treatments-xml/data/61/51/D1/6151D1301E4FFF8663A6FB2BA9E2DCC8.xml

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<mods:title id="BE8F286DE630C72BEDCD2924ABAE02BD">A taxonomic revision of the African leaf chafer genus Asthenopholis Brenske, 1898 (Coleoptera: Scarabaeidae: Melolonthinae): a SEM study</mods:title>
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<paragraph id="E94760261E4FFF9363A6FB2BAF50DB93" blockId="13.[151,1047,1234,1295]" box="[151,625,1234,1260]" pageId="13" pageNumber="14">
<heading id="B20FD74A1E4FFF9363A6FB2BAF50DB93" bold="true" box="[151,625,1234,1260]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
<taxonomicName id="2EF81BA51E4FFF9363A6FB2BAF50DB93" authority="Moser, 1913" authorityName="Moser" authorityYear="1913" box="[151,625,1234,1260]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ugandensis">
<emphasis id="DB8CBC341E4FFF9363A6FB2BAF50DB93" bold="true" box="[151,625,1234,1260]" pageId="13" pageNumber="14">
<emphasis id="DB8CBC341E4FFF9363A6FB2BACF5DB93" bold="true" box="[151,468,1234,1260]" italics="true" pageId="13" pageNumber="14">Asthenopholis ugandensis</emphasis>
<bibRefCitation id="8D691DD71E4FFF9362EAFB2BAF50DB93" author="Moser" box="[475,625,1234,1260]" pageId="13" pageNumber="44" refString="Moser, J. (1913) Beitrag zur kenntnis der Melolonthiden II. Annales de la Societe Entomologique de Belgique, 57 (5 - 12), 331 - 345." type="journal article" year="1913">Moser, 1913</bibRefCitation>
</emphasis>
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</paragraph>
<paragraph id="E94760261E4FFF9363A6FB0CA936DA70" blockId="13.[151,1047,1234,1295]" box="[151,1047,1269,1295]" pageId="13" pageNumber="14">(Figs. 4, 23, 30–31, 45, 69–73, 100–101, 119–121, 140–141, 159–161, 177).</paragraph>
</subSubSection>
<subSubSection id="A1E233AD1E4FFF9363A6FABBACCDDA06" pageId="13" pageNumber="14" type="reference_group">
<paragraph id="E94760261E4FFF9363A6FABBACCDDA06" blockId="13.[151,1436,1346,1401]" pageId="13" pageNumber="14">
<treatmentCitationGroup id="C9E847081E4FFF9363A6FABBACCDDA06" pageId="13" pageNumber="14">
<taxonomicName id="2EF81BA51E4FFF9363A6FABBAF61DA26" authority="Moser, 1913" authorityName="Moser" authorityYear="1913" box="[151,576,1346,1369]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ugandensis">
<emphasis id="DB8CBC341E4FFF9363A6FABBAC8CDA26" box="[151,429,1346,1369]" italics="true" pageId="13" pageNumber="14">Asthenopholis ugandensis</emphasis>
<bibRefCitation id="8D691DD71E4FFF936284FABBAF61DA26" author="Moser" box="[437,576,1346,1369]" pageId="13" pageNumber="44" refString="Moser, J. (1913) Beitrag zur kenntnis der Melolonthiden II. Annales de la Societe Entomologique de Belgique, 57 (5 - 12), 331 - 345." type="journal article" year="1913">Moser, 1913</bibRefCitation>
</taxonomicName>
: (Length 
<quantity id="2E00CDC31E4FFF936196FABBAE3CDA26" box="[679,797,1346,1369]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="13" pageNumber="14" unit="mm" value="21.0" valueMax="22.0" valueMin="20.0">20–22 mm</quantity>
, 
<collectingCountry id="91EF20B61E4FFF936018FABBAE5EDA26" box="[809,895,1346,1369]" name="Uganda" pageId="13" pageNumber="14">Uganda</collectingCountry>
, Entebbe, 
<typeStatus id="3643DE841E4FFF9360C1FABBA973DA26" box="[1008,1106,1346,1369]" pageId="13" pageNumber="14" type="lectotype">lectotype</typeStatus>
and 
<typeStatus id="3643DE841E4FFF9367B9FABBA803DA26" box="[1160,1314,1346,1369]" pageId="13" pageNumber="14" type="paralectotype">paralectotypes</typeStatus>
in ZMHB) 341–342. 
<treatmentCitation id="685946371E4FFF936200FA9BACC9DA06" author="Kolbe" box="[305,488,1378,1401]" page="352" pageId="13" pageNumber="14" year="1914">
<bibRefCitation id="8D691DD71E4FFF936200FA9BAC93DA06" author="Kolbe" box="[305,434,1378,1401]" pageId="13" pageNumber="43" refString="Kolbe, H. (1914) Scarabaeiden. In: Schubotz, H. (ed.) Wissenschaftliche Ergebnisse der Deutschen Zentral-Afrika- Expedition 1907 - 1908. Band V, Zoologie III, Lieferung 3. Klinkhardt &amp; Biermann, Leipzig, pp. 261 - 416." type="journal article" year="1914">Kolbe 1914</bibRefCitation>
: 352
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="A1E233AD1E4FFF9063A6FA51A9E5DE59" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="diagnosis">
<paragraph id="E94760261E4FFF9363A6FA51A997DA95" blockId="13.[151,1437,1448,2016]" pageId="13" pageNumber="14">
<emphasis id="DB8CBC341E4FFF9363A6FA51AC38DABD" bold="true" box="[151,281,1448,1474]" pageId="13" pageNumber="14">Diagnosis.</emphasis>
The smallest (
<quantity id="2E00CDC31E4FFF9362FAFA50AF6EDABC" box="[459,591,1449,1475]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="13" pageNumber="14" unit="mm" value="19.0" valueMax="20.0" valueMin="18.0">18–20 mm</quantity>
) known north-eastern African 
<taxonomicName id="2EF81BA51E4FFF9360F2FA50A94CDABD" box="[963,1133,1449,1474]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="DB8CBC341E4FFF9360F2FA50A94CDABD" box="[963,1133,1449,1474]" italics="true" pageId="13" pageNumber="14">Asthenopholis</emphasis>
</taxonomicName>
species. Brown to black, scutellum bare. Cuticular setae and scales finer and whiter (Fig. 4) than 
<taxonomicName id="2EF81BA51E4FFF9360D3FA29A96FDA96" box="[994,1102,1488,1513]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="crassa">
<emphasis id="DB8CBC341E4FFF9360D3FA29A96FDA96" box="[994,1102,1488,1513]" italics="true" pageId="13" pageNumber="14">A. crassa</emphasis>
</taxonomicName>
(Fig. 3).
</paragraph>
<paragraph id="E94760261E4FFF9063F7FA0FA9E5DE59" blockId="13.[151,1437,1448,2016]" lastBlockId="14.[151,1437,152,990]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<emphasis id="DB8CBC341E4FFF9363F7FA0FAC5DD96F" bold="true" box="[198,380,1526,1552]" pageId="13" pageNumber="14">Redescription.</emphasis>
<emphasis id="DB8CBC341E4FFF9362B5FA0EAC92D96F" box="[388,435,1527,1552]" italics="true" pageId="13" pageNumber="14">Size</emphasis>
(n = 
<specimenCount id="FFFEABAF1E4FFF9362C2FA0FAF73D96F" box="[499,594,1526,1552]" pageId="13" pageNumber="14" type="male">5 males</specimenCount>
; n = 
<specimenCount id="FFFEABAF1E4FFF9361BCFA0FAE25D96F" box="[653,772,1526,1552]" pageId="13" pageNumber="14" type="female">2 females</specimenCount>
). Length: males, mean 
<quantity id="2E00CDC31E4FFF936724FA0FA9A0D96F" box="[1045,1153,1526,1552]" metricMagnitude="-2" metricUnit="m" metricValue="1.89" pageId="13" pageNumber="14" unit="mm" value="18.9">18.9 mm</quantity>
, (range 18.0– 
<quantity id="2E00CDC31E4FFF936612FA0FA8AED96F" box="[1315,1423,1526,1552]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" pageId="13" pageNumber="14" unit="mm" value="19.5">19.5 mm</quantity>
), females, 
<quantity id="2E00CDC31E4FFF936230F9E4AC4CD948" box="[257,365,1565,1591]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" pageId="13" pageNumber="14" unit="mm" value="19.5">19.5 mm</quantity>
, (19.0–20.0 mm). Width: males, mean 
<quantity id="2E00CDC31E4FFF936006F9E4AE82D948" box="[823,931,1565,1591]" metricMagnitude="-2" metricUnit="m" metricValue="1.06" pageId="13" pageNumber="14" unit="mm" value="10.6">10.6 mm</quantity>
, (range 10.0–11.0 mm), females, 11.0 mm, (11.0 mm). 
<emphasis id="DB8CBC341E4FFF93622EF9BDAC53D922" box="[287,370,1604,1629]" italics="true" pageId="13" pageNumber="14">Colour</emphasis>
(Fig. 4): Brown to black cuticle covered by white scales and tan setae. 
<emphasis id="DB8CBC341E4FFF936781F9BDA9D1D922" box="[1200,1264,1604,1629]" italics="true" pageId="13" pageNumber="14">Head</emphasis>
(Figs. 30–31): Entire head covered with off-white setae. Clypeus with anterior margin slightly reflexed, corners rounded inward near eye; clypeal margin weakly elevated; clypeal suture present but indistinct. Raised protuberance situated laterally near eye; surrounded by clump of stiff, finer setae intermixed with scales; setae double the scale length. 
<emphasis id="DB8CBC341E4FFF93620CF926AC99D987" box="[317,440,1759,1784]" italics="true" pageId="13" pageNumber="14">Pronotum</emphasis>
(Figs. 69–73): Pronotum ratio 1.56:1 (n = 53); 1.55:1 (n = 2Ƥ). Row of short, lanceolate scales along anterior medial margin of pronotum; disc covered by off-white scales, directed posteriorly. Lateral edges with an obtuse rounded angle in the middle; anterior and posterior margins smooth, lateral margins incised by setiferous punctures (setae finer than on disc), forming an irregularly, flattened serrated edge. 
<emphasis id="DB8CBC341E4FFF936278F880ACE0D8ED" box="[329,449,1913,1938]" italics="true" pageId="13" pageNumber="14">Scutellum</emphasis>
(Figs. 100–101): Triangular, posterior apex rounded, surface smooth and almost without setae, punctation or prominent sculpture under light microscope, imbricate under SEM and with one or two fine setae along the margins. 
<emphasis id="DB8CBC341E4FFF936160F83EAFBDD89F" box="[593,668,1991,2016]" italics="true" pageId="13" pageNumber="14">Elytra</emphasis>
(Figs. 119–121): Elytral striae very weak, almost absent, about three per elytron. Elytra covered with fine off-white scales, uniformly dispersed, except scale-less circular patches situated posterior-laterally (ranging from 0–4 per elytron). 
<taxonomicName id="2EF81BA51E4CFF906085FF46A904DFA7" box="[948,1061,191,216]" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="15" phylum="Chordata" rank="genus">
<emphasis id="DB8CBC341E4CFF906085FF46A904DFA7" box="[948,1061,191,216]" italics="true" pageId="14" pageNumber="15">Pygidium</emphasis>
</taxonomicName>
(Figs. 140–141): Covered with straight, almost parallel sided fine setae; bare area medially on anterior edge; setiferous puncture rims not contiguous, with setae inserted almost on edge. 
<emphasis id="DB8CBC341E4CFF9061F6FEF5AE52DE5A" box="[711,883,268,293]" italics="true" pageId="14" pageNumber="15">Male genitalia</emphasis>
illustrated in Figs. 159–161.
</paragraph>
</subSubSection>
<subSubSection id="A1E233AD1E4CFF9063F4FECBA82BDE0C" pageId="14" pageNumber="15" type="distribution">
<paragraph id="E94760261E4CFF9063F4FECBA9AEDE33" blockId="14.[151,1437,152,990]" box="[197,1167,306,332]" pageId="14" pageNumber="15">
<emphasis id="DB8CBC341E4CFF9063F4FECBAC44DE33" bold="true" box="[197,357,306,332]" pageId="14" pageNumber="15">Distribution.</emphasis>
Known from two localities in 
<collectingCountry id="91EF20B61E4CFF9061FCFECBAE09DE33" box="[717,808,306,332]" name="Uganda" pageId="14" pageNumber="15">Uganda</collectingCountry>
and one in 
<collectingCountry id="91EF20B61E4CFF906080FECBA93BDE33" box="[945,1050,306,332]" name="Tanzania" pageId="14" pageNumber="15">Tanzania</collectingCountry>
(Fig. 23).
</paragraph>
<paragraph id="E94760261E4CFF9063F4FEA1A82BDE0C" blockId="14.[151,1437,152,990]" box="[197,1290,344,371]" pageId="14" pageNumber="15">
<emphasis id="DB8CBC341E4CFF9063F4FEA1AE20DE0D" bold="true" box="[197,769,344,370]" pageId="14" pageNumber="15">Natural history, pest status, and conservation.</emphasis>
Unknown. Phenology indicated in Fig. 177.
</paragraph>
</subSubSection>
<subSubSection id="A1E233AD1E4CFF9063F4FE86AF68DEBF" pageId="14" pageNumber="15" type="discussion">
<paragraph id="E94760261E4CFF9063F4FE86AF68DEBF" blockId="14.[151,1437,152,990]" pageId="14" pageNumber="15">
<emphasis id="DB8CBC341E4CFF9063F4FE86AC60DEE6" bold="true" box="[197,321,383,409]" pageId="14" pageNumber="15">Remarks.</emphasis>
Four specimens in the ZMHB collection with the exact label data as Moser’s two 
<typeStatus id="3643DE841E4CFF906603FE79A8BDDEE5" box="[1330,1436,384,410]" pageId="14" pageNumber="15" type="syntype">syntypes</typeStatus>
have been labelled as 
<typeStatus id="3643DE841E4CFF9062A9FE5FAF64DEBF" box="[408,581,422,448]" pageId="14" pageNumber="15" type="paralectotype">paralectotypes</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="A1E233AD1E4CFF9063F4FE34A866DCA1" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="E94760261E4CFF9063F4FE34ADDCDCEF" blockId="14.[151,1437,152,990]" pageId="14" pageNumber="15">
<emphasis id="DB8CBC341E4CFF9063F4FE34A902DE98" bold="true" box="[197,1059,461,487]" pageId="14" pageNumber="15">
<typeStatus id="3643DE841E4CFF9063F4FE34AC24DE98" box="[197,261,461,487]" pageId="14" pageNumber="15">Type</typeStatus>
material examined (
<taxonomicName id="2EF81BA51E4CFF906126FE34A932DE98" authority="Moser, 1913" authorityName="Moser" authorityYear="1913" box="[535,1043,461,487]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="ugandensis">
<emphasis id="DB8CBC341E4CFF906126FE34AE47DE98" bold="true" box="[535,870,461,487]" italics="true" pageId="14" pageNumber="15">Asthenopholis ugandensis</emphasis>
<bibRefCitation id="8D691DD71E4CFF90605EFE34A932DE98" author="Moser" box="[879,1043,461,487]" pageId="14" pageNumber="44" refString="Moser, J. (1913) Beitrag zur kenntnis der Melolonthiden II. Annales de la Societe Entomologique de Belgique, 57 (5 - 12), 331 - 345." type="journal article" year="1913">Moser, 1913</bibRefCitation>
</taxonomicName>
):
</emphasis>
<typeStatus id="3643DE841E4CFF90671AFE34A986DE98" box="[1067,1191,461,487]" pageId="14" pageNumber="15" type="lectotype">Lectotype</typeStatus>
(
<emphasis id="DB8CBC341E4CFF90678BFE34A8ABDE98" bold="true" box="[1210,1418,461,487]" pageId="14" pageNumber="15">here designated</emphasis>
): 
<emphasis id="DB8CBC341E4CFF9063A6FE0AAC39DD72" bold="true" box="[151,280,499,525]" pageId="14" pageNumber="15">
<collectingCountry id="91EF20B61E4CFF9063A6FE0AAC39DD72" box="[151,280,499,525]" name="Uganda" pageId="14" pageNumber="15">UGANDA</collectingCountry>
</emphasis>
, “Entebbe, 
<collectingCountry id="91EF20B61E4CFF906294FE0DAF25DD71" box="[421,516,500,526]" name="Uganda" pageId="14" pageNumber="15">Uganda</collectingCountry>
, Centr. Afr. [same data on reverse of label] / 
<taxonomicName id="2EF81BA51E4CFF90672CFE0DA8B6DD71" authority="Mos." authorityName="Mos." box="[1053,1431,500,526]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="ugandensis">
<emphasis id="DB8CBC341E4CFF90672CFE0DA870DD72" box="[1053,1361,500,525]" italics="true" pageId="14" pageNumber="15">Asthenopholis ugandensis</emphasis>
Mos.
</taxonomicName>
, 
<typeStatus id="3643DE841E4CFF9063A6FDE3ADEEDD4B" box="[151,207,538,564]" pageId="14" pageNumber="15">Type</typeStatus>
3 [symbol, written on faded white card by Moser] / 
<typeStatus id="3643DE841E4CFF906076FDE3AEA0DD4B" box="[839,897,538,564]" pageId="14" pageNumber="15">Type</typeStatus>
[printed on reddish brown card] / Zool. Mus. Berlin / 0 2602 [JduGH photo number] / 
<typeStatus id="3643DE841E4CFF90614FFDB8AFD4DD24" box="[638,757,577,603]" pageId="14" pageNumber="15" type="lectotype">Lectotype</typeStatus>
3, 
<taxonomicName id="2EF81BA51E4CFF906015FDB8A9DDDD24" authority="Moser, 1913" authorityName="Moser" authorityYear="1913" box="[804,1276,577,603]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="ugandensis">
<emphasis id="DB8CBC341E4CFF906015FDB8A97ADD25" box="[804,1115,577,602]" italics="true" pageId="14" pageNumber="15">Asthenopholis ugandensis</emphasis>
<bibRefCitation id="8D691DD71E4CFF906752FDB8A9DDDD24" author="Moser" box="[1123,1276,577,603]" pageId="14" pageNumber="44" refString="Moser, J. (1913) Beitrag zur kenntnis der Melolonthiden II. Annales de la Societe Entomologique de Belgique, 57 (5 - 12), 331 - 345." type="journal article" year="1913">Moser, 1913</bibRefCitation>
</taxonomicName>
, des. J. du G. Harrison 2009” [specimen condition: 
<taxonomicName id="2EF81BA51E4CFF906160FD91AFC4DDFD" box="[593,741,616,642]" class="Insecta" family="Dermestidae" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="family">Dermestidae</taxonomicName>
damage as follows: pinhole enlarged in right elytron; holes in abdominal sternites; left hand side tarsomere chewed] (13 
<collectionCode id="8FE9F8E31E4CFF9060A4FD77AED3DDD7" box="[917,1010,654,680]" pageId="14" pageNumber="15">ZMHB</collectionCode>
). 
<typeStatus id="3643DE841E4CFF906736FD77A99BDDD7" box="[1031,1210,654,680]" pageId="14" pageNumber="15" type="paralectotype">Paralectotypes</typeStatus>
(5). One 
<typeStatus id="3643DE841E4CFF906602FD77A8BCDDD7" box="[1331,1437,654,680]" pageId="14" pageNumber="15" type="paratype">paratype</typeStatus>
labelled “Entebbe, 
<collectingCountry id="91EF20B61E4CFF906247FD4CACF5DDB0" box="[374,468,693,719]" name="Uganda" pageId="14" pageNumber="15">Uganda</collectingCountry>
, Centr. Afr. [same data on reverse of label] / 
<taxonomicName id="2EF81BA51E4CFF9060D1FD4CA877DDB0" authority="Mos." authorityName="Mos." box="[992,1366,693,719]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="ugandensis">
<emphasis id="DB8CBC341E4CFF9060D1FD4CA830DDB1" box="[992,1297,693,718]" italics="true" pageId="14" pageNumber="15">Asthenopholis ugandensis</emphasis>
Mos.
