431 lines
57 KiB
XML
431 lines
57 KiB
XML
<document id="BC4CEACD0645B3028EAC2B08B378900D" ID-CLB-Dataset="7762" ID-DOI="10.5852/ejt.2021.739.1263" ID-GBIF-Dataset="79e0ebb9-6412-4821-879a-9ec760579681" ID-ISSN="2118-9773" ID-Zenodo-Dep="4600344" ID-ZooBank="C333D3B5-7B4B-4A53-8548-E74DFEE9EBC2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615533921823" checkinUser="plazi" docAuthor="Linský, Marek, Čiamporová-Zaťovičová, Zuzana & Čiampor, Fedor" docDate="2021" docId="03B187D7E82BFFC3FDC6FEF7FAFEF8BC" docLanguage="en" docName="ejt-739_1-35.pdf" docOrigin="European Journal of Taxonomy 739 (1)" docSource="http://dx.doi.org/10.5852/ejt.2021.739.1263" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Onychelmis splendida Linský & Čiamporová-Zaťovičová & Čiampor 2021, sp. nov." docType="treatment" docUuid="56128631-3C7F-4D7E-941E-DDA1DC318912" docUuidSource="ZooBank" docVersion="8" lastPageNumber="20" masterDocId="FF88FFAFE83AFFD0FFA2FFFFF916FF92" masterDocTitle="A revision of Onychelmis Hinton, 1941 (Coleoptera: Elmidae), with description of new species, DNA barcoding and notes on the geography of the genus" masterLastPageNumber="35" masterPageNumber="1" pageNumber="18" updateTime="1698933319914" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="D6747BCFFD6DEE2AD30D3A99AE644E53">A revision of Onychelmis Hinton, 1941 (Coleoptera: Elmidae), with description of new species, DNA barcoding and notes on the geography of the genus</mods:title>
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<mods:namePart id="A59D4A3B4DD103D1AB924F43DB54D188">Linský, Marek</mods:namePart>
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<mods:affiliation id="4A53440EA25EC584448D12DF21CE5521">Zoology Lab, Institute of Botany, Plant Science and Biodiversity Centre, Slovak Academy of Sciences, Dúbravská cesta 9, SK- 84523, Bratislava, Slovakia. & Department of Zoology, Faculty of Natural Science, Comenius University, Mlynská dolina B- 1, SK- 84215 Bratislava, Slovakia.</mods:affiliation>
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<mods:nameIdentifier id="CC68562641BE1F15AE4710928E7EBE00" type="email">marek.linsky@gmail.com</mods:nameIdentifier>
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<mods:namePart id="F7C752DBC2225CC5B1AAD33E5E4E0A28">Čiamporová-Zaťovičová, Zuzana</mods:namePart>
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<mods:affiliation id="A8DADE54622EFAF0D9DD73D3DECF9359">Zoology Lab, Institute of Botany, Plant Science and Biodiversity Centre, Slovak Academy of Sciences, Dúbravská cesta 9, SK- 84523, Bratislava, Slovakia. & Department of Ecology, Faculty of Natural Science, Comenius University, Mlynská dolina B- 1, SK- 84215 Bratislava, Slovakia.</mods:affiliation>
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<mods:nameIdentifier id="A6E2FFD623C7D2042EA1B878FF1359D4" type="email">zuzana.zatovicova@savba.sk</mods:nameIdentifier>
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<mods:namePart id="AF38FEC53C230A7366D224DC32E82AA2">Čiampor, Fedor</mods:namePart>
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<mods:affiliation id="873DA794CE7EA27BD7B97591AA8EB5C6">Zoology Lab, Institute of Botany, Plant Science and Biodiversity Centre, Slovak Academy of Sciences, Dúbravská cesta 9, SK- 84523, Bratislava, Slovakia.</mods:affiliation>
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<mods:nameIdentifier id="E95D243786AE2C92A1601EFFC85110CF" type="email">f.ciampor@savba.sk</mods:nameIdentifier>
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<mods:title id="E6DB3593099E294CCC14FAB88299134B">European Journal of Taxonomy</mods:title>
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<mods:part id="597945AE7664BDBC1A30D08F6FAD7BFC">
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<mods:date id="FDCF717FEB6DEFA79BFF5996ED508977">2021</mods:date>
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<mods:number id="C5F887D86F15DED670E37A2DCD098D90">2021-03-11</mods:number>
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<mods:url id="7BDA98CA947667FA5D8359E2B92769FC">http://dx.doi.org/10.5852/ejt.2021.739.1263</mods:url>
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<treatment id="03B187D7E82BFFC3FDC6FEF7FAFEF8BC" ID-DOI="http://doi.org/10.5281/zenodo.4602503" ID-GBIF-Taxon="180331475" ID-Zenodo-Dep="4602503" LSID="urn:lsid:zoobank.org:act:56128631-3C7F-4D7E-941E-DDA1DC318912" httpUri="http://treatment.plazi.org/id/03B187D7E82BFFC3FDC6FEF7FAFEF8BC" lastPageId="19" lastPageNumber="20" pageId="17" pageNumber="18">
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<subSubSection id="C302654AE82BFFC1FDC6FEF7FAC6FEB1" box="[612,976,264,291]" pageId="17" pageNumber="18" type="nomenclature">
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<paragraph id="8BA736C1E82BFFC1FDC6FEF7FAC6FEB1" blockId="17.[367,1220,264,362]" box="[612,976,264,291]" pageId="17" pageNumber="18">
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<heading id="D0EF81ADE82BFFC1FDC6FEF7FAC6FEB1" box="[612,976,264,291]" centered="true" fontSize="11" level="2" pageId="17" pageNumber="18" reason="2">
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<taxonomicName id="4C184D42E82BFFC1FDC6FEF7FA78FEB1" authority="Linský & Čiamporová-Zaťovičová & Čiampor, 2021" authorityName="Linský & Čiamporová-Zaťovičová & Čiampor" authorityYear="2021" box="[612,878,264,291]" class="Insecta" family="Elmidae" genus="Onychelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="splendida" status="sp. nov.">
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<emphasis id="B96CEAD3E82BFFC1FDC6FEF7FA78FEB1" bold="true" box="[612,878,264,291]" italics="true" pageId="17" pageNumber="18">Onychelmis splendida</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25F57A8E82BFFC1FCD4FEF6FAC6FEB1" box="[886,976,265,291]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C302654AE82BFFC1FECDFED3FD04FEFB" pageId="17" pageNumber="18" type="description">
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<paragraph id="8BA736C1E82BFFC1FECDFED3FDD2FED5" blockId="17.[367,1220,264,362]" box="[367,1220,300,327]" pageId="17" pageNumber="18">
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<uri id="FF893AC3E82BFFC1FECDFED3FDD2FED5" box="[367,1220,300,327]" pageId="17" pageNumber="18">
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urn:lsid:zoobank.org:act:
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<uuid id="FFBE0C14E82BFFC1FD2FFED3FDD2FED5" box="[653,1220,300,327]" pageId="17" pageNumber="18">56128631-3C7F-4D7E-941E-DDA1DC318912</uuid>
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</uri>
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</paragraph>
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<paragraph id="8BA736C1E82BFFC1FD83FEB0FD04FEFB" blockId="17.[367,1220,264,362]" box="[545,1042,335,362]" pageId="17" pageNumber="18">
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<figureCitation id="13232A44E82BFFC1FD83FEB0FB97FEFB" box="[545,641,335,361]" captionStart="Fig" captionStartId="11.[189,232,1892,1918]" captionTargetBox="[246,1344,271,1850]" captionTargetId="figure-12@11.[240,1348,265,1854]" captionTargetPageId="11" captionText="Fig. 3. Habitus of Onychelmis spp. A. O. leleupi Delève, 1968, ♂ (PUCE). B. O. lobata sp. nov., holotype, ♂ (PUCE). C. O. onorei sp. nov., holotype, ♂ (PUCE). D. O. splendida sp. nov., holotype, ♂ (PUCE). E. O. lenkae sp. nov., paratype, ♂ (PUCE). F. O. minor sp. nov., holotype, ♂ (PUCE). Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4600354" httpUri="https://zenodo.org/record/4600354/files/figure.png" pageId="17" pageNumber="18">Figs 3D</figureCitation>
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,
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<figureCitation id="13232A44E82BFFC1FD2CFEB0FBA4FEF8" box="[654,690,335,362]" captionStart="Fig" captionStartId="17.[189,232,1928,1954]" captionTargetBox="[189,1398,1019,1889]" captionTargetId="figure-149@17.[189,1398,1019,1889]" captionTargetPageId="17" captionText="Fig. 5. Venter and metaventrites of male Onychelmis spp. A, C. O. lenkae sp. nov., paratype, ♂ (PUCE). B. O. splendida sp. nov., paratype, ♂ (PUCE). Arrows show setal tufts. Scale bars: A = 0.5 mm; B–C = 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.4600360" httpUri="https://zenodo.org/record/4600360/files/figure.png" pageId="17" pageNumber="18">5B</figureCitation>
