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<mods:title id="547952A36C0B0E454508EE56267963B9">Revision of the wolf spider genus Diahogna Roewer, 1960 (Araneae, Lycosidae)</mods:title>
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<subSubSection id="C36C65545057B35987E1378CFCFEFA4A" box="[510,809,1434,1458]" pageId="3" pageNumber="275" type="nomenclature">
<paragraph id="8BC936DF5057B35987E1378CFCFEFA4A" blockId="3.[430,903,1369,1458]" box="[510,809,1434,1458]" pageId="3" pageNumber="275">
<heading id="D08181B35057B35987E1378CFCFEFA4A" box="[510,809,1434,1458]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="275" reason="2">
<taxonomicName id="4C764D5C5057B35987E1378CFCFEFA4A" ID-CoL="444G" authority="Roewer, 1960" authorityName="Roewer" authorityYear="1960" box="[510,809,1434,1458]" class="Arachnida" family="Lycosidae" genus="Diahogna" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="275" phylum="Arthropoda" rank="genus">
<emphasis id="B902EACD5057B35987E1378CFDABFA4A" bold="true" box="[510,636,1434,1458]" italics="true" pageId="3" pageNumber="275">Diahogna</emphasis>
<bibRefCitation id="EFE74B2E5057B359849A378CFCFEFA4A" author="Roewer CF" box="[645,809,1434,1458]" pageId="3" pageNumber="275" pagination="519 - 1040" refId="ref9730" refString="Roewer CF. 1960. Araneae Lycosiformae II (Lycosidae) (Fortsetzung und Schluss). Exploration du Parc National de l'Upemba-Mission GF de Witte 55: 519 - 1040. (Imprint date 1959)." type="journal article" year="1960">Roewer, 1960</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<paragraph id="8BC936DF5057B359869537CAFC71FA0B" blockId="3.[138,934,1499,1523]" box="[138,934,1499,1523]" pageId="3" pageNumber="275">
<emphasis id="B902EACD5057B359869537CAFEC0FA0B" box="[138,279,1499,1523]" italics="true" pageId="3" pageNumber="275">
<typeStatus id="54CD887D5057B359869537CAFF17FA0B" box="[138,192,1500,1523]" pageId="3" pageNumber="275">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C764D5C5057B359873C37CAFD53FA0B" authority="Karsch, 1878" authorityName=", Karsch" authorityYear="1878" box="[291,644,1499,1523]" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="275" phylum="Arthropoda" rank="species" species="martensii">
<emphasis id="B902EACD5057B359873C37CAFE0DFA0B" box="[291,474,1499,1523]" italics="true" pageId="3" pageNumber="275">Lycosa martensii</emphasis>
<bibRefCitation id="EFE74B2E5057B35987FC37CDFD53FA0B" author="Karsch F" box="[483,644,1499,1523]" pageId="3" pageNumber="275" pagination="771 - 826" refId="ref9206" refString="Karsch F. 1878. Exotisch-araneologisches. Zeitschrift fur die Gesamten Naturwissenschaften 51: 332 - 333, 771 - 826." type="journal article" year="1878">Karsch, 1878</bibRefCitation>
</taxonomicName>
. By original designation.