</taxonomicName>
, 
<typeStatus id="3643DE841E4CFF906653FD4CA8BDDDB0" box="[1378,1436,693,719]" pageId="14" pageNumber="15">Type</typeStatus>
Ƥ [symbol, written on faded white card by Moser] / 
<typeStatus id="3643DE841E4CFF9061CCFD25AE16DD89" box="[765,823,732,758]" pageId="14" pageNumber="15">Type</typeStatus>
[printed on reddish brown card] / Zool. Mus. Berlin / 
<typeStatus id="3643DE841E4CFF906396FCFBAC6BDC63" box="[167,330,770,796]" pageId="14" pageNumber="15" type="paralectotype">Paralectotype</typeStatus>
Ƥ, 
<taxonomicName id="2EF81BA51E4CFF906248FCFBAE70DC63" authority="Moser, 1913" authorityName="Moser" authorityYear="1913" box="[377,849,770,796]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="ugandensis">
<emphasis id="DB8CBC341E4CFF906248FCFBAF91DC64" box="[377,688,770,795]" italics="true" pageId="14" pageNumber="15">Asthenopholis ugandensis</emphasis>
<bibRefCitation id="8D691DD71E4CFF906188FCFBAE70DC63" author="Moser" box="[697,849,770,796]" pageId="14" pageNumber="44" refString="Moser, J. (1913) Beitrag zur kenntnis der Melolonthiden II. Annales de la Societe Entomologique de Belgique, 57 (5 - 12), 331 - 345." type="journal article" year="1913">Moser, 1913</bibRefCitation>
</taxonomicName>
, des. J. du G. Harrison 2009” (1Ƥ 
<collectionCode id="8FE9F8E31E4CFF9067C5FCFBA86EDC63" box="[1268,1359,770,796]" pageId="14" pageNumber="15">ZMHB</collectionCode>
). Four 
<typeStatus id="3643DE841E4CFF9063A6FCD0AC28DC3C" box="[151,265,809,835]" pageId="14" pageNumber="15" type="paratype">paratypes</typeStatus>
labelled “Entebbe, 
<collectingCountry id="91EF20B61E4CFF9062C6FCD0AF76DC3C" box="[503,599,809,835]" name="Uganda" pageId="14" pageNumber="15">Uganda</collectingCountry>
, Centr. Afr.” [same data on reverse of label], [identical locality labels and pins as the 
<typeStatus id="3643DE841E4CFF906268FCA9ACEADC15" box="[345,459,848,874]" pageId="14" pageNumber="15" type="lectotype">lectotype</typeStatus>
and the other 
<typeStatus id="3643DE841E4CFF9061B1FCA9AE06DC15" box="[640,807,848,874]" pageId="14" pageNumber="15" type="paralectotype">paralectotype</typeStatus>
but without any written 
<typeStatus id="3643DE841E4CFF90676CFCA9A9B3DC15" box="[1117,1170,848,874]" pageId="14" pageNumber="15">type</typeStatus>
labels] (33 1Ƥ 
<collectionCode id="8FE9F8E31E4CFF90666FFCA9ADD0DCEF" pageId="14" pageNumber="15">SEM ZMHB</collectionCode>
).
</paragraph>
<paragraph id="E94760261E4CFF9063F4FC64A866DCA1" blockId="14.[151,1437,152,990]" pageId="14" pageNumber="15">
<emphasis id="DB8CBC341E4CFF9063F4FC64AF6EDCC8" bold="true" box="[197,591,925,951]" pageId="14" pageNumber="15">Additional material examined</emphasis>
(13). 
<emphasis id="DB8CBC341E4CFF906193FC64AE60DCC8" bold="true" box="[674,833,925,951]" pageId="14" pageNumber="15">
<collectingCountry id="91EF20B61E4CFF906193FC64AE60DCC8" box="[674,833,925,951]" name="Tanzania" pageId="14" pageNumber="15">TANZANIA</collectingCountry>
</emphasis>
. Manyara See [Lake], 12.[19]61, H. u. B. Frey / 
<taxonomicName id="2EF81BA51E4CFF9063A6FC3DAF3DDCA1" authority="Moser" authorityName="Moser" box="[151,540,964,990]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="ugandensis">
<emphasis id="DB8CBC341E4CFF9063A6FC3DACE9DCA2" box="[151,456,964,989]" italics="true" pageId="14" pageNumber="15">Asthenopholis ugandensis</emphasis>
Moser
</taxonomicName>
, female, J. Decelle det., 1969 / Comp. Typ. 
<taxonomicName id="2EF81BA51E4CFF90672FFC3DA983DCA2" box="[1054,1186,964,989]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="ugandensis">
<emphasis id="DB8CBC341E4CFF90672FFC3DA983DCA2" box="[1054,1186,964,989]" italics="true" pageId="14" pageNumber="15">ugandensis</emphasis>
</taxonomicName>
(13 
<collectionCode id="8FE9F8E31E4CFF9067EEFC3DA81ADCA1" box="[1247,1339,964,990]" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="14" pageNumber="15">MRAC</collectionCode>
).
</paragraph>
</subSubSection>
<paragraph id="E94760261E4CFF9063A6FBCEA860DBE4" pageId="14" pageNumber="15">
<table id="9BF892861E4C006163A6FBCEA860DBE4" box="[151,1345,1079,1179]" gridcols="6" gridrows="3" pageId="14" pageNumber="15">
<tr id="57C862641E4C006163A6FBCEA860DB2E" box="[151,1345,1079,1105]" gridrow="0" pageId="14" pageNumber="15" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="14190B181E4C006163A6FBCEAF80DB2E" box="[151,673,1079,1105]" gridcol="0" gridrow="0" pageId="14" pageNumber="15">
<taxonomicName id="2EF81BA51E4CFF9063A6FBCEAF6BDB2E" authority="Brenske, 1898" authorityName="Brenske" authorityYear="1898" box="[151,586,1079,1105]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="DB8CBC341E4CFF9063A6FBCEAF6BDB2E" bold="true" box="[151,586,1079,1105]" pageId="14" pageNumber="15">
<emphasis id="DB8CBC341E4CFF9063A6FBCEACB4DB2E" bold="true" box="[151,405,1079,1105]" italics="true" pageId="14" pageNumber="15">Asthenopholis minor</emphasis>
Brenske, 1898
</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="57C862641E4C006163A6FBA3A860DB0B" box="[151,1345,1114,1140]" gridrow="1" pageId="14" pageNumber="15" rowspan-5="1">
<th id="14190B181E4C006163A6FBA3AF80DB0B" box="[151,673,1114,1140]" gridcol="0" gridrow="1" pageId="14" pageNumber="15">(Figs. 5, 22, 32–33, 46–47, 74–78, 102–103,</th>
<td id="14190B181E4C00616199FBA3AE37DB0B" box="[680,790,1114,1140]" gridcol="1" gridrow="1" pageId="14" pageNumber="15">122–124,</td>
<td id="14190B181E4C0061602CFBA3AEAADB0B" box="[797,907,1114,1140]" gridcol="2" gridrow="1" pageId="14" pageNumber="15">142–143,</td>
<td id="14190B181E4C006160A3FBA3A9A5DB0B" box="[914,1156,1114,1140]" gridcol="3" gridrow="1" pageId="14" pageNumber="15">162–164, 178) black</td>
<td id="14190B181E4C006167BAFBA3A820DB0B" box="[1163,1281,1114,1140]" gridcol="4" gridrow="1" pageId="14" pageNumber="15">form</td>
</tr>
<tr id="57C862641E4C006163A6FB78A860DBE4" box="[151,1345,1153,1179]" gridrow="2" pageId="14" pageNumber="15">
<th id="14190B181E4C006163A6FB78AF80DBE4" box="[151,673,1153,1179]" gridcol="0" gridrow="2" pageId="14" pageNumber="15">(Figs. 7, 22, 36–37, 50–51, 84–88, 106–107,</th>
<td id="14190B181E4C00616199FB78AE37DBE4" box="[680,790,1153,1179]" gridcol="1" gridrow="2" pageId="14" pageNumber="15">128–130,</td>
<td id="14190B181E4C0061602CFB78AEAADBE4" box="[797,907,1153,1179]" gridcol="2" gridrow="2" pageId="14" pageNumber="15">146–147,</td>
<td id="14190B181E4C006160A3FB78A9A5DBE4" box="[914,1156,1153,1179]" gridcol="3" gridrow="2" pageId="14" pageNumber="15">168–170, 178) black</td>
<td id="14190B181E4C006167BAFB78A820DBE4" box="[1163,1281,1153,1179]" gridcol="4" gridrow="2" pageId="14" pageNumber="15">and russet</td>
<td id="14190B181E4C00616639FB78A860DBE4" box="[1288,1345,1153,1179]" gridcol="5" gridrow="2" pageId="14" pageNumber="15">form</td>
</tr>
</table>
</paragraph>
<subSubSection id="A1E233AD1E4CFF9063A6FB37AECADA3A" pageId="14" pageNumber="15" type="reference_group">
<paragraph id="E94760261E4CFF9063A6FB37AC27DA5A" blockId="14.[151,1437,1230,1349]" pageId="14" pageNumber="15">
<treatmentCitationGroup id="C9E847081E4CFF9063A6FB37AC27DA5A" pageId="14" pageNumber="15">
<taxonomicName id="2EF81BA51E4CFF9063A6FB37AF3DDB9A" authority="Brenske, 1898" authorityName="Brenske" authorityYear="1898" box="[151,540,1230,1253]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="DB8CBC341E4CFF9063A6FB37AC54DB9A" box="[151,373,1230,1253]" italics="true" pageId="14" pageNumber="15">Asthenopholis minor</emphasis>
<bibRefCitation id="8D691DD71E4CFF90624CFB37AF3DDB9A" author="Brenske" box="[381,540,1230,1253]" pageId="14" pageNumber="43" refString="Brenske, E. (1898) Melolonthiden aus Afrika. Stettiner Entomologische Zeitung, 59 (10 - 12) (iv), 333 - 394." type="journal article" year="1898">Brenske, 1898</bibRefCitation>
</taxonomicName>
: (Length 16, width 8 mill., Transvaal, male 
<typeStatus id="3643DE841E4CFF9060C6FB37A978DB9A" box="[1015,1113,1230,1253]" pageId="14" pageNumber="15" type="lectotype">lectotype</typeStatus>
in ZMHB) 390; 
<treatmentCitation id="685946371E4CFF906623FB37ADC4DA7A" author="Arrow" page="97" pageId="14" pageNumber="15" year="1902">
<bibRefCitation id="8D691DD71E4CFF906623FB37A8B9DB9A" author="Arrow" box="[1298,1432,1230,1253]" pageId="14" pageNumber="43" refString="Arrow, G. J. (1902) On rutelid and melolonthid beetles from Mashonaland and East Africa. The Annals and Magazine of Natural History, series 7, volume 9, 89 - 101." type="journal article" year="1902">Arrow 1902</bibRefCitation>
: 97
</treatmentCitation>
; 
<treatmentCitation id="685946371E4CFF9063C2FB17ACF9DA7A" author="Peringuey" box="[243,472,1262,1285]" page="287" pageId="14" pageNumber="15" year="1904">
<bibRefCitation id="8D691DD71E4CFF9063C2FB17AC83DA7A" author="Peringuey" box="[243,418,1262,1285]" pageId="14" pageNumber="44" refString="Peringuey, L. (1904) Descriptive catalogue of the Coleoptera of South Africa (Lucanidae and Scarabaeidae). Transactions of the South African Philosophical Society, 13, 1 - 293 + 4 plates." type="book chapter" year="1904">Péringuey 1904</bibRefCitation>
: 287
</treatmentCitation>
(as a synonym of 
<taxonomicName id="2EF81BA51E4CFF90619CFB17AE66DA7A" box="[685,839,1262,1285]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="subfasciata">
<emphasis id="DB8CBC341E4CFF90619CFB17AE66DA7A" box="[685,839,1262,1285]" italics="true" pageId="14" pageNumber="15">A. subfasciata</emphasis>
</taxonomicName>
); 
<treatmentCitation id="685946371E4CFF906050FB17A976DA7A" author="Dalla" box="[865,1111,1262,1285]" page="182" pageId="14" pageNumber="15" year="1912">
<bibRefCitation id="8D691DD71E4CFF906050FB17A900DA7A" author="Dalla" box="[865,1057,1262,1285]" pageId="14" pageNumber="43" refString="Dalla Torre, K. W. von (1912) Scarabaeidae: Melolonthinae III. Melolonthini, Pars 49, Vol. 20 (4), pp. 135 - 290. In: S. Schenkling (ed.), Coleopterorum Catalogus, W. Junk, Berlin, 450 pp." type="book chapter" year="1912">Dalla Torre 1912</bibRefCitation>
: 182
</treatmentCitation>
(catalogue); 
<treatmentCitation id="685946371E4CFF9067DAFB17A8BDDA7A" author="Arrow" box="[1259,1436,1262,1285]" page="60" pageId="14" pageNumber="15" year="1917">
<bibRefCitation id="8D691DD71E4CFF9067DAFB17A852DA7A" author="Arrow" box="[1259,1395,1262,1285]" pageId="14" pageNumber="43" refString="Arrow, G. J. (1917) Some systematic notes on melolonthine Coleoptera. The Annals and Magazine of Natural History, series 8, volume 19, 59 - 65." type="journal article" year="1917">Arrow 1917</bibRefCitation>
: 60
</treatmentCitation>
(key).
</treatmentCitationGroup>
</paragraph>
<paragraph id="E94760261E4CFF9063A6FAD7AECADA3A" blockId="14.[151,1437,1230,1349]" box="[151,1003,1326,1349]" pageId="14" pageNumber="15">
<treatmentCitationGroup id="C9E847081E4CFF9063A6FAD7AECADA3A" box="[151,1003,1326,1349]" pageId="14" pageNumber="15">
<taxonomicName id="2EF81BA51E4CFF9063A6FAD7ACDCDA3A" authority="Brenske" authorityName="Brenske" box="[151,509,1326,1349]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="bechuana">
<emphasis id="DB8CBC341E4CFF9063A6FAD7ACBDDA3A" box="[151,412,1326,1349]" italics="true" pageId="14" pageNumber="15">Asthenopholis bechuana</emphasis>
Brenske
</taxonomicName>
, in litt.: 
<treatmentCitation id="685946371E4CFF906164FAD7AE12DA3A" author="Peringuey" box="[597,819,1326,1349]" page="287" pageId="14" pageNumber="15" year="1904">
<bibRefCitation id="8D691DD71E4CFF906164FAD7AE20DA3A" author="Peringuey" box="[597,769,1326,1349]" pageId="14" pageNumber="44" refString="Peringuey, L. (1904) Descriptive catalogue of the Coleoptera of South Africa (Lucanidae and Scarabaeidae). Transactions of the South African Philosophical Society, 13, 1 - 293 + 4 plates." type="book chapter" year="1904">Péringuey 1904</bibRefCitation>
: 287
</treatmentCitation>
(no description).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="A1E233AD1E4CFF9063A6FA8CA96BD97C" pageId="14" pageNumber="15" type="diagnosis">
<paragraph id="E94760261E4CFF9063A6FA8CA96BD97C" blockId="14.[151,1437,1397,2003]" pageId="14" pageNumber="15">
<emphasis id="DB8CBC341E4CFF9063A6FA8CAC39DAF0" bold="true" box="[151,280,1397,1423]" pageId="14" pageNumber="15">Diagnosis.</emphasis>
The external morphology of this species is the most variable within the genus. The head, thorax, and scutellum are black; or with elytra reddish brown; medium sized species (14.0–19.0 mm). Teneral adults appear more brown than black, probably due to incomplete melanisation of the head, thorax, and scutellum. Scales on the thorax, scutellum, and elytra are fine, white, and of equal diameter.
</paragraph>
</subSubSection>
<subSubSection id="A1E233AD1E4CFF9163F7F9F6A9F0DB2D" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="E94760261E4CFF9163F7F9F6A9C7DE59" blockId="14.[151,1437,1397,2003]" lastBlockId="15.[151,1438,152,2034]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<emphasis id="DB8CBC341E4CFF9063F7F9F6AC5DD956" bold="true" box="[198,380,1551,1577]" pageId="14" pageNumber="15">Redescription.</emphasis>
<emphasis id="DB8CBC341E4CFF9062B7F9E9AC94D956" box="[390,437,1552,1577]" italics="true" pageId="14" pageNumber="15">Size</emphasis>
(n = 
<specimenCount id="FFFEABAF1E4CFF9062CCF9E9AF4CD955" box="[509,621,1552,1578]" pageId="14" pageNumber="15" type="male">15 males</specimenCount>
; n = 
<specimenCount id="FFFEABAF1E4CFF906181F9E9AE18D955" box="[688,825,1552,1578]" pageId="14" pageNumber="15" type="female">10 females</specimenCount>
). Length: males, mean 
<quantity id="2E00CDC31E4CFF906765F9E9A9E2D955" box="[1108,1219,1552,1578]" metricMagnitude="-2" metricUnit="m" metricValue="1.66" pageId="14" pageNumber="15" unit="mm" value="16.6">16.6 mm</quantity>
, (range 14.5–18.0 mm), females, mean 
<quantity id="2E00CDC31E4CFF9062A2F9CFAF21D92F" box="[403,512,1590,1616]" metricMagnitude="-2" metricUnit="m" metricValue="1.6199999999999999" pageId="14" pageNumber="15" unit="mm" value="16.2">16.2 mm</quantity>
, (range 14.0–19.00 mm). Width: males, mean 
<quantity id="2E00CDC31E4CFF90671CF9CFA9ADD92F" box="[1069,1164,1590,1616]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" pageId="14" pageNumber="15" unit="mm" value="8.8">8.8 mm</quantity>
, (range 8.0–10.0 mm), females, mean 
<quantity id="2E00CDC31E4CFF906266F9A4AC9BD908" box="[343,442,1629,1655]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="14" pageNumber="15" unit="mm" value="8.4">8.4 mm</quantity>
, (range 7.0–10.00 mm). 
<emphasis id="DB8CBC341E4CFF9061C4F9A4AE6FD909" box="[757,846,1629,1654]" italics="true" pageId="14" pageNumber="15">Colour</emphasis>
(Figs. 5, 7): Cuticle of completely sclerotised specimens is black and covered by small, blunt ended white scales. However, populations with black head, thorax, and scutellum and reddish brown elytra exist, and there are other populations with glossy brown individuals with tan coloured setae. The various colour forms are listed separately in the material examined section. 
<emphasis id="DB8CBC341E4CFF9063CDF901AC1CD86E" box="[252,317,1784,1809]" italics="true" pageId="14" pageNumber="15">Head</emphasis>
(Figs. 32–33, 36–37, 46–47, 50–51): Head uniformly covered with fine, white setae; longer setae on eye protuberance. Clypeus slightly reflexed in the middle. 
<emphasis id="DB8CBC341E4CFF906098F8E6A93CD847" box="[937,1053,1823,1848]" italics="true" pageId="14" pageNumber="15">Pronotum</emphasis>
(Figs. 74–78, 84–88): Pronotum ratio 1.55:1 (n = 153); 1.56:1 (n = 10Ƥ). Pronotum covered with small lanceolate scales; lateral margin finely serrated almost straight posteriad, with basolateral angles not protruding; longer setae inserted within serrations. Row of longer and finer setae along the anterior edge. 
<emphasis id="DB8CBC341E4CFF9060FFF86AA969D8D3" box="[974,1096,1939,1964]" italics="true" pageId="14" pageNumber="15">Scutellum</emphasis>
(Figs. 102–103, 106–107): Scutellum triangular, apex blunt and covered with two 
<typeStatus id="3643DE841E4CFF906069F840AEBAD8AC" box="[856,923,1977,2003]" pageId="14" pageNumber="15">types</typeStatus>
of setae; i.e., long and fine, shorter and lanceolate; unsetose area anteriomedially and along lateral edges. 
