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<document id="3334265D51C4FD6D57A5174F6B745F92" ID="10.11646/zootaxa.4150.2.5" ID-DOI="10.11646/zootaxa.4150.2.5" ID-GBIF-Dataset="4fb51315-d658-4ef6-b8d1-7bb64ca24250" ID-ISSN="1175-5326" ID-Zenodo-Dep="266388" ID-ZooBank="BBD78FA7-BC15-4120-BD4C-0B215257C351" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1470897312730" checkinUser="plazi" docAuthor="Bogorodsky, Sergey V., Suzuki, Toshiyuki &amp; Mal, Ahmad O." docDate="2016" docId="03E487E3FFF1A61E1B8974F7E8A4F8D5" docLanguage="en" docName="zootaxa.4150.2.5.pdf" docOrigin="Zootaxa 4150 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trimma fishelsoni Goren 1985" docType="treatment" docVersion="8" lastPageNumber="181" masterDocId="FFDDFF9BFFFBA6131B1E7566EB5AFFCA" masterDocTitle="Description of a new species of Priolepis (Perciformes: Gobiidae) from the Red Sea, a new record of Priolepis compita, and a distributional range extension of Trimma fishelsoni" masterLastPageNumber="184" masterPageNumber="168" pageNumber="178" updateTime="1698381815852" updateUser="plazi">
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<mods:title id="EB4E60BF119AE7513749F55FC1C0D0EE">Description of a new species of Priolepis (Perciformes: Gobiidae) from the Red Sea, a new record of Priolepis compita, and a distributional range extension of Trimma fishelsoni</mods:title>
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<mods:namePart id="741546B02CCBC521A0D1DC810523B0EC">Bogorodsky, Sergey V.</mods:namePart>
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<mods:namePart id="B5D0B3AA042980C5E84C3AA54428A413">Suzuki, Toshiyuki</mods:namePart>
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<mods:namePart id="D7206F4682DEAA67CAC384AC8472BFEC">Mal, Ahmad O.</mods:namePart>
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<treatment id="03E487E3FFF1A61E1B8974F7E8A4F8D5" ID-DOI="http://doi.org/10.5281/zenodo.5694119" ID-GBIF-Taxon="124329209" ID-Zenodo-Dep="5694119" LSID="urn:lsid:plazi:treatment:03E487E3FFF1A61E1B8974F7E8A4F8D5" httpUri="http://treatment.plazi.org/id/03E487E3FFF1A61E1B8974F7E8A4F8D5" lastPageId="13" lastPageNumber="181" pageId="10" pageNumber="178">
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<taxonomicName id="4C4D4D76FFF1A6191B8974F7E94EFE61" ID-CoL="58V5J" authority="Goren, 1985" authorityName="Goren" authorityYear="1985" box="[151,532,401,427]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="178" phylum="Chordata" rank="species" species="fishelsoni">
<emphasis id="B939EAE7FFF1A6191B8974F7E94EFE61" bold="true" box="[151,532,401,427]" pageId="10" pageNumber="178">
<emphasis id="B939EAE7FFF1A6191B8974F7EA2EFE61" bold="true" box="[151,372,401,427]" italics="true" pageId="10" pageNumber="178">Trimma fishelsoni</emphasis>
<bibRefCitation id="EFDC4B04FFF1A6191A6574F7E94EFE61" author="Goren" box="[379,532,401,427]" pageId="10" pageNumber="178" refString="Goren, M. (1985) Trimma fishelsoni, a new gobiid fish from the Gulf of Elat, northern Red Sea. Israel Journal of Zoology, 33 (1 / 2), 63 - 67." type="journal article" year="1985">Goren, 1985</bibRefCitation>
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<paragraph id="8BF236F5FFF1A6191B8974D2EAF9FE07" blockId="10.[151,532,401,496]" box="[151,419,436,461]" pageId="10" pageNumber="178">Fishelsons Pygmygoby</paragraph>
</subSubSection>
<subSubSection id="C357657EFFF1A6191B8974B1EAF7FE3A" box="[151,429,471,496]" pageId="10" pageNumber="178" type="description">
<paragraph id="8BF236F5FFF1A6191B8974B1EAF7FE3A" blockId="10.[151,532,401,496]" box="[151,429,471,496]" pageId="10" pageNumber="178">
<figureCitation id="13762A70FFF1A6191B8974B1EA58FE3A" box="[151,258,471,496]" captionStart="FIGURE 4" captionStartId="11.[151,250,1954,1976]" captionTargetBox="[151,1436,752,1933]" captionTargetId="figure@11.[151,1436,739,1933]" captionTargetPageId="11" captionText="FIGURE 4. Trimma fishelsoni. A: SMF 35735, fresh specimen, 20.0 mm SL, female, Al Lith, Saudi Arabia, Red Sea; B: SMF 35737, preserved specimen, 21.6 mm SL, male, Al Lith, Saudi Arabia, Red Sea. Photo by S. V. Bogorodsky (A) and T. Suzuki (B)." httpUri="https://zenodo.org/record/266392/files/figure.png" pageId="10" pageNumber="178">Figures 4</figureCitation>
,
<figureCitation id="13762A70FFF1A6191A1074B1EA44FE3A" box="[270,286,471,496]" captionStart="FIGURE 5" captionStartId="12.[151,251,1939,1961]" captionTargetBox="[281,1337,196,1913]" captionTargetId="figure@12.[254,1337,193,1915]" captionTargetPageId="12" captionText="FIGURE 5. Trimma fishelsoni, SMF 35737, cephalic sensory papillae of head. Dorsal (top), lateral (middle) and ventral (bottom) views of head. Dots represent the sensory papillae; AN = anterior nares, PN = posterior nares; arrows show position of lower end of gill opening. Photo and drawing by T. Suzuki." httpUri="https://zenodo.org/record/266393/files/figure.png" pageId="10" pageNumber="178">5</figureCitation>
,
<figureCitation id="13762A70FFF1A6191A3474B1EA62FE3A" box="[298,312,471,496]" captionStart="FIGURE 6" captionStartId="13.[151,250,1557,1579]" captionTargetBox="[150,1435,193,1535]" captionTargetId="figure@13.[150,1436,193,1536]" captionTargetPageId="13" captionText="FIGURE 6. Underwater photographs of alive individuals. A: Trimma fishelsoni, Dahab, Gulf of Aqaba, Red Sea; B: Trimma taylori, Al Lith, Saudi Arabia, Red Sea. Photos by S. V. Bogorodsky." httpUri="https://zenodo.org/record/266394/files/figure.png" pageId="10" pageNumber="178">6</figureCitation>
A,
<tableCitation id="C6CF034EFFF1A6191A4474B1EAF7FE3A" box="[346,429,471,496]" captionStart="TABLE 1" captionStartId="9.[151,239,152,175]" captionTargetBox="[159,1387,237,1519]" captionTargetPageId="9" captionText="TABLE 1. Measurements expressed as percentage of standard length and counts for Priolepis melanops n. sp. and Trimma fishselsoni." httpUri="http://table.plazi.org/id/DF32667DFFF2A61A1B8975FEEA32FF1A" pageId="10" pageNumber="178" tableUuid="DF32667DFFF2A61A1B8975FEEA32FF1A">Table 1</tableCitation>
</paragraph>
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<taxonomicName id="4C4D4D76FFF1A6191B897747EAA1FDFC" ID-CoL="58V5J" authority="Goren, 1985: 64" authorityName="Goren" authorityPageNumber="64" authorityYear="1985" box="[151,507,544,566]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="178" phylum="Chordata" rank="species" species="fishelsoni">
<emphasis id="B939EAE7FFF1A6191B897747EA17FDFC" box="[151,333,544,566]" italics="true" pageId="10" pageNumber="178">Trimma fishelsoni</emphasis>
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<bibRefCitation id="EFDC4B04FFF1A6191A4D7746EA8FFDFC" author="Goren" box="[339,469,544,566]" pageId="10" pageNumber="178" refString="Goren, M. (1985) Trimma fishelsoni, a new gobiid fish from the Gulf of Elat, northern Red Sea. Israel Journal of Zoology, 33 (1 / 2), 63 - 67." type="journal article" year="1985">Goren, 1985</bibRefCitation>
