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<document id="4A6ECB32CA31E8F8F93FABEFB43C5EAD" ID-CLB-Dataset="2380" ID-DOI="10.5281/zenodo.5739643" ID-GBIF-Dataset="f9439543-76ca-43e9-aca5-0a8d3899a2b1" ID-ISSN="0374-1036" ID-Zenodo-Dep="5739643" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1638252251656" checkinUser="marcus" docAuthor="Makranczy, György" docDate="2013" docId="03CEDF09FFFDF34BD1B390A9D9D9FE41" docLanguage="en" docName="ActaEntMusNatPra.2013.53.1.177-208.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 53 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Thinodromus tibialis" docType="treatment" docVersion="8" lastPageNumber="187" masterDocId="FFF7A771FFF4F340D0219333DB72FFF2" masterDocTitle="Review of the southern African species of Thinodromus (Coleoptera: Staphylinidae: Oxytelinae)" masterLastPageNumber="208" masterPageNumber="177" pageNumber="186" updateTime="1699471184067" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="DD7DA8A9747993043132C1A5FAFAAC26">Review of the southern African species of Thinodromus (Coleoptera: Staphylinidae: Oxytelinae)</mods:title>
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<mods:namePart id="9A5677EF8E2C5CD0D148E1F8548B7663">Makranczy, György</mods:namePart>
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<treatment id="03CEDF09FFFDF34BD1B390A9D9D9FE41" ID-DOI="http://doi.org/10.5281/zenodo.5911271" ID-GBIF-Taxon="192662470" ID-Zenodo-Dep="5911271" LSID="urn:lsid:plazi:treatment:03CEDF09FFFDF34BD1B390A9D9D9FE41" httpUri="http://treatment.plazi.org/id/03CEDF09FFFDF34BD1B390A9D9D9FE41" lastPageId="11" lastPageNumber="187" pageId="9" pageNumber="186">
<subSubSection id="C37D3D94FFFDF349D1B390A9D852FC41" box="[402,800,922,947]" pageId="9" pageNumber="186" type="nomenclature">
<paragraph id="8BD86E1FFFFDF349D1B390A9D852FC41" blockId="9.[402,800,922,947]" box="[402,800,922,947]" pageId="9" pageNumber="186">
<heading id="D090D973FFFDF349D1B390A9D852FC41" bold="true" box="[402,800,922,947]" centered="true" fontSize="8" level="2" pageId="9" pageNumber="186" reason="3">
<taxonomicName id="4C67159CFFFDF349D1B390A9D852FC41" ID-CoL="56H6F" authority="(Fauvel, 1907)" baseAuthorityName="Fauvel" baseAuthorityYear="1907" box="[402,800,922,947]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="186" phylum="Arthropoda" rank="species" species="tibialis">
<emphasis id="B913B20DFFFDF349D1B390A9D852FC41" bold="true" box="[402,800,922,947]" pageId="9" pageNumber="186">
<emphasis id="B913B20DFFFDF349D1B390A9D90AFC41" bold="true" box="[402,632,922,947]" italics="true" pageId="9" pageNumber="186">Thinodromus tibialis</emphasis>
(
<bibRefCitation id="EFF613EEFFFDF349D2A790A8D86BFC41" author="FAUVEL A." box="[646,793,923,947]" pageId="9" pageNumber="186" pagination="10 - 70" refId="ref14219" refString="FAUVEL A. 1907: Voyage de M. Ch. Alluaud dans l'Afrique Orientale. Staphylinidae. Revue d ' Entomologie 26: 10 - 70." type="journal article" year="1907">Fauvel, 1907</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37D3D94FFFDF349D1D3908AD9B2FC3E" box="[498,704,953,973]" pageId="9" pageNumber="186" type="description">
<paragraph id="8BD86E1FFFFDF349D1D3908AD9B2FC3E" blockId="9.[113,1090,953,1316]" box="[498,704,953,973]" pageId="9" pageNumber="186">
(
<figureCitation id="135C729AFFFDF349D1D9908AD929FC3F" box="[504,603,953,973]" captionStart="Figs 2833. 2831" captionStartId="16.[151,188,1545,1564]" captionTargetBox="[151,1126,219,1496]" captionTargetId="figure-10@16.[151,1126,219,1497]" captionTargetPageId="16" captionText="Figs 2833. 2831 Thinodromus tibialis (Fauvel, 1907) (28 head and pronotum; 29 female antenna; 30 male antenna; 31 elytron and abdominal base); 3233 T. gildenkovi sp. nov. (32 head and pronotum; 33 elytron and abdominal base). Scale bar: 0.45 mm for 2831, 0.55 mm for 3233." figureDoi="http://doi.org/10.5281/zenodo.5739659" httpUri="https://zenodo.org/record/5739659/files/figure.png" pageId="9" pageNumber="186">Figs 2831</figureCitation>
,
<figureCitation id="135C729AFFFDF349D242908AD9E9FC3E" box="[611,667,953,972]" captionStart-0="Figs 5257. 5254" captionStart-1="Figs 5863. 5859" captionStartId-0="20.[151,188,1520,1539]" captionStartId-1="21.[113,150,1545,1564]" captionTargetBox-0="[154,1102,220,1466]" captionTargetBox-1="[115,1063,221,1464]" captionTargetId-0="figure-10@20.[151,1103,220,1467]" captionTargetId-1="figure-10@21.[113,1065,219,1466]" captionTargetPageId-0="20" captionTargetPageId-1="21" captionText-0="Figs 5257. 5254 Thinodromus rhodesianus (Scheerpeltz, 1974), male (5253 aedeagus; 54 paramere); 5557 T. tibialis (Fauvel, 1907), male (5556 aedeagus; 57 paramere).Abbreviations:AC apical copulatory sclerite; BA basal sclerites, BM basomedial sclerites, MA medioapical sclerites ML medial lamellae. Scale bar: 0.10 mm for 5557, 0.12 mm for 5254." captionText-1="Figs 5863. 5859 Thinodromus tibialis (Fauvel, 1907), male (58 sternite VIII; 59 tergite X); 6061 T. meridionalis sp. nov., male (60 sternite VIII; 61 tergite X); 6263 T. facilis Gildenkov, 2000, male (62 sternite VIII; 63 tergite X). Scale bar: 0.10 mm for 5961 and 63, 0.15 mm for 58 and 62." figureDoi-0="http://doi.org/10.5281/zenodo.5739671" figureDoi-1="http://doi.org/10.5281/zenodo.5739673" httpUri-0="https://zenodo.org/record/5739671/files/figure.png" httpUri-1="https://zenodo.org/record/5739673/files/figure.png" pageId="9" pageNumber="186">5559</figureCitation>