<emphasis id="DB8CBC341E4DFF9160ADFF61AEC4DFCE" box="[924,997,152,177]" italics="true" pageId="15" pageNumber="16">Elytra</emphasis>
(Figs. 122–124, 128–130): Scales on the elytra are fine, white, and spatulate; suture and costae only indistinct; with or without denudate maculae on elytra. 
<taxonomicName id="2EF81BA51E4DFF916222FF1CACA6DF81" box="[275,391,229,254]" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageId="15" pageNumber="16" phylum="Chordata" rank="genus">
<emphasis id="DB8CBC341E4DFF916222FF1CACA6DF81" box="[275,391,229,254]" italics="true" pageId="15" pageNumber="16">Pygidium</emphasis>
</taxonomicName>
(Figs. 142–143, 146–147): 
<taxonomicName id="2EF81BA51E4DFF9161EAFF1CAE71DF80" box="[731,848,229,255]" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageId="15" pageNumber="39" phylum="Chordata" rank="genus">Pygidium</taxonomicName>
flat and uniformly covered by fine, white setae. 
<emphasis id="DB8CBC341E4DFF9163A6FEF5AC63DE5A" box="[151,322,268,293]" italics="true" pageId="15" pageNumber="16">Male genitalia</emphasis>
illustrated in Figs. 162–164 (black form) and 168–170 (black and russet form).
</paragraph>
<paragraph id="E94760261E4DFF9163F4FECBAC44DDFD" blockId="15.[151,1438,152,2034]" pageId="15" pageNumber="16">
<emphasis id="DB8CBC341E4DFF9163F4FECBAF2DDE33" bold="true" box="[197,524,306,332]" pageId="15" pageNumber="16">Morphological variability.</emphasis>
Resolution of specimens to be included and excluded from 
<taxonomicName id="2EF81BA51E4DFF9167F8FECBA80EDE34" box="[1225,1327,306,331]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="DB8CBC341E4DFF9167F8FECBA80EDE34" box="[1225,1327,306,331]" italics="true" pageId="15" pageNumber="16">A. minor</emphasis>
</taxonomicName>
has been complicated. The external morphology allows one to readily sort the specimens into four main morphospecies, but when one looks for consistent morphological characters to differentiate these into species the boundaries between all specimens included here become obscure. Consequently, all these specimens have been identified as 
<taxonomicName id="2EF81BA51E4DFF916243FE34ACFBDE99" box="[370,474,461,486]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="DB8CBC341E4DFF916243FE34ACFBDE99" box="[370,474,461,486]" italics="true" pageId="15" pageNumber="16">A. minor</emphasis>
</taxonomicName>
, but for practical purposes the phenotypes mentioned are listed separately in the material examined to facilitate their study at the molecular level by others, which may provide the evidence to recognize further species from 
<emphasis id="DB8CBC341E4DFF916129FDE3AFA8DD4C" box="[536,649,538,563]" italics="true" pageId="15" pageNumber="16">
<taxonomicName id="2EF81BA51E4DFF916129FDE3AFA4DD4C" box="[536,645,538,563]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="minor">A. minor</taxonomicName>
.
</emphasis>
The four main morphospecies are labelled as follows: 
<taxonomicName id="2EF81BA51E4DFF916600FDE3A8BCDD4C" box="[1329,1437,538,563]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="DB8CBC341E4DFF916600FDE3A8BCDD4C" box="[1329,1437,538,563]" italics="true" pageId="15" pageNumber="16">A. minor</emphasis>
</taxonomicName>
(narrow black and red form); 
<taxonomicName id="2EF81BA51E4DFF9162CAFDB8AF42DD25" box="[507,611,577,602]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="DB8CBC341E4DFF9162CAFDB8AF42DD25" box="[507,611,577,602]" italics="true" pageId="15" pageNumber="16">A. minor</emphasis>
</taxonomicName>
(broad black and red form); 
<taxonomicName id="2EF81BA51E4DFF91608FFDB8A906DD25" box="[958,1063,577,602]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="DB8CBC341E4DFF91608FFDB8A906DD25" box="[958,1063,577,602]" italics="true" pageId="15" pageNumber="16">A. minor</emphasis>
</taxonomicName>
(black form); 
<taxonomicName id="2EF81BA51E4DFF9167E7FDB8A81FDD25" box="[1238,1342,577,602]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="DB8CBC341E4DFF9167E7FDB8A81FDD25" box="[1238,1342,577,602]" italics="true" pageId="15" pageNumber="16">A. minor</emphasis>
</taxonomicName>
(brown and glossy form).
</paragraph>
<paragraph id="E94760261E4DFF9163F4FD77AC95DC15" blockId="15.[151,1438,152,2034]" pageId="15" pageNumber="16">
An explanation for the cuticular colour range observed within this species is pigment polyphenism, where the concentration of developmental hormones varies depending on environmental and physiological conditions during the beetles’ ecdysis. This in turn has an effect upon the degree of melanisation of the adult beetle, allowing for both melanic and non-melanic individuals within the same population, or different populations, of the same species (see reviews by 
<bibRefCitation id="8D691DD71E4DFF9161C9FCD0A936DC3C" author="Emlen" box="[760,1047,809,835]" pageId="15" pageNumber="43" refString="Emlen, D. J. &amp; Nijhout, H. F. (2000) The development and evolution of exaggerated morphologies in insects. Annual Review of Entomology, 45, 661 - 708." type="journal article" year="2000">Emlen &amp; Nijhout 2000</bibRefCitation>
, 
<bibRefCitation id="8D691DD71E4DFF916715FCD0A842DC3C" author="Hartfelder" box="[1060,1379,809,835]" pageId="15" pageNumber="43" refString="Hartfelder, K. &amp; Emlen, D. J. (2005) Endocrine control of insect polyphenism. Comprehensive Molecular Insect Science, 3, 651 - 703." type="journal article" year="2005">Hartfelder &amp; Emlen 2005</bibRefCitation>
, and 
<bibRefCitation id="8D691DD71E4DFF9163A6FCA9AC07DC15" author="Verma" box="[151,294,848,874]" pageId="15" pageNumber="44" refString="Verma, K. K. (2007) Polyphenism in insects and the juvenile hormone. Journal of Bioscience, 32 (2), 415 - 420." type="journal article" year="2007">Verma 2007</bibRefCitation>
for details).
</paragraph>
<paragraph id="E94760261E4DFF9163F4FC8FA9F0DB2D" blockId="15.[151,1438,152,2034]" pageId="15" pageNumber="16">
A detailed examination of the external morphology, male genitalia, and their inclusion within the 
<collectionCode id="8FE9F8E31E4DFF91666EFC8FA8BDDCEF" box="[1375,1436,886,912]" pageId="15" pageNumber="16">SEM</collectionCode>
study phase of the revision, did however reveal the existence of two species within 
<taxonomicName id="2EF81BA51E4DFF916741FC64A9F7DCC9" box="[1136,1238,925,950]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="DB8CBC341E4DFF916741FC64A9F7DCC9" box="[1136,1238,925,950]" italics="true" pageId="15" pageNumber="16">A. minor</emphasis>
</taxonomicName>
, i.e. (i) 
<taxonomicName id="2EF81BA51E4DFF916604FC64A8BDDCC9" box="[1333,1436,925,950]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="DB8CBC341E4DFF916604FC64A8BDDCC9" box="[1333,1436,925,950]" italics="true" pageId="15" pageNumber="16">A. minor</emphasis>
</taxonomicName>
and (ii) 
<taxonomicName id="2EF81BA51E4DFF9163C5FC3DAC46DCA2" box="[244,359,964,989]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="crypsis">
<emphasis id="DB8CBC341E4DFF9163C5FC3DAC46DCA2" box="[244,359,964,989]" italics="true" pageId="15" pageNumber="16">A. crypsis</emphasis>
</taxonomicName>
. Thus, this is a situation where specimens that appear to be different based on their external morphology represent a morphologically variable species, and another that appears to be the same species includes a cryptic species. Additional sampling of 
<taxonomicName id="2EF81BA51E4DFF916020FBE8AE5CDB55" box="[785,893,1041,1066]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="DB8CBC341E4DFF916020FBE8AE5CDB55" box="[785,893,1041,1066]" italics="true" pageId="15" pageNumber="16">A. minor</emphasis>
</taxonomicName>
across the geographic range and molecular analyses are recommended to further resolve the species limits and within-species-variation.
</paragraph>
</subSubSection>
<subSubSection id="A1E233AD1E4DFF8E63F4FBA7AC26D88D" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="distribution">
<paragraph id="E94760261E4DFF9163F4FBA7A943DA1F" blockId="15.[151,1438,152,2034]" pageId="15" pageNumber="16">
<emphasis id="DB8CBC341E4DFF9163F4FBA7AC44DB07" bold="true" box="[197,357,1118,1144]" pageId="15" pageNumber="16">Distribution.</emphasis>
The known distribution of 
<taxonomicName id="2EF81BA51E4DFF916196FBA6AE2CDB07" box="[679,781,1119,1144]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="DB8CBC341E4DFF916196FBA6AE2CDB07" box="[679,781,1119,1144]" italics="true" pageId="15" pageNumber="16">A. minor</emphasis>
</taxonomicName>
is centred within and around KwaZulu– 
<collectingCountry id="91EF20B61E4DFF9167D6FBA7A806DB07" box="[1255,1319,1118,1144]" name="South Africa" pageId="15" pageNumber="16">Natal</collectingCountry>
(Fig. 22). Locality data indicate that this species has the widest distribution of any South African 
<taxonomicName id="2EF81BA51E4DFF9167ABFB7CA861DBE1" box="[1178,1344,1157,1182]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="DB8CBC341E4DFF9167ABFB7CA861DBE1" box="[1178,1344,1157,1182]" italics="true" pageId="15" pageNumber="16">Asthenopholis</emphasis>
</taxonomicName>
species with records from the following provinces and countries: 
<collectingCountry id="91EF20B61E4DFF916043FB55A938DBB9" box="[882,1049,1196,1222]" name="South Africa" pageId="15" pageNumber="16">South Africa</collectingCountry>
: Western Cape (1 record) [viz, Vlottenburg (J. A. Nagle)]; Eastern Cape (1) [Stutterheim (J. A. Nagle)]; Free State (2); Limpopo (2) [Nylstroom (ex student collection)]; Mpumalanga (7); KwaZulu– 
<collectingCountry id="91EF20B61E4DFF916091FB00AEC1DA6C" box="[928,992,1273,1299]" name="South Africa" pageId="15" pageNumber="16">Natal</collectingCountry>
(25); 
<collectingCountry id="91EF20B61E4DFF91671BFB00A9A8DA6C" box="[1066,1161,1273,1299]" name="Lesotho" pageId="15" pageNumber="16">Lesotho</collectingCountry>
(1) and 
<collectingCountry id="91EF20B61E4DFF9167DEFB00A84BDA6C" box="[1263,1386,1273,1299]" name="Swaziland" pageId="15" pageNumber="16">Swaziland</collectingCountry>
(5). However, the validity of the localities included within square brackets above require confirmation, as they appear to be outliers, but without further collecting this cannot be determined here.
</paragraph>
<paragraph id="E94760261E4DFF9163F4FA94A8B7D99C" blockId="15.[151,1438,152,2034]" pageId="15" pageNumber="16">
The specimen labelled “Greyton” is mapped to “Greytown” in KwaZulu– 
<collectingCountry id="91EF20B61E4DFF916770FA94A9A3DAF8" box="[1089,1154,1389,1415]" name="South Africa" pageId="15" pageNumber="16">Natal</collectingCountry>
and not Greyton in the Western Cape, as it originates from the Pietermaritzburg collection and this locality is more likely than the Western Cape one. However, with 
<taxonomicName id="2EF81BA51E4DFF91617FFA42AF9CDAAB" box="[590,701,1467,1492]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="DB8CBC341E4DFF91617FFA42AF9CDAAB" box="[590,701,1467,1492]" italics="true" pageId="15" pageNumber="16">A. minor</emphasis>
</taxonomicName>
(broad black and red form) records from Vlottenburg and Stutterheim future collecting may prove the Greyton (Western Cape) locality correct. The Vlottenburg and Stutterheim localities originate from three specimens collected by J. A. Nagle, an amateur entomologist who evidently worked alone, and whose collection of insects was donated to the Transvaal Museum (B. Dombrowsky, TMSA; personal communication 2007). Nagle’s collection is characterised by series of all insect orders, with varying dates but from the same locality, thus the Nagle Western and East Cape 
<taxonomicName id="2EF81BA51E4DFF9163A6F95AAC70D9C3" box="[151,337,1699,1724]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="DB8CBC341E4DFF9163A6F95AAC70D9C3" box="[151,337,1699,1724]" italics="true" pageId="15" pageNumber="16">Asthenopholis</emphasis>
</taxonomicName>
specimens may actually have been collected at Winkelspruit (
<geoCoordinate id="8CCC06E11E4DFF9167BAF95AA9D6D9C2" box="[1163,1271,1699,1725]" direction="south" orientation="latitude" pageId="15" pageNumber="16" precision="925" value="-30.1">30°06’S</geoCoordinate>
<geoCoordinate id="8CCC06E11E4DFF916630F95AA84FD9C2" box="[1281,1390,1699,1725]" direction="east" orientation="longitude" pageId="15" pageNumber="16" precision="925" value="30.85">30°51’E</geoCoordinate>
) or Pietermaritzburg (
<geoCoordinate id="8CCC06E11E4DFF91625DF930ACECD99C" box="[364,461,1737,1763]" direction="south" orientation="latitude" pageId="15" pageNumber="16" precision="925" value="-29.566668">29°34’S</geoCoordinate>
<geoCoordinate id="8CCC06E11E4DFF9162E5F930AF16D99C" box="[468,567,1737,1763]" direction="east" orientation="longitude" pageId="15" pageNumber="16" precision="925" value="30.35">30°21’E</geoCoordinate>
) both in KwaZulu– 
<collectingCountry id="91EF20B61E4DFF916011F930AE40D99C" box="[800,865,1737,1763]" name="South Africa" pageId="15" pageNumber="16">Natal</collectingCountry>
, where J. A. Nagle often collected other insects.
</paragraph>
<paragraph id="E94760261E4DFF9163F4F909ADCDD88D" blockId="15.[151,1438,152,2034]" pageId="15" pageNumber="16">
Three specimens of this species were found in the Berlin collection, i.e. the 
<typeStatus id="3643DE841E4DFF916778F909A999D875" box="[1097,1208,1776,1802]" pageId="15" pageNumber="16" type="lectotype">lectotype</typeStatus>
, 
<typeStatus id="3643DE841E4DFF9167F5F909A844D875" box="[1220,1381,1776,1802]" pageId="15" pageNumber="16" type="paralectotype">paralectotype</typeStatus>
, and one non-type. Both 
<typeStatus id="3643DE841E4DFF9162B1F8EFAC9FD84F" box="[384,446,1814,1840]" pageId="15" pageNumber="16">types</typeStatus>
are labelled as coming from the “Transvaal”, while the third specimen is labelled as coming from Port 
<collectingCountry id="91EF20B61E4DFF916244F8C4AC97D828" box="[373,438,1853,1879]" name="South Africa" pageId="15" pageNumber="16">Natal</collectingCountry>
(= Durban). Based on the species known distribution, the Transvaal specimens are regarded here as incorrectly labelled. Based on the known distribution of 
<taxonomicName id="2EF81BA51E4DFF9160C0F89DA976D802" box="[1009,1111,1892,1917]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="DB8CBC341E4DFF9160C0F89DA976D802" box="[1009,1111,1892,1917]" italics="true" pageId="15" pageNumber="16">A. minor</emphasis>
</taxonomicName>
, the specimen records from “Betchuanen Land” would be incorrectly labelled specimens, because to date no 
<taxonomicName id="2EF81BA51E4DFF916769F872A9DED8DB" box="[1112,1279,1931,1956]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="DB8CBC341E4DFF916769F872A9DED8DB" box="[1112,1279,1931,1956]" italics="true" pageId="15" pageNumber="16">Asthenopholis</emphasis>
</taxonomicName>
species have been positively recorded from 
<collectingCountry id="91EF20B61E4DFF916129F848AFB3D8B4" box="[536,658,1969,1995]" name="Botswana" pageId="15" pageNumber="16">Botswana</collectingCountry>
(previously 
<collectingCountry id="91EF20B61E4DFF916000F848AEC1D8B4" box="[817,992,1969,1995]" name="Botswana" pageId="15" pageNumber="16">Bechuanaland</collectingCountry>
Protectorate [
<bibRefCitation id="8D691DD71E4DFF9167BEF848ADF9D88D" author="Broadley" pageId="15" pageNumber="43" refString="Broadley, D. G. &amp; Minshull, J. I. (1986) A gazetteer of African countries, their constituent parts, and their synonyms. Arnoldia Zimbabwe, 9 (26), 333 - 342." type="journal article" year="1986">Broadley &amp; Minshull 1986</bibRefCitation>
]).
</paragraph>
<paragraph id="E94760261E52FF8E63A6F88BAC26D88D" blockId="16.[151,1436,1906,2034]" pageId="16" pageNumber="17">
<emphasis id="DB8CBC341E52FF8E63A6F88BAC2DD8F5" bold="true" box="[151,268,1906,1930]" pageId="16" pageNumber="17">PLATE 4.</emphasis>
Figs. 13–20: 
<taxonomicName id="2EF81BA51E52FF8E6295F88AAF88D8F5" box="[420,681,1907,1930]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="adspersa">
<emphasis id="DB8CBC341E52FF8E6295F88AAF88D8F5" box="[420,681,1907,1930]" italics="true" pageId="16" pageNumber="17">Asthenopholis adspersa</emphasis>
</taxonomicName>
(adult male). 13. labium in ventral view; 14. labium in ventral view showing sutural line; 15. lateral view of left mandible; 16. lateral view of right mandible; 17. epipharynx in dorsal view; 18. surface of right mandible’s molar lobe; 19. ventral view of left maxillae; 20. dorsal view of right maxillae. Scale bars = 100 µm.
</paragraph>
</subSubSection>
<subSubSection id="A1E233AD1E53FF8B63A6F88FA84AD9E6" lastPageId="21" lastPageNumber="22" pageId="17" pageNumber="18" type="description">
<paragraph id="E94760261E53FF8F63A6F88FAFCFD8AC" blockId="17.[151,1436,1910,2003]" pageId="17" pageNumber="18">
<emphasis id="DB8CBC341E53FF8F63A6F88FAC28D8F1" bold="true" box="[151,265,1910,1934]" pageId="17" pageNumber="18">PLATE 5.</emphasis>
Figs. 21–22: Known distribution of South African 
<taxonomicName id="2EF81BA51E53FF8F6016F88EAE9FD8F1" box="[807,958,1911,1934]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="DB8CBC341E53FF8F6016F88EAE9FD8F1" box="[807,958,1911,1934]" italics="true" pageId="17" pageNumber="18">Asthenopholis</emphasis>
</taxonomicName>
species. 21. 
<taxonomicName id="2EF81BA51E53FF8F6774F88EA9D7D8F1" box="[1093,1270,1911,1934]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="adspersa">
<emphasis id="DB8CBC341E53FF8F6774F88EA9E2D8F1" box="[1093,1219,1911,1934]" italics="true" pageId="17" pageNumber="18">A. adspersa</emphasis>
()
</taxonomicName>
; 
<taxonomicName id="2EF81BA51E53FF8F6632F88EA860D8F1" box="[1283,1345,1911,1934]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="rex" status="sp. nov.">
<emphasis id="DB8CBC341E53FF8F6632F88EA860D8F1" box="[1283,1345,1911,1934]" italics="true" pageId="17" pageNumber="18">A. rex</emphasis>
</taxonomicName>
<emphasis id="DB8CBC341E53FF8F6676F88FA8BDD8F1" bold="true" box="[1351,1436,1910,1934]" pageId="17" pageNumber="18">
<taxonomicNameLabel id="C0BF014F1E53FF8F6676F88FA8BDD8F1" box="[1351,1436,1910,1934]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(); and 
<taxonomicName id="2EF81BA51E53FF8F63CFF863ACB9D8CE" box="[254,408,1946,1969]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="subfasciata">
<emphasis id="DB8CBC341E53FF8F63CFF863ACB9D8CE" box="[254,408,1946,1969]" italics="true" pageId="17" pageNumber="18">A. subfasciata</emphasis>
</taxonomicName>
(ˑ). 22. 
