1971; in part)</bibRefCitation>
,
<bibRefCitation id="8C154B638342EC69FBF9F8F3D8BFF833" author="Meyer" box="[1071,1432,1916,1939]" pageId="8" pageNumber="16" refString="Meyer, J. R. &amp; Wilson, L. D. (1973) A distributional checklist of the turtles, crocodilians, and lizards of Honduras. Contributions in Science, Natural History Museum, Los Angeles County, 244, 1 - 39." type="journal article" year="1973">Meyer and Wilson (1973; in part)</bibRefCitation>
;
<bibRefCitation id="8C154B638342EC69FF11F813DF31F813" author="Fitch" box="[199,534,1948,1971]" pageId="8" pageNumber="16" refString="Fitch, H. S. &amp; Seigel, R. A. (1984) Ecological and taxonomic notes on Nicaraguan anoles. Milwaukee Public Museum Contributions in Biology and Geology, 57, 1 - 13." type="journal article" year="1984">Fitch and Seigel (1984; in part)</bibRefCitation>
,
<bibRefCitation id="8C154B638342EC69FDF4F813DFBDF813" author="Lee" box="[546,666,1948,1971]" pageId="8" pageNumber="16" refString="Lee, J. C. (1996) The amphibians and reptiles of the Yucatan Peninsula. Cornell University Press, Ithaca, New York, U. S. A., 500 pp." type="book" year="1996">Lee (1996)</bibRefCitation>
,
<bibRefCitation id="8C154B638342EC69FD73F813DE50F813" author="Poe" box="[677,887,1948,1971]" pageId="8" pageNumber="17" refString="Poe, S. (2004) Phylogeny of anoles. Herpetological Monographs, 18, 37 - 89." type="journal article" year="2004">Poe (2004; in part.)</bibRefCitation>
.
</paragraph>
<paragraph id="E83B36928342EC69FF41F833DE37F873" blockId="8.[151,1436,1884,2035]" box="[151,784,1980,2003]" pageId="8" pageNumber="9">
<taxonomicName id="2F844D118342EC69FF41F833DF56F873" authority="Etheridge (1959)" authorityName="Etheridge" authorityYear="1959" box="[151,625,1980,2003]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="subSpecies" species="pentaprion" subSpecies="beckeri">
<emphasis id="DAF0EA808342EC69FF41F833DC8BF873" box="[151,428,1980,2003]" italics="true" pageId="8" pageNumber="9">Anolis pentaprion beckeri</emphasis>
:
<bibRefCitation id="8C154B638342EC69FE6CF833DF56F873" author="Etheridge" box="[442,625,1980,2003]" pageId="8" pageNumber="16" refString="Etheridge, R. (1959) The Relationships of the Anoles (Reptilia: Sauria: Iguanidae). An Interpretation Based on Skeletal Morphology. Ph. D. Dissertation, University of Michigan, Ann Arbor, Michigan, U. S. A." type="book" year="1959">Etheridge (1959)</bibRefCitation>
</taxonomicName>
,
<bibRefCitation id="8C154B638342EC69FDAAF833DE2BF873" author="Stuart" box="[636,780,1980,2003]" pageId="8" pageNumber="17" refString="Stuart, L. C. (1963) A checklist of the herpetofauna of Guatemala. Miscellaneous Publications Museum of Zoology University of Michigan, 122, 1 - 150." type="journal article" year="1963">Stuart (1963)</bibRefCitation>
.
</paragraph>
<paragraph id="E83B36928342EC69FF41F853DE92F853" blockId="8.[151,1436,1884,2035]" box="[151,949,2012,2035]" pageId="8" pageNumber="9">
<taxonomicName id="2F844D118342EC69FF41F853DC9FF853" authority="Kohler" authorityName="Kohler" box="[151,440,2012,2035]" class="Squamata" family="Dactyloidae" genus="Norops" higherTaxonomySource="CoL" kingdom="Animalia" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="pentaprion">
<emphasis id="DAF0EA808342EC69FF41F853DC47F853" box="[151,352,2012,2035]" italics="true" pageId="8" pageNumber="9">Norops pentaprion</emphasis>
:
<bibRefCitation id="8C154B638342EC69FEB8F853DC9FF853" author="Kohler" box="[366,440,2012,2035]" pageId="8" pageNumber="16" refString="Kohler, G. (2001) Anfibios y Reptiles de Nicaragua. Herpeton, Offenbach, Germany, 208 pp." type="book" year="2001">Köhler</bibRefCitation>
</taxonomicName>
(2001; in part),
<bibRefCitation id="8C154B638342EC69FDB0F853DE72F853" author="McCranie" box="[614,853,2012,2035]" pageId="8" pageNumber="16" refString="McCranie, J. R., Townsend, J. H. &amp; Wilson, L. D. (2006) The amphibians and reptiles of the Honduran Mosquitia. - Krieger Publishing Company, Malabar, Florida, U. S. A., 291 pp." type="book" year="2006">
McCranie
<emphasis id="DAF0EA808342EC69FD01F853DE2AF853" box="[727,781,2012,2035]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
(2006
</bibRefCitation>
; in part).
