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<document id="47F68E7F8F9E717D5F45DAF96692C02D" ID-CLB-Dataset="7762" ID-DOI="10.5852/ejt.2021.739.1263" ID-GBIF-Dataset="79e0ebb9-6412-4821-879a-9ec760579681" ID-ISSN="2118-9773" ID-Zenodo-Dep="4600344" ID-ZooBank="C333D3B5-7B4B-4A53-8548-E74DFEE9EBC2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615533921823" checkinUser="plazi" docAuthor="Linský, Marek, Čiamporová-Zaťovičová, Zuzana &amp; Čiampor, Fedor" docDate="2021" docId="03B187D7E83CFFD8FD99FD66FA1DFA00" docLanguage="en" docName="ejt-739_1-35.pdf" docOrigin="European Journal of Taxonomy 739 (1)" docSource="http://dx.doi.org/10.5852/ejt.2021.739.1263" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Onychelmis longicollis" docType="treatment" docVersion="8" lastPageNumber="9" masterDocId="FF88FFAFE83AFFD0FFA2FFFFF916FF92" masterDocTitle="A revision of Onychelmis Hinton, 1941 (Coleoptera: Elmidae), with description of new species, DNA barcoding and notes on the geography of the genus" masterLastPageNumber="35" masterPageNumber="1" pageNumber="7" updateTime="1698933319914" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="51910072A31248D1156EC0233A5B66C3">A revision of Onychelmis Hinton, 1941 (Coleoptera: Elmidae), with description of new species, DNA barcoding and notes on the geography of the genus</mods:title>
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<mods:namePart id="6AE3CC43195B3C89987F3903D61CA6EE">Linský, Marek</mods:namePart>
<mods:nameIdentifier id="05750E19CF9644985DCBBEF87D72F4FC" type="LSID">E2AC74B9-FA5F-4D43-9168-4513D3E80A04</mods:nameIdentifier>
<mods:affiliation id="3AFA70881F857FBF41BF4606446D9A67">Zoology Lab, Institute of Botany, Plant Science and Biodiversity Centre, Slovak Academy of Sciences, Dúbravská cesta 9, SK- 84523, Bratislava, Slovakia. &amp; Department of Zoology, Faculty of Natural Science, Comenius University, Mlynská dolina B- 1, SK- 84215 Bratislava, Slovakia.</mods:affiliation>
<mods:nameIdentifier id="C2AB2C7AB85A28C32440A407011A0F54" type="email">marek.linsky@gmail.com</mods:nameIdentifier>
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<mods:namePart id="162970E9DD74F2DFD294A7A850B17A7C">Čiamporová-Zaťovičová, Zuzana</mods:namePart>
<mods:nameIdentifier id="DA2912B3C3B7BD2DB9D6E45877EB3402" type="LSID">8D6A1FDB-06AB-44A7-B1F9-3FC86B7C9804</mods:nameIdentifier>
<mods:affiliation id="EEC9C62A8B92B3F2916A3B22621F6CC2">Zoology Lab, Institute of Botany, Plant Science and Biodiversity Centre, Slovak Academy of Sciences, Dúbravská cesta 9, SK- 84523, Bratislava, Slovakia. &amp; Department of Ecology, Faculty of Natural Science, Comenius University, Mlynská dolina B- 1, SK- 84215 Bratislava, Slovakia.</mods:affiliation>
<mods:nameIdentifier id="6951FF76B3F5B3BAC6FDCF44333C0FAF" type="email">zuzana.zatovicova@savba.sk</mods:nameIdentifier>
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<mods:namePart id="1E1C463D3D402196195317CA880B0E3B">Čiampor, Fedor</mods:namePart>
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<mods:affiliation id="0C813742152E015E1E126B5EE0D1C800">Zoology Lab, Institute of Botany, Plant Science and Biodiversity Centre, Slovak Academy of Sciences, Dúbravská cesta 9, SK- 84523, Bratislava, Slovakia.</mods:affiliation>
<mods:nameIdentifier id="BC84EFB20514DD79BB8544C0BDE2AAB0" type="email">f.ciampor@savba.sk</mods:nameIdentifier>
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<mods:title id="0070AE3DAB3670A342E60543C0B869BB">European Journal of Taxonomy</mods:title>
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<mods:date id="3C4E8EC353D7087D7F512137C2D32A1B">2021</mods:date>
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<mods:number id="A5DE700FCE6DF788AECFA1BFD33DDED8">739</mods:number>
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<mods:classification id="FF0616705DC93E432230039C68C3F4AD">journal article</mods:classification>
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<treatment id="03B187D7E83CFFD8FD99FD66FA1DFA00" ID-DOI="http://doi.org/10.5281/zenodo.4602495" ID-GBIF-Taxon="180331470" ID-Zenodo-Dep="4602495" LSID="urn:lsid:plazi:treatment:03B187D7E83CFFD8FD99FD66FA1DFA00" httpUri="http://treatment.plazi.org/id/03B187D7E83CFFD8FD99FD66FA1DFA00" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7">
<subSubSection id="C302654AE83CFFD6FD99FD66FAEEFD21" box="[571,1016,665,692]" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="8BA736C1E83CFFD6FD99FD66FAEEFD21" blockId="6.[571,1016,665,727]" box="[571,1016,665,692]" pageId="6" pageNumber="7">
<heading id="D0EF81ADE83CFFD6FD99FD66FAEEFD21" box="[571,1016,665,692]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="7" reason="2">