<taxonomicName id="2EF81BA51E53FF8F6132F863AF4FD8CE" box="[515,622,1946,1969]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="crypsis" status="sp. nov.">
<emphasis id="DB8CBC341E53FF8F6132F863AF4FD8CE" box="[515,622,1946,1969]" italics="true" pageId="17" pageNumber="18">A. crypsis</emphasis>
</taxonomicName>
<emphasis id="DB8CBC341E53FF8F6147F860AFECD8CE" bold="true" box="[630,717,1945,1969]" pageId="17" pageNumber="18">
<taxonomicNameLabel id="C0BF014F1E53FF8F6147F860AFECD8CE" box="[630,717,1945,1969]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(); 
<taxonomicName id="2EF81BA51E53FF8F6038F863AEBDD8CF" box="[777,924,1945,1969]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="DB8CBC341E53FF8F6038F863AE49D8CE" box="[777,872,1946,1969]" italics="true" pageId="17" pageNumber="18">A. minor</emphasis>
()
</taxonomicName>
; and 
<taxonomicName id="2EF81BA51E53FF8F60E6F863A951D8CE" box="[983,1136,1946,1969]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="DB8CBC341E53FF8F60E6F863A951D8CE" box="[983,1136,1946,1969]" italics="true" pageId="17" pageNumber="18">Asthenopholis</emphasis>
</taxonomicName>
species near to 
<taxonomicName id="2EF81BA51E53FF8F662EF863A84AD8CE" box="[1311,1387,1946,1969]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="crypsis">
<emphasis id="DB8CBC341E53FF8F662EF863A84AD8CE" box="[1311,1387,1946,1969]" italics="true" pageId="17" pageNumber="18">crypsis</emphasis>
</taxonomicName>
(Ο). Questioned localities or determinations indicated by (?).
</paragraph>
<paragraph id="E94760261E50FF8C63A6FA28AE92D974" blockId="18.[151,1435,1489,1547]" pageId="18" pageNumber="19">
<emphasis id="DB8CBC341E50FF8C63A6FA28AC28DA96" bold="true" box="[151,265,1489,1513]" pageId="18" pageNumber="19">PLATE 6.</emphasis>
Fig. 23: All locality records of 
<taxonomicName id="2EF81BA51E50FF8C616AFA28AFD3DA97" box="[603,754,1489,1512]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="DB8CBC341E50FF8C616AFA28AFD3DA97" box="[603,754,1489,1512]" italics="true" pageId="18" pageNumber="19">Asthenopholis</emphasis>
</taxonomicName>
species plotted for 
<collectingCountry id="91EF20B61E50FF8C60F5FA28A96CDA97" box="[964,1101,1489,1512]" name="South Africa" pageId="18" pageNumber="19">South Africa</collectingCountry>
(); and known distribution of East African 
<taxonomicName id="2EF81BA51E50FF8C6212FA0DAC9BD974" box="[291,442,1524,1547]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="DB8CBC341E50FF8C6212FA0DAC9BD974" box="[291,442,1524,1547]" italics="true" pageId="18" pageNumber="19">Asthenopholis</emphasis>
</taxonomicName>
species; 
<taxonomicName id="2EF81BA51E50FF8C612DFA0DAF93D974" box="[540,690,1524,1547]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="crassa">
<emphasis id="DB8CBC341E50FF8C612DFA0DAF5ED974" box="[540,639,1524,1547]" italics="true" pageId="18" pageNumber="19">A. crassa</emphasis>
()
</taxonomicName>
; and 
<taxonomicName id="2EF81BA51E50FF8C61DAFA0DAE8ED974" box="[747,943,1524,1547]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="ugandensis">
<emphasis id="DB8CBC341E50FF8C61DAFA0DAEA1D974" box="[747,896,1524,1547]" italics="true" pageId="18" pageNumber="19">A. ugandensis</emphasis>
()
</taxonomicName>
.
</paragraph>
<paragraph id="E94760261E50FF8C63F4F9C4AE84D901" blockId="18.[151,1437,1597,2010]" pageId="18" pageNumber="19">
<emphasis id="DB8CBC341E50FF8C63F4F9C4ACF0D928" bold="true" box="[197,465,1597,1623]" pageId="18" pageNumber="19">Conservation status.</emphasis>
The species appears to be endemic to grassland in 
<collectingCountry id="91EF20B61E50FF8C677FF9C4A9D1D928" box="[1102,1264,1597,1623]" name="South Africa" pageId="18" pageNumber="19">South Africa</collectingCountry>
, 
<collectingCountry id="91EF20B61E50FF8C67CCF9C4A842D928" box="[1277,1379,1597,1623]" name="Lesotho" pageId="18" pageNumber="19">Lesotho</collectingCountry>
, and 
<collectingCountry id="91EF20B61E50FF8C63A6F99DAC35D901" box="[151,276,1636,1662]" name="Swaziland" pageId="18" pageNumber="19">Swaziland</collectingCountry>
, and occurs in some of the Drakensberg National Parks.
</paragraph>
<paragraph id="E94760261E50FF8C63F4F973A99FD9B4" blockId="18.[151,1437,1597,2010]" pageId="18" pageNumber="19">
Populations occurring within areas of intensive agriculture, e.g. sugarcane in KwaZulu– 
<collectingCountry id="91EF20B61E50FF8C6612F973A846D9DB" box="[1315,1383,1674,1700]" name="South Africa" pageId="18" pageNumber="19">Natal</collectingCountry>
and 
<collectingCountry id="91EF20B61E50FF8C63A6F948AC35D9B4" box="[151,276,1713,1739]" name="Swaziland" pageId="18" pageNumber="19">Swaziland</collectingCountry>
, could be threatened by the repeated use of pre-plant anti-white grub chemicals.
</paragraph>
<paragraph id="E94760261E50FF8C63F4F921ACB0D8A5" blockId="18.[151,1437,1597,2010]" pageId="18" pageNumber="19">
<emphasis id="DB8CBC341E50FF8C63F4F921ACADD98D" bold="true" box="[197,396,1752,1778]" pageId="18" pageNumber="19">Natural history.</emphasis>
Martin Krüger collected eight males and a single female of 
<taxonomicName id="2EF81BA51E50FF8C6762F921A99BD98E" box="[1107,1210,1752,1777]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="DB8CBC341E50FF8C6762F921A99BD98E" box="[1107,1210,1752,1777]" italics="true" pageId="18" pageNumber="19">A. minor</emphasis>
</taxonomicName>
(brown and glossy form) swarming over a lawn at 12: 
<specimenCount id="FFFEABAF1E50FF8C6104F906AF55D866" box="[565,628,1791,1817]" pageId="18" pageNumber="19" type="generic">30 p.</specimenCount>
m., indicating that this species is active during the day, but perhaps only while mating. Twice, Mike Way recorded swarming during the day in two forms of 
<taxonomicName id="2EF81BA51E50FF8C6794F8DCA82AD841" box="[1189,1291,1829,1854]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="DB8CBC341E50FF8C6794F8DCA82AD841" box="[1189,1291,1829,1854]" italics="true" pageId="18" pageNumber="19">A. minor</emphasis>
</taxonomicName>
(black form and the broad black and red form). While a series of 
<taxonomicName id="2EF81BA51E50FF8C6035F8B5AE4AD81A" box="[772,875,1868,1893]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="DB8CBC341E50FF8C6035F8B5AE4AD81A" box="[772,875,1868,1893]" italics="true" pageId="18" pageNumber="19">A. minor</emphasis>
</taxonomicName>
(narrow black and red form) were taken during the Maluti Drakensberg Transfronteir Survey where sampling was undertaken during the day. An 
<taxonomicName id="2EF81BA51E50FF8C6602F88AA8BDD8F3" box="[1331,1436,1907,1932]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="DB8CBC341E50FF8C6602F88AA8BDD8F3" box="[1331,1436,1907,1932]" italics="true" pageId="18" pageNumber="19">A. minor</emphasis>
</taxonomicName>
(broad black and red form) (E-Y: 765) was collected from a species of 
<taxonomicName id="2EF81BA51E50FF8C60D4F860A914D8CD" box="[997,1077,1945,1970]" class="Magnoliopsida" family="Proteaceae" genus="Protea" kingdom="Plantae" order="Proteales" pageId="18" pageNumber="19" phylum="Tracheophyta" rank="genus">
<emphasis id="DB8CBC341E50FF8C60D4F860A914D8CD" box="[997,1077,1945,1970]" italics="true" pageId="18" pageNumber="19">Protea</emphasis>
</taxonomicName>
at Cathedral Peak. Phenology indicated in Fig. 178.
</paragraph>
<paragraph id="E94760261E51FF8D63A6F853AFC4D89B" blockId="19.[151,1437,1962,2020]" pageId="19" pageNumber="20">
<emphasis id="DB8CBC341E51FF8D63A6F853AC2FD8BD" bold="true" box="[151,270,1962,1986]" pageId="19" pageNumber="20">PLATE 7.</emphasis>
Figs. 24–31: Front view of head. 24–25 
<taxonomicName id="2EF81BA51E51FF8D61EDF852AE41D8BD" box="[732,864,1963,1986]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="adspersa">
<emphasis id="DB8CBC341E51FF8D61EDF852AE41D8BD" box="[732,864,1963,1986]" italics="true" pageId="19" pageNumber="20">A. adspersa</emphasis>
</taxonomicName>
; 26–27 
<taxonomicName id="2EF81BA51E51FF8D608CF852A921D8BD" box="[957,1024,1963,1986]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="rex" status="sp. nov.">
<emphasis id="DB8CBC341E51FF8D608CF852A921D8BD" box="[957,1024,1963,1986]" italics="true" pageId="19" pageNumber="20">A. rex</emphasis>
</taxonomicName>
<emphasis id="DB8CBC341E51FF8D6736F853A943D8BD" bold="true" box="[1031,1122,1962,1986]" pageId="19" pageNumber="20">
<taxonomicNameLabel id="C0BF014F1E51FF8D6736F853A943D8BD" box="[1031,1122,1962,1986]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; 28–29 
<taxonomicName id="2EF81BA51E51FF8D678EF852A806D8BD" box="[1215,1319,1963,1986]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="crassa">
<emphasis id="DB8CBC341E51FF8D678EF852A806D8BD" box="[1215,1319,1963,1986]" italics="true" pageId="19" pageNumber="20">A. crassa</emphasis>
</taxonomicName>
; 30–31 
<taxonomicName id="2EF81BA51E51FF8D66B5F852AC2ED89B" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="ugandensis">
<emphasis id="DB8CBC341E51FF8D66B5F852AC2ED89B" italics="true" pageId="19" pageNumber="20">A. ugandensis</emphasis>
</taxonomicName>
. Scale bars on left = 
<quantity id="2E00CDC31E51FF8D62C0F834AF13D89B" box="[497,562,1997,2020]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="19" pageNumber="20" unit="mm" value="1.0">1 mm</quantity>
; right = 100 µm.
</paragraph>
<paragraph id="E94760261E56FF8A63A6F864A972D8A8" blockId="20.[151,1436,1949,2007]" pageId="20" pageNumber="21">
<emphasis id="DB8CBC341E56FF8A63A6F864AC2AD8CA" bold="true" box="[151,267,1949,1973]" pageId="20" pageNumber="21">PLATE 8.</emphasis>
Figs. 32–39: Front view of head. 32–33 
<taxonomicName id="2EF81BA51E56FF8A61FDF867AE0DD8CA" box="[716,812,1950,1973]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="DB8CBC341E56FF8A61FDF867AE0DD8CA" box="[716,812,1950,1973]" italics="true" pageId="20" pageNumber="21">A. minor</emphasis>
</taxonomicName>
(black form); 34–35 
<taxonomicName id="2EF81BA51E56FF8A6726F867A9A3D8CA" box="[1047,1154,1950,1973]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="crypsis" status="sp. nov.">
<emphasis id="DB8CBC341E56FF8A6726F867A9A3D8CA" box="[1047,1154,1950,1973]" italics="true" pageId="20" pageNumber="21">A. crypsis</emphasis>
</taxonomicName>
<emphasis id="DB8CBC341E56FF8A67BBF864A9C3D8CA" bold="true" box="[1162,1250,1949,1973]" pageId="20" pageNumber="21">
<taxonomicNameLabel id="C0BF014F1E56FF8A67BBF864A9C3D8CA" box="[1162,1250,1949,1973]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; 36–37 
<taxonomicName id="2EF81BA51E56FF8A660DF867A8BDD8CA" box="[1340,1436,1950,1973]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="DB8CBC341E56FF8A660DF867A8BDD8CA" box="[1340,1436,1950,1973]" italics="true" pageId="20" pageNumber="21">A. minor</emphasis>
</taxonomicName>
(black and russet form); 38–39 
<taxonomicName id="2EF81BA51E56FF8A62D7F839AF5CD8A8" box="[486,637,1984,2007]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="subfasciata">
<emphasis id="DB8CBC341E56FF8A62D7F839AF5CD8A8" box="[486,637,1984,2007]" italics="true" pageId="20" pageNumber="21">A. subfasciata</emphasis>
</taxonomicName>
. Scale bars on left = 
<quantity id="2E00CDC31E56FF8A606EF839AE81D8A8" box="[863,928,1984,2007]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="20" pageNumber="21" unit="mm" value="1.0">1 mm</quantity>
; right = 100 µm.
</paragraph>
<paragraph id="E94760261E57FF8B63A6F9A6A84AD9E6" blockId="21.[151,1436,1631,1690]" pageId="21" pageNumber="22">
<emphasis id="DB8CBC341E57FF8B63A6F9A6AC28D908" bold="true" box="[151,265,1631,1655]" pageId="21" pageNumber="22">PLATE 9.</emphasis>
Figs. 40–45: Dorsal view of head and pronotal apex of: 40 
<taxonomicName id="2EF81BA51E57FF8B604EF999AEDCD908" box="[895,1021,1632,1655]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="adspersa">
<emphasis id="DB8CBC341E57FF8B604EF999AEDCD908" box="[895,1021,1632,1655]" italics="true" pageId="21" pageNumber="22">A. adspersa</emphasis>
</taxonomicName>
; 42 
<taxonomicName id="2EF81BA51E57FF8B671DF999A94BD908" box="[1068,1130,1632,1655]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="rex" status="sp. nov.">
<emphasis id="DB8CBC341E57FF8B671DF999A94BD908" box="[1068,1130,1632,1655]" italics="true" pageId="21" pageNumber="22">A. rex</emphasis>
</taxonomicName>
<emphasis id="DB8CBC341E57FF8B6741F9A6A9E4D908" bold="true" box="[1136,1221,1631,1655]" pageId="21" pageNumber="22">
<taxonomicNameLabel id="C0BF014F1E57FF8B6741F9A6A9E4D908" box="[1136,1221,1631,1655]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; 44 
<taxonomicName id="2EF81BA51E57FF8B67C5F999A877D908" box="[1268,1366,1632,1655]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="crassa">
<emphasis id="DB8CBC341E57FF8B67C5F999A877D908" box="[1268,1366,1632,1655]" italics="true" pageId="21" pageNumber="22">A. crassa</emphasis>
</taxonomicName>
; 45 
<taxonomicName id="2EF81BA51E57FF8B66B4F999AC2ED9E6" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="ugandensis">
<emphasis id="DB8CBC341E57FF8B66B4F999AC2ED9E6" italics="true" pageId="21" pageNumber="22">A. ugandensis</emphasis>
</taxonomicName>
. 41, 43 area to left of right hand eye enlarged: 41. 
<taxonomicName id="2EF81BA51E57FF8B601AF97BAE88D9E6" box="[811,937,1666,1689]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="adspersa">
<emphasis id="DB8CBC341E57FF8B601AF97BAE88D9E6" box="[811,937,1666,1689]" italics="true" pageId="21" pageNumber="22">A. adspersa</emphasis>
</taxonomicName>
; 43. 
<taxonomicName id="2EF81BA51E57FF8B60EEF97BA93CD9E6" box="[991,1053,1666,1689]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="rex" status="sp. nov.">
<emphasis id="DB8CBC341E57FF8B60EEF97BA93CD9E6" box="[991,1053,1666,1689]" italics="true" pageId="21" pageNumber="22">A. rex</emphasis>
</taxonomicName>
<emphasis id="DB8CBC341E57FF8B6715F97BA95BD9E5" bold="true" box="[1060,1146,1666,1690]" pageId="21" pageNumber="22">
<taxonomicNameLabel id="C0BF014F1E57FF8B6715F97BA95BD9E5" box="[1060,1146,1666,1690]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; Scale bars = 100 µm.
</paragraph>
</subSubSection>
<subSubSection id="A1E233AD1E57FF8B63F4F919AF1CD89D" pageId="21" pageNumber="22" type="materials_examined">
<paragraph id="E94760261E57FF8B63F4F919AF1CD89D" blockId="21.[151,1436,1760,2018]" pageId="21" pageNumber="22">
<emphasis id="DB8CBC341E57FF8B63F4F919AC70D985" bold="true" box="[197,337,1760,1786]" pageId="21" pageNumber="22">Pest status.</emphasis>
A long series of laboratory reared and field collected specimens from 
<collectionCode id="8FE9F8E31E57FF8B67A0F919A9C5D985" box="[1169,1252,1760,1786]" pageId="21" pageNumber="22">SASRI</collectionCode>
(South African Sugar Research Institute) (previously 
<collectionCode id="8FE9F8E31E57FF8B6159F8FFAFEAD85F" box="[616,715,1798,1824]" pageId="21" pageNumber="22">SASEX</collectionCode>
, South African Sugar Experiment Station) were examined. 
<collectionCode id="8FE9F8E31E57FF8B63A6F8D4ADCAD838" box="[151,235,1837,1863]" pageId="21" pageNumber="22">SASRI</collectionCode>
accession numbers for material of 
<taxonomicName id="2EF81BA51E57FF8B61BCF8D4AFD4D839" box="[653,757,1837,1862]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="DB8CBC341E57FF8B61BCF8D4AFD4D839" box="[653,757,1837,1862]" italics="true" pageId="21" pageNumber="22">A. minor</emphasis>
</taxonomicName>
include 41, 828, 833 and 1853. The 
<collectionCode id="8FE9F8E31E57FF8B679CF8D4A820D838" box="[1197,1281,1837,1863]" pageId="21" pageNumber="22">SASRI</collectionCode>
pair labelled only “1996 lab reared” 
<collectionCode id="8FE9F8E31E57FF8B6285F8ADAF32D811" box="[436,531,1876,1902]" pageId="21" pageNumber="22">SASEX</collectionCode>
828 are considerably larger than the mean size for 
<taxonomicName id="2EF81BA51E57FF8B67B2F8ADA9CAD812" box="[1155,1259,1876,1901]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="DB8CBC341E57FF8B67B2F8ADA9CAD812" box="[1155,1259,1876,1901]" italics="true" pageId="21" pageNumber="22">A. minor</emphasis>
</taxonomicName>
, but it is quite probable that this is due to being well fed under laboratory conditions rather than having emerged naturally. See material examined for localities from where specimens were collected during the 
<collectionCode id="8FE9F8E31E57FF8B678BF858A831D8C4" box="[1210,1296,1953,1979]" pageId="21" pageNumber="22">SASRI</collectionCode>
white grub survey undertaken over 
<date id="9D4646E61E57FF8B6280F831AF19D89D" box="[433,568,1992,2018]" pageId="21" pageNumber="22" value="1996" valueMax="1997">1996–1997</date>
.