,
<figureCitation id="135C729AFFFDF349D282908AD9CBFC3E" box="[675,697,953,972]" captionStart="Figs 8185. 8183" captionStartId="25.[113,150,1532,1551]" captionTargetBox="[128,1073,222,1462]" captionTargetId="figure-10@25.[124,1076,219,1466]" captionTargetPageId="25" captionText="Figs 8185. 8183 female ringstructures.81 Thinodromus tibialis (Fauvel, 1907); 82 T. dasys Gildenkov, 2000; 83 T. nigerius Gildenkov, 2000. 8485 female ringstructures with spermathecae. 84 T. facilis Gildenkov, 2000; 85 T. gildenkovi sp. nov. Scale bar: 0.10 mm for 85, 0.12 mm for 81 and 8384, 0.14 mm for 82." figureDoi="http://doi.org/10.5281/zenodo.5739681" httpUri="https://zenodo.org/record/5739681/files/figure.png" pageId="9" pageNumber="186">81</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37D3D94FFFDF349D05090EEDA96FBFB" pageId="9" pageNumber="186" type="reference_group">
<paragraph id="8BD86E1FFFFDF349D05090EEDAB0FC1D" blockId="9.[113,1090,953,1316]" box="[113,450,988,1007]" pageId="9" pageNumber="186">
<treatmentCitationGroup id="AB774931FFFDF349D05090EEDAB0FC1D" box="[113,450,988,1007]" pageId="9" pageNumber="186">
<treatmentCitation id="0AC6480EFFFDF349D05090EEDACCFC1D" author="FAUVEL A." box="[113,446,988,1007]" page="13" pageId="9" pageNumber="186" year="1907">
<taxonomicName id="4C67159CFFFDF349D05090EEDACCFC1D" ID-CoL="599XS" authority="Fauvel, 1907: 13" authorityName="Fauvel" authorityPageNumber="13" authorityYear="1907" box="[113,446,988,1007]" class="Insecta" family="Staphylinidae" genus="Trogophloeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="186" phylum="Arthropoda" rank="species" species="tibialis">
<emphasis id="B913B20DFFFDF349D05090EEDA54FC1D" box="[113,294,989,1007]" italics="true" pageId="9" pageNumber="186">Trogophloeus tibialis</emphasis>
<bibRefCitation id="EFF613EEFFFDF349D10D90EFDACCFC1D" author="FAUVEL A." box="[300,446,988,1007]" pageId="9" pageNumber="186" pagination="10 - 70" refId="ref14219" refString="FAUVEL A. 1907: Voyage de M. Ch. Alluaud dans l'Afrique Orientale. Staphylinidae. Revue d ' Entomologie 26: 10 - 70." type="journal article" year="1907">Fauvel, 1907: 13</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD86E1FFFFDF349D05090C4DA96FBFB" blockId="9.[113,1090,953,1316]" box="[113,484,1014,1034]" pageId="9" pageNumber="186">
<treatmentCitationGroup id="AB774931FFFDF349D05090C4DA96FBFB" box="[113,484,1014,1034]" pageId="9" pageNumber="186">
<treatmentCitation id="0AC6480EFFFDF349D05090C4DAADFBFB" author="HERMAN L. H." box="[113,479,1014,1034]" page=")" pageId="9" pageNumber="186" year="1970">
<taxonomicName id="4C67159CFFFDF349D05090C4DAADFBFB" ID-CoL="56H6F" authority=": HERMAN (1970: 387)" authorityName="HERMAN" authorityPageNumber="387" authorityYear="1970" box="[113,479,1014,1034]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="186" phylum="Arthropoda" rank="species" species="tibialis">
<emphasis id="B913B20DFFFDF349D05090C4DA51FBFB" box="[113,291,1015,1033]" italics="true" pageId="9" pageNumber="186">Thinodromus tibialis</emphasis>
:
<bibRefCitation id="EFF613EEFFFDF349D10F90C4DAADFBFB" author="HERMAN L. H." box="[302,479,1014,1034]" pageId="9" pageNumber="186" pagination="343 - 454" refId="ref14325" refString="HERMAN L. H. 1970: Phylogeny and reclassification of the genera of the rove-beetle subfamily Oxytelinae of the World (Coleoptera, Staphylinidae). Bulletin of the American Museum of Natural History 142 (5): 343 - 454." type="journal article" year="1970">HERMAN (1970: 387)</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C37D3D94FFFDF349D0509713D902FAD1" pageId="9" pageNumber="186" type="materials_examined">
<paragraph id="8BD86E1FFFFDF349D0509713D902FAD1" blockId="9.[113,1090,953,1316]" pageId="9" pageNumber="186">
<emphasis id="B913B20DFFFDF349D0509713DA26FBC1" bold="true" box="[113,340,1055,1075]" pageId="9" pageNumber="186">Type material examined.</emphasis>
<materialsCitation id="3B0F6442FFFDF349D1789712DF30FB9B" ID-GBIF-Occurrence="3458217330" collectionCode="ISNB" determinerName="Makranczy" elevation="1900" latitude="-0.7202777" location="Naivasha" longLatPrecision="21" longitude="36.42861" municipality="Naivasha" pageId="9" pageNumber="186" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
<typeStatus id="54DCD0BDFFFDF349D1789712DAC0FBC0" box="[345,434,1057,1076]" pageId="9" pageNumber="186" type="lectotype">LECTOTYPE</typeStatus>
(here designated):
<specimenCount id="9D61A596FFFDF349D2769728D914FBC6" box="[599,614,1051,1076]" pageId="9" pageNumber="186" type="male"></specimenCount>
, “
<collectingMunicipality id="6BBCF465FFFDF349D25B9713D9B8FBC1" box="[634,714,1056,1075]" pageId="9" pageNumber="186">Naivasha</collectingMunicipality>
[approx.