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,
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<figureCitation id="13232A44E82BFFC1FD1DFEB0FBF3FEFB" box="[703,741,335,361]" captionStart="Fig" captionStartId="18.[189,232,1893,1919]" captionTargetBox="[199,1389,1223,1846]" captionTargetId="figure-257@18.[189,1398,1213,1855]" captionTargetPageId="18" captionText="Fig. 6. Tomentum on pro- and mesofemora of Onychelmis spp. A. O. leleupi Delève, 1968. B. O. lenkae sp. nov. C. O. lobata sp. nov. D. O. longicollis (Sharp, 1882). E. O. minor sp. nov. F. O. onorei sp. nov. G. O. splendida sp. nov. H. O. whiteheadi Spangler & Santiago, 1991. Drawings are schematic, bodies are divided to identical halves: left = male, right = female (note: females unknown for D and H)." figureDoi="http://doi.org/10.5281/zenodo.4600362" httpUri="https://zenodo.org/record/4600362/files/figure.png" pageId="17" pageNumber="18">6G</figureCitation>
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,
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<figureCitation id="13232A44E82BFFC1FD53FEB0FBE9FEFB" box="[753,767,335,361]" captionStart="Fig" captionStartId="22.[189,232,1759,1785]" captionTargetBox="[208,1384,280,1715]" captionTargetId="figure-16@22.[189,1398,265,1721]" captionTargetPageId="22" captionText="Fig. 9. Aedeagi of Onychelmis spp. A. O. onorei sp. nov., ventral view. B. O. onorei sp. nov., lateral view. C. O. splendida sp. nov., ventral view. D. O. splendida sp. nov., lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4600370" httpUri="https://zenodo.org/record/4600370/files/figure.png" pageId="17" pageNumber="18">9</figureCitation>
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C–D, 11B, 11F, 12A–B
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</paragraph>
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</subSubSection>
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<subSubSection id="C302654AE82BFFC1FF1FFE70F845FDC9" pageId="17" pageNumber="18" type="diagnosis">
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<paragraph id="8BA736C1E82BFFC1FF1FFE70F8D0FE3B" blockId="17.[189,454,399,425]" box="[189,454,399,425]" pageId="17" pageNumber="18">
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<heading id="D0EF81ADE82BFFC1FF1FFE70F8D0FE3B" bold="true" box="[189,454,399,425]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
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<emphasis id="B96CEAD3E82BFFC1FF1FFE70F8D0FE3B" bold="true" box="[189,454,399,425]" pageId="17" pageNumber="18">Differential diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BA736C1E82BFFC1FF1FFE43F845FDC9" blockId="17.[188,1399,444,603]" pageId="17" pageNumber="18">
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<taxonomicName id="4C184D42E82BFFC1FF1FFE43F8D7FE44" authority="Linský & Čiamporová-Zaťovičová & Čiampor, 2021" authorityName="Linský & Čiamporová-Zaťovičová & Čiampor" authorityYear="2021" box="[189,449,444,470]" class="Insecta" family="Elmidae" genus="Onychelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="splendida" status="sp. nov.">
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<emphasis id="B96CEAD3E82BFFC1FF1FFE43F8D7FE44" box="[189,449,444,470]" italics="true" pageId="17" pageNumber="18">Onychelmis splendida</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25F57A8E82BFFC1FE65FE43FB34FE44" box="[455,546,444,470]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
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can be distinguished from all species of the genus by the combination of the following characters: 1) moderate size (CL:
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<quantity id="4CE09B24E82BFFC1FD45FE22FA83FE65" box="[743,917,477,503]" metricMagnitude="-3" metricUnit="m" metricValue="1.385" metricValueMax="1.4" metricValueMin="1.37" pageId="17" pageNumber="18" unit="mm" value="1.385" valueMax="1.4" valueMin="1.37">1.37–1.40 mm</quantity>
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); 2) pro- and mesofemora with tomentum reaching to around middle in males, and behind middle in females (
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<figureCitation id="13232A44E82BFFC1FBA3FE01FD72FD8B" box="[1025,1124,510,537]" captionStart="Fig" captionStartId="18.[189,232,1893,1919]" captionTargetBox="[199,1389,1223,1846]" captionTargetId="figure-257@18.[189,1398,1213,1855]" captionTargetPageId="18" captionText="Fig. 6. Tomentum on pro- and mesofemora of Onychelmis spp. A. O. leleupi Delève, 1968. B. O. lenkae sp. nov. C. O. lobata sp. nov. D. O. longicollis (Sharp, 1882). E. O. minor sp. nov. F. O. onorei sp. nov. G. O. splendida sp. nov. H. O. whiteheadi Spangler & Santiago, 1991. Drawings are schematic, bodies are divided to identical halves: left = male, right = female (note: females unknown for D and H)." figureDoi="http://doi.org/10.5281/zenodo.4600362" httpUri="https://zenodo.org/record/4600362/files/figure.png" pageId="17" pageNumber="18">Fig. 6G</figureCitation>
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); 3) humeri produced; 4) prominent carina on sixth interval absent; 5) elytral punctures absent; 6) shape of aedeagus lanceolate (
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<figureCitation id="13232A44E82BFFC1FF67FDBEF81CFDC9" box="[197,266,577,603]" captionStart="Fig" captionStartId="22.[189,232,1759,1785]" captionTargetBox="[208,1384,280,1715]" captionTargetId="figure-16@22.[189,1398,265,1721]" captionTargetPageId="22" captionText="Fig. 9. Aedeagi of Onychelmis spp. A. O. onorei sp. nov., ventral view. B. O. onorei sp. nov., lateral view. C. O. splendida sp. nov., ventral view. D. O. splendida sp. nov., lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4600370" httpUri="https://zenodo.org/record/4600370/files/figure.png" pageId="17" pageNumber="18">Fig. 9</figureCitation>
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C–D).
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</paragraph>
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</subSubSection>
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<subSubSection id="C302654AE82BFFC1FF1FFD7EF81EFD7A" pageId="17" pageNumber="18" type="etymology">
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<paragraph id="8BA736C1E82BFFC1FF1FFD7EF852FD09" blockId="17.[189,1399,641,744]" box="[189,324,641,667]" pageId="17" pageNumber="18">
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<heading id="D0EF81ADE82BFFC1FF1FFD7EF852FD09" bold="true" box="[189,324,641,667]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
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<emphasis id="B96CEAD3E82BFFC1FF1FFD7EF852FD09" bold="true" box="[189,324,641,667]" pageId="17" pageNumber="18">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BA736C1E82BFFC1FF1FFD52F81EFD7A" blockId="17.[189,1399,641,744]" pageId="17" pageNumber="18">
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The specific epithet
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<taxonomicName id="4C184D42E82BFFC1FE0DFD52FB37FD55" authorityName="Linský & Čiamporová-Zaťovičová & Čiampor" authorityYear="2021" box="[431,545,685,711]" class="Insecta" family="Elmidae" genus="Onychelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="splendida">
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<emphasis id="B96CEAD3E82BFFC1FE0DFD52FB37FD55" box="[431,545,685,711]" italics="true" pageId="17" pageNumber="18">splendida</emphasis>
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</taxonomicName>
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is from the Latin ʻ
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<emphasis id="B96CEAD3E82BFFC1FCA4FD52FA95FD55" box="[774,899,685,711]" italics="true" pageId="17" pageNumber="18">splendidus</emphasis>
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ʼ (ʻshinyʼ) referring to the glabrous, shiny elytra.