</paragraph>
</subSubSection>
<subSubSection id="A1E233AD1E54FF8863F4FF6EAFDBDD4B" pageId="22" pageNumber="23" type="discussion">
<paragraph id="E94760261E54FF8863F4FF6EACBBDE59" blockId="22.[151,1437,151,487]" pageId="22" pageNumber="23">
<emphasis id="DB8CBC341E54FF8863F4FF6EAC1FDFCE" bold="true" box="[197,318,151,177]" pageId="22" pageNumber="23">Remarks.</emphasis>
<bibRefCitation id="8D691DD71E54FF886274FF61AF31DFCD" author="Peringuey" box="[325,528,152,178]" pageId="22" pageNumber="44" refString="Peringuey, L. (1904) Descriptive catalogue of the Coleoptera of South Africa (Lucanidae and Scarabaeidae). Transactions of the South African Philosophical Society, 13, 1 - 293 + 4 plates." type="book chapter" year="1904">Péringuey (1904)</bibRefCitation>
was unable to differentiate between the morphological variability at intra and interspecific levels in 
<taxonomicName id="2EF81BA51E54FF8862AEFF46AF66DFA7" box="[415,583,191,216]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="DB8CBC341E54FF8862AEFF46AF66DFA7" box="[415,583,191,216]" italics="true" pageId="22" pageNumber="23">Asthenopholis</emphasis>
</taxonomicName>
and thus placed 
<taxonomicName id="2EF81BA51E54FF886023FF46AE5ADFA7" box="[786,891,191,216]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="DB8CBC341E54FF886023FF46AE5ADFA7" box="[786,891,191,216]" italics="true" pageId="22" pageNumber="23">A. minor</emphasis>
</taxonomicName>
as conspecific with 
<emphasis id="DB8CBC341E54FF886741FF46A800DFA7" box="[1136,1313,191,216]" italics="true" pageId="22" pageNumber="23">
<taxonomicName id="2EF81BA51E54FF886741FF46A83CDFA7" box="[1136,1309,191,216]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="subfasciata">A. subfasciata</taxonomicName>
.
</emphasis>
However, both the distribution (Fig. 22) and external morphology of these two species differ (see species key and respective diagnoses).
</paragraph>
<paragraph id="E94760261E54FF8863F4FECBA91FDE98" blockId="22.[151,1437,151,487]" pageId="22" pageNumber="23">
The 
<typeStatus id="3643DE841E54FF8863CBFECBAC19DE33" box="[250,312,306,332]" pageId="22" pageNumber="23">types</typeStatus>
series examined comprises two incompletely sclerotised males. However, they match well with recently collected specimens of 
<taxonomicName id="2EF81BA51E54FF88612CFEA0AFA4DE0D" box="[541,645,345,370]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="DB8CBC341E54FF88612CFEA0AFA4DE0D" box="[541,645,345,370]" italics="true" pageId="22" pageNumber="23">A. minor</emphasis>
</taxonomicName>
, which include both completely (black cuticle) and incompletely (reddish black cuticle) sclerotised specimens. Brenske’s specimen of “ 
<taxonomicName id="2EF81BA51E54FF886738FE79A98BDEE6" box="[1033,1194,384,410]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="bechuana">
<emphasis id="DB8CBC341E54FF886738FE79A93ADEE6" box="[1033,1051,384,409]" italics="true" pageId="22" pageNumber="23">A</emphasis>
. 
<emphasis id="DB8CBC341E54FF88671DFE79A98BDEE6" box="[1068,1194,384,409]" italics="true" pageId="22" pageNumber="23">bechuana</emphasis>
</taxonomicName>
” is a female, while Péringuey’s specimen of “ 
<taxonomicName id="2EF81BA51E54FF8862ECFE5EAF5FDEBF" box="[477,638,422,448]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="bechuana">
<emphasis id="DB8CBC341E54FF8862ECFE5EACCEDEBF" box="[477,495,423,448]" italics="true" pageId="22" pageNumber="23">A</emphasis>
. 
<emphasis id="DB8CBC341E54FF886130FE5EAF5FDEBF" box="[513,638,423,448]" italics="true" pageId="22" pageNumber="23">bechuana</emphasis>
</taxonomicName>
” is a male. Although 
<bibRefCitation id="8D691DD71E54FF8860BCFE5FA944DEBF" author="Peringuey" box="[909,1125,422,448]" pageId="22" pageNumber="44" refString="Peringuey, L. (1904) Descriptive catalogue of the Coleoptera of South Africa (Lucanidae and Scarabaeidae). Transactions of the South African Philosophical Society, 13, 1 - 293 + 4 plates." type="book chapter" year="1904">Péringuey (1904)</bibRefCitation>
attributed this species to Brenske (in litt.), it was never actually described by Brenske, who died in 1904.
</paragraph>
<paragraph id="E94760261E54FF8863A6FDE3AFDBDD4B" blockId="22.[151,1437,538,1570]" box="[151,762,538,564]" pageId="22" pageNumber="23">
<taxonomicName id="2EF81BA51E54FF8863A6FDE3ACABDD4C" box="[151,394,538,563]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="DB8CBC341E54FF8863A6FDE3ACABDD4C" box="[151,394,538,563]" italics="true" pageId="22" pageNumber="23">Asthenopholis minor</emphasis>
</taxonomicName>
- narrow black and russet form
</paragraph>
</subSubSection>
<subSubSection id="A1E233AD1E54FF8663F7FDB9A9E2DCC8" lastPageId="24" lastPageNumber="25" pageId="22" pageNumber="23" type="materials_examined">
<paragraph id="E94760261E54FF8863F7FDB9A906DCEF" blockId="22.[151,1437,538,1570]" pageId="22" pageNumber="23">
<emphasis id="DB8CBC341E54FF8863F7FDB9AEEDDD25" bold="true" box="[198,972,576,603]" pageId="22" pageNumber="23">
<typeStatus id="3643DE841E54FF8863F7FDB9AC22DD25" box="[198,259,576,602]" pageId="22" pageNumber="23">Type</typeStatus>
material examined (
<taxonomicName id="2EF81BA51E54FF886134FDB8AE9FDD25" authority="Brenske, 1898" authorityName="Brenske" authorityYear="1898" box="[517,958,576,603]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="DB8CBC341E54FF886134FDB8AE23DD24" bold="true" box="[517,770,577,603]" italics="true" pageId="22" pageNumber="23">Asthenopholis minor</emphasis>
<bibRefCitation id="8D691DD71E54FF886038FDB9AE9FDD25" author="Brenske" box="[777,958,576,602]" pageId="22" pageNumber="43" refString="Brenske, E. (1898) Melolonthiden aus Afrika. Stettiner Entomologische Zeitung, 59 (10 - 12) (iv), 333 - 394." type="journal article" year="1898">Brenske, 1898</bibRefCitation>
</taxonomicName>
):
</emphasis>
<typeStatus id="3643DE841E54FF8860E2FDB8A969DD24" box="[979,1096,577,603]" pageId="22" pageNumber="23" type="lectotype">Lectotype</typeStatus>
(
<emphasis id="DB8CBC341E54FF886768FDB9A83DDD25" bold="true" box="[1113,1308,576,602]" pageId="22" pageNumber="23">here designated</emphasis>
): 
<collectingCountry id="91EF20B61E54FF886604FDB9AC2CDDFE" name="South Africa" pageId="22" pageNumber="23">
<emphasis id="DB8CBC341E54FF886604FDB9AC2CDDFE" bold="true" pageId="22" pageNumber="23">SOUTH AFRICA</emphasis>
</collectingCountry>
, “Transvaal / [male genitalia card, but genitalia missing] / 
<taxonomicName id="2EF81BA51E54FF8860DDFD91A9CDDDFE" box="[1004,1260,616,641]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="DB8CBC341E54FF8860DDFD91A9CDDDFE" box="[1004,1260,616,641]" italics="true" pageId="22" pageNumber="23">Asthenopholis minor</emphasis>
</taxonomicName>
<typeStatus id="3643DE841E54FF8867C7FD91A80EDDFD" box="[1270,1327,616,642]" pageId="22" pageNumber="23">type</typeStatus>
. 3 Brsk. [written by Brenske] / Coll. Brenske / 
<typeStatus id="3643DE841E54FF886142FD77AF8EDDD7" box="[627,687,654,680]" pageId="22" pageNumber="23">Type</typeStatus>
[printed on reddish brown card] / Zool. Mus. Berlin / 0 2602 [JduGH photograph number] / Lecotype 3 
<taxonomicName id="2EF81BA51E54FF8861A4FD4CA91CDDB0" authority="Brenske, 1898" authorityName="Brenske" authorityYear="1898" box="[661,1085,693,719]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="DB8CBC341E54FF8861A4FD4CAEA8DDB1" box="[661,905,693,718]" italics="true" pageId="22" pageNumber="23">Asthenopholis minor</emphasis>
<bibRefCitation id="8D691DD71E54FF8860A1FD4CA91CDDB0" author="Brenske" box="[912,1085,693,719]" pageId="22" pageNumber="43" refString="Brenske, E. (1898) Melolonthiden aus Afrika. Stettiner Entomologische Zeitung, 59 (10 - 12) (iv), 333 - 394." type="journal article" year="1898">Brenske, 1898</bibRefCitation>
</taxonomicName>
. des. J. du G. Harrison, 2009” [specimen condition; right hand side antennal club missing, left hand side protarsi and mesotibia missing, dermestid perforation in abdominal sternites] (13 
<collectionCode id="8FE9F8E31E54FF8861D0FCFBAE1ADC63" box="[737,827,770,796]" pageId="22" pageNumber="23">ZMHB</collectionCode>
); One 
<typeStatus id="3643DE841E54FF8860B4FCFBA909DC63" box="[901,1064,770,796]" pageId="22" pageNumber="23" type="paralectotype">paralectotype</typeStatus>
: “Transvaal, D. Heyne [written] / [male genitalia card mounted] / 
<taxonomicName id="2EF81BA51E54FF88612DFCD0AE2FDC3D" box="[540,782,809,834]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="DB8CBC341E54FF88612DFCD0AE2FDC3D" box="[540,782,809,834]" italics="true" pageId="22" pageNumber="23">Asthenopholis minor</emphasis>
</taxonomicName>
cotype. Brsk [written by Brenske] / Coll. Brenske / Co- [written in black] 
<typeStatus id="3643DE841E54FF88625FFCA9AC88DC15" box="[366,425,848,874]" pageId="22" pageNumber="23">Type</typeStatus>
[printed on reddish brown card] / Zool. Mus. Berlin / Paralecotype 3 
<taxonomicName id="2EF81BA51E54FF8867C5FCA9ACB0DCEF" authority="Brenske, 1898" authorityName="Brenske" authorityYear="1898" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="DB8CBC341E54FF8867C5FCA9ADFCDCEF" italics="true" pageId="22" pageNumber="23">Asthenopholis minor</emphasis>
<bibRefCitation id="8D691DD71E54FF8863D5FC8FACB0DCEF" author="Brenske" box="[228,401,886,912]" pageId="22" pageNumber="43" refString="Brenske, E. (1898) Melolonthiden aus Afrika. Stettiner Entomologische Zeitung, 59 (10 - 12) (iv), 333 - 394." type="journal article" year="1898">Brenske, 1898</bibRefCitation>
</taxonomicName>
. des. J. du G. Harrison, 2009 / JduGH128” (13 
<collectionCode id="8FE9F8E31E54FF8860F1FC8FA93BDCEF" box="[960,1050,886,912]" pageId="22" pageNumber="23">ZMHB</collectionCode>
).
</paragraph>
<paragraph id="E94760261E54FF8863F4FC64A81FD95D" blockId="22.[151,1437,538,1570]" pageId="22" pageNumber="23">
<emphasis id="DB8CBC341E54FF8863F4FC64AF1CDCC8" bold="true" box="[197,573,925,951]" pageId="22" pageNumber="23">Additional material examined</emphasis>
(with similar morphology to the 
<typeStatus id="3643DE841E54FF8860F4FC64A911DCC8" box="[965,1072,925,951]" pageId="22" pageNumber="23" type="lectotype">lectotype</typeStatus>
and 
<typeStatus id="3643DE841E54FF88675BFC64A82BDCC8" box="[1130,1290,925,951]" pageId="22" pageNumber="23" type="paralectotype">paralectotype</typeStatus>
) (Σ16, 153, 1Ƥ). 
<collectingCountry id="91EF20B61E54FF8863E7FC3AAC9ADCA2" box="[214,443,963,989]" name="South Africa" pageId="22" pageNumber="23">
<emphasis id="DB8CBC341E54FF8863E7FC3AAC9ADCA2" bold="true" box="[214,443,963,989]" pageId="22" pageNumber="23">SOUTH AFRICA</emphasis>
</collectingCountry>
,? Empangeni, 1916, W.J. Haygarth / 
<collectionCode id="8FE9F8E31E54FF8860B8FC3DAEFEDCA1" box="[905,991,964,990]" httpUri="http://grbio.org/cool/i1wq-4y1w" name="Durban Museum" pageId="22" pageNumber="23">DMSA</collectionCode>
<collectionCode id="8FE9F8E31E54FF8860D9FC3DA905DCA1" LSID="urn:lsid:biocol.org:col:15629" box="[1000,1060,964,990]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15629" name="Universidad Nacional de Colombia" pageId="22" pageNumber="23">COL</collectionCode>
11994 /? 
<taxonomicName id="2EF81BA51E54FF8867ADFC3DA80EDCA2" box="[1180,1327,964,989]" class="Insecta" family="Melolonthidae" genus="Coniopholis" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="DB8CBC341E54FF8867ADFC3DA80EDCA2" box="[1180,1327,964,989]" italics="true" pageId="22" pageNumber="23">Coniopholis</emphasis>
</taxonomicName>
sp. nova [incorrect genus] / JduGH101 (13 
<collectionCode id="8FE9F8E31E54FF88610CFC13AFB9DB7B" box="[573,664,1002,1028]" httpUri="http://grbio.org/cool/i1wq-4y1w" name="Durban Museum" pageId="22" pageNumber="23">DMSA</collectionCode>
); nr [near] Eshowe, W.J. Haygarth, Ac. No. 1980 / 
<collectionCode id="8FE9F8E31E54FF886632FC13A878DB7B" box="[1283,1369,1002,1028]" httpUri="http://grbio.org/cool/i1wq-4y1w" name="Durban Museum" pageId="22" pageNumber="23">DMSA</collectionCode>
<collectionCode id="8FE9F8E31E54FF886650FC13A8BCDB7B" LSID="urn:lsid:biocol.org:col:15629" box="[1377,1437,1002,1028]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15629" name="Universidad Nacional de Colombia" pageId="22" pageNumber="23">COL</collectionCode>
11860 / JduGH100 (13 
<collectionCode id="8FE9F8E31E54FF886283FBE8AF2DDB54" box="[434,524,1041,1067]" httpUri="http://grbio.org/cool/i1wq-4y1w" name="Durban Museum" pageId="22" pageNumber="23">DMSA</collectionCode>
); Qachas Nek, 
<collectingCountry id="91EF20B61E54FF88618FFBE8AE63DB54" box="[702,834,1041,1067]" name="Lesotho" pageId="22" pageNumber="23">Basutoland</collectingCountry>
[
<collectingCountry id="91EF20B61E54FF88607EFBE8AE97DB54" box="[847,950,1041,1067]" name="Lesotho" pageId="22" pageNumber="23">Lesotho</collectingCountry>
], 
<date id="9D4646E61E54FF8860F8FBE8A95FDB54" box="[969,1150,1041,1067]" pageId="22" pageNumber="23" value="1939-01-08" valueMax="1939-01-13" valueMin="1939-01-08">8–13 Jan. 1939</date>
, L. Bevis / 
<collectionCode id="8FE9F8E31E54FF886634FBE8A87BDB54" box="[1285,1370,1041,1067]" httpUri="http://grbio.org/cool/i1wq-4y1w" name="Durban Museum" pageId="22" pageNumber="23">DMSA</collectionCode>
<collectionCode id="8FE9F8E31E54FF886650FBE8A8BDDB54" LSID="urn:lsid:biocol.org:col:15629" box="[1377,1436,1041,1067]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15629" name="Universidad Nacional de Colombia" pageId="22" pageNumber="23">COL</collectionCode>
11995 / JduGH78 (13 
<collectionCode id="8FE9F8E31E54FF886281FBC1AF2CDB2D" box="[432,525,1080,1106]" httpUri="http://grbio.org/cool/i1wq-4y1w" name="Durban Museum" pageId="22" pageNumber="23">DMSA</collectionCode>
); Qachas Nek, 
<date id="9D4646E61E54FF8861F6FBC1AE41DB2D" box="[711,864,1080,1106]" pageId="22" pageNumber="23" value="2005-12-02">2 Dec. 2005</date>
, 1716 m, grassland, Site No. 35, active flying, 
<collectionCode id="8FE9F8E31E54FF8863A6FBA7ADCADB07" box="[151,235,1118,1144]" pageId="22" pageNumber="23">MDTP</collectionCode>
2705 / JduGH556 (13 
<collectionCode id="8FE9F8E31E54FF88613CFBA7AF47DB07" box="[525,614,1118,1144]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="22" pageNumber="23">TMSA</collectionCode>
); Qachas Nek, 
<date id="9D4646E61E54FF886011FBA7AE9BDB07" box="[800,954,1118,1144]" pageId="22" pageNumber="23" value="2005-12-02">2 Dec. 2005</date>
, 1716 m, grassland, Site No. 35, active flying, 
<collectionCode id="8FE9F8E31E54FF8863C1FB7CAC62DBE0" box="[240,323,1157,1183]" pageId="22" pageNumber="23">MDTP</collectionCode>
3967 / JduGH557 (13 
<collectionCode id="8FE9F8E31E54FF886155FB7CAF9CDBE0" box="[612,701,1157,1183]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="22" pageNumber="23">TMSA</collectionCode>
); Qachas Nek, 
<date id="9D4646E61E54FF886047FB7CA92FDBE0" box="[886,1038,1157,1183]" pageId="22" pageNumber="23" value="2005-12-02">2 Dec. 2005</date>
, 1716 m, grassland, Site No. 35, active flying, 
<collectionCode id="8FE9F8E31E54FF88620AFB55ACACDBB9" box="[315,397,1196,1222]" pageId="22" pageNumber="23">MDTP</collectionCode>
5003 / JduGH558 (13 
<collectionCode id="8FE9F8E31E54FF886196FB55AFDFDBB9" box="[679,766,1196,1222]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="22" pageNumber="23">TMSA</collectionCode>
); Qachas Nek, 
<date id="9D4646E61E54FF886083FB55A967DBB9" box="[946,1094,1196,1222]" pageId="22" pageNumber="23" value="2005-12-02">2 Dec. 2005</date>
, 1716 m, grassland, Site No. 35, active flying, 
<collectionCode id="8FE9F8E31E54FF886256FB2BAC98DB93" box="[359,441,1234,1260]" pageId="22" pageNumber="23">MDTP</collectionCode>
59051 / JduGH559 (13 
<collectionCode id="8FE9F8E31E54FF8861EEFB2BAE17DB93" box="[735,822,1234,1260]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="22" pageNumber="23">TMSA</collectionCode>
); Qachas Nek, 
<date id="9D4646E61E54FF8860D8FB2BA95ADB93" box="[1001,1147,1234,1260]" pageId="22" pageNumber="23" value="2005-12-02">2 Dec. 2005</date>
, 1692 m, grassland, Site No. 36, active flying, 
<collectionCode id="8FE9F8E31E54FF8862AFFB00ACD1DA6C" box="[414,496,1273,1299]" pageId="22" pageNumber="23">MDTP</collectionCode>
6254 / JduGH560 (13 
<collectionCode id="8FE9F8E31E54FF88603DFB00AE42DA6C" box="[780,867,1273,1299]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="22" pageNumber="23">TMSA</collectionCode>
); Qachas Nek, 
<date id="9D4646E61E54FF886728FB00A98CDA6C" box="[1049,1197,1273,1299]" pageId="22" pageNumber="23" value="2005-12-02">2 Dec. 2005</date>
, 1692 m, grassland, Site No. 36, active flying, 
<collectionCode id="8FE9F8E31E54FF8862DAFAD9AF60DA45" box="[491,577,1312,1338]" pageId="22" pageNumber="23">MDTP</collectionCode>
6784 / JduGH561 (13 
<collectionCode id="8FE9F8E31E54FF88605CFAD9AEE8DA45" box="[877,969,1312,1338]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="22" pageNumber="23">TMSA</collectionCode>
); Qachas Nek, 
<date id="9D4646E61E54FF8867B8FAD9A806DA45" box="[1161,1319,1312,1338]" pageId="22" pageNumber="23" value="2005-12-02">2 Dec. 2005</date>
, 1692 m, grassland, Site No. 36, active flying, 
<collectionCode id="8FE9F8E31E54FF88617FFABFAFBEDA1F" box="[590,671,1350,1376]" pageId="22" pageNumber="23">MDTP</collectionCode>
7092 / JduGH562 (13 
<collectionCode id="8FE9F8E31E54FF886089FABFA92EDA1F" box="[952,1039,1350,1376]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="22" pageNumber="23">TMSA</collectionCode>
); Qachas Nek, 
<date id="9D4646E61E54FF8867F3FABFA875DA1F" box="[1218,1364,1350,1376]" pageId="22" pageNumber="23" value="2005-12-02">2 Dec. 2005</date>
, 1692 m, grassland, Site No. 36, general, 
<collectionCode id="8FE9F8E31E54FF886103FA94AFA2DAF8" box="[562,643,1389,1415]" pageId="22" pageNumber="23">MDTP</collectionCode>
6546 / JduGH563 (13 
<collectionCode id="8FE9F8E31E54FF8860A6FA94AECFDAF8" box="[919,1006,1389,1415]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="22" pageNumber="23">TMSA</collectionCode>
); Qachas Nek, 
<date id="9D4646E61E54FF8867AEFA94A80EDAF8" box="[1183,1327,1389,1415]" pageId="22" pageNumber="23" value="2005-12-02">2 Dec. 2005</date>
, 1692 m, grassland, Site No. 36, general, 
<collectionCode id="8FE9F8E31E54FF886127FA6DAF49DAD1" box="[534,616,1428,1454]" pageId="22" pageNumber="23">MDTP</collectionCode>
58585 / JduGH564 (13 
<collectionCode id="8FE9F8E31E54FF8860A2FA6DAECADAD1" box="[915,1003,1428,1454]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="22" pageNumber="23">TMSA</collectionCode>
); 
<collectingCountry id="91EF20B61E54FF886731FA6DA94DDAD1" box="[1024,1132,1428,1454]" name="Saint Lucia" pageId="22" pageNumber="23">St. Lucia</collectingCountry>
Lake, H.W. Bell-Marley, 
<date id="9D4646E61E54FF8863A6FA43AC2DDAAB" box="[151,268,1466,1492]" pageId="22" pageNumber="23" value="1922-10">Oct. 1922</date>
/ 
<taxonomicName id="2EF81BA51E54FF886215FA42AC95DAAB" box="[292,436,1467,1492]" class="Insecta" family="Melolonthidae" genus="Coniopholis" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="DB8CBC341E54FF886215FA42AC95DAAB" box="[292,436,1467,1492]" italics="true" pageId="22" pageNumber="23">Coniopholis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C0BF014F1E54FF88628CFA43AF38DAAB" box="[445,537,1466,1492]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
[incorrect genus] / 
<collectionCode id="8FE9F8E31E54FF886032FA43AE79DAAB" box="[771,856,1466,1492]" httpUri="http://grbio.org/cool/i1wq-4y1w" name="Durban Museum" pageId="22" pageNumber="23">DMSA</collectionCode>
<collectionCode id="8FE9F8E31E54FF886051FA43AEBADAAB" LSID="urn:lsid:biocol.org:col:15629" box="[864,923,1466,1492]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15629" name="Universidad Nacional de Colombia" pageId="22" pageNumber="23">COL</collectionCode>
11990 / JduGH98 (1Ƥ 
<collectionCode id="8FE9F8E31E54FF886785FA43A82FDAAB" box="[1204,1294,1466,1492]" httpUri="http://grbio.org/cool/i1wq-4y1w" name="Durban Museum" pageId="22" pageNumber="23">DMSA</collectionCode>
); Umfolozi, 
<date id="9D4646E61E54FF8863A6FA18AC37DA84" box="[151,278,1505,1531]" pageId="22" pageNumber="23" value="1956-10">Oct. 1956</date>
, Zumpt / 
<taxonomicName id="2EF81BA51E54FF8862A1FA18A935DA84" authority="Brenske Det. J. Decelle, 1969" authorityName="Brenske Det. J. Decelle" authorityYear="1969" box="[400,1044,1505,1531]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="DB8CBC341E54FF8862A1FA18AFB5DA85" box="[400,660,1505,1530]" italics="true" pageId="22" pageNumber="23">Asthenopholis minor</emphasis>
Brenske Det. J. Decelle, 1969
</taxonomicName>
/ JduGH460 (13 
<collectionCode id="8FE9F8E31E54FF8867C6FA18A877DA84" box="[1271,1366,1505,1531]" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="22" pageNumber="23">MRAC</collectionCode>
), (13 
<collectionCode id="8FE9F8E31E54FF8863A6F9F1ADCFD95D" box="[151,238,1544,1570]" pageId="22" pageNumber="23">ZSMG</collectionCode>
); Zululand, W.J. Haygarth, Ac. No. 
