treatments-xml/data/03/D9/29/03D92963143F9C20FEB9F8851B0538C1.xml
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<document id="44A29F58248F8701F10CACC50E9C923D" ID-DOI="10.5281/zenodo.7162582" ID-ISSN="1536-9307" ID-Zenodo-Dep="7162582" ID-ZooBank="48B30467-7DD1-4194-8585-847CC07E15D4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1665231705657" checkinUser="felipe" docAuthor="Kovařík, František &amp; Lowe, Graeme" docDate="2021" docId="03D92963143F9C20FEB9F8851B0538C1" docLanguage="en" docName="Euscorpius.344.1-14.pdf" docOrigin="Euscorpius 344" docStyle="DocumentStyle:5E96F6379CE20B0E8E4F9E7DF8E1C0DF.2:Euscorpius.2020-.journal_article" docStyleId="5E96F6379CE20B0E8E4F9E7DF8E1C0DF" docStyleName="Euscorpius.2020-.journal_article" docStyleVersion="2" docTitle="Lanzatus huluul Kovařík &amp; Lowe 2021, sp. n." docType="treatment" docVersion="3" lastPageNumber="11" masterDocId="FFE0511B143B9C2CFF8BFF881A613E1D" masterDocTitle="Scorpions of the Horn of Africa (Arachnida: Scorpiones) Part XXVII Lanzatus huluul sp n from Somaliland (Buthidae)" masterLastPageNumber="12" masterPageNumber="1" pageNumber="3" updateTime="1699366030113" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="4D37FA5653F89274637C09B5B1F2DD16">Scorpions of the Horn of Africa (Arachnida: Scorpiones) Part XXVII Lanzatus huluul sp n from Somaliland (Buthidae)</mods:title>
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<mods:date id="9B8B255B9044BF84740C0BAC2223A732">2021</mods:date>
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<heading id="D0872F19143F9C28FEB9F8851823393A" box="[306,578,1804,1831]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="3" reason="2">
<taxonomicName id="4C70E3F6143F9C28FEB9F8851B9B393A" ID-CoL="992Q5" authority="Kovařík &amp; Lowe, 2021" authorityName="Kovařík &amp; Lowe" authorityYear="2021" box="[306,506,1804,1831]" class="Arachnida" family="Buthidae" genus="Lanzatus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="huluul" status="sp. nov.">Lanzatus huluul</taxonomicName>
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.
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.
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(
<figureCitation id="134B84F0143F9C28FE96F8A61BCC395B" box="[285,429,1838,1862]" captionStart-0="Figures 12" captionStart-1="Figures 36" captionStart-2="Figures 716" captionStart-3="Figures 1737" captionStart-4="Figures 3845" captionStart-5="Figures 4651" captionStartId-0="3.[116,193,1967,1989]" captionStartId-1="5.[115,192,2016,2038]" captionStartId-2="6.[115,192,1960,1982]" captionStartId-3="7.[114,191,1898,1920]" captionStartId-4="8.[115,192,1948,1970]" captionStartId-5="9.[115,192,1066,1088]" captionTargetBox-0="[113,1474,225,1960]" captionTargetBox-1="[133,1455,229,2008]" captionTargetBox-2="[113,1467,243,1945]" captionTargetBox-3="[116,1459,237,1887]" captionTargetBox-4="[115,1476,242,1931]" captionTargetBox-5="[126,1464,253,1043]" captionTargetId-0="figure-58@3.[113,1474,225,1962]" captionTargetId-1="figure-53@5.[113,1474,223,2012]" captionTargetId-2="figure-98@6.[113,1474,234,1948]" captionTargetId-3="figure-126@7.[116,1477,234,1890]" captionTargetId-4="figure-73@8.[115,1476,242,1934]" captionTargetId-5="figure-519@9.[113,1474,234,1055]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="9" captionText-0="Figures 12. Lanzatus specimens in vivo habitus. Figure 1. Lanzatus somalilandus Kovařík et Lowe, 2016, male from Somaliland, Shanshade village. Figure 2. Lanzatus huluul sp. n., female paratYpe. Images taken in the laboratorY with different sand/background than in realitY is on the original localities." captionText-1="Figures 36: Lanzatus huluul sp. n., habitus. Figures 34. Male holotYpe in dorsal (3) and ventral (4) views. Figures 56. Female paratYpe in dorsal (5) and ventral (6) views. Scale bar: 5 mm." captionText-2="Figures 716: Lanzatus huluul sp. n., metasoma and telson. Figures 711. HolotYpe male, metasoma V and telson (7) and telson (8) in lateral views and metasoma and telson in lateral (9), dorsal (10), and ventral (11) views. Figures 1216. ParatYpe female, metasoma V and telson (12) and telson (13) in lateral views and metasoma and telson in lateral (14), dorsal (15), and ventral (16) views.Scale bars: 5 mm (911 and 1416)." captionText-3="Figures 1737: Lanzatus huluul sp. n., right pedipalp. Figures 1726. HolotYpe male, chela in dorsal (17), external (18) and ventral (19) views, patella in dorsal (20), external (21) and ventral (22) views, femur and trochanter in dorsal (23), internal (24) and ventral (25) views, dentate margins of movable finger (26). Figures 2737. ParatYpe female, chela in dorsal (27), external (28) and ventral (29) views, patella in dorsal (30), external (31) and ventral (32) views, femur and trochanter in dorsal (33), internal (34) and ventral (35) views. dentate margins of movable (36) and fixed (37) fingers. Trichobothrial pattern indicated in Figures 2731 and 3334 bY white circles." captionText-4="Figures 3845: Lanzatus huluul sp. n. Figures 3839, 4245. HolotYpe male, carapace and tergites IIV (38), sternopectinal area and sternites (39), and left legs IIV, retrolateral aspect (4245). Figures 4041. ParatYpe female, carapace and tergites IV (40) and sternopectinal area and sternites (41)." captionText-5="Figures 4651: Comparison of telsons of Lanzatus species. Figures 4647. Lanzatus huluul sp. n., female paratYpe (46) and male holotYpe (47). Figures 48. Lanzatus somalicus Kovařík, 2001, male paratYpe. Figures 4951. Lanzatus somalilandus Kovařík et Lowe, 2016, female paratYpe (49), male holotYpe (50) and male from Somaliland, Shanshade Village. (51)." figureDoi-0="http://doi.org/10.5281/zenodo.7162584" figureDoi-1="http://doi.org/10.5281/zenodo.7162588" figureDoi-2="http://doi.org/10.5281/zenodo.7162590" figureDoi-3="http://doi.org/10.5281/zenodo.7162592" figureDoi-4="http://doi.org/10.5281/zenodo.7162594" figureDoi-5="http://doi.org/10.5281/zenodo.7162596" httpUri-0="https://zenodo.org/record/7162584/files/figure.png" httpUri-1="https://zenodo.org/record/7162588/files/figure.png" httpUri-2="https://zenodo.org/record/7162590/files/figure.png" httpUri-3="https://zenodo.org/record/7162592/files/figure.png" httpUri-4="https://zenodo.org/record/7162594/files/figure.png" httpUri-5="https://zenodo.org/record/7162596/files/figure.png" pageId="4" pageNumber="3">Figures 247</figureCitation>
,
<figureCitation id="134B84F0143F9C28FE33F8A61B9C395B" box="[440,509,1838,1862]" captionStart-0="Figures 5253" captionStart-1="Figures 5455" captionStartId-0="10.[111,188,1981,2003]" captionStartId-1="11.[115,192,1982,2004]" captionTargetBox-0="[212,1338,238,1965]" captionTargetBox-1="[115,1476,233,1977]" captionTargetId-0="figure-48@10.[144,1458,223,1974]" captionTargetId-1="figure-43@11.[115,1476,230,1979]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="Figures 5253: Lanzatus huluul sp. n., left hemispermatophore. HolotYpe male, whole hemispermatophore (52) and capsule, compressed to separate lobes (53), convex views. Scale bars: 1 mm (52), 200 μm (53)." captionText-1="Figures 5455: Figure 54. Map showing known distribution of the genus Lanzatus. Figure 55. Somaliland, vicinitY of Huluul Village, tYpe localitY of Lanzatus huluul sp. n." figureDoi-0="http://doi.org/10.5281/zenodo.7162598" figureDoi-1="http://doi.org/10.5281/zenodo.7162600" httpUri-0="https://zenodo.org/record/7162598/files/figure.png" httpUri-1="https://zenodo.org/record/7162600/files/figure.png" pageId="4" pageNumber="3">5254</figureCitation>
,
<tableCitation id="C6F2ADCE143F9C28FD8CF8A61834395B" box="[519,597,1838,1862]" captionStart="Table 1" captionStartId="4.[110,166,991,1012]" captionText="Table 1. Comparative measurements of tYpes of Lanzatus huluul sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="4" pageNumber="3">Table 1</tableCitation>
)
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<location id="8EAFCEAE143F9C28FD87F82318EC39DE" LSID="urn:lsid:plazi:treatment:03D92963143F9C20FEB9F8851B0538C1:8EAFCEAE143F9C28FD87F82318EC39DE" box="[524,653,1963,1987]" latitude="9.977614" longLatPrecision="5" longitude="46.6932" municipality="Huluul Village" name="Somaliland" pageId="4" pageNumber="3">Somaliland</location>
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a. s. l.