: 64
</treatmentCitation>
</taxonomicName>
,
<figureCitation id="13762A70FFF1A619191B7746E93EFDFC" box="[517,612,544,566]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3. A" captionStartId-0="4.[151,250,1942,1964]" captionStartId-1="5.[151,250,1939,1961]" captionStartId-2="8.[151,250,1955,1977]" captionTargetBox-0="[151,1436,896,1913]" captionTargetBox-1="[276,1303,193,1860]" captionTargetBox-2="[151,1436,733,1932]" captionTargetId-0="figure@4.[151,1436,895,1921]" captionTargetId-1="figure@5.[264,1303,193,1894]" captionTargetId-2="figure@8.[151,1436,733,1934]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="8" captionText-0="FIGURE 1. Priolepis melanops, n. sp., SMF 35728, holotype, female, 24.0 mm SL, Al Lith, Saudi Arabia, Red Sea A: freshly collected; B: preserved; C: ventral view of region between pelvic and anal fins; red line indicates basal membrane, yellow lines show fourth and fifth pelvic-fin rays. Photos by S. V. Bogorodsky (A) and T. Suzuki (B, C)." captionText-1="FIGURE 2. Priolepis melanops n. sp., SMF 35728, holotype, cephalic sensory papillae of head. Dorsal (top), lateral (middle) and ventral (bottom) views of head. Dots represent the sensory papillae; AN = anterior nares, PN = posterior nares; arrow shows position of lower end of gill opening. Photo and drawing by T. Suzuki." captionText-2="FIGURE 3. A: Priolepis compita, live individual, Sharm el Moya, at entrance of the Gulf of Aqaba, Red Sea; B: Priolepis semidoliata, fresh specimen, BPBM 41242, 21.0 mm SL, Dahab, Gulf of Aqaba, Red Sea. Photos by S. V. Bogorodsky." httpUri-0="https://zenodo.org/record/266389/files/figure.png" httpUri-1="https://zenodo.org/record/266390/files/figure.png" httpUri-2="https://zenodo.org/record/266391/files/figure.png" pageId="10" pageNumber="178">Figs. 13</figureCitation>
(&quot;University Reef&quot;,
<collectingRegion id="4989F817FFF1A619182B7746E831FDFC" box="[821,875,544,566]" country="Egypt" name="Shamal Sina'" pageId="10" pageNumber="178">Sinai</collectingRegion>
,
<collectingCountry id="F35A7665FFF1A619186B7747E8EEFDFC" box="[885,948,545,566]" name="Egypt" pageId="10" pageNumber="178">Egypt</collectingCountry>
,
<location id="8E92602EFFF1A61918A17746EF0EFDFC" LSID="urn:lsid:plazi:treatment:03E487E3FFF1A61E1B8974F7E8A4F8D5:8E92602EFFF1A61918A17746EF0EFDFC" box="[959,1108,544,566]" country="Egypt" name="Gulf of Aqaba" pageId="10" pageNumber="178" stateProvince="Sinai">Gulf of Aqaba</location>
,
<collectingRegion id="4989F817FFF1A6191F407746EFE9FDFC" box="[1118,1203,544,566]" country="Egypt" name="Al Bahr al Ahmar" pageId="10" pageNumber="178">Red Sea</collectingRegion>
)
</materialsCitation>
-
<materialsCitation id="3B253CA8FFF1A6191FD17746E802FDB9" ID-GBIF-Occurrence="1291547734" country="Egypt" location="Gulf of Aqaba" pageId="10" pageNumber="178" specimenCount="1" stateProvince="Red Sea">
<treatmentCitation id="0AEC10E4FFF1A6191FD17746EBBAFD9E" author="Goren" page="66" pageId="10" pageNumber="178" year="1994">
<bibRefCitation id="EFDC4B04FFF1A6191FD17746EEC3FDFC" author="Goren" box="[1231,1433,544,566]" pageId="10" pageNumber="178" refString="Goren, M. &amp; Dor, M. (1994) An Updated Checklist of the Fishes of the Red Sea; CLOFRES II. Jerusalem; Israel Academy of Sciences and Humanities, XII + 120 pp." type="book" year="1994">Goren &amp; Dor, 1994</bibRefCitation>
: 66
</treatmentCitation>
(
<collectingRegion id="4989F817FFF1A6191BF07758EA1CFD9E" box="[238,326,574,596]" country="Egypt" name="Al Bahr al Ahmar" pageId="10" pageNumber="178">Red Sea</collectingRegion>
, checklist);
<treatmentCitation id="0AEC10E4FFF1A6191ADF7758E9ECFD9E" author="Winterbottom" box="[449,694,574,596]" page="96" pageId="10" pageNumber="178" year="1995">
<bibRefCitation id="EFDC4B04FFF1A6191ADF7758E9CBFD9E" author="Winterbottom" box="[449,657,574,596]" pageId="10" pageNumber="178" refString="Winterbottom, R. (1995) Red Sea gobiid fishes of the genus Trimma, with the description of two new species. Revue franzaise d'Aquariologie Herpйtologie, 22 (3 / 4), 93 - 98." type="journal article" year="1995">Winterbottom, 1995</bibRefCitation>
: 96
</treatmentCitation>
(
<location id="8E92602EFFF1A61919DD7758E802FD9E" LSID="urn:lsid:plazi:treatment:03E487E3FFF1A61E1B8974F7E8A4F8D5:8E92602EFFF1A61919DD7758E802FD9E" box="[707,856,574,596]" country="Egypt" name="Gulf of Aqaba" pageId="10" pageNumber="178" stateProvince="Red Sea">Gulf of Aqaba</location>
to southern
<collectingCountry id="F35A7665FFF1A61918C97759EF49FD9E" box="[983,1043,575,596]" name="Egypt" pageId="10" pageNumber="178">Egypt</collectingCountry>
at Ras Banas);
<treatmentCitation id="0AEC10E4FFF1A6191FAB7758EA7AFDB9" author="Golani" page="49" pageId="10" pageNumber="178" year="2010">
<bibRefCitation id="EFDC4B04FFF1A6191FAB7758EBA4FDB9" author="Golani" pageId="10" pageNumber="178" refString="Golani, D. &amp; Bogorodsky, S. V. (2010) The fishes of the Red Sea - reappraisal and updated checklist. Zootaxa, 2463, 1 - 135." type="journal article" year="2010">Golani &amp; Bogorodsky, 2010</bibRefCitation>
: 49
</treatmentCitation>
(
<collectingRegion id="4989F817FFF1A6191A33773BEADFFDB9" box="[301,389,605,627]" country="Egypt" name="Al Bahr al Ahmar" pageId="10" pageNumber="178">Red Sea</collectingRegion>
, checklist); Hoese
<emphasis id="B939EAE7FFF1A619195A7738E92BFDB9" box="[580,625,605,627]" italics="true" pageId="10" pageNumber="178">et al</emphasis>
., 2015: 538 (
<collectingRegion id="4989F817FFF1A61919E9773BE80AFDB9" box="[759,848,605,627]" country="Egypt" name="Al Bahr al Ahmar" pageId="10" pageNumber="178">Red Sea</collectingRegion>
)
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C357657EFFF1A6191B8977F9E89AFCEF" pageId="10" pageNumber="178" type="materials_examined">
<paragraph id="8BF236F5FFF1A6191B8977F9E89AFCEF" blockId="10.[151,1437,671,2029]" pageId="10" pageNumber="178">
<materialsCitation id="3B253CA8FFF1A6191B8977F9E974FCCA" ID-GBIF-Occurrence="1291547731" collectingDate="2014-11-17" collectionCode="SMF" collectorName="S. V. Bogorodsky &amp; T. J. Alpermann" country="Saudi Arabia" elevation="0" location="Material" municipality="Al Lith" pageId="10" pageNumber="178" specimenCount="1">
<emphasis id="B939EAE7FFF1A6191B8977F9EADBFD72" bold="true" box="[151,385,671,696]" pageId="10" pageNumber="178">
<location id="8E92602EFFF1A6191B8977F9EBA5FD72" LSID="urn:lsid:plazi:treatment:03E487E3FFF1A61E1B8974F7E8A4F8D5:8E92602EFFF1A6191B8977F9EBA5FD72" box="[151,255,671,696]" country="Saudi Arabia" municipality="Al Lith" name="Material" pageId="10" pageNumber="178">Material</location>
examined.
