554 lines
100 KiB
XML
554 lines
100 KiB
XML
<document id="ED7E6117EE54A084CA879526313B3290" ID-CLB-Dataset="9742" ID-DOI="10.5852/ejt.2020.720.1119" ID-GBIF-Dataset="60b5a286-c831-49ba-b499-ad61183e7f36" ID-Zenodo-Dep="4085203" ID-ZooBank="A32297C8-00D2-4A71-837A-CE49107C1F27" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1602601192165" checkinUser="valdenar" docAuthor="Wesener, Thomas" docDate="2020" docId="03B187B1B717FFA92523374BFF6C23AC" docLanguage="en" docName="ejt_720_wesener.pdf" docOrigin="European Journal of Taxonomy 720" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Dichromatobolus elephantulus Wesener 2020, gen. et sp. nov." docType="treatment" docUuid="A768C2C3-047C-43A9-8F6B-47EDD40371B4" docUuidSource="ZooBank" docVersion="8" lastPageNumber="117" masterDocId="FF88FFC9B714FFA327203643FF952113" masterDocTitle="Dichromatobolus, a new genus of spirobolidan millipedes from Madagascar (Spirobolida, Pachybolidae)" masterLastPageNumber="120" masterPageNumber="107" pageNumber="110" updateTime="1698846057459" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="59F4465D7FA12EF8662F6E4D8D69D82F">
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<mods:title id="98211356B451EABB38194D9DC6DF5EDB">Dichromatobolus, a new genus of spirobolidan millipedes from Madagascar (Spirobolida, Pachybolidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="C94E162E1EAE9C6A8C834DD0B07FCC98" type="personal">
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<mods:roleTerm id="EFF49B1130B7E105F0B7582528E787F9">Author</mods:roleTerm>
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<mods:namePart id="847D0987FB9AB9406AA57C5AB1E16B99">Wesener, Thomas</mods:namePart>
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<mods:affiliation id="93381B4FC125E4DFC4F760AB2EAB8ABC">Zoological Research Museum Alexander Koenig (ZFMK), Leibniz Institute for Animal Biodiversity, Adenauerallee 160, D- 53113, Bonn, Germany.</mods:affiliation>
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<mods:nameIdentifier id="0E7F3D2E72AC519FF2F21E01713DB55B" type="email">t.wesener@leibniz-zfmk.de</mods:nameIdentifier>
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<mods:typeOfResource id="EE185E007080813F827391E480738716">text</mods:typeOfResource>
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<mods:title id="8556452A2E87653EFEAD1C7E4942DFDB">European Journal of Taxonomy</mods:title>
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<mods:part id="7CF03A7CEBAF408DAD8B380C524C4FE7">
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<mods:date id="7CC6B9408F253613C2FF2D013924BE92">2020</mods:date>
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<mods:number id="61B35324A36DADED9FA00072B5B12F0C">2020-10-12</mods:number>
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<mods:detail id="1B55B067228E5B087BE1F0EE2B0D8071" type="volume">
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<mods:number id="834947EF0DF9A3B1711DF7373CF6A868">720</mods:number>
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<mods:classification id="3A539B87A22996F3A1A8F8BA2B47C9B2">journal article</mods:classification>
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<mods:identifier id="E1E975DDE69B85589015E006210A5FB5" type="DOI">10.5852/ejt.2020.720.1119</mods:identifier>
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<treatment id="03B187B1B717FFA92523374BFF6C23AC" ID-DOI="http://doi.org/10.5281/zenodo.4324177" ID-GBIF-Taxon="168338728" ID-Zenodo-Dep="4324177" LSID="urn:lsid:zoobank.org:act:A768C2C3-047C-43A9-8F6B-47EDD40371B4" httpUri="http://treatment.plazi.org/id/03B187B1B717FFA92523374BFF6C23AC" lastPageId="10" lastPageNumber="117" pageId="3" pageNumber="110">
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<subSubSection id="C302652CB717FFA02523374BFBA42030" box="[515,1073,264,291]" pageId="3" pageNumber="110" type="nomenclature">
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<paragraph id="8BA736A7B717FFA02523374BFBA42030" blockId="3.[365,1223,264,368]" box="[515,1073,264,291]" pageId="3" pageNumber="110">
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<heading id="D0EF81CBB717FFA02523374BFBA42030" box="[515,1073,264,291]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="110" reason="2">
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<taxonomicName id="4C184D24B717FFA02523374BFCEC2030" ID-CoL="5a94bc9b-53c2-4fe0-8333-cc7ca7fa548e" authority="Wesener, 2020" authorityName="Wesener" authorityYear="2020" box="[515,889,264,291]" class="Diplopoda" family="Pachybolidae" genus="Dichromatobolus" kingdom="Animalia" order="Spirobolida" pageId="3" pageNumber="110" phylum="Arthropoda" rank="species" species="elephantulus" status="gen. et sp. nov.">
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<emphasis id="B96CEAB5B717FFA02523374BFCEC2030" bold="true" box="[515,889,264,291]" italics="true" pageId="3" pageNumber="110">Dichromatobolus elephantulus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25F57CEB717FFA024A1374AFBA42030" box="[897,1073,265,291]" pageId="3" pageNumber="110" rank="species">gen. et sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C302652CB717FFA0264D376CFCD82063" pageId="3" pageNumber="110" type="description">
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<paragraph id="8BA736A7B717FFA0264D376CFB52205A" blockId="3.[365,1223,264,368]" box="[365,1223,303,329]" pageId="3" pageNumber="110">
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<uri id="FF893AA5B717FFA0264D376CFB52205A" box="[365,1223,303,329]" pageId="3" pageNumber="110">
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urn:lsid:zoobank.org:act:
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<uuid id="FFBE0C72B717FFA025AE376CFB52205A" box="[654,1223,303,329]" pageId="3" pageNumber="110">A768C2C3-047C-43A9-8F6B-47EDD40371B4</uuid>
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</uri>
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</paragraph>
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<paragraph id="8BA736A7B717FFA025C73715FCD82063" blockId="3.[365,1223,264,368]" box="[743,845,342,368]" pageId="3" pageNumber="110">
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<figureCitation id="13232A22B717FFA025C73715FCD82063" box="[743,845,342,368]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStartId-0="4.[189,232,1786,1812]" captionStartId-1="5.[189,232,1822,1848]" captionStartId-2="6.[189,232,1964,1990]" captionStartId-3="7.[189,232,1784,1810]" captionStartId-4="8.[189,232,1892,1918]" captionStartId-5="9.[189,232,1963,1989]" captionTargetBox-0="[272,1315,265,1748]" captionTargetBox-1="[285,1303,265,1782]" captionTargetBox-2="[189,1397,1311,1924]" captionTargetBox-3="[253,1335,268,1747]" captionTargetBox-4="[189,1398,1021,1852]" captionTargetBox-5="[204,1383,279,1926]" captionTargetId-0="figure-13@4.[272,1315,265,1748]" captionTargetId-1="figure-11@5.[285,1303,265,1782]" captionTargetId-2="figure-314@6.[189,1398,1311,1925]" captionTargetId-3="figure-11@7.[253,1335,265,1747]" captionTargetId-4="figure-239@8.[189,1398,1017,1852]" captionTargetId-5="figure-12@9.[204,1383,265,1926]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="9" captionText-0="Fig. 1. Dichromatobolus elephantulus gen. et sp. nov. A. Living ♂ (red; FMNH-INS 55888) and ♀ (grey; FMNH-INS 55887) from Odenwald Exoten 2007 import, photo by P. Decker. B–C. Living immature ♂, 2019 import (ZFMK MYR8708), photo by L. Moritz. B. Head. C. Whole body. D–F. ♂ (ZFMK MYR8745), multi-layer photographs. D. Anterior body, lateral view; E. Leg pairs 1–7, ventral view. F. Posterior body end with telson. Abbreviations: av = paraprocts; C = lips of paraprocts; Ca = cardines; Co = collum (ring 1); Gn = gnathochilarium; Go = gonopods; IL = incisura lateralis; ms = marginal setae; P-M = parasitic(?) mites; Pre = epiproct; St = stipes; sub = hypoproct. Not to scale." captionText-1="Fig. 2. Dichromatobolus elephantulus gen. et sp. nov., ♂ (ZFMK MYR9896), SEM. A. Right antenna. B. Disc, apical view. C. Border of antennomere 5 to 6 with sensilla basiconica. D. Gnathochilarium, internal surface. E. Left mandible, mesal view. Abbreviations: 4iT = four-combed inner tooth; ac = apical cone; cP = central pads; d = disc; Endo = endochilarium; eP = lateral palpus; eT = external tooth; iP = inner palpus; LT = lateral tooth; mp = molar plate; pL = pectinate lamellae; sb = sensilla bascionica; numbers refer to antennomere number." captionText-2="Fig. 3. Dichromatobolus elephantulus gen. et sp. nov., ♂ (ZFMK MYR9896), SEM. A. Limbus of midbody ring. B. Phoretic(?) mite at body ring. Abbreviations: pm = tergite margin; sc = cuticular scales." captionText-3="Fig. 4. Dichromatobolus elephantulus gen. et sp. nov., holotype, ♂ (CAS MG-54C-78), drawings. A. Gnathochilarium, ventral view. B. Second leg pair, posterior view. C. First leg pair, posterior view. D. Left midbody leg pair, posterior view. E. Anterior gonopod, anterior view. F. Anterior gonopod, posterior view. Abbreviations: cp = coxite process; cP = central pads; Cx = coxite; eP = lateral palpus; Fe = femur; iP = inner palpus; LL = lamellae lingualis; Me = mentum; Po = postfemur; Pre = prefemur; St = sternite; St-Cx = sterno-coxite; T = telopodite; Ta = tarsus; Ti = tibia; Tp = telopodite process. Scale bars = 1 mm." captionText-4="Fig. 5. Dichromatobolus elephantulus gen. et sp. nov., holotype, ♂ (CAS MG-54C-78), left posterior gonopod, drawings. A. Anterior view. B. Posterior view. C. Detail of telopodite, posterior view. Abbreviations: Apo = apodeme; Cx = coxite; sg =efferent dUct; scl = small mesal sclerite; T = telopodite. Scale bars = 1 mm." captionText-5="Fig. 6. Distribution of Dichromatobolus elephantulus gen. et sp. nov. Vegetation color marks modified from Moat & Smith (2007)." figureDoi-0="http://doi.org/10.5281/zenodo.4085205" figureDoi-1="http://doi.org/10.5281/zenodo.4085207" figureDoi-2="http://doi.org/10.5281/zenodo.4085209" figureDoi-3="http://doi.org/10.5281/zenodo.4085212" figureDoi-4="http://doi.org/10.5281/zenodo.4085214" figureDoi-5="http://doi.org/10.5281/zenodo.4085216" httpUri-0="https://zenodo.org/record/4085205/files/figure.png" httpUri-1="https://zenodo.org/record/4085207/files/figure.png" httpUri-2="https://zenodo.org/record/4085209/files/figure.png" httpUri-3="https://zenodo.org/record/4085212/files/figure.png" httpUri-4="https://zenodo.org/record/4085214/files/figure.png" httpUri-5="https://zenodo.org/record/4085216/files/figure.png" pageId="3" pageNumber="110">Figs 1–6</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C302652CB717FFA0279D37D8FE4B2392" pageId="3" pageNumber="110" type="diagnosis">
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<paragraph id="8BA736A7B717FFA0279D37D8FEA320A6" blockId="3.[189,310,411,437]" box="[189,310,411,437]" pageId="3" pageNumber="110">
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<heading id="D0EF81CBB717FFA0279D37D8FEA320A6" bold="true" box="[189,310,411,437]" fontSize="11" level="3" pageId="3" pageNumber="110" reason="3">
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<emphasis id="B96CEAB5B717FFA0279D37D8FEA320A6" bold="true" box="[189,310,411,437]" pageId="3" pageNumber="110">Diagnosis</emphasis>
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</heading>
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||
</paragraph>
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<paragraph id="8BA736A7B717FFA0279D378EFE4B2392" blockId="3.[189,1398,461,641]" pageId="3" pageNumber="110">
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||
Body
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<quantity id="4CE09B42B717FFA02625378EFE1320F4" box="[261,390,461,487]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="3" pageNumber="110" unit="mm" value="70.0" valueMax="80.0" valueMin="60.0">60–80 mm</quantity>
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long, massive, bulky spirobolidan. Epiproct not projecting. Males red, females grey, ozopore openings in both sexes marked by a black circle. Males with tarsal pads, coxa projections absent. Legs and antennae short. Anterior gonopods, telopodite process well-developed and at apex curved laterally. Posterior gonopod slender and elongated, telopodite simple, rectangular, apically with opening of efferent dUct.