<geoCoordinate id="EE5308D8FFFDF349D33C9713D8F2FBC1" box="[797,896,1055,1075]" degrees="00" direction="south" minutes="43" orientation="latitude" pageId="9" pageNumber="186" precision="15" seconds="13" value="-0.7202777">00°4313″S</geoCoordinate>
,
<geoCoordinate id="EE5308D8FFFDF349D3AB9713D89DFBC6" box="[906,1007,1055,1076]" degrees="36" direction="east" minutes="25" orientation="longitude" pageId="9" pageNumber="186" precision="15" seconds="43" value="36.42861">36°2543″E</geoCoordinate>
,
<quantity id="4C9FC3FAFFFDF349D3D99713DF4AFBC1" box="[1016,1080,1056,1075]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="9" pageNumber="186" unit="m" value="1900.0">
<elevation id="004A892CFFFDF349D3D99713DF4AFBC1" box="[1016,1080,1056,1075]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="9" pageNumber="186" unit="m" value="1900.0">1900m</elevation>
</quantity>
]; 12 \ tibialis; Fvl. \ Coll. et det A. Fauvel;
<taxonomicName id="4C67159CFFFDF349D1F19708D93BFBBC" box="[464,585,1083,1102]" class="Insecta" family="Staphylinidae" genus="Trogophloeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="186" phylum="Arthropoda" rank="genus">Trogophloeus</taxonomicName>
;
<taxonomicName id="4C67159CFFFDF349D2709708D9B1FBBC" authority="Fauv." authorityName="Fauv." box="[593,707,1083,1102]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="186" phylum="Arthropoda" rank="species" species="tibialis">tibialis Fauv.</taxonomicName>
; R.I.Sc.N.B. 17.479 \ Ex-Typis \ Lectotypus;
<taxonomicName id="4C67159CFFFDF349D0509766DB9AFB9A" box="[113,232,1109,1128]" class="Insecta" family="Staphylinidae" genus="Trogophloeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="186" phylum="Arthropoda" rank="genus">Trogophloeus</taxonomicName>
;
<taxonomicName id="4C67159CFFFDF349D0CE9766DA1AFB9A" authority="Fauvel" authorityName="Fauvel" box="[239,360,1109,1128]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="186" phylum="Arthropoda" rank="species" species="tibialis">tibialis Fauvel</taxonomicName>
; des. Makranczy,2001 \
<taxonomicName id="4C67159CFFFDF349D2129766D9DAFB9A" authorityName="Kraatz" authorityYear="1857" box="[563,680,1109,1128]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="186" phylum="Arthropoda" rank="genus">Thinodromus</taxonomicName>
;
<taxonomicName id="4C67159CFFFDF349D28F9766D847FB9A" authority="(Fauvel)" baseAuthorityName="Fauvel" baseAuthorityYear="1907" box="[686,821,1109,1128]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="186" phylum="Arthropoda" rank="species" species="tibialis">tibialis (Fauvel)</taxonomicName>
; det.
<determinerName id="7E0482DBFFFDF349D37E9766D8CDFB9A" box="[863,959,1109,1128]" pageId="9" pageNumber="186">Makranczy</determinerName>
, 2001” (
<collectionCode id="ED76F6DAFFFDF349D4229765DF45FB9B" box="[1027,1079,1110,1129]" pageId="9" pageNumber="186">ISNB</collectionCode>
).