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(LocalitY No.
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(LocalitY No. 21SI),
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<specimenCount id="9D7653FC143F9C28FCBBFBE319333A9E" box="[816,850,1131,1155]" pageId="4" pageNumber="3" type="male">1♂</specimenCount>
(
<typeStatus id="54CB26D7143F9C28FCD5FBE319A13A9E" box="[862,960,1131,1155]" pageId="4" pageNumber="3" type="holotype">holotYpe</typeStatus>
, DNA No. 1998)
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(
<typeStatus id="54CB26D7143F9C28FB23FBE31F683A9E" box="[1192,1289,1131,1155]" pageId="4" pageNumber="3" type="paratype">paratYpe</typeStatus>
, DNA No. 2035), leg.
<collectorName id="2685FDA3143F9C28FCD5FB0319B33ABE" box="[862,978,1163,1187]" pageId="4" pageNumber="3">F. Kovařík</collectorName>
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<subSubSection id="C36ACBFE143F9C28FCBBFB431F383AFF" box="[816,1369,1227,1251]" pageId="4" pageNumber="3" type="etymology">
<paragraph id="8BCF9875143F9C28FCBBFB431F383AFF" blockId="4.[816,1369,1227,1251]" box="[816,1369,1227,1251]" pageId="4" pageNumber="3">ETYMOLOGY. Named after the village of occurrence.</paragraph>
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<subSubSection id="C36ACBFE143F9C28FCBBFA831EE6389E" pageId="4" pageNumber="3" type="diagnosis">
<paragraph id="8BCF9875143F9C28FCBBFA831EE6389E" blockId="4.[816,1475,1291,1667]" pageId="4" pageNumber="3">
DIAGNOSIS. Total length 20.7 (male)
<quantity id="4C883590143F9C28FB64FA831F353B3F" box="[1263,1364,1291,1315]" metricMagnitude="-2" metricUnit="m" metricValue="2.11" pageId="4" pageNumber="3" unit="mm" value="21.1">21.1 mm</quantity>
(female). Base color uniformlY Yellowish orange with black onlY around the eYes. Fifth metasomal segment slightlY marbled in graYish black. Pectine teeth number
<quantity id="4C883590143F9C28FB54FAE31F213B9E" box="[1247,1344,1387,1411]" metricMagnitude="-1" metricUnit="m" metricValue="5.461" metricValueMax="5.588" metricValueMin="5.334" pageId="4" pageNumber="3" unit="in" value="21.5" valueMax="22.0" valueMin="21.0">2122 in</quantity>
both sexes. Pedipalp chela and patella with inconspicuous smooth carinae. Pedipalp chela smooth and narrow. Sternites finelY granulated, without carinae. Tergites IVI finelY granulated with one carina present or indicated. Metasomal segments smooth to finelY granulated, without complete carinae, with rounded edges. Telson extremelY elongated, smooth, without aculear ring. Telson length/ width ratio
<quantity id="4C883590143F9C28FB54F9C31F4A387E" box="[1247,1323,1611,1635]" metricMagnitude="-1" metricUnit="m" metricValue="1.52908" pageId="4" pageNumber="3" unit="in" value="6.02">6.02 in</quantity>
male,
<quantity id="4C883590143F9C28FAFDF9C31FA3387E" box="[1398,1474,1611,1635]" metricMagnitude="-1" metricUnit="m" metricValue="1.35382" pageId="4" pageNumber="3" unit="in" value="5.33">5.33 in</quantity>
female. Aculeus weaklY curved.
</paragraph>
</subSubSection>
<subSubSection id="C36ACBFE143F9C20FCBBF9231B0538C1" lastPageId="12" lastPageNumber="11" pageId="4" pageNumber="3" type="description">
<paragraph id="8BCF9875143F9C28FCBBF9231E55397E" blockId="4.[816,1475,1707,2051]" pageId="4" pageNumber="3">
DESCRIPTION. The adults are 20.7 (male)
<quantity id="4C883590143F9C28FA8AF9231F0138DF" box="[1281,1376,1707,1731]" metricMagnitude="-2" metricUnit="m" metricValue="2.11" pageId="4" pageNumber="3" unit="mm" value="21.1">21.1 mm</quantity>
(female) long. The habitus is shown in
<figureCitation id="134B84F0143F9C28FB03F9431E9238FE" box="[1160,1267,1739,1763]" captionStart-0="Figures 12" captionStart-1="Figures 36" captionStartId-0="3.[116,193,1967,1989]" captionStartId-1="5.[115,192,2016,2038]" captionTargetBox-0="[113,1474,225,1960]" captionTargetBox-1="[133,1455,229,2008]" captionTargetId-0="figure-58@3.[113,1474,225,1962]" captionTargetId-1="figure-53@5.[113,1474,223,2012]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionText-0="Figures 12. Lanzatus specimens in vivo habitus. Figure 1. Lanzatus somalilandus Kovařík et Lowe, 2016, male from Somaliland, Shanshade village. Figure 2. Lanzatus huluul sp. n., female paratYpe. Images taken in the laboratorY with different sand/background than in realitY is on the original localities." captionText-1="Figures 36: Lanzatus huluul sp. n., habitus. Figures 34. Male holotYpe in dorsal (3) and ventral (4) views. Figures 56. Female paratYpe in dorsal (5) and ventral (6) views. Scale bar: 5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7162584" figureDoi-1="http://doi.org/10.5281/zenodo.7162588" httpUri-0="https://zenodo.org/record/7162584/files/figure.png" httpUri-1="https://zenodo.org/record/7162588/files/figure.png" pageId="4" pageNumber="3">Figs. 26</figureCitation>
. For positions and distribution of trichobothria of pedipalps see
<figureCitation id="134B84F0143F9C28FA9DF9631917393E" captionStart="Figures 1737" captionStartId="7.[114,191,1898,1920]" captionTargetBox="[116,1459,237,1887]" captionTargetId="figure-126@7.[116,1477,234,1890]" captionTargetPageId="7" captionText="Figures 1737: Lanzatus huluul sp. n., right pedipalp. Figures 1726. HolotYpe male, chela in dorsal (17), external (18) and ventral (19) views, patella in dorsal (20), external (21) and ventral (22) views, femur and trochanter in dorsal (23), internal (24) and ventral (25) views, dentate margins of movable finger (26). Figures 2737. ParatYpe female, chela in dorsal (27), external (28) and ventral (29) views, patella in dorsal (30), external (31) and ventral (32) views, femur and trochanter in dorsal (33), internal (34) and ventral (35) views. dentate margins of movable (36) and fixed (37) fingers. Trichobothrial pattern indicated in Figures 2731 and 3334 bY white circles." figureDoi="http://doi.org/10.5281/zenodo.7162592" httpUri="https://zenodo.org/record/7162592/files/figure.png" pageId="4" pageNumber="3">Figs. 2731 and 3334</figureCitation>
. Sexual dimorphism: adult male integument matte, female glossY, pedipalp patella narrower in male (
<figureCitation id="134B84F0143F9C28FAE3F8A31FDB395E" box="[1384,1466,1835,1859]" captionStart="Figures 1737" captionStartId="7.[114,191,1898,1920]" captionTargetBox="[116,1459,237,1887]" captionTargetId="figure-126@7.[116,1477,234,1890]" captionTargetPageId="7" captionText="Figures 1737: Lanzatus huluul sp. n., right pedipalp. Figures 1726. HolotYpe male, chela in dorsal (17), external (18) and ventral (19) views, patella in dorsal (20), external (21) and ventral (22) views, femur and trochanter in dorsal (23), internal (24) and ventral (25) views, dentate margins of movable finger (26). Figures 2737. ParatYpe female, chela in dorsal (27), external (28) and ventral (29) views, patella in dorsal (30), external (31) and ventral (32) views, femur and trochanter in dorsal (33), internal (34) and ventral (35) views. dentate margins of movable (36) and fixed (37) fingers. Trichobothrial pattern indicated in Figures 2731 and 3334 bY white circles." figureDoi="http://doi.org/10.5281/zenodo.7162592" httpUri="https://zenodo.org/record/7162592/files/figure.png" pageId="4" pageNumber="3">Fig. 20</figureCitation>
) than in female (
<figureCitation id="134B84F0143F9C28FC53F8C31E46397E" box="[984,1063,1867,1891]" captionStart="Figures 1737" captionStartId="7.[114,191,1898,1920]" captionTargetBox="[116,1459,237,1887]" captionTargetId="figure-126@7.[116,1477,234,1890]" captionTargetPageId="7" captionText="Figures 1737: Lanzatus huluul sp. n., right pedipalp. Figures 1726. HolotYpe male, chela in dorsal (17), external (18) and ventral (19) views, patella in dorsal (20), external (21) and ventral (22) views, femur and trochanter in dorsal (23), internal (24) and ventral (25) views, dentate margins of movable finger (26). Figures 2737. ParatYpe female, chela in dorsal (27), external (28) and ventral (29) views, patella in dorsal (30), external (31) and ventral (32) views, femur and trochanter in dorsal (33), internal (34) and ventral (35) views. dentate margins of movable (36) and fixed (37) fingers. Trichobothrial pattern indicated in Figures 2731 and 3334 bY white circles." figureDoi="http://doi.org/10.5281/zenodo.7162592" httpUri="https://zenodo.org/record/7162592/files/figure.png" pageId="4" pageNumber="3">Fig. 30</figureCitation>
).