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</paragraph>
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</subSubSection>
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<subSubSection id="C302652CB717FFA0279D34EFFCE7220D" pageId="3" pageNumber="110" type="etymology">
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<paragraph id="8BA736A7B717FFA0279D34EFFED123D5" blockId="3.[189,324,684,710]" box="[189,324,684,710]" pageId="3" pageNumber="110">
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<heading id="D0EF81CBB717FFA0279D34EFFED123D5" bold="true" box="[189,324,684,710]" fontSize="11" level="3" pageId="3" pageNumber="110" reason="3">
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<emphasis id="B96CEAB5B717FFA0279D34EFFED123D5" bold="true" box="[189,324,684,710]" pageId="3" pageNumber="110">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BA736A7B717FFA0279D349DFCE7220D" blockId="3.[189,1398,734,799]" pageId="3" pageNumber="110">Elephantulus, diminutive noun in apposition, after the resemblance to an elephant in the female grey color, as well as a hint referring to the width of the species.</paragraph>
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</subSubSection>
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<subSubSection id="C302652CB717FFA0279D350AFBBC2474" pageId="3" pageNumber="110" type="materials_examined">
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<heading id="D0EF81CBB717FFA0279D350AFEF92270" bold="true" box="[189,364,841,867]" fontSize="11" level="3" pageId="3" pageNumber="110" reason="3">
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<emphasis id="B96CEAB5B717FFA0279D350AFEF92270" bold="true" box="[189,364,841,867]" pageId="3" pageNumber="110">Type material</emphasis>
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<paragraph id="8BA736A7B717FFA027CA3538FEC92286" blockId="3.[189,1399,891,995]" box="[234,348,891,917]" pageId="3" pageNumber="110">
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<emphasis id="B96CEAB5B717FFA027CA3538FEC92286" bold="true" box="[234,348,891,917]" pageId="3" pageNumber="110">
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<typeStatus id="54A38805B717FFA027CA3538FEC92286" box="[234,348,891,917]" pageId="3" pageNumber="110" type="holotype">Holotype</typeStatus>
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</emphasis>
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</paragraph>
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<paragraph id="8BA736A7B717FFA0279D35E1FAF322F0" blockId="3.[189,1399,891,995]" pageId="3" pageNumber="110">
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<materialsCitation id="3B703CFAB717FFA0279D35E1FAF322F0" ID-GBIF-Occurrence="2874722308" collectingDate="2011-01-10" collectingDateMax="2011-01-17" collectingDateMin="2011-01-10" collectionCode="CAS" collectorName="M. Irwin & R. Harin Hala" country="Madagascar" elevation="130" latitude="-24.650333" location="Itampolo" longLatPrecision="12" longitude="43.963165" pageId="3" pageNumber="110" specimenCode="CASENT 9068311" specimenCount="1" typeStatus="holotype">
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<collectingCountry id="F30F7637B717FFA0279D35E1FE1B22AF" box="[189,398,930,956]" name="Madagascar" pageId="3" pageNumber="110">MADAGASCAR</collectingCountry>
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• ♂;
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E of
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<location id="8EC7607CB717FFA0257F35E2FD5822A8" LSID="urn:lsid:plazi:treatment:03B187B1B717FFA92523374BFF6C23AC:8EC7607CB717FFA0257F35E2FD5822A8" box="[607,717,929,955]" country="Madagascar" latitude="-24.650333" longLatPrecision="12" longitude="43.963165" name="Itampolo" pageId="3" pageNumber="110">Itampolo</location>
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;
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<geoCoordinate id="EE2C5060B717FFA025FA35E3FCF322AF" box="[730,870,928,956]" degrees="24" direction="south" minutes="39.02" orientation="latitude" pageId="3" pageNumber="110" precision="9" value="-24.650333">24°39.02′ S</geoCoordinate>
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,
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<geoCoordinate id="EE2C5060B717FFA0245435E3FB9622AF" box="[884,1027,928,956]" degrees="43" direction="east" minutes="57.79" orientation="longitude" pageId="3" pageNumber="110" precision="9" value="43.963165">43°57.79′ E</geoCoordinate>
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;
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<quantity id="4CE09B42B717FFA0233135E1FBC922AF" box="[1041,1116,930,956]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="110" unit="m" value="130.0">130 m</quantity>
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a.s.l.
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</elevation>
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;
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<date id="FFA61067B717FFA0238C35E1FAE722AF" box="[1196,1394,930,956]" pageId="3" pageNumber="110" value="2011-01-10" valueMax="2011-01-17" valueMin="2011-01-10">
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<collectingDate id="EFE2E98FB717FFA0238C35E1FAE722AF" box="[1196,1394,930,956]" pageId="3" pageNumber="110" value="2011-01-10" valueMax="2011-01-17" valueMin="2011-01-10">10–17 Jan. 2011</collectingDate>
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</date>
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;
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<collectorName id="26ED5371B717FFA0279D358AFEBD22F1" box="[189,296,968,995]" pageId="3" pageNumber="110">M. Irwin</collectorName>
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,
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<collectorName id="26ED5371B717FFA02614358AFE4222F1" box="[308,471,968,995]" pageId="3" pageNumber="110">R. Harin Hala</collectorName>
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leg.;
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<collectingMethod id="52594EB0B717FFA0253A358BFD3A22F0" box="[538,687,968,995]" pageId="3" pageNumber="110">Malaise trap</collectingMethod>
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; path in dry forest;
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<collectionCode id="ED09AE62B717FFA024B4358BFC5822F1" box="[916,973,968,994]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="3" pageNumber="110" type="Museum">CAS</collectionCode>
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MG-54C-78 (
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<specimenCode id="DBBE9EDCB717FFA02354358BFACF22F0" box="[1140,1370,968,995]" collectionCode="CASENT" pageId="3" pageNumber="110">CASENT 9068311</specimenCode>
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).
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<paragraph id="8BA736A7B717FFA0279D324FFE6C2535" blockId="3.[189,505,1036,1062]" box="[189,505,1036,1062]" pageId="3" pageNumber="110">
|
||
<heading id="D0EF81CBB717FFA0279D324FFE6C2535" bold="true" box="[189,505,1036,1062]" fontSize="11" level="3" pageId="3" pageNumber="110" reason="3">
|
||
<emphasis id="B96CEAB5B717FFA0279D324FFE6C2535" bold="true" box="[189,505,1036,1062]" pageId="3" pageNumber="110">Other material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA736A7B717FFA0279D327CFBBC2474" blockId="3.[189,1399,1085,1383]" pageId="3" pageNumber="110">
|
||
<materialsCitation id="3B703CFAB717FFA0279D327CFDF725B4" ID-GBIF-Occurrence="2874722305" collectingDate="2002-02-21" collectingDateMax="2002-02-25" collectingDateMin="2002-02-21" collectionCode="CAS" collectorName="Fisher & Griswold" country="Madagascar" elevation="80" latitude="-24.65361" location="Mahafaly" longLatPrecision="21" longitude="43.996666" municipality="Province" pageId="3" pageNumber="110" specimenCode="BLF 5762" specimenCount="1" stateProvince="Toliara">
|
||
<collectingCountry id="F30F7637B717FFA0279D327CFE1B254A" box="[189,398,1087,1113]" name="Madagascar" pageId="3" pageNumber="110">MADAGASCAR</collectingCountry>
|
||
• 2 ♂♂;
|
||
<collectingMunicipality id="6BC3ACDDB717FFA026CF327CFDC2254A" box="[495,599,1087,1113]" pageId="3" pageNumber="110">Province</collectingMunicipality>
|
||
<collectingRegion id="49DCF845B717FFA0257C327CFD27254A" box="[604,690,1087,1113]" country="Madagascar" name="Toliara" pageId="3" pageNumber="110">Toliara</collectingRegion>
|
||
,
|
||
<location id="8EC7607CB717FFA0259B327CFCBE254A" LSID="urn:lsid:plazi:treatment:03B187B1B717FFA92523374BFF6C23AC:8EC7607CB717FFA0259B327CFCBE254A" box="[699,811,1087,1113]" country="Madagascar" latitude="-24.65361" longLatPrecision="21" longitude="43.996666" municipality="Province" name="Mahafaly" pageId="3" pageNumber="110" stateProvince="Toliara">Mahafaly</location>
|
||
PlateaU,
|
||
<locationDeviation id="9A152F85B717FFA024B3327CFB4F254A" box="[915,1242,1087,1113]" location="Itampolo" pageId="3" pageNumber="110">
|
||
<quantity id="4CE09B42B717FFA024B3327CFC77254A" box="[915,994,1087,1113]" metricMagnitude="3" metricUnit="m" metricValue="6.2" pageId="3" pageNumber="110" unit="km" value="6.2">6.2 km</quantity>
|
||
74°ENE of Itampolo
|
||
</locationDeviation>
|
||
;
|
||
<geoCoordinate id="EE2C5060B717FFA023C3327EFAE4254A" box="[1251,1393,1085,1113]" degrees="24" direction="south" minutes="39" orientation="latitude" pageId="3" pageNumber="110" precision="15" seconds="13" value="-24.65361">24°39′13″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE2C5060B717FFA0279D3227FED92593" box="[189,332,1124,1152]" degrees="43" direction="east" minutes="59" orientation="longitude" pageId="3" pageNumber="110" precision="15" seconds="48" value="43.996666">43°59′48″ E</geoCoordinate>
|
||
;
|
||
<elevation id="0035D194B717FFA026753226FE5F256C" box="[341,458,1125,1152]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="110" unit="m" value="80.0">
|
||
<quantity id="4CE09B42B717FFA026753226FE1B2593" box="[341,398,1125,1152]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="110" unit="m" value="80.0">80 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
;
|
||
<date id="FFA61067B717FFA026F43226FD02256C" box="[468,663,1125,1152]" pageId="3" pageNumber="110" value="2002-02-21" valueMax="2002-02-25" valueMin="2002-02-21">
|
||
<collectingDate id="EFE2E98FB717FFA026F43226FD02256C" box="[468,663,1125,1152]" pageId="3" pageNumber="110" value="2002-02-21" valueMax="2002-02-25" valueMin="2002-02-21">21–25 Feb. 2002</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26ED5371B717FFA025803226FD7E256C" box="[672,747,1125,1151]" pageId="3" pageNumber="110">Fisher</collectorName>
|
||
,
|
||
<collectorName id="26ED5371B717FFA025D33226FCF5256C" box="[755,864,1125,1151]" pageId="3" pageNumber="110">Griswold</collectorName>
|
||
<emphasis id="B96CEAB5B717FFA024453225FC09256C" box="[869,924,1125,1151]" italics="true" pageId="3" pageNumber="110">et al.</emphasis>