</paragraph>
<paragraph id="8BC936DF5057B3598695343AFF24F9BC" blockId="3.[138,243,1580,1604]" box="[138,243,1580,1604]" pageId="3" pageNumber="275">
<emphasis id="B902EACD5057B3598695343AFF24F9BC" box="[138,243,1580,1604]" italics="true" pageId="3" pageNumber="275">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BC936DF5057B35C8695344BFEEDFE27" blockId="3.[138,1185,1629,1685]" lastBlockId="6.[125,1172,136,479]" lastPageId="6" lastPageNumber="278" pageId="3" pageNumber="275">
<taxonomicName id="4C764D5C5057B3598695344BFF21F98D" authorityName="Roewer" authorityYear="1960" box="[138,246,1629,1653]" class="Arachnida" family="Lycosidae" genus="Diahogna" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="275" phylum="Arthropoda" rank="genus">
<emphasis id="B902EACD5057B3598695344BFF21F98D" box="[138,246,1629,1653]" italics="true" pageId="3" pageNumber="275">Diahogna</emphasis>
</taxonomicName>
differs from all other wolf spider genera by the following combination of somatic and genitalic characters. The dorsal carapace profile is straight in lateral view (
<figureCitation id="134D2A5A5057B3598204346BFB44F96D" box="[1051,1171,1661,1685]" captionStart="Figure 2" captionStartId="5.[138,198,1584,1603]" captionTargetBox="[138,1177,185,1548]" captionTargetId="figure-0@5.[138,1185,183,1548]" captionTargetPageId="5" captionText="Figure 2. Male of Diahogna martensii (Karsch) from Cape Naturaliste, Tasmania, Australia (QVMAG 13:42218). (A, B) Carapace profile, dorsal and lateral views, respectively; (C) eye pattern in frontal view; (D) left male pedipalp, apical part of bulbus; (E, F) left male pedipalp, ventral and lateral views, respectively. Scale bar: 3.93 mm (A); 4.17 mm (B); 1.16 mm (C); 1.04 mm (D); 0.77 mm (E, F)." figureDoi="http://doi.org/10.5281/zenodo.4581381" httpUri="https://zenodo.org/record/4581381/files/figure.png" pageId="3" pageNumber="275">Figure 2B</figureCitation>
), the carapace itself is uniformly brown with a lighter area medially, sometimes in the form of a Y-shaped median band (
<figureCitation id="134D2A5A5052B35C87D032BEFDFCFF38" box="[463,555,168,192]" captionStart="Figure 1" captionStartId="4.[125,185,1558,1577]" captionTargetBox="[266,1032,179,1519]" captionTargetId="figure-1@4.[266,1033,179,1522]" captionTargetPageId="4" captionText="Figure 1. Photographs (dorsal view) of species of Diahogna. (A) Female of D. martensii (Karsch) from Blackmans Lagoon, Tasmania, Australia (QVMAG 13:42205); (B) female of D. exculta (L. Koch) from OBrians Gap, Wollongong, New South Wales, Australia (AM KS58056); (C) female of D. hildegardae n. sp. from Tambo River at Doctors Flat, Victoria, Australia (WAM T48116); (D) male holotype of D. pisauroides n. sp. from Adelaide River, Northern Territory, Australia (NTMAG A485). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4581379" httpUri="https://zenodo.org/record/4581379/files/figure.png" pageId="6" pageNumber="278">Figure 1</figureCitation>
AD). The row of the AE is straight or slightly procurved and wider than the row of the PME (
<figureCitation id="134D2A5A5052B35C84AE32DEFCFCFF18" box="[689,811,200,224]" captionStart="Figure 2" captionStartId="5.[138,198,1584,1603]" captionTargetBox="[138,1177,185,1548]" captionTargetId="figure-0@5.[138,1185,183,1548]" captionTargetPageId="5" captionText="Figure 2. Male of Diahogna martensii (Karsch) from Cape Naturaliste, Tasmania, Australia (QVMAG 13:42218). (A, B) Carapace profile, dorsal and lateral views, respectively; (C) eye pattern in frontal view; (D) left male pedipalp, apical part of bulbus; (E, F) left male pedipalp, ventral and lateral views, respectively. Scale bar: 3.93 mm (A); 4.17 mm (B); 1.16 mm (C); 1.04 mm (D); 0.77 mm (E, F)." figureDoi="http://doi.org/10.5281/zenodo.4581381" httpUri="https://zenodo.org/record/4581381/files/figure.png" pageId="6" pageNumber="278">Figure 2C</figureCitation>
). The row of the PLE is much wider than that of the PME giving the spider a pisaurid appearance (