</materialsCitation>
<materialsCitation id="3B0F6442FFFDF349D0509742DAD4FB4B" ID-GBIF-Occurrence="3458217322" collectingDate="1903-12" collectionCode="ISNB, MNHN" collectorName="A. Fauvel" determinerName="Makranczy" latitude="4.1666665" longLatPrecision="1307" longitude="10.166667" pageId="9" pageNumber="186" specimenCount="5" specimenCount-female="2" typeStatus="paralectotype">
<typeStatus id="54DCD0BDFFFDF349D0509742DB8AFB70" box="[113,248,1137,1156]" pageId="9" pageNumber="186" type="paralectotype">PARALECTOTYPES</typeStatus>
(
<specimenCount id="9D61A596FFFDF349D1239743DA1FFB71" box="[258,365,1136,1155]" pageId="9" pageNumber="186" type="generic">3 specimens</specimenCount>
):“Naivasha; 12 \ Coll.et det
<collectorName id="26920BC9FFFDF349D2769743D9D9FB71" box="[599,683,1136,1155]" pageId="9" pageNumber="186">A.Fauvel</collectorName>
;
<taxonomicName id="4C67159CFFFDF349D2939743D85AFB71" box="[690,808,1136,1155]" class="Insecta" family="Staphylinidae" genus="Trogophloeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="186" phylum="Arthropoda" rank="genus">Trogophloeus</taxonomicName>
;
<taxonomicName id="4C67159CFFFDF349D30E9743D8EFFB76" authority="Fauv." authorityName="Fauv." box="[815,925,1136,1156]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="186" phylum="Arthropoda" rank="species" species="tibialis">tibialis Fauv.</taxonomicName>
;
<collectorName id="26920BC9FFFDF349D3839743D8AFFB71" box="[930,989,1136,1155]" pageId="9" pageNumber="186">R.I.Sc.</collectorName>
N.B. 17.479 \ Ex-Typis \
<typeStatus id="54DCD0BDFFFDF349D0FA97B8DA29FB6C" box="[219,347,1163,1182]" pageId="9" pageNumber="186" type="paralectotype">Paralectotypus</typeStatus>
;
<taxonomicName id="4C67159CFFFDF349D14597B8DAAFFB6C" box="[356,477,1163,1182]" class="Insecta" family="Staphylinidae" genus="Trogophloeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="186" phylum="Arthropoda" rank="genus">Trogophloeus</taxonomicName>
;
<taxonomicName id="4C67159CFFFDF349D1C497B8D913FB6C" authority="Fauvel" authorityName="Fauvel" box="[485,609,1163,1182]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="186" phylum="Arthropoda" rank="species" species="tibialis">tibialis Fauvel</taxonomicName>
; des. Makranczy, 2001 \
<taxonomicName id="4C67159CFFFDF349D31497B8D8D9FB6C" authorityName="Kraatz" authorityYear="1857" box="[821,939,1163,1182]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="186" phylum="Arthropoda" rank="genus">Thinodromus</taxonomicName>
;
<taxonomicName id="4C67159CFFFDF349D39297B8DF4FFB6C" authority="(Fauvel)" baseAuthorityName="Fauvel" baseAuthorityYear="1907" box="[947,1085,1163,1182]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="186" phylum="Arthropoda" rank="species" species="tibialis">tibialis (Fauvel)</taxonomicName>
; det.
<determinerName id="7E0482DBFFFDF349D0B79796DB85FB4A" box="[150,247,1189,1208]" pageId="9" pageNumber="186">Makranczy</determinerName>
, 2001” (
<specimenCount id="9D61A596FFFDF349D1619795DA2CFB4A" box="[320,350,1183,1209]" pageId="9" pageNumber="186" type="female">1 ♀</specimenCount>
,
<collectionCode id="ED76F6DAFFFDF349D1489795DAEFFB4B" box="[361,413,1190,1209]" pageId="9" pageNumber="186">ISNB</collectionCode>
),
</materialsCitation>
<materialsCitation id="3B0F6442FFFDF349D18A9796D8A1FB1C" ID-GBIF-Occurrence="3458217310" collectingDate="1903-12" collectionCode="ISNB, MNHN" collectorName="A. Fauvel " determinerName="Makranczy" latitude="4.1666665" longLatPrecision="1307" longitude="10.166667" pageId="9" pageNumber="186" specimenCount="5" specimenCount-female="2" typeStatus="paralectotype">
“Afrique
<superScript id="7C12C357FFFDF349D1DD9795D969FB42" attach="left" box="[508,539,1188,1209]" fontSize="5" pageId="9" pageNumber="186">Orle</superScript>
Anglaise; Naivasha; (Rift-Valley);
<collectorName id="26920BC9FFFDF349D3699796D8DDFB4A" box="[840,943,1189,1208]" pageId="9" pageNumber="186">Ch.Alluaud</collectorName>
<date id="FFD948DFFFFDF349D3959795DF75FB4B" box="[948,1031,1190,1209]" pageId="9" pageNumber="186" value="1903-12">
<collectingDate id="EF9DB137FFFDF349D3959795DF75FB4B" box="[948,1031,1190,1209]" pageId="9" pageNumber="186" value="1903-12">XII. 1903</collectingDate>
</date>
\ Coll. et det
<collectorName id="26920BC9FFFDF349D08397F3DB85FB21" box="[162,247,1216,1235]" pageId="9" pageNumber="186">A. Fauvel</collectorName>
;
<taxonomicName id="4C67159CFFFDF349D0DF97F3DA05FB21" box="[254,375,1216,1235]" class="Insecta" family="Staphylinidae" genus="Trogophloeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="186" phylum="Arthropoda" rank="genus">Trogophloeus</taxonomicName>
;
<taxonomicName id="4C67159CFFFDF349D15F97F3DA9CFB26" authority="Fauv." authorityName="Fauv." box="[382,494,1216,1236]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="186" phylum="Arthropoda" rank="species" species="tibialis">tibialis Fauv.</taxonomicName>
;
<collectorName id="26920BC9FFFDF349D1D597F3D942FB21" box="[500,560,1216,1235]" pageId="9" pageNumber="186">R.I.Sc.</collectorName>
N.B. 17.479 \ Ex-Typis \
<typeStatus id="54DCD0BDFFFDF349D32197F3D8F2FB21" box="[768,896,1216,1235]" pageId="9" pageNumber="186" type="paralectotype">Paralectotypus</typeStatus>
;
<taxonomicName id="4C67159CFFFDF349D3A697F3DF72FB21" box="[903,1024,1216,1235]" class="Insecta" family="Staphylinidae" genus="Trogophloeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="186" phylum="Arthropoda" rank="genus">Trogophloeus</taxonomicName>
;
<taxonomicName id="4C67159CFFFDF349D42697F3DBDCFB1C" authority="Fauvel" authorityName="Fauvel" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="186" phylum="Arthropoda" rank="species" species="tibialis">tibialis Fauvel</taxonomicName>
; des. Makranczy, 2001 \
<taxonomicName id="4C67159CFFFDF349D1A797E8DA8EFB1C" authorityName="Kraatz" authorityYear="1857" box="[390,508,1243,1262]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="186" phylum="Arthropoda" rank="genus">Thinodromus</taxonomicName>
;
<taxonomicName id="4C67159CFFFDF349D22497E8D9E2FB1C" authority="(Fauvel)" baseAuthorityName="Fauvel" baseAuthorityYear="1907" box="[517,656,1243,1262]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="186" phylum="Arthropoda" rank="species" species="tibialis">tibialis (Fauvel)</taxonomicName>
; det.