|
||
leg.; spiny forest thicket; hand collecting;
|
||
<collectionCode id="ED09AE62B717FFA0279D32CFFF6325B5" box="[189,246,1164,1190]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="3" pageNumber="110" type="Museum">CAS</collectionCode>
|
||
<specimenCode id="DBBE9EDCB717FFA027DD32CFFEE025B4" box="[253,373,1164,1191]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="3" pageNumber="110" type="Museum">BLF 5762</specimenCode>
|
||
(GCKMP-FM-060)
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B703CFAB717FFA0255A32CFFDF125E7" ID-GBIF-Occurrence="2874722302" collectingDate="2005-04-19" collectorName="Voahangy Soarimalala" country="Madagascar" elevation="50" latitude="-24.188334" location="Soarano village" longLatPrecision="125" longitude="43.77833" municipality="Atsimo-Andrefena" pageId="3" pageNumber="110" specimenCount="2" specimenCount-female="2" stateProvince="Toliara">
|
||
<specimenCount id="9D1EFD2EB717FFA0255A32CFFD2825B5" box="[634,701,1164,1190]" pageId="3" pageNumber="110" type="female">2 ♀♀</specimenCount>
|
||
;
|
||
<collectingRegion id="49DCF845B717FFA025EB32CFFCB425B5" box="[715,801,1164,1190]" country="Madagascar" name="Toliara" pageId="3" pageNumber="110">Toliara</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6BC3ACDDB717FFA0240A32CFFB9C25B5" box="[810,1033,1164,1190]" pageId="3" pageNumber="110">Atsimo-Andrefena</collectingMunicipality>
|
||
,
|
||
<locationDeviation id="9A152F85B717FFA0233532CFFAA525B5" box="[1045,1328,1164,1191]" location="Efoetse" pageId="3" pageNumber="110">
|
||
<quantity id="4CE09B42B717FFA0233532CFFBE125B5" box="[1045,1140,1164,1191]" metricMagnitude="4" metricUnit="m" metricValue="1.35" pageId="3" pageNumber="110" unit="km" value="13.5">13.5 km</quantity>
|
||
SSE of Efoetse
|
||
</locationDeviation>
|
||
,
|
||
<quantity id="4CE09B42B717FFA0221B32CFFAE325B5" box="[1339,1398,1164,1190]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="110" unit="km" value="2.0">2 km</quantity>
|
||
E of
|
||
<location id="8EC7607CB717FFA027D432F1FE3825DF" LSID="urn:lsid:plazi:treatment:03B187B1B717FFA92523374BFF6C23AC:8EC7607CB717FFA027D432F1FE3825DF" box="[244,429,1202,1228]" country="Madagascar" latitude="-24.188334" longLatPrecision="125" longitude="43.77833" municipality="Atsimo-Andrefena" name="Soarano village" pageId="3" pageNumber="110" stateProvince="Toliara">Soarano village</location>
|
||
;
|
||
<geoCoordinate id="EE2C5060B717FFA0269832F2FDA725DF" box="[440,562,1201,1229]" degrees="24" direction="south" minutes="11.3" orientation="latitude" pageId="3" pageNumber="110" precision="92" value="-24.188334">24°11.3′ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE2C5060B717FFA0251E32F2FD2925DE" box="[574,700,1201,1229]" degrees="43" direction="east" minutes="46.7" orientation="longitude" pageId="3" pageNumber="110" precision="92" value="43.77833">43°46.7′ E</geoCoordinate>
|
||
;
|
||
<elevation id="0035D194B717FFA025E732F1FCD525DF" box="[711,832,1202,1229]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="110" unit="m" value="50.0">
|
||
<quantity id="4CE09B42B717FFA025E732F1FC9425DE" box="[711,769,1202,1229]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="110" unit="m" value="50.0">50 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
;
|
||
<date id="FFA61067B717FFA0246C32F1FC7325DE" box="[844,998,1202,1229]" pageId="3" pageNumber="110" value="2005-04-19">
|
||
<collectingDate id="EFE2E98FB717FFA0246C32F1FC7325DE" box="[844,998,1202,1229]" pageId="3" pageNumber="110" value="2005-04-19">19 Apr. 2005</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26ED5371B717FFA024D032F1FB6225DF" box="[1008,1271,1202,1228]" pageId="3" pageNumber="110">Voahangy Soarimalala</collectorName>
|
||
leg.; spiny bUsh,
|
||
<collectingMethod id="52594EB0B717FFA02624329AFECC25E0" box="[260,345,1241,1267]" pageId="3" pageNumber="110">pitfalls</collectingMethod>
|
||
; FMNH-INS 3196507
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B703CFAB717FFA025A0329AFE932409" ID-GBIF-Occurrence="2874722303" collectorName="Germany" country="Madagascar" location="Odenwald Exoten" pageId="3" pageNumber="110" specimenCount="1">
|
||
<specimenCount id="9D1EFD2EB717FFA025A0329AFD5025E0" box="[640,709,1241,1267]" pageId="3" pageNumber="110" type="male">7 ♂♂</specimenCount>
|
||
;
|
||
<location id="8EC7607CB717FFA025F4329AFC3D25E0" LSID="urn:lsid:plazi:treatment:03B187B1B717FFA92523374BFF6C23AC:8EC7607CB717FFA025F4329AFC3D25E0" box="[724,936,1241,1267]" country="Madagascar" name="Odenwald Exoten" pageId="3" pageNumber="110">Odenwald Exoten</location>
|
||
import to
|
||
<collectorName id="26ED5371B717FFA02307329AFB0125E0" box="[1063,1172,1241,1267]" pageId="3" pageNumber="110">
|
||
<collectingCountry id="F30F7637B717FFA02307329AFB0125E0" box="[1063,1172,1241,1267]" name="Germany" pageId="3" pageNumber="110">Germany</collectingCountry>
|
||
</collectorName>
|
||
2007; FMNH-INS 55888
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B703CFAB717FFA0263D3343FC1E2409" ID-GBIF-Occurrence="2874722301" box="[285,907,1279,1306]" collectorName="Germany" country="Madagascar" location="Odenwald Exoten" pageId="3" pageNumber="110" specimenCount="17" specimenCount-female="17">
|
||
<specimenCount id="9D1EFD2EB717FFA0263D3343FEF82409" box="[285,365,1280,1306]" pageId="3" pageNumber="110" type="female">17 ♀♀</specimenCount>
|
||
; same data as for previoUs; FMNH-INS 55887
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B703CFAB717FFA024823343FDC62453" ID-GBIF-Occurrence="2874722306" collectionCode="ZFMK" collectorName="Germany" country="Madagascar" location="Odenwald Exoten" pageId="3" pageNumber="110" specimenCode="MYR9897, MYR9898, MYR9899" specimenCount="3" specimenCount-female="3">
|
||
<specimenCount id="9D1EFD2EB717FFA024823343FC712409" box="[930,996,1280,1306]" pageId="3" pageNumber="110" type="female">3 ♀♀</specimenCount>
|
||
; same data as for previoUs;
|
||
<collectionCode id="ED09AE62B717FFA022053343FAE32409" box="[1317,1398,1280,1306]" collectionName="Germany, Bonn, Zoologische Forschungsinstitut und Museum "Alexander Koenig"" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="3" pageNumber="110" type="Museum">ZFMK</collectionCode>
|
||
<specimenCode id="DBBE9EDCB717FFA0279D3365FEAB2453" box="[189,318,1318,1344]" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="3" pageNumber="110" type="Museum">MYR9897</specimenCode>
|
||
,
|
||
<specimenCode id="DBBE9EDCB717FFA026693365FE5C2453" box="[329,457,1318,1344]" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="3" pageNumber="110" type="Museum">MYR9898</specimenCode>
|
||
,
|
||
<specimenCode id="DBBE9EDCB717FFA026F63365FDC62453" box="[470,595,1318,1344]" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="3" pageNumber="110" type="Museum">MYR9899</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B703CFAB717FFA0254C3365FB5B2453" ID-GBIF-Occurrence="2874722307" box="[620,1230,1318,1345]" collectionCode="ZFMK" collectorName="Germany" country="Madagascar" location="Odenwald Exoten" pageId="3" pageNumber="110" specimenCode="MYR9896" specimenCount="1">
|
||
<specimenCount id="9D1EFD2EB717FFA0254C3365FD252452" box="[620,688,1318,1345]" pageId="3" pageNumber="110" type="male">2 ♂♂</specimenCount>
|
||
; same data as for previoUs;
|
||
<collectionCode id="ED09AE62B717FFA024D83364FBDC2452" box="[1016,1097,1319,1345]" collectionName="Germany, Bonn, Zoologische Forschungsinstitut und Museum "Alexander Koenig"" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="3" pageNumber="110" type="Museum">ZFMK</collectionCode>
|
||
<specimenCode id="DBBE9EDCB717FFA023713365FB5B2453" box="[1105,1230,1318,1344]" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="3" pageNumber="110" type="Museum">MYR9896</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B703CFAB717FFA023C83365FBB02474" ID-GBIF-Occurrence="2874722304" collectionCode="ZFMK" collectorName="France" country="Madagascar" pageId="3" pageNumber="110" specimenCode="MYR8708, MYR8745" specimenCount="3" specimenCount-male="1">
|
||
<specimenCount id="9D1EFD2EB717FFA023C83365FAE52452" box="[1256,1392,1318,1345]" pageId="3" pageNumber="110" type="male">2 imm. ♂♂</specimenCount>
|
||
; pet trade, ebugz.eu; import to
|
||
<collectingCountry id="F30F7637B717FFA0253E330EFDFB2474" box="[542,622,1357,1383]" name="France" pageId="3" pageNumber="110">France</collectingCountry>
|
||
2019;
|
||
<collectionCode id="ED09AE62B717FFA0259F330EFC852474" box="[703,784,1357,1383]" collectionName="Germany, Bonn, Zoologische Forschungsinstitut und Museum "Alexander Koenig"" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="3" pageNumber="110" type="Museum">ZFMK</collectionCode>
|
||
<specimenCode id="DBBE9EDCB717FFA02438330EFC0D2474" box="[792,920,1357,1383]" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="3" pageNumber="110" type="Museum">MYR8708</specimenCode>
|
||
,
|
||
<specimenCode id="DBBE9EDCB717FFA02484330FFBB02474" box="[932,1061,1356,1383]" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="3" pageNumber="110" type="Museum">MYR8745</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C302652CB717FFAB279D33D2FB8B22D9" lastPageId="8" lastPageNumber="115" pageId="3" pageNumber="110" type="description">
|
||
<paragraph id="8BA736A7B717FFA0279D33D2FEDB24B8" blockId="3.[189,334,1425,1451]" box="[189,334,1425,1451]" pageId="3" pageNumber="110">
|
||
<heading id="D0EF81CBB717FFA0279D33D2FEDB24B8" bold="true" box="[189,334,1425,1451]" fontSize="11" level="3" pageId="3" pageNumber="110" reason="3">
|
||
<emphasis id="B96CEAB5B717FFA0279D33D2FEDB24B8" bold="true" box="[189,334,1425,1451]" pageId="3" pageNumber="110">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA736A7B717FFA0279D3380FAE22717" blockId="3.[189,1399,1475,1541]" pageId="3" pageNumber="110">
|
||
MEASUREMENTS. Male
|
||
<typeStatus id="54A38805B717FFA026E83380FDA624CE" box="[456,563,1475,1501]" pageId="3" pageNumber="110" type="holotype">holotype</typeStatus>
|
||
: 47 +0 body rings,
|
||
<quantity id="4CE09B42B717FFA024363380FCFC24CD" box="[790,873,1475,1502]" metricMagnitude="-2" metricUnit="m" metricValue="6.2" pageId="3" pageNumber="110" unit="mm" value="62.0">62 mm</quantity>
|
||
long,
|
||
<quantity id="4CE09B42B717FFA024933387FB9824CD" box="[947,1037,1476,1502]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="3" pageNumber="110" unit="mm" value="7.7">7.7 mm</quantity>
|
||
wide. Widest male (CAS BLF 5762), broken,
|
||
<quantity id="4CE09B42B717FFA0264F33A9FE5D2717" box="[367,456,1514,1540]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="3" pageNumber="110" unit="mm" value="8.2">8.2 mm</quantity>
|
||
wide. Female (ZFMK MYR9899): 48 +0 body rings,
|
||
<quantity id="4CE09B42B717FFA0231D33A9FB1A2717" box="[1085,1167,1514,1540]" metricMagnitude="-2" metricUnit="m" metricValue="7.6" pageId="3" pageNumber="110" unit="mm" value="76.0">76 mm</quantity>
|
||
long, 9.0 mm wide.