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</paragraph>
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</subSubSection>
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<subSubSection id="C302654AE82BFFC2FF1FFCF1FA8EFDC9" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="materials_examined">
|
||
<paragraph id="8BA736C1E82BFFC1FF1FFCF1F8BCFCBA" blockId="17.[189,426,782,808]" box="[189,426,782,808]" pageId="17" pageNumber="18">
|
||
<heading id="D0EF81ADE82BFFC1FF1FFCF1F8BCFCBA" bold="true" box="[189,426,782,808]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
|
||
<emphasis id="B96CEAD3E82BFFC1FF1FFCF1F8BCFCBA" bold="true" box="[189,426,782,808]" pageId="17" pageNumber="18">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA736C1E82BFFC1FF48FCC4FB6BFC29" blockId="17.[189,1399,827,957]" pageId="17" pageNumber="18">
|
||
<materialsCitation id="3B703C9CE82BFFC1FF48FCC4FB6BFC29" ID-GBIF-Occurrence="3051860313" collectedFrom="confluence of two larger streams, with gravel, stones, boulders, Ciampor & Ciamporova-Zatovicova lgt." collectingDate="2013-08-17" collectionCode="PUCE" country="Ecuador" county="Ecuador" elevation="1109" latitude="-0.7247222" location="Sumaco" longLatPrecision="1" longitude="-77.76706" municipality="Coca" pageId="17" pageNumber="18" specimenCount="1" stateProvince="Napo prov." typeStatus="holotype">
|
||
<emphasis id="B96CEAD3E82BFFC1FF48FCC4F84AFCC7" bold="true" box="[234,348,827,853]" pageId="17" pageNumber="18">
|
||
<typeStatus id="54A38863E82BFFC1FF48FCC4F84AFCC7" box="[234,348,827,853]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="F30F7651E82BFFC1FF1FFCA3F85DFCE4" box="[189,331,860,886]" name="Ecuador" pageId="17" pageNumber="18">ECUADOR</collectingCountry>
|
||
•
|
||
<specimenCount id="9D1EFD48E82BFFC1FECEFCA2F894FCE5" box="[364,386,861,887]" pageId="17" pageNumber="18" type="male">♂</specimenCount>
|
||
“Ecuador,
|
||
<collectingRegion id="49DCF823E82BFFC1FDACFCA2FB81FCE5" box="[526,663,861,887]" country="Ecuador" name="Napo" pageId="17" pageNumber="18">Napo prov.</collectingRegion>
|
||
,
|
||
<locationDeviation id="9A152FE3E82BFFC1FD05FCA3FA51FCE4" box="[679,839,860,887]" pageId="17" pageNumber="18">road to Coca</locationDeviation>
|
||
,
|
||
<location id="8EC7601AE82BFFC1FCF5FCA3FAA0FCE4" LSID="urn:lsid:plazi:treatment:03B187D7E82BFFC3FDC6FEF7FAFEF8BC:8EC7601AE82BFFC1FCF5FCA3FAA0FCE4" box="[855,950,860,886]" country="Ecuador" county="Ecuador" latitude="-0.7247222" longLatPrecision="1" longitude="-77.76706" municipality="Coca" name="Sumaco" pageId="17" pageNumber="18" stateProvince="Napo prov.">Sumaco</location>
|
||
env.,
|
||
<geoCoordinate id="EE2C5006E82BFFC1FBA6FCA5FDB8FCE4" box="[1028,1198,858,886]" degrees="00" direction="south" minutes="43" orientation="latitude" pageId="17" pageNumber="18" precision="1" seconds="29.0" value="-0.7247222">00°43′29.0″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE2C5006E82BFFC1FB1DFCA5FC64FCE5" box="[1215,1394,858,887]" degrees="77" direction="west" minutes="46" orientation="longitude" pageId="17" pageNumber="18" precision="1" seconds="01.4" value="-77.76706">77°46′01.4″ W</geoCoordinate>
|
||
,
|
||
<elevation id="0035D1F2E82BFFC1FF1FFC81F840FC05" box="[189,342,893,920]" metricMagnitude="3" metricUnit="m" metricValue="1.109" pageId="17" pageNumber="18" unit="m" value="1109.0">
|
||
<quantity id="4CE09B24E82BFFC1FF1FFC81F800FC0A" box="[189,278,894,920]" metricMagnitude="3" metricUnit="m" metricValue="1.109" pageId="17" pageNumber="18" unit="m" value="1109.0">1109 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
,
|
||
<date id="FFA61001E82BFFC1FEC7FC81F8FBFC0A" box="[357,493,894,920]" pageId="17" pageNumber="18" value="2013-08-17">
|
||
<collectingDate id="EFE2E9E9E82BFFC1FEC7FC81F8FBFC0A" box="[357,493,894,920]" pageId="17" pageNumber="18" value="2013-08-17">17. 8. 2013</collectingDate>
|
||
</date>
|
||
,
|
||
<collectedFrom id="3804753CE82BFFC1FE59FC82FB07FC29" pageId="17" pageNumber="18">confluence of two larger streams, with gravel, stones, boulders, Čiampor & Čiamporová-Zaťovičová lgt.</collectedFrom>
|
||
”;
|
||
<collectionCode id="ED09AE04E82BFFC1FD88FC5EFB6FFC29" box="[554,633,929,955]" pageId="17" pageNumber="18">PUCE</collectionCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<caption id="DF676649E82BFFC1FF1FF877F808F87B" ID-DOI="http://doi.org/10.5281/zenodo.4600360" ID-Zenodo-Dep="4600360" httpUri="https://zenodo.org/record/4600360/files/figure.png" pageId="17" pageNumber="18" startId="17.[189,232,1928,1954]" targetBox="[189,1398,1019,1889]" targetPageId="17">
|
||
<paragraph id="8BA736C1E82BFFC1FF1FF877F808F87B" blockId="17.[189,1399,1928,2025]" pageId="17" pageNumber="18">
|
||
<emphasis id="B96CEAD3E82BFFC1FF1FF877F81CF830" bold="true" box="[189,266,1928,1954]" pageId="17" pageNumber="18">Fig. 5.</emphasis>
|
||
Venter and metaventrites of male
|
||
<taxonomicName id="4C184D42E82BFFC1FD34F877FA36F830" authorityName="Hinton" authorityYear="1941" box="[662,800,1928,1954]" class="Insecta" family="Elmidae" genus="Onychelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96CEAD3E82BFFC1FD34F877FA36F830" box="[662,800,1928,1954]" italics="true" pageId="17" pageNumber="18">Onychelmis</emphasis>
|
||
</taxonomicName>
|
||
spp.
|
||
<emphasis id="B96CEAD3E82BFFC1FCFCF877FA65F830" bold="true" box="[862,883,1928,1954]" pageId="17" pageNumber="18">A</emphasis>
|
||
,
|
||
<emphasis id="B96CEAD3E82BFFC1FC23F877FA80F830" bold="true" box="[897,918,1928,1954]" pageId="17" pageNumber="18">C</emphasis>
|
||
.
|
||
<taxonomicName id="4C184D42E82BFFC1FC06F877FD02F830" authority="Linský & Čiamporová-Zaťovičová & Čiampor, 2021" authorityName="Linský & Čiamporová-Zaťovičová & Čiampor" authorityYear="2021" box="[932,1044,1928,1954]" class="Insecta" family="Elmidae" genus="Onychelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="lenkae" status="sp. nov.">
|
||
<emphasis id="B96CEAD3E82BFFC1FC06F877FD02F830" box="[932,1044,1928,1954]" italics="true" pageId="17" pageNumber="18">O. lenkae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25F57A8E82BFFC1FBBBF876FD60F831" box="[1049,1142,1929,1955]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, paratype, ♂ (PUCE).
|
||
<emphasis id="B96CEAD3E82BFFC1FF1FF854F9C7F857" bold="true" box="[189,209,1963,1989]" pageId="17" pageNumber="18">B</emphasis>
|
||
.