</emphasis>
<collectionCode id="ED5CAE30FFF1A6191A9677C6EA9AFD72" box="[392,448,672,696]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="10" pageNumber="178">SMF</collectionCode>
35735 (KAU 14-857), 20.0 mm SL, female, stained with alizarin red,
<collectingCountry id="F35A7665FFF1A6191FC077F9EE2EFD72" box="[1246,1396,671,696]" name="Saudi Arabia" pageId="10" pageNumber="178">Saudi Arabia</collectingCountry>
,
<collectingMunicipality id="6B96AC8FFFF1A6191E9E77F9EB93FD17" pageId="10" pageNumber="178">Al Lith</collectingMunicipality>
, reef wall to depth of about
<elevation id="0060D1C6FFF1A619191377A2E9BCFD16" box="[525,742,708,733]" isEnumeration="true" pageId="10" pageNumber="178">
<quantity id="4CB59B10FFF1A619191377A2E929FD16" box="[525,627,708,733]" metricMagnitude="1" metricUnit="m" metricValue="3.75" metricValueMax="4.0" metricValueMin="3.5" pageId="10" pageNumber="178" unit="m" value="37.5" valueMax="40.0" valueMin="35.0">3540 m</quantity>
,
<quantity id="4CB59B10FFF1A619199E77A2E9BCFD16" box="[640,742,708,732]" metricMagnitude="1" metricUnit="m" metricValue="2.8" metricValueMax="2.9" metricValueMin="2.7" pageId="10" pageNumber="178" unit="m" value="28.0" valueMax="29.0" valueMin="27.0">2729 m</quantity>
</elevation>
, St. 41, 20°11'632''N, 40°03'137''E,
<collectingDate id="EFB7E9DDFFF1A6191F9F77A2EE05FD16" box="[1153,1375,708,733]" pageId="10" pageNumber="178" value="2014-11-17">
<date id="FFF31035FFF1A6191F9F77A2EE05FD16" box="[1153,1375,708,733]" pageId="10" pageNumber="178" value="2014-11-17">17 November 2014</date>
</collectingDate>
, coll.
<collectorName id="26B85323FFF1A6191B89778EEA0CFCCA" box="[151,342,743,768]" pageId="10" pageNumber="178">S.V. Bogorodsky</collectorName>
&amp;
<collectorName id="26B85323FFF1A6191A62778EE974FCCA" box="[380,558,743,768]" pageId="10" pageNumber="178">T.J. Alpermann</collectorName>
</materialsCitation>
; SMF 35737,
<quantity id="4CB59B10FFF1A61919D17781E86EFD35" box="[719,820,743,768]" metricMagnitude="-2" metricUnit="m" metricValue="2.16" pageId="10" pageNumber="178" unit="mm" value="21.6">21.6 mm</quantity>
SL, male, and KAUMM 368 (KAU14-858),
<quantity id="4CB59B10FFF1A6191E267781EEC7FD35" box="[1336,1437,743,768]" metricMagnitude="-2" metricUnit="m" metricValue="1.54" pageId="10" pageNumber="178" unit="mm" value="15.4">15.4 mm</quantity>
SL, sex unknown, collection data the same as for specimen SMF 35735.
</paragraph>
</subSubSection>
<subSubSection id="C357657EFFF1A6191BD97649E83AFB32" pageId="10" pageNumber="178" type="diagnosis">
<paragraph id="8BF236F5FFF1A6191BD97649E964FBA2" blockId="10.[151,1437,671,2029]" pageId="10" pageNumber="178">
<emphasis id="B939EAE7FFF1A6191BD97649EA60FC82" bold="true" box="[199,314,815,840]" pageId="10" pageNumber="178">Diagnosis</emphasis>
.
<taxonomicName id="4C4D4D76FFF1A6191A527657E940FC82" box="[332,538,815,840]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="178" phylum="Chordata" rank="species" species="fishelsoni">
<emphasis id="B939EAE7FFF1A6191A527657E940FC82" box="[332,538,815,840]" italics="true" pageId="10" pageNumber="178">Trimma fishelsoni</emphasis>
</taxonomicName>
differs from the other species of the genus in the following combination of characters: dorsal-fin rays VI + I,78, second spine elongate and filamentous; anal-fin rays I,78; pectoral-fin rays 1417 (usually 1516); longitudinal scale series 2528; median predorsal scales 914, scales reaching to interorbital area. Head scaled, with 12 rows of scales on cheek and 34 rows on opercle. Bony interorbital width broad, about half pupil diameter. Pelvic fins separated, without basal membrane or frenum, fifth ray unbranched, its length 4656% length of fourth ray. Pale greenish yellow densely dotted with melanophores on postorbital head, along back, and on lower half of body, with faint broad purple stripe above midside of body. Dorsal and anal fins broadly striped with pale yellow and blue, caudal fin yellow with blue margins and with two large bright yellow blotches in vertical alignment at base.
</paragraph>
<paragraph id="8BF236F5FFF1A6191BD97112EFB9FB1F" blockId="10.[151,1437,671,2029]" pageId="10" pageNumber="178">
<emphasis id="B939EAE7FFF1A6191BD97112EAEFFB46" box="[199,437,1140,1165]" italics="true" pageId="10" pageNumber="178">Head papilla pattern</emphasis>
. This species has a reduced transverse pattern of sensory papilla rows, shown in
<figureCitation id="13762A70FFF1A6191E4C7112EECDFB47" box="[1362,1431,1140,1165]" captionStart="FIGURE 5" captionStartId="12.[151,251,1939,1961]" captionTargetBox="[281,1337,196,1913]" captionTargetId="figure@12.[254,1337,193,1915]" captionTargetPageId="12" captionText="FIGURE 5. Trimma fishelsoni, SMF 35737, cephalic sensory papillae of head. Dorsal (top), lateral (middle) and ventral (bottom) views of head. Dots represent the sensory papillae; AN = anterior nares, PN = posterior nares; arrows show position of lower end of gill opening. Photo and drawing by T. Suzuki." httpUri="https://zenodo.org/record/266393/files/figure.png" pageId="10" pageNumber="178">Fig. 5</figureCitation>
, with number in each row (in parentheses) as follows:
<emphasis id="B939EAE7FFF1A61919EE71FFE9A4FB7A" box="[752,766,1177,1200]" italics="true" pageId="10" pageNumber="178">a</emphasis>
(6),
<emphasis id="B939EAE7FFF1A619182A71F1E818FB7A" box="[820,834,1175,1200]" italics="true" pageId="10" pageNumber="178">b</emphasis>
(4),