|
||
</paragraph>
|
||
<paragraph id="8BA736A7B717FFA0279D306CFE7327AF" blockId="3.[189,1399,1582,1725]" pageId="3" pageNumber="110">
|
||
COLORATION. Sexually dimorphic (
|
||
<figureCitation id="13232A22B717FFA02574306DFD24275A" box="[596,689,1582,1609]" captionStart="Fig" captionStartId="4.[189,232,1786,1812]" captionTargetBox="[272,1315,265,1748]" captionTargetId="figure-13@4.[272,1315,265,1748]" captionTargetPageId="4" captionText="Fig. 1. Dichromatobolus elephantulus gen. et sp. nov. A. Living ♂ (red; FMNH-INS 55888) and ♀ (grey; FMNH-INS 55887) from Odenwald Exoten 2007 import, photo by P. Decker. B–C. Living immature ♂, 2019 import (ZFMK MYR8708), photo by L. Moritz. B. Head. C. Whole body. D–F. ♂ (ZFMK MYR8745), multi-layer photographs. D. Anterior body, lateral view; E. Leg pairs 1–7, ventral view. F. Posterior body end with telson. Abbreviations: av = paraprocts; C = lips of paraprocts; Ca = cardines; Co = collum (ring 1); Gn = gnathochilarium; Go = gonopods; IL = incisura lateralis; ms = marginal setae; P-M = parasitic(?) mites; Pre = epiproct; St = stipes; sub = hypoproct. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4085205" httpUri="https://zenodo.org/record/4085205/files/figure.png" pageId="3" pageNumber="110">Fig. 1A</figureCitation>
|
||
). Male: head and body reddish (
|
||
<figureCitation id="13232A22B717FFA02306306DFBF3275A" box="[1062,1126,1582,1609]" captionStart="Fig" captionStartId="4.[189,232,1786,1812]" captionTargetBox="[272,1315,265,1748]" captionTargetId="figure-13@4.[272,1315,265,1748]" captionTargetPageId="4" captionText="Fig. 1. Dichromatobolus elephantulus gen. et sp. nov. A. Living ♂ (red; FMNH-INS 55888) and ♀ (grey; FMNH-INS 55887) from Odenwald Exoten 2007 import, photo by P. Decker. B–C. Living immature ♂, 2019 import (ZFMK MYR8708), photo by L. Moritz. B. Head. C. Whole body. D–F. ♂ (ZFMK MYR8745), multi-layer photographs. D. Anterior body, lateral view; E. Leg pairs 1–7, ventral view. F. Posterior body end with telson. Abbreviations: av = paraprocts; C = lips of paraprocts; Ca = cardines; Co = collum (ring 1); Gn = gnathochilarium; Go = gonopods; IL = incisura lateralis; ms = marginal setae; P-M = parasitic(?) mites; Pre = epiproct; St = stipes; sub = hypoproct. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4085205" httpUri="https://zenodo.org/record/4085205/files/figure.png" pageId="3" pageNumber="110">Fig. 1</figureCitation>
|
||
A–E), telson bright red (
|
||
<figureCitation id="13232A22B717FFA027E63016FE89277C" box="[198,284,1621,1647]" captionStart="Fig" captionStartId="4.[189,232,1786,1812]" captionTargetBox="[272,1315,265,1748]" captionTargetId="figure-13@4.[272,1315,265,1748]" captionTargetPageId="4" captionText="Fig. 1. Dichromatobolus elephantulus gen. et sp. nov. A. Living ♂ (red; FMNH-INS 55888) and ♀ (grey; FMNH-INS 55887) from Odenwald Exoten 2007 import, photo by P. Decker. B–C. Living immature ♂, 2019 import (ZFMK MYR8708), photo by L. Moritz. B. Head. C. Whole body. D–F. ♂ (ZFMK MYR8745), multi-layer photographs. D. Anterior body, lateral view; E. Leg pairs 1–7, ventral view. F. Posterior body end with telson. Abbreviations: av = paraprocts; C = lips of paraprocts; Ca = cardines; Co = collum (ring 1); Gn = gnathochilarium; Go = gonopods; IL = incisura lateralis; ms = marginal setae; P-M = parasitic(?) mites; Pre = epiproct; St = stipes; sub = hypoproct. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4085205" httpUri="https://zenodo.org/record/4085205/files/figure.png" pageId="3" pageNumber="110">Fig. 1F</figureCitation>
|
||
), ozopores marked by black spot. Legs and antennae black (
|
||
<figureCitation id="13232A22B717FFA024F43016FB81277C" box="[980,1044,1621,1647]" captionStart="Fig" captionStartId="4.[189,232,1786,1812]" captionTargetBox="[272,1315,265,1748]" captionTargetId="figure-13@4.[272,1315,265,1748]" captionTargetPageId="4" captionText="Fig. 1. Dichromatobolus elephantulus gen. et sp. nov. A. Living ♂ (red; FMNH-INS 55888) and ♀ (grey; FMNH-INS 55887) from Odenwald Exoten 2007 import, photo by P. Decker. B–C. Living immature ♂, 2019 import (ZFMK MYR8708), photo by L. Moritz. B. Head. C. Whole body. D–F. ♂ (ZFMK MYR8745), multi-layer photographs. D. Anterior body, lateral view; E. Leg pairs 1–7, ventral view. F. Posterior body end with telson. Abbreviations: av = paraprocts; C = lips of paraprocts; Ca = cardines; Co = collum (ring 1); Gn = gnathochilarium; Go = gonopods; IL = incisura lateralis; ms = marginal setae; P-M = parasitic(?) mites; Pre = epiproct; St = stipes; sub = hypoproct. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4085205" httpUri="https://zenodo.org/record/4085205/files/figure.png" pageId="3" pageNumber="110">Fig. 1</figureCitation>
|
||
B–F). Female: Head and body including telson grey (
|
||
<figureCitation id="13232A22B717FFA026E5303FFDB72785" box="[453,546,1660,1686]" captionStart="Fig" captionStartId="4.[189,232,1786,1812]" captionTargetBox="[272,1315,265,1748]" captionTargetId="figure-13@4.[272,1315,265,1748]" captionTargetPageId="4" captionText="Fig. 1. Dichromatobolus elephantulus gen. et sp. nov. A. Living ♂ (red; FMNH-INS 55888) and ♀ (grey; FMNH-INS 55887) from Odenwald Exoten 2007 import, photo by P. Decker. B–C. Living immature ♂, 2019 import (ZFMK MYR8708), photo by L. Moritz. B. Head. C. Whole body. D–F. ♂ (ZFMK MYR8745), multi-layer photographs. D. Anterior body, lateral view; E. Leg pairs 1–7, ventral view. F. Posterior body end with telson. Abbreviations: av = paraprocts; C = lips of paraprocts; Ca = cardines; Co = collum (ring 1); Gn = gnathochilarium; Go = gonopods; IL = incisura lateralis; ms = marginal setae; P-M = parasitic(?) mites; Pre = epiproct; St = stipes; sub = hypoproct. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4085205" httpUri="https://zenodo.org/record/4085205/files/figure.png" pageId="3" pageNumber="110">Fig. 1A</figureCitation>
|
||
), ozopores marked by black spot. Posterior margin of body rings darker. Legs and antennae black.