<determinerName id="7E0482DBFFFDF349D29E97E8D852FB1C" box="[703,800,1243,1262]" pageId="9" pageNumber="186">Makranczy</determinerName>
, 2001” (
<specimenCount id="9D61A596FFFDF349D34A97E8D8F8FB1C" box="[875,906,1237,1262]" pageId="9" pageNumber="186" type="female">1 ♀</specimenCount>
,
<collectionCode id="ED76F6DAFFFDF349D3B797E8D8B8FB1C" box="[918,970,1243,1262]" pageId="9" pageNumber="186">ISNB</collectionCode>
),
</materialsCitation>
<materialsCitation id="3B0F6442FFFDF349D3F897E8D91EFAD1" ID-GBIF-Occurrence="3458217327" collectingDate="1903-12" collectionCode="ISNB, MNHN" collectorName="Jarrige" determinerName="Makranczy" latitude="4.1666665" longLatPrecision="1307" longitude="10.166667" pageId="9" pageNumber="186" specimenCount="5" specimenCount-female="2" typeStatus="paralectotype">
“Afrique or. Anglaise; Naivasha; Rift-Valley [
<geoCoordinate id="EE5308D8FFFDF349D1AD97C5DAA5FAFB" box="[396,471,1269,1289]" degrees="04" direction="north" minutes="10" orientation="latitude" pageId="9" pageNumber="186" precision="925" value="4.1666665">04°10N</geoCoordinate>
,
<geoCoordinate id="EE5308D8FFFDF349D1FE97C5D955FAFB" box="[479,551,1269,1289]" degrees="10" direction="east" minutes="10" orientation="longitude" pageId="9" pageNumber="186" precision="925" value="10.166667">10°10E</geoCoordinate>
];
<collectorName id="26920BC9FFFDF349D21797C6D9ECFAFA" box="[566,670,1269,1288]" pageId="9" pageNumber="186">Ch. Alluaud</collectorName>
<date id="FFD948DFFFFDF349D28397C5D982FAFB" box="[674,752,1270,1289]" pageId="9" pageNumber="186" value="1903-12">
<collectingDate id="EF9DB137FFFDF349D28397C5D982FAFB" box="[674,752,1270,1289]" pageId="9" pageNumber="186" value="1903-12">XII.1903</collectingDate>
</date>
\
<typeStatus id="54DCD0BDFFFDF349D32197C6D8F2FAFA" box="[768,896,1269,1288]" pageId="9" pageNumber="186" type="paralectotype">Paralectotypus</typeStatus>
;
<taxonomicName id="4C67159CFFFDF349D3A997C6D88CFAFA" authorityName="Kraatz" authorityYear="1857" box="[904,1022,1269,1288]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="186" phylum="Arthropoda" rank="genus">Thinodromus</taxonomicName>
;
<taxonomicName id="4C67159CFFFDF349D42697C6DBDCFAD1" authority="Fauvel" authorityName="Fauvel" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="186" phylum="Arthropoda" rank="species" species="tibialis">tibialis Fauvel</taxonomicName>
; des. Makranczy, 2001” (1 ♂, coll.
<collectorName id="26920BC9FFFDF349D1FC9623D96BFAD1" box="[477,537,1296,1315]" pageId="9" pageNumber="186">Jarrige</collectorName>
,
<collectionCode id="ED76F6DAFFFDF349D2009623D914FAD1" box="[545,614,1296,1315]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="186">MNHN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37D3D94FFFDF34AD0509608D8CBFB66" lastPageId="10" lastPageNumber="187" pageId="9" pageNumber="186" type="description">
<paragraph id="8BD86E1FFFFDF34AD0509608D8CBFB66" blockId="9.[113,1089,1339,1588]" lastBlockId="10.[151,1127,220,1588]" lastPageId="10" lastPageNumber="187" pageId="9" pageNumber="186">
<emphasis id="B913B20DFFFDF349D0509608DA68FAA1" bold="true" box="[113,282,1339,1363]" pageId="9" pageNumber="186">Redescription.</emphasis>
<emphasis id="B913B20DFFFDF349D100960EDACCFAA6" box="[289,446,1341,1364]" italics="true" pageId="9" pageNumber="186">Measurements</emphasis>
(in mm, n = 3): HW = 0.49 (0.480.50); TW = 0.47 (0.46 0.47); PW = 0.59 (0.570.60); SW = 0.75 (0.740.76); AW = 0.82 (0.810.83); HL = 0.34 (0.330.35); EL = 0.19 (0.180.20); TL = 0.03 (0.0250.04); PL = 0.42 (0.410.425); SL = 0.73 (0.720.75); SC = 0.70 (0.690.71); FB = 1.55 (1.521.58); BL = 3.04 (2.953.10).