</paragraph>
</subSubSection>
<caption id="BCFB661A8343EC68FF41F85AD9AAF84C" box="[151,1165,2005,2029]" httpUri="https://zenodo.org/record/193529/files/figure.png" pageId="9" pageNumber="10" targetBox="[346,1244,209,1972]" targetPageId="9">
<paragraph id="E83B36928343EC68FF41F85AD9AAF84C" blockId="9.[151,1165,2005,2029]" box="[151,1165,2005,2029]" pageId="9" pageNumber="10">
<emphasis id="DAF0EA808343EC68FF41F85ADC3EF84D" bold="true" box="[151,281,2005,2029]" pageId="9" pageNumber="10">FIGURE 5.</emphasis>
Head of holotype of
<taxonomicName id="2F844D118343EC68FE2AF859DFFAF84D" box="[508,733,2006,2029]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="charlesmyersi">
<emphasis id="DAF0EA808343EC68FE2AF859DFFAF84D" box="[508,733,2006,2029]" italics="true" pageId="9" pageNumber="10">Anolis charlesmyersi</emphasis>
</taxonomicName>
(SMF 89688). Scale bars equal 1.0 mm.
</paragraph>
</caption>
<subSubSection id="A09E65198340EC6BFF41FF18D801FC8C" pageId="10" pageNumber="11" type="diagnosis">
<paragraph id="E83B36928340EC6BFF41FF18D801FC8C" blockId="10.[151,1438,151,2007]" pageId="10" pageNumber="11">
<emphasis id="DAF0EA808340EC6BFF41FF18DC3AFF11" bold="true" box="[151,285,151,177]" pageId="10" pageNumber="11">Diagnosis.</emphasis>
A medium-sized species (SVL in largest specimen examined
<quantity id="2F7C9B778340EC6BFBC9FF17D9AAFF12" box="[1055,1165,152,178]" metricMagnitude="-2" metricUnit="m" metricValue="6.06" pageId="10" pageNumber="11" unit="mm" value="60.6">60.6 mm</quantity>
in males,
<quantity id="2F7C9B778340EC6BFAD8FF17D85BFF12" box="[1294,1404,152,178]" metricMagnitude="-2" metricUnit="m" metricValue="5.54" pageId="10" pageNumber="11" unit="mm" value="55.4">55.4 mm</quantity>
in females) of the genus
<taxonomicName id="2F844D118340EC6BFE7AFF32DCDCFF76" box="[428,507,189,214]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">
<emphasis id="DAF0EA808340EC6BFE7AFF32DCDCFF76" box="[428,507,189,214]" italics="true" pageId="10" pageNumber="11">Anolis</emphasis>
</taxonomicName>
(sensu
<bibRefCitation id="8C154B638340EC6BFD8AFF32DFFCFF77" author="Poe" box="[604,731,189,215]" pageId="10" pageNumber="17" refString="Poe, S. (2004) Phylogeny of anoles. Herpetological Monographs, 18, 37 - 89." type="journal article" year="2004">Poe, 2004</bibRefCitation>
) that differs from all Central American anoles except
<emphasis id="DAF0EA808340EC6BFA54FF32DE8AFF5B" italics="true" pageId="10" pageNumber="11">
<taxonomicName id="2F844D118340EC6BFA54FF32DC64FF5B" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="charlesmyersi">A. charlesmyersi</taxonomicName>
,
<taxonomicName id="2F844D118340EC6BFE86FF6DDCF5FF5B" box="[336,466,226,251]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="cristifer">A. cristifer</taxonomicName>
,
<taxonomicName id="2F844D118340EC6BFE08FF6DDF48FF5B" box="[478,623,226,251]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="fungosus">A. fungosus</taxonomicName>
,
<taxonomicName id="2F844D118340EC6BFDAAFF6DDE02FF5B" box="[636,805,226,251]" class="Squamata" family="Dactyloidae" genus="Norops" higherTaxonomySource="CoL" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="pentaprion">A. pentaprion</taxonomicName>
,
<taxonomicName id="2F844D118340EC6BFCE4FF6DDE8FFF5B" box="[818,936,226,251]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="salvini">A. salvini</taxonomicName>
,
</emphasis>
and
<taxonomicName id="2F844D118340EC6BFC3FFF6DD957FF5B" box="[1001,1136,226,251]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="utilensis">
<emphasis id="DAF0EA808340EC6BFC3FFF6DD957FF5B" box="[1001,1136,226,251]" italics="true" pageId="10" pageNumber="11">A. utilensis</emphasis>
</taxonomicName>
by having (1) extremely short hind legs (fourth toe of adpressed hindlimb usually reaching to level of tympanum; ratio shank length / head length 0.490.84); (2) smooth, juxtaposed ventral scales; (3) a relatively short tail (ratio tail length / SVL 1.01.5).