<taxonomicName id="4C184D42E83CFFD6FD99FD66FAEEFD21" ID-CoL="49QNX" authority="(Sharp, 1882)" baseAuthorityName="Sharp" baseAuthorityYear="1882" box="[571,1016,665,692]" class="Insecta" family="Elmidae" genus="Onychelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="longicollis">
<emphasis id="B96CEAD3E83CFFD6FD99FD66FA46FD26" bold="true" box="[571,848,665,692]" italics="true" pageId="6" pageNumber="7">Onychelmis longicollis</emphasis>
(
<bibRefCitation id="EF894B30E83CFFD6FCFDFD66FAE6FD21" author="Sharp D." box="[863,1008,665,691]" pageId="6" pageNumber="7" pagination="1 - 144" refId="ref17769" refString="Sharp D. 1882. Insecta, Coleoptera: Haliplidae, Dytiscidae, Gyrinidae, Hydrophilidae, Heteroceridae, Parnidae, Georissidae, Cyathoceridae. Biologia Centrali-Americana 1 (2): 1 - 144." type="journal article" year="1882">Sharp, 1882</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C302654AE83CFFD6FDD4FD43FAA8FD45" box="[630,958,700,727]" pageId="6" pageNumber="7" type="description">
<paragraph id="8BA736C1E83CFFD6FDD4FD43FAA8FD45" blockId="6.[571,1016,665,727]" box="[630,958,700,727]" pageId="6" pageNumber="7">
<figureCitation id="13232A44E83CFFD6FDD4FD43FBA8FD45" box="[630,702,700,727]" captionStart="Fig" captionStartId="7.[188,231,1928,1954]" captionTargetBox="[248,1355,704,1872]" captionTargetId="figure-147@7.[189,1398,684,1890]" captionTargetPageId="7" captionText="Fig. 2. Habitus of Onychelmis spp. A–B. O. longicollis (Sharp, 1882), holotype, ♂ (NHM). A. Dorsal view. B. Ventral view. C–D. O. whiteheadi Spangler &amp; Santiago, 1991, holotype, ♂ (NMNH). C. Dorsal view. D. Ventral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4600350" httpUri="https://zenodo.org/record/4600350/files/figure.png" pageId="6" pageNumber="7">Figs 2</figureCitation>
A–B, 6D, 7C–D, 12B
</paragraph>
</subSubSection>
<subSubSection id="C302654AE83CFFD6FF1FFD01FA56FCEF" pageId="6" pageNumber="7" type="reference_group">
<paragraph id="8BA736C1E83CFFD6FF1FFD01FB4DFC8A" blockId="6.[189,603,766,792]" box="[189,603,766,792]" pageId="6" pageNumber="7">
<treatmentCitationGroup id="AB0811EFE83CFFD6FF1FFD01FB4DFC8A" box="[189,603,766,792]" pageId="6" pageNumber="7">
<treatmentCitation id="0AB910D0E83CFFD6FF1FFD01FB41FC8A" author="Sharp D." box="[189,599,766,792]" page="138" pageId="6" pageNumber="7" year="1882">
<taxonomicName id="4C184D42E83CFFD6FF1FFD01FB41FC8A" authority="Sharp, 1882: 138" authorityName="Sharp" authorityPageNumber="138" authorityYear="1882" box="[189,599,766,792]" class="Insecta" family="Elmidae" genus="Elmis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="longicollis">
<emphasis id="B96CEAD3E83CFFD6FF1FFD01F895FC8A" box="[189,387,766,792]" italics="true" pageId="6" pageNumber="7">Elmis longicollis</emphasis>
<bibRefCitation id="EF894B30E83CFFD6FE28FD01FB41FC8A" author="Sharp D." box="[394,599,766,792]" pageId="6" pageNumber="7" pagination="1 - 144" refId="ref17769" refString="Sharp D. 1882. Insecta, Coleoptera: Haliplidae, Dytiscidae, Gyrinidae, Hydrophilidae, Heteroceridae, Parnidae, Georissidae, Cyathoceridae. Biologia Centrali-Americana 1 (2): 1 - 144." type="journal article" year="1882">Sharp, 1882: 138</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA736C1E83CFFD6FF1FFCC0FA56FCEF" blockId="6.[189,1398,831,893]" pageId="6" pageNumber="7">
<treatmentCitationGroup id="AB0811EFE83CFFD6FF1FFCC0FA56FCEF" pageId="6" pageNumber="7">
<taxonomicName id="4C184D42E83CFFD6FF1FFCC0F8DBFCCB" ID-CoL="49QNX" baseAuthorityName="Sharp" baseAuthorityYear="1882" box="[189,461,831,857]" class="Insecta" family="Elmidae" genus="Onychelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="longicollis">
<emphasis id="B96CEAD3E83CFFD6FF1FFCC0F8DBFCCB" box="[189,461,831,857]" italics="true" pageId="6" pageNumber="7">Onychelmis longicollis</emphasis>
</taxonomicName>
–
<treatmentCitation id="0AB910D0E83CFFD6FE53FCC0FBAEFCCB" author="Hinton H. E." box="[497,696,831,858]" page="67" pageId="6" pageNumber="7" year="1941">
<bibRefCitation id="EF894B30E83CFFD6FE53FCC0FBAEFCCB" author="Hinton H. E." box="[497,696,831,858]" pageId="6" pageNumber="7" pagination="65 - 104" refId="ref16569" refString="Hinton H. E. 1941. New genera and species of Elmidae (Coleoptera). Transactions of the Royal Entomological Society of London 91 (3): 65 - 104. https: // doi. org / 10.1111 / j. 1365 - 2311.1941. tb 01044. x" type="journal article" year="1941">Hinton 1941: 67</bibRefCitation>
</treatmentCitation>
, figs 7–8. —
<treatmentCitation id="0AB910D0E83CFFD6FCFDFCC0FDCCFCC8" author="Spangler P. &amp; Santiago S." box="[863,1242,831,858]" page="497" pageId="6" pageNumber="7" year="1991">