</paragraph>
<paragraph id="8BCF9875143F9C28FCBBF8E31EC0361E" blockId="4.[816,1475,1707,2051]" pageId="4" pageNumber="3">
<emphasis id="B9044467143F9C28FCBBF8E319CA399E" bold="true" box="[816,939,1899,1923]" pageId="4" pageNumber="3">Coloration</emphasis>
(
<figureCitation id="134B84F0143F9C28FC32F8E31E41399E" box="[953,1056,1899,1923]" captionStart-0="Figures 12" captionStart-1="Figures 36" captionStartId-0="3.[116,193,1967,1989]" captionStartId-1="5.[115,192,2016,2038]" captionTargetBox-0="[113,1474,225,1960]" captionTargetBox-1="[133,1455,229,2008]" captionTargetId-0="figure-58@3.[113,1474,225,1962]" captionTargetId-1="figure-53@5.[113,1474,223,2012]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionText-0="Figures 12. Lanzatus specimens in vivo habitus. Figure 1. Lanzatus somalilandus Kovařík et Lowe, 2016, male from Somaliland, Shanshade village. Figure 2. Lanzatus huluul sp. n., female paratYpe. Images taken in the laboratorY with different sand/background than in realitY is on the original localities." captionText-1="Figures 36: Lanzatus huluul sp. n., habitus. Figures 34. Male holotYpe in dorsal (3) and ventral (4) views. Figures 56. Female paratYpe in dorsal (5) and ventral (6) views. Scale bar: 5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7162584" figureDoi-1="http://doi.org/10.5281/zenodo.7162588" httpUri-0="https://zenodo.org/record/7162584/files/figure.png" httpUri-1="https://zenodo.org/record/7162588/files/figure.png" pageId="4" pageNumber="3">Figs. 26</figureCitation>
). The base color is uniformlY Yellowish orange with black onlY around the eYes, tergites darker. Tarsomeres of legs white. Fifth metasomal segment slightlY marbled in graYish black on anterior 3/5. Chelicerae Yellow with orange reticulation in female.
</paragraph>
<caption id="DF0FC8FD143E9C29FFF8F8681831360E" ID-DOI="http://doi.org/10.5281/zenodo.7162588" ID-Zenodo-Dep="7162588" httpUri="https://zenodo.org/record/7162588/files/figure.png" pageId="5" pageNumber="4" startId="5.[115,192,2016,2038]" targetBox="[133,1455,229,2008]" targetPageId="5">
<paragraph id="8BCF9875143E9C29FFF8F8681831360E" blockId="5.[115,1473,2016,2067]" pageId="5" pageNumber="4">
<emphasis id="B9044467143E9C29FFF8F8681A8839EB" bold="true" box="[115,233,2016,2038]" pageId="5" pageNumber="4">Figures 36</emphasis>
:
<taxonomicName id="4C70E3F6143E9C29FF7EF8691BEF39EB" authority="Kovařík &amp; Lowe, 2021" authorityName="Kovařík &amp; Lowe" authorityYear="2021" box="[245,398,2017,2038]" class="Arachnida" family="Buthidae" genus="Lanzatus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="huluul" status="sp. nov.">
<emphasis id="B9044467143E9C29FF7EF8691BEF39EB" box="[245,398,2017,2038]" italics="true" pageId="5" pageNumber="4">Lanzatus huluul</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A237F91C143E9C29FE1FF8691BA839EB" box="[404,457,2017,2038]" pageId="5" pageNumber="4" rank="species">
<emphasis id="B9044467143E9C29FE1FF8691BCA39EB" bold="true" box="[404,427,2017,2038]" pageId="5" pageNumber="4">sp</emphasis>
.
<emphasis id="B9044467143E9C29FE3CF8691BA539EB" bold="true" box="[439,452,2017,2038]" pageId="5" pageNumber="4">n</emphasis>
.
</taxonomicNameLabel>
, habitus.
<emphasis id="B9044467143E9C29FDADF86818FC39EB" bold="true" box="[550,669,2016,2038]" pageId="5" pageNumber="4">Figures 34</emphasis>
. Male holotYpe in dorsal (3) and ventral (4) views.
<emphasis id="B9044467143E9C29FB03F8681E9F39EB" bold="true" box="[1160,1278,2016,2038]" pageId="5" pageNumber="4">Figures 56</emphasis>
. Female paratYpe in dorsal (5) and ventral (6) views. Scale bar: 5 mm.
</paragraph>
</caption>
<caption id="DF0FC8FD143D9C2AFFF8F8201FA339E5" ID-DOI="http://doi.org/10.5281/zenodo.7162590" ID-Zenodo-Dep="7162590" httpUri="https://zenodo.org/record/7162590/files/figure.png" pageId="6" pageNumber="5" startId="6.[115,192,1960,1982]" targetBox="[113,1467,243,1945]" targetPageId="6">
<paragraph id="8BCF9875143D9C2AFFF8F8201FA339E5" blockId="6.[115,1474,1960,2040]" pageId="6" pageNumber="5">
<emphasis id="B9044467143D9C2AFFF8F8201A9439A0" bold="true" box="[115,245,1960,1982]" pageId="6" pageNumber="5">Figures 716</emphasis>
:
<taxonomicName id="4C70E3F6143D9C2AFE89F8211BFA39A0" authority="Kovařík &amp; Lowe, 2021" authorityName="Kovařík &amp; Lowe" authorityYear="2021" box="[258,411,1960,1982]" class="Arachnida" family="Buthidae" genus="Lanzatus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="huluul" status="sp. nov.">
<emphasis id="B9044467143D9C2AFE89F8211BFA39A0" box="[258,411,1960,1982]" italics="true" pageId="6" pageNumber="5">Lanzatus huluul</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A237F91C143D9C2AFE2AF8201BB739A0" box="[417,470,1960,1981]" pageId="6" pageNumber="5" rank="species">
<emphasis id="B9044467143D9C2AFE2AF8201BD939A0" bold="true" box="[417,440,1960,1981]" pageId="6" pageNumber="5">sp</emphasis>
.
<emphasis id="B9044467143D9C2AFE48F8201BB139A0" bold="true" box="[451,464,1960,1981]" pageId="6" pageNumber="5">n</emphasis>
.
</taxonomicNameLabel>
, metasoma and telson.
<emphasis id="B9044467143D9C2AFD3CF820195939A0" bold="true" box="[695,824,1960,1982]" pageId="6" pageNumber="5">Figures 711</emphasis>
. HolotYpe male, metasoma V and telson (7) and telson (8) in lateral views and metasoma and telson in lateral (9), dorsal (10), and ventral (11) views.
<emphasis id="B9044467143D9C2AFCFCF84D1E6439C7" bold="true" box="[887,1029,1989,2011]" pageId="6" pageNumber="5">Figures 1216</emphasis>
. ParatYpe female, metasoma V and telson (12) and telson (13) in lateral views and metasoma and telson in lateral (14), dorsal (15), and ventral (16) views. Scale bars: 5 mm (911 and 1416).
</paragraph>
</caption>
<caption id="DF0FC8FD143C9C2BFFF9F8E21EDA39E9" ID-DOI="http://doi.org/10.5281/zenodo.7162592" ID-Zenodo-Dep="7162592" httpUri="https://zenodo.org/record/7162592/files/figure.png" pageId="7" pageNumber="6" startId="7.[114,191,1898,1920]" targetBox="[116,1459,237,1887]" targetPageId="7">
<paragraph id="8BCF9875143C9C2BFFF9F8E21EDA39E9" blockId="7.[114,1473,1898,2036]" pageId="7" pageNumber="6">
<emphasis id="B9044467143C9C2BFFF9F8E21B623962" bold="true" box="[114,259,1898,1920]" pageId="7" pageNumber="6">Figures 1737</emphasis>
:
<taxonomicName id="4C70E3F6143C9C2BFE99F8E31BCC3962" authority="Kovařík &amp; Lowe, 2021" authorityName="Kovařík &amp; Lowe" authorityYear="2021" box="[274,429,1898,1920]" class="Arachnida" family="Buthidae" genus="Lanzatus" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="huluul" status="sp. nov.">
<emphasis id="B9044467143C9C2BFE99F8E31BCC3962" box="[274,429,1898,1920]" italics="true" pageId="7" pageNumber="6">Lanzatus huluul</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A237F91C143C9C2BFE3DF8E31B8C399D" box="[438,493,1899,1920]" pageId="7" pageNumber="6" rank="species">
<emphasis id="B9044467143C9C2BFE3DF8E31BAC399D" bold="true" box="[438,461,1899,1920]" pageId="7" pageNumber="6">sp</emphasis>
.