<taxonomicName id="2F844D118340EC6BFEDCFEDDDC90FECB" box="[266,439,338,363]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="beckeri">
<emphasis id="DAF0EA808340EC6BFEDCFEDDDC90FECB" box="[266,439,338,363]" italics="true" pageId="10" pageNumber="11">Anolis beckeri</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2F844D118340EC6BFD81FEDDDE07FECB" box="[599,800,338,363]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="charlesmyersi">
<emphasis id="DAF0EA808340EC6BFD81FEDDDE07FECB" box="[599,800,338,363]" italics="true" pageId="10" pageNumber="11">A. charlesmyersi</emphasis>
</taxonomicName>
by having a pink dewlap in life with purple pigment between rows of gorgetal scales and regular rows of numerous small narrowly spaced gorgetal scales (versus a brick red dewlap with few large widely spaced gorgetal scales scattered across the dewlap).
<taxonomicName id="2F844D118340EC6BFB24FE12D8BBFE16" box="[1266,1436,413,438]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="beckeri">
<emphasis id="DAF0EA808340EC6BFB24FE12D8BBFE16" box="[1266,1436,413,438]" italics="true" pageId="10" pageNumber="11">Anolis beckeri</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2F844D118340EC6BFEFAFE4DDC8CFE7B" box="[300,427,450,475]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="cristifer">
<emphasis id="DAF0EA808340EC6BFEFAFE4DDC8CFE7B" box="[300,427,450,475]" italics="true" pageId="10" pageNumber="11">A. cristifer</emphasis>
</taxonomicName>
by having a double row of slightly enlarged middorsal scales, not forming a serrated crest (versus a serrated middorsal crest, at least on posterior portion of dorsum, present in
<taxonomicName id="2F844D118340EC6BFB6DFE67D81EFDA1" box="[1211,1337,488,513]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="cristifer">
<emphasis id="DAF0EA808340EC6BFB6DFE67D81EFDA1" box="[1211,1337,488,513]" italics="true" pageId="10" pageNumber="11">A. cristifer</emphasis>
</taxonomicName>
).
<taxonomicName id="2F844D118340EC6BFA87FE67DDC9FD86" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="beckeri">
<emphasis id="DAF0EA808340EC6BFA87FE67DDC9FD86" italics="true" pageId="10" pageNumber="11">Anolis beckeri</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2F844D118340EC6BFE5AFD82DF30FD86" box="[396,535,525,550]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="fungosus">
<emphasis id="DAF0EA808340EC6BFE5AFD82DF30FD86" box="[396,535,525,550]" italics="true" pageId="10" pageNumber="11">A. fungosus</emphasis>
</taxonomicName>
by lacking bony parietal protuberances (versus having a pair of small bony parietal protuberances posterior and lateral to interparietal plate).
<taxonomicName id="2F844D118340EC6BFC43FDBDD91AFDEB" box="[917,1085,562,587]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="beckeri">
<emphasis id="DAF0EA808340EC6BFC43FDBDD91AFDEB" box="[917,1085,562,587]" italics="true" pageId="10" pageNumber="11">Anolis beckeri</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2F844D118340EC6BFB01FDBDD85FFDEB" box="[1239,1400,562,587]" class="Squamata" family="Dactyloidae" genus="Norops" higherTaxonomySource="CoL" kingdom="Animalia" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="pentaprion">
<emphasis id="DAF0EA808340EC6BFB01FDBDD85FFDEB" box="[1239,1400,562,587]" italics="true" pageId="10" pageNumber="11">A. pentaprion</emphasis>
</taxonomicName>
by lacking a pronounced serrated caudal crest (versus having a pronounced serrated caudal crest) and by having gorgetal rows with 410 widely spaced scales (versus gorgetal rows with 1725 narrowly spaced scales).
