treatments-xml/data/5F/23/87/5F2387E7712FFE8DFF5FFED5FEE21FA5.xml

<document id="8006B2CDF4AD36E293927007B98E6A4D" ID-DOI="10.11646/zootaxa.4796.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="3896810" ID-ZooBank="66DD1FEB-6BDE-4AEB-8A7B-96594371E9C5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1592309422476" checkinUser="plazi" docAuthor="Machado, Renato Jose Pires &amp; Oswald, John David" docDate="2020" docId="5F2387E7712FFE8DFF5FFED5FEE21FA5" docLanguage="en" docName="zootaxa.4796.1.1.pdf.imf" docOrigin="Zootaxa 4796 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Riekoleon longitudinalis Machado &amp; Oswald 2020, new species" docType="treatment" docUuid="9C389775-39FE-431E-9844-081C21D80773" docUuidSource="ZooBank" docVersion="4" lastPageNumber="304" masterDocId="A31AFF9F7003FFA2FFC8FFA5FFE8183D" masterDocTitle="Morphological phylogeny and taxonomic revision of the former antlion subtribe Periclystina (Neuroptera: Myrmeleontidae: Dendroleontinae)" masterLastPageNumber="322" masterPageNumber="1" pageNumber="301" updateTime="1700276817129" updateUser="plazi">
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<mods:title id="CC3F2E5822D146A04FF8C40D35D41721">Morphological phylogeny and taxonomic revision of the former antlion subtribe Periclystina (Neuroptera: Myrmeleontidae: Dendroleontinae)</mods:title>
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<mods:namePart id="741D8D08FA3BD5E06BD7A268DB796B28">Machado, Renato Jose Pires</mods:namePart>
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<mods:namePart id="E17795AA2AB24B5C33D1A7158DF1B93C">Oswald, John David</mods:namePart>
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<mods:date id="40BB5DCB173185C623E8941FC78513E8">2020</mods:date>
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<mods:number id="58DC9765AFCF51D8537D2B4BD2510E69">2020-06-16</mods:number>
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<paragraph id="D73536F1712FFE8EFF5FFED5FD3419B7" blockId="300.[151,1098,368,467]" box="[151,732,368,395]" pageId="300" pageNumber="301">
<heading id="8C7D819D712FFE8EFF5FFED5FD3419B7" bold="true" box="[151,732,368,395]" fontSize="11" level="1" pageId="300" pageNumber="301" reason="1">
<emphasis id="E5FEEAE3712FFE8EFF5FFED5FD3419B7" bold="true" box="[151,732,368,395]" pageId="300" pageNumber="301">
<taxonomicName id="108A4D72712FFE8EFF5FFED5FDD719B6" authority="Machado" authorityName="Machado &amp; Oswald" authorityYear="2020" box="[151,575,368,395]" class="Insecta" family="Myrmeleontidae" genus="Riekoleon" kingdom="Animalia" order="Neuroptera" pageId="300" pageNumber="301" phylum="Arthropoda" rank="species" species="longitudinalis" status="sp. nov.">
<emphasis id="E5FEEAE3712FFE8EFF5FFED5FE2919B6" bold="true" box="[151,449,368,395]" italics="true" pageId="300" pageNumber="301">Riekoleon longitudinalis</emphasis>
Machado
</taxonomicName>
, 
<taxonomicNameLabel id="FECD5798712FFE8EFD84FED4FD3419B7" box="[588,732,368,395]" pageId="300" pageNumber="301" rank="species">new species</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<paragraph id="D73536F1712FFE8EFF5FFE31FEA21992" blockId="300.[151,1098,368,467]" box="[151,330,404,431]" pageId="300" pageNumber="301">
(
<figureCitation id="4FB12A74712FFE8EFF57FE31FEAA1992" box="[159,322,404,431]" captionStart-0="FIGURE 160" captionStart-1="FIGURE 161" captionStart-2="FIGURE 162" captionStartId-0="301.[151,250,1907,1932]" captionStartId-1="302.[151,250,1878,1903]" captionStartId-2="303.[151,250,1271,1296]" captionTargetBox-0="[153,1436,457,1868]" captionTargetBox-1="[153,1429,185,1853]" captionTargetBox-2="[151,1434,181,1246]" captionTargetId-0="figure@301.[151,1436,453,1882]" captionTargetId-1="figure@302.[151,1436,181,1853]" captionTargetId-2="figure@303.[151,1435,181,1246]" captionTargetPageId-0="301" captionTargetPageId-1="302" captionTargetPageId-2="303" captionText-0="FIGURE 160. Riekoleon longitudinalis sp. nov.: a) head, anterior; b) head and thorax, dorsal; c) wings, dorsal." captionText-1="FIGURE 161. Riekoleon longitudinalis sp. nov.: female: a) terminalia, lateral; b) terminalia, ventral (note pregenital plate hidden beneath sternite). Scale bars = 1mm." captionText-2="FIGURE 162. Distribution of Riekoleon species (part)." figureDoi-0="http://doi.org/10.5281/zenodo.3897142" figureDoi-1="http://doi.org/10.5281/zenodo.3897150" figureDoi-2="http://doi.org/10.5281/zenodo.3897152" httpUri-0="https://zenodo.org/record/3897142/files/figure.png" httpUri-1="https://zenodo.org/record/3897150/files/figure.png" httpUri-2="https://zenodo.org/record/3897152/files/figure.png" pageId="300" pageNumber="301">Figs. 160–162</figureCitation>
)
</paragraph>
<paragraph id="D73536F1712FFE8EFF5FFE1DFBA219EE" blockId="300.[151,1098,368,467]" box="[151,1098,440,467]" pageId="300" pageNumber="301">
http://zoobank.org/NomenclaturalActs/ 
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</paragraph>
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<subSubSection id="9F90657A712FFE8EFF5FFDA5FDCE1A5F" pageId="300" pageNumber="301" type="diagnosis">
<paragraph id="D73536F1712FFE8EFF5FFDA5FDCE1A5F" blockId="300.[151,1437,512,2015]" pageId="300" pageNumber="301">
<emphasis id="E5FEEAE3712FFE8EFF5FFDA5FEFA1A26" bold="true" box="[151,274,512,539]" pageId="300" pageNumber="301">Diagnosis.</emphasis>
Forewing: first fork of RP in proximal half of wing, dark longitudinal line extending across radial area; hind wing with subapical mark as large as rhegmal mark; female posterior gonapophysis longer than wide; female anterior gonapophysis a large plate.