<bibRefCitation id="EF894B30E83CFFD6FCFDFCC0FDCCFCC8" author="Spangler P. &amp; Santiago S." box="[863,1242,831,858]" pageId="6" pageNumber="7" pagination="495 - 498" refId="ref17990" refString="Spangler P. &amp; Santiago S. 1991. A new species and new records from Colombia of the water beetle genus Onychelmis Hinton (Coleoptera: Elmidae: Elminae). Proceedings of the Entomological Society of Washington 93 (2): 495 - 498." type="journal article" year="1991">
Spangler &amp;
<collectingRegion id="49DCF823E83CFFD6FC4DFCC0FD40FCCB" box="[1007,1110,831,857]" country="Chile" name="Region Metropolitana" pageId="6" pageNumber="7">Santiago</collectingRegion>
1991: 497
</bibRefCitation>
</treatmentCitation>
, figs 3–4. —
<treatmentCitation id="0AB910D0E83CFFD6FF48FC9CFB6FFCEF" author="Gonzalez-Cordoba M. &amp; Zuniga M. C. &amp; Manzo V. &amp; Giraldo L. P. &amp; Chara J." box="[234,633,867,893]" page="11" pageId="6" pageNumber="7" year="2016">
<bibRefCitation id="EF894B30E83CFFD6FF48FC9CFB6FFCEF" author="Gonzalez-Cordoba M. &amp; Zuniga M. C. &amp; Manzo V. &amp; Giraldo L. P. &amp; Chara J." box="[234,633,867,893]" pageId="6" pageNumber="7" pagination="1 - 17" refId="ref16402" refString="Gonzalez-Cordoba M., Zuniga M. C., Manzo V., Giraldo L. P. &amp; Chara J. 2016. Notelmis Hinton y Onychelmis Hinton (Coleoptera: Elmidae: Elminae) de Colombia: Claves taxonomicas ilustradas. Boletin del Museo de Entomologia de la Universidad del Valle 16 (2): 1 - 17." type="journal article" year="2016">
González-Córdoba
<emphasis id="B96CEAD3E83CFFD6FE6DFC9CFB1CFCEF" box="[463,522,867,893]" italics="true" pageId="6" pageNumber="7">et al.</emphasis>
2016: 11
</bibRefCitation>
</treatmentCitation>
, figs 5a–b, 7c–b.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C302654AE83CFFD6FF1FFC5BF845FBE8" pageId="6" pageNumber="7" type="diagnosis">
<paragraph id="8BA736C1E83CFFD6FF1FFC5BF8D0FC2C" blockId="6.[189,454,932,958]" box="[189,454,932,958]" pageId="6" pageNumber="7">
<heading id="D0EF81ADE83CFFD6FF1FFC5BF8D0FC2C" bold="true" box="[189,454,932,958]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="B96CEAD3E83CFFD6FF1FFC5BF8D0FC2C" bold="true" box="[189,454,932,958]" pageId="6" pageNumber="7">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BA736C1E83CFFD6FF1FFC2CF845FBE8" blockId="6.[188,1399,979,1146]" pageId="6" pageNumber="7">
<taxonomicName id="4C184D42E83CFFD6FF1FFC2CF8DBFC7F" baseAuthorityName="Sharp" baseAuthorityYear="1882" box="[189,461,979,1005]" class="Insecta" family="Elmidae" genus="Onychelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="longicollis">
<emphasis id="B96CEAD3E83CFFD6FF1FFC2CF8DBFC7F" box="[189,461,979,1005]" italics="true" pageId="6" pageNumber="7">Onychelmis longicollis</emphasis>
</taxonomicName>
can be distinguished from all species of the genus by the combination of the following characters: 1) smaller size (CL:
<quantity id="4CE09B24E83CFFD6FD6CFC09FA54FB83" box="[718,834,1014,1041]" metricMagnitude="-3" metricUnit="m" metricValue="1.27" pageId="6" pageNumber="7" unit="mm" value="1.27">1.27 mm</quantity>
); 2) tomentum on profemora reaching behind middle, on mesofemora to around middle (
<figureCitation id="13232A44E83CFFD6FD0FFBE6FA1CFBA6" box="[685,778,1049,1076]" captionStart="Fig" captionStartId="18.[189,232,1893,1919]" captionTargetBox="[199,1389,1223,1846]" captionTargetId="figure-257@18.[189,1398,1213,1855]" captionTargetPageId="18" captionText="Fig. 6. Tomentum on pro- and mesofemora of Onychelmis spp. A. O. leleupi Delève, 1968. B. O. lenkae sp. nov. C. O. lobata sp. nov. D. O. longicollis (Sharp, 1882). E. O. minor sp. nov. F. O. onorei sp. nov. G. O. splendida sp. nov. H. O. whiteheadi Spangler &amp; Santiago, 1991. Drawings are schematic, bodies are divided to identical halves: left = male, right = female (note: females unknown for D and H)." figureDoi="http://doi.org/10.5281/zenodo.4600362" httpUri="https://zenodo.org/record/4600362/files/figure.png" pageId="6" pageNumber="7">Fig. 6D</figureCitation>
); 3) humeri feebly developed; 4) prominent carina on sixth elytral interval present; 5) elytral punctures moderately impressed; 6) shape of aedeagus lanceolate (
<figureCitation id="13232A44E83CFFD6FF67FB9FF81FFBE8" box="[197,265,1120,1146]" captionStart="Fig" captionStartId="20.[189,232,1926,1952]" captionTargetBox="[252,1337,285,1878]" captionTargetId="figure-16@20.[234,1353,265,1886]" captionTargetPageId="20" captionText="Fig. 7. Aedeagi of Onychelmis spp. A. O. whiteheadi Spangler &amp; Santiago, 1991, ventral view. B. O. whiteheadi, lateral view. C. O. longicollis (Sharp, 1882), ventral view. D. O. longicollis, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4600366" httpUri="https://zenodo.org/record/4600366/files/figure.png" pageId="6" pageNumber="7">Fig. 7</figureCitation>
C–D).