<emphasis id="B9044467143C9C2BFE50F8E31B89399D" bold="true" box="[475,488,1899,1920]" pageId="7" pageNumber="6">n</emphasis>
.
</taxonomicNameLabel>
, right pedipalp.
<emphasis id="B9044467143C9C2BFD19F8E219423962" bold="true" box="[658,803,1898,1920]" pageId="7" pageNumber="6">Figures 1726</emphasis>
. HolotYpe male, chela in dorsal (17), external (18) and ventral (19) views, patella in dorsal (20), external (21) and ventral (22) views, femur and trochanter in dorsal (23), internal (24) and ventral (25) views, dentate margins of movable finger (26).
<emphasis id="B9044467143C9C2BFE7CF82C18E739A4" bold="true" box="[503,646,1956,1978]" pageId="7" pageNumber="6">Figures 2737</emphasis>
. ParatYpe female, chela in dorsal (27), external (28) and ventral (29) views, patella in dorsal (30), external (31) and ventral (32) views, femur and trochanter in dorsal (33), internal (34) and ventral (35) views. dentate margins of movable (36) and fixed (37) fingers. Trichobothrial pattern indicated in Figures 2731 and 3334 bY white circles.
</paragraph>
</caption>
<caption id="DF0FC8FD14339C24FFF8F8141B4139F6" ID-DOI="http://doi.org/10.5281/zenodo.7162594" ID-Zenodo-Dep="7162594" httpUri="https://zenodo.org/record/7162594/files/figure.png" pageId="8" pageNumber="7" startId="8.[115,192,1948,1970]" targetBox="[115,1476,242,1931]" targetPageId="8">
<paragraph id="8BCF987514339C24FFF8F8141B4139F6" blockId="8.[115,1473,1948,2028]" pageId="8" pageNumber="7">
<emphasis id="B904446714339C24FFF8F8141B6039AC" bold="true" box="[115,257,1948,1970]" pageId="8" pageNumber="7">Figures 3845</emphasis>
:
<taxonomicName id="4C70E3F614339C24FE86F8151BC739AF" authority="Kovařík &amp; Lowe, 2021" authorityName="Kovařík &amp; Lowe" authorityYear="2021" box="[269,422,1949,1970]" class="Arachnida" family="Buthidae" genus="Lanzatus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="huluul" status="sp. nov.">
<emphasis id="B904446714339C24FE86F8151BC739AF" box="[269,422,1949,1970]" italics="true" pageId="8" pageNumber="7">Lanzatus huluul</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A237F91C14339C24FE20F8151B8139AF" box="[427,480,1949,1970]" pageId="8" pageNumber="7" rank="species">
<emphasis id="B904446714339C24FE20F8151BA339AF" bold="true" box="[427,450,1949,1970]" pageId="8" pageNumber="7">sp</emphasis>
.
<emphasis id="B904446714339C24FE46F8151BBB39AF" bold="true" box="[461,474,1949,1970]" pageId="8" pageNumber="7">n</emphasis>
.
</taxonomicNameLabel>
<emphasis id="B904446714339C24FE6EF814181539AC" bold="true" box="[485,628,1948,1970]" pageId="8" pageNumber="7">Figures 3839</emphasis>
,
<emphasis id="B904446714339C24FDF4F81418DA39AC" bold="true" box="[639,699,1948,1970]" pageId="8" pageNumber="7">4245</emphasis>
. HolotYpe male, carapace and tergites IIV (38), sternopectinal area and sternites (39), and left legs IIV, retrolateral aspect (4245).
<emphasis id="B904446714339C24FDE7F831189D39D3" bold="true" box="[620,764,1977,1999]" pageId="8" pageNumber="7">Figures 4041</emphasis>
. ParatYpe female, carapace and tergites IV (40) and sternopectinal area and sternites (41).
</paragraph>
</caption>
<caption id="DF0FC8FD14329C25FFF8FBA219D23A67" ID-DOI="http://doi.org/10.5281/zenodo.7162596" ID-Zenodo-Dep="7162596" httpUri="https://zenodo.org/record/7162596/files/figure.png" pageId="9" pageNumber="8" startId="9.[115,192,1066,1088]" targetBox="[126,1464,253,1043]" targetPageId="9">
<paragraph id="8BCF987514329C25FFF8FBA219D23A67" blockId="9.[115,1473,1066,1146]" pageId="9" pageNumber="8">
<emphasis id="B904446714329C25FFF8FBA21B633A22" bold="true" box="[115,258,1066,1088]" pageId="9" pageNumber="8">Figures 4651</emphasis>
: Comparison of telsons of
<taxonomicName id="4C70E3F614329C25FD80FBA318D43A5D" box="[523,693,1066,1088]" class="Arachnida" family="Buthidae" genus="Lanzatus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B904446714329C25FD80FBA318033A5D" box="[523,610,1067,1088]" italics="true" pageId="9" pageNumber="8">Lanzatus</emphasis>
species.
</taxonomicName>
<emphasis id="B904446714329C25FD30FBA2192A3A22" bold="true" box="[699,843,1066,1088]" pageId="9" pageNumber="8">Figures 4647</emphasis>
.
<taxonomicName id="4C70E3F614329C25FCD3FBA319933A22" authority="Kovařík &amp; Lowe, 2021" authorityName="Kovařík &amp; Lowe" authorityYear="2021" box="[856,1010,1066,1088]" class="Arachnida" family="Buthidae" genus="Lanzatus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="huluul" status="sp. nov.">
<emphasis id="B904446714329C25FCD3FBA319933A22" box="[856,1010,1066,1088]" italics="true" pageId="9" pageNumber="8">Lanzatus huluul</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A237F91C14329C25FC72FBA21E4E3A5D" box="[1017,1071,1066,1088]" pageId="9" pageNumber="8" rank="species">
<emphasis id="B904446714329C25FC72FBA21E713A22" bold="true" box="[1017,1040,1066,1087]" pageId="9" pageNumber="8">sp</emphasis>
.
<emphasis id="B904446714329C25FB96FBA21E4B3A22" bold="true" box="[1053,1066,1066,1087]" pageId="9" pageNumber="8">n</emphasis>
.
</taxonomicNameLabel>
, female paratYpe (46) and male holotYpe (47).
<emphasis id="B904446714329C25FF2CFBCF1B723A41" bold="true" box="[167,275,1095,1117]" pageId="9" pageNumber="8">Figures 48</emphasis>
.
<taxonomicName id="4C70E3F614329C25FEABFBC018103A41" authority="Kovarik, 2001" authorityName="Kovarik" authorityYear="2001" box="[288,625,1095,1117]" class="Arachnida" family="Buthidae" genus="Lanzatus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="somalicus">
<emphasis id="B904446714329C25FEABFBC01BBC3A41" box="[288,477,1095,1117]" italics="true" pageId="9" pageNumber="8">Lanzatus somalicus</emphasis>
<bibRefCitation id="EFE1E58414329C25FE6FFBCF18103A41" author="KOVARIK" box="[484,625,1095,1117]" pageId="9" pageNumber="8" pagination="41 - 44" refId="ref3964" refString="KOVARIK, F. 2001. Lanzatus somalicus gen. n. et sp. n. (Scorpiones: Buthidae) from Somalia. Acta Societas Zoologicae Bohemicae, 65 (1): 41 - 44." type="journal article" year="2001">Kovařík, 2001</bibRefCitation>
</taxonomicName>
, male paratYpe.
<emphasis id="B904446714329C25FC9AFBCF19C03A41" bold="true" box="[785,929,1095,1117]" pageId="9" pageNumber="8">Figures 4951</emphasis>
.
<taxonomicName id="4C70E3F614329C25FC26FBC01F143A40" authority="Kovarik et Lowe, 2016" authorityName="Kovarik et Lowe" authorityYear="2016" box="[941,1397,1095,1117]" class="Arachnida" family="Buthidae" genus="Lanzatus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="somalilandus">
<emphasis id="B904446714329C25FC26FBC01EEA3A41" box="[941,1163,1095,1117]" italics="true" pageId="9" pageNumber="8">Lanzatus somalilandus</emphasis>
Kovařík et Lowe, 2016
</taxonomicName>
, female paratYpe (49), male holotYpe (50) and male from Somaliland, Shanshade Village. (51).