</paragraph>
</subSubSection>
<subSubSection id="9F90657A712FFE8FFF0FFDC9FD7C1992" lastPageId="301" lastPageNumber="302" pageId="300" pageNumber="301" type="description">
<paragraph id="D73536F1712FFE8EFF0FFDC9FC2C1ABA" blockId="300.[151,1437,512,2015]" box="[199,964,620,647]" pageId="300" pageNumber="301">
<emphasis id="E5FEEAE3712FFE8EFF0FFDC9FEB11ABA" bold="true" box="[199,345,620,647]" pageId="300" pageNumber="301">Description.</emphasis>
Lengths: forewing: 
<quantity id="10729B14712FFE8EFDF4FDC8FD501ABA" box="[572,696,621,647]" metricMagnitude="-2" metricUnit="m" metricValue="3.25" metricValueMax="3.3" metricValueMin="3.2" pageId="300" pageNumber="301" unit="mm" value="32.5" valueMax="33.0" valueMin="32.0">32–33 mm</quantity>
; hind wing: 
<quantity id="10729B14712FFE8EFC8BFDC8FC571ABA" box="[835,959,621,647]" metricMagnitude="-2" metricUnit="m" metricValue="2.95" metricValueMax="3.0" metricValueMin="2.9" pageId="300" pageNumber="301" unit="mm" value="29.5" valueMax="30.0" valueMin="29.0">29–30 mm</quantity>
.
</paragraph>
<paragraph id="D73536F1712FFE8EFF0FFD35FA7C1BF7" blockId="300.[151,1437,512,2015]" pageId="300" pageNumber="301">
Head (
<figureCitation id="4FB12A74712FFE8EFEDCFD35FE691A96" box="[276,385,656,683]" captionStart="FIGURE 160" captionStartId="301.[151,250,1907,1932]" captionTargetBox="[153,1436,457,1868]" captionTargetId="figure@301.[151,1436,453,1882]" captionTargetPageId="301" captionText="FIGURE 160. Riekoleon longitudinalis sp. nov.: a) head, anterior; b) head and thorax, dorsal; c) wings, dorsal." figureDoi="http://doi.org/10.5281/zenodo.3897142" httpUri="https://zenodo.org/record/3897142/files/figure.png" pageId="300" pageNumber="301">Figs. 160</figureCitation>
a–b): 
<emphasis id="E5FEEAE3712FFE8EFE09FD35FDF21A97" box="[449,538,656,682]" italics="true" pageId="300" pageNumber="301">Labrum</emphasis>
pale with a line of elongate setae. 
<emphasis id="E5FEEAE3712FFE8EFC66FD35FBE01A97" box="[942,1032,656,682]" italics="true" pageId="300" pageNumber="301">Clypeus</emphasis>
pale; with few elongate pale setae. 
<emphasis id="E5FEEAE3712FFE8EFF5FFD10FF311AF3" box="[151,217,693,718]" italics="true" pageId="300" pageNumber="301">Frons</emphasis>
beneath antennae pale but remaining areas black; set with few pale setae. 
<emphasis id="E5FEEAE3712FFE8EFBC6FD10FBA31AF2" box="[1038,1099,693,719]" italics="true" pageId="300" pageNumber="301">Gena</emphasis>
pale. 
<emphasis id="E5FEEAE3712FFE8EFB44FD10FB3A1AF3" box="[1164,1234,693,718]" italics="true" pageId="300" pageNumber="301">Vertex</emphasis>
raised; in anterior view pale except for two black longitudinal marks around the central region; in dorsal view pale except for two black longitudinal lines surrounding central area, and a black transverse medial line; covered with short black setae. 
<emphasis id="E5FEEAE3712FFE8EFF5FFC85FF0E1B07" box="[151,230,800,826]" italics="true" pageId="300" pageNumber="301">Ocular</emphasis>
setae absent. 
<emphasis id="E5FEEAE3712FFE8EFE49FC84FE031B06" box="[385,491,801,827]" italics="true" pageId="300" pageNumber="301">Antennae</emphasis>
almost twice longer than pronotum; basal flagellomeres almost as long as wide, apical ones wider than long; scape, pedicel and first flagellomere black, remaining segments light brown; flagellum covered with short black setae; scape and pedicel set with short pale setae; torular membrane black. 
<emphasis id="E5FEEAE3712FFE8EFB39FCCDFA8F1BBF" box="[1265,1383,872,898]" italics="true" pageId="300" pageNumber="301">Mandibles</emphasis>
pale with tip black. 
<emphasis id="E5FEEAE3712FFE8EFEF4FC29FE911B9B" box="[316,377,908,934]" italics="true" pageId="300" pageNumber="301">Palpi</emphasis>
, maxillary and labial pale, except for basal maxillary segment black, and area around palpimacula opening slightly darker; apical labial palpomere fusiform, palpimacula opening oval-shaped, located medially.