<figureCitation id="134D2A5A5052B35C85B332FEFBC4FEF8" box="[940,1043,232,256]" captionStart="Figure 1" captionStartId="4.[125,185,1558,1577]" captionTargetBox="[266,1032,179,1519]" captionTargetId="figure-1@4.[266,1033,179,1522]" captionTargetPageId="4" captionText="Figure 1. Photographs (dorsal view) of species of Diahogna. (A) Female of D. martensii (Karsch) from Blackmans Lagoon, Tasmania, Australia (QVMAG 13:42205); (B) female of D. exculta (L. Koch) from OBrians Gap, Wollongong, New South Wales, Australia (AM KS58056); (C) female of D. hildegardae n. sp. from Tambo River at Doctors Flat, Victoria, Australia (WAM T48116); (D) male holotype of D. pisauroides n. sp. from Adelaide River, Northern Territory, Australia (NTMAG A485). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4581379" httpUri="https://zenodo.org/record/4581379/files/figure.png" pageId="6" pageNumber="278">Figures 1</figureCitation>
AD, 2A). The abdomen is uniformly brown to olive-grey, a lanceolate heart mark can be present. Unique within the Australasian
<taxonomicName id="4C764D5C5052B35C849F3331FD20FEC7" authorityName="Sundevall" authorityYear="1833" box="[640,759,295,319]" class="Arachnida" family="Lycosidae" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="278" phylum="Arthropoda" rank="family">Lycosidae</taxonomicName>
, in particular in contrast to the affiliated
<taxonomicName id="4C764D5C5052B35C86E93351FE93FEA7" authorityName="Thorell" authorityYear="1877" box="[246,324,327,351]" class="Arachnida" family="Lycosidae" genus="Artoria" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="278" phylum="Arthropoda" rank="genus">
<emphasis id="B902EACD5052B35C86E93351FE93FEA7" box="[246,324,327,351]" italics="true" pageId="6" pageNumber="278">Artoria</emphasis>
</taxonomicName>
,
<taxonomicName id="4C764D5C5052B35C877E3351FE0BFEA7" authorityName="Roewer" authorityYear="1960" box="[353,476,327,351]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="278" phylum="Arthropoda" rank="genus">
<emphasis id="B902EACD5052B35C877E3351FE0BFEA7" box="[353,476,327,351]" italics="true" pageId="6" pageNumber="278">Tetralycosa</emphasis>
</taxonomicName>
,
<taxonomicName id="4C764D5C5052B35C87E73351FDA5FEA7" authorityName="L. Koch" authorityYear="1878" box="[504,626,327,351]" class="Arachnida" family="Lycosidae" genus="Anoteropsis" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="278" phylum="Arthropoda" rank="genus">
<emphasis id="B902EACD5052B35C87E73351FDA5FEA7" box="[504,626,327,351]" italics="true" pageId="6" pageNumber="278">Anoteropsis</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C764D5C5052B35C84D1335EFCE7FEA7" authorityName="Vink" authorityYear="2002" box="[718,816,328,351]" class="Arachnida" family="Lycosidae" genus="Notocosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="278" phylum="Arthropoda" rank="genus">
<emphasis id="B902EACD5052B35C84D1335EFCE7FEA7" box="[718,816,328,351]" italics="true" pageId="6" pageNumber="278">Notocosa</emphasis>
</taxonomicName>
, the tegulum of the male pedipalp is drawn out apically, i.e. it has a distinct apical tip (
<figureCitation id="134D2A5A5052B35C85773371FC3FFE87" box="[872,1000,359,383]" captionStart="Figure 2" captionStartId="5.[138,198,1584,1603]" captionTargetBox="[138,1177,185,1548]" captionTargetId="figure-0@5.[138,1185,183,1548]" captionTargetPageId="5" captionText="Figure 2. Male of Diahogna martensii (Karsch) from Cape Naturaliste, Tasmania, Australia (QVMAG 13:42218). (A, B) Carapace profile, dorsal and lateral views, respectively; (C) eye pattern in frontal view; (D) left male pedipalp, apical part of bulbus; (E, F) left male pedipalp, ventral and lateral views, respectively. Scale bar: 3.93 mm (A); 4.17 mm (B); 1.16 mm (C); 1.04 mm (D); 0.77 mm (E, F)." figureDoi="http://doi.org/10.5281/zenodo.4581381" httpUri="https://zenodo.org/record/4581381/files/figure.png" pageId="6" pageNumber="278">Figures 2E</figureCitation>
,
<figureCitation id="134D2A5A5052B35C85E73371FBCBFE87" box="[1016,1052,359,383]" captionStart="Figure 5" captionStartId="12.[125,186,1554,1573]" captionTargetBox="[125,1172,351,1513]" captionTargetId="figure-44@12.[125,1174,351,1518]" captionTargetPageId="12" captionText="Figure 5. Diahogna exculta (L. Koch), male from Noumea, New Caledonia (MHNP 3877) and from OBrians Gap, Wollongong, New South Wales, Australia (AM KS58056). (A, B) Left pedipalp, ventral and lateral views, respectively; (C) left male pedipalp, apical part of bulbus; (D, E) female epigyne, ventral and dorsal views, respectively. Scale bar: 0.81 mm (A, B); 0.63 mm (C); 0.67 mm (D, E)." figureDoi="http://doi.org/10.5281/zenodo.4581387" httpUri="https://zenodo.org/record/4581387/files/figure.png" pageId="6" pageNumber="278">5A</figureCitation>