</paragraph>
</caption>
<paragraph id="8BCF987514329C25FFFAFB111B8D382C" blockId="9.[113,772,1177,2033]" pageId="9" pageNumber="8">
<emphasis id="B904446714329C25FFFAFB111BFC3AAC" bold="true" box="[113,413,1177,1201]" pageId="9" pageNumber="8">Carapace and mesosoma</emphasis>
(
<figureCitation id="134B84F014329C25FE3AFB11185C3AAC" box="[433,573,1177,1201]" captionStart="Figures 3845" captionStartId="8.[115,192,1948,1970]" captionTargetBox="[115,1476,242,1931]" captionTargetId="figure-73@8.[115,1476,242,1934]" captionTargetPageId="8" captionText="Figures 3845: Lanzatus huluul sp. n. Figures 3839, 4245. HolotYpe male, carapace and tergites IIV (38), sternopectinal area and sternites (39), and left legs IIV, retrolateral aspect (4245). Figures 4041. ParatYpe female, carapace and tergites IV (40) and sternopectinal area and sternites (41)." figureDoi="http://doi.org/10.5281/zenodo.7162594" httpUri="https://zenodo.org/record/7162594/files/figure.png" pageId="9" pageNumber="8">Figs. 3841</figureCitation>
). Entire carapace finelY granulated. Carinae absent. Anterior margin of carapace almost straight. Median ocular tubercle large, width 35% of carapace posterior width. Median eYes prominent, diameter 16% of carapace length. SuperciliarY carinae finelY granulated. Tergites finelY granulated, asetose, with one median carina more stronglY indicated in the female. Pectinal tooth count
<quantity id="4C88359014329C25FEDAFAF11BE63B8C" box="[337,391,1401,1425]" metricMagnitude="-1" metricUnit="m" metricValue="5.334" pageId="9" pageNumber="8" unit="in" value="21.0">21 in</quantity>
male and
<quantity id="4C88359014329C25FE7DFAF118343B8C" box="[502,597,1401,1425]" metricMagnitude="-1" metricUnit="m" metricValue="5.461" metricValueMax="5.588" metricValueMin="5.334" pageId="9" pageNumber="8" unit="in" value="21.5" valueMax="22.0" valueMin="21.0">2122 in</quantity>
female. Pectine marginal tips extend to half of the fifth sternite in both sexes. Pectines with 3 marginal lamellae and 89 middle lamellae. All lamellae and fulcra bear numerous setae. All sternites finelY granulated (more so in female), setose, without carinae, posterior margins smooth.
</paragraph>
<paragraph id="8BCF987514329C25FFFAF9B1183239EC" blockId="9.[113,772,1177,2033]" pageId="9" pageNumber="8">
<emphasis id="B904446714329C25FFFAF9B11B3D384C" bold="true" box="[113,348,1593,1617]" pageId="9" pageNumber="8">Hemispermatophore</emphasis>
(
<figureCitation id="134B84F014329C25FEF2F9B1186B384C" box="[377,522,1593,1617]" captionStart="Figures 5253" captionStartId="10.[111,188,1981,2003]" captionTargetBox="[212,1338,238,1965]" captionTargetId="figure-48@10.[144,1458,223,1974]" captionTargetPageId="10" captionText="Figures 5253: Lanzatus huluul sp. n., left hemispermatophore. HolotYpe male, whole hemispermatophore (52) and capsule, compressed to separate lobes (53), convex views. Scale bars: 1 mm (52), 200 μm (53)." figureDoi="http://doi.org/10.5281/zenodo.7162598" httpUri="https://zenodo.org/record/7162598/files/figure.png" pageId="9" pageNumber="8">Figs. 5253</figureCitation>
). Flagelliform, trunk elongate, ca. 6.3 times length of capsule region; flagellum with linear pars recta and pars reflecta, separate from sperm hemiduct lobes. Proximal pars recta with narrow laminate expansion along anterior margin. Sperm hemiduct divided into 3 lobes: posterior lobe long, broad, laminate, gentlY tapered; median lobe small, narrow, filamentous in distal 2/3; anterior lobe tapered, acuminate. Posterior margin of median lobe slightlY overlapping posterior lobe, the two partiallY joined along an axial suture or carina. Basal lobe a broad, curved scoop, obliquelY-angled. The capsule is verY similar to that of
<taxonomicName id="4C70E3F614329C25FFFAF8121B7A39AC" authorityName="Kovarik et Lowe" authorityYear="2016" box="[113,283,1945,1969]" class="Arachnida" family="Buthidae" genus="Lanzatus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="somalilandus">
<emphasis id="B904446714329C25FFFAF8121B7A39AC" box="[113,283,1945,1969]" italics="true" pageId="9" pageNumber="8">L. somalilandus</emphasis>
</taxonomicName>
, which has sperm hemiduct lobes of the same shape, and also bears an oblique, scoop-like basal lobe (cf. Kovařík &amp; Njoroge, 2021:16, figs. 111115).
</paragraph>
<paragraph id="8BCF987514329C25FCBBFB111EBF384C" blockId="9.[816,1475,1177,2033]" pageId="9" pageNumber="8">
<emphasis id="B904446714329C25FCBBFB111E513AAC" bold="true" box="[816,1072,1177,1201]" pageId="9" pageNumber="8">Metasoma and telson</emphasis>
(
<figureCitation id="134B84F014329C25FBCEFB111EA03AAC" box="[1093,1217,1177,1201]" captionStart="Figures 716" captionStartId="6.[115,192,1960,1982]" captionTargetBox="[113,1467,243,1945]" captionTargetId="figure-98@6.[113,1474,234,1948]" captionTargetPageId="6" captionText="Figures 716: Lanzatus huluul sp. n., metasoma and telson. Figures 711. HolotYpe male, metasoma V and telson (7) and telson (8) in lateral views and metasoma and telson in lateral (9), dorsal (10), and ventral (11) views. Figures 1216. ParatYpe female, metasoma V and telson (12) and telson (13) in lateral views and metasoma and telson in lateral (14), dorsal (15), and ventral (16) views.Scale bars: 5 mm (911 and 1416)." figureDoi="http://doi.org/10.5281/zenodo.7162590" httpUri="https://zenodo.org/record/7162590/files/figure.png" pageId="9" pageNumber="8">Figs. 716</figureCitation>
). Metasomal segments smooth and setose, bearing numerous long macrosetae, without carinae, with rounded edges. Dorsal surfaces of metasoma IIV bearing a shallow longitudinal groove. First metasomal segment finelY granulated in both sexes, metasoma IIIII smooth (female) or denselY and verY finelY granulated (males), metasoma IVV smooth in both sexes. Posterior metasomal segments successivelY more narrow than anterior segments. Metasoma V non-uniform in width, tapering posteriorlY. Telson extremelY elongated, smooth, without aculear ring. Posterior ventral aspect of vesicle concave in lateral profile, transitioning smoothlY to aculeus without a constriction. Aculeus stout, weaklY curved. Telson length/ width ratio
<quantity id="4C88359014329C25FC20F9B11997384C" box="[939,1014,1593,1617]" metricMagnitude="-1" metricUnit="m" metricValue="1.52908" pageId="9" pageNumber="8" unit="in" value="6.02">6.02 in</quantity>
male,
<quantity id="4C88359014329C25FBB6F9B11EE9384C" box="[1085,1160,1593,1617]" metricMagnitude="-1" metricUnit="m" metricValue="1.35382" pageId="9" pageNumber="8" unit="in" value="5.33">5.33 in</quantity>
female.
</paragraph>
<paragraph id="8BCF987514329C25FCBBF9D11E26398C" blockId="9.[816,1475,1177,2033]" pageId="9" pageNumber="8">
<emphasis id="B904446714329C25FCBBF9D119C8386C" bold="true" box="[816,937,1625,1649]" pageId="9" pageNumber="8">Chelicerae</emphasis>
. Manus longer than wide, dorsal surface smooth, glossY, with strong anterior marginal and dorsointernal carinae. Fingers robust, with tYpical buthid dentition (Vachon, 1963). Dorsal margin of movable finger armed with 5 teeth: dorsal distal tine, subdistal, median and apparentlY 2 small basal teeth fused in bicusp. Ventral margin of movable finger with 3 teeth: ventral distal tine, median and basal teeth. Fixed finger margin with 4 teeth: distal tine, subdistal, median and basal teeth fused into bicusp. Ventral surface of fixed finger armed with a single tooth.