<emphasis id="B913B20DFFFDF349D050968EDA42FA26" box="[113,304,1469,1492]" italics="true" pageId="9" pageNumber="186">Lustre and colour</emphasis>
: Moderately shining, due to setation of body and traces of microsculpture on elytra. Head and abdomen blackish dark brown but apices of tergites (because of more transparency) reddish. Pronotum and elytra reddish dark brown, elytral apex appears somewhat darker. Mouthparts and antennae uniformly medium to dark brown, first antennomere conspicuously brighter, orange/light brown. Legs medium brown, middle of tibiae darkest.
<emphasis id="B913B20DFFFEF34AD0B693CEDA01FEE6" box="[151,371,253,276]" italics="true" pageId="10" pageNumber="187">Shape and sculpture</emphasis>
: Head (
<figureCitation id="135C729AFFFEF34AD1E993CFD965FEE6" box="[456,535,252,276]" captionStart="Figs 2833. 2831" captionStartId="16.[151,188,1545,1564]" captionTargetBox="[151,1126,219,1496]" captionTargetId="figure-10@16.[151,1126,219,1497]" captionTargetPageId="16" captionText="Figs 2833. 2831 Thinodromus tibialis (Fauvel, 1907) (28 head and pronotum; 29 female antenna; 30 male antenna; 31 elytron and abdominal base); 3233 T. gildenkovi sp. nov. (32 head and pronotum; 33 elytron and abdominal base). Scale bar: 0.45 mm for 2831, 0.55 mm for 3233." figureDoi="http://doi.org/10.5281/zenodo.5739659" httpUri="https://zenodo.org/record/5739659/files/figure.png" pageId="10" pageNumber="187">Fig. 28</figureCitation>
) quite small and short, eyes occupy most of side, leaving a small but significant, remarkably rounded temple. Pronotum transverse, first half of sides and anterior corners broadly rounded, posterior constriction of sides towards base a bit stronger than in most congeners, posterior corner more gently rounded, conspicuous. Horseshoe-shaped impression rather deep and sides (while also deep, gradually constricting) obliquely run towards margins at 1/4 of pronotal length (but not reach it). Anterior half of disc centre with three smaller impressions, not well marked. Deflexed margin very thin but apparent along whole length of sides and in the same extent along basal edge. Elytra (
<figureCitation id="135C729AFFFEF34AD41B92EFDBC1FDE6" captionStart="Figs 2833. 2831" captionStartId="16.[151,188,1545,1564]" captionTargetBox="[151,1126,219,1496]" captionTargetId="figure-10@16.[151,1126,219,1497]" captionTargetPageId="16" captionText="Figs 2833. 2831 Thinodromus tibialis (Fauvel, 1907) (28 head and pronotum; 29 female antenna; 30 male antenna; 31 elytron and abdominal base); 3233 T. gildenkovi sp. nov. (32 head and pronotum; 33 elytron and abdominal base). Scale bar: 0.45 mm for 2831, 0.55 mm for 3233." figureDoi="http://doi.org/10.5281/zenodo.5739659" httpUri="https://zenodo.org/record/5739659/files/figure.png" pageId="10" pageNumber="187">Fig. 31</figureCitation>
) combined a little broader than long, dilated posteriorly; with a shallow indefinitely bordered impression posteriad scutellum plus very slight, indefinite, oblique depression from shoulder area to middle of suture. Posterior elytral margin with membranous lobe only apparent in outer half, conspicuously pulled out near outer corners. Apex of abdominal tergite VII with palisade fringe (widest medially).
<emphasis id="B913B20DFFFEF34AD2E8914EDF6FFD66" box="[713,1053,637,660]" italics="true" pageId="10" pageNumber="187">Punctation and microsculpture</emphasis>
: Head with medium large but rather dense punctation, interspaces a fraction of puncture diameters, but shiny. Pronotum with on average slightly larger punctures but also larger interspaces; only appearance of microsculpture the slightly scabrous posterior pronotal corners. Elytra with even larger (and more uniform) punctures, even less dense (except scutellar area and shoulders) than on pronotum. Abdominal terga medially as deeply punctured as elytra but punctures become smaller towards apices; almost without trace of microsculpture in grooves behind basal ridges.
<emphasis id="B913B20DFFFEF34AD1EC906ED93FFC86" box="[461,589,861,884]" italics="true" pageId="10" pageNumber="187">Pubescence</emphasis>
: Whole body with fairly long, semi-erect setation, but only apices of tergites with really long setae. Sizes of setae do not differ greatly on the forebody, but abdominal terga have much finer and smaller setae also near the bases.