|
||
</paragraph>
|
||
<paragraph id="8BA736A7B717FFA0279D30A4FB9626FB" blockId="3.[189,1399,1767,2025]" pageId="3" pageNumber="110">
|
||
HEAD. In male, each eye with 45–50 ommatidia arranged in 10 vertical rows with: 4, 5, 6, 7, 7, 6, 5, 4, 3, 2 ommatidia, respectively (
|
||
<figureCitation id="13232A22B717FFA02562314EFD52263B" box="[578,711,1805,1832]" captionStart="Fig" captionStartId="4.[189,232,1786,1812]" captionTargetBox="[272,1315,265,1748]" captionTargetId="figure-13@4.[272,1315,265,1748]" captionTargetPageId="4" captionText="Fig. 1. Dichromatobolus elephantulus gen. et sp. nov. A. Living ♂ (red; FMNH-INS 55888) and ♀ (grey; FMNH-INS 55887) from Odenwald Exoten 2007 import, photo by P. Decker. B–C. Living immature ♂, 2019 import (ZFMK MYR8708), photo by L. Moritz. B. Head. C. Whole body. D–F. ♂ (ZFMK MYR8745), multi-layer photographs. D. Anterior body, lateral view; E. Leg pairs 1–7, ventral view. F. Posterior body end with telson. Abbreviations: av = paraprocts; C = lips of paraprocts; Ca = cardines; Co = collum (ring 1); Gn = gnathochilarium; Go = gonopods; IL = incisura lateralis; ms = marginal setae; P-M = parasitic(?) mites; Pre = epiproct; St = stipes; sub = hypoproct. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4085205" httpUri="https://zenodo.org/record/4085205/files/figure.png" pageId="3" pageNumber="110">Fig. 1B, D</figureCitation>
|
||
). Incisura lateralis open (
|
||
<figureCitation id="13232A22B717FFA024D8314EFBC2263B" box="[1016,1111,1805,1832]" captionStart="Fig" captionStartId="4.[189,232,1786,1812]" captionTargetBox="[272,1315,265,1748]" captionTargetId="figure-13@4.[272,1315,265,1748]" captionTargetPageId="4" captionText="Fig. 1. Dichromatobolus elephantulus gen. et sp. nov. A. Living ♂ (red; FMNH-INS 55888) and ♀ (grey; FMNH-INS 55887) from Odenwald Exoten 2007 import, photo by P. Decker. B–C. Living immature ♂, 2019 import (ZFMK MYR8708), photo by L. Moritz. B. Head. C. Whole body. D–F. ♂ (ZFMK MYR8745), multi-layer photographs. D. Anterior body, lateral view; E. Leg pairs 1–7, ventral view. F. Posterior body end with telson. Abbreviations: av = paraprocts; C = lips of paraprocts; Ca = cardines; Co = collum (ring 1); Gn = gnathochilarium; Go = gonopods; IL = incisura lateralis; ms = marginal setae; P-M = parasitic(?) mites; Pre = epiproct; St = stipes; sub = hypoproct. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4085205" httpUri="https://zenodo.org/record/4085205/files/figure.png" pageId="3" pageNumber="110">Fig. 1D</figureCitation>
|
||
). Labrum with standard three irregular teeth and single row of 10–12 stout marginal setae (
|
||
<figureCitation id="13232A22B717FFA024E23177FB8E265D" box="[962,1051,1844,1870]" captionStart="Fig" captionStartId="4.[189,232,1786,1812]" captionTargetBox="[272,1315,265,1748]" captionTargetId="figure-13@4.[272,1315,265,1748]" captionTargetPageId="4" captionText="Fig. 1. Dichromatobolus elephantulus gen. et sp. nov. A. Living ♂ (red; FMNH-INS 55888) and ♀ (grey; FMNH-INS 55887) from Odenwald Exoten 2007 import, photo by P. Decker. B–C. Living immature ♂, 2019 import (ZFMK MYR8708), photo by L. Moritz. B. Head. C. Whole body. D–F. ♂ (ZFMK MYR8745), multi-layer photographs. D. Anterior body, lateral view; E. Leg pairs 1–7, ventral view. F. Posterior body end with telson. Abbreviations: av = paraprocts; C = lips of paraprocts; Ca = cardines; Co = collum (ring 1); Gn = gnathochilarium; Go = gonopods; IL = incisura lateralis; ms = marginal setae; P-M = parasitic(?) mites; Pre = epiproct; St = stipes; sub = hypoproct. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4085205" httpUri="https://zenodo.org/record/4085205/files/figure.png" pageId="3" pageNumber="110">Fig. 1E</figureCitation>
|
||
). Clypeus with two setiferous foveolae on each side. Antennae short, protruding back to ring
|
||
<quantity id="4CE09B42B717FFA024BF3118FC592666" box="[927,972,1883,1909]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="3" pageNumber="110" unit="in" value="3.0">3 in</quantity>
|
||
males (
|
||
<figureCitation id="13232A22B717FFA023073118FB172666" box="[1063,1154,1883,1909]" captionStart="Fig" captionStartId="4.[189,232,1786,1812]" captionTargetBox="[272,1315,265,1748]" captionTargetId="figure-13@4.[272,1315,265,1748]" captionTargetPageId="4" captionText="Fig. 1. Dichromatobolus elephantulus gen. et sp. nov. A. Living ♂ (red; FMNH-INS 55888) and ♀ (grey; FMNH-INS 55887) from Odenwald Exoten 2007 import, photo by P. Decker. B–C. Living immature ♂, 2019 import (ZFMK MYR8708), photo by L. Moritz. B. Head. C. Whole body. D–F. ♂ (ZFMK MYR8745), multi-layer photographs. D. Anterior body, lateral view; E. Leg pairs 1–7, ventral view. F. Posterior body end with telson. Abbreviations: av = paraprocts; C = lips of paraprocts; Ca = cardines; Co = collum (ring 1); Gn = gnathochilarium; Go = gonopods; IL = incisura lateralis; ms = marginal setae; P-M = parasitic(?) mites; Pre = epiproct; St = stipes; sub = hypoproct. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4085205" httpUri="https://zenodo.org/record/4085205/files/figure.png" pageId="3" pageNumber="110">Fig. 1B</figureCitation>
|
||
). Relative lengths of antennomeres: 1<< 2>> 3=4>5= 6, second antennomere twice as long as others (
|
||
<figureCitation id="13232A22B717FFA023B631C2FB632688" box="[1174,1270,1921,1947]" captionStart="Fig" captionStartId="5.[189,232,1822,1848]" captionTargetBox="[285,1303,265,1782]" captionTargetId="figure-11@5.[285,1303,265,1782]" captionTargetPageId="5" captionText="Fig. 2. Dichromatobolus elephantulus gen. et sp. nov., ♂ (ZFMK MYR9896), SEM. A. Right antenna. B. Disc, apical view. C. Border of antennomere 5 to 6 with sensilla basiconica. D. Gnathochilarium, internal surface. E. Left mandible, mesal view. Abbreviations: 4iT = four-combed inner tooth; ac = apical cone; cP = central pads; d = disc; Endo = endochilarium; eP = lateral palpus; eT = external tooth; iP = inner palpus; LT = lateral tooth; mp = molar plate; pL = pectinate lamellae; sb = sensilla bascionica; numbers refer to antennomere number." figureDoi="http://doi.org/10.5281/zenodo.4085207" httpUri="https://zenodo.org/record/4085207/files/figure.png" pageId="3" pageNumber="110">Fig. 2A</figureCitation>
|
||
). Terminal antennomere with four large sensory cones located together inside a membranous area (
|
||
<figureCitation id="13232A22B717FFA0222631EBFAFC26D1" box="[1286,1385,1960,1986]" captionStart="Fig" captionStartId="5.[189,232,1822,1848]" captionTargetBox="[285,1303,265,1782]" captionTargetId="figure-11@5.[285,1303,265,1782]" captionTargetPageId="5" captionText="Fig. 2. Dichromatobolus elephantulus gen. et sp. nov., ♂ (ZFMK MYR9896), SEM. A. Right antenna. B. Disc, apical view. C. Border of antennomere 5 to 6 with sensilla basiconica. D. Gnathochilarium, internal surface. E. Left mandible, mesal view. Abbreviations: 4iT = four-combed inner tooth; ac = apical cone; cP = central pads; d = disc; Endo = endochilarium; eP = lateral palpus; eT = external tooth; iP = inner palpus; LT = lateral tooth; mp = molar plate; pL = pectinate lamellae; sb = sensilla bascionica; numbers refer to antennomere number." figureDoi="http://doi.org/10.5281/zenodo.4085207" httpUri="https://zenodo.org/record/4085207/files/figure.png" pageId="3" pageNumber="110">Fig. 2B</figureCitation>
|
||
). Antennomere 5 and 6 latero-apically with sensilla basiconica (
|
||
<figureCitation id="13232A22B717FFA024B9318DFC6326FB" box="[921,1014,1998,2024]" captionStart="Fig" captionStartId="5.[189,232,1822,1848]" captionTargetBox="[285,1303,265,1782]" captionTargetId="figure-11@5.[285,1303,265,1782]" captionTargetPageId="5" captionText="Fig. 2. Dichromatobolus elephantulus gen. et sp. nov., ♂ (ZFMK MYR9896), SEM. A. Right antenna. B. Disc, apical view. C. Border of antennomere 5 to 6 with sensilla basiconica. D. Gnathochilarium, internal surface. E. Left mandible, mesal view. Abbreviations: 4iT = four-combed inner tooth; ac = apical cone; cP = central pads; d = disc; Endo = endochilarium; eP = lateral palpus; eT = external tooth; iP = inner palpus; LT = lateral tooth; mp = molar plate; pL = pectinate lamellae; sb = sensilla bascionica; numbers refer to antennomere number." figureDoi="http://doi.org/10.5281/zenodo.4085207" httpUri="https://zenodo.org/record/4085207/files/figure.png" pageId="3" pageNumber="110">Fig. 2C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF67662FB710FFA7279D30B9FB4326F4" ID-DOI="http://doi.org/10.5281/zenodo.4085205" ID-Zenodo-Dep="4085205" httpUri="https://zenodo.org/record/4085205/files/figure.png" pageId="4" pageNumber="111" startId="4.[189,232,1786,1812]" targetBox="[272,1315,265,1748]" targetPageId="4">
|
||
<paragraph id="8BA736A7B710FFA7279D30B9FB4326F4" blockId="4.[189,1399,1786,2024]" pageId="4" pageNumber="111">
|
||
<emphasis id="B96CEAB5B710FFA7279D30B9FE9C2607" bold="true" box="[189,265,1786,1812]" pageId="4" pageNumber="111">Fig. 1.</emphasis>
|
||
<taxonomicName id="4C184D24B710FFA7262E30B9FDE32607" authority="Wesener, 2020" authorityName="Wesener" authorityYear="2020" box="[270,630,1786,1812]" class="Diplopoda" family="Pachybolidae" genus="Dichromatobolus" kingdom="Animalia" order="Spirobolida" pageId="4" pageNumber="111" phylum="Arthropoda" rank="species" species="elephantulus" status="gen. et sp. nov.">
|
||
<emphasis id="B96CEAB5B710FFA7262E30B9FDE32607" box="[270,630,1786,1812]" italics="true" pageId="4" pageNumber="111">Dichromatobolus elephantulus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25F57CEB710FFA7255B30B8FCB32606" box="[635,806,1787,1813]" pageId="4" pageNumber="111" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="B96CEAB5B710FFA7240B30B9FCD52607" bold="true" box="[811,832,1786,1812]" pageId="4" pageNumber="111">A</emphasis>
|
||
. Living ♂ (red; FMNH-INS 55888) and ♀ (grey; FMNH-INS 55887) from Odenwald Exoten 2007 import, photo by P. Decker.
|
||
<emphasis id="B96CEAB5B710FFA72347315DFB0B2624" bold="true" box="[1127,1182,1821,1848]" pageId="4" pageNumber="111">B–C</emphasis>
|
||
. Living immature ♂, 2019 import (ZFMK MYR8708), photo by L. Moritz.
|
||
<emphasis id="B96CEAB5B710FFA724583102FC192648" bold="true" box="[888,908,1857,1883]" pageId="4" pageNumber="111">B</emphasis>
|
||
. Head.
|
||
<emphasis id="B96CEAB5B710FFA724CC3103FB942649" bold="true" box="[1004,1025,1856,1882]" pageId="4" pageNumber="111">C</emphasis>
|
||
. Whole body.
|
||
<emphasis id="B96CEAB5B710FFA723943102FB7F2648" bold="true" box="[1204,1258,1857,1883]" pageId="4" pageNumber="111">D–F</emphasis>
|
||
. ♂ (ZFMK MYR8745), multi-layer photographs.
|
||
<emphasis id="B96CEAB5B710FFA725AC3127FD34266D" bold="true" box="[652,673,1892,1918]" pageId="4" pageNumber="111">D</emphasis>
|
||
. Anterior body, lateral view;
|
||
<emphasis id="B96CEAB5B710FFA723233127FB82266D" bold="true" box="[1027,1047,1892,1918]" pageId="4" pageNumber="111">E</emphasis>
|
||
. Leg pairs 1–7, ventral view.
|
||
<emphasis id="B96CEAB5B710FFA7279D31C4FF5A26B2" bold="true" box="[189,207,1927,1953]" pageId="4" pageNumber="111">F</emphasis>
|
||
. Posterior body end with telson. Abbreviations: av = paraprocts; C = lips of paraprocts; Ca = cardines; Co = collum (ring 1); Gn = gnathochilarium; Go = gonopods; IL = incisura lateralis; ms = marginal setae; P-M = parasitic(?) mites; Pre = epiproct; St = stipes; sub = hypoproct. Not to scale.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF67662FB711FFA6279D315DFD1026FB" ID-DOI="http://doi.org/10.5281/zenodo.4085207" ID-Zenodo-Dep="4085207" httpUri="https://zenodo.org/record/4085207/files/figure.png" pageId="5" pageNumber="112" startId="5.[189,232,1822,1848]" targetBox="[285,1303,265,1782]" targetPageId="5">
|
||
<paragraph id="8BA736A7B711FFA6279D315DFD1026FB" blockId="5.[189,1399,1822,2025]" pageId="5" pageNumber="112">
|
||
<emphasis id="B96CEAB5B711FFA6279D315DFE9E262B" bold="true" box="[189,267,1822,1848]" pageId="5" pageNumber="112">Fig. 2.</emphasis>
|
||
<taxonomicName id="4C184D24B711FFA62633315DFDE8262B" authority="Wesener, 2020" authorityName="Wesener" authorityYear="2020" box="[275,637,1822,1848]" class="Diplopoda" family="Pachybolidae" genus="Dichromatobolus" kingdom="Animalia" order="Spirobolida" pageId="5" pageNumber="112" phylum="Arthropoda" rank="species" species="elephantulus" status="gen. et sp. nov.">
|
||
<emphasis id="B96CEAB5B711FFA62633315DFDE8262B" box="[275,637,1822,1848]" italics="true" pageId="5" pageNumber="112">Dichromatobolus elephantulus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25F57CEB711FFA625A4315CFCAD262A" box="[644,824,1823,1849]" pageId="5" pageNumber="112" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
, ♂ (ZFMK MYR9896), SEM.