<taxonomicName id="2F844D118340EC6BFF41FD2DDC60FD1B" box="[151,327,674,699]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="beckeri">
<emphasis id="DAF0EA808340EC6BFF41FD2DDC60FD1B" box="[151,327,674,699]" italics="true" pageId="10" pageNumber="11">Anolis beckeri</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2F844D118340EC6BFE3CFD2DDF7AFD1B" box="[490,605,674,699]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="salvini">
<emphasis id="DAF0EA808340EC6BFE3CFD2DDF7AFD1B" box="[490,605,674,699]" italics="true" pageId="10" pageNumber="11">A. salvini</emphasis>
</taxonomicName>
by having smooth, obliquely conical ventral scales (versus keeled), lacking a dark interorbital bar (versus present), and lacking enlarged postcloacal scales (versus males with a pair of greatly enlarged postcloacal scales).
<taxonomicName id="2F844D118340EC6BFD7FFD62DE72FCA6" box="[681,853,749,774]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="beckeri">
<emphasis id="DAF0EA808340EC6BFD7FFD62DE72FCA6" box="[681,853,749,774]" italics="true" pageId="10" pageNumber="11">Anolis beckeri</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2F844D118340EC6BFC20FD62D95BFCA6" box="[1014,1148,749,774]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="utilensis">
<emphasis id="DAF0EA808340EC6BFC20FD62D95BFCA6" box="[1014,1148,749,774]" italics="true" pageId="10" pageNumber="11">A. utilensis</emphasis>
</taxonomicName>
by having the proximal subdigital scales of toes differentiated as slightly broadened lamellae (versus those scales granular).
</paragraph>
</subSubSection>
<subSubSection id="A09E65198340EC6AFF12FCB8D985FE82" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="description">
<paragraph id="E83B36928340EC6AFF12FCB8DF93FF77" blockId="10.[151,1438,151,2007]" lastBlockId="11.[151,1436,152,290]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<emphasis id="DAF0EA808340EC6BFF12FCB8DC46FCF1" bold="true" box="[196,353,823,849]" pageId="10" pageNumber="11">Description.</emphasis>
SVL 46.560.6 (51.0±5.0) mm in males, 38.055.4 (58.3±9.6) mm in females; for other morphometric data see Table 1; dorsal head scales in internasal, prefrontal, and frontal areas rugose; scales in parietal area rugose to smooth; shallow frontal and parietal depressions present; 58 postrostrals; anterior nasal divided, lower scale contacting rostral and first supralabial; 59 internasals; canthal ridge sharply defined; scales comprising supraorbital semicircles ridged, especially anterior ones, largest scale in semicircles larger than largest supraocular scale; supraorbital semicircles well defined; supraorbital semicircles broadly in contact with each other or separated by one scale at narrowest point; 13 scales separating supraorbital semicircles and interparietal at narrowest point; interparietal well defined, irregular in outline, longer than wide, larger than size of ear opening; about 411 enlarged, smooth to faintly keeled supraocular scales on each side, in 23 rows; enlarged supraoculars completely separated from supraorbital semicircles; three short superciliaries, posterior one shortest; usually 3 enlarged canthals; 614 scales between second canthals; 79 scales present between posterior canthals; loreal region slightly concave, 1830 mostly smooth or rugose (some keeled) loreal scales in a maximum of 34 horizontal rows; 710 supralabials and 7 10 infralabials to level below center of eye; suboculars smooth to rugose, in broad contact with supralabials; ear opening vertically oval; scales anterior to ear opening slightly larger than those posterior to ear opening; 57 postmentals, outer pair largest; gular scales not keeled; male dewlap moderately large, extending to beyond level of axillae onto chest; male dewlap with 56 horizontal gorgetal-sternal scale rows, about 1618 scales per row; female dewlap well developed, extending to level of axillae; female dewlap with 47 horizontal gorgetal-sternal scale rows, with about 915 scales per row; about 2 middorsal scale rows slightly enlarged, mostly smooth, dorsal scales lateral to middorsal series grading into granular lateral scales; no enlarged scales among granular laterals; lateral scales granular, slightly smaller than largest dorsal scales; about 95104 dorsal scales along vertebral midline between levels of axillae and groin in males, about