<date id="9D4646E61E54FF8861A2F9F1AE36D95D" box="[659,791,1544,1570]" pageId="22" pageNumber="23" value="1881" valueMax="1916">1881–1916</date>
/ 
<collectionCode id="8FE9F8E31E54FF88601FF9F1AEA2D95D" box="[814,899,1544,1570]" httpUri="http://grbio.org/cool/i1wq-4y1w" name="Durban Museum" pageId="22" pageNumber="23">DMSA</collectionCode>
<collectionCode id="8FE9F8E31E54FF8860BBF9F1AEE4D95D" LSID="urn:lsid:biocol.org:col:15629" box="[906,965,1544,1570]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15629" name="Universidad Nacional de Colombia" pageId="22" pageNumber="23">COL</collectionCode>
11861 / JduGH97 (13 
<collectionCode id="8FE9F8E31E54FF8867E9F9F1A813D95D" box="[1240,1330,1544,1570]" httpUri="http://grbio.org/cool/i1wq-4y1w" name="Durban Museum" pageId="22" pageNumber="23">DMSA</collectionCode>
).
</paragraph>
<paragraph id="E94760261E54FF8863A6F9ACAFBED910" blockId="22.[151,1438,1621,2034]" box="[151,671,1621,1647]" pageId="22" pageNumber="23">
<taxonomicName id="2EF81BA51E54FF8863A6F9ACACABD911" box="[151,394,1621,1646]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="DB8CBC341E54FF8863A6F9ACACABD911" box="[151,394,1621,1646]" italics="true" pageId="22" pageNumber="23">Asthenopholis minor</emphasis>
</taxonomicName>
- black and russet form
</paragraph>
<paragraph id="E94760261E54FF8863F7F982A8BDD88D" blockId="22.[151,1438,1621,2034]" pageId="22" pageNumber="23">
<emphasis id="DB8CBC341E54FF8863F7F982ACE3D9EA" bold="true" box="[198,450,1659,1685]" pageId="22" pageNumber="23">Material examined</emphasis>
(Σ148, 593, 9Ƥ, 80 unsexed). 
<emphasis id="DB8CBC341E54FF8860B6F982A957D9EA" bold="true" box="[903,1142,1659,1685]" pageId="22" pageNumber="23">
<collectingCountry id="91EF20B61E54FF8860B6F982A957D9EA" box="[903,1142,1659,1685]" name="South Africa" pageId="22" pageNumber="23">SOUTH AFRICA</collectingCountry>
</emphasis>
, Betchuanen, Land. / 
<taxonomicName id="2EF81BA51E54FF8863A6F95AACD5D9C3" box="[151,500,1699,1724]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="betschuanana">
<emphasis id="DB8CBC341E54FF8863A6F95AACD5D9C3" box="[151,500,1699,1724]" italics="true" pageId="22" pageNumber="23">Asthenopholis betschuanana</emphasis>
</taxonomicName>
[
<emphasis id="DB8CBC341E54FF886136F95AAF08D9C3" box="[519,553,1699,1724]" italics="true" pageId="22" pageNumber="23">sic</emphasis>
] 
<typeStatus id="3643DE841E54FF88610DF95BAF59D9C3" box="[572,632,1698,1724]" pageId="22" pageNumber="23">Type</typeStatus>
Brsk. / Coll. Brenske / 
<typeStatus id="3643DE841E54FF8860ACF95BAEF8D9C3" box="[925,985,1698,1724]" pageId="22" pageNumber="23">Type</typeStatus>
/ Zool. Mus. Berlin / 0 2610 [JduGH photograph number] / JduGH108 (1Ƥ 
<collectionCode id="8FE9F8E31E54FF88616FF930AF99D99C" box="[606,696,1737,1763]" pageId="22" pageNumber="23">ZMHB</collectionCode>
). 
<collectingCountry id="91EF20B61E54FF8861FDF930AE10D99C" box="[716,817,1737,1763]" name="South Africa" pageId="22" pageNumber="23">NATAL</collectingCountry>
., Frere. // A. 
<collectingCountry id="91EF20B61E54FF8860FAF930A91BD99C" box="[971,1082,1737,1763]" name="Marshall Islands" pageId="22" pageNumber="23">Marshall</collectingCountry>
., 1892. / 227 [written in black ink] / [mouthparts, card mounted by Péringuey] / [aedeagus, point mounted by Péringuey] / 
<taxonomicName id="2EF81BA51E54FF8867C3F909AC4CD84F" authority="Brsk." authorityName="Brsk." class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="beschuanus">
<emphasis id="DB8CBC341E54FF8867C3F909AC02D84F" italics="true" pageId="22" pageNumber="23">Asthenopholis beschuanus</emphasis>
Brsk.
</taxonomicName>
in litt., = 
<taxonomicName id="2EF81BA51E54FF8862D9F8EEAFBDD84F" authority="Brsk." authorityName="Brsk." box="[488,668,1814,1840]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="DB8CBC341E54FF8862D9F8EEAF73D84F" box="[488,594,1815,1840]" italics="true" pageId="22" pageNumber="23">A. minor</emphasis>
Brsk.
</taxonomicName>
the “…” [illegible] elytra [written in black ink by Péringuey] / SAM-COL-A035867 / JduGH109 (13 
<collectionCode id="8FE9F8E31E54FF8861BEF8C4AFCFD828" box="[655,750,1853,1879]" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="22" pageNumber="23">SAMC</collectionCode>
). Bethel, Beste, illegible, 
<taxonomicName id="2EF81BA51E54FF886773F8C4A8BCD828" authority="Brsk." authorityName="Brsk." box="[1090,1437,1853,1879]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="DB8CBC341E54FF886773F8C4A86FD829" box="[1090,1358,1853,1878]" italics="true" pageId="22" pageNumber="23">Asthenopholis minor</emphasis>
Brsk.
</taxonomicName>
[handwritten in black ink by Brenske] / JduGH102 (13 1Ƥ 
<collectionCode id="8FE9F8E31E54FF886086F89DA93BD801" box="[951,1050,1892,1918]" pageId="22" pageNumber="23">ZMHB</collectionCode>
); Botha's H.[ill], 
<date id="9D4646E61E54FF886634F89DA8A8D801" box="[1285,1417,1892,1918]" pageId="22" pageNumber="23" value="1897-02">Feb. 1897</date>
/ 
<taxonomicName id="2EF81BA51E54FF8863A6F872AF6DD8DB" authority="Blanch." authorityName="Blanch." box="[151,588,1930,1956]" class="Insecta" family="Dermestidae" genus="Cisthenopholis" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="subfasciata">
<emphasis id="DB8CBC341E54FF8863A6F872ACC4D8DB" box="[151,485,1931,1956]" italics="true" pageId="22" pageNumber="23">Cisthenopholis subfasciata</emphasis>
Blanch.
</taxonomicName>
[incorrect genus and species] / JduGH81 (13 
<collectionCode id="8FE9F8E31E54FF8867BFF873A9CAD8DB" box="[1166,1259,1930,1956]" httpUri="http://grbio.org/cool/i1wq-4y1w" name="Durban Museum" pageId="22" pageNumber="23">DMSA</collectionCode>
); Botha's Hill, G.A.K. 
<collectingCountry id="91EF20B61E54FF8863C1F848AC7AD8B4" box="[240,347,1969,1995]" name="Marshall Islands" pageId="22" pageNumber="23">Marshall</collectingCountry>
, B.M. 
<date id="9D4646E61E54FF88629BF848AF31D8B4" box="[426,528,1969,1995]" pageId="22" pageNumber="23" value="1912" valueMax="1995">1912–95</date>
/ JduGH82 (13 
<collectionCode id="8FE9F8E31E54FF8861E5F848AE10D8B4" box="[724,817,1969,1995]" pageId="22" pageNumber="23">BMNH</collectionCode>
); Carolina, 
<date id="9D4646E61E54FF886088F848A91ED8B4" box="[953,1087,1969,1995]" pageId="22" pageNumber="23" value="1931-12-19">19.12.1931</date>
, J.L. Swierstra / 
<taxonomicName id="2EF81BA51E54FF886632F848A84BD8B5" box="[1283,1386,1969,1994]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="DB8CBC341E54FF886632F848A84BD8B5" box="[1283,1386,1969,1994]" italics="true" pageId="22" pageNumber="23">A. minor</emphasis>
</taxonomicName>
det. A.V. Evans / JduGH88 (13 
<collectionCode id="8FE9F8E31E54FF8862C0F821AF6AD88D" box="[497,587,2008,2034]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="22" pageNumber="23">TMSA</collectionCode>
); Cathedral Peak, 
<date id="9D4646E61E54FF88601DF821AE96D88D" box="[812,951,2008,2034]" pageId="22" pageNumber="23" value="2003-11-25">25.11.2003</date>
, E-Y: 3571, grassland, M. Burger &amp; R.
</paragraph>
<paragraph id="E94760261E55FF8963A6FF61AFCBDA84" blockId="23.[151,1438,150,1531]" pageId="23" pageNumber="24">
Müller / JduGH519 (23 
<collectionCode id="8FE9F8E31E55FF8962F8FF61AF02DFCD" box="[457,547,152,178]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="23" pageNumber="24">TMSA</collectionCode>
); Cathedral Peak, 
<date id="9D4646E61E55FF896035FF61AEB1DFCD" box="[772,912,152,178]" pageId="23" pageNumber="24" value="1974-11-20">20.11.1974</date>
, E-Y: 765, singled on 
<taxonomicName id="2EF81BA51E55FF896795FF61A8AADFCD" authority="L. Schulze" authorityName="L. Schulze" box="[1188,1419,152,178]" class="Magnoliopsida" family="Proteaceae" genus="Protea" kingdom="Plantae" order="Proteales" pageId="23" pageNumber="24" phylum="Tracheophyta" rank="genus">
<emphasis id="DB8CBC341E55FF896795FF61A9D7DFCE" box="[1188,1270,152,177]" italics="true" pageId="23" pageNumber="24">Protea</emphasis>
, L. Schulze
</taxonomicName>
/ 
<taxonomicName id="2EF81BA51E55FF8963A6FF46AF0ADFA7" authority="Brenske Det." authorityName="Brenske Det." box="[151,555,190,216]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="23" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="DB8CBC341E55FF8963A6FF46ACAFDFA7" box="[151,398,191,216]" italics="true" pageId="23" pageNumber="24">Asthenopholis minor</emphasis>
Brenske det.
</taxonomicName>
A.V. Evans / JduGH64 (13 
<collectionCode id="8FE9F8E31E55FF8960B3FF47AEF8DFA7" box="[898,985,190,216]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="23" pageNumber="24">TMSA</collectionCode>
); Drakensberg, Cathedral Peak Hotel, 
<quantity id="2E00CDC31E55FF8963A6FF1CADD6DF80" box="[151,247,229,255]" metricMagnitude="3" metricUnit="m" metricValue="1.46304" pageId="23" pageNumber="24" unit="ft" value="4800.0">4,800 ft</quantity>
., 
<date id="9D4646E61E55FF896236FF1CACADDF80" box="[263,396,229,255]" pageId="23" pageNumber="24" value="1951-12-12">12.xii.1951</date>
/ E.A.C.L.E. Schelpe B.M. 1952–336 / Swarming &amp; mating on lawn. / JduGH75 (63 1Ƥ 
<collectionCode id="8FE9F8E31E55FF8963A6FEF5ADD5DE59" box="[151,244,268,294]" pageId="23" pageNumber="24">BMNH</collectionCode>
), ditto but (unsexed 65 [of these 58 are typical form with black head, thorax and scutellum, while 11 are a lighter brown, but these may be teneral individuals, i.e. not completely sclerotised] 
<collectionCode id="8FE9F8E31E55FF89661AFECBA8AEDE33" box="[1323,1423,306,332]" pageId="23" pageNumber="24">BMNH</collectionCode>
); Drakensberg, Cathedral Peak Hotel, 
<quantity id="2E00CDC31E55FF896170FEA0AF80DE0C" box="[577,673,345,371]" metricMagnitude="3" metricUnit="m" metricValue="1.46304" pageId="23" pageNumber="24" unit="ft" value="4800.0">4,800 ft</quantity>
., 
<date id="9D4646E61E55FF896180FEA0AE17DE0C" box="[689,822,345,371]" pageId="23" pageNumber="24" value="1951-12-12">12.xii.1951</date>
/ E.A.C.L.E. Schelpe B.M. 1952–336 / Swarming &amp; mating on lawn. / Coll. Mus. Tervuren [repeat locality data label] / JduGH76 (13 
<collectionCode id="8FE9F8E31E55FF89676CFE79A998DEE5" box="[1117,1209,384,410]" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="23" pageNumber="24">MRAC</collectionCode>
); Durban, illegible, 1891 / 
<taxonomicName id="2EF81BA51E55FF8963C2FE5EACFDDEBF" authority="Bl." authorityName="Bl." box="[243,476,422,448]" class="Insecta" family="Dermestidae" genus="Cisthenopholis" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="subfasciata">
<emphasis id="DB8CBC341E55FF8963C2FE5EAC8DDEBF" box="[243,428,423,448]" italics="true" pageId="23" pageNumber="24">A. subfasciata</emphasis>
Bl.