<emphasis id="B939EAE7FFF1A619186971FFE8D9FB7A" box="[887,899,1177,1200]" italics="true" pageId="10" pageNumber="178">c</emphasis>
(6),
<emphasis id="B939EAE7FFF1A61918A771FFE889FB7A" box="[953,979,1177,1200]" italics="true" pageId="10" pageNumber="178">cp</emphasis>
(1),
<emphasis id="B939EAE7FFF1A6191F1771FFEF7AFB7A" box="[1033,1056,1177,1200]" italics="true" pageId="10" pageNumber="178">cs</emphasis>
(2),
<emphasis id="B939EAE7FFF1A6191F4871F1EF3EFB7A" box="[1110,1124,1175,1200]" italics="true" pageId="10" pageNumber="178">d</emphasis>
(7),
<emphasis id="B939EAE7FFF1A6191F8471F1EFEBFB7A" box="[1178,1201,1175,1200]" italics="true" pageId="10" pageNumber="178">d</emphasis>
(9),
<emphasis id="B939EAE7FFF1A6191FF971FFEE58FB7A" box="[1255,1282,1177,1200]" italics="true" pageId="10" pageNumber="178">ea</emphasis>
(13),
<emphasis id="B939EAE7FFF1A6191E5B71FFEE3AFB7A" box="[1349,1376,1177,1200]" italics="true" pageId="10" pageNumber="178">ep</emphasis>
(14),
<emphasis id="B939EAE7FFF1A6191B8971DAEBC5FB1E" box="[151,159,1212,1236]" italics="true" pageId="10" pageNumber="178">f</emphasis>
(3),
<emphasis id="B939EAE7FFF1A6191BCA71DBEBB8FB1E" box="[212,226,1213,1236]" italics="true" pageId="10" pageNumber="178">g</emphasis>
(1),
<emphasis id="B939EAE7FFF1A6191A0671DBEA74FB1E" box="[280,302,1213,1236]" italics="true" pageId="10" pageNumber="178">ia</emphasis>
(8),
<emphasis id="B939EAE7FFF1A6191A7D71DBEA23FB1E" box="[355,377,1213,1236]" italics="true" pageId="10" pageNumber="178">ip</emphasis>
(7),
<emphasis id="B939EAE7FFF1A6191AB171DBEA99FB1E" box="[431,451,1213,1236]" italics="true" pageId="10" pageNumber="178">m</emphasis>
(1),
<emphasis id="B939EAE7FFF1A6191AE771DBE95DFB1E" box="[505,519,1213,1236]" italics="true" pageId="10" pageNumber="178">n</emphasis>
(4),
<emphasis id="B939EAE7FFF1A619192271DBE908FB1E" box="[572,594,1213,1236]" italics="true" pageId="10" pageNumber="178">ot</emphasis>
(12),
<emphasis id="B939EAE7FFF1A619198B71DBE9F4FB1E" box="[661,686,1213,1236]" italics="true" pageId="10" pageNumber="178">os</emphasis>
(6),
<emphasis id="B939EAE7FFF1A61919FA71DBE9A0FB1E" box="[740,762,1213,1236]" italics="true" pageId="10" pageNumber="178">oi</emphasis>
(4),
<emphasis id="B939EAE7FFF1A619183171DBE867FB1E" box="[815,829,1213,1236]" italics="true" pageId="10" pageNumber="178">p</emphasis>
(7),
<emphasis id="B939EAE7FFF1A619186D71DBE824FB1E" box="[883,894,1213,1236]" italics="true" pageId="10" pageNumber="178">r</emphasis>
(3),
<emphasis id="B939EAE7FFF1A61918AA71DBE8E5FB1E" box="[948,959,1213,1236]" italics="true" pageId="10" pageNumber="178">s</emphasis>
(1),
<emphasis id="B939EAE7FFF1A61918EA71DBEF58FB1E" box="[1012,1026,1213,1236]" italics="true" pageId="10" pageNumber="178">u</emphasis>
(5),
<emphasis id="B939EAE7FFF1A6191F2671DBEF1EFB1E" box="[1080,1092,1213,1236]" italics="true" pageId="10" pageNumber="178">x</emphasis>
(9), and
<emphasis id="B939EAE7FFF1A6191FB771DBEFEEFB1E" box="[1193,1204,1213,1236]" italics="true" pageId="10" pageNumber="178">z</emphasis>
(6).
</paragraph>
<paragraph id="8BF236F5FFF1A6191BD971B9E83AFB32" blockId="10.[151,1437,671,2029]" box="[199,864,1247,1272]" pageId="10" pageNumber="178">
Coloration of freshly collected specimen shown in
<figureCitation id="13762A70FFF1A619181C71B9E81FFB32" box="[770,837,1247,1272]" captionStart="FIGURE 4" captionStartId="11.[151,250,1954,1976]" captionTargetBox="[151,1436,752,1933]" captionTargetId="figure@11.[151,1436,739,1933]" captionTargetPageId="11" captionText="FIGURE 4. Trimma fishelsoni. A: SMF 35735, fresh specimen, 20.0 mm SL, female, Al Lith, Saudi Arabia, Red Sea; B: SMF 35737, preserved specimen, 21.6 mm SL, male, Al Lith, Saudi Arabia, Red Sea. Photo by S. V. Bogorodsky (A) and T. Suzuki (B)." httpUri="https://zenodo.org/record/266392/files/figure.png" pageId="10" pageNumber="178">Fig. 4</figureCitation>
A.
</paragraph>
</subSubSection>
<subSubSection id="C357657EFFF1A6191BD97062EE35FA1A" pageId="10" pageNumber="178" type="distribution">
<paragraph id="8BF236F5FFF1A6191BD97062EE35FA1A" blockId="10.[151,1437,671,2029]" pageId="10" pageNumber="178">
<emphasis id="B939EAE7FFF1A6191BD97062EAAAFAD7" bold="true" box="[199,496,1284,1309]" pageId="10" pageNumber="178">Distribution and habitat.</emphasis>
<taxonomicName id="4C4D4D76FFF1A6191AE87063E9E5FAD7" box="[502,703,1284,1309]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="178" phylum="Chordata" rank="species" species="fishelsoni">
<emphasis id="B939EAE7FFF1A6191AE87063E9E5FAD7" box="[502,703,1284,1309]" italics="true" pageId="10" pageNumber="178">Trimma fishelsoni</emphasis>
</taxonomicName>
was originally described based on
<specimenCount id="9D4BFD7CFFF1A6191F597062EF85FAD7" box="[1095,1247,1284,1309]" pageId="10" pageNumber="178" type="generic">20 specimens</specimenCount>
from the
<collectingRegion id="4989F817FFF1A6191E557062EE25FAD7" box="[1355,1407,1284,1309]" country="Papua New Guinea" name="Gulf" pageId="10" pageNumber="178">Gulf</collectingRegion>
of
<collectingRegion id="4989F817FFF1A6191B897041EBBAFA8A" box="[151,224,1319,1344]" country="Jordan" name="Al 'Aqabah" pageId="10" pageNumber="178">Aqaba</collectingRegion>
by
<bibRefCitation id="EFDC4B04FFF1A6191A0F704EEAF5FA8A" author="Goren" box="[273,431,1319,1344]" pageId="10" pageNumber="178" refString="Goren, M. (1985) Trimma fishelsoni, a new gobiid fish from the Gulf of Elat, northern Red Sea. Israel Journal of Zoology, 33 (1 / 2), 63 - 67." type="journal article" year="1985">Goren (1985)</bibRefCitation>
.