<quantity id="2F7C9B778340EC6BFA92F9C7DD89F927" metricMagnitude="0" metricUnit="m" metricValue="2.6289" metricValueMax="3.1242" metricValueMin="2.1336" pageId="10" pageNumber="11" unit="in" value="103.5" valueMax="123.0" valueMin="84.0">84123 in</quantity>
females; about 5680 dorsal scales along vertebral midline contained in one head length in males, about
<quantity id="2F7C9B778340EC6BFF41F91DDC27F90C" box="[151,256,1682,1708]" metricMagnitude="0" metricUnit="m" metricValue="1.4605" metricValueMax="1.778" metricValueMin="1.143" pageId="10" pageNumber="11" unit="in" value="57.5" valueMax="70.0" valueMin="45.0">4570 in</quantity>
females; ventral scales on midsection slightly larger than largest dorsal scales; ventral body scales smooth, obliquely conical, juxtaposed; about 6786 ventral scales along midventral line between levels of axilla and groin in males, about
<quantity id="2F7C9B778340EC6BFDC7F952DF5EF957" box="[529,633,1757,1783]" metricMagnitude="0" metricUnit="m" metricValue="2.0193" metricValueMax="2.3368" metricValueMin="1.7018" pageId="10" pageNumber="11" unit="in" value="79.5" valueMax="92.0" valueMin="67.0">6792 in</quantity>
females; about 3460 ventral scales contained in one head length in males, about
<quantity id="2F7C9B778340EC6BFEE0F88DDCB8F8BC" box="[310,415,1794,1820]" metricMagnitude="0" metricUnit="m" metricValue="1.1938" metricValueMax="1.524" metricValueMin="0.8635999999999999" pageId="10" pageNumber="11" unit="in" value="47.0" valueMax="60.0" valueMin="34.0">3460 in</quantity>
females; about 146178 scales around midbody in males, about
<quantity id="2F7C9B778340EC6BFB7EF88DD808F8BC" box="[1192,1327,1794,1820]" metricMagnitude="0" metricUnit="m" metricValue="3.9116" metricValueMax="4.5212" metricValueMin="3.3019999999999996" pageId="10" pageNumber="11" unit="in" value="154.0" valueMax="178.0" valueMin="130.0">130178 in</quantity>
females; tubelike axillary pocket absent; precloacal scales not keeled; enlarged postcloacal scales absent; tail oval at about midlength; tail height/tail width 0.801.25; basal subcaudal scales smooth to faintly keeled; lateral caudal scales keeled, homogeneous, although an indistinct division in segments is discernible; dorsal medial caudal scale row enlarged, keeled, not forming a crest; 2 median subcaudal scale rows distinctly enlarged, keeled; most scales on lateral surface of antebrachium smooth; 1832 subdigital lamellae on Phalanges IIIV of Toe IV of hind limbs; 710 subdigital scales on Phalanx I of Toe IV of hind limbs; longest toe of adpressed hind limb usually reaching to shoulder region.
</paragraph>
<paragraph id="E83B36928341EC6AFF12FF6DD985FE82" blockId="11.[151,1436,152,290]" pageId="11" pageNumber="12">
<emphasis id="DAF0EA808341EC6AFF12FF6DDF29FF5C" bold="true" box="[196,526,226,252]" pageId="11" pageNumber="12">Geographic distribution:</emphasis>
Along the Caribbean versant of Nuclear Central
<collectingCountry id="909376028341EC6AFBAEFF6DD9C5FF5C" box="[1144,1250,226,252]" name="United States of America" pageId="11" pageNumber="12">America</collectingCountry>
from Tabasco,
<collectingCountry id="909376028341EC6AFF41FE87DDD3FE82" box="[151,244,264,290]" name="Mexico" pageId="11" pageNumber="12">Mexico</collectingCountry>
, to northern
<collectingCountry id="909376028341EC6AFE5EFE87DF24FE82" box="[392,515,264,290]" name="Nicaragua" pageId="11" pageNumber="12">Nicaragua</collectingCountry>
, including the southern portion of the Yucatán Peninsula.
</paragraph>
</subSubSection>
</treatment>
</document>