|
||
<taxonomicName id="4C184D42E82BFFC1FF7DF853F863F857" authority="Linský & Čiamporová-Zaťovičová & Čiampor, 2021" authorityName="Linský & Čiamporová-Zaťovičová & Čiampor" authorityYear="2021" box="[223,373,1963,1990]" class="Insecta" family="Elmidae" genus="Onychelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="splendida" status="sp. nov.">
|
||
<emphasis id="B96CEAD3E82BFFC1FF7DF853F863F857" box="[223,373,1963,1990]" italics="true" pageId="17" pageNumber="18">O. splendida</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25F57A8E82BFFC1FEDFF853F8CCF854" box="[381,474,1964,1990]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, paratype, ♂ (PUCE). Arrows show setal tufts. Scale bars: A = 0.5 mm; B–C = 0.3 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA736C1E828FFC2FF48FEF6F84CFE64" blockId="18.[189,1399,265,502]" pageId="18" pageNumber="19">
|
||
<materialsCitation id="3B703C9CE828FFC2FF48FEF6FD76FED4" ID-GBIF-Occurrence="3051860323" collectedFrom="confluence of two larger streams, with gravel, stones, boulders, Ciampor & Ciamporova-Zatovicova lgt." collectingDate="2013-08-17" collectionCode="PUCE, CCB" country="Ecuador" county="Ecuador" elevation="1109" latitude="-0.7247222" location="ECUADOR" longLatPrecision="1" longitude="-77.76706" municipality="Coca" pageId="18" pageNumber="19" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Napo" typeStatus="paratype">
|
||
<emphasis id="B96CEAD3E828FFC2FF48FEF6F871FEB1" bold="true" box="[234,359,265,291]" pageId="18" pageNumber="19">
|
||
<typeStatus id="54A38863E828FFC2FF48FEF6F871FEB1" box="[234,359,265,291]" pageId="18" pageNumber="19" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="F30F7651E828FFC2FF1FFED3F85DFED4" box="[189,331,300,326]" name="Ecuador" pageId="18" pageNumber="19">ECUADOR</collectingCountry>
|
||
•
|
||
<specimenCount id="9D1EFD48E828FFC2FEC4FED3F8BDFED4" box="[358,427,300,326]" pageId="18" pageNumber="19" type="male">3 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D1EFD48E828FFC2FE1BFED3F8E8FED4" box="[441,510,300,326]" pageId="18" pageNumber="19" type="female">3 ♀♀</specimenCount>
|
||
; same collection data as for holotype;
|
||
<collectionCode id="ED09AE04E828FFC2FC69FED3FD00FED4" box="[971,1046,300,326]" pageId="18" pageNumber="19">PUCE</collectionCode>
|
||
/
|
||
<collectionCode id="ED09AE04E828FFC2FB87FED3FD76FED4" box="[1061,1120,300,326]" country="India" httpUri="http://grbio.org/cool/e3tf-9780" name="Central College, Bangalore" pageId="18" pageNumber="19">CCB</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B703C9CE828FFC2FBD9FED3FB26FE22" ID-GBIF-Occurrence="3051860316" collectedFrom="fast flowing, with boulders, stones, gravel, submerged wood, Ciampor & Ciamporova- Zatovicova lgt." collectingDate="2013-08-17" collectionCode="PUCE, CCB" country="Ecuador" county="Ecuador" elevation="1138" latitude="-0.7071389" location="Sumaco" longLatPrecision="1" longitude="-77.719444" municipality="Coca" pageId="18" pageNumber="19" specimenCount="1" specimenCount-male="1" stateProvince="Napo prov." typeStatus="paratype">
|
||
<specimenCount id="9D1EFD48E828FFC2FBD9FED3FDBDFED4" box="[1147,1195,300,326]" pageId="18" pageNumber="19" type="male">1 ♂</specimenCount>
|
||
; “Ecuador,
|
||
<collectingRegion id="49DCF823E828FFC2FA95FED3F9EDFEF8" country="Ecuador" name="Napo" pageId="18" pageNumber="19">Napo prov.</collectingRegion>
|
||
,
|
||
<locationDeviation id="9A152FE3E828FFC2FEAAFEB0F8B7FEFB" box="[264,417,335,362]" pageId="18" pageNumber="19">road to Coca</locationDeviation>
|
||
,
|
||
<location id="8EC7601AE828FFC2FE0CFEB0FB1BFEFB" LSID="urn:lsid:plazi:treatment:03B187D7E82BFFC3FDC6FEF7FAFEF8BC:8EC7601AE828FFC2FE0CFEB0FB1BFEFB" box="[430,525,335,361]" country="Ecuador" county="Ecuador" latitude="-0.7071389" longLatPrecision="1" longitude="-77.719444" municipality="Coca" name="Sumaco" pageId="18" pageNumber="19" stateProvince="Napo prov.">Sumaco</location>
|
||
env.,
|
||
<geoCoordinate id="EE2C5006E828FFC2FDF6FEB2FBECFEFB" box="[596,762,333,361]" degrees="00" direction="south" minutes="42" orientation="latitude" pageId="18" pageNumber="19" precision="1" seconds="25.7" value="-0.7071389">00°42′25.7″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE2C5006E828FFC2FCAAFEB2FAAEFEF8" box="[776,952,333,362]" degrees="77" direction="west" minutes="43" orientation="longitude" pageId="18" pageNumber="19" precision="1" seconds="10.0" value="-77.719444">77°43′10.0″ W</geoCoordinate>
|
||
,
|
||
<elevation id="0035D1F2E828FFC2FC66FEB0FD4AFEFB" box="[964,1116,335,362]" metricMagnitude="3" metricUnit="m" metricValue="1.138" pageId="18" pageNumber="19" unit="m" value="1138.0">
|
||
<quantity id="4CE09B24E828FFC2FC66FEB0FD0AFEF8" box="[964,1052,335,362]" metricMagnitude="3" metricUnit="m" metricValue="1.138" pageId="18" pageNumber="19" unit="m" value="1138.0">1138 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
,
|
||
<date id="FFA61001E828FFC2FBC8FEB0FDE4FEFB" box="[1130,1266,335,361]" pageId="18" pageNumber="19" value="2013-08-17">
|
||
<collectingDate id="EFE2E9E9E828FFC2FBC8FEB0FDE4FEFB" box="[1130,1266,335,361]" pageId="18" pageNumber="19" value="2013-08-17">17. 8. 2013</collectingDate>
|
||
</date>
|
||
, stream ca.