<bibRefCitation id="EFDC4B04FFF1A6191AA37041E9F4FA8A" author="Winterbottom" box="[445,686,1319,1344]" pageId="10" pageNumber="178" refString="Winterbottom, R. (1995) Red Sea gobiid fishes of the genus Trimma, with the description of two new species. Revue franzaise d'Aquariologie Herpйtologie, 22 (3 / 4), 93 - 98." type="journal article" year="1995">Winterbottom (1995)</bibRefCitation>
extended the range of distribution south to Ras Banas, southern
<collectingCountry id="F35A7665FFF1A6191B89702BEB84FAAF" box="[151,222,1357,1381]" name="Egypt" pageId="10" pageNumber="178">Egypt</collectingCountry>
. The range is here extended to the central
<collectingRegion id="4989F817FFF1A61919A8702AE84FFAAE" box="[694,789,1356,1381]" country="Egypt" name="Al Bahr al Ahmar" pageId="10" pageNumber="178">Red Sea</collectingRegion>
, at Al Lith, about
<quantity id="4CB59B10FFF1A61918FE702AEF6FFAAF" box="[992,1077,1356,1381]" metricMagnitude="5" metricUnit="m" metricValue="2.5" pageId="10" pageNumber="178" unit="km" value="250.0">250 km</quantity>
south from Jeddah. The species is typically seen hovering a few centimeters above the substratum, often upside down, forming small groups with each goby about
<quantity id="4CB59B10FFF1A6191A4A70F2EA98FA66" box="[340,450,1428,1452]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="10" pageNumber="178" unit="cm" value="20.0" valueMax="30.0" valueMin="10.0">1030 cm</quantity>
from other. Reported from coral reefs at depths of
<quantity id="4CB59B10FFF1A61918E770F2EF05FA66" box="[1017,1119,1428,1453]" metricMagnitude="1" metricUnit="m" metricValue="3.1" metricValueMax="4.7" metricValueMin="1.5" pageId="10" pageNumber="178" unit="m" value="31.0" valueMax="47.0" valueMin="15.0">1547 m</quantity>
; the specimens from Al Lith were collected from a cave at a depth of
<quantity id="4CB59B10FFF1A619194070DEE999FA1A" box="[606,707,1464,1488]" metricMagnitude="1" metricUnit="m" metricValue="2.8" metricValueMax="2.9" metricValueMin="2.7" pageId="10" pageNumber="178" unit="m" value="28.0" valueMax="29.0" valueMin="27.0">2729 m</quantity>
, together with
<taxonomicName id="4C4D4D76FFF1A619187470D1EF19FA1A" box="[874,1091,1463,1488]" class="Actinopterygii" family="Gobiidae" genus="Priolepis" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="178" phylum="Chordata" rank="species" species="melanops" status="sp. nov.">
<emphasis id="B939EAE7FFF1A619187470D1EF19FA1A" box="[874,1091,1463,1488]" italics="true" pageId="10" pageNumber="178">Priolepis melanops</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20A579CFFF1A6191F5470D1EFD2FA1A" box="[1098,1160,1463,1488]" pageId="10" pageNumber="178" rank="species">
<emphasis id="B939EAE7FFF1A6191F5470D1EFD2FA1A" bold="true" box="[1098,1160,1463,1488]" pageId="10" pageNumber="178">n. sp.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="4C4D4D76FFF1A6191FA170DFEE32FA1A" box="[1215,1384,1463,1488]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="178" phylum="Chordata" rank="species" species="taylori">
<emphasis id="B939EAE7FFF1A6191FA170DFEE32FA1A" box="[1215,1384,1463,1488]" italics="true" pageId="10" pageNumber="178">Trimma taylori</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C357657EFFF1A6191BD970BAE9D0F84A" pageId="10" pageNumber="178" type="discussion">
<paragraph id="8BF236F5FFF1A6191BD970BAEAD3F94F" blockId="10.[151,1437,671,2029]" pageId="10" pageNumber="178">
<emphasis id="B939EAE7FFF1A6191BD970BAEA6EFA3F" bold="true" box="[199,308,1500,1525]" pageId="10" pageNumber="178">Remarks</emphasis>
. The three specimens from Al Lith generally matches the description and diagnosis provided by
<bibRefCitation id="EFDC4B04FFF1A6191B897366EA6AF9D2" author="Goren" box="[151,304,1535,1560]" pageId="10" pageNumber="178" refString="Goren, M. (1985) Trimma fishelsoni, a new gobiid fish from the Gulf of Elat, northern Red Sea. Israel Journal of Zoology, 33 (1 / 2), 63 - 67." type="journal article" year="1985">Goren (1985)</bibRefCitation>
and
<bibRefCitation id="EFDC4B04FFF1A6191A747099E906F9D2" author="Winterbottom" box="[362,604,1535,1560]" pageId="10" pageNumber="178" refString="Winterbottom, R. (1995) Red Sea gobiid fishes of the genus Trimma, with the description of two new species. Revue franzaise d'Aquariologie Herpйtologie, 22 (3 / 4), 93 - 98." type="journal article" year="1995">Winterbottom (1995)</bibRefCitation>
. They differ, however, in having all pectoral-fin rays unbranched (versus upper and lower 57 rays unbranched). More specimens from southern part of the distribution are needed to determine whether unbranched rays represent intraspecific variation or are related to separate northern and southern populations.
</paragraph>
<paragraph id="8BF236F5FFF1A6191BD973E9E9D0F84A" blockId="10.[151,1437,671,2029]" pageId="10" pageNumber="178">
The colour of live fish (
<figureCitation id="13762A70FFF1A6191AD073E9E94AF962" box="[462,528,1679,1704]" captionStart="FIGURE 6" captionStartId="13.[151,250,1557,1579]" captionTargetBox="[150,1435,193,1535]" captionTargetId="figure@13.[150,1436,193,1536]" captionTargetPageId="13" captionText="FIGURE 6. Underwater photographs of alive individuals. A: Trimma fishelsoni, Dahab, Gulf of Aqaba, Red Sea; B: Trimma taylori, Al Lith, Saudi Arabia, Red Sea. Photos by S. V. Bogorodsky." httpUri="https://zenodo.org/record/266394/files/figure.png" pageId="10" pageNumber="178">Fig. 6</figureCitation>
A) is slightly different from that of freshly collected specimens (
<figureCitation id="13762A70FFF1A6191FC273E9EE45F962" box="[1244,1311,1679,1704]" captionStart="FIGURE 4" captionStartId="11.[151,250,1954,1976]" captionTargetBox="[151,1436,752,1933]" captionTargetId="figure@11.[151,1436,739,1933]" captionTargetPageId="11" captionText="FIGURE 4. Trimma fishelsoni. A: SMF 35735, fresh specimen, 20.0 mm SL, female, Al Lith, Saudi Arabia, Red Sea; B: SMF 35737, preserved specimen, 21.6 mm SL, male, Al Lith, Saudi Arabia, Red Sea. Photo by S. V. Bogorodsky (A) and T. Suzuki (B)." httpUri="https://zenodo.org/record/266392/files/figure.png" pageId="10" pageNumber="178">Fig. 4</figureCitation>
A): a broad stripe above midside of body dark purple; iris mainly dusky yellow with an oblique blue line across upper edge of pupil; each dorsal and anal fins with yellow stripe, rest of fin blue; and caudal fin with blue margins.
<bibRefCitation id="EFDC4B04FFF1A6191E1A73BEEEC6F93A" author="Goren" box="[1284,1436,1751,1776]" pageId="10" pageNumber="178" refString="Goren, M. (1985) Trimma fishelsoni, a new gobiid fish from the Gulf of Elat, northern Red Sea. Israel Journal of Zoology, 33 (1 / 2), 63 - 67." type="journal article" year="1985">Goren (1985)</bibRefCitation>
described presence of four faint saddle-like dusky blotches, one under each dorsal fin and two on the caudal peduncle which were apparent in preserved specimen; such blotches not visible in live gobies or in fresh specimens. Body along back in preserved specimens from Al Lith (
<figureCitation id="13762A70FFF1A61918857222E8BBF896" box="[923,993,1860,1885]" captionStart="FIGURE 4" captionStartId="11.[151,250,1954,1976]" captionTargetBox="[151,1436,752,1933]" captionTargetId="figure@11.[151,1436,739,1933]" captionTargetPageId="11" captionText="FIGURE 4. Trimma fishelsoni. A: SMF 35735, fresh specimen, 20.0 mm SL, female, Al Lith, Saudi Arabia, Red Sea; B: SMF 35737, preserved specimen, 21.6 mm SL, male, Al Lith, Saudi Arabia, Red Sea. Photo by S. V. Bogorodsky (A) and T. Suzuki (B)." httpUri="https://zenodo.org/record/266392/files/figure.png" pageId="10" pageNumber="178">Fig. 4</figureCitation>
B) densely covered with melanophores but not form patches or saddle-like blotches.