</taxonomicName>
[misidentification] / SAM-COL-A035869 / JduGH105 (13 
<collectionCode id="8FE9F8E31E55FF8967DEFE5FA86FDEBF" box="[1263,1358,422,448]" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="23" pageNumber="24">SAMC</collectionCode>
); East Griqualand, [New] Amalfi, W.J. Haygarth, Ac. No. 1981 / JduGH65 (23 
<collectionCode id="8FE9F8E31E55FF896721FE34A94ADE98" box="[1040,1131,461,487]" httpUri="http://grbio.org/cool/i1wq-4y1w" name="Durban Museum" pageId="23" pageNumber="24">DMSA</collectionCode>
); Eshowe, 
<date id="9D4646E61E55FF8967DEFE34A856DE98" box="[1263,1399,461,487]" pageId="23" pageNumber="24" value="1996-11-15">15.11.1996</date>
, T. Prince, “The bugs were found in a cane field previously infected with white grub.” / JduGH69 (43 
<collectionCode id="8FE9F8E31E55FF896671FE0DA8AEDD71" box="[1344,1423,500,526]" pageId="23" pageNumber="24">SASR</collectionCode>
); Eshowe, Sunrise Farm, 
<date id="9D4646E61E55FF89629FFDE3AF07DD4B" box="[430,550,538,564]" pageId="23" pageNumber="24" value="1996-04-12">4.12.1996</date>
, M.J. Way, Govender, Zuma, in [sugar]cane / JduGH66 (5 
<collectionCode id="8FE9F8E31E55FF8967E6FDE3A808DD4B" LSID="urn:lsid:biocol.org:col:34248" box="[1239,1321,538,564]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="23" pageNumber="24">SANC</collectionCode>
; 5 
<collectionCode id="8FE9F8E31E55FF89667BFDE3A8B6DD4B" box="[1354,1431,538,564]" pageId="23" pageNumber="24">SASR</collectionCode>
; 5 
<collectionCode id="8FE9F8E31E55FF89639CFDB8AC25DD24" box="[173,260,577,603]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="23" pageNumber="24">TMSA</collectionCode>
); Eshowe, Sunrise Farm, 
<date id="9D4646E61E55FF896101FDB8AF96DD24" box="[560,695,577,603]" pageId="23" pageNumber="24" value="1996-12-12">12.12.1996</date>
, Way &amp; Zuma / JduGH67 (13 
<collectionCode id="8FE9F8E31E55FF896719FDB8A957DD24" box="[1064,1142,577,603]" pageId="23" pageNumber="24">SASR</collectionCode>
); Eshowe, Sunrise Farm, 
<date id="9D4646E61E55FF8963A6FD91AC05DDFD" box="[151,292,616,642]" pageId="23" pageNumber="24" value="1997-03-18">18.03.1997</date>
, M.J. Way, lab reared, brown form, [date of emergence] 
<date id="9D4646E61E55FF8960D5FD91A94CDDFD" box="[996,1133,616,642]" pageId="23" pageNumber="24" value="1997-03-18">18.03.1997</date>
/ JduGH70 (1Ƥ 
<collectionCode id="8FE9F8E31E55FF89660CFD91A8AEDDFD" box="[1341,1423,616,642]" pageId="23" pageNumber="24">SASR</collectionCode>
); Ezulwini [near Mbabane], M.J. Way, No. 3 [white grub 
<typeStatus id="3643DE841E55FF89600DFD77AE57DDD7" box="[828,886,654,680]" pageId="23" pageNumber="24">type</typeStatus>
]; 97 white grub survey, lab reared / 3rd instar exuvia / JduGH68 (13 
<collectionCode id="8FE9F8E31E55FF89628CFD4CAF2EDDB0" box="[445,527,693,719]" pageId="23" pageNumber="24">SASR</collectionCode>
); Greyton [Greytown],?, illegible, /? demouoler Decelle [pencil note] / dissected male / JduGH71 (13 
<collectionCode id="8FE9F8E31E55FF896138FD25AF49DD89" LSID="urn:lsid:biocol.org:col:34988" box="[521,616,732,758]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="24">MNHN</collectionCode>
); Inchanga, K.H. Barnard, // K.H. Barnard 
<date id="9D4646E61E55FF896755FD25A9FEDD89" box="[1124,1247,732,758]" pageId="23" pageNumber="24" value="1917-11">Nov. 1917</date>
/ 
<taxonomicName id="2EF81BA51E55FF8967C7FD25AC73DC63" authority="Brenske" authorityName="Brenske" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="DB8CBC341E55FF8967C7FD25ADC3DC64" italics="true" pageId="23" pageNumber="24">Asthenopholis minor</emphasis>
Brenske
</taxonomicName>
<taxonomicName id="2EF81BA51E55FF89626BFCFBAC9CDC63" box="[346,445,770,796]" pageId="23" pageNumber="24" rank="variety" variety="det">var. det</taxonomicName>
. R.D. Pope 1951 / SAM-COL-A035866 / JduGH84 (13 
<collectionCode id="8FE9F8E31E55FF8967BEFCFBA9CDDC63" box="[1167,1260,770,796]" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="23" pageNumber="24">SAMC</collectionCode>
); Kastrol Nek [Kastrolnek Pass], 
<date id="9D4646E61E55FF896249FCD0ACCCDC3C" box="[376,493,809,835]" pageId="23" pageNumber="24" value="1922-01">Jan. 1922</date>
, A. Roberts &amp; G. van Dam / 
<taxonomicName id="2EF81BA51E55FF896077FCD0AE8FDC3D" box="[838,942,809,834]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="DB8CBC341E55FF896077FCD0AE8FDC3D" box="[838,942,809,834]" italics="true" pageId="23" pageNumber="24">A. minor</emphasis>
</taxonomicName>
det. A.V. Evans / JduGH92 (13 
<collectionCode id="8FE9F8E31E55FF896609FCD0A8AEDC3C" box="[1336,1423,809,835]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="23" pageNumber="24">TMSA</collectionCode>
); Kestell Dist.[rict], Qua–Qua N.[ational] P.[ark], Honingkloof, 
<date id="9D4646E61E55FF8960A0FCA9A934DC15" box="[913,1045,848,874]" pageId="23" pageNumber="24" value="1993-11-23">23.xi.1993</date>
, D. de Swardt / NMBH-38713 / JduGH527 (13 
<collectionCode id="8FE9F8E31E55FF896263FC8FAC8BDCEF" box="[338,426,886,912]" pageId="23" pageNumber="24">BMSA</collectionCode>
). Nagana Res. Lab. H.H. Curson, 1922, Pres. By Imp. Bur. Ent. Brit. Mus. 1923–234 / Ant. 9-jtd. [jointed] [Arrow's hand] / JduGH80 (13 
<collectionCode id="8FE9F8E31E55FF89602CFC64AE5CDCC8" box="[797,893,925,951]" pageId="23" pageNumber="24">BMNH</collectionCode>
); Nylstroom, 
<date id="9D4646E61E55FF896717FC64A992DCC8" box="[1062,1203,925,951]" pageId="23" pageNumber="24" value="1987-04-13">13.04.1987</date>
, 
<collectionCode id="8FE9F8E31E55FF8967F1FC64A832DCC8" box="[1216,1299,925,951]" httpUri="http://grbio.org/cool/73qr-ciny" name="University of Pretoria" pageId="23" pageNumber="24">UPSA</collectionCode>
: S. Gothe / JduGH96 (1Ƥ 
<collectionCode id="8FE9F8E31E55FF896274FC3DACBEDCA1" LSID="urn:lsid:biocol.org:col:34802" box="[325,415,964,990]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="23" pageNumber="24">LACM</collectionCode>
); Ostafrika, Coll. Kraatz / Moser det. / 
<taxonomicName id="2EF81BA51E55FF89605DFC3DA98BDCA2" box="[876,1194,964,989]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="subfasciatus">
<emphasis id="DB8CBC341E55FF89605DFC3DA98BDCA2" box="[876,1194,964,989]" italics="true" pageId="23" pageNumber="24">Asthenopholis subfasciatus</emphasis>
</taxonomicName>
[misidentification] / JduGH104 (13 1Ƥ 
<collectionCode id="8FE9F8E31E55FF8962A0FC13ACC0DB7B" box="[401,481,1002,1028]" pageId="23" pageNumber="24">DEIG</collectionCode>
); Oliviershoek Pass, 
<date id="9D4646E61E55FF8961DEFC13AE50DB7B" box="[751,881,1002,1028]" pageId="23" pageNumber="24" value="1989-02-12">2.12.1989</date>
, V-V: 382, [T.] van Viegen &amp; [P.] Stobbia / JduGH551 (43 
<collectionCode id="8FE9F8E31E55FF896267FBE8AC8BDB54" box="[342,426,1041,1067]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="23" pageNumber="24">TMSA</collectionCode>
) Olivier[s]hoek Pass / 
<quantity id="2E00CDC31E55FF89618CFBE8AE44DB54" box="[701,869,1041,1067]" metricMagnitude="3" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="23" pageNumber="24" unit="m" value="1900.0" valueMax="2000.0" valueMin="1800.0">1800–2000 m</quantity>
, 
<date id="9D4646E61E55FF896043FBE8AECDDB54" box="[882,1004,1041,1067]" pageId="23" pageNumber="24" value="1989-02-12">2.12.1989</date>
, T. van Viegen / JduGH63 (103 13 
<collectionCode id="8FE9F8E31E55FF8963A6FBC1ADF7DB2D" box="[151,214,1080,1106]" pageId="23" pageNumber="24">SEM</collectionCode>
<collectionCode id="8FE9F8E31E55FF8963EFFBC1AC19DB2D" box="[222,312,1080,1106]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="23" pageNumber="24">TMSA</collectionCode>
); Port 
<collectingCountry id="91EF20B61E55FF8962BAFBC1ACEFDB2D" box="[395,462,1080,1106]" name="South Africa" pageId="23" pageNumber="24">Natal</collectingCountry>
[= Durban] / 36369, Fry Coll. 1905–100, Fry Coll. 1905–100 / JduGH99 (13 
<collectionCode id="8FE9F8E31E55FF8963A6FBA7ADD4DB07" box="[151,245,1118,1144]" pageId="23" pageNumber="24">BMNH</collectionCode>
); Scottburgh [near Port Shepstone], 
<date id="9D4646E61E55FF896192FBA7AE0BDB07" box="[675,810,1118,1144]" pageId="23" pageNumber="24" value="1926-03-14">14.03.1926</date>
, R.H.R. Stevenson, Brit. Mus. 1955–734 / JduGH77 (13 
<collectionCode id="8FE9F8E31E55FF8963FDFB7CAC08DBE0" box="[204,297,1157,1183]" pageId="23" pageNumber="24">BMNH</collectionCode>
); Stutterheim, 2.01.1959, J.A. Nagle, [1 dark, 1 light spec.] / JduGH74 (23 
<collectionCode id="8FE9F8E31E55FF8967ACFB7CA9D5DBE0" box="[1181,1268,1157,1183]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="23" pageNumber="24">TMSA</collectionCode>
); Ulundi [near Nkandla], G.A.K. 
<collectingCountry id="91EF20B61E55FF89625AFB55ACF7DBB9" box="[363,470,1196,1222]" name="Marshall Islands" pageId="23" pageNumber="24">Marshall</collectingCountry>
, B.M. 
<date id="9D4646E61E55FF896114FB55AFAADBB9" box="[549,651,1196,1222]" pageId="23" pageNumber="24" value="1912" valueMax="1995">1912–95</date>
/ 31/1 [date] / JduGH83 (13 
<collectionCode id="8FE9F8E31E55FF8960D2FB55A961DBB9" box="[995,1088,1196,1222]" pageId="23" pageNumber="24">BMNH</collectionCode>
); Umzimkulu [near Kokstad], 
<date id="9D4646E61E55FF8963A6FB2BAC3FDB93" box="[151,286,1234,1260]" pageId="23" pageNumber="24" value="1979-12-28">28.12.1979</date>
, leg. A.E.B. / JduGH79 (13 
<collectionCode id="8FE9F8E31E55FF896146FB2BAFF3DB93" LSID="urn:lsid:biocol.org:col:34802" box="[631,722,1234,1260]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="23" pageNumber="24">LACM</collectionCode>
); Vlottenburg [Station, Cape Town], 
<date id="9D4646E61E55FF8967B8FB2BA820DB93" box="[1161,1281,1234,1260]" pageId="23" pageNumber="24" value="1958-06-11">6.11.1958</date>
, J.A. Nagle / JduGH73 (23 
<collectionCode id="8FE9F8E31E55FF89627EFB00AC8BDA6C" box="[335,426,1273,1299]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="23" pageNumber="24">TMSA</collectionCode>
); Wakkerstroom, 
<date id="9D4646E61E55FF8961B7FB00AE34DA6C" box="[646,789,1273,1299]" pageId="23" pageNumber="24" value="1987-01-13">13.01.1987</date>
, L. Pretorius, SE 2730 Ac / JduGH95 (1Ƥ 
<collectionCode id="8FE9F8E31E55FF896600FB00A8AEDA6C" LSID="urn:lsid:biocol.org:col:34802" box="[1329,1423,1273,1299]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="23" pageNumber="24">LACM</collectionCode>
); Wakkerstroom, 
<date id="9D4646E61E55FF896263FAD9ACE4DA45" box="[338,453,1312,1338]" pageId="23" pageNumber="24" value="1922-01">Jan. 1922</date>
, A. Roberts &amp; G. van Dam / JduGH86 (13 1Ƥ 
<collectionCode id="8FE9F8E31E55FF8960C8FAD9A972DA45" LSID="urn:lsid:biocol.org:col:34802" box="[1017,1107,1312,1338]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="23" pageNumber="24">LACM</collectionCode>
); Wakkerstroom, 
<date id="9D4646E61E55FF896612FAD9A8B9DA45" box="[1315,1432,1312,1338]" pageId="23" pageNumber="24" value="1922-01">Jan. 1922</date>
, A. Roberts &amp; G. van Dam / JduGH85 (23 
<collectionCode id="8FE9F8E31E55FF8961B8FABFAFC1DA1F" box="[649,736,1350,1376]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="23" pageNumber="24">TMSA</collectionCode>
); Wakkerstroom, 
<date id="9D4646E61E55FF89609CFABFA915DA1F" box="[941,1076,1350,1376]" pageId="23" pageNumber="24" value="1987-12-01">12.01.1987</date>
, B.L. Veenemans, SE 2730 Ac / JduGH94 (1Ƥ 
<collectionCode id="8FE9F8E31E55FF896262FA94AC8CDAF8" LSID="urn:lsid:biocol.org:col:34802" box="[339,429,1389,1415]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="23" pageNumber="24">LACM</collectionCode>
); Wakkerstroom, Martins Dam, 
<date id="9D4646E61E55FF896017FA94AE8DDAF8" box="[806,940,1389,1415]" pageId="23" pageNumber="24" value="1995-12-31">31.12.1995</date>
, R. Stals, 
<geoCoordinate id="8CCC06E11E55FF896710FA94A952DAF8" box="[1057,1139,1389,1415]" direction="south" orientation="latitude" pageId="23" pageNumber="24" precision="555" value="-27.2">27.20S</geoCoordinate>
<geoCoordinate id="8CCC06E11E55FF89674BFA94A9EFDAF8" box="[1146,1230,1389,1415]" direction="east" orientation="longitude" pageId="23" pageNumber="24" precision="555" value="30.1">30.10E</geoCoordinate>
MPU / JduGH93 (13 
<collectionCode id="8FE9F8E31E55FF8963FDFA6DAC3EDAD1" LSID="urn:lsid:biocol.org:col:34248" box="[204,287,1428,1454]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="23" pageNumber="24">SANC</collectionCode>
); White River, H.L.L.F. E. Tvl. 
<date id="9D4646E61E55FF8961A4FA6DAE2FDAD1" box="[661,782,1428,1454]" pageId="23" pageNumber="24" value="1906-12">Dec. 1906</date>
/ 
<taxonomicName id="2EF81BA51E55FF896014FA6DAEAADAD2" box="[805,907,1428,1453]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="DB8CBC341E55FF896014FA6DAEAADAD2" box="[805,907,1428,1453]" italics="true" pageId="23" pageNumber="24">A. minor</emphasis>
</taxonomicName>
det. A.V. Evans / JduGH87 (1Ƥ 
<collectionCode id="8FE9F8E31E55FF89663FFA6DA844DAD1" box="[1294,1381,1428,1454]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="23" pageNumber="24">TMSA</collectionCode>
). 
<emphasis id="DB8CBC341E55FF896649FA6AAC1EDAAB" bold="true" pageId="23" pageNumber="24">No locality data.</emphasis>
<collectingCountry id="91EF20B61E55FF896279FA43ACABDAAB" box="[328,394,1466,1492]" name="South Africa" pageId="23" pageNumber="24">Natal</collectingCountry>
/ JduGH103 (13 
<collectionCode id="8FE9F8E31E55FF896156FA43AFE2DAAB" box="[615,707,1466,1492]" pageId="23" pageNumber="24">ZMHB</collectionCode>
); no data / JduGH107 (13 
<collectionCode id="8FE9F8E31E55FF896721FA43A94CDAAB" box="[1040,1133,1466,1492]" httpUri="http://grbio.org/cool/i1wq-4y1w" name="Durban Museum" pageId="23" pageNumber="24">DMSA</collectionCode>
); Not legible, 1.01.1897, 
<collectingCountry id="91EF20B61E55FF8963A6FA18ADDEDA84" box="[151,255,1505,1531]" name="Marshall Islands" pageId="23" pageNumber="24">Marshall</collectingCountry>
Coll. 
<date id="9D4646E61E55FF896276FA18AC8EDA84" box="[327,431,1505,1531]" pageId="23" pageNumber="24" value="1912" valueMax="1995">1912–95</date>
/ JduGH106 (13 
<collectionCode id="8FE9F8E31E55FF8961B0FA18AFFFDA84" box="[641,734,1505,1531]" pageId="23" pageNumber="24">BMNH</collectionCode>
).
</paragraph>
<paragraph id="E94760261E55FF8963A6F9D6AF03D937" blockId="23.[151,1437,1582,2034]" box="[151,546,1582,1608]" pageId="23" pageNumber="24">
<taxonomicName id="2EF81BA51E55FF8963A6F9D6ACABD937" box="[151,394,1583,1608]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="DB8CBC341E55FF8963A6F9D6ACABD937" box="[151,394,1583,1608]" italics="true" pageId="23" pageNumber="24">Asthenopholis minor</emphasis>
</taxonomicName>
- black form
</paragraph>
<paragraph id="E94760261E55FF8663F7F9ACAEBCDEBF" blockId="23.[151,1437,1582,2034]" lastBlockId="24.[151,1437,152,448]" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
<emphasis id="DB8CBC341E55FF8963F7F9ACAC94D910" bold="true" box="[198,437,1621,1647]" pageId="23" pageNumber="24">Material examined</emphasis>
(Σ89, 773, 12Ƥ). 