|
||
<quantity id="4CE09B24E828FFC2FF1FFE8DF81CFE1F" box="[189,266,370,397]" metricMagnitude="0" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="18" pageNumber="19" unit="m" value="2.5" valueMax="3.0" valueMin="2.0">2–3 m</quantity>
|
||
wide,
|
||
<collectedFrom id="3804753CE828FFC2FEC2FE8DF86AFE22" pageId="18" pageNumber="19">fast flowing, with boulders, stones, gravel, submerged wood, Čiampor & Čiamporová- Zaťovičová lgt.</collectedFrom>
|
||
”;
|
||
<collectionCode id="ED09AE04E828FFC2FE38FE69F8F3FE22" box="[410,485,406,432]" pageId="18" pageNumber="19">PUCE</collectionCode>
|
||
/
|
||
<collectionCode id="ED09AE04E828FFC2FE57FE69FB26FE22" box="[501,560,406,432]" country="India" httpUri="http://grbio.org/cool/e3tf-9780" name="Central College, Bangalore" pageId="18" pageNumber="19">CCB</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B703C9CE828FFC2FDF1FE69F843FE64" ID-GBIF-Occurrence="3051860303" collectedFrom="small waterfall, Ciampor & Ciamporova-Zatovicova lgt." collectingDate="2013-08-17" collectionCode="PUCE, CCB" country="Ecuador" elevation="1129" latitude="-0.7276666" location="Napo prov." longLatPrecision="1" longitude="-77.76559" municipality="Ecuador" pageId="18" pageNumber="19" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Napo prov." typeStatus="paratype">
|
||
<specimenCount id="9D1EFD48E828FFC2FDF1FE69FB93FE22" box="[595,645,406,432]" pageId="18" pageNumber="19" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D1EFD48E828FFC2FD3AFE69FBF6FE22" box="[664,736,406,432]" pageId="18" pageNumber="19" type="female">3 ♀♀</specimenCount>
|
||
; “Ecuador,
|
||
<collectingRegion id="49DCF823E828FFC2FCD4FE69FD16FE22" box="[886,1024,406,432]" country="Ecuador" name="Napo" pageId="18" pageNumber="19">Napo prov.</collectingRegion>
|
||
,
|
||
<locationDeviation id="9A152FE3E828FFC2FBB3FE69FDA2FE22" box="[1041,1204,406,432]" pageId="18" pageNumber="19">road to Coca</locationDeviation>
|
||
,
|
||
<geoCoordinate id="EE2C5006E828FFC2FB64FE6BFC67FE22" box="[1222,1393,404,432]" degrees="00" direction="south" minutes="43" orientation="latitude" pageId="18" pageNumber="19" precision="1" seconds="39.6" value="-0.7276666">00°43′39.6″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE2C5006E828FFC2FF1FFE48F878FE46" box="[189,366,439,468]" degrees="77" direction="west" minutes="45" orientation="longitude" pageId="18" pageNumber="19" precision="1" seconds="56.1" value="-77.76559">77°45′56.1″ W</geoCoordinate>
|
||
,
|
||
<elevation id="0035D1F2E828FFC2FED9FE46FB00FE41" box="[379,534,441,468]" metricMagnitude="3" metricUnit="m" metricValue="1.129" pageId="18" pageNumber="19" unit="m" value="1129.0">
|
||
<quantity id="4CE09B24E828FFC2FED9FE46F8C3FE46" box="[379,469,441,468]" metricMagnitude="3" metricUnit="m" metricValue="1.129" pageId="18" pageNumber="19" unit="m" value="1129.0">1129 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
,
|
||
<date id="FFA61001E828FFC2FD87FE46FBB9FE41" box="[549,687,441,467]" pageId="18" pageNumber="19" value="2013-08-17">
|
||
<collectingDate id="EFE2E9E9E828FFC2FD87FE46FBB9FE41" box="[549,687,441,467]" pageId="18" pageNumber="19" value="2013-08-17">17. 8. 2013</collectingDate>
|
||
</date>
|
||
,
|
||
<collectedFrom id="3804753CE828FFC2FD1FFE46FC73FE41" box="[701,1381,441,470]" pageId="18" pageNumber="19">small waterfall, Čiampor & Čiamporová-Zaťovičová lgt.</collectedFrom>
|
||
”;
|
||
<collectionCode id="ED09AE04E828FFC2FF1FFE23F81EFE64" box="[189,264,476,502]" pageId="18" pageNumber="19">PUCE</collectionCode>
|
||
/
|
||
<collectionCode id="ED09AE04E828FFC2FEB5FE23F843FE64" box="[279,341,476,502]" country="India" httpUri="http://grbio.org/cool/e3tf-9780" name="Central College, Bangalore" pageId="18" pageNumber="19">CCB</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA736C1E828FFC2FF48FDE1F8B0FDAA" blockId="18.[189,920,542,604]" box="[234,422,542,568]" pageId="18" pageNumber="19">
|
||
<emphasis id="B96CEAD3E828FFC2FF48FDE1F8B0FDAA" bold="true" box="[234,422,542,568]" pageId="18" pageNumber="19">Other material</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BA736C1E828FFC2FF1FFDBEFA8EFDC9" blockId="18.[189,920,542,604]" box="[189,920,577,604]" pageId="18" pageNumber="19">
|
||
<materialsCitation id="3B703C9CE828FFC2FF1FFDBEFA85FDC9" ID-GBIF-Occurrence="3051860324" box="[189,915,577,604]" collectedFrom="confluence of two larger streams, with gravel, stones, boulders, Ciampor & Ciamporova-Zatovicova lgt." collectingDate="2013-08-17" collectionCode="CCB" country="Ecuador" county="Ecuador" elevation="1109" latitude="-0.7247222" location="ECUADOR" longLatPrecision="1" longitude="-77.76706" municipality="Coca" pageId="18" pageNumber="19" specimenCount="1" specimenCount-male="1" stateProvince="Napo">
|
||
<collectingCountry id="F30F7651E828FFC2FF1FFDBEF85DFDC9" box="[189,331,577,603]" name="Ecuador" pageId="18" pageNumber="19">ECUADOR</collectingCountry>
|
||
•
|
||
<specimenCount id="9D1EFD48E828FFC2FEC6FDBEF884FDCE" box="[356,402,577,604]" pageId="18" pageNumber="19" type="male">1 ♂</specimenCount>
|
||
; same collection data as for holotype;
|
||
<collectionCode id="ED09AE04E828FFC2FCF7FDBEFA85FDC9" box="[853,915,577,603]" country="India" httpUri="http://grbio.org/cool/e3tf-9780" name="Central College, Bangalore" pageId="18" pageNumber="19">CCB</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C302654AE828FFC3FF1FFD7DFB9BF9B9" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="description">
|
||
<paragraph id="8BA736C1E828FFC2FF1FFD7DF858FD0E" blockId="18.[189,334,642,668]" box="[189,334,642,668]" pageId="18" pageNumber="19">
|
||
<heading id="D0EF81ADE828FFC2FF1FFD7DF858FD0E" bold="true" box="[189,334,642,668]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
|
||
<emphasis id="B96CEAD3E828FFC2FF1FFD7DF858FD0E" bold="true" box="[189,334,642,668]" pageId="18" pageNumber="19">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA736C1E828FFC2FF48FD4EF83CFD59" blockId="18.[189,1399,689,857]" box="[234,298,689,715]" pageId="18" pageNumber="19">
|
||
<emphasis id="B96CEAD3E828FFC2FF48FD4EF83CFD59" bold="true" box="[234,298,689,715]" pageId="18" pageNumber="19">Male</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BA736C1E828FFC2FF1FFD2AFA84FCCA" blockId="18.[189,1399,689,857]" pageId="18" pageNumber="19">
|
||
BODY (
|
||
<figureCitation id="13232A44E828FFC2FEB5FD2BF86FFD7D" box="[279,377,724,751]" captionStart="Fig" captionStartId="11.[189,232,1892,1918]" captionTargetBox="[246,1344,271,1850]" captionTargetId="figure-12@11.[240,1348,265,1854]" captionTargetPageId="11" captionText="Fig. 3. Habitus of Onychelmis spp. A. O. leleupi Delève, 1968, ♂ (PUCE). B. O. lobata sp. nov., holotype, ♂ (PUCE). C. O. onorei sp. nov., holotype, ♂ (PUCE). D. O. splendida sp. nov., holotype, ♂ (PUCE). E. O. lenkae sp. nov., paratype, ♂ (PUCE). F. O. minor sp. nov., holotype, ♂ (PUCE). Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4600354" httpUri="https://zenodo.org/record/4600354/files/figure.png" pageId="18" pageNumber="19">Fig. 3D</figureCitation>
|
||
). Obovate, black dorsally, brown ventrally; length
|
||
<quantity id="4CE09B24E828FFC2FC4DFD2AFDB7FD7D" box="[1007,1185,725,751]" metricMagnitude="-3" metricUnit="m" metricValue="1.385" metricValueMax="1.4" metricValueMin="1.37" pageId="18" pageNumber="19" unit="mm" value="1.385" valueMax="1.4" valueMin="1.37">1.37–1.40 mm</quantity>
|
||
; width
|
||
<quantity id="4CE09B24E828FFC2FAA3FD2BF9F8FC80" metricMagnitude="-4" metricUnit="m" metricValue="6.6" metricValueMax="6.8" metricValueMin="6.4" pageId="18" pageNumber="19" unit="mm" value="0.66" valueMax="0.68" valueMin="0.64">0.64–0.68 mm</quantity>
|
||
; dorsum convex, glabrous with sparse, light yellowish-brown setae. Hairy or scale-like tomentum distributed on following areas: genae, sides of prosternum, mesoventrite, metaventrite and abdomen, epipleura, and medial and lateral portions on bases of femora.