|
||
<emphasis id="B96CEAB5B711FFA62387315DFB29262B" bold="true" box="[1191,1212,1822,1848]" pageId="5" pageNumber="112">A</emphasis>
|
||
. Right antenna.
|
||
<emphasis id="B96CEAB5B711FFA6279D3102FF442648" bold="true" box="[189,209,1857,1883]" pageId="5" pageNumber="112">B</emphasis>
|
||
. Disc, apical view.
|
||
<emphasis id="B96CEAB5B711FFA626E03102FE402648" bold="true" box="[448,469,1857,1883]" pageId="5" pageNumber="112">C</emphasis>
|
||
. Border of antennomere 5 to 6 with sensilla basiconica.
|
||
<emphasis id="B96CEAB5B711FFA623A23102FB022648" bold="true" box="[1154,1175,1857,1883]" pageId="5" pageNumber="112">D</emphasis>
|
||
. Gnathochilarium, internal surface.
|
||
<emphasis id="B96CEAB5B711FFA626A83126FE09266C" bold="true" box="[392,412,1893,1919]" pageId="5" pageNumber="112">E</emphasis>
|
||
. Left mandible, mesal view. Abbreviations: 4iT = four-combed inner tooth; ac = apical cone; cP = central pads; d = disc; Endo = endochilarium; eP = lateral palpus; eT = external tooth; iP = inner palpus; LT = lateral tooth; mp = molar plate; pL = pectinate lamellae; sb = sensilla bascionica; numbers refer to antennomere number.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA736A7B712FFA5279D374AFB122061" blockId="6.[189,1399,265,370]" pageId="6" pageNumber="113">
|
||
GNATHOCHILARIUM. Lamellae linguales each with two standard setae located one behind the other. Stipites each with three apical setae. Basal part of mentum not divided, but with several ridges (
|
||
<figureCitation id="13232A22B712FFA5222C3773FAFF2058" box="[1292,1386,304,331]" captionStart="Fig" captionStartId="7.[189,232,1784,1810]" captionTargetBox="[253,1335,268,1747]" captionTargetId="figure-11@7.[253,1335,265,1747]" captionTargetPageId="7" captionText="Fig. 4. Dichromatobolus elephantulus gen. et sp. nov., holotype, ♂ (CAS MG-54C-78), drawings. A. Gnathochilarium, ventral view. B. Second leg pair, posterior view. C. First leg pair, posterior view. D. Left midbody leg pair, posterior view. E. Anterior gonopod, anterior view. F. Anterior gonopod, posterior view. Abbreviations: cp = coxite process; cP = central pads; Cx = coxite; eP = lateral palpus; Fe = femur; iP = inner palpus; LL = lamellae lingualis; Me = mentum; Po = postfemur; Pre = prefemur; St = sternite; St-Cx = sterno-coxite; T = telopodite; Ta = tarsus; Ti = tibia; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4085212" httpUri="https://zenodo.org/record/4085212/files/figure.png" pageId="6" pageNumber="113">Fig. 4A</figureCitation>
|
||
). Endochilarium and central pads shaped like in other
|
||
<taxonomicName id="4C184D24B712FFA52407371BFC252061" authorityName="Cook" authorityYear="1895" box="[807,944,344,370]" class="Diplopoda" kingdom="Animalia" order="Spirobolida" pageId="6" pageNumber="113" phylum="Arthropoda" rank="order">Spirobolida</taxonomicName>
|
||
species (
|
||
<figureCitation id="13232A22B712FFA5233B371BFBEF2061" box="[1051,1146,344,370]" captionStart="Fig" captionStartId="5.[189,232,1822,1848]" captionTargetBox="[285,1303,265,1782]" captionTargetId="figure-11@5.[285,1303,265,1782]" captionTargetPageId="5" captionText="Fig. 2. Dichromatobolus elephantulus gen. et sp. nov., ♂ (ZFMK MYR9896), SEM. A. Right antenna. B. Disc, apical view. C. Border of antennomere 5 to 6 with sensilla basiconica. D. Gnathochilarium, internal surface. E. Left mandible, mesal view. Abbreviations: 4iT = four-combed inner tooth; ac = apical cone; cP = central pads; d = disc; Endo = endochilarium; eP = lateral palpus; eT = external tooth; iP = inner palpus; LT = lateral tooth; mp = molar plate; pL = pectinate lamellae; sb = sensilla bascionica; numbers refer to antennomere number." figureDoi="http://doi.org/10.5281/zenodo.4085207" httpUri="https://zenodo.org/record/4085207/files/figure.png" pageId="6" pageNumber="113">Fig. 2D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA736A7B712FFA5279D37DDFCA2233C" blockId="6.[189,1399,414,559]" pageId="6" pageNumber="113">
|
||
MANDIBLE. In male, stipes with projecting sharp edge (
|
||
<figureCitation id="13232A22B712FFA5247237DDFC2720AB" box="[850,946,414,440]" captionStart="Fig" captionStartId="4.[189,232,1786,1812]" captionTargetBox="[272,1315,265,1748]" captionTargetId="figure-13@4.[272,1315,265,1748]" captionTargetPageId="4" captionText="Fig. 1. Dichromatobolus elephantulus gen. et sp. nov. A. Living ♂ (red; FMNH-INS 55888) and ♀ (grey; FMNH-INS 55887) from Odenwald Exoten 2007 import, photo by P. Decker. B–C. Living immature ♂, 2019 import (ZFMK MYR8708), photo by L. Moritz. B. Head. C. Whole body. D–F. ♂ (ZFMK MYR8745), multi-layer photographs. D. Anterior body, lateral view; E. Leg pairs 1–7, ventral view. F. Posterior body end with telson. Abbreviations: av = paraprocts; C = lips of paraprocts; Ca = cardines; Co = collum (ring 1); Gn = gnathochilarium; Go = gonopods; IL = incisura lateralis; ms = marginal setae; P-M = parasitic(?) mites; Pre = epiproct; St = stipes; sub = hypoproct. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4085205" httpUri="https://zenodo.org/record/4085205/files/figure.png" pageId="6" pageNumber="113">Fig. 1D</figureCitation>
|
||
). Gnathal lobe, external tooth simple, rounded; inner tooth with four cusps (
|
||
<figureCitation id="13232A22B712FFA525A03786FD4820F3" box="[640,733,453,480]" captionStart="Fig" captionStartId="5.[189,232,1822,1848]" captionTargetBox="[285,1303,265,1782]" captionTargetId="figure-11@5.[285,1303,265,1782]" captionTargetPageId="5" captionText="Fig. 2. Dichromatobolus elephantulus gen. et sp. nov., ♂ (ZFMK MYR9896), SEM. A. Right antenna. B. Disc, apical view. C. Border of antennomere 5 to 6 with sensilla basiconica. D. Gnathochilarium, internal surface. E. Left mandible, mesal view. Abbreviations: 4iT = four-combed inner tooth; ac = apical cone; cP = central pads; d = disc; Endo = endochilarium; eP = lateral palpus; eT = external tooth; iP = inner palpus; LT = lateral tooth; mp = molar plate; pL = pectinate lamellae; sb = sensilla bascionica; numbers refer to antennomere number." figureDoi="http://doi.org/10.5281/zenodo.4085207" httpUri="https://zenodo.org/record/4085207/files/figure.png" pageId="6" pageNumber="113">Fig. 2E</figureCitation>
|
||
). Six pectinate lamellae. Mesal margin of pectinate area (intermediate area) with circa four rows of small, slender spines. Molar plate with numerous (8 or 9) transverse furrows (
|
||
<figureCitation id="13232A22B712FFA526873456FD97233C" box="[423,514,533,559]" captionStart="Fig" captionStartId="5.[189,232,1822,1848]" captionTargetBox="[285,1303,265,1782]" captionTargetId="figure-11@5.[285,1303,265,1782]" captionTargetPageId="5" captionText="Fig. 2. Dichromatobolus elephantulus gen. et sp. nov., ♂ (ZFMK MYR9896), SEM. A. Right antenna. B. Disc, apical view. C. Border of antennomere 5 to 6 with sensilla basiconica. D. Gnathochilarium, internal surface. E. Left mandible, mesal view. Abbreviations: 4iT = four-combed inner tooth; ac = apical cone; cP = central pads; d = disc; Endo = endochilarium; eP = lateral palpus; eT = external tooth; iP = inner palpus; LT = lateral tooth; mp = molar plate; pL = pectinate lamellae; sb = sensilla bascionica; numbers refer to antennomere number." figureDoi="http://doi.org/10.5281/zenodo.4085207" httpUri="https://zenodo.org/record/4085207/files/figure.png" pageId="6" pageNumber="113">Fig. 2E</figureCitation>
|
||
). Female not investigated.