</paragraph>
</subSubSection>
<subSubSection id="C302654AE83CFFD6FF1FFB5EFC61FAA3" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="8BA736C1E83CFFD6FF1FFB5EF8BCFB29" blockId="6.[189,426,1185,1211]" box="[189,426,1185,1211]" pageId="6" pageNumber="7">
<heading id="D0EF81ADE83CFFD6FF1FFB5EF8BCFB29" bold="true" box="[189,426,1185,1211]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="B96CEAD3E83CFFD6FF1FFB5EF8BCFB29" bold="true" box="[189,426,1185,1211]" pageId="6" pageNumber="7">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BA736C1E83CFFD6FF48FB2FFC61FAA3" blockId="6.[189,1399,1232,1329]" pageId="6" pageNumber="7">
<materialsCitation id="3B703C9CE83CFFD6FF48FB2FFC61FAA3" ID-GBIF-Occurrence="3051860309" collectionCode="NHM" collectorName="B. C. A. Col." country="Panama" elevation="915" location="Champion." municipality="Champion" pageId="6" pageNumber="7" specimenCount="1" stateProvince="Chiriqui" typeStatus="holotype">
<emphasis id="B96CEAD3E83CFFD6FF48FB2FF84AFB78" bold="true" box="[234,348,1232,1258]" pageId="6" pageNumber="7">
<typeStatus id="54A38863E83CFFD6FF48FB2FF84AFB78" box="[234,348,1232,1258]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F30F7651E83CFFD6FF1FFB0CF82EFA9F" box="[189,312,1267,1293]" name="Panama" pageId="6" pageNumber="7">PANAMA</collectingCountry>
•
<specimenCount id="9D1EFD48E83CFFD6FEF7FB0BF87BFA9C" box="[341,365,1268,1294]" pageId="6" pageNumber="7" type="male">♂</specimenCount>
; “
<taxonomicName id="4C184D42E83CFFD6FE24FB0CFB45FA9F" box="[390,595,1267,1293]" class="Insecta" family="Elmidae" genus="Elmis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="longicollis">Elmis longicollis</taxonomicName>
Type D.S. V.de
<collectingRegion id="49DCF823E83CFFD6FCBBFB0CFA6DFA9F" box="[793,891,1267,1293]" country="Panama" name="Chiriqui" pageId="6" pageNumber="7">Chiriqui</collectingRegion>
<quantity id="4CE09B24E83CFFD6FC26FB0CFAE7FA9F" box="[900,1009,1267,1293]" metricMagnitude="2" metricUnit="m" metricValue="9.150096" metricValueMax="18.288" metricValueMin="0.012192" pageId="6" pageNumber="7" unit="ft" value="3002.0" valueMax="6000.0" valueMin="4.0">
<elevation id="0035D1F2E83CFFD6FC26FB0CFAE7FA9F" box="[900,1009,1267,1293]" metricMagnitude="2" metricUnit="m" metricValue="9.150096" metricValueMax="18.288" metricValueMin="0.012192" pageId="6" pageNumber="7" unit="ft" value="3002.0" valueMax="6000.0" valueMin="4.0">4-6000ft.</elevation>
</quantity>
<collectingRegion id="49DCF823E83CFFD6FC58FB0BFD4DFA9C" box="[1018,1115,1268,1294]" country="Panama" name="Panama" pageId="6" pageNumber="7">Panama</collectingRegion>
.
<collectingMunicipality id="6BC3ACBBE83CFFD6FBC5FB0CFDF7FA9F" box="[1127,1249,1267,1293]" pageId="6" pageNumber="7">Champion</collectingMunicipality>
// ♂ // Type [red disc] // V. de
<collectingRegion id="49DCF823E83CFFD6FE2AFAE9F8FCFAA2" box="[392,490,1302,1328]" country="Panama" name="Chiriqui" pageId="6" pageNumber="7">Chiriqui</collectingRegion>
<quantity id="4CE09B24E83CFFD6FE52FAE9FB99FAA2" box="[496,655,1302,1328]" metricMagnitude="3" metricUnit="m" metricValue="1.524" metricValueMax="1.8288" metricValueMin="1.2192" pageId="6" pageNumber="7" unit="ft" value="5000.0" valueMax="6000.0" valueMin="4000.0">
<elevation id="0035D1F2E83CFFD6FE52FAE9FB99FAA2" box="[496,655,1302,1328]" metricMagnitude="3" metricUnit="m" metricValue="1.524" metricValueMax="1.8288" metricValueMin="1.2192" pageId="6" pageNumber="7" unit="ft" value="5000.0" valueMax="6000.0" valueMin="4000.0">4000-6000 ft.</elevation>
</quantity>
<location id="8EC7601AE83CFFD6FD37FAE9FA01FAA2" LSID="urn:lsid:plazi:treatment:03B187D7E83CFFD8FD99FD66FA1DFA00:8EC7601AE83CFFD6FD37FAE9FA01FAA2" box="[661,791,1302,1328]" country="Panama" municipality="Champion" name="Champion." pageId="6" pageNumber="7" stateProvince="Chiriqui">Champion.</location>
//
<collectorName id="26ED5317E83CFFD6FC91FAE9FAA8FAA2" box="[819,958,1302,1328]" pageId="6" pageNumber="7">B.C.A. Col.</collectorName>
<date id="FFA61001E83CFFD6FC61FAE8FAE6FAA2" box="[963,1008,1302,1329]" pageId="6" pageNumber="7">I. 2.</date>