<emphasis id="B913B20DFFFEF34AD0B6908ED935FC26" box="[151,583,957,980]" italics="true" pageId="10" pageNumber="187">Primary and secondary sexual features</emphasis>
: Female antennae (
<figureCitation id="135C729AFFFEF34AD301908FD802FC26" box="[800,880,956,980]" captionStart="Figs 2833. 2831" captionStartId="16.[151,188,1545,1564]" captionTargetBox="[151,1126,219,1496]" captionTargetId="figure-10@16.[151,1126,219,1497]" captionTargetPageId="16" captionText="Figs 2833. 2831 Thinodromus tibialis (Fauvel, 1907) (28 head and pronotum; 29 female antenna; 30 male antenna; 31 elytron and abdominal base); 3233 T. gildenkovi sp. nov. (32 head and pronotum; 33 elytron and abdominal base). Scale bar: 0.45 mm for 2831, 0.55 mm for 3233." figureDoi="http://doi.org/10.5281/zenodo.5739659" httpUri="https://zenodo.org/record/5739659/files/figure.png" pageId="10" pageNumber="187">Fig. 29</figureCitation>
) moderately elongate, middle antennomeres (articles 45) roughly as long as wide, penultimate antennomeres (articles 910) slightly transverse. Male antennae (
<figureCitation id="135C729AFFFEF34AD2EA90CFD86EFBE6" box="[715,796,1020,1044]" captionStart="Figs 2833. 2831" captionStartId="16.[151,188,1545,1564]" captionTargetBox="[151,1126,219,1496]" captionTargetId="figure-10@16.[151,1126,219,1497]" captionTargetPageId="16" captionText="Figs 2833. 2831 Thinodromus tibialis (Fauvel, 1907) (28 head and pronotum; 29 female antenna; 30 male antenna; 31 elytron and abdominal base); 3233 T. gildenkovi sp. nov. (32 head and pronotum; 33 elytron and abdominal base). Scale bar: 0.45 mm for 2831, 0.55 mm for 3233." figureDoi="http://doi.org/10.5281/zenodo.5739659" httpUri="https://zenodo.org/record/5739659/files/figure.png" pageId="10" pageNumber="187">Fig. 30</figureCitation>
) just a little bit more slender. Male: MA of aedeagal internal sac elongate, curved, spoon-like, ML less transverse than in other species, BM very elongate, gently broadening medially, BA rather short, more or less straight, AC reverse V-shaped but at tip obtuse-angled, arms curved inwads (
<figureCitation id="135C729AFFFEF34AD3FA976FDF28FB81" box="[987,1114,1115,1140]" captionStart="Figs 5257. 5254" captionStartId="20.[151,188,1520,1539]" captionTargetBox="[154,1102,220,1466]" captionTargetId="figure-10@20.[151,1103,220,1467]" captionTargetPageId="20" captionText="Figs 5257. 5254 Thinodromus rhodesianus (Scheerpeltz, 1974), male (5253 aedeagus; 54 paramere); 5557 T. tibialis (Fauvel, 1907), male (5556 aedeagus; 57 paramere).Abbreviations:AC apical copulatory sclerite; BA basal sclerites, BM basomedial sclerites, MA medioapical sclerites ML medial lamellae. Scale bar: 0.10 mm for 5557, 0.12 mm for 5254." figureDoi="http://doi.org/10.5281/zenodo.5739671" httpUri="https://zenodo.org/record/5739671/files/figure.png" pageId="10" pageNumber="187">Figs 5557</figureCitation>
), sternite VIII (
<figureCitation id="135C729AFFFEF34AD10D974FDA0CFB61" box="[300,382,1147,1172]" captionStart="Figs 5863. 5859" captionStartId="21.[113,150,1545,1564]" captionTargetBox="[115,1063,221,1464]" captionTargetId="figure-10@21.[113,1065,219,1466]" captionTargetPageId="21" captionText="Figs 5863. 5859 Thinodromus tibialis (Fauvel, 1907), male (58 sternite VIII; 59 tergite X); 6061 T. meridionalis sp. nov., male (60 sternite VIII; 61 tergite X); 6263 T. facilis Gildenkov, 2000, male (62 sternite VIII; 63 tergite X). Scale bar: 0.10 mm for 5961 and 63, 0.15 mm for 58 and 62." figureDoi="http://doi.org/10.5281/zenodo.5739673" httpUri="https://zenodo.org/record/5739673/files/figure.png" pageId="10" pageNumber="187">Fig. 58</figureCitation>
), tergite X (
<figureCitation id="135C729AFFFEF34AD220974FD921FB61" box="[513,595,1147,1172]" captionStart="Figs 5863. 5859" captionStartId="21.[113,150,1545,1564]" captionTargetBox="[115,1063,221,1464]" captionTargetId="figure-10@21.[113,1065,219,1466]" captionTargetPageId="21" captionText="Figs 5863. 5859 Thinodromus tibialis (Fauvel, 1907), male (58 sternite VIII; 59 tergite X); 6061 T. meridionalis sp. nov., male (60 sternite VIII; 61 tergite X); 6263 T. facilis Gildenkov, 2000, male (62 sternite VIII; 63 tergite X). Scale bar: 0.10 mm for 5961 and 63, 0.15 mm for 58 and 62." figureDoi="http://doi.org/10.5281/zenodo.5739673" httpUri="https://zenodo.org/record/5739673/files/figure.png" pageId="10" pageNumber="187">Fig. 59</figureCitation>
); female: ringstructure (
<figureCitation id="135C729AFFFEF34AD37A974FD8DEFB66" box="[859,940,1148,1172]" captionStart="Figs 8185. 8183" captionStartId="25.[113,150,1532,1551]" captionTargetBox="[128,1073,222,1462]" captionTargetId="figure-10@25.[124,1076,219,1466]" captionTargetPageId="25" captionText="Figs 8185. 8183 female ringstructures.81 Thinodromus tibialis (Fauvel, 1907); 82 T. dasys Gildenkov, 2000; 83 T. nigerius Gildenkov, 2000. 8485 female ringstructures with spermathecae. 84 T. facilis Gildenkov, 2000; 85 T. gildenkovi sp. nov. Scale bar: 0.10 mm for 85, 0.12 mm for 81 and 8384, 0.14 mm for 82." figureDoi="http://doi.org/10.5281/zenodo.5739681" httpUri="https://zenodo.org/record/5739681/files/figure.png" pageId="10" pageNumber="187">Fig. 81</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37D3D94FFFEF34AD0B697A8D805FA26" pageId="10" pageNumber="187" type="diagnosis">