,
<figureCitation id="134D2A5A5052B35C82333371FB86FE87" box="[1068,1105,359,383]" captionStart="Figure 7" captionStartId="17.[138,198,1611,1630]" captionTargetBox="[152,1185,895,1575]" captionTargetId="figure-130@17.[138,1187,895,1575]" captionTargetPageId="17" captionText="Figure 7. Diahogna pisauroides n. sp., male holotype from Adelaide River, Northern Territory, Australia (NTMAG A485). (A, B) Left pedipalp, ventral and lateral views, respectively; (C) left male pedipalp, apical part of bulbus. Scale bar: 0.68 mm (A, B); 0.37 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.4581391" httpUri="https://zenodo.org/record/4581391/files/figure.png" pageId="6" pageNumber="278">7A</figureCitation>
, 9A). The tegular apophysis points apically or retrolaterally and has a truncated tip. The epigyne forms a heavily sclerotised plate without a clearly distinguishable median septum.
</paragraph>
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<paragraph id="8BC936DF5050B35E86623400FCF5F96B" blockId="4.[125,1172,1558,1683]" pageId="4" pageNumber="276">
Figure 1. Photographs (dorsal view) of species of
<taxonomicName id="4C764D5C5050B35E84553400FD77F9D1" authorityName="Roewer" authorityYear="1960" box="[586,672,1558,1577]" class="Arachnida" family="Lycosidae" genus="Diahogna" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="276" phylum="Arthropoda" rank="genus">
<emphasis id="B902EACD5050B35E84553400FD77F9D1" box="[586,672,1558,1577]" italics="true" pageId="4" pageNumber="276">Diahogna</emphasis>
</taxonomicName>
. (A) Female of
<taxonomicName id="4C764D5C5050B35E852B3400FC20F9D1" authority="(Karsch)" baseAuthorityName="Karsch" box="[820,1015,1558,1577]" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="276" phylum="Arthropoda" rank="species" species="martensii">
<emphasis id="B902EACD5050B35E852B3400FC77F9D1" box="[820,928,1558,1577]" italics="true" pageId="4" pageNumber="276">D. martensii</emphasis>
(Karsch)
</taxonomicName>
from Blackmans Lagoon, Tasmania, Australia (QVMAG 13:42205); (B) female of
<taxonomicName id="4C764D5C5050B35E851E3427FC1EF9BC" authority="(L. Koch)" baseAuthorityName="L. Koch" box="[769,969,1585,1604]" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="276" phylum="Arthropoda" rank="species" species="exculta">
<emphasis id="B902EACD5050B35E851E3427FC89F9BC" box="[769,862,1585,1604]" italics="true" pageId="4" pageNumber="276">D. exculta</emphasis>
(L. Koch)
</taxonomicName>
from OBrians Gap, Wollongong, New South Wales, Australia (AM KS58056); (C) female of
<taxonomicName id="4C764D5C5050B35E8538345DFC70F9A6" authority="Framenau, 2006" authorityName="Framenau" authorityYear="2006" box="[807,935,1611,1630]" class="Arachnida" family="Lycosidae" genus="Diahogna" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="276" phylum="Arthropoda" rank="species" species="hildegardae" status="sp. nov.">
<emphasis id="B902EACD5050B35E8538345DFC70F9A6" box="[807,935,1611,1630]" italics="true" pageId="4" pageNumber="276">D. hildegardae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157B65050B35E85B2345DFC36F9A6" box="[941,993,1611,1630]" pageId="4" pageNumber="276" rank="species">n. sp.</taxonomicNameLabel>
from Tambo River at Doctors Flat, Victoria, Australia (WAM T48116); (D) male holotype of
<taxonomicName id="4C764D5C5050B35E85533470FC1EF981" authority="Framenau, 2006" authorityName="Framenau" authorityYear="2006" box="[844,969,1638,1657]" class="Arachnida" family="Lycosidae" genus="Diahogna" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="276" phylum="Arthropoda" rank="species" species="pisauroides" status="sp. nov.">
<emphasis id="B902EACD5050B35E85533470FC1EF981" box="[844,969,1638,1657]" italics="true" pageId="4" pageNumber="276">D. pisauroides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157B65050B35E85CE3470FBD0F981" box="[977,1031,1638,1657]" pageId="4" pageNumber="276" rank="species">n. sp.</taxonomicNameLabel>
from Adelaide River, Northern Territory, Australia (NTMAG A485). Scale bar: 1 mm.