<taxonomicName id="4C184D42E83CFFD6FC55FAE9FC03FAA2" authority="Sharp." authorityName="Sharp." box="[1015,1301,1302,1328]" class="Insecta" family="Elmidae" genus="Elmis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="longicollis">Elmis longicollis Sharp.</taxonomicName>
”;
<collectionCode id="ED09AE04E83CFFD6FA89FAE8FC65FAA3" box="[1323,1395,1303,1329]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="6" pageNumber="7">NHM</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C302654AE83CFFD8FF1FFAA8F82FFC32" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7" type="description">
<paragraph id="8BA736C1E83CFFD6FF1FFAA8F87DFAE3" blockId="6.[189,363,1367,1393]" box="[189,363,1367,1393]" pageId="6" pageNumber="7">
<heading id="D0EF81ADE83CFFD6FF1FFAA8F87DFAE3" bold="true" box="[189,363,1367,1393]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="B96CEAD3E83CFFD6FF1FFAA8F87DFAE3" bold="true" box="[189,363,1367,1393]" pageId="6" pageNumber="7">Redescription</emphasis>
</heading>
</paragraph>
<paragraph id="8BA736C1E83CFFD6FF48FA79F896F9BF" blockId="6.[189,1399,1414,1581]" pageId="6" pageNumber="7">
<emphasis id="B96CEAD3E83CFFD6FF48FA79F83CFA32" bold="true" box="[234,298,1414,1440]" pageId="6" pageNumber="7">Male</emphasis>
(
<typeStatus id="54A38863E83CFFD6FE9BFA79F8B5FA33" box="[313,419,1414,1441]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
) BODY. Obovate (
<figureCitation id="13232A44E83CFFD6FE27FA56F8DBFA56" box="[389,461,1449,1476]" captionStart="Fig" captionStartId="7.[188,231,1928,1954]" captionTargetBox="[248,1355,704,1872]" captionTargetId="figure-147@7.[189,1398,684,1890]" captionTargetPageId="7" captionText="Fig. 2. Habitus of Onychelmis spp. A–B. O. longicollis (Sharp, 1882), holotype, ♂ (NHM). A. Dorsal view. B. Ventral view. C–D. O. whiteheadi Spangler &amp; Santiago, 1991, holotype, ♂ (NMNH). C. Dorsal view. D. Ventral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4600350" httpUri="https://zenodo.org/record/4600350/files/figure.png" pageId="6" pageNumber="7">Fig. 2</figureCitation>
A–B); length
<quantity id="4CE09B24E83CFFD6FDD1FA55FBF4FA56" box="[627,738,1450,1476]" metricMagnitude="-3" metricUnit="m" metricValue="1.27" pageId="6" pageNumber="7" unit="mm" value="1.27">1.27 mm</quantity>
; width
<quantity id="4CE09B24E83CFFD6FC9FFA56FABAFA56" box="[829,940,1449,1476]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="7" unit="mm" value="0.6">0.60 mm</quantity>
; dorsum conveX, glabrous. Body with sparse, light yellowish setae. Hairy or scale-like tomentum distributed on following areas: genae, sides of prosternum, mesoventrite, metaventrite and abdomen, epipleura, and medial and lateral portions of bases of femora.
</paragraph>
<paragraph id="8BA736C1E83CFFD6FF1FF9AAFD1CF900" blockId="6.[188,1398,1621,1682]" pageId="6" pageNumber="7">COLOUR. Head, pronotum and elytra black; venter dark brown with reddish tinge; trochanters, tibiae and tarsi brown; first two segments of antennae and tarsal claws pale brown.</paragraph>
<paragraph id="8BA736C1E83CFFD6FF1FF945FDBEF816" blockId="6.[188,1399,1721,1924]" pageId="6" pageNumber="7">
HEAD. Partly retractable into prothoraX. Antennae with only 2 segments remaining; pedicel about twice as long as scape. Labrum with anterior margin almost straight; anterolateral angles broadly arcuate with numerous golden, recumbent hair-like setae; clypeus shorter and wider than labrum, about 3.5 times as wide as long; anterior margin slightly concave, anterolateral angles rounded; frontoclypeal suture almost straight. Eyes well developed, HW:
<quantity id="4CE09B24E83CFFD6FDCCF8B9FBCAF8F3" box="[622,732,1862,1889]" metricMagnitude="-4" metricUnit="m" metricValue="2.9" pageId="6" pageNumber="7" unit="mm" value="0.29">0.29 mm</quantity>
, ID:
<quantity id="4CE09B24E83CFFD6FCB9F8B9FA9FF8F3" box="[795,905,1862,1889]" metricMagnitude="-4" metricUnit="m" metricValue="1.4" pageId="6" pageNumber="7" unit="mm" value="0.14">0.14 mm</quantity>
, suboval in lateral view, protruding from head outline in dorsal view, circumocular surface raised. Frons convex between eyes.