</paragraph>
<paragraph id="D73536F1712FFE8EFF0FFC71FB821CFA" blockId="300.[151,1437,512,2015]" pageId="300" pageNumber="301">
Thorax (
<figureCitation id="4FB12A74712FFE8EFEE1FC71FE741BD2" box="[297,412,980,1007]" captionStart="FIGURE 160" captionStartId="301.[151,250,1907,1932]" captionTargetBox="[153,1436,457,1868]" captionTargetId="figure@301.[151,1436,453,1882]" captionTargetPageId="301" captionText="FIGURE 160. Riekoleon longitudinalis sp. nov.: a) head, anterior; b) head and thorax, dorsal; c) wings, dorsal." figureDoi="http://doi.org/10.5281/zenodo.3897142" httpUri="https://zenodo.org/record/3897142/files/figure.png" pageId="300" pageNumber="301">Fig. 160b</figureCitation>
): 
<emphasis id="E5FEEAE3712FFE8EFE79FC70FDC81BD3" box="[433,544,981,1006]" italics="true" pageId="300" pageNumber="301">Pronotum</emphasis>
about as long as wide; posterior margin as wide as anterior; subapical furrow present; pale except by a broad black longitudinal streak that enclose a thin pale line on medial area, two thin black lateral lines on posterior area (lines interrupted on their medial length), and posterior corners black; beset with many short black setae and a few long setae on margins. 
<emphasis id="E5FEEAE3712FFE8EFCE1FBE5FBFB1C67" box="[809,1043,1088,1114]" italics="true" pageId="300" pageNumber="301">Pterothoracic notum</emphasis>
pale except for a broad black line that extends from pronotum throughout pterothorax; mesoscutum with two thin longitudinal black lines; covered with short black setae. 
<emphasis id="E5FEEAE3712FFE8EFE5DFB2DFD971C9F" box="[405,639,1160,1186]" italics="true" pageId="300" pageNumber="301">Pterothoracic pleura</emphasis>
mostly black but ventral area of ventral segments pale; dorsal segments covered with black setae and ventral segments with white setae; Miller’s organ present.
</paragraph>
<paragraph id="D73536F1712FFE8EFF0FFB75FB401E13" blockId="300.[151,1437,512,2015]" pageId="300" pageNumber="301">
Wings (
<figureCitation id="4FB12A74712FFE8EFED6FB75FE651CD6" box="[286,397,1232,1259]" captionStart="FIGURE 160" captionStartId="301.[151,250,1907,1932]" captionTargetBox="[153,1436,457,1868]" captionTargetId="figure@301.[151,1436,453,1882]" captionTargetPageId="301" captionText="FIGURE 160. Riekoleon longitudinalis sp. nov.: a) head, anterior; b) head and thorax, dorsal; c) wings, dorsal." figureDoi="http://doi.org/10.5281/zenodo.3897142" httpUri="https://zenodo.org/record/3897142/files/figure.png" pageId="300" pageNumber="301">Fig. 160c</figureCitation>
): Rather narrow with tip acute, anterior Banksian line present in both wings, posterior absent; veins mostly brown, but with some veins intercalated with white spots, beset with short black setae. 
<emphasis id="E5FEEAE3712FFE8EFAD0FB51FF1C1D0F" italics="true" pageId="300" pageNumber="301">Male pilula axillaris</emphasis>
present. 
<emphasis id="E5FEEAE3712FFE8EFEA8FABCFE241D0F" box="[352,460,1305,1330]" italics="true" pageId="300" pageNumber="301">Forewing</emphasis>
membrane mostly hyaline but with a longitudinal brown line starting on costal area base, continuing throughout mediocubital area and extending over gradate crossveins up to apex, line interrupted by some small hyaline areas; there is also a small stigmal brown mark, and tiny brown marks on posterior and apical margins; pterostigma with apex white and base brown; CuA fork located between RP origin and first fork; three presectoral crossveins, distal crossvein with a medial veinlet connecting to RP; subcostal veinlets simple, but few distal sometimes forked; posterior area narrower than prefork area at CuA fork level. 
<emphasis id="E5FEEAE3712FFE8EFB9EFA69FB231DDB" box="[1110,1227,1484,1510]" italics="true" pageId="300" pageNumber="301">Hind wing</emphasis>
membrane mostly hyaline except for two large brown spots, an apical and other on rhegmal area; pterostigma mostly white; MP fork located between RP origin and first fork; subcostal venlets simple; one presectoral crossvein.
</paragraph>
<paragraph id="D73536F1712FFE8EFF0FF99CFD451F87" blockId="300.[151,1437,512,2015]" pageId="300" pageNumber="301">
Legs (
<figureCitation id="4FB12A74712FFE8EFEC5F99DFE911E6E" box="[269,377,1592,1619]" captionStart="FIGURE 160" captionStartId="301.[151,250,1907,1932]" captionTargetBox="[153,1436,457,1868]" captionTargetId="figure@301.[151,1436,453,1882]" captionTargetPageId="301" captionText="FIGURE 160. Riekoleon longitudinalis sp. nov.: a) head, anterior; b) head and thorax, dorsal; c) wings, dorsal." figureDoi="http://doi.org/10.5281/zenodo.3897142" httpUri="https://zenodo.org/record/3897142/files/figure.png" pageId="300" pageNumber="301">Figs. 160</figureCitation>
a–b): 
<emphasis id="E5FEEAE3712FFE8EFE70F99DFD841E6F" box="[440,620,1592,1619]" italics="true" pageId="300" pageNumber="301">All pairs of legs</emphasis>
, femur elongate (&gt; 2.5x length of coxa); tibia slightly longer than femur and about twice longer than tarsi; tibial spurs extending to T2 apex; all tarsomeres about same size, except for T5 slightly longer; claws about half of T5 length; coxa and trochanter set with white setae and few black setae; femur with internal surface covered with short white setae but remaining areas covered with short black setae, and scattered long black and white setae; tibia and tarsi set with short black setae, and scattered long black setae, mainly at tibia; T5 ventrally with two rows of thick, long, black setae. 
<emphasis id="E5FEEAE3712FFE8EFC8BF949FC661F3B" box="[835,910,1772,1798]" italics="true" pageId="300" pageNumber="301">Proleg</emphasis>
sense hair absent; tibia with antennal cleaning setae ventroapically; coxa, trochanter and femur with internal surface pale but external dark brown, femur apex completely dark brown, tibia mostly pale except for three dark brown marks on external surface; tarsi brown but T5 slightly lighter. 