</paragraph>
</caption>
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<paragraph id="8BC936DF5051B35F86953426FD34F96B" blockId="5.[138,1186,1584,1683]" pageId="5" pageNumber="277">
Figure 2. Male of
<taxonomicName id="4C764D5C5051B35F872A3426FDE0F9BB" authority="(Karsch)" baseAuthorityName="Karsch" baseAuthorityYear="1878" box="[309,567,1584,1603]" class="Arachnida" family="Lycosidae" genus="Diahogna" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="277" phylum="Arthropoda" rank="species" species="martensii">
<emphasis id="B902EACD5051B35F872A3426FE37F9BB" box="[309,480,1584,1603]" italics="true" pageId="5" pageNumber="277">Diahogna martensii</emphasis>
(Karsch)
</taxonomicName>
from Cape Naturaliste, Tasmania, Australia (QVMAG 13:42218). (A, B) Carapace profile, dorsal and lateral views, respectively; (C) eye pattern in frontal view; (D) left male pedipalp, apical part of bulbus; (E, F) left male pedipalp, ventral and lateral views, respectively. Scale bar: 3.93 mm (A); 4.17 mm (B); 1.16 mm (C); 1.04 mm (D); 0.77 mm (E, F).
</paragraph>
</caption>
<paragraph id="8BC936DF5052B35C86623032FF22FDC4" blockId="6.[125,245,548,572]" box="[125,245,548,572]" pageId="6" pageNumber="278">
<emphasis id="B902EACD5052B35C86623032FF22FDC4" box="[125,245,548,572]" italics="true" pageId="6" pageNumber="278">Description</emphasis>
</paragraph>
<paragraph id="8BC936DF5052B35C86623042FEC4FBF2" blockId="6.[125,1172,596,1257]" pageId="6" pageNumber="278">
Medium-sized wolf spiders (TL ca 6.014.0 mm). Males smaller than females. Carapace longer than wide, dorsal profile straight in lateral view (
<figureCitation id="134D2A5A5052B35C85193065FCA9FD73" box="[774,894,627,651]" captionStart="Figure 2" captionStartId="5.[138,198,1584,1603]" captionTargetBox="[138,1177,185,1548]" captionTargetId="figure-0@5.[138,1185,183,1548]" captionTargetPageId="5" captionText="Figure 2. Male of Diahogna martensii (Karsch) from Cape Naturaliste, Tasmania, Australia (QVMAG 13:42218). (A, B) Carapace profile, dorsal and lateral views, respectively; (C) eye pattern in frontal view; (D) left male pedipalp, apical part of bulbus; (E, F) left male pedipalp, ventral and lateral views, respectively. Scale bar: 3.93 mm (A); 4.17 mm (B); 1.16 mm (C); 1.04 mm (D); 0.77 mm (E, F)." figureDoi="http://doi.org/10.5281/zenodo.4581381" httpUri="https://zenodo.org/record/4581381/files/figure.png" pageId="6" pageNumber="278">Figure 2B</figureCitation>
). Head flanks in frontal view a gentle slope (
<figureCitation id="134D2A5A5052B35C87773085FE35FD53" box="[360,482,659,683]" captionStart="Figure 2" captionStartId="5.[138,198,1584,1603]" captionTargetBox="[138,1177,185,1548]" captionTargetId="figure-0@5.[138,1185,183,1548]" captionTargetPageId="5" captionText="Figure 2. Male of Diahogna martensii (Karsch) from Cape Naturaliste, Tasmania, Australia (QVMAG 13:42218). (A, B) Carapace profile, dorsal and lateral views, respectively; (C) eye pattern in frontal view; (D) left male pedipalp, apical part of bulbus; (E, F) left male pedipalp, ventral and lateral views, respectively. Scale bar: 3.93 mm (A); 4.17 mm (B); 1.16 mm (C); 1.04 mm (D); 0.77 mm (E, F)." figureDoi="http://doi.org/10.5281/zenodo.4581381" httpUri="https://zenodo.org/record/4581381/files/figure.png" pageId="6" pageNumber="278">Figure 2C</figureCitation>