|
||
</paragraph>
|
||
<paragraph id="8BA736C1E828FFC2FF1FFC7FF8CCFC72" blockId="18.[188,1398,896,992]" pageId="18" pageNumber="19">COLOUR. Head, pronotum and elytra black; venter dark brown to black with reddish tinge; coxae, femora and tarsi dark brown with reddish tinge; trochanters and tibiae brown; basal segments of antennae and tarsal claws pale brown.</paragraph>
|
||
<paragraph id="8BA736C1E828FFC3FF1FFBF7F80FFE22" blockId="18.[189,1399,1032,1165]" lastBlockId="19.[189,1399,265,432]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
|
||
HEAD. Partly retractable into prothoraX, dorsally shiny. Antennae filiform, 11-segmented; pedicel about twice as long as scape; remaining segments about 4 times as long as scape and pedicel combined; segments 3–10 subrectangular, subequal in length; terminal segment longest, suboval, with pointed apex. Labrum with anterior margin very slightly emarginate medially; anterolateral angles broadly arcuate with numerous golden, recumbent hair-like setae; clypeus shorter and wider than labrum, about 3.5 times as wide as long, anterior margin slightly concave, anterolateral angles rounded; frontoclypeal suture almost straight. Eyes well developed, HW:
|
||
<quantity id="4CE09B24E829FFC3FCA9FEB0FAAEFEF8" box="[779,952,335,362]" metricMagnitude="-4" metricUnit="m" metricValue="3.05" metricValueMax="3.1" metricValueMin="3.0" pageId="19" pageNumber="20" unit="mm" value="0.305" valueMax="0.31" valueMin="0.3">0.30–0.31 mm</quantity>
|
||
, ID:
|
||
<quantity id="4CE09B24E829FFC3FC51FEB0FDB7FEF8" box="[1011,1185,335,362]" metricMagnitude="-4" metricUnit="m" metricValue="1.75" metricValueMax="1.7999999999999998" metricValueMin="1.7" pageId="19" pageNumber="20" unit="mm" value="0.175" valueMax="0.18" valueMin="0.17">0.17–0.18 mm</quantity>
|
||
, suboval in lateral view, protruding from head outline in dorsal view, circumocular surface raised. Frons between eyes convex.
|
||
</paragraph>
|
||
<caption id="DF676649E828FFC2FF1FF89AFC3FF878" ID-DOI="http://doi.org/10.5281/zenodo.4600362" ID-Zenodo-Dep="4600362" httpUri="https://zenodo.org/record/4600362/files/figure.png" pageId="18" pageNumber="19" startId="18.[189,232,1893,1919]" targetBox="[199,1389,1223,1846]" targetPageId="18">
|
||
<paragraph id="8BA736C1E828FFC2FF1FF89AFC3FF878" blockId="18.[188,1399,1893,2026]" pageId="18" pageNumber="19">
|
||
<emphasis id="B96CEAD3E828FFC2FF1FF89AF81CF8ED" bold="true" box="[189,266,1893,1919]" pageId="18" pageNumber="19">Fig. 6.</emphasis>
|
||
Tomentum on pro- and mesofemora of
|
||
<taxonomicName id="4C184D42E828FFC2FD75F89AFA77F8ED" authorityName="Hinton" authorityYear="1941" box="[727,865,1893,1919]" class="Insecta" family="Elmidae" genus="Onychelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96CEAD3E828FFC2FD75F89AFA77F8ED" box="[727,865,1893,1919]" italics="true" pageId="18" pageNumber="19">Onychelmis</emphasis>
|
||
</taxonomicName>
|
||
spp.
|
||
<emphasis id="B96CEAD3E828FFC2FC3DF89AFAA2F8ED" bold="true" box="[927,948,1893,1919]" pageId="18" pageNumber="19">A</emphasis>
|
||
.
|
||
<taxonomicName id="4C184D42E828FFC2FC60F899FDCCF8ED" authority="Deleve, 1968" authorityName="Deleve" authorityYear="1968" box="[962,1242,1893,1920]" class="Insecta" family="Elmidae" genus="Onychelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="leleupi">
|
||
<emphasis id="B96CEAD3E828FFC2FC60F899FD23F8ED" box="[962,1077,1893,1920]" italics="true" pageId="18" pageNumber="19">O. leleupi</emphasis>
|
||
<bibRefCitation id="EF894B30E828FFC2FB99F89AFDCCF8ED" author="Deleve J." box="[1083,1242,1893,1919]" pageId="18" pageNumber="19" pagination="211 - 272" refId="ref16205" refString="Deleve J. 1968. Coleoptera Elminthidae. In: Leleup N. (ed.) Mission zoologique belge aux Iles Galapagos et en Ecuador (N. et J. Leleup, 1964 - 1965). Resultats scientifiques 1: 211 - 272. Institut Royal des Sciences Naturelles de Belgique, Brussels." type="journal article" year="1968">Delève, 1968</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B96CEAD3E828FFC2FB47F899FDEFF812" bold="true" box="[1253,1273,1894,1920]" pageId="18" pageNumber="19">B</emphasis>
|
||
.
|
||
<taxonomicName id="4C184D42E828FFC2FAA5F899FC61F8ED" authority="Linský & Čiamporová-Zaťovičová & Čiampor, 2021" authorityName="Linský & Čiamporová-Zaťovičová & Čiampor" authorityYear="2021" box="[1287,1399,1893,1920]" class="Insecta" family="Elmidae" genus="Onychelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="lenkae" status="sp. nov.">
|
||
<emphasis id="B96CEAD3E828FFC2FAA5F899FC61F8ED" box="[1287,1399,1893,1920]" italics="true" pageId="18" pageNumber="19">O. lenkae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25F57A8E828FFC2FF1EF876F801F831" box="[188,279,1929,1955]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="B96CEAD3E828FFC2FEBCF877F825F830" bold="true" box="[286,307,1928,1954]" pageId="18" pageNumber="19">C</emphasis>
|
||
.
|
||
<taxonomicName id="4C184D42E828FFC2FEE0F876F8A6F831" authority="Linský & Čiamporová-Zaťovičová & Čiampor, 2021" authorityName="Linský & Čiamporová-Zaťovičová & Čiampor" authorityYear="2021" box="[322,432,1929,1955]" class="Insecta" family="Elmidae" genus="Onychelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="lobata" status="sp. nov.">
|
||
<emphasis id="B96CEAD3E828FFC2FEE0F876F8A6F831" box="[322,432,1929,1955]" italics="true" pageId="18" pageNumber="19">O. lobata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25F57A8E828FFC2FE15F876FB07F831" box="[439,529,1929,1955]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="B96CEAD3E828FFC2FDBAF876FB3BF831" bold="true" box="[536,557,1929,1955]" pageId="18" pageNumber="19">D</emphasis>
|
||
.
|
||
<taxonomicName id="4C184D42E828FFC2FD9EF876FA91F831" authority="(Sharp, 1882)" baseAuthorityName="Sharp" baseAuthorityYear="1882" box="[572,903,1929,1955]" class="Insecta" family="Elmidae" genus="Elmis" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="longicollis">
|
||
<emphasis id="B96CEAD3E828FFC2FD9EF876FBCAF831" box="[572,732,1929,1955]" italics="true" pageId="18" pageNumber="19">O. longicollis</emphasis>
|
||
(
|
||
<bibRefCitation id="EF894B30E828FFC2FD49F876FA68F831" author="Sharp D." box="[747,894,1929,1955]" pageId="18" pageNumber="19" pagination="1 - 144" refId="ref17769" refString="Sharp D. 1882. Insecta, Coleoptera: Haliplidae, Dytiscidae, Gyrinidae, Hydrophilidae, Heteroceridae, Parnidae, Georissidae, Cyathoceridae. Biologia Centrali-Americana 1 (2): 1 - 144." type="journal article" year="1882">Sharp, 1882</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B96CEAD3E828FFC2FC30F876FAB0F831" bold="true" box="[914,934,1929,1955]" pageId="18" pageNumber="19">E</emphasis>
|
||
.