</paragraph>
<paragraph id="8BA736C1E83CFFD8FF1FF854FD06FDD1" blockId="6.[189,1399,1963,2025]" lastBlockId="8.[188,1400,265,580]" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7">
THORAX. Pronotum widest behind middle, PW:
<quantity id="4CE09B24E83CFFD6FCA1F854FA64F854" box="[771,882,1963,1990]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" pageId="6" pageNumber="7" unit="mm" value="0.41">0.41 mm</quantity>
, PL:
<quantity id="4CE09B24E83CFFD6FC14F854FD33F854" box="[950,1061,1963,1990]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="6" pageNumber="7" unit="mm" value="0.43">0.43 mm</quantity>
; surface shiny, with narrow reticulation along basal margin and posterolateral angles, with dense tiny punctures; sublateral carinae absent, fine raised line indistinct; disc conveX, divided by a broad, deep transverse impression before middle; two prescutellar foveae separated by a raised line extending from base to apical discal half, connecting both halves and merging into them; anterior margin arcuate; posterior margin bisinuate; sides of pronotum convex before and after transverse constriction; lateral margins narrowly rimmed; posterolateral angles orthogonal;anterolateral angles slightly produced.Hypomeron finely microreticulate, widest in middle. Prosternum moderately long in front of procoXae, with anterior margin concave; sides raised around procoxae, forming carinae, not reaching anterior margin, prosternal process long, moderately broad and with posterior margin broadly rounded. Mesoventrite coarse, short and wide, with deep triangular groove for reception of prosternal process; posterior margin around mesocoxae raised. Metaventrite slightly wider than long, rugose, more or less shiny with setigerous punctures; disc convex with deep, medial, triangular depression in posterior half; discrimen in basal ¾; with one prebasal fovea on each side of metaventrite. Elytra (EL:
<quantity id="4CE09B24E832FFD8FD0BFED2FA03FEDA" box="[681,789,301,328]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" pageId="8" pageNumber="9" unit="mm" value="0.85">0.85 mm</quantity>
, EW:
<quantity id="4CE09B24E832FFD8FCFDFED2FADCFEDA" box="[863,970,301,328]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="8" pageNumber="9" unit="mm" value="0.6">0.60 mm</quantity>
) conveX, widest in anterior ¾; sides strongly declivous; surface shiny, with dense tiny punctures; elytral margin narrowly rimmed; humeri feebly developed; epipleuron tapering posteriorly. Prominent carina on siXth interval well developed, reaching ⅘ of elytron. Elytra with ten rows of small punctures, moderately impressed, separated by 1–2 times puncture diameter, diminishing toward lateral margins and elytral apeX. Scutellum subovate, flat. Legs moderately long; femora clavate; tibiae longest, apically with cleaning fringes. Tarsi 5-segmented, first four segments each with one fine pale, recumbent seta, fifth segment slightly shorter than remaining segments combined; claws with a large subbasal and smaller basal teeth.
</paragraph>
<caption id="DF676649E83DFFD7FF1EF877FBA3F87B" ID-DOI="http://doi.org/10.5281/zenodo.4600350" ID-Zenodo-Dep="4600350" httpUri="https://zenodo.org/record/4600350/files/figure.png" pageId="7" pageNumber="8" startId="7.[188,231,1928,1954]" targetBox="[248,1355,704,1872]" targetPageId="7">
<paragraph id="8BA736C1E83DFFD7FF1EF877FBA3F87B" blockId="7.[188,1398,1928,2025]" pageId="7" pageNumber="8">
<emphasis id="B96CEAD3E83DFFD7FF1EF877F81DF830" bold="true" box="[188,267,1928,1954]" pageId="7" pageNumber="8">Fig. 2.</emphasis>
Habitus of
<taxonomicName id="4C184D42E83DFFD7FE38F877FB32F830" authorityName="Hinton" authorityYear="1941" box="[410,548,1928,1954]" class="Insecta" family="Elmidae" genus="Onychelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B96CEAD3E83DFFD7FE38F877FB32F830" box="[410,548,1928,1954]" italics="true" pageId="7" pageNumber="8">Onychelmis</emphasis>
</taxonomicName>
spp.
<emphasis id="B96CEAD3E83DFFD7FDC6F877FB8AF830" bold="true" box="[612,668,1928,1955]" pageId="7" pageNumber="8">A–B</emphasis>
.
<taxonomicName id="4C184D42E83DFFD7FD09F877FAEDF830" authority="(Sharp, 1882)" baseAuthorityName="Sharp" baseAuthorityYear="1882" box="[683,1019,1928,1954]" class="Insecta" family="Elmidae" genus="Elmis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="longicollis">
<emphasis id="B96CEAD3E83DFFD7FD09F877FA5AF830" box="[683,844,1928,1954]" italics="true" pageId="7" pageNumber="8">O. longicollis</emphasis>
(
<bibRefCitation id="EF894B30E83DFFD7FCFCF877FAE5F830" author="Sharp D." box="[862,1011,1928,1954]" pageId="7" pageNumber="8" pagination="1 - 144" refId="ref17769" refString="Sharp D. 1882. Insecta, Coleoptera: Haliplidae, Dytiscidae, Gyrinidae, Hydrophilidae, Heteroceridae, Parnidae, Georissidae, Cyathoceridae. Biologia Centrali-Americana 1 (2): 1 - 144." type="journal article" year="1882">Sharp, 1882</bibRefCitation>
)
</taxonomicName>
, holotype, ♂ (NHM).
<emphasis id="B96CEAD3E83DFFD7FAA1F877FC0EF830" bold="true" box="[1283,1304,1928,1954]" pageId="7" pageNumber="8">A</emphasis>
. Dorsal view.
<emphasis id="B96CEAD3E83DFFD7FEA0F854F800F857" bold="true" box="[258,278,1963,1989]" pageId="7" pageNumber="8">B</emphasis>
. Ventral view.
<emphasis id="B96CEAD3E83DFFD7FE60F854F8EDF857" bold="true" box="[450,507,1963,1990]" pageId="7" pageNumber="8">C–D</emphasis>
.
<taxonomicName id="4C184D42E83DFFD7FDABF853FAF9F857" authority="Spangler &amp; Santiago, 1991" authorityName="Spangler &amp; Santiago" authorityYear="1991" box="[521,1007,1963,1990]" class="Insecta" family="Elmidae" genus="Onychelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="whiteheadi">
<emphasis id="B96CEAD3E83DFFD7FDABF853FBBAF857" box="[521,684,1963,1990]" italics="true" pageId="7" pageNumber="8">O. whiteheadi</emphasis>
<bibRefCitation id="EF894B30E83DFFD7FD13F854FAF9F857" author="Spangler P. &amp; Santiago S." box="[689,1007,1963,1989]" pageId="7" pageNumber="8" pagination="495 - 498" refId="ref17990" refString="Spangler P. &amp; Santiago S. 1991. A new species and new records from Colombia of the water beetle genus Onychelmis Hinton (Coleoptera: Elmidae: Elminae). Proceedings of the Entomological Society of Washington 93 (2): 495 - 498." type="journal article" year="1991">Spangler &amp; Santiago, 1991</bibRefCitation>
</taxonomicName>
, holotype, ♂ (NMNH).