). Carapace brown, medially lighter (sometimes in the form of a Y-shaped median band) and with marginal or submarginal light bands enhanced by white setae. Anterior median eyes generally larger than anterior lateral eyes (of equal size in male
<taxonomicName id="4C764D5C5052B35C86A230E2FE8EFCF3" authorityName="Framenau" authorityYear="2006" box="[189,345,755,779]" class="Arachnida" family="Lycosidae" genus="Diahogna" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="278" phylum="Arthropoda" rank="species" species="pisauroides">
<emphasis id="B902EACD5052B35C86A230E2FE8EFCF3" box="[189,345,755,779]" italics="true" pageId="6" pageNumber="278">D. pisauroides</emphasis>
</taxonomicName>
), row of anterior eyes wider than row of posterior median eyes; row of anterior eyes straight or slightly procurved. Chelicerae with three promarginal and three (rarely two or four) retromarginal teeth. Labium generally longer than wide (except female
<taxonomicName id="4C764D5C5052B35C86623145FF27FC93" box="[125,240,851,875]" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="278" phylum="Arthropoda" rank="species" species="exculta">
<emphasis id="B902EACD5052B35C86623145FF27FC93" box="[125,240,851,875]" italics="true" pageId="6" pageNumber="278">D. exculta</emphasis>
</taxonomicName>
and male
<taxonomicName id="4C764D5C5052B35C87703145FDDBFC93" authorityName="Framenau" authorityYear="2006" box="[367,524,851,875]" class="Arachnida" family="Lycosidae" genus="Diahogna" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="278" phylum="Arthropoda" rank="species" species="pisauroides">
<emphasis id="B902EACD5052B35C87703145FDDBFC93" box="[367,524,851,875]" italics="true" pageId="6" pageNumber="278">D. pisauroides</emphasis>
</taxonomicName>
). Abdomen olive-grey with light lanceolate heart mark formed by white setae. Leg formula
<collectionCode id="ED67AE1A5052B35C84593164FDB4FC72" box="[582,611,882,906]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="6" pageNumber="278">VI</collectionCode>
.I.II&gt;III (
<collectionCode id="ED67AE1A5052B35C84E83164FCC1FC72" box="[759,790,882,906]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="6" pageNumber="278">VI</collectionCode>
.III.I.II in male
<taxonomicName id="4C764D5C5052B35C82133165FB55FC72" box="[1036,1154,882,906]" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="278" phylum="Arthropoda" rank="species" species="exculta">
<emphasis id="B902EACD5052B35C82133165FB55FC72" box="[1036,1154,882,906]" italics="true" pageId="6" pageNumber="278">D. exculta</emphasis>
</taxonomicName>
). Spination of legs: femur: two or three dorsal, one apicoprolateral, zero or two retrolateral; patella: zero or one prolateral; tibia one to three ventral pairs (basal pairs sometimes reduced to one or no spine), zero or one prolateral; metatarsus: three ventral pairs, one apicoventral.