</paragraph>
<paragraph id="8BCF987514329C20FCBBF811187B3FFC" blockId="9.[816,1475,1177,2033]" lastBlockId="12.[113,772,233,737]" lastPageId="12" lastPageNumber="11" pageId="9" pageNumber="8">
<emphasis id="B904446714329C25FCBBF81119FE39AC" bold="true" box="[816,927,1945,1969]" pageId="9" pageNumber="8">Pedipalps</emphasis>
(
<figureCitation id="134B84F014329C25FC24F8111E5539AC" box="[943,1076,1945,1969]" captionStart="Figures 1737" captionStartId="7.[114,191,1898,1920]" captionTargetBox="[116,1459,237,1887]" captionTargetId="figure-126@7.[116,1477,234,1890]" captionTargetPageId="7" captionText="Figures 1737: Lanzatus huluul sp. n., right pedipalp. Figures 1726. HolotYpe male, chela in dorsal (17), external (18) and ventral (19) views, patella in dorsal (20), external (21) and ventral (22) views, femur and trochanter in dorsal (23), internal (24) and ventral (25) views, dentate margins of movable finger (26). Figures 2737. ParatYpe female, chela in dorsal (27), external (28) and ventral (29) views, patella in dorsal (30), external (31) and ventral (32) views, femur and trochanter in dorsal (33), internal (34) and ventral (35) views. dentate margins of movable (36) and fixed (37) fingers. Trichobothrial pattern indicated in Figures 2731 and 3334 bY white circles." figureDoi="http://doi.org/10.5281/zenodo.7162592" httpUri="https://zenodo.org/record/7162592/files/figure.png" pageId="9" pageNumber="8">Figs. 1737</figureCitation>
). Pedipalps smooth, sparselY hirsute, with inconspicuous smooth carinae, onlY the femur with one (male) or two (female) dorsal carinae composed of strong granules. Pedipalp chela narrow, chela length/ width ratio
<quantity id="4C88359014379C20FD69FF611AE73F3C" metricMagnitude="-1" metricUnit="m" metricValue="1.6256" pageId="12" pageNumber="11" unit="in" value="6.4">6.4 in</quantity>
male,
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female. Dentate margins of chela fingers with distinct granules divided into 7 rows (including short apical row of 3 granules) on movable finger, 6 rows on fixed finger. Granule rows oblique except for long proximal row, which is parallel to finger axis. All rows flanked bY a single internal accessorY granule. Both fingers with enlarged terminal denticle, and 12 subterminal denticles.
</paragraph>
<caption id="DF0FC8FD14319C26FFE4F835197F39ED" ID-DOI="http://doi.org/10.5281/zenodo.7162598" ID-Zenodo-Dep="7162598" httpUri="https://zenodo.org/record/7162598/files/figure.png" pageId="10" pageNumber="9" startId="10.[111,188,1981,2003]" targetBox="[212,1338,238,1965]" targetPageId="10">
<paragraph id="8BCF987514319C26FFE4F835197F39ED" blockId="10.[111,1470,1981,2032]" pageId="10" pageNumber="9">
<emphasis id="B904446714319C26FFE4F8351B6639CF" bold="true" box="[111,263,1981,2003]" pageId="10" pageNumber="9">Figures 5253:</emphasis>
<taxonomicName id="4C70E3F614319C26FE85F8361BC939CE" authority="Kovařík &amp; Lowe, 2021" authorityName="Kovařík &amp; Lowe" authorityYear="2021" box="[270,424,1982,2003]" class="Arachnida" family="Buthidae" genus="Lanzatus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="huluul" status="sp. nov.">
<emphasis id="B904446714319C26FE85F8361BC939CE" box="[270,424,1982,2003]" italics="true" pageId="10" pageNumber="9">Lanzatus huluul</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A237F91C14319C26FE24F8361B8439CE" box="[431,485,1982,2003]" pageId="10" pageNumber="9" rank="species">
<emphasis id="B904446714319C26FE24F8361BA739CE" bold="true" box="[431,454,1982,2003]" pageId="10" pageNumber="9">sp</emphasis>
.
<emphasis id="B904446714319C26FE59F8361BBE39CE" bold="true" box="[466,479,1982,2003]" pageId="10" pageNumber="9">n</emphasis>
.
</taxonomicNameLabel>
, left hemispermatophore. HolotYpe male, whole hemispermatophore (52) and capsule, compressed to separate lobes (53), convex views. Scale bars: 1 mm (52), 200 μm (53).
</paragraph>
</caption>
<caption id="DF0FC8FD14309C27FFF8F8361BD639ED" ID-DOI="http://doi.org/10.5281/zenodo.7162600" ID-Zenodo-Dep="7162600" httpUri="https://zenodo.org/record/7162600/files/figure.png" pageId="11" pageNumber="10" startId="11.[115,192,1982,2004]" targetBox="[115,1476,233,1977]" targetPageId="11">
<paragraph id="8BCF987514309C27FFF8F8361BD639ED" blockId="11.[115,1472,1982,2033]" pageId="11" pageNumber="10">
<emphasis id="B904446714309C27FFF8F8361B6339CE" bold="true" box="[115,258,1982,2004]" pageId="11" pageNumber="10">Figures 5455</emphasis>
:
<emphasis id="B904446714309C27FE9BF8361B1339CE" bold="true" box="[272,370,1982,2004]" pageId="11" pageNumber="10">Figure 54</emphasis>
. Map showing known distribution of the genus
<taxonomicName id="4C70E3F614309C27FCCEF83719FD39C9" authorityName="Kovarik" authorityYear="2001" box="[837,924,1983,2004]" class="Arachnida" family="Buthidae" genus="Lanzatus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B904446714309C27FCCEF83719FD39C9" box="[837,924,1983,2004]" italics="true" pageId="11" pageNumber="10">Lanzatus</emphasis>
</taxonomicName>
.
<emphasis id="B904446714309C27FC23F8361E6A39CE" bold="true" box="[936,1035,1982,2004]" pageId="11" pageNumber="10">Figure 55</emphasis>
. Somaliland, vicinitY of Huluul Village, tYpe localitY of
<taxonomicName id="4C70E3F614309C27FF6AF8541B1B39ED" authority="Kovařík &amp; Lowe, 2021" authorityName="Kovařík &amp; Lowe" authorityYear="2021" box="[225,378,2011,2033]" class="Arachnida" family="Buthidae" genus="Lanzatus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="huluul" status="sp. nov.">
<emphasis id="B904446714309C27FF6AF8541B1B39ED" box="[225,378,2011,2033]" italics="true" pageId="11" pageNumber="10">Lanzatus huluul</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A237F91C14309C27FE0AF8531BD639ED" box="[385,439,2011,2032]" pageId="11" pageNumber="10" rank="species">
<emphasis id="B904446714309C27FE0AF8531BF939ED" bold="true" box="[385,408,2011,2032]" pageId="11" pageNumber="10">sp</emphasis>
.
<emphasis id="B904446714309C27FE28F8531BD139ED" bold="true" box="[419,432,2011,2032]" pageId="11" pageNumber="10">n</emphasis>
.
</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BCF987514379C20FFFAFE611AA73CDC" blockId="12.[113,772,233,737]" pageId="12" pageNumber="11">
<emphasis id="B904446714379C20FFFAFE611AC73C1C" bold="true" box="[113,166,489,513]" pageId="12" pageNumber="11">Legs</emphasis>
(
<figureCitation id="134B84F014379C20FF35FE611B263C1C" box="[190,327,489,513]" captionStart="Figures 3845" captionStartId="8.[115,192,1948,1970]" captionTargetBox="[115,1476,242,1931]" captionTargetId="figure-73@8.[115,1476,242,1934]" captionTargetPageId="8" captionText="Figures 3845: Lanzatus huluul sp. n. Figures 3839, 4245. HolotYpe male, carapace and tergites IIV (38), sternopectinal area and sternites (39), and left legs IIV, retrolateral aspect (4245). Figures 4041. ParatYpe female, carapace and tergites IV (40) and sternopectinal area and sternites (41)." figureDoi="http://doi.org/10.5281/zenodo.7162594" httpUri="https://zenodo.org/record/7162594/files/figure.png" pageId="12" pageNumber="11">Figs. 4245</figureCitation>
). Legs IIV with tibial spurs absent, retrolateral and prolateral pedal spurs present.All legs smooth, without distinct carinae. Telotarsi bear two rows of fine macrosetae on their ventral surfaces, and several additional macrosetae on other surfaces. Basitarsi of legs IIII with 4 to 7 macrosetae, not arranged in regular rows (bristle combs absent).
</paragraph>
<paragraph id="8BCF987514379C20FFFAFD411BC93CFC" blockId="12.[113,772,233,737]" box="[113,424,713,737]" pageId="12" pageNumber="11">
<emphasis id="B904446714379C20FFFAFD411B793CFC" bold="true" box="[113,280,713,737]" pageId="12" pageNumber="11">Measurements</emphasis>
. See
<tableCitation id="C6F2ADCE14379C20FED9FD411BC33CFC" box="[338,418,713,737]" captionStart="Table 1" captionStartId="4.[110,166,991,1012]" captionText="Table 1. Comparative measurements of tYpes of Lanzatus huluul sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="12" pageNumber="11">Table 1</tableCitation>
.
</paragraph>
<paragraph id="8BCF987514379C20FFFAFC811BE53A9C" blockId="12.[113,771,777,1153]" pageId="12" pageNumber="11">
AFFINITIES.