<emphasis id="E5FEEAE3712FFE8EFEA6F8FDFE251F4F" box="[366,461,1880,1906]" italics="true" pageId="300" pageNumber="301">Mesoleg</emphasis>
with same colour pattern of proleg, except for coxa, trochanter and femur entirely pale (femur apex brown). 
<emphasis id="E5FEEAE3712FFE8EFE41F8D9FE0D1FAB" box="[393,485,1916,1942]" italics="true" pageId="300" pageNumber="301">Metaleg</emphasis>
with femur, tibia and T1 slightly longer than in other legs; colour pattern similar to mesoleg, except for tibia pale with apex brown.
</paragraph>
<paragraph id="D73536F1712FFE8FFF0FF861FC2D188E" blockId="300.[151,1437,512,2015]" lastBlockId="301.[151,1437,153,431]" lastPageId="301" lastPageNumber="302" pageId="300" pageNumber="301">Abdomen: Tergites mostly dark brown with some irregular pale marks; beset with short black setae. Sternites mostly pale with scattered dark marks; set mostly with short white setae.</paragraph>
<paragraph id="D73536F1712EFE8FFF0FFF18FDED18EA" blockId="301.[151,1437,153,431]" box="[199,517,189,215]" pageId="301" pageNumber="302">Male Terminalia: Unknown.</paragraph>
<paragraph id="D73536F1712EFE8FFF0FFF44FD7C1992" blockId="301.[151,1437,153,431]" pageId="301" pageNumber="302">
Female Terminalia (
<figureCitation id="4FB12A74712EFE8FFE61FF44FDFB18C1" box="[425,531,225,252]" captionStart="FIGURE 161" captionStartId="302.[151,250,1878,1903]" captionTargetBox="[153,1429,185,1853]" captionTargetId="figure@302.[151,1436,181,1853]" captionTargetPageId="302" captionText="FIGURE 161. Riekoleon longitudinalis sp. nov.: female: a) terminalia, lateral; b) terminalia, ventral (note pregenital plate hidden beneath sternite). Scale bars = 1mm." figureDoi="http://doi.org/10.5281/zenodo.3897150" httpUri="https://zenodo.org/record/3897150/files/figure.png" pageId="301" pageNumber="302">Figs. 161</figureCitation>
a–b): 
<emphasis id="E5FEEAE3712EFE8FFD9CFF47FD2918C6" box="[596,705,226,251]" italics="true" pageId="301" pageNumber="302">Ectoproct</emphasis>
posterior margin rounded covered, with thin elongate setae. 
<emphasis id="E5FEEAE3712EFE8FFAA6FF47FE811922" italics="true" pageId="301" pageNumber="302">Lateral gonapophyses</emphasis>
rounded, smaller than ectoproct, covered with elongate setae. 7 
<emphasis id="E5FEEAE3712EFE8FFBE1FEA1FB671922" box="[1065,1167,260,287]" italics="true" pageId="301" pageNumber="302">
<superScript id="20FF9BB9712EFE8FFBE1FEA1FBDE192F" attach="left" box="[1065,1078,260,274]" fontSize="6" pageId="301" pageNumber="302">th</superScript>
sternite
</emphasis>
distal margin straight in ventral view; set with long black setae. 
<emphasis id="E5FEEAE3712EFE8FFD9EFE8CFCE3197E" box="[598,779,297,323]" italics="true" pageId="301" pageNumber="302">Pregenital plate</emphasis>
shaped a small arrowhead. 
<emphasis id="E5FEEAE3712EFE8FFB8DFE8FFABD197E" box="[1093,1365,297,323]" italics="true" pageId="301" pageNumber="302">Posterior gonapophyses</emphasis>
broad and elongate, semi rectangular; covered with long black setae, and numerous long cavisetae at apex. 
<emphasis id="E5FEEAE3712EFE8FFB32FEE8FA8C195A" box="[1274,1380,332,359]" italics="true" pageId="301" pageNumber="302">
9 
<superScript id="20FF9BB9712EFE8FFAC0FEE9FAFD1967" attach="left" box="[1288,1301,332,346]" fontSize="6" pageId="301" pageNumber="302">th</superScript>
tergite
</emphasis>
with a membranous digitiform process. 
<emphasis id="E5FEEAE3712EFE8FFDD5FED4FCCA19B6" box="[541,802,369,395]" italics="true" pageId="301" pageNumber="302">Anterior gonapophyses</emphasis>
large rounded plate, covered with many long setae. 
<emphasis id="E5FEEAE3712EFE8FFAA2FED7FED51992" italics="true" pageId="301" pageNumber="302">Ventral membrane</emphasis>
gonapophyseal plates present.
</paragraph>
</subSubSection>
<caption id="83F56679712EFE8FFF5FF8D6FAE81FB1" ID-DOI="http://doi.org/10.5281/zenodo.3897142" ID-Zenodo-Dep="3897142" box="[151,1280,1907,1932]" httpUri="https://zenodo.org/record/3897142/files/figure.png" pageId="301" pageNumber="302" startId="301.[151,250,1907,1932]" targetBox="[153,1436,457,1868]" targetPageId="301">
<paragraph id="D73536F1712EFE8FFF5FF8D6FAE81FB1" blockId="301.[151,1280,1907,1932]" box="[151,1280,1907,1932]" pageId="301" pageNumber="302">
<emphasis id="E5FEEAE3712EFE8FFF5FF8D6FEC51FB1" bold="true" box="[151,301,1907,1932]" pageId="301" pageNumber="302">FIGURE 160.</emphasis>
<taxonomicName id="108A4D72712EFE8FFEFBF8D6FDC31FB6" authority="Machado &amp; Oswald, 2020" authorityName="Machado &amp; Oswald" authorityYear="2020" box="[307,555,1907,1931]" class="Insecta" family="Myrmeleontidae" genus="Riekoleon" kingdom="Animalia" order="Neuroptera" pageId="301" pageNumber="302" phylum="Arthropoda" rank="species" species="longitudinalis" status="sp. nov.">
<emphasis id="E5FEEAE3712EFE8FFEFBF8D6FDC31FB6" box="[307,555,1907,1931]" italics="true" pageId="301" pageNumber="302">Riekoleon longitudinalis</emphasis>
</taxonomicName>
<emphasis id="E5FEEAE3712EFE8FFDF9F8D1FD6A1FB1" bold="true" box="[561,642,1908,1932]" pageId="301" pageNumber="302">
<taxonomicNameLabel id="FECD5798712EFE8FFDF9F8D1FD6A1FB1" box="[561,642,1908,1932]" pageId="301" pageNumber="302" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: a) head, anterior; b) head and thorax, dorsal; c) wings, dorsal.