<emphasis id="B96CEAD3E83DFFD7FAA7F854FC0CF857" bold="true" box="[1285,1306,1963,1989]" pageId="7" pageNumber="8">C</emphasis>
. Dorsal view.
<emphasis id="B96CEAD3E83DFFD7FEA1F831F80EF87A" bold="true" box="[259,280,1998,2024]" pageId="7" pageNumber="8">D</emphasis>
. Ventral view. Scale bar = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BA736C1E832FFD8FF1FFD94F85BFCAB" blockId="8.[189,1399,619,825]" pageId="8" pageNumber="9">
ABDOMEN. With 5 ventrites. First ventrite with basal margin broadly rounded; fifth ventrite longest, apically setose, with posterior margin broadly arcuate. Aedeagus (
<figureCitation id="13232A44E832FFD8FC72FD70FD00FD3B" box="[976,1046,655,681]" captionStart="Fig" captionStartId="20.[189,232,1926,1952]" captionTargetBox="[252,1337,285,1878]" captionTargetId="figure-16@20.[234,1353,265,1886]" captionTargetPageId="20" captionText="Fig. 7. Aedeagi of Onychelmis spp. A. O. whiteheadi Spangler &amp; Santiago, 1991, ventral view. B. O. whiteheadi, lateral view. C. O. longicollis (Sharp, 1882), ventral view. D. O. longicollis, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4600366" httpUri="https://zenodo.org/record/4600366/files/figure.png" pageId="8" pageNumber="9">Fig. 7</figureCitation>
C–D) elongate. Penis without fibula; corona in apical half; in ventral view conveX, from apical half tapering toward broad rounded apeX; in lateral view slender, evenly narrowed from base to slightly curved apeX. Parameres absent. Phallobase about 1.5 times as long as penis, subparallel, in ventral view straight, slightly curved in lateral view.
</paragraph>
<paragraph id="8BA736C1E832FFD8FF48FC9DF853FCEE" blockId="8.[189,325,866,928]" box="[234,325,866,892]" pageId="8" pageNumber="9">
<emphasis id="B96CEAD3E832FFD8FF48FC9DF853FCEE" bold="true" box="[234,325,866,892]" pageId="8" pageNumber="9">Female</emphasis>
</paragraph>
<paragraph id="8BA736C1E832FFD8FF1FFC79F82FFC32" blockId="8.[189,325,866,928]" box="[189,313,902,928]" pageId="8" pageNumber="9">Unknown.</paragraph>
</subSubSection>
<subSubSection id="C302654AE832FFD8FF1FFC37FC10FB33" pageId="8" pageNumber="9" type="distribution">
<paragraph id="8BA736C1E832FFD8FF1FFC37F840FC70" blockId="8.[189,342,968,994]" box="[189,342,968,994]" pageId="8" pageNumber="9">
<heading id="D0EF81ADE832FFD8FF1FFC37F840FC70" bold="true" box="[189,342,968,994]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="B96CEAD3E832FFD8FF1FFC37F840FC70" bold="true" box="[189,342,968,994]" pageId="8" pageNumber="9">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BA736C1E832FFD8FF1FFC08FC10FB33" blockId="8.[189,1399,1015,1186]" pageId="8" pageNumber="9">
The species is reported from three countries (
<figureCitation id="13232A44E832FFD8FD4BFC08FA4EFB83" box="[745,856,1015,1041]" captionStart="Fig" captionStartId="28.[189,232,1916,1942]" captionTargetBox="[189,1398,1158,1876]" captionTargetId="figure-274@28.[189,1398,1158,1876]" captionTargetPageId="28" captionText="Fig. 12. Distribution map of Onychelmis Hinton, 1941. A. New species described within this study. B. Published records of Onychelmis and potential distribution of the genus." figureDoi="http://doi.org/10.5281/zenodo.4600376" httpUri="https://zenodo.org/record/4600376/files/figure.png" pageId="8" pageNumber="9">Fig. 12B</figureCitation>
),
<collectingCountry id="F30F7651E832FFD8FCD3FC08FAF3FB83" box="[881,997,1015,1041]" name="Colombia" pageId="8" pageNumber="9">Colombia</collectingCountry>
(
<bibRefCitation id="EF894B30E832FFD8FC55FC08FC7CFB80" author="Gonzalez-Cordoba M. &amp; Zuniga M. C. &amp; Manzo V. &amp; Giraldo L. P. &amp; Chara J." box="[1015,1386,1015,1042]" pageId="8" pageNumber="9" pagination="1 - 17" refId="ref16402" refString="Gonzalez-Cordoba M., Zuniga M. C., Manzo V., Giraldo L. P. &amp; Chara J. 2016. Notelmis Hinton y Onychelmis Hinton (Coleoptera: Elmidae: Elminae) de Colombia: Claves taxonomicas ilustradas. Boletin del Museo de Entomologia de la Universidad del Valle 16 (2): 1 - 17." type="journal article" year="2016">
González-Córdoba
<emphasis id="B96CEAD3E832FFD8FB43FC07FC36FB83" box="[1249,1312,1015,1041]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
2016
</bibRefCitation>
),
<collectingCountry id="F30F7651E832FFD8FF1FFBE4F820FBA7" box="[189,310,1051,1077]" name="Nicaragua" pageId="8" pageNumber="9">Nicaragua</collectingCountry>
(Shepard pers. com.) and
<collectingCountry id="F30F7651E832FFD8FD21FBE3FBF6FBA4" box="[643,736,1052,1078]" name="Panama" pageId="8" pageNumber="9">Panama</collectingCountry>
(
<bibRefCitation id="EF894B30E832FFD8FD56FBE4FA90FBA7" author="Sharp D." box="[756,902,1051,1077]" pageId="8" pageNumber="9" pagination="1 - 144" refId="ref17769" refString="Sharp D. 1882. Insecta, Coleoptera: Haliplidae, Dytiscidae, Gyrinidae, Hydrophilidae, Heteroceridae, Parnidae, Georissidae, Cyathoceridae. Biologia Centrali-Americana 1 (2): 1 - 144." type="journal article" year="1882">Sharp 1882</bibRefCitation>