</paragraph>
</subSubSection>
<subSubSection id="C357657EFFF1A6181BD972EBEF1DFD07" lastPageId="11" lastPageNumber="179" pageId="10" pageNumber="178" type="reference_group">
<paragraph id="8BF236F5FFF1A6181BD972EBE859FE8A" blockId="10.[151,1437,671,2029]" lastBlockId="11.[151,1436,151,717]" lastPageId="11" lastPageNumber="179" pageId="10" pageNumber="178">
<taxonomicName id="4C4D4D76FFF1A6191BD972EBEAC8F86F" ID-CoL="58V5J" box="[199,402,1932,1957]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="178" phylum="Chordata" rank="species" species="fishelsoni">
<emphasis id="B939EAE7FFF1A6191BD972EBEAC8F86F" box="[199,402,1932,1957]" italics="true" pageId="10" pageNumber="178">Trimma fishelsoni</emphasis>
</taxonomicName>
may be distinguished from other
<collectingRegion id="4989F817FFF1A619180872EAE82EF86E" box="[790,884,1932,1957]" country="Egypt" name="Al Bahr al Ahmar" pageId="10" pageNumber="178">Red Sea</collectingRegion>
congeners by a combination of scaled predorsal area, cheek and opercle, a very long second dorsal-fin spine, a moderately broad interorbital area, no spots on the head and body, body with an indistinct, dark purple stripe above midside, and each dorsal and anal fin with a yellow band.
<taxonomicName id="4C4D4D76FFF0A6181A2B75FFE958FF7A" ID-CoL="58V5J" box="[309,514,151,176]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="179" phylum="Chordata" rank="species" species="fishelsoni">
<emphasis id="B939EAE7FFF0A6181A2B75FFE958FF7A" box="[309,514,151,176]" italics="true" pageId="11" pageNumber="179">Trimma fishelsoni</emphasis>
</taxonomicName>
is assigned to the
<taxonomicName id="4C4D4D76FFF0A61819FD75FFE80DFF7A" ID-CoL="58V7Q" box="[739,855,153,176]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="179" phylum="Chordata" rank="species" species="tevegae">
<emphasis id="B939EAE7FFF0A61819FD75FFE80DFF7A" box="[739,855,153,176]" italics="true" pageId="11" pageNumber="179">T. tevegae</emphasis>
</taxonomicName>
species group and is most similar to
<taxonomicName id="4C4D4D76FFF0A6181E0D75FFEA93FF1E" ID-CoL="58V5T" authority="Winterbottom &amp; Zur, 2007" authorityName="Winterbottom &amp; Zur" authorityYear="2007" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="179" phylum="Chordata" rank="species" species="gigantum">
<emphasis id="B939EAE7FFF0A6181E0D75FFEEC6FF7A" box="[1299,1436,153,176]" italics="true" pageId="11" pageNumber="179">T. gigantum</emphasis>
<bibRefCitation id="EFDC4B04FFF0A6181B8975DAEA93FF1E" author="Winterbottom" box="[151,457,188,213]" pageId="11" pageNumber="179" refString="Winterbottom, R. &amp; Zur, M. (2007) Three new species of genus Trimma from Palau, Western Pacific (Percomorpha; Gobiidae). aqua, International Journal of Ichthyology, 13, 13 - 24." type="journal article" year="2007">Winterbottom &amp; Zur, 2007</bibRefCitation>
</taxonomicName>
, described from
<collectingCountry id="F35A7665FFF0A618199B75DAE99CFF1F" box="[645,710,188,213]" name="Palau" pageId="11" pageNumber="179">Palau</collectingCountry>
, western Pacific. It differs in having fewer predorsal scales (11 14 versus 1418), in the presence of two yellow blotches at the caudal-fin base (versus only thin yellow line), and in having the lower half of the body and pectoral-fin base densely pigmented with melanophores (versus a few scattered melanophores on body and pectoral-fin base).
</paragraph>
<paragraph id="8BF236F5FFF0A6181BD9742AEF1DFD07" blockId="11.[151,1436,151,717]" pageId="11" pageNumber="179">
The similarity of
<taxonomicName id="4C4D4D76FFF0A6181A97742BE909FEAF" ID-CoL="58V5J" box="[393,595,332,357]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="179" phylum="Chordata" rank="species" species="fishelsoni">
<emphasis id="B939EAE7FFF0A6181A97742BE909FEAF" box="[393,595,332,357]" italics="true" pageId="11" pageNumber="179">Trimma fishelsoni</emphasis>
</taxonomicName>
with
<taxonomicName id="4C4D4D76FFF0A618198C742BE858FEAE" ID-CoL="58V7Q" box="[658,770,333,356]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="179" phylum="Chordata" rank="species" species="tevegae">
<emphasis id="B939EAE7FFF0A618198C742BE858FEAE" box="[658,770,333,356]" italics="true" pageId="11" pageNumber="179">T. tevegae</emphasis>
</taxonomicName>
led confusion in identification of specimens collected from the vicinity of Jeddah, Saudi Arabia.
<materialsCitation id="3B253CA8FFF0A618192B7409EF1DFD07" ID-GBIF-Occurrence="1291547732" collectionCode="ANSP, WAM" collectorName="J. E. Randall &amp; Three &amp; Randall &amp; Vine &amp; Jeddah &amp; D. Hoese &amp; Recently" country="Maldives" location="Winterbottom" municipality="In" pageId="11" pageNumber="179" specimenCode="ANSP 158831, WAM 25787.008" specimenCount="1" stateProvince="Red Sea">
<bibRefCitation id="EFDC4B04FFF0A618192B7409E878FE42" author="Winterbottom" box="[565,802,367,392]" pageId="11" pageNumber="179" refString="Winterbottom, R. (1995) Red Sea gobiid fishes of the genus Trimma, with the description of two new species. Revue franzaise d'Aquariologie Herpйtologie, 22 (3 / 4), 93 - 98." type="journal article" year="1995">
<location id="8E92602EFFF0A618192B7409E98BFE42" LSID="urn:lsid:plazi:treatment:03E487E3FFF1A61E1B8974F7E8A4F8D5:8E92602EFFF0A618192B7409E98BFE42" box="[565,721,367,392]" country="Maldives" municipality="In" name="Winterbottom" pageId="11" pageNumber="179" stateProvince="Red Sea">Winterbottom</location>
(1995)
</bibRefCitation>
included
<taxonomicName id="4C4D4D76FFF0A61818917417E8A5FE42" ID-CoL="58V7Q" box="[911,1023,369,392]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="179" phylum="Chordata" rank="species" species="tevegae">
<emphasis id="B939EAE7FFF0A61818917417E8A5FE42" box="[911,1023,369,392]" italics="true" pageId="11" pageNumber="179">T. tevegae</emphasis>
</taxonomicName>
in his review of the
<collectingRegion id="4989F817FFF0A6181FFC7409EE65FE42" box="[1250,1343,367,392]" country="Egypt" name="Al Bahr al Ahmar" pageId="11" pageNumber="179">Red Sea</collectingRegion>
<taxonomicName id="4C4D4D76FFF0A6181E5B7417EEC6FE42" ID-CoL="827C" box="[1349,1436,369,392]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="179" phylum="Chordata" rank="genus">
<emphasis id="B939EAE7FFF0A6181E5B7417EEC6FE42" box="[1349,1436,369,392]" italics="true" pageId="11" pageNumber="179">Trimma</emphasis>
</taxonomicName>
(
<collectionCode id="ED5CAE30FFF0A6181B8074F2EBBDFE66" box="[158,231,404,428]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="11" pageNumber="179">ANSP</collectionCode>
, without catalogue number) and used an underwater photograph of
<taxonomicName id="4C4D4D76FFF0A61818F674F3EB88FE1A" ID-CoL="58V4Z" authority="Yoshino &amp; Araga, 1975" authorityName="Yoshino &amp; Araga" authorityYear="1975" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="179" phylum="Chordata" rank="species" species="caudomaculatum">
<emphasis id="B939EAE7FFF0A61818F674F3EF9FFE67" box="[1000,1221,404,429]" italics="true" pageId="11" pageNumber="179">T. caudomaculatum</emphasis>
<bibRefCitation id="EFDC4B04FFF0A6181FD074F2EB88FE1A" author="Yoshino" pageId="11" pageNumber="179" refString="Yoshino, T. &amp; Araga, C. (1975) Trimma caudomaculata. In: Masuda, H., Araga, C. &amp; Yoshino, T. (Eds.), Coastal fishes of southern Japan. Tokai University Press, Tokyo, 379 pp." type="book" year="1975">Yoshino &amp; Araga, 1975</bibRefCitation>
</taxonomicName>
, taken by
<collectorName id="26B85323FFF0A6181A5D74DEEA91FE1A" box="[323,459,439,464]" pageId="11" pageNumber="179">J.E. Randall</collectorName>
at the
<collectingCountry id="F35A7665FFF0A618190B74D1E9DAFE1A" box="[533,640,439,464]" name="Maldives" pageId="11" pageNumber="179">Maldives</collectingCountry>
, as
<taxonomicName id="4C4D4D76FFF0A61819B774DFE843FE1A" ID-CoL="58V7Q" box="[681,793,441,464]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="179" phylum="Chordata" rank="species" species="tevegae">
<emphasis id="B939EAE7FFF0A61819B774DFE843FE1A" box="[681,793,441,464]" italics="true" pageId="11" pageNumber="179">T. tevegae</emphasis>
</taxonomicName>
.