</paragraph>
</caption>
<caption id="83F56679712DFE8CFF5FF8F3FDDC1FAE" ID-DOI="http://doi.org/10.5281/zenodo.3897150" ID-Zenodo-Dep="3897150" httpUri="https://zenodo.org/record/3897150/files/figure.png" pageId="302" pageNumber="303" startId="302.[151,250,1878,1903]" targetBox="[153,1429,185,1853]" targetPageId="302">
<paragraph id="D73536F1712DFE8CFF5FF8F3FDDC1FAE" blockId="302.[151,1436,1878,1939]" pageId="302" pageNumber="303">
<emphasis id="E5FEEAE3712DFE8CFF5FF8F3FEC41F52" bold="true" box="[151,300,1878,1903]" pageId="302" pageNumber="303">FIGURE 161.</emphasis>
<taxonomicName id="108A4D72712DFE8CFEFAF8F2FDC21F52" authority="Machado &amp; Oswald, 2020" authorityName="Machado &amp; Oswald" authorityYear="2020" box="[306,554,1879,1903]" class="Insecta" family="Myrmeleontidae" genus="Riekoleon" kingdom="Animalia" order="Neuroptera" pageId="302" pageNumber="303" phylum="Arthropoda" rank="species" species="longitudinalis" status="sp. nov.">
<emphasis id="E5FEEAE3712DFE8CFEFAF8F2FDC21F52" box="[306,554,1879,1903]" italics="true" pageId="302" pageNumber="303">Riekoleon longitudinalis</emphasis>
</taxonomicName>
<emphasis id="E5FEEAE3712DFE8CFDF8F8F2FD691F52" bold="true" box="[560,641,1879,1903]" pageId="302" pageNumber="303">
<taxonomicNameLabel id="FECD5798712DFE8CFDF8F8F2FD691F52" box="[560,641,1879,1903]" pageId="302" pageNumber="303" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: 
<emphasis id="E5FEEAE3712DFE8CFD46F8F3FD3E1F52" bold="true" box="[654,726,1878,1903]" pageId="302" pageNumber="303">female</emphasis>
: a) terminalia, lateral; b) terminalia, ventral (note pregenital plate hidden beneath sternite). Scale bars = 1mm.
</paragraph>
</caption>
<caption id="83F56679712CFE8DFF5FFB52FD211D2D" ID-DOI="http://doi.org/10.5281/zenodo.3897152" ID-Zenodo-Dep="3897152" box="[151,713,1271,1296]" httpUri="https://zenodo.org/record/3897152/files/figure.png" pageId="303" pageNumber="304" startId="303.[151,250,1271,1296]" targetBox="[151,1434,181,1246]" targetPageId="303">
<paragraph id="D73536F1712CFE8DFF5FFB52FD211D2D" blockId="303.[151,713,1271,1296]" box="[151,713,1271,1296]" pageId="303" pageNumber="304">
<emphasis id="E5FEEAE3712CFE8DFF5FFB52FEC51D2D" bold="true" box="[151,301,1271,1296]" pageId="303" pageNumber="304">FIGURE 162.</emphasis>
Distribution of 
<taxonomicName id="108A4D72712CFE8DFE07FB5DFDDC1D2D" authorityName="New" authorityYear="1985" box="[463,564,1272,1296]" class="Insecta" family="Myrmeleontidae" genus="Riekoleon" kingdom="Animalia" order="Neuroptera" pageId="303" pageNumber="304" phylum="Arthropoda" rank="genus">
<emphasis id="E5FEEAE3712CFE8DFE07FB5DFDDC1D2D" box="[463,564,1272,1296]" italics="true" pageId="303" pageNumber="304">Riekoleon</emphasis>
</taxonomicName>
species (part).
</paragraph>
</caption>
<subSubSection id="9F90657A712CFE8DFF0FFA98FC051D40" pageId="303" pageNumber="304" type="distribution">
<paragraph id="D73536F1712CFE8DFF0FFA98FAFB1D65" blockId="303.[151,1437,1341,1945]" box="[199,1299,1341,1369]" pageId="303" pageNumber="304">
<emphasis id="E5FEEAE3712CFE8DFF0FFA98FEB11D65" bold="true" box="[199,345,1341,1368]" pageId="303" pageNumber="304">Distribution</emphasis>
(
<figureCitation id="4FB12A74712CFE8DFEA1FA9BFE201D64" box="[361,456,1342,1369]" captionStart="FIGURE 162" captionStartId="303.[151,250,1271,1296]" captionTargetBox="[151,1434,181,1246]" captionTargetId="figure@303.[151,1435,181,1246]" captionTargetPageId="303" captionText="FIGURE 162. Distribution of Riekoleon species (part)." figureDoi="http://doi.org/10.5281/zenodo.3897152" httpUri="https://zenodo.org/record/3897152/files/figure.png" pageId="303" pageNumber="304">Fig. 162</figureCitation>
). 
<collectingCountry id="AF9D7661712CFE8DFE16FA9BFDA11D65" box="[478,585,1342,1368]" name="Australia" pageId="303" pageNumber="304">Australia</collectingCountry>
: QLD. Known from sparse records in northeastern 
<collectingRegion id="154EF813712CFE8DFB4FFA9BFAE71D65" box="[1159,1295,1342,1368]" country="Australia" name="Queensland" pageId="303" pageNumber="304">Queensland</collectingRegion>
.