</paragraph>
<paragraph id="8BC936DF5052B35C86873604FCB7FB11" blockId="6.[125,1172,596,1257]" pageId="6" pageNumber="278">
Tegulum longitudinally divided, retrolateral part narrow in ventral view. Tegular apophysis located retrolaterally at apical tip of tegulum, without basal lobe. Embolus originating prolaterally on palea and curving ventrally around it, long and slender. Basoembolic apophysis forms a white or transparent, membranous lobe. Terminal apophysis with bulging base. Epigyne with a heavily sclerotised plate, with two small (
<taxonomicName id="4C764D5C5052B35C82683684FF1EFB31" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="278" phylum="Arthropoda" rank="species" species="exculta">
<emphasis id="B902EACD5052B35C82683684FF1EFB31" italics="true" pageId="6" pageNumber="278">D. exculta</emphasis>
</taxonomicName>
,
<taxonomicName id="4C764D5C5052B35C86C336A4FEA9FB31" authorityName="Framenau" authorityYear="2006" box="[220,382,1201,1225]" class="Arachnida" family="Lycosidae" genus="Diahogna" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="278" phylum="Arthropoda" rank="species" species="hildegardae">
<emphasis id="B902EACD5052B35C86C336A4FEA9FB31" box="[220,382,1201,1225]" italics="true" pageId="6" pageNumber="278">D. hildegardae</emphasis>
</taxonomicName>
) or a single, wide opening (
<taxonomicName id="4C764D5C5052B35C84C936A4FCB7FB31" baseAuthorityName="Karsch" baseAuthorityYear="1878" box="[726,864,1201,1225]" class="Arachnida" family="Lycosidae" genus="Diahogna" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="278" phylum="Arthropoda" rank="species" species="martensii">
<emphasis id="B902EACD5052B35C84C936A4FCB7FB31" box="[726,864,1201,1225]" italics="true" pageId="6" pageNumber="278">D. martensii</emphasis>
</taxonomicName>
). Spermathecae round or ovoid, copulatory ducts distinctly narrower than spermathecae.
</paragraph>
<paragraph id="8BC936DF5052B35C86623738FF0BFABE" blockId="6.[125,220,1326,1350]" box="[125,220,1326,1350]" pageId="6" pageNumber="278">
<emphasis id="B902EACD5052B35C86623738FF0BFABE" box="[125,220,1326,1350]" italics="true" pageId="6" pageNumber="278">Remarks</emphasis>
</paragraph>
<paragraph id="8BC936DF5052B35C86623748FE11F96D" blockId="6.[125,1173,1374,1685]" pageId="6" pageNumber="278">
A recent molecular phylogeny inferred from 12S rRNA and
<collectionCode id="ED67AE1A5052B35C85203748FC4CFA8E" box="[831,923,1374,1398]" pageId="6" pageNumber="278">NADH</collectionCode>
1 including 70 worldwide lycosid species with the majority from the Australasian region, included
<taxonomicName id="4C764D5C5052B35C85E53769FB43FA6E" authorityName="Framenau" authorityYear="2006" box="[1018,1172,1406,1430]" class="Arachnida" family="Lycosidae" genus="Diahogna" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="278" phylum="Arthropoda" rank="species" species="pisauroides">
<emphasis id="B902EACD5052B35C85E53769FB43FA6E" box="[1018,1172,1406,1430]" italics="true" pageId="6" pageNumber="278">D. pisauroides</emphasis>
</taxonomicName>
(as New Genus
<specimenCount id="9D70FD565052B35C87573788FE55FA4E" box="[328,386,1438,1462]" pageId="6" pageNumber="278" type="generic">2 sp.</specimenCount>
) as representative of
<taxonomicName id="4C764D5C5052B35C84983788FC21FA4E" authority="(Murphy et al. 2006)" baseAuthorityName="Murphy" baseAuthorityYear="2006" box="[647,1014,1438,1462]" class="Arachnida" family="Lycosidae" genus="Diahogna" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="278" phylum="Arthropoda" rank="genus">
<emphasis id="B902EACD5052B35C84983788FD25FA4E" box="[647,754,1438,1462]" italics="true" pageId="6" pageNumber="278">Diahogna</emphasis>
(
<bibRefCitation id="EFE74B2E5052B35C851C3788FC3AFA4E" author="Murphy NP &amp; Framenau VW &amp; Donellan SC &amp; Harvey MS &amp; Park Y-C &amp; Austin AD" box="[771,1005,1438,1462]" pageId="6" pageNumber="278" pagination="583 - 602" refId="ref9468" refString="Murphy NP, Framenau VW, Donellan SC, Harvey MS, Park Y-C, Austin AD. 2006. Phylogenetic reconstruction of the wolf spiders (Araneae: Lycosidae) using sequences from the 12 S rRNA, 28 rRNA and NADH 1 genes: implications for classification, biogeography and the evolution of web building behavior. Molecular Phylogenetics and Evolution 38: 583 - 602." type="journal article" year="2006">Murphy et al. 2006</bibRefCitation>
)
</taxonomicName>
. Independent of the phylogenetic analysis used (parsimony, Bayesian),
<taxonomicName id="4C764D5C5052B35C852D37A8FC06FA2E" authorityName="Framenau" authorityYear="2006" box="[818,977,1470,1494]" class="Arachnida" family="Lycosidae" genus="Diahogna" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="278" phylum="Arthropoda" rank="species" species="pisauroides">
<emphasis id="B902EACD5052B35C852D37A8FC06FA2E" box="[818,977,1470,1494]" italics="true" pageId="6" pageNumber="278">D. pisauroides</emphasis>
</taxonomicName>
represented the sister taxon of
<taxonomicName id="4C764D5C5052B35C872937C8FD04FA0E" authority="(L. Koch, 1876)" baseAuthorityName="L. Koch" baseAuthorityYear="1876" box="[310,723,1502,1526]" class="Arachnida" family="Lycosidae" genus="Tetralycosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="278" phylum="Arthropoda" rank="species" species="oraria">
<emphasis id="B902EACD5052B35C872937C8FE29FA0E" box="[310,510,1502,1526]" italics="true" pageId="6" pageNumber="278">Tetralycosa oraria</emphasis>
(L.