). However, the
<typeStatus id="54A38863E832FFD8FBEFFBE4FDA1FBA7" box="[1101,1207,1051,1077]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
, first illustrated herein, differs from the non-type
<taxonomicName id="4C184D42E832FFD8FDE0FBBFFBF4FBCB" baseAuthorityName="Sharp" baseAuthorityYear="1882" box="[578,738,1087,1114]" class="Insecta" family="Elmidae" genus="Elmis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="longicollis">
<emphasis id="B96CEAD3E832FFD8FDE0FBBFFBF4FBCB" box="[578,738,1087,1114]" italics="true" pageId="8" pageNumber="9">O. longicollis</emphasis>
</taxonomicName>
from
<collectingCountry id="F30F7651E832FFD8FC8BFBC0FA8BFBCB" box="[809,925,1087,1113]" name="Colombia" pageId="8" pageNumber="9">Colombia</collectingCountry>
depicted in
<bibRefCitation id="EF894B30E832FFD8FB8FFBC0FC61FBCB" author="Spangler P. &amp; Santiago S." box="[1069,1399,1087,1113]" pageId="8" pageNumber="9" pagination="495 - 498" refId="ref17990" refString="Spangler P. &amp; Santiago S. 1991. A new species and new records from Colombia of the water beetle genus Onychelmis Hinton (Coleoptera: Elmidae: Elminae). Proceedings of the Entomological Society of Washington 93 (2): 495 - 498." type="journal article" year="1991">Spangler &amp; Santiago (1991)</bibRefCitation>
and later followed by
<bibRefCitation id="EF894B30E832FFD8FE6CFB9CFA47FBEC" author="Gonzalez-Cordoba M. &amp; Zuniga M. C. &amp; Manzo V. &amp; Giraldo L. P. &amp; Chara J." box="[462,849,1123,1150]" pageId="8" pageNumber="9" pagination="1 - 17" refId="ref16402" refString="Gonzalez-Cordoba M., Zuniga M. C., Manzo V., Giraldo L. P. &amp; Chara J. 2016. Notelmis Hinton y Onychelmis Hinton (Coleoptera: Elmidae: Elminae) de Colombia: Claves taxonomicas ilustradas. Boletin del Museo de Entomologia de la Universidad del Valle 16 (2): 1 - 17." type="journal article" year="2016">
González-Córdoba
<emphasis id="B96CEAD3E832FFD8FD15FB9BFBE3FBEF" box="[695,757,1123,1149]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
(2016)
</bibRefCitation>
, and it is possible that they in fact represent different species, and therefore the correct distribution of
<taxonomicName id="4C184D42E832FFD8FCC2FB77FD16FB33" baseAuthorityName="Sharp" baseAuthorityYear="1882" box="[864,1024,1159,1186]" class="Insecta" family="Elmidae" genus="Elmis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="longicollis">
<emphasis id="B96CEAD3E832FFD8FCC2FB77FD16FB33" box="[864,1024,1159,1186]" italics="true" pageId="8" pageNumber="9">O. longicollis</emphasis>
</taxonomicName>
needs to be reviewed.
</paragraph>
</subSubSection>
<subSubSection id="C302654AE832FFD8FF1FFB35FA1DFA00" pageId="8" pageNumber="9" type="discussion">
<paragraph id="8BA736C1E832FFD8FF1FFB35F839FB76" blockId="8.[189,303,1226,1252]" box="[189,303,1226,1252]" pageId="8" pageNumber="9">
<heading id="D0EF81ADE832FFD8FF1FFB35F839FB76" bold="true" box="[189,303,1226,1252]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="B96CEAD3E832FFD8FF1FFB35F839FB76" bold="true" box="[189,303,1226,1252]" pageId="8" pageNumber="9">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BA736C1E832FFD8FF1FFAF3FA1DFA00" blockId="8.[188,1398,1292,1426]" pageId="8" pageNumber="9">
As stated by
<bibRefCitation id="EF894B30E832FFD8FEF0FAF3F8F8FAB4" author="Sharp D." box="[338,494,1292,1318]" pageId="8" pageNumber="9" pagination="1 - 144" refId="ref17769" refString="Sharp D. 1882. Insecta, Coleoptera: Haliplidae, Dytiscidae, Gyrinidae, Hydrophilidae, Heteroceridae, Parnidae, Georissidae, Cyathoceridae. Biologia Centrali-Americana 1 (2): 1 - 144." type="journal article" year="1882">Sharp (1882)</bibRefCitation>
, the
<typeStatus id="54A38863E832FFD8FD80FAF3FB43FAB4" box="[546,597,1292,1318]" pageId="8" pageNumber="9">type</typeStatus>
specimen is in poor condition and some characters like tibial fringes, which
<bibRefCitation id="EF894B30E832FFD8FEADFACFF8ADFAD8" author="Hinton H. E." box="[271,443,1328,1354]" pageId="8" pageNumber="9" pagination="65 - 104" refId="ref16569" refString="Hinton H. E. 1941. New genera and species of Elmidae (Coleoptera). Transactions of the Royal Entomological Society of London 91 (3): 65 - 104. https: // doi. org / 10.1111 / j. 1365 - 2311.1941. tb 01044. x" type="journal article" year="1941">Hinton (1941)</bibRefCitation>
incorrectly interpreted as plastron, are difficult to properly observe. Antennal segments 3–11, the left hind leg and the tarsus of the left mid leg are missing, the left elytron is broken, and the left foreleg is glued to the mounting point.
</paragraph>
</subSubSection>
</treatment>
</document>
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