</paragraph>
<paragraph id="D73536F1712CFE8DFF0FFAC4FC051D40" blockId="303.[151,1437,1341,1945]" box="[199,1005,1377,1405]" pageId="303" pageNumber="304">
<emphasis id="E5FEEAE3712CFE8DFF0FFAC4FE2F1D41" bold="true" box="[199,455,1377,1404]" pageId="303" pageNumber="304">Adult activity period.</emphasis>
Records for September, November, and January.
</paragraph>
</subSubSection>
<subSubSection id="9F90657A712CFE8DFF0FFA20FDB11D9D" box="[199,601,1413,1440]" pageId="303" pageNumber="304" type="biology_ecology">
<paragraph id="D73536F1712CFE8DFF0FFA20FDB11D9D" blockId="303.[151,1437,1341,1945]" box="[199,601,1413,1440]" pageId="303" pageNumber="304">
<emphasis id="E5FEEAE3712CFE8DFF0FFA20FECF1D9D" bold="true" box="[199,295,1413,1440]" pageId="303" pageNumber="304">Biology.</emphasis>
Unknown, larva unknown.
</paragraph>
</subSubSection>
<subSubSection id="9F90657A712CFE8DFF0FFA0FFD2E1EFC" pageId="303" pageNumber="304" type="etymology">
<paragraph id="D73536F1712CFE8DFF0FFA0FFBE61DD5" blockId="303.[151,1437,1341,1945]" pageId="303" pageNumber="304">
<emphasis id="E5FEEAE3712CFE8DFF0FFA0FFEA01DF9" bold="true" box="[199,328,1450,1476]" pageId="303" pageNumber="304">Etymology</emphasis>
. 
<emphasis id="E5FEEAE3712CFE8DFE9FFA0FFE3B1DF9" box="[343,467,1450,1476]" italics="true" pageId="303" pageNumber="304">longitudin-</emphasis>
(from Latin longitudinis, lengths) + - 
<emphasis id="E5FEEAE3712CFE8DFCB1FA0FFC491DF9" box="[889,929,1450,1476]" italics="true" pageId="303" pageNumber="304">alis</emphasis>
(from Latin -alis, pertaining to), in reference to the long, dark, longitudinal markings present on the pronotum and forewing.
</paragraph>
<paragraph id="D73536F1712CFE8DFF0FFA54FD281E69" blockId="303.[151,1437,1341,1945]" pageId="303" pageNumber="304">
<emphasis id="E5FEEAE3712CFE8DFF0FFA54FE441E31" bold="true" box="[199,428,1521,1548]" pageId="303" pageNumber="304">Name-bearing type</emphasis>
. 
<emphasis id="E5FEEAE3712CFE8DFE72FA57FD3E1E31" box="[442,726,1522,1548]" italics="true" pageId="303" pageNumber="304">
<taxonomicName id="108A4D72712CFE8DFE72FA57FD381E31" authorityName="Machado &amp; Oswald" authorityYear="2020" box="[442,720,1522,1548]" class="Insecta" family="Myrmeleontidae" genus="Riekoleon" kingdom="Animalia" order="Neuroptera" pageId="303" pageNumber="304" phylum="Arthropoda" rank="species" species="longitudinalis">Riekoleon longitudinalis</taxonomicName>
:
</emphasis>
<materialsCitation id="67E23CAC712CFE8DFD15FA57FD281E69" collectingDate="1990-01-04" collectorName="M. S. &amp; B. J. Moulds. Condition" location="Routh Creek" pageId="303" pageNumber="304" specimenCount="1" stateProvince="Queensland" typeStatus="holotype">
<typeStatus id="08318853712CFE8DFD15FA57FCAD1E31" box="[733,837,1522,1548]" pageId="303" pageNumber="304" type="holotype">Holotype</typeStatus>
(by present designation), female, AMSA. 
<emphasis id="E5FEEAE3712CFE8DFAE8FA54FF211E0D" bold="true" pageId="303" pageNumber="304">AUSTRA- LIA</emphasis>
: 
<emphasis id="E5FEEAE3712CFE8DFF13F9B0FE811E0D" bold="true" box="[219,361,1557,1584]" pageId="303" pageNumber="304">
<collectingRegion id="154EF813712CFE8DFF13F9B0FE811E0D" box="[219,361,1557,1584]" country="Australia" name="Queensland" pageId="303" pageNumber="304">Queensland</collectingRegion>
</emphasis>
: 
<emphasis id="E5FEEAE3712CFE8DFEB2F9B3FDE01E0D" box="[378,520,1558,1584]" italics="true" pageId="303" pageNumber="304">
<location id="D255602A712CFE8DFEB2F9B3FDE01E0D" LSID="urn:lsid:plazi:treatment:5F2387E7712FFE8DFF5FFED5FEE21FA5:D255602A712CFE8DFEB2F9B3FDE01E0D" box="[378,520,1558,1584]" name="Routh Creek" pageId="303" pageNumber="304" stateProvince="Queensland">Routh Creek</location>
</emphasis>
: 
<locationDeviation id="C6872FD3712CFE8DFDD2F9B3FCDE1E0D" box="[538,822,1558,1585]" location="Georgetown" pageId="303" pageNumber="304">
<quantity id="10729B14712CFE8DFDD2F9B3FD8C1E0D" box="[538,612,1558,1584]" metricMagnitude="4" metricUnit="m" metricValue="2.1" pageId="303" pageNumber="304" unit="km" value="21.0">21 km</quantity>
E of Georgetown
</locationDeviation>
, 
<date id="A3341031712CFE8DFC8CF9B3FC551E0D" box="[836,957,1558,1584]" pageId="303" pageNumber="304" value="1990-01-04">
<collectingDate id="B370E9D9712CFE8DFC8CF9B3FC551E0D" box="[836,957,1558,1584]" pageId="303" pageNumber="304" value="1990-01-04">4.jan.1990</collectingDate>
</date>
, 
<collectorName id="7A7F5327712CFE8DFC03F9B3FAD41E0D" box="[971,1340,1558,1585]" pageId="303" pageNumber="304">M.S. &amp; B.J. Moulds. Condition</collectorName>
: pinned; good; but antennae missing; terminalia dissected.