<bibRefCitation id="EFE74B2E5052B35C842437C8FD1BFA0E" author="Koch L" box="[571,716,1502,1526]" pageId="6" pageNumber="278" pagination="741 - 888" refId="ref9268" refString="Koch L. 1876. Die Arachniden Australiens, nach der Natur beschrieben und abgebildet. Nuremburg: Bauer and Raspe. p 741 - 888." type="book chapter" year="1876">Koch, 1876</bibRefCitation>
)
</taxonomicName>
within a distinct Australasian/Pacific clade at the subfamily level. Morphologically, this clade is characterised by the presence of a unique basoembolic apophysis on the male pedipalp and had previously been argued to represent a new lycosid subfamily (
<bibRefCitation id="EFE74B2E5052B35C8442342BFCF4F9AD" author="Framenau VW" box="[605,803,1597,1621]" pageId="6" pageNumber="278" pagination="209 - 235" refId="ref8959" refString="Framenau VW. 2002. Review of the wolf spider genus Artoria Thorell (Araneae, Lycosidae). Invertebrate Systematics 16: 209 - 235." type="journal article" year="2002">Framenau 2002</bibRefCitation>
;
<bibRefCitation id="EFE74B2E5052B35C8528342BFC6CF9AD" author="Vink CJ" box="[823,955,1597,1621]" pageId="6" pageNumber="278" refId="ref9813" refString="Vink CJ. 2002. Lycosidae (Arachnida: Araneae). Lincoln (New Zealand): Manaaki Whenua Press. 94 p. (Fauna of New Zealand; 44)." type="book" year="2002">Vink 2002</bibRefCitation>
; Framenau et al. forthcoming). A formal description of this subfamily is forthcoming as part of my revision of the Australian
<taxonomicName id="4C764D5C5052B35C8755346BFE15F96D" authorityName="Sundevall" authorityYear="1833" box="[330,450,1661,1685]" class="Arachnida" family="Lycosidae" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="278" phylum="Arthropoda" rank="family">Lycosidae</taxonomicName>
.
</paragraph>
<paragraph id="8BC936DF5053B35D8695329FFEDCFF59" blockId="7.[138,267,137,161]" box="[138,267,137,161]" pageId="7" pageNumber="279">
<emphasis id="B902EACD5053B35D8695329FFEDCFF59" box="[138,267,137,161]" italics="true" pageId="7" pageNumber="279">Distribution</emphasis>
</paragraph>
<paragraph id="8BC936DF5053B35D869532AFFE3BFF29" blockId="7.[138,492,185,209]" box="[138,492,185,209]" pageId="7" pageNumber="279">
<collectingCountry id="F361764F5053B35D869532AFFF25FF29" box="[138,242,185,209]" name="Australia" pageId="7" pageNumber="279">Australia</collectingCountry>
and
<collectingCountry id="F361764F5053B35D873032AFFE3FFF29" box="[303,488,185,209]" name="New Caledonia" pageId="7" pageNumber="279">New Caledonia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>