<paragraph id="8BD86E1FFFFEF34AD0B697A8D805FA26" blockId="10.[151,1127,220,1588]" pageId="10" pageNumber="187">
<emphasis id="B913B20DFFFEF34AD0B697A8DAE2FB41" bold="true" box="[151,400,1179,1203]" pageId="10" pageNumber="187">Differential diagnosis.</emphasis>
<taxonomicName id="4C67159CFFFEF34AD1B497AED900FB46" box="[405,626,1181,1204]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="187" phylum="Arthropoda" rank="species" species="tibialis">
<emphasis id="B913B20DFFFEF34AD1B497AED900FB46" box="[405,626,1181,1204]" italics="true" pageId="10" pageNumber="187">Thinodromus tibialis</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4C67159CFFFEF34AD31097AED8A3FB46" authorityName="Makranczy" authorityYear="2013" box="[817,977,1181,1204]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="187" phylum="Arthropoda" rank="species" species="meridionalis">
<emphasis id="B913B20DFFFEF34AD31097AED8A3FB46" box="[817,977,1181,1204]" italics="true" pageId="10" pageNumber="187">T. meridionalis</emphasis>
</taxonomicName>
, both with the frontal edge of the parameres gently pulled ahead while being evenly rounded, the parameres of these species are unusually and evenly broad. Other shared features include a spoon-like broadened apical formation of MA, conspicuous laterally enlarged flap-like appendages of a sclerite adjacent to the otherwise rather small ML. The most important genitalic differences for both male and female are listed under
<taxonomicName id="4C67159CFFFEF34AD240960ED876FAA6" authority="Makranczy, 2013" authorityName="Makranczy" authorityYear="2013" box="[609,772,1341,1364]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="187" phylum="Arthropoda" rank="species" species="meridionalis" status="sp. nov.">
<emphasis id="B913B20DFFFEF34AD240960ED876FAA6" box="[609,772,1341,1364]" italics="true" pageId="10" pageNumber="187">T. meridionalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2200F76FFFEF34AD32D960FD813FAA1" box="[780,865,1340,1363]" pageId="10" pageNumber="187" rank="species">sp. nov.</taxonomicNameLabel>
Other slight differences are deemed to be due to the specimens being too few, rather old and in less than perfect condition. This is especially true for
<taxonomicName id="4C67159CFFFEF34AD201964ED9F6FA66" box="[544,644,1405,1428]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="187" phylum="Arthropoda" rank="species" species="tibialis">
<emphasis id="B913B20DFFFEF34AD201964ED9F6FA66" box="[544,644,1405,1428]" italics="true" pageId="10" pageNumber="187">T. tibialis</emphasis>
</taxonomicName>
, where all known specimens are more than a 100 years old and plates of the terminalia seem to be with membranous apical edges slightly deformed/shrunk, and do not allow solid conclusions for a diagnosis.
</paragraph>
</subSubSection>
<subSubSection id="C37D3D94FFFEF34AD0B696E8DF3CFA06" box="[151,1102,1499,1524]" pageId="10" pageNumber="187" type="distribution">
<paragraph id="8BD86E1FFFFEF34AD0B696E8DF3CFA06" blockId="10.[151,1127,220,1588]" box="[151,1102,1499,1524]" pageId="10" pageNumber="187">
<emphasis id="B913B20DFFFEF34AD0B696E8DA58FA01" bold="true" box="[151,298,1499,1523]" pageId="10" pageNumber="187">Distribution.</emphasis>
The species is still only known from its original
<typeStatus id="54DCD0BDFFFEF34AD31596EFD810FA06" box="[820,866,1500,1524]" pageId="10" pageNumber="187">type</typeStatus>
material from
<collectingCountry id="F3702E8FFFFEF34AD42096EFDF38FA06" box="[1025,1098,1500,1524]" name="Kenya" pageId="10" pageNumber="187">Kenya</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C37D3D94FFFEF34BD0B696CFD9D9FE41" lastPageId="11" lastPageNumber="188" pageId="10" pageNumber="187" type="discussion">
<paragraph id="8BD86E1FFFFEF34BD0B696CFD9D9FE41" blockId="10.[151,1127,220,1588]" lastBlockId="11.[113,1089,219,435]" lastPageId="11" lastPageNumber="188" pageId="10" pageNumber="187">
<emphasis id="B913B20DFFFEF34AD0B696CFDB8DF9E6" bold="true" box="[151,255,1532,1556]" pageId="10" pageNumber="187">Remarks</emphasis>
. When describing the species, A. Fauvel likely received the series of specimens from Ch. Alluaud with fewer sets of locality labels, and hence hand-copied the collecting data to the specimens he retained in his collection, now deposited in ISNB. The single specimen examined by me in MNHN (being likely part of the original Alluaud collection) does not have any identification label by Fauvel, but clearly bears the original locality label according to which Fauvel prepared the hand-written copies for the specimens now deposited in ISNB. As all specimens are conspecific and there is no contradictory evidence, I consider all mentioned specimens as
<typeStatus id="54DCD0BDFFFFF34BD120924FDA10FE66" box="[257,354,380,404]" pageId="11" pageNumber="188" type="syntype">syntypes</typeStatus>
. Because additional
<typeStatus id="54DCD0BDFFFFF34BD219924FD9E7FE66" box="[568,661,380,404]" pageId="11" pageNumber="188" type="syntype">syntypes</typeStatus>
may still exist, I am designating here the single male from Fauvels collection as the
<typeStatus id="54DCD0BDFFFFF34BD26392A8D9D5FE41" box="[578,679,411,435]" pageId="11" pageNumber="188" type="lectotype">lectotype</typeStatus>
.
</paragraph>
</subSubSection>
</treatment>
</document>