<emphasis id="DB8CBC341E55FF8961AAF9ACAE5FD910" bold="true" box="[667,894,1621,1647]" pageId="23" pageNumber="24">
<collectingCountry id="91EF20B61E55FF8961AAF9ACAE5FD910" box="[667,894,1621,1647]" name="South Africa" pageId="23" pageNumber="24">SOUTH AFRICA</collectingCountry>
</emphasis>
, KwaZulu– 
<collectingCountry id="91EF20B61E55FF89673FF9ACA96ED910" box="[1038,1103,1621,1647]" name="South Africa" pageId="23" pageNumber="24">Natal</collectingCountry>
, Bennett Farm, 
<date id="9D4646E61E55FF896621F9ACA8B6D910" box="[1296,1431,1621,1647]" pageId="23" pageNumber="24" value="1997-03-18">18.03.1997</date>
, K., Naidoe, 
<collectionCode id="8FE9F8E31E55FF89621AF985AC57D9E9" box="[299,374,1660,1686]" pageId="23" pageNumber="24">SASR</collectionCode>
<taxonomicName id="2EF81BA51E55FF89624FF985AF08D9EA" box="[382,553,1660,1685]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis id="DB8CBC341E55FF89624FF985AF08D9EA" box="[382,553,1660,1685]" italics="true" pageId="23" pageNumber="24">Asthenopholis</emphasis>
</taxonomicName>
larval species No. 3 / JduGH126 (13 
<collectionCode id="8FE9F8E31E55FF8960CCF985A96CD9E9" box="[1021,1101,1660,1686]" pageId="23" pageNumber="24">SASR</collectionCode>
); Emoyeni, 
<date id="9D4646E61E55FF8967D1F985A87BD9E9" box="[1248,1370,1660,1686]" pageId="23" pageNumber="24" value="1994-10">Oct. 1994</date>
, S.ex 4300 [4 of series only probably from this locality] / JduGH127 (63 1 
<collectionCode id="8FE9F8E31E55FF8960F6F95BA97FD9C3" box="[967,1118,1698,1724]" pageId="23" pageNumber="24">SEM SASR</collectionCode>
); Emoyeni, 
<date id="9D4646E61E55FF8967DCF95BA853D9C3" box="[1261,1394,1698,1724]" pageId="23" pageNumber="24" value="1987-11-26">26.11.1987</date>
, A. Carnegie, Ac.S.ex 833 / JduGH124 (13 
<collectionCode id="8FE9F8E31E55FF896192F930AE22D99C" LSID="urn:lsid:biocol.org:col:34802" box="[675,771,1737,1763]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="23" pageNumber="24">LACM</collectionCode>
); Emoyeni, 
<date id="9D4646E61E55FF8960AFF930A910D99C" box="[926,1073,1737,1763]" pageId="23" pageNumber="24" value="1987-11-26">26.11.1987</date>
, A. Carnegie, Ac.S.ex 833 / JduGH125 (43 
<collectionCode id="8FE9F8E31E55FF89626FF909AC90D875" box="[350,433,1776,1802]" pageId="23" pageNumber="24">SASR</collectionCode>
); Emoyeni, 
<date id="9D4646E61E55FF896179F909AFE6D875" box="[584,711,1776,1802]" pageId="23" pageNumber="24" value="1996-07-11">7.11.1996</date>
, Way &amp; Govender, field collected adults, in [sugar]cane / JduGH129 (103 
<collectionCode id="8FE9F8E31E55FF896250F8EFAC8FD84F" box="[353,430,1814,1840]" pageId="23" pageNumber="24">SASR</collectionCode>
), (83 
<collectionCode id="8FE9F8E31E55FF8962C9F8EFAF6ED84F" box="[504,591,1814,1840]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="23" pageNumber="24">TMSA</collectionCode>
); Emoyeni, 
<date id="9D4646E61E55FF8961EAF8EFAE40D84F" box="[731,865,1814,1840]" pageId="23" pageNumber="24" value="1996-04-17">17.04.1996</date>
, M.J. Way, laboratory reared larvae No. 3 [white grub morphospecies 3], emergence dates vary / JduGH130 (103 1 
<collectionCode id="8FE9F8E31E55FF8960F3F8C4A924D828" box="[962,1029,1853,1879]" pageId="23" pageNumber="24">SEM</collectionCode>
5Ƥ 
<collectionCode id="8FE9F8E31E55FF89670DF8C4A9ACD828" box="[1084,1165,1853,1879]" pageId="23" pageNumber="24">SASR</collectionCode>
); Emoyeni, M.J. Way, larvae collected 8.05.1996 emerged dates vary / JduGH131 (63 2Ƥ 
<collectionCode id="8FE9F8E31E55FF896088F89DA926D801" box="[953,1031,1892,1918]" pageId="23" pageNumber="24">SASR</collectionCode>
); Emoyeni, M.J. Way / JduGH132 (43 2Ƥ 
<collectionCode id="8FE9F8E31E55FF8963CAF873AC6BD8DB" box="[251,330,1930,1956]" pageId="23" pageNumber="24">SASR</collectionCode>
); Emoyeni, 
<quantity id="2E00CDC31E55FF8962E9F873AF05D8DB" box="[472,548,1930,1956]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="23" pageNumber="24" unit="km" value="15.0">15 km</quantity>
WSW of Mtunzini, 
<date id="9D4646E61E55FF896029F873AEBED8DB" box="[792,927,1930,1956]" pageId="23" pageNumber="24" value="1987-11-26">26.11.1987</date>
, A. Carnegie, AcSex 833 / JduGH123 (13 1Ƥ 
<collectionCode id="8FE9F8E31E55FF8963F2F848AC37D8B4" LSID="urn:lsid:biocol.org:col:34248" box="[195,278,1969,1995]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="23" pageNumber="24">SANC</collectionCode>
); Malvern, illegible, 
<date id="9D4646E61E55FF89613DF848AFA7D8B4" box="[524,646,1969,1995]" pageId="23" pageNumber="24" value="1899-12">Dec. 1899</date>
/ JduGH494 (13 
<collectionCode id="8FE9F8E31E55FF89606BF848AE98D8B4" LSID="urn:lsid:biocol.org:col:34988" box="[858,953,1969,1995]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="24">MNHN</collectionCode>
); Malvern, G.A.K. 
<collectingCountry id="91EF20B61E55FF8967A8F848A825D8B4" box="[1177,1284,1969,1995]" name="Marshall Islands" pageId="23" pageNumber="24">Marshall</collectingCountry>
, Comp. Typ. 
<taxonomicName id="2EF81BA51E55FF8963A6F821ADC1D88D" box="[151,224,2008,2034]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="38" phylum="Arthropoda" rank="species" species="minor">Minor</taxonomicName>
[Decelle label] / JduGH493 (13 
<collectionCode id="8FE9F8E31E55FF896155F821AFE1D88D" box="[612,704,2008,2034]" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="23" pageNumber="24">MRAC</collectionCode>
); Malvern, G.A.K. 
<collectingCountry id="91EF20B61E55FF8960ACF821A929D88D" box="[925,1032,2008,2034]" name="Marshall Islands" pageId="23" pageNumber="24">Marshall</collectingCountry>
, 
<date id="9D4646E61E55FF896722F821A9A8D88D" box="[1043,1161,2008,2034]" pageId="23" pageNumber="24" value="1897-10">
Oct. 
<quantity id="2E00CDC31E55FF89677DF821A9A8D88D" box="[1100,1161,2008,2034]" metricMagnitude="5" metricUnit="m" metricValue="5.7841895999999995" pageId="23" pageNumber="24" unit="ft" value="1897700.0">1897</quantity>
</date>
, 700 ft. / 
<taxonomicName id="2EF81BA51E55FF8667C7F821AC68DFCD" authority="Brenske" authorityName="Brenske" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" lastPageId="24" lastPageNumber="25" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="DB8CBC341E55FF8667C7F821ADFFDFCE" italics="true" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">Asthenopholis minor</emphasis>
Brenske
</taxonomicName>
, comp. with 
<typeStatus id="3643DE841E5AFF8662D2FF61AF37DFCD" box="[483,534,152,178]" pageId="24" pageNumber="25">type</typeStatus>
G.J.A. [handwritten by Arrow, and refer to 
<bibRefCitation id="8D691DD71E5AFF866710FF61A9D4DFCD" author="Arrow" box="[1057,1269,152,178]" pageId="24" pageNumber="43" refString="Arrow, G. J. (1917) Some systematic notes on melolonthine Coleoptera. The Annals and Magazine of Natural History, series 8, volume 19, 59 - 65." type="journal article" year="1917">Arrow (1917: 60)</bibRefCitation>
] / 1901–179 / JduGH489 (23 
<collectionCode id="8FE9F8E31E5AFF866251FF47ACE3DFA7" box="[352,450,190,216]" pageId="24" pageNumber="25">BMNH</collectionCode>
); Malvern, G.A.K. 
<collectingCountry id="91EF20B61E5AFF866185FF47AE06DFA7" box="[692,807,190,216]" name="Marshall Islands" pageId="24" pageNumber="25">Marshall</collectingCountry>
, 
<date id="9D4646E61E5AFF866005FF47AE9EDFA7" box="[820,959,190,216]" pageId="24" pageNumber="25" value="1897-09">Sept. 1897</date>
, 1901–179 / JduGH490 (13 
<collectionCode id="8FE9F8E31E5AFF86661CFF47A8AEDFA7" box="[1325,1423,190,216]" pageId="24" pageNumber="25">BMNH</collectionCode>
); Malvern, G.A.K. 
<collectingCountry id="91EF20B61E5AFF86625DFF1CACFDDF80" box="[364,476,229,255]" name="Marshall Islands" pageId="24" pageNumber="25">Marshall</collectingCountry>
, 
<date id="9D4646E61E5AFF8662D8FF1CAF44DF80" box="[489,613,229,255]" pageId="24" pageNumber="25" value="1897-10">Oct. 1897</date>
, 700– 
<quantity id="2E00CDC31E5AFF866181FF1CAFDEDF80" box="[688,767,229,255]" metricMagnitude="2" metricUnit="m" metricValue="2.4384" pageId="24" pageNumber="25" unit="ft" value="800.0">800 ft</quantity>
., 
<date id="9D4646E61E5AFF866020FF1CAE5ADF80" box="[785,891,229,255]" pageId="24" pageNumber="25" value="1912" valueMax="1995">1912–95</date>
/ JduGH491 (13 
<collectionCode id="8FE9F8E31E5AFF86676BFF1CA99BDF80" box="[1114,1210,229,255]" pageId="24" pageNumber="25">BMNH</collectionCode>
); Malvern, G.A.K. 
<collectingCountry id="91EF20B61E5AFF8663A6FEF5AC28DE59" box="[151,265,268,294]" name="Marshall Islands" pageId="24" pageNumber="25">Marshall</collectingCountry>
, 
<taxonomicName id="2EF81BA51E5AFF866227FEF5AF68DE59" authority="Br." authorityName="Br." box="[278,585,268,294]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="DB8CBC341E5AFF866227FEF5AF3BDE5A" box="[278,538,268,293]" italics="true" pageId="24" pageNumber="25">Asthenopholis minor</emphasis>
Br.
</taxonomicName>
/ male genitalia [dissected by Péringuey] / SAMC-COL-A035871 / JduGH492 (13 
<collectionCode id="8FE9F8E31E5AFF866266FECBAC91DE33" box="[343,432,306,332]" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="24" pageNumber="25">SAMC</collectionCode>
); Mtunzini, 
<date id="9D4646E61E5AFF866172FECBAF9DDE33" box="[579,700,306,332]" pageId="24" pageNumber="25" value="1983-02-11">2.11.1983</date>
, R. Heathcote, antennae dimorphic in this series / JduGH122 (173 13 1Ƥ 
<collectionCode id="8FE9F8E31E5AFF866203FEA0ACA7DE0C" LSID="urn:lsid:biocol.org:col:34248" box="[306,390,345,371]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="24" pageNumber="25">SANC</collectionCode>
); Port 
<collectingCountry id="91EF20B61E5AFF8662E1FEA0AF2EDE0C" box="[464,527,345,371]" name="South Africa" pageId="24" pageNumber="25">Natal</collectingCountry>
[= Durban], D. Schulze / JduGH495 (13 
<collectionCode id="8FE9F8E31E5AFF8660C8FEA0A972DE0C" box="[1017,1107,345,371]" pageId="24" pageNumber="25">ZMHB</collectionCode>
). 
<emphasis id="DB8CBC341E5AFF866754FEA1A810DE0D" bold="true" box="[1125,1329,344,370]" pageId="24" pageNumber="25">No locality data.</emphasis>
<collectionCode id="8FE9F8E31E5AFF866609FEA0A8B9DE0C" box="[1336,1432,345,371]" pageId="24" pageNumber="25">SASEX</collectionCode>
, “1996 lab reared” / Dc SEX 828 [this pair are very large for 
<taxonomicName id="2EF81BA51E5AFF866057FE79AEEDDEE6" box="[870,972,384,409]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="DB8CBC341E5AFF866057FE79AEEDDEE6" box="[870,972,384,409]" italics="true" pageId="24" pageNumber="25">A. minor</emphasis>
</taxonomicName>
, but perhaps this is as a result of being lab reared rather than field emerged] / JduGH497 (13 1Ƥ 
<collectionCode id="8FE9F8E31E5AFF866073FE5FAEB0DEBF" box="[834,913,422,448]" pageId="24" pageNumber="25">SASR</collectionCode>
).
</paragraph>
<paragraph id="E94760261E5AFF8663A6FE0DAF90DD71" blockId="24.[151,1437,500,951]" box="[151,689,500,526]" pageId="24" pageNumber="25">
<taxonomicName id="2EF81BA51E5AFF8663A6FE0DACABDD72" box="[151,394,500,525]" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="DB8CBC341E5AFF8663A6FE0DACABDD72" box="[151,394,500,525]" italics="true" pageId="24" pageNumber="25">Asthenopholis minor</emphasis>
</taxonomicName>
- brown and glossy form
</paragraph>
<paragraph id="E94760261E5AFF8663F7FDE3A9E2DCC8" blockId="24.[151,1437,500,951]" pageId="24" pageNumber="25">
<emphasis id="DB8CBC341E5AFF8663F7FDE3AC9ADD4B" bold="true" box="[198,443,538,564]" pageId="24" pageNumber="25">Material examined</emphasis>
(Σ52, 473, 5Ƥ). 
<collectingCountry id="91EF20B61E5AFF866196FDE3AEAEDD4B" box="[679,911,538,564]" name="South Africa" pageId="24" pageNumber="25">
<emphasis id="DB8CBC341E5AFF866196FDE3AEAEDD4B" bold="true" box="[679,911,538,564]" pageId="24" pageNumber="25">SOUTH AFRICA</emphasis>
</collectingCountry>
, Kopees Kraal, Lydenburg, 
<date id="9D4646E61E5AFF8667C8FDE3A850DD4B" box="[1273,1393,538,564]" pageId="24" pageNumber="25" value="1996-05-12">5.12.1996</date>
, S. Endrődy-Younga, UV light, E-Y: 3252 / JduGH456 (153 1 
<collectionCode id="8FE9F8E31E5AFF866053FDB8AE84DD24" box="[866,933,577,603]" pageId="24" pageNumber="25">SEM</collectionCode>
2Ƥ 
<collectionCode id="8FE9F8E31E5AFF8660EDFDB8A914DD24" box="[988,1077,577,603]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="24" pageNumber="25">TMSA</collectionCode>
); Nqutu, 1960, A.H. Newton, C.I.E. Coll. 17034 / JduGH461 (73 1 
<collectionCode id="8FE9F8E31E5AFF86615EFD91AE3EDDFD" box="[623,799,616,642]" pageId="24" pageNumber="25">SEM BMNH</collectionCode>
); Pietersburg University, 
<date id="9D4646E61E5AFF866753FD91A9C3DDFD" box="[1122,1250,616,642]" pageId="24" pageNumber="25" value="1976-07-11">7.11.1976</date>
, P.E. Reavell / JduGH458 (23 
<collectionCode id="8FE9F8E31E5AFF866265FD77AC8ADDD7" box="[340,427,654,680]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="24" pageNumber="25">TMSA</collectionCode>
); Vryheid, 
<date id="9D4646E61E5AFF86611CFD77AF93DDD7" box="[557,690,654,680]" pageId="24" pageNumber="25" value="1995-12-11">12.11.1995</date>
, M. Krüger, swarming over lawn at 12: 
<specimenCount id="FFFEABAF1E5AFF8667B3FD77A9E1DDD7" box="[1154,1216,654,680]" pageId="24" pageNumber="25" type="generic">30 p.</specimenCount>
m. [important data as it indicates diurnal activity]; JduGH455 (83 1Ƥ 
<collectionCode id="8FE9F8E31E5AFF86601CFD4CAEA8DDB0" box="[813,905,693,719]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="24" pageNumber="25">TMSA</collectionCode>
), (53 
<collectionCode id="8FE9F8E31E5AFF8660EBFD4CA91BDDB0" box="[986,1082,693,719]" pageId="24" pageNumber="25">MKSA</collectionCode>
); Zukobene, 
<collectingCountry id="91EF20B61E5AFF8667D1FD4CA809DDB0" box="[1248,1320,693,719]" name="South Africa" pageId="24" pageNumber="25">Natal</collectingCountry>
, [19]03 / JduGH459 (13 
<collectionCode id="8FE9F8E31E5AFF86626AFD25AC95DD89" box="[347,436,732,758]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="24" pageNumber="25">TMSA</collectionCode>
). 
<emphasis id="DB8CBC341E5AFF8662F9FD22AF5FDD8A" bold="true" box="[456,638,731,757]" pageId="24" pageNumber="25">
<collectingCountry id="91EF20B61E5AFF8662F9FD22AF5FDD8A" box="[456,638,731,757]" name="Swaziland" pageId="24" pageNumber="25">SWAZILAND</collectingCountry>
</emphasis>
. Mhlume, 
<date id="9D4646E61E5AFF866034FD25AEAEDD89" box="[773,911,732,758]" pageId="24" pageNumber="25" value="1986-11-25">25.11.1986</date>
, A. Carnegie, 
<collectionCode id="8FE9F8E31E5AFF866770FD25A981DD89" box="[1089,1184,732,758]" pageId="24" pageNumber="25">SASEX</collectionCode>
828 / 
<taxonomicName id="2EF81BA51E5AFF8667C1FD25ACCFDC63" authority="Det. A.V. Evans, 1987" authorityName="Det. A.V. Evans" authorityYear="1987" class="Insecta" family="Melolonthidae" genus="Asthenopholis" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="DB8CBC341E5AFF8667C1FD25ADFCDC64" italics="true" pageId="24" pageNumber="25">Asthenopholis minor</emphasis>
Det. A.V. Evans, 1987
</taxonomicName>
/ JduGH463 (13 1Ƥ 
<collectionCode id="8FE9F8E31E5AFF8661C1FCFBAE62DC63" LSID="urn:lsid:biocol.org:col:34248" box="[752,835,770,796]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="24" pageNumber="25">SANC</collectionCode>
); Mhlume, 
<date id="9D4646E61E5AFF8660FAFCFBA970DC63" box="[971,1105,770,796]" pageId="24" pageNumber="25" value="1986-11-25">25.11.1986</date>
, A. Carnegie, 
<collectionCode id="8FE9F8E31E5AFF8667CAFCFBA879DC63" box="[1275,1368,770,796]" pageId="24" pageNumber="25">SASEX</collectionCode>
828 / JduGH462 (23 1 
<collectionCode id="8FE9F8E31E5AFF866258FCD0AF2BDC3C" box="[361,522,809,835]" pageId="24" pageNumber="25">SEM SASR</collectionCode>
); Mhlume, S. Date, 
<date id="9D4646E61E5AFF866039FCD0AEACDC3C" box="[776,909,809,835]" pageId="24" pageNumber="25" value="1964-11">Nov. 1964</date>
, 
<collectionCode id="8FE9F8E31E5AFF8660AAFCD0AEDDDC3C" box="[923,1020,809,835]" pageId="24" pageNumber="25">SASEX</collectionCode>
41 / JduGH464 (13 1Ƥ 
<collectionCode id="8FE9F8E31E5AFF866606FCD0A8AEDC3C" LSID="urn:lsid:biocol.org:col:34248" box="[1335,1423,809,835]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="24" pageNumber="25">SANC</collectionCode>
); Mhlume, S. Date, 
<collectionCode id="8FE9F8E31E5AFF86625EFCA9ACEDDC15" box="[367,460,848,874]" pageId="24" pageNumber="25">SASEX</collectionCode>
41 / JduGH465 (23 
<collectionCode id="8FE9F8E31E5AFF8661F4FCA9AE32DC15" box="[709,787,848,874]" pageId="24" pageNumber="25">SASR</collectionCode>
); 
<collectingCountry id="91EF20B61E5AFF866019FCA9AE84DC15" box="[808,933,848,874]" name="Swaziland" pageId="24" pageNumber="25">Swaziland</collectingCountry>
, 6.06.1996– 
<date id="9D4646E61E5AFF866707FCA9A999DC15" box="[1078,1208,848,874]" pageId="24" pageNumber="25" value="1996-11-15">15.11.1996</date>
[dates are of larval collection to adult emergence], 
<collectionCode id="8FE9F8E31E5AFF86612CFC8FAF5DDCEF" box="[541,636,886,912]" pageId="24" pageNumber="25">SASEX</collectionCode>
/ JduGH496 (13 
<collectionCode id="8FE9F8E31E5AFF86606AFC8FAE8DDCEF" box="[859,940,886,912]" pageId="24" pageNumber="25">SASR</collectionCode>
); Tabankulu (Field B), 
<date id="9D4646E61E5AFF8667FDFC8FA876DCEF" box="[1228,1367,886,912]" pageId="24" pageNumber="25" value="2006-11-23">23.11.2006</date>
, D.E. Conlong, light trap near [sugar]cane, white grub adult, SEX 1853 / JduGH554 (23 
<collectionCode id="8FE9F8E31E5AFF866759FC64A996DCC8" box="[1128,1207,925,951]" pageId="24" pageNumber="25">SASR</collectionCode>
).
</paragraph>
</subSubSection>
</treatment>
</document>