</materialsCitation>
</paragraph>
<paragraph id="D73536F1712CFE8DFF0FF9FBFD2E1EFC" blockId="303.[151,1437,1341,1945]" pageId="303" pageNumber="304">
<materialsCitation id="67E23CAC712CFE8DFF0FF9FBFD2A1EFC" collectingDate="1998-11-25" collectionCode="SAMA, ANIC" collectorName="Sept., A. M &amp; E. D. Edwards &amp; H. Sutrisno" country="Australia" determinerName="T. R. New" elevation="820" latitude="-17.39" location="Millstream Falls" municipality="Magnetic Island" pageId="303" pageNumber="304" specimenCount="4" specimenCount-female="4" stateProvince="Queensland" typeStatus="paratype">
<emphasis id="E5FEEAE3712CFE8DFF0FF9FBFED71E45" bold="true" box="[199,319,1630,1656]" pageId="303" pageNumber="304">
<typeStatus id="08318853712CFE8DFF0FF9FBFED71E45" box="[199,319,1630,1656]" pageId="303" pageNumber="304" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="C18CFD78712CFE8DFE86F9FBFE981E45" box="[334,368,1630,1656]" pageId="303" pageNumber="304" type="female">2♀</specimenCount>
). 
<emphasis id="E5FEEAE3712CFE8DFE4DF9F8FDC21E45" bold="true" box="[389,554,1629,1656]" pageId="303" pageNumber="304">
<collectingCountry id="AF9D7661712CFE8DFE4DF9F8FDC21E45" box="[389,554,1629,1656]" name="Australia" pageId="303" pageNumber="304">AUSTRALIA</collectingCountry>
</emphasis>
: 
<emphasis id="E5FEEAE3712CFE8DFDFFF9F8FD261E45" bold="true" box="[567,718,1629,1656]" pageId="303" pageNumber="304">
<collectingRegion id="154EF813712CFE8DFDFFF9F8FD211E45" box="[567,713,1629,1656]" country="Australia" name="Queensland" pageId="303" pageNumber="304">Queensland</collectingRegion>
:
</emphasis>
<collectingMunicipality id="3751AC8B712CFE8DFD1BF9FBFC631E45" box="[723,907,1630,1656]" pageId="303" pageNumber="304">Magnetic Island</collectingMunicipality>
: 
<collectorName id="7A7F5327712CFE8DFC5CF9FBFBE71E45" box="[916,1039,1630,1656]" pageId="303" pageNumber="304">Sept., A.M</collectorName>
. Lea, 
<emphasis id="E5FEEAE3712CFE8DFB86F9FBFA8F1E45" box="[1102,1383,1630,1657]" italics="true" pageId="303" pageNumber="304">
<taxonomicName id="108A4D72712CFE8DFB86F9FBFB2B1E45" authorityName="Banks" authorityYear="1913" box="[1102,1219,1630,1656]" class="Insecta" family="Myrmeleontidae" genus="Glenoleon" kingdom="Animalia" order="Neuroptera" pageId="303" pageNumber="304" phylum="Arthropoda" rank="genus">Glenoleon</taxonomicName>
sp? 
<taxonomicName id="108A4D72712CFE8DFB32F9FBFA8F1E45" authorityName="Machado &amp; Oswald" authorityYear="2020" baseAuthorityName="Tillyard" baseAuthorityYear="1916" box="[1274,1383,1630,1656]" class="Insecta" family="Myrmeleontidae" genus="Austrogymnocnemia" kingdom="Animalia" order="Neuroptera" pageId="303" pageNumber="304" phylum="Arthropoda" rank="species" species="berthoudi">berthoudi</taxonomicName>
</emphasis>
, det. 
<determinerName id="22E9DA35712CFE8DFF5FF926FEEA1EA0" box="[151,258,1667,1693]" pageId="303" pageNumber="304">T.R. New</determinerName>
1982 (
<specimenCount id="C18CFD78712CFE8DFE98F927FE9A1EA1" box="[336,370,1666,1692]" pageId="303" pageNumber="304" type="female">1♀</specimenCount>
, 
<collectionCode id="B19BAE34712CFE8DFEB6F927FE3A1EA1" box="[382,466,1666,1692]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="303" pageNumber="304" type="Museum">SAMA</collectionCode>
, pinned); 
<location id="D255602A712CFE8DFDF6F927FD121EA1" LSID="urn:lsid:plazi:treatment:5F2387E7712FFE8DFF5FFED5FEE21FA5:D255602A712CFE8DFDF6F927FD121EA1" box="[574,762,1666,1692]" country="Australia" latitude="-17.39" municipality="Magnetic Island" name="Millstream Falls" pageId="303" pageNumber="304" stateProvince="Queensland">Millstream Falls</location>
: 
<locationDeviation id="C6872FD3712CFE8DFCCAF924FC121EA1" box="[770,1018,1665,1693]" location="Ravenshoe" pageId="303" pageNumber="304">
<quantity id="10729B14712CFE8DFCCAF924FCD21EA1" box="[770,826,1665,1692]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="303" pageNumber="304" unit="km" value="5.0">5 km</quantity>
W of Ravenshoe
</locationDeviation>
, 
<quantity id="10729B14712CFE8DFBCBF927FBAD1EA1" box="[1027,1093,1666,1692]" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="303" pageNumber="304" unit="m" value="820.0">
<elevation id="5CA7D1C2712CFE8DFBCBF927FBAD1EA1" box="[1027,1093,1666,1692]" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="303" pageNumber="304" unit="m" value="820.0">820m</elevation>
</quantity>
, 
<geoCoordinate id="B2BE5036712CFE8DFB87F924FB441EA1" box="[1103,1196,1665,1692]" degrees="17.39" direction="south" orientation="latitude" pageId="303" pageNumber="304" precision="555" value="-17.39">17.39S–</geoCoordinate>
145.27W, 
<date id="A3341031712CFE8DFAD5F924FA701EA1" box="[1309,1432,1665,1692]" pageId="303" pageNumber="304" value="1998-11-25">