|
||
<taxonomicName id="4C184D42E828FFC2FC16F876FD08F830" authority="Linský & Čiamporová-Zaťovičová & Čiampor, 2021" authorityName="Linský & Čiamporová-Zaťovičová & Čiampor" authorityYear="2021" box="[948,1054,1929,1955]" class="Insecta" family="Elmidae" genus="Onychelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="minor" status="sp. nov.">
|
||
<emphasis id="B96CEAD3E828FFC2FC16F876FD08F830" box="[948,1054,1929,1955]" italics="true" pageId="18" pageNumber="19">O. minor</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25F57A8E828FFC2FB87F876FD69F831" box="[1061,1151,1929,1955]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="B96CEAD3E828FFC2FB24F876FD8EF831" bold="true" box="[1158,1176,1929,1955]" pageId="18" pageNumber="19">F</emphasis>
|
||
.
|
||
<taxonomicName id="4C184D42E828FFC2FB04F876FC00F830" authority="Linský & Čiamporová-Zaťovičová & Čiampor, 2021" authorityName="Linský & Čiamporová-Zaťovičová & Čiampor" authorityYear="2021" box="[1190,1302,1929,1955]" class="Insecta" family="Elmidae" genus="Onychelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="onorei" status="sp. nov.">
|
||
<emphasis id="B96CEAD3E828FFC2FB04F876FC00F830" box="[1190,1302,1929,1955]" italics="true" pageId="18" pageNumber="19">O. onorei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25F57A8E828FFC2FABEF876FC61F831" box="[1308,1399,1929,1955]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="B96CEAD3E828FFC2FF1FF854F9C2F857" bold="true" box="[189,212,1963,1989]" pageId="18" pageNumber="19">G</emphasis>
|
||
.
|
||
<taxonomicName id="4C184D42E828FFC2FF41F853F86FF854" authority="Linský & Čiamporová-Zaťovičová & Čiampor, 2021" authorityName="Linský & Čiamporová-Zaťovičová & Čiampor" authorityYear="2021" box="[227,377,1964,1990]" class="Insecta" family="Elmidae" genus="Onychelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="splendida" status="sp. nov.">
|
||
<emphasis id="B96CEAD3E828FFC2FF41F853F86FF854" box="[227,377,1964,1990]" italics="true" pageId="18" pageNumber="19">O. splendida</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25F57A8E828FFC2FE22F853F8CDF854" box="[384,475,1964,1990]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="B96CEAD3E828FFC2FE40F853F8EFF854" bold="true" box="[482,505,1964,1990]" pageId="18" pageNumber="19">H</emphasis>
|
||
.
|
||
<taxonomicName id="4C184D42E828FFC2FDAAF853FAE0F854" authority="Spangler & Santiago, 1991" authorityName="Spangler & Santiago" authorityYear="1991" box="[520,1014,1964,1990]" class="Insecta" family="Elmidae" genus="Onychelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="whiteheadi">
|
||
<emphasis id="B96CEAD3E828FFC2FDAAF853FBBBF854" box="[520,685,1964,1990]" italics="true" pageId="18" pageNumber="19">O. whiteheadi</emphasis>
|
||
<bibRefCitation id="EF894B30E828FFC2FD16F853FAE0F854" author="Spangler P. & Santiago S." box="[692,1014,1964,1990]" pageId="18" pageNumber="19" pagination="495 - 498" refId="ref17990" refString="Spangler P. & Santiago S. 1991. A new species and new records from Colombia of the water beetle genus Onychelmis Hinton (Coleoptera: Elmidae: Elminae). Proceedings of the Entomological Society of Washington 93 (2): 495 - 498." type="journal article" year="1991">Spangler & Santiago, 1991</bibRefCitation>
|
||
</taxonomicName>
|
||
. Drawings are schematic, bodies are divided to identical halves: left = male, right = female (note: females unknown for D and H).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA736C1E829FFC3FF1FFE28FB69FB46" blockId="19.[187,1400,471,1236]" pageId="19" pageNumber="20">
|
||
THORAX. Pronotum widest behind middle, PW:
|
||
<quantity id="4CE09B24E829FFC3FD53FE28FA89FE60" box="[753,927,471,498]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" metricValueMax="4.3" metricValueMin="4.1" pageId="19" pageNumber="20" unit="mm" value="0.42" valueMax="0.43" valueMin="0.41">0.41–0.43 mm</quantity>
|
||
, PL:
|
||
<quantity id="4CE09B24E829FFC3FC7CFE28FD9AFE60" box="[990,1164,471,498]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" metricValueMax="4.9" metricValueMin="4.699999999999999" pageId="19" pageNumber="20" unit="mm" value="0.48" valueMax="0.49" valueMin="0.47">0.47–0.49 mm</quantity>
|
||
; surface shiny, with narrow reticulation along basal margin and posterolateral angles, with dense tiny punctures; sublateral carinae absent, fine raised line indistinct; disc conveX, divided by a broad, deep transverse impression before middle; two prescutellar foveae separated by a raised line extending from base to apical discal half, connecting pronotal halves and merging into them; anterior margin arcuate; posterior margin bisinuate; sides of pronotum convex before and after transverse constriction; lateral margins narrowly rimmed; posterolateral angles orthogonal; anterolateral angles slightly produced. Hypomeron finely microreticulate, widest in middle. Prosternum moderately long in front of procoXae, with anterior margin concave; sides raised around procoxae, forming carinae, not reaching anterior margin, prosternal process long, moderately broad, with posterior margin broadly rounded. Mesoventrite coarse, short and wide, with deep triangular groove for reception of prosternal process; posterior margin around mesocoxae raised. Metaventrite slightly wider than long, shiny; disc convex with very shallow to indistinct, medial, triangular depression in posterior half; discrimen in basal ¾, very thin, almost indistinct; with one prebasal fovea on each side of metaventrite (
|
||
<figureCitation id="13232A44E829FFC3FDC7FC5EFBD4FC2E" box="[613,706,929,956]" captionStart="Fig" captionStartId="17.[189,232,1928,1954]" captionTargetBox="[189,1398,1019,1889]" captionTargetId="figure-149@17.[189,1398,1019,1889]" captionTargetPageId="17" captionText="Fig. 5. Venter and metaventrites of male Onychelmis spp. A, C. O. lenkae sp. nov., paratype, ♂ (PUCE). B. O. splendida sp. nov., paratype, ♂ (PUCE). Arrows show setal tufts. Scale bars: A = 0.5 mm; B–C = 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.4600360" httpUri="https://zenodo.org/record/4600360/files/figure.png" pageId="19" pageNumber="20">Fig. 5B</figureCitation>
|
||
). Elytra (EL:
|
||
<quantity id="4CE09B24E829FFC3FCC4FC5EFD02FC29" box="[870,1044,929,955]" metricMagnitude="-4" metricUnit="m" metricValue="9.05" metricValueMax="9.2" metricValueMin="8.9" pageId="19" pageNumber="20" unit="mm" value="0.905" valueMax="0.92" valueMin="0.89">0.89–0.92 mm</quantity>
|
||
, EW:
|
||
<quantity id="4CE09B24E829FFC3FBFFFC5EFC1DFC29" box="[1117,1291,929,955]" metricMagnitude="-4" metricUnit="m" metricValue="6.6" metricValueMax="6.8" metricValueMin="6.4" pageId="19" pageNumber="20" unit="mm" value="0.66" valueMax="0.68" valueMin="0.64">0.64–0.68 mm</quantity>
|
||
) convex, widest in anterior ¾; sides strongly declivous; surface shiny, with dense tiny punctures; elytral margin narrowly rimmed; humeri protruding from outline; epipleuron tapering posteriorly; sublateral carina absent; strial punctures absent (in few specimens a partial striation is present). Scutellum subovate, flat. Legs moderately long; femora clavate; tibiae longest. Protibiae with anterior cleaning fringe on apical ⅓; mesotibiae with two cleaning fringes – anterior on apical ⅕ and posterior on apical ⅓; metatibiae with posterior cleaning fringe on apical ⅓. Tarsi 5-segmented, first four segments each with one fine pale, recumbent seta, fifth segment slightly shorter than remaining segments combined; claws with a large subbasal and smaller basal teeth.