<taxonomicName id="4C70E3F614379C20FF7FFC821BC23D3C" authority="Kovařík &amp; Lowe, 2021" authorityName="Kovařík &amp; Lowe" authorityYear="2021" box="[244,419,777,801]" class="Arachnida" family="Buthidae" genus="Lanzatus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="huluul" status="sp. nov.">
<emphasis id="B904446714379C20FF7FFC821BC23D3C" box="[244,419,777,801]" italics="true" pageId="12" pageNumber="11">Lanzatus huluul</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A237F91C14379C20FE25FC811B8E3D3C" box="[430,495,777,801]" pageId="12" pageNumber="11" rank="species">
<emphasis id="B904446714379C20FE25FC811BA63D3C" bold="true" box="[430,455,777,801]" pageId="12" pageNumber="11">sp</emphasis>
.
<emphasis id="B904446714379C20FE52FC811B893D3C" bold="true" box="[473,488,777,801]" pageId="12" pageNumber="11">n</emphasis>
.
</taxonomicNameLabel>
is differentiated from
<taxonomicName id="4C70E3F614379C20FD65FC821ABD3D5C" authorityName="Kovarik" authorityYear="2001" class="Arachnida" family="Buthidae" genus="Lanzatus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="somalicus">
<emphasis id="B904446714379C20FD65FC821ABD3D5C" italics="true" pageId="12" pageNumber="11">L. somalicus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C70E3F614379C20FE99FCA21BDF3D5C" authorityName="Kovarik et Lowe" authorityYear="2016" box="[274,446,809,833]" class="Arachnida" family="Buthidae" genus="Lanzatus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="somalilandus">
<emphasis id="B904446714379C20FE99FCA21BDF3D5C" box="[274,446,809,833]" italics="true" pageId="12" pageNumber="11">L. somalilandus</emphasis>
</taxonomicName>
mainlY bY the morphologY of the telson (
<figureCitation id="134B84F014379C20FF69FCC11B003D7C" box="[226,353,841,865]" captionStart="Figures 4651" captionStartId="9.[115,192,1066,1088]" captionTargetBox="[126,1464,253,1043]" captionTargetId="figure-519@9.[113,1474,234,1055]" captionTargetPageId="9" captionText="Figures 4651: Comparison of telsons of Lanzatus species. Figures 4647. Lanzatus huluul sp. n., female paratYpe (46) and male holotYpe (47). Figures 48. Lanzatus somalicus Kovařík, 2001, male paratYpe. Figures 4951. Lanzatus somalilandus Kovařík et Lowe, 2016, female paratYpe (49), male holotYpe (50) and male from Somaliland, Shanshade Village. (51)." figureDoi="http://doi.org/10.5281/zenodo.7162596" httpUri="https://zenodo.org/record/7162596/files/figure.png" pageId="12" pageNumber="11">Figs. 4651</figureCitation>
), which is extremelY elongated, smooth, without an aculear ring, and with the aculeus slightlY curved in
<taxonomicName id="4C70E3F614379C20FFFAFC021AAB3DBC" authority="Kovařík &amp; Lowe, 2021" authorityName="Kovařík &amp; Lowe" authorityYear="2021" box="[113,202,905,929]" class="Arachnida" family="Buthidae" genus="Lanzatus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="huluul" status="sp. nov.">
<emphasis id="B904446714379C20FFFAFC021AAB3DBC" box="[113,202,905,929]" italics="true" pageId="12" pageNumber="11">L huluul</emphasis>
</taxonomicName>
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<emphasis id="B904446714379C20FF5BFC011A883DBC" bold="true" box="[208,233,905,929]" pageId="12" pageNumber="11">sp</emphasis>
.
<emphasis id="B904446714379C20FF7DFC011B643DBC" bold="true" box="[246,261,905,929]" pageId="12" pageNumber="11">n</emphasis>
.
</taxonomicNameLabel>
Telson length/ width ratio
<quantity id="4C88359014379C20FDA2FC0118123DBC" box="[553,627,905,929]" metricMagnitude="-1" metricUnit="m" metricValue="1.52908" pageId="12" pageNumber="11" unit="in" value="6.02">6.02 in</quantity>
male,
<quantity id="4C88359014379C20FD32FC0119623DBC" box="[697,771,905,929]" metricMagnitude="-1" metricUnit="m" metricValue="1.35382" pageId="12" pageNumber="11" unit="in" value="5.33">5.33 in</quantity>
female in
<taxonomicName id="4C70E3F614379C20FF57FC221B573DDC" authority="Kovařík &amp; Lowe, 2021" authorityName="Kovařík &amp; Lowe" authorityYear="2021" box="[220,310,937,961]" class="Arachnida" family="Buthidae" genus="Lanzatus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="huluul" status="sp. nov.">
<emphasis id="B904446714379C20FF57FC221B573DDC" box="[220,310,937,961]" italics="true" pageId="12" pageNumber="11">L huluul</emphasis>
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<emphasis id="B904446714379C20FEB6FC211B373DDC" bold="true" box="[317,342,937,961]" pageId="12" pageNumber="11">sp</emphasis>
.
<emphasis id="B904446714379C20FEEFFC211B123DDC" bold="true" box="[356,371,937,961]" pageId="12" pageNumber="11">n</emphasis>
.
</taxonomicNameLabel>
In
<taxonomicName id="4C70E3F614379C20FE16FC2218453DDC" authorityName="Kovarik" authorityYear="2001" box="[413,548,937,961]" class="Arachnida" family="Buthidae" genus="Lanzatus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="somalicus">
<emphasis id="B904446714379C20FE16FC2218453DDC" box="[413,548,937,961]" italics="true" pageId="12" pageNumber="11">L. somalicus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C70E3F614379C20FDD3FC2219623DDC" authorityName="Kovarik et Lowe" authorityYear="2016" box="[600,771,937,961]" class="Arachnida" family="Buthidae" genus="Lanzatus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="somalilandus">
<emphasis id="B904446714379C20FDD3FC2219623DDC" box="[600,771,937,961]" italics="true" pageId="12" pageNumber="11">L. somalilandus</emphasis>
</taxonomicName>
the aculeus is stronglY curved. The telson length/ width ratio is
<quantity id="4C88359014379C20FF01FC611AB53A1C" box="[138,212,1001,1025]" metricMagnitude="-1" metricUnit="m" metricValue="1.00076" pageId="12" pageNumber="11" unit="in" value="3.94">3.94 in</quantity>
the male, and
<quantity id="4C88359014379C20FEE4FC611BD83A1C" box="[367,441,1001,1025]" metricMagnitude="-2" metricUnit="m" metricValue="9.1694" pageId="12" pageNumber="11" unit="in" value="3.61">3.61 in</quantity>
the female of
<taxonomicName id="4C70E3F614379C20FDD9FC62189D3A1C" authorityName="Kovarik et Lowe" authorityYear="2016" box="[594,764,1001,1025]" class="Arachnida" family="Buthidae" genus="Lanzatus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="somalilandus">
<emphasis id="B904446714379C20FDD9FC62189D3A1C" box="[594,764,1001,1025]" italics="true" pageId="12" pageNumber="11">L. somalilandus</emphasis>
</taxonomicName>
, and
<quantity id="4C88359014379C20FF14FB811A8B3A3C" box="[159,234,1033,1057]" metricMagnitude="-2" metricUnit="m" metricValue="8.763" pageId="12" pageNumber="11" unit="in" value="3.45">3.45 in</quantity>
a male
<typeStatus id="54CB26D714379C20FECBFB821BFD3A3C" box="[320,412,1034,1057]" pageId="12" pageNumber="11" type="paratype">paratYpe</typeStatus>
of
<taxonomicName id="4C70E3F614379C20FE4BFB8218293A3C" authorityName="Kovarik" authorityYear="2001" box="[448,584,1033,1057]" class="Arachnida" family="Buthidae" genus="Lanzatus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="somalicus">
<emphasis id="B904446714379C20FE4BFB8218293A3C" box="[448,584,1033,1057]" italics="true" pageId="12" pageNumber="11">L. somalicus</emphasis>
</taxonomicName>
. Basitarsi of legs IIII have macrosetae arranged in regular rows (bristle combs present) in
<taxonomicName id="4C70E3F614379C20FF7FFBC21BC53A7C" authorityName="Kovarik et Lowe" authorityYear="2016" box="[244,420,1097,1121]" class="Arachnida" family="Buthidae" genus="Lanzatus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="somalilandus">
<emphasis id="B904446714379C20FF7FFBC21B623A7C" box="[244,259,1098,1121]" italics="true" pageId="12" pageNumber="11">L</emphasis>
.