<collectingMunicipality id="6B96AC8FFFF0A618183674DEE865FE05" box="[808,831,440,463]" pageId="11" pageNumber="179">In</collectingMunicipality>
fact, six specimens of this lot (
<specimenCode id="DBEB9E8EFFF0A6181F8774DEEE1AFE1A" box="[1177,1344,439,464]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="11" pageNumber="179">
<collectionCode id="ED5CAE30FFF0A6181F8774DEEFB8FE1A" box="[1177,1250,440,464]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="11" pageNumber="179">ANSP</collectionCode>
158831
</specimenCode>
, labeled
<taxonomicName id="4C4D4D76FFF0A6181B8974BBEBB7FE3E" ID-CoL="827C" box="[151,237,477,500]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="179" phylum="Chordata" rank="genus">
<emphasis id="B939EAE7FFF0A6181B8974BBEBB7FE3E" box="[151,237,477,500]" italics="true" pageId="11" pageNumber="179">Trimma</emphasis>
</taxonomicName>
sp.) are
<taxonomicName id="4C4D4D76FFF0A6181A5374BBE952FE3F" ID-CoL="58V5N" box="[333,520,476,501]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="179" phylum="Chordata" rank="species" species="flavicaudatum">
<emphasis id="B939EAE7FFF0A6181A5374BBE952FE3F" box="[333,520,476,501]" italics="true" pageId="11" pageNumber="179">T. flavicaudatum</emphasis>
</taxonomicName>
(Rick Winterbottom, pers. comm.).
<collectorName id="26B85323FFF0A618188374BAE885FE3F" box="[925,991,476,501]" pageId="11" pageNumber="179">Three</collectorName>
specimens from the
<collectingRegion id="4989F817FFF0A6181FD874BBEE7BFE3E" box="[1222,1313,477,500]" country="Papua New Guinea" name="Western" pageId="11" pageNumber="179">Western</collectingRegion>
Australian Museum (
<specimenCode id="DBEB9E8EFFF0A6181A167766EA8AFDD2" box="[264,464,511,536]" collectionCode="WAM" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="11" pageNumber="179">
<collectionCode id="ED5CAE30FFF0A6181A167766EA15FDD2" box="[264,335,512,536]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="11" pageNumber="179">WAM</collectionCode>
25787.008
</specimenCode>
) were collected by
<collectorName id="26B85323FFF0A61819AF7499E852FDD2" box="[689,776,511,536]" pageId="11" pageNumber="179">Randall</collectorName>
and
<collectorName id="26B85323FFF0A618185F7499E82CFDD2" box="[833,886,511,536]" pageId="11" pageNumber="179">Vine</collectorName>
in 1977 from
<collectorName id="26B85323FFF0A6181F027499EF36FDD2" box="[1052,1132,511,536]" pageId="11" pageNumber="179">Jeddah</collectorName>
, and were identified by
<collectorName id="26B85323FFF0A6181E9F7766EB87FDF6" pageId="11" pageNumber="179">D. Hoese</collectorName>
as
<taxonomicName id="4C4D4D76FFF0A6181A1F7743EA2BFDF6" ID-CoL="58V7Q" box="[257,369,549,572]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="179" phylum="Chordata" rank="species" species="tevegae">
<emphasis id="B939EAE7FFF0A6181A1F7743EA2BFDF6" box="[257,369,549,572]" italics="true" pageId="11" pageNumber="179">T. tevegae</emphasis>
</taxonomicName>
.
<collectorName id="26B85323FFF0A6181A617742EAB9FDF7" box="[383,483,548,573]" pageId="11" pageNumber="179">Recently</collectorName>
specimens of this lot were examined and re-identified as
<taxonomicName id="4C4D4D76FFF0A6181F7D7743EFB3FDF7" ID-CoL="58V5J" box="[1123,1257,548,573]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="179" phylum="Chordata" rank="species" species="fishelsoni">
<emphasis id="B939EAE7FFF0A6181F7D7743EFB3FDF7" box="[1123,1257,548,573]" italics="true" pageId="11" pageNumber="179">T. fishelsoni</emphasis>
</taxonomicName>
(Winterbottom, pers. comm.).
<taxonomicName id="4C4D4D76FFF0A6181A5C772FEAA1FDAA" ID-CoL="58V7Q" box="[322,507,585,608]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="179" phylum="Chordata" rank="species" species="tevegae">
<emphasis id="B939EAE7FFF0A6181A5C772FEAA1FDAA" box="[322,507,585,608]" italics="true" pageId="11" pageNumber="179">Trimma tevegae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4D4D76FFF0A6181922772FE844FDAA" ID-CoL="58V4Z" box="[572,798,583,608]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="179" phylum="Chordata" rank="species" species="caudomaculatum">
<emphasis id="B939EAE7FFF0A6181922772FE844FDAA" box="[572,798,583,608]" italics="true" pageId="11" pageNumber="179">T. caudomaculatum</emphasis>
</taxonomicName>
thus currently not known from the
<collectingRegion id="4989F817FFF0A6181FCE7721EE6FFDAA" box="[1232,1333,583,608]" country="Egypt" name="Al Bahr al Ahmar" pageId="11" pageNumber="179">Red Sea</collectingRegion>
.
<taxonomicName id="4C4D4D76FFF0A6181E5B772FEA58FD4F" ID-CoL="58V5J" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="179" phylum="Chordata" rank="species" species="fishelsoni">
<emphasis id="B939EAE7FFF0A6181E5B772FEA58FD4F" italics="true" pageId="11" pageNumber="179">Trimma fishelsoni</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C4D4D76FFF0A6181A84770BE92DFD4F" ID-CoL="58V4Z" box="[410,631,620,645]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="179" phylum="Chordata" rank="species" species="caudomaculatum">
<emphasis id="B939EAE7FFF0A6181A84770BE92DFD4F" box="[410,631,620,645]" italics="true" pageId="11" pageNumber="179">T. caudomaculatum</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4D4D76FFF0A61819AF770BE879FD4E" ID-CoL="58V7Q" box="[689,803,621,644]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="179" phylum="Chordata" rank="species" species="tevegae">
<emphasis id="B939EAE7FFF0A61819AF770BE879FD4E" box="[689,803,621,644]" italics="true" pageId="11" pageNumber="179">T. tevegae</emphasis>
</taxonomicName>
in having more pectoral-fin rays (usually 1516 versus usually 14), and in lacking a dark blotch on the caudal peduncle (dark blotch, covering the posterior caudal peduncle and the basal two-fifths of the caudal fin, present in the other two species).
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DF32667DFFF0A6181B8972C4EB9AF83C" httpUri="https://zenodo.org/record/266392/files/figure.png" pageId="11" pageNumber="179" targetBox="[151,1436,752,1933]" targetPageId="11">
<paragraph id="8BF236F5FFF0A6181B8972C4EB9AF83C" blockId="11.[151,1436,1954,2038]" pageId="11" pageNumber="179">
<emphasis id="B939EAE7FFF0A6181B8972C4EA48F87D" bold="true" box="[151,274,1954,1976]" pageId="11" pageNumber="179">FIGURE 4.</emphasis>
<taxonomicName id="4C4D4D76FFF0A6181A0772C5EA88F872" box="[281,466,1954,1976]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="179" phylum="Chordata" rank="species" species="fishelsoni">
<emphasis id="B939EAE7FFF0A6181A0772C5EA88F872" box="[281,466,1954,1976]" italics="true" pageId="11" pageNumber="179">Trimma fishelsoni</emphasis>
</taxonomicName>
.