|
||
</paragraph>
|
||
<paragraph id="8BA736A7B712FFA5279D3418FC6E2366" blockId="6.[189,1019,602,629]" box="[189,1019,602,629]" pageId="6" pageNumber="113">
|
||
COLLUM. Smooth, laterally not protruding as far as ring 2 (
|
||
<figureCitation id="13232A22B712FFA5244D3419FC7B2366" box="[877,1006,602,629]" captionStart="Fig" captionStartId="4.[189,232,1786,1812]" captionTargetBox="[272,1315,265,1748]" captionTargetId="figure-13@4.[272,1315,265,1748]" captionTargetPageId="4" captionText="Fig. 1. Dichromatobolus elephantulus gen. et sp. nov. A. Living ♂ (red; FMNH-INS 55888) and ♀ (grey; FMNH-INS 55887) from Odenwald Exoten 2007 import, photo by P. Decker. B–C. Living immature ♂, 2019 import (ZFMK MYR8708), photo by L. Moritz. B. Head. C. Whole body. D–F. ♂ (ZFMK MYR8745), multi-layer photographs. D. Anterior body, lateral view; E. Leg pairs 1–7, ventral view. F. Posterior body end with telson. Abbreviations: av = paraprocts; C = lips of paraprocts; Ca = cardines; Co = collum (ring 1); Gn = gnathochilarium; Go = gonopods; IL = incisura lateralis; ms = marginal setae; P-M = parasitic(?) mites; Pre = epiproct; St = stipes; sub = hypoproct. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4085205" httpUri="https://zenodo.org/record/4085205/files/figure.png" pageId="6" pageNumber="113">Fig. 1B, D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA736A7B712FFA5279D34E3FAD5221A" blockId="6.[189,1399,672,778]" pageId="6" pageNumber="113">
|
||
BODY RINGS. Ozopores marked by black color, starting at ring 6, located a half diameter before suture between mesozona and metazona. Rings with smooth, but irregular coriaceous surface, ventrally on metazona with transverse ridges. Limbus simple, with cuticular scales and wavy margin (
|
||
<figureCitation id="13232A22B712FFA523F534ACFAA6221A" box="[1237,1331,751,777]" captionStart="Fig" captionStartId="6.[189,232,1964,1990]" captionTargetBox="[189,1397,1311,1924]" captionTargetId="figure-314@6.[189,1398,1311,1925]" captionTargetPageId="6" captionText="Fig. 3. Dichromatobolus elephantulus gen. et sp. nov., ♂ (ZFMK MYR9896), SEM. A. Limbus of midbody ring. B. Phoretic(?) mite at body ring. Abbreviations: pm = tergite margin; sc = cuticular scales." figureDoi="http://doi.org/10.5281/zenodo.4085209" httpUri="https://zenodo.org/record/4085209/files/figure.png" pageId="6" pageNumber="113">Fig. 3A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA736A7B712FFA5279D3576FB262264" blockId="6.[189,1398,821,888]" pageId="6" pageNumber="113">
|
||
TELSON. Paraprocts with weak lips, little micropunctation towards edges (
|
||
<figureCitation id="13232A22B712FFA5236F3576FB38225C" box="[1103,1197,821,847]" captionStart="Fig" captionStartId="4.[189,232,1786,1812]" captionTargetBox="[272,1315,265,1748]" captionTargetId="figure-13@4.[272,1315,265,1748]" captionTargetPageId="4" captionText="Fig. 1. Dichromatobolus elephantulus gen. et sp. nov. A. Living ♂ (red; FMNH-INS 55888) and ♀ (grey; FMNH-INS 55887) from Odenwald Exoten 2007 import, photo by P. Decker. B–C. Living immature ♂, 2019 import (ZFMK MYR8708), photo by L. Moritz. B. Head. C. Whole body. D–F. ♂ (ZFMK MYR8745), multi-layer photographs. D. Anterior body, lateral view; E. Leg pairs 1–7, ventral view. F. Posterior body end with telson. Abbreviations: av = paraprocts; C = lips of paraprocts; Ca = cardines; Co = collum (ring 1); Gn = gnathochilarium; Go = gonopods; IL = incisura lateralis; ms = marginal setae; P-M = parasitic(?) mites; Pre = epiproct; St = stipes; sub = hypoproct. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4085205" httpUri="https://zenodo.org/record/4085205/files/figure.png" pageId="6" pageNumber="113">Fig. 1F</figureCitation>
|
||
). Epiproct wellrounded, not protruding above paraproct (
|
||
<figureCitation id="13232A22B712FFA52589351EFD6F226B" box="[681,762,861,888]" captionStart="Fig" captionStartId="4.[189,232,1786,1812]" captionTargetBox="[272,1315,265,1748]" captionTargetId="figure-13@4.[272,1315,265,1748]" captionTargetPageId="4" captionText="Fig. 1. Dichromatobolus elephantulus gen. et sp. nov. A. Living ♂ (red; FMNH-INS 55888) and ♀ (grey; FMNH-INS 55887) from Odenwald Exoten 2007 import, photo by P. Decker. B–C. Living immature ♂, 2019 import (ZFMK MYR8708), photo by L. Moritz. B. Head. C. Whole body. D–F. ♂ (ZFMK MYR8745), multi-layer photographs. D. Anterior body, lateral view; E. Leg pairs 1–7, ventral view. F. Posterior body end with telson. Abbreviations: av = paraprocts; C = lips of paraprocts; Ca = cardines; Co = collum (ring 1); Gn = gnathochilarium; Go = gonopods; IL = incisura lateralis; ms = marginal setae; P-M = parasitic(?) mites; Pre = epiproct; St = stipes; sub = hypoproct. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4085205" httpUri="https://zenodo.org/record/4085205/files/figure.png" pageId="6" pageNumber="113">Fig.1F</figureCitation>
|
||
). Hypoproct inconspicuous (
|
||
<figureCitation id="13232A22B712FFA5236C351EFB312264" box="[1100,1188,861,887]" captionStart="Fig" captionStartId="4.[189,232,1786,1812]" captionTargetBox="[272,1315,265,1748]" captionTargetId="figure-13@4.[272,1315,265,1748]" captionTargetPageId="4" captionText="Fig. 1. Dichromatobolus elephantulus gen. et sp. nov. A. Living ♂ (red; FMNH-INS 55888) and ♀ (grey; FMNH-INS 55887) from Odenwald Exoten 2007 import, photo by P. Decker. B–C. Living immature ♂, 2019 import (ZFMK MYR8708), photo by L. Moritz. B. Head. C. Whole body. D–F. ♂ (ZFMK MYR8745), multi-layer photographs. D. Anterior body, lateral view; E. Leg pairs 1–7, ventral view. F. Posterior body end with telson. Abbreviations: av = paraprocts; C = lips of paraprocts; Ca = cardines; Co = collum (ring 1); Gn = gnathochilarium; Go = gonopods; IL = incisura lateralis; ms = marginal setae; P-M = parasitic(?) mites; Pre = epiproct; St = stipes; sub = hypoproct. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4085205" httpUri="https://zenodo.org/record/4085205/files/figure.png" pageId="6" pageNumber="113">Fig. 1F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA736A7B712FFA5279D35E0FC542590" blockId="6.[189,1399,930,1155]" pageId="6" pageNumber="113">
|
||
LEGS. Leg 1 with a large cylindrical coxa, twice as long as other podomeres (
|
||
<figureCitation id="13232A22B712FFA5231E35E1FB0E22AE" box="[1086,1179,930,957]" captionStart="Fig" captionStartId="7.[189,232,1784,1810]" captionTargetBox="[253,1335,268,1747]" captionTargetId="figure-11@7.[253,1335,265,1747]" captionTargetPageId="7" captionText="Fig. 4. Dichromatobolus elephantulus gen. et sp. nov., holotype, ♂ (CAS MG-54C-78), drawings. A. Gnathochilarium, ventral view. B. Second leg pair, posterior view. C. First leg pair, posterior view. D. Left midbody leg pair, posterior view. E. Anterior gonopod, anterior view. F. Anterior gonopod, posterior view. Abbreviations: cp = coxite process; cP = central pads; Cx = coxite; eP = lateral palpus; Fe = femur; iP = inner palpus; LL = lamellae lingualis; Me = mentum; Po = postfemur; Pre = prefemur; St = sternite; St-Cx = sterno-coxite; T = telopodite; Ta = tarsus; Ti = tibia; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4085212" httpUri="https://zenodo.org/record/4085212/files/figure.png" pageId="6" pageNumber="113">Fig. 4C</figureCitation>
|
||
). Tarsus with three pairs of ventral spines, without an apical spine beyond claw. Leg 2 with a strongly elongated coxa, fused to sternite. Tarsus with four pairs of ventral spines and a short apical spine (
|
||
<figureCitation id="13232A22B712FFA5230A35B1FB13251F" box="[1066,1158,1010,1036]" captionStart="Fig" captionStartId="7.[189,232,1784,1810]" captionTargetBox="[253,1335,268,1747]" captionTargetId="figure-11@7.[253,1335,265,1747]" captionTargetPageId="7" captionText="Fig. 4. Dichromatobolus elephantulus gen. et sp. nov., holotype, ♂ (CAS MG-54C-78), drawings. A. Gnathochilarium, ventral view. B. Second leg pair, posterior view. C. First leg pair, posterior view. D. Left midbody leg pair, posterior view. E. Anterior gonopod, anterior view. F. Anterior gonopod, posterior view. Abbreviations: cp = coxite process; cP = central pads; Cx = coxite; eP = lateral palpus; Fe = femur; iP = inner palpus; LL = lamellae lingualis; Me = mentum; Po = postfemur; Pre = prefemur; St = sternite; St-Cx = sterno-coxite; T = telopodite; Ta = tarsus; Ti = tibia; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4085212" httpUri="https://zenodo.org/record/4085212/files/figure.png" pageId="6" pageNumber="113">Fig. 4B</figureCitation>
|
||
). Midbody legs with a rectangular coxa, as long as other podomeres. Each podomere ventrally with a single or a pair of apical setae, tarsus in males with a ventral pad, 6 pairs of ventral and 5 pairs of lateral spines (
|
||
<figureCitation id="13232A22B712FFA523923202FA852548" box="[1202,1296,1089,1115]" captionStart="Fig" captionStartId="7.[189,232,1784,1810]" captionTargetBox="[253,1335,268,1747]" captionTargetId="figure-11@7.[253,1335,265,1747]" captionTargetPageId="7" captionText="Fig. 4. Dichromatobolus elephantulus gen. et sp. nov., holotype, ♂ (CAS MG-54C-78), drawings. A. Gnathochilarium, ventral view. B. Second leg pair, posterior view. C. First leg pair, posterior view. D. Left midbody leg pair, posterior view. E. Anterior gonopod, anterior view. F. Anterior gonopod, posterior view. Abbreviations: cp = coxite process; cP = central pads; Cx = coxite; eP = lateral palpus; Fe = femur; iP = inner palpus; LL = lamellae lingualis; Me = mentum; Po = postfemur; Pre = prefemur; St = sternite; St-Cx = sterno-coxite; T = telopodite; Ta = tarsus; Ti = tibia; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4085212" httpUri="https://zenodo.org/record/4085212/files/figure.png" pageId="6" pageNumber="113">Fig. 4D</figureCitation>
|
||
). Length of midbody legs circa 0.7 times body diameter in males (
|
||
<figureCitation id="13232A22B712FFA52479322BFC262590" box="[857,947,1128,1155]" captionStart="Fig" captionStartId="4.[189,232,1786,1812]" captionTargetBox="[272,1315,265,1748]" captionTargetId="figure-13@4.[272,1315,265,1748]" captionTargetPageId="4" captionText="Fig. 1. Dichromatobolus elephantulus gen. et sp. nov. A. Living ♂ (red; FMNH-INS 55888) and ♀ (grey; FMNH-INS 55887) from Odenwald Exoten 2007 import, photo by P. Decker. B–C. Living immature ♂, 2019 import (ZFMK MYR8708), photo by L. Moritz. B. Head. C. Whole body. D–F. ♂ (ZFMK MYR8745), multi-layer photographs. D. Anterior body, lateral view; E. Leg pairs 1–7, ventral view. F. Posterior body end with telson. Abbreviations: av = paraprocts; C = lips of paraprocts; Ca = cardines; Co = collum (ring 1); Gn = gnathochilarium; Go = gonopods; IL = incisura lateralis; ms = marginal setae; P-M = parasitic(?) mites; Pre = epiproct; St = stipes; sub = hypoproct. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4085205" httpUri="https://zenodo.org/record/4085205/files/figure.png" pageId="6" pageNumber="113">Fig. 1E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA736A7B712FFA527CA32EDFDB325DB" blockId="6.[189,1085,1198,1264]" box="[234,550,1198,1224]" pageId="6" pageNumber="113">
|
||
<emphasis id="B96CEAB5B712FFA527CA32EDFDB325DB" bold="true" box="[234,550,1198,1224]" pageId="6" pageNumber="113">Female sexual characters</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BA736A7B712FFA5279D3295FBA825E3" blockId="6.[189,1085,1198,1264]" box="[189,1085,1238,1264]" pageId="6" pageNumber="113">
|
||
Body color grey (
|
||
<figureCitation id="13232A22B712FFA526AC3295FE7C25E3" box="[396,489,1238,1264]" captionStart="Fig" captionStartId="4.[189,232,1786,1812]" captionTargetBox="[272,1315,265,1748]" captionTargetId="figure-13@4.[272,1315,265,1748]" captionTargetPageId="4" captionText="Fig. 1. Dichromatobolus elephantulus gen. et sp. nov. A. Living ♂ (red; FMNH-INS 55888) and ♀ (grey; FMNH-INS 55887) from Odenwald Exoten 2007 import, photo by P. Decker. B–C. Living immature ♂, 2019 import (ZFMK MYR8708), photo by L. Moritz. B. Head. C. Whole body. D–F. ♂ (ZFMK MYR8745), multi-layer photographs. D. Anterior body, lateral view; E. Leg pairs 1–7, ventral view. F. Posterior body end with telson. Abbreviations: av = paraprocts; C = lips of paraprocts; Ca = cardines; Co = collum (ring 1); Gn = gnathochilarium; Go = gonopods; IL = incisura lateralis; ms = marginal setae; P-M = parasitic(?) mites; Pre = epiproct; St = stipes; sub = hypoproct. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4085205" httpUri="https://zenodo.org/record/4085205/files/figure.png" pageId="6" pageNumber="113">Fig. 1A</figureCitation>
|
||
). Female vulva simple, resembling a bivalve shell.