<collectingDate id="B370E9D9712CFE8DFAD5F924FA701EA1" box="[1309,1432,1665,1692]" pageId="303" pageNumber="304" value="1998-11-25">25.xi.1998</collectingDate>
</date>
, 
<collectorName id="7A7F5327712CFE8DFF5FF902FEDC1EFD" box="[151,308,1702,1729]" pageId="303" pageNumber="304">E.D. Edwards</collectorName>
&amp; 
<collectorName id="7A7F5327712CFE8DFE9FF902FE3F1EFD" box="[343,471,1702,1729]" pageId="303" pageNumber="304">H. Sutrisno</collectorName>
(
<specimenCount id="C18CFD78712CFE8DFE22F903FDE31EFD" box="[490,523,1702,1728]" pageId="303" pageNumber="304" type="female">1♀</specimenCount>
, 
<collectionCode id="B19BAE34712CFE8DFDDFF903FDB61EFD" box="[535,606,1702,1728]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="303" pageNumber="304">ANIC</collectionCode>
, pinned)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="9F90657A712CFE8DFF0FF96CFEE21FA5" pageId="303" pageNumber="304" type="discussion">
<paragraph id="D73536F1712CFE8DFF0FF96CFEE21FA5" blockId="303.[151,1437,1341,1945]" pageId="303" pageNumber="304">
<emphasis id="E5FEEAE3712CFE8DFF0FF96CFEA71ED9" bold="true" box="[199,335,1737,1764]" pageId="303" pageNumber="304">Comments.</emphasis>
<taxonomicName id="108A4D72712CFE8DFE9DF96FFD8F1ED9" authority="Machado &amp; Oswald, 2020" authorityName="Machado &amp; Oswald" authorityYear="2020" box="[341,615,1738,1764]" class="Insecta" family="Myrmeleontidae" genus="Riekoleon" kingdom="Animalia" order="Neuroptera" pageId="303" pageNumber="304" phylum="Arthropoda" rank="species" species="longitudinalis" status="sp. nov.">
<emphasis id="E5FEEAE3712CFE8DFE9DF96FFD8F1ED9" box="[341,615,1738,1764]" italics="true" pageId="303" pageNumber="304">Riekoleon longitudinalis</emphasis>
</taxonomicName>
<emphasis id="E5FEEAE3712CFE8DFDA4F96FFD2D1ED9" bold="true" box="[620,709,1738,1764]" pageId="303" pageNumber="304">
<taxonomicNameLabel id="FECD5798712CFE8DFDA4F96FFD2D1ED9" box="[620,709,1738,1764]" pageId="303" pageNumber="304" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is the largest species in the genus based on forewing length and is easily identified by the dark longitudinal lines of the forewing and pronotum. The short pronotal setae, and large and rectangular female posterior gonapophysis place this species close to 
<taxonomicName id="108A4D72712CFE8DFC6FF8B6FBE21F11" box="[935,1034,1810,1836]" class="Insecta" family="Myrmeleontidae" genus="Riekoleon" kingdom="Animalia" order="Neuroptera" pageId="303" pageNumber="304" phylum="Arthropoda" rank="species" species="rudda">
<emphasis id="E5FEEAE3712CFE8DFC6FF8B6FBE21F11" box="[935,1034,1810,1836]" italics="true" pageId="303" pageNumber="304">R. rudda</emphasis>
</taxonomicName>
. The shape of the dark longitudinal line of the forewing is similar to 
<taxonomicName id="108A4D72712CFE8DFDC6F892FC801F6D" authorityName="Machado &amp; Oswald" authorityYear="2020" baseAuthorityName="Tillyard" baseAuthorityYear="1916" box="[526,872,1846,1873]" class="Insecta" family="Myrmeleontidae" genus="Austrogymnocnemia" kingdom="Animalia" order="Neuroptera" pageId="303" pageNumber="304" phylum="Arthropoda" rank="species" species="berthoudi">
<emphasis id="E5FEEAE3712CFE8DFDC6F892FC801F6D" box="[526,872,1846,1873]" italics="true" pageId="303" pageNumber="304">Austrogymnocnemia berthoudi</emphasis>
</taxonomicName>
, but the large apical spots in the hind wing (of 
<taxonomicName id="108A4D72712CFE8DFA4CF892FEDB1F49" authorityName="Machado &amp; Oswald" authorityYear="2020" class="Insecta" family="Myrmeleontidae" genus="Riekoleon" kingdom="Animalia" order="Neuroptera" pageId="303" pageNumber="304" phylum="Arthropoda" rank="species" species="longitudinalis">
<emphasis id="E5FEEAE3712CFE8DFA4CF892FEDB1F49" italics="true" pageId="303" pageNumber="304">R. longitudinalis</emphasis>
</taxonomicName>
) and the general form of the female terminalia clearly separate them. For phylogenetic relationships see 
<figureCitation id="4FB12A74712CFE8DFF0AF8DBFEEE1FA5" box="[194,262,1918,1944]" captionStart="FIGURE 4" captionStartId="30.[151,250,1373,1398]" captionTargetBox="[151,1436,348,1348]" captionTargetId="figure@30.[151,1436,345,1348]" captionTargetPageId="30" captionText="FIGURE 4. Strict consensus tree of morphology-based parsimony analysis of species belonging to the former subtribe Periclystina (ingroup, names in black), and outgroups (names in red). Former Glenoleon species are shown on blue branches, former Austrogymnocnemia species on green branches. Numbers above branches are Bremer support values; numbers below branches are bootstrap values (shown where&gt;50%)." figureDoi="http://doi.org/10.5281/zenodo.3896822" httpUri="https://zenodo.org/record/3896822/files/figure.png" pageId="303" pageNumber="304">Fig. 4</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>