<emphasis id="B939EAE7FFF0A6181AC172C4EAABF87D" bold="true" box="[479,497,1954,1975]" pageId="11" pageNumber="179">A</emphasis>
: SMF 35735, fresh specimen, 20.0 mm SL, female, Al Lith, Saudi Arabia, Red Sea;
<emphasis id="B939EAE7FFF0A6181E5572C5EE06F872" bold="true" box="[1355,1372,1955,1976]" pageId="11" pageNumber="179">B</emphasis>
: SMF 35737, preserved specimen, 21.6 mm SL, male, Al Lith, Saudi Arabia, Red Sea. Photo by S.V. Bogorodsky (A) and T. Suzuki (B).
</paragraph>
</caption>
<caption id="DF32667DFFF7A61F1B8972F5E842F82C" httpUri="https://zenodo.org/record/266393/files/figure.png" pageId="12" pageNumber="180" targetBox="[281,1337,196,1913]" targetPageId="12">
<paragraph id="8BF236F5FFF7A61F1B8972F5E842F82C" blockId="12.[151,1436,1939,2022]" pageId="12" pageNumber="180">
<emphasis id="B939EAE7FFF7A61F1B8972F5EA4DF862" bold="true" box="[151,279,1939,1961]" pageId="12" pageNumber="180">FIGURE 5.</emphasis>
<taxonomicName id="4C4D4D76FFF7A61F1A3E72F2EA86F863" box="[288,476,1939,1961]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="180" phylum="Chordata" rank="species" species="fishelsoni">
<emphasis id="B939EAE7FFF7A61F1A3E72F2EA86F863" box="[288,476,1939,1961]" italics="true" pageId="12" pageNumber="180">Trimma fishelsoni</emphasis>
</taxonomicName>
, SMF 35737, cephalic sensory papillae of head. Dorsal (top), lateral (middle) and ventral (bottom) views of head. Dots represent the sensory papillae; AN= anterior nares, PN= posterior nares; arrows show position of lower end of gill opening. Photo and drawing by T. Suzuki.
</paragraph>
</caption>
<caption id="DF32667DFFF6A61E1B897373E811F980" httpUri="https://zenodo.org/record/266394/files/figure.png" pageId="13" pageNumber="181" targetBox="[150,1435,193,1535]" targetPageId="13">
<paragraph id="8BF236F5FFF6A61E1B897373E811F980" blockId="13.[151,1436,1557,1610]" pageId="13" pageNumber="181">
<emphasis id="B939EAE7FFF6A61E1B897373EA4EF9E0" bold="true" box="[151,276,1557,1579]" pageId="13" pageNumber="181">FIGURE 6.</emphasis>
Underwater photographs of alive individuals.
<emphasis id="B939EAE7FFF6A61E19EC7373E85EF9E0" bold="true" box="[754,772,1557,1578]" pageId="13" pageNumber="181">A</emphasis>
:
<taxonomicName id="4C4D4D76FFF6A61E180D7370EF47F9E1" authority="Dahab" authorityName="Dahab" box="[787,1053,1557,1579]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="181" phylum="Chordata" rank="species" species="fishelsoni">
<emphasis id="B939EAE7FFF6A61E180D7370E890F9E1" box="[787,970,1557,1579]" italics="true" pageId="13" pageNumber="181">Trimma fishelsoni</emphasis>
, Dahab
</taxonomicName>
, Gulf of Aqaba, Red Sea;
<emphasis id="B939EAE7FFF6A61E1E307370EE65F9E1" bold="true" box="[1326,1343,1558,1579]" pageId="13" pageNumber="181">B</emphasis>
:
<taxonomicName id="4C4D4D76FFF6A61E1E537370EA6EF980" authority="Al Lith" authorityName="Al Lith" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="181" phylum="Chordata" rank="species" species="taylori">
<emphasis id="B939EAE7FFF6A61E1E537370EB81F980" italics="true" pageId="13" pageNumber="181">Trimma taylori</emphasis>
, Al Lith
</taxonomicName>
, Saudi Arabia, Red Sea. Photos by S.V. Bogorodsky.
</paragraph>
</caption>
<subSubSection id="C357657EFFF6A61E1BD97310E8A4F8D5" pageId="13" pageNumber="181" type="description">
<paragraph id="8BF236F5FFF6A61E1BD97310E8A4F8D5" blockId="13.[151,1436,1654,1824]" pageId="13" pageNumber="181">
The three specimens of
<taxonomicName id="4C4D4D76FFF6A61E1ACA731EE900F945" box="[468,602,1654,1679]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="181" phylum="Chordata" rank="species" species="fishelsoni">
<emphasis id="B939EAE7FFF6A61E1ACA731EE900F945" box="[468,602,1654,1679]" italics="true" pageId="13" pageNumber="181">T. fishelsoni</emphasis>
</taxonomicName>
from Al Lith were collected together with
<taxonomicName id="4C4D4D76FFF6A61E1F21731EEFFEF945" box="[1087,1188,1654,1679]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="181" phylum="Chordata" rank="species" species="taylori">
<emphasis id="B939EAE7FFF6A61E1F21731EEFFEF945" box="[1087,1188,1654,1679]" italics="true" pageId="13" pageNumber="181">T. taylori</emphasis>
</taxonomicName>
(
<figureCitation id="13762A70FFF6A61E1FA87310EFA2F945" box="[1206,1272,1654,1679]" captionStart="FIGURE 6" captionStartId="13.[151,250,1557,1579]" captionTargetBox="[150,1435,193,1535]" captionTargetId="figure@13.[150,1436,193,1536]" captionTargetPageId="13" captionText="FIGURE 6. Underwater photographs of alive individuals. A: Trimma fishelsoni, Dahab, Gulf of Aqaba, Red Sea; B: Trimma taylori, Al Lith, Saudi Arabia, Red Sea. Photos by S. V. Bogorodsky." httpUri="https://zenodo.org/record/266394/files/figure.png" pageId="13" pageNumber="181">Fig. 6</figureCitation>
B) close to the entrance of a small cave on a reef wall. These two species can be easily confused under water. In the
<collectingRegion id="4989F817FFF6A61E1FE573FCEE75F979" box="[1275,1327,1690,1715]" country="Papua New Guinea" name="Gulf" pageId="13" pageNumber="181">Gulf</collectingRegion>
of
<collectingRegion id="4989F817FFF6A61E1E4D73FCEEC6F979" box="[1363,1436,1690,1715]" country="Jordan" name="Al 'Aqabah" pageId="13" pageNumber="181">Aqaba</collectingRegion>
they are not observed together, with individuals of
<taxonomicName id="4C4D4D76FFF6A61E19C573A6E83EF91D" box="[731,868,1726,1751]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="181" phylum="Chordata" rank="species" species="fishelsoni">
<emphasis id="B939EAE7FFF6A61E19C573A6E83EF91D" box="[731,868,1726,1751]" italics="true" pageId="13" pageNumber="181">T. fishelsoni</emphasis>
</taxonomicName>
often swimming in a dispersed group close to the substrate and quickly retiring inside a cave or holes when alarmed, whereas
<taxonomicName id="4C4D4D76FFF6A61E1F227382EFFDF931" box="[1084,1191,1762,1787]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="181" phylum="Chordata" rank="species" species="taylori">
<emphasis id="B939EAE7FFF6A61E1F227382EFFDF931" box="[1084,1191,1762,1787]" italics="true" pageId="13" pageNumber="181">T. taylori</emphasis>
</taxonomicName>
forms small groups swimming at some distance from reef, and usually swim away when alarmed.
</paragraph>
</subSubSection>
</treatment>
</document>