|
||
</paragraph>
|
||
<caption id="DF67662FB712FFA5279D31EFFAE226FA" ID-DOI="http://doi.org/10.5281/zenodo.4085209" ID-Zenodo-Dep="4085209" httpUri="https://zenodo.org/record/4085209/files/figure.png" pageId="6" pageNumber="113" startId="6.[189,232,1964,1990]" targetBox="[189,1397,1311,1924]" targetPageId="6">
|
||
<paragraph id="8BA736A7B712FFA5279D31EFFAE226FA" blockId="6.[189,1399,1964,2026]" pageId="6" pageNumber="113">
|
||
<emphasis id="B96CEAB5B712FFA5279D31EFFE9B26D5" bold="true" box="[189,270,1964,1990]" pageId="6" pageNumber="113">Fig. 3.</emphasis>
|
||
<taxonomicName id="4C184D24B712FFA5263A31EFFD1226D5" authority="Wesener, 2020" authorityName="Wesener" authorityYear="2020" box="[282,647,1964,1990]" class="Diplopoda" family="Pachybolidae" genus="Dichromatobolus" kingdom="Animalia" order="Spirobolida" pageId="6" pageNumber="113" phylum="Arthropoda" rank="species" species="elephantulus" status="gen. et sp. nov.">
|
||
<emphasis id="B96CEAB5B712FFA5263A31EFFD1226D5" box="[282,647,1964,1990]" italics="true" pageId="6" pageNumber="113">Dichromatobolus elephantulus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25F57CEB712FFA525B231EFFCC426D5" box="[658,849,1964,1990]" pageId="6" pageNumber="113" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
, ♂ (ZFMK MYR9896), SEM.
|
||
<emphasis id="B96CEAB5B712FFA523F131EFFB7326D5" bold="true" box="[1233,1254,1964,1990]" pageId="6" pageNumber="113">A</emphasis>
|
||
. Limbus of midbody ring.
|
||
<emphasis id="B96CEAB5B712FFA52648318CFEE926FA" bold="true" box="[360,380,1999,2025]" pageId="6" pageNumber="113">B</emphasis>
|
||
. Phoretic(?) mite at body ring. Abbreviations: pm = tergite margin; sc = cuticular scales.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF67662FB713FFA4279D30BBFECD26F5" ID-DOI="http://doi.org/10.5281/zenodo.4085212" ID-Zenodo-Dep="4085212" httpUri="https://zenodo.org/record/4085212/files/figure.png" pageId="7" pageNumber="114" startId="7.[189,232,1784,1810]" targetBox="[253,1335,268,1747]" targetPageId="7">
|
||
<paragraph id="8BA736A7B713FFA4279D30BBFECD26F5" blockId="7.[189,1399,1784,2022]" pageId="7" pageNumber="114">
|
||
<emphasis id="B96CEAB5B713FFA4279D30BBFE9A2601" bold="true" box="[189,271,1784,1810]" pageId="7" pageNumber="114">Fig. 4.</emphasis>
|
||
<taxonomicName id="4C184D24B713FFA4263C30BBFD1F2601" authority="Wesener, 2020" authorityName="Wesener" authorityYear="2020" box="[284,650,1784,1810]" class="Diplopoda" family="Pachybolidae" genus="Dichromatobolus" kingdom="Animalia" order="Spirobolida" pageId="7" pageNumber="114" phylum="Arthropoda" rank="species" species="elephantulus" status="gen. et sp. nov.">
|
||
<emphasis id="B96CEAB5B713FFA4263C30BBFD1F2601" box="[284,650,1784,1810]" italics="true" pageId="7" pageNumber="114">Dichromatobolus elephantulus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25F57CEB713FFA425B730BAFCCD2600" box="[663,856,1785,1811]" pageId="7" pageNumber="114" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
, holotype, ♂ (CAS MG-54C-78), drawings.
|
||
<emphasis id="B96CEAB5B713FFA4279D3158FF472626" bold="true" box="[189,210,1819,1845]" pageId="7" pageNumber="114">A</emphasis>
|
||
. Gnathochilarium, ventral view.
|
||
<emphasis id="B96CEAB5B713FFA42579315FFDF82625" bold="true" box="[601,621,1820,1846]" pageId="7" pageNumber="114">B</emphasis>
|
||
. Second leg pair, posterior view.
|
||
<emphasis id="B96CEAB5B713FFA424DC3158FB842626" bold="true" box="[1020,1041,1819,1845]" pageId="7" pageNumber="114">C</emphasis>
|
||
. First leg pair, posterior view.
|
||
<emphasis id="B96CEAB5B713FFA4279D317CFF47264A" bold="true" box="[189,210,1855,1881]" pageId="7" pageNumber="114">D</emphasis>
|
||
. Left midbody leg pair, posterior view.
|
||
<emphasis id="B96CEAB5B713FFA4259D317CFD44264A" bold="true" box="[701,721,1855,1881]" pageId="7" pageNumber="114">E</emphasis>
|
||
. Anterior gonopod, anterior view.
|
||
<emphasis id="B96CEAB5B713FFA42358317CFB1F264A" bold="true" box="[1144,1162,1855,1881]" pageId="7" pageNumber="114">F</emphasis>
|
||
. Anterior gonopod, posterior view. Abbreviations: cp = coxite process; cP = central pads; Cx = coxite; eP = lateral palpus; Fe = femur; iP = inner palpus; LL = lamellae lingualis; Me = mentum; Po = postfemur; Pre = prefemur; St = sternite; St-Cx = sterno-coxite; T = telopodite; Ta = tarsus; Ti = tibia; Tp = telopodite process. Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA736A7B71CFFAB27CA374AFD9E2030" blockId="8.[189,1079,265,330]" box="[234,523,265,291]" pageId="8" pageNumber="115">
|
||
<emphasis id="B96CEAB5B71CFFAB27CA374AFD9E2030" bold="true" box="[234,523,265,291]" pageId="8" pageNumber="115">Male sexual characters</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BA736A7B71CFFAB279D376CFBA2205A" blockId="8.[189,1079,265,330]" box="[189,1079,303,330]" pageId="8" pageNumber="115">
|
||
Tarsal pads present (
|
||
<figureCitation id="13232A22B71CFFAB268D376CFD98205A" box="[429,525,303,329]" captionStart="Fig" captionStartId="7.[189,232,1784,1810]" captionTargetBox="[253,1335,268,1747]" captionTargetId="figure-11@7.[253,1335,265,1747]" captionTargetPageId="7" captionText="Fig. 4. Dichromatobolus elephantulus gen. et sp. nov., holotype, ♂ (CAS MG-54C-78), drawings. A. Gnathochilarium, ventral view. B. Second leg pair, posterior view. C. First leg pair, posterior view. D. Left midbody leg pair, posterior view. E. Anterior gonopod, anterior view. F. Anterior gonopod, posterior view. Abbreviations: cp = coxite process; cP = central pads; Cx = coxite; eP = lateral palpus; Fe = femur; iP = inner palpus; LL = lamellae lingualis; Me = mentum; Po = postfemur; Pre = prefemur; St = sternite; St-Cx = sterno-coxite; T = telopodite; Ta = tarsus; Ti = tibia; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4085212" httpUri="https://zenodo.org/record/4085212/files/figure.png" pageId="8" pageNumber="115">Fig. 4D</figureCitation>
|
||
). Coxae 3–7 without coxal processes (
|
||
<figureCitation id="13232A22B71CFFAB24EE376CFBBC205A" box="[974,1065,303,329]" captionStart="Fig" captionStartId="4.[189,232,1786,1812]" captionTargetBox="[272,1315,265,1748]" captionTargetId="figure-13@4.[272,1315,265,1748]" captionTargetPageId="4" captionText="Fig. 1. Dichromatobolus elephantulus gen. et sp. nov. A. Living ♂ (red; FMNH-INS 55888) and ♀ (grey; FMNH-INS 55887) from Odenwald Exoten 2007 import, photo by P. Decker. B–C. Living immature ♂, 2019 import (ZFMK MYR8708), photo by L. Moritz. B. Head. C. Whole body. D–F. ♂ (ZFMK MYR8745), multi-layer photographs. D. Anterior body, lateral view; E. Leg pairs 1–7, ventral view. F. Posterior body end with telson. Abbreviations: av = paraprocts; C = lips of paraprocts; Ca = cardines; Co = collum (ring 1); Gn = gnathochilarium; Go = gonopods; IL = incisura lateralis; ms = marginal setae; P-M = parasitic(?) mites; Pre = epiproct; St = stipes; sub = hypoproct. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4085205" httpUri="https://zenodo.org/record/4085205/files/figure.png" pageId="8" pageNumber="115">Fig. 1E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA736A7B71CFFAB279D3737FC4B233A" blockId="8.[189,1399,372,553]" pageId="8" pageNumber="115">
|
||
Anterior gonopod sternite massive (
|
||
<figureCitation id="13232A22B71CFFAB254E3737FD5E209D" box="[622,715,372,398]" captionStart="Fig" captionStartId="7.[189,232,1784,1810]" captionTargetBox="[253,1335,268,1747]" captionTargetId="figure-11@7.[253,1335,265,1747]" captionTargetPageId="7" captionText="Fig. 4. Dichromatobolus elephantulus gen. et sp. nov., holotype, ♂ (CAS MG-54C-78), drawings. A. Gnathochilarium, ventral view. B. Second leg pair, posterior view. C. First leg pair, posterior view. D. Left midbody leg pair, posterior view. E. Anterior gonopod, anterior view. F. Anterior gonopod, posterior view. Abbreviations: cp = coxite process; cP = central pads; Cx = coxite; eP = lateral palpus; Fe = femur; iP = inner palpus; LL = lamellae lingualis; Me = mentum; Po = postfemur; Pre = prefemur; St = sternite; St-Cx = sterno-coxite; T = telopodite; Ta = tarsus; Ti = tibia; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4085212" httpUri="https://zenodo.org/record/4085212/files/figure.png" pageId="8" pageNumber="115">Fig. 4E</figureCitation>
|
||
), elongated into wide, well-rounded lobe (
|
||
<figureCitation id="13232A22B71CFFAB23EE3737FAB9209D" box="[1230,1324,372,398]" captionStart="Fig" captionStartId="7.[189,232,1784,1810]" captionTargetBox="[253,1335,268,1747]" captionTargetId="figure-11@7.[253,1335,265,1747]" captionTargetPageId="7" captionText="Fig. 4. Dichromatobolus elephantulus gen. et sp. nov., holotype, ♂ (CAS MG-54C-78), drawings. A. Gnathochilarium