<emphasis id="B904446714379C20FE9DFBC11BC53A7C" box="[278,420,1097,1121]" italics="true" pageId="12" pageNumber="11">somalilandus</emphasis>
</taxonomicName>
, and irregularlY (bristle combs absent) in
<taxonomicName id="4C70E3F614379C20FF6BFBE21B203A9C" authority="Kovařík &amp; Lowe, 2021" authorityName="Kovařík &amp; Lowe" authorityYear="2021" box="[224,321,1129,1153]" class="Arachnida" family="Buthidae" genus="Lanzatus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="huluul" status="sp. nov.">
<emphasis id="B904446714379C20FF6BFBE21A8E3A9C" box="[224,239,1130,1153]" italics="true" pageId="12" pageNumber="11">L</emphasis>
.
<emphasis id="B904446714379C20FF76FBE11B203A9C" box="[253,321,1129,1153]" italics="true" pageId="12" pageNumber="11">huluul</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A237F91C14379C20FECCFBE11BE53A9C" box="[327,388,1129,1153]" pageId="12" pageNumber="11" rank="species">
<emphasis id="B904446714379C20FECCFBE11B013A9C" bold="true" box="[327,352,1129,1153]" pageId="12" pageNumber="11">sp</emphasis>
.
<emphasis id="B904446714379C20FEE5FBE11B1C3A9C" bold="true" box="[366,381,1129,1153]" pageId="12" pageNumber="11">n</emphasis>
.
</taxonomicNameLabel>
</paragraph>
<paragraph id="8BCF987514379C20FFFAFB211AFD3AFC" blockId="12.[113,771,1193,1249]" pageId="12" pageNumber="11">
DISTRIBUTION. Known onlY from the
<typeStatus id="54CB26D714379C20FD8CFB2218543ADC" box="[519,565,1194,1217]" pageId="12" pageNumber="11">tYpe</typeStatus>
localitY (
<figureCitation id="134B84F014379C20FD16FB211AEE3AFC" captionStart="Figures 5455" captionStartId="11.[115,192,1982,2004]" captionTargetBox="[115,1476,233,1977]" captionTargetId="figure-43@11.[115,1476,230,1979]" captionTargetPageId="11" captionText="Figures 5455: Figure 54. Map showing known distribution of the genus Lanzatus. Figure 55. Somaliland, vicinitY of Huluul Village, tYpe localitY of Lanzatus huluul sp. n." figureDoi="http://doi.org/10.5281/zenodo.7162600" httpUri="https://zenodo.org/record/7162600/files/figure.png" pageId="12" pageNumber="11">Figs. 54 55</figureCitation>
).
</paragraph>
<paragraph id="8BCF987514379C20FFFAFA811B0538C1" blockId="12.[113,771,1289,1756]" pageId="12" pageNumber="11">
COMMENTS ON LOCALITY AND LIFE STRATEGY. The
<typeStatus id="54CB26D714379C20FD23FA8218813B3C" box="[680,736,1290,1313]" pageId="12" pageNumber="11">tYpes</typeStatus>
of
<taxonomicName id="4C70E3F614379C20FFFAFAA21AB73B5C" authority="Kovařík &amp; Lowe, 2021" authorityName="Kovařík &amp; Lowe" authorityYear="2021" box="[113,214,1321,1345]" class="Arachnida" family="Buthidae" genus="Lanzatus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="huluul" status="sp. nov.">
<emphasis id="B904446714379C20FFFAFAA21AE13B5C" box="[113,128,1322,1345]" italics="true" pageId="12" pageNumber="11">L</emphasis>
.
<emphasis id="B904446714379C20FF19FAA11AB73B5C" box="[146,214,1321,1345]" italics="true" pageId="12" pageNumber="11">huluul</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A237F91C14379C20FF6AFAA11B433B5C" box="[225,290,1321,1345]" pageId="12" pageNumber="11" rank="species">
<emphasis id="B904446714379C20FF6AFAA11A9B3B5C" bold="true" box="[225,250,1321,1345]" pageId="12" pageNumber="11">sp</emphasis>
.
<emphasis id="B904446714379C20FE87FAA11B7A3B5C" bold="true" box="[268,283,1321,1345]" pageId="12" pageNumber="11">n</emphasis>
.
</taxonomicNameLabel>
were collected on a consolidated substrate that can support construction of stable burrows (
<figureCitation id="134B84F014379C20FD0EFAC118B63B7C" box="[645,727,1353,1377]" captionStart="Figures 5455" captionStartId="11.[115,192,1982,2004]" captionTargetBox="[115,1476,233,1977]" captionTargetId="figure-43@11.[115,1476,230,1979]" captionTargetPageId="11" captionText="Figures 5455: Figure 54. Map showing known distribution of the genus Lanzatus. Figure 55. Somaliland, vicinitY of Huluul Village, tYpe localitY of Lanzatus huluul sp. n." figureDoi="http://doi.org/10.5281/zenodo.7162600" httpUri="https://zenodo.org/record/7162600/files/figure.png" pageId="12" pageNumber="11">Fig. 55</figureCitation>
). In contrast, the related
<taxonomicName id="4C70E3F614379C20FEDCFAE218673B9C" authorityName="Kovarik et Lowe" authorityYear="2016" box="[343,518,1385,1409]" class="Arachnida" family="Buthidae" genus="Lanzatus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="somalilandus">
<emphasis id="B904446714379C20FEDCFAE218673B9C" box="[343,518,1385,1409]" italics="true" pageId="12" pageNumber="11">L. somalilandus</emphasis>
</taxonomicName>
was found on aeolian sand in red desert (Kovařík et. al., 2016: 89, fig. 40). This habitat difference correlates with the presence or absence of psammophilous adaptation (bristle combs) in the two species. A similarlY elongated telson and tapered metasoma V is also found in the ultrapsammophile vaejovid,
<taxonomicName id="4C70E3F614379C20FD04F9811BCA385C" authority="(Williams, 1969)" baseAuthorityName="Williams" baseAuthorityYear="1969" class="Arachnida" family="Vaejovidae" genus="Vejovoidus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="longiunguis">
<emphasis id="B904446714379C20FD04F9811A8E385C" italics="true" pageId="12" pageNumber="11">Vejovoidus longiunguis</emphasis>
(
<bibRefCitation id="EFE1E58414379C20FF77F9A11BC2385C" author="WILLIAMS" box="[252,419,1577,1601]" pageId="12" pageNumber="11" pagination="285 - 291" refId="ref4725" refString="WILLIAMS, S. C. 1969. A new species of Syntropis from Baja California Sur, Mexico with notes on its biologY. The Pan-Pacific Entomologist, 45 (4): 285 - 291." type="journal article" year="1969">Williams, 1969</bibRefCitation>
)
</taxonomicName>
.
<bibRefCitation id="EFE1E58414379C20FE38F9A118F5385C" author="WILLIAMS" box="[435,660,1577,1601]" pageId="12" pageNumber="11" pagination="285 - 291" refId="ref4725" refString="WILLIAMS, S. C. 1969. A new species of Syntropis from Baja California Sur, Mexico with notes on its biologY. The Pan-Pacific Entomologist, 45 (4): 285 - 291." type="journal article" year="1969">Williams (1969: 290)</bibRefCitation>
suggested that in this species “
<emphasis id="B904446714379C20FEC7F9C11AC83883" italics="true" pageId="12" pageNumber="11">streamlining of the metasoma and telson is perhaps an adaptation for escaping from being buried in the loose sand.</emphasis>
” However, this hYpothesis is not supported in
<taxonomicName id="4C70E3F614379C20FDF5F90F18BB3883" authorityName="Kovarik" authorityYear="2001" box="[638,730,1671,1694]" class="Arachnida" family="Buthidae" genus="Lanzatus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B904446714379C20FDF5F90F18BB3883" box="[638,730,1671,1694]" italics="true" pageId="12" pageNumber="11">Lanzatus</emphasis>
</taxonomicName>
, for which the non-psammophilic species,
<taxonomicName id="4C70E3F614379C20FE73F92D183538A1" authority="Kovařík &amp; Lowe, 2021" authorityName="Kovařík &amp; Lowe" authorityYear="2021" box="[504,596,1700,1724]" class="Arachnida" family="Buthidae" genus="Lanzatus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="huluul" status="sp. nov.">
<emphasis id="B904446714379C20FE73F92D186738A1" box="[504,518,1701,1724]" italics="true" pageId="12" pageNumber="11">L</emphasis>
.
<emphasis id="B904446714379C20FD98F92C183538A1" box="[531,596,1700,1724]" italics="true" pageId="12" pageNumber="11">huluul</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A237F91C14379C20FDD0F92C18F538A1" box="[603,660,1700,1724]" pageId="12" pageNumber="11" rank="species">
<emphasis id="B904446714379C20FDD0F92C181238A1" bold="true" box="[603,627,1700,1724]" pageId="12" pageNumber="11">sp</emphasis>
.
<emphasis id="B904446714379C20FD0BF92C18EF38A1" bold="true" box="[640,654,1700,1724]" pageId="12" pageNumber="11">n</emphasis>
.
</taxonomicNameLabel>
, possesses the more slender telson.
</paragraph>
</subSubSection>
</treatment>
</document>