|
||
</paragraph>
|
||
<paragraph id="8BA736C1E829FFC3FF1FFB03FA98FA31" blockId="19.[189,1398,1276,1443]" pageId="19" pageNumber="20">
|
||
ABDOMEN. With 5 ventrites. First ventrite with basal margin broadly rounded, fifth ventrite longest, apically setose, with posterior margin broadly arcuate. Aedeagus (
|
||
<figureCitation id="13232A44E829FFC3FC66FAE0FD1FFAAB" box="[964,1033,1311,1337]" captionStart="Fig" captionStartId="22.[189,232,1759,1785]" captionTargetBox="[208,1384,280,1715]" captionTargetId="figure-16@22.[189,1398,265,1721]" captionTargetPageId="22" captionText="Fig. 9. Aedeagi of Onychelmis spp. A. O. onorei sp. nov., ventral view. B. O. onorei sp. nov., lateral view. C. O. splendida sp. nov., ventral view. D. O. splendida sp. nov., lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4600370" httpUri="https://zenodo.org/record/4600370/files/figure.png" pageId="19" pageNumber="20">Fig. 9</figureCitation>
|
||
C–D) elongate. Penis elongate, without fibula; corona in basal half; in ventral view concave with rounded apeX; in lateral view evenly narrowed from base to curved apeX. Parameres absent. Phallobase almost 2 times as long as penis, parallel-sided, in ventral view straight, curved in lateral view.
|
||
</paragraph>
|
||
<paragraph id="8BA736C1E829FFC3FF48FA35F853FA76" blockId="19.[189,1398,1482,1579]" box="[234,325,1482,1508]" pageId="19" pageNumber="20">
|
||
<emphasis id="B96CEAD3E829FFC3FF48FA35F853FA76" bold="true" box="[234,325,1482,1508]" pageId="19" pageNumber="20">Female</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BA736C1E829FFC3FF1FFA12FB9BF9B9" blockId="19.[189,1398,1482,1579]" pageId="19" pageNumber="20">EXternally similar to male, eXcept metaventrite without prebasal foveae, fifth ventrite more elongate and extension of femoral tomentum greater.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C302654AE829FFC3FF1FF9ADFD5DF92C" pageId="19" pageNumber="20" type="biology_ecology">
|
||
<paragraph id="8BA736C1E829FFC3FF1FF9ADF80AF9FE" blockId="19.[189,284,1618,1644]" box="[189,284,1618,1644]" pageId="19" pageNumber="20">
|
||
<heading id="D0EF81ADE829FFC3FF1FF9ADF80AF9FE" bold="true" box="[189,284,1618,1644]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
|
||
<emphasis id="B96CEAD3E829FFC3FF1FF9ADF80AF9FE" bold="true" box="[189,284,1618,1644]" pageId="19" pageNumber="20">Biology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA736C1E829FFC3FF1FF97FFD5DF92C" blockId="19.[189,1398,1664,1726]" pageId="19" pageNumber="20">
|
||
Collected from stream ca
|
||
<quantity id="4CE09B24E829FFC3FE4BF97EFB03F909" box="[489,533,1665,1691]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="19" pageNumber="20" unit="m" value="7.0">7 m</quantity>
|
||
wide in deforested area with riffles and pools, close to the confluence with a larger stream, substrate composed of boulders, stones and gravel (
|
||
<figureCitation id="13232A44E829FFC3FC74F95BFD28F92C" box="[982,1086,1700,1726]" captionStart="Fig" captionStartId="27.[189,232,1709,1735]" captionTargetBox="[193,1396,460,1667]" captionTargetId="figure-63@27.[189,1398,456,1669]" captionTargetPageId="27" captionText="Fig. 11. Sampling sites for Onychelmis spp. in Ecuador. A. Morona-Santiago Prov., Limon env., Rio Yungantza, 02°59′49.3″ S, 78°29′18.9″ W, 1522 m a.s.l. B. Napo Prov., road to Coca, 00°43′39.6″ S, 77°45′56.1″ W, 1129 m a.s.l. C. Morona-Santiago Prov., Indanza env., 03°08′38.2″ S, 78°32′10.5″ W, 1299 m a.s.l. D. Morona-Santiago Prov., Santa Rosa de Mamanguy env., Cascada la Encañada, 03°05′14.7″ S, 78°24′36.0″ W, 698 m a.s.l. E. Pastaza Prov., Rio Jantunpaccha, 01°26′30.6″ S, 77°52′25.5″ W, 1011 m a.s.l. F. Napo Prov., road to Coca, Sumaco env., 00°43′29.0″ S, 77°46′01.4″ W, 1109 m a.s.l." figureDoi="http://doi.org/10.5281/zenodo.4600374" httpUri="https://zenodo.org/record/4600374/files/figure.png" pageId="19" pageNumber="20">Fig. 11F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C302654AE829FFC3FF1FF91AFAFEF8BC" box="[189,1000,1765,1838]" pageId="19" pageNumber="20" type="distribution">
|
||
<paragraph id="8BA736C1E829FFC3FF1FF91AF840F96D" blockId="19.[189,342,1765,1791]" box="[189,342,1765,1791]" pageId="19" pageNumber="20">
|
||
<heading id="D0EF81ADE829FFC3FF1FF91AF840F96D" bold="true" box="[189,342,1765,1791]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
|
||
<emphasis id="B96CEAD3E829FFC3FF1FF91AF840F96D" bold="true" box="[189,342,1765,1791]" pageId="19" pageNumber="20">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA736C1E829FFC3FF1FF8EBFAFEF8BC" blockId="19.[189,1000,1812,1838]" box="[189,1000,1812,1838]" pageId="19" pageNumber="20">
|
||
Known from three localities in
|
||
<collectingRegion id="49DCF823E829FFC3FD8EF8EBFBCDF8BC" box="[556,731,1812,1838]" country="Ecuador" name="Napo" pageId="19" pageNumber="20">Napo Province</collectingRegion>
|
||
(
|
||
<figureCitation id="13232A44E829FFC3FD49F8EBFA60F8BC" box="[747,886,1812,1838]" captionStart="Fig" captionStartId="27.[189,232,1709,1735]" captionTargetBox="[193,1396,460,1667]" captionTargetId="figure-63@27.[189,1398,456,1669]" captionTargetPageId="27" captionText="Fig. 11. Sampling sites for Onychelmis spp. in Ecuador. A. Morona-Santiago Prov., Limon env., Rio Yungantza, 02°59′49.3″ S, 78°29′18.9″ W, 1522 m a.s.l. B. Napo Prov., road to Coca, 00°43′39.6″ S, 77°45′56.1″ W, 1129 m a.s.l. C. Morona-Santiago Prov., Indanza env., 03°08′38.2″ S, 78°32′10.5″ W, 1299 m a.s.l. D. Morona-Santiago Prov., Santa Rosa de Mamanguy env., Cascada la Encañada, 03°05′14.7″ S, 78°24′36.0″ W, 698 m a.s.l. E. Pastaza Prov., Rio Jantunpaccha, 01°26′30.6″ S, 77°52′25.5″ W, 1011 m a.s.l. F. Napo Prov., road to Coca, Sumaco env., 00°43′29.0″ S, 77°46′01.4″ W, 1109 m a.s.l." figureDoi="http://doi.org/10.5281/zenodo.4600374" httpUri="https://zenodo.org/record/4600374/files/figure.png" pageId="19" pageNumber="20">Figs 11B, F</figureCitation>
|
||
,
|
||
<figureCitation id="13232A44E829FFC3FC20F8EBFA89F8BC" box="[898,927,1812,1838]" captionStart="Fig" captionStartId="28.[189,232,1916,1942]" captionTargetBox="[189,1398,1158,1876]" captionTargetId="figure-274@28.[189,1398,1158,1876]" captionTargetPageId="28" captionText="Fig. 12. Distribution map of Onychelmis Hinton, 1941. A. New species described within this study. B. Published records of Onychelmis and potential distribution of the genus." figureDoi="http://doi.org/10.5281/zenodo.4600376" httpUri="https://zenodo.org/record/4600376/files/figure.png" pageId="19" pageNumber="20">12</figureCitation>
|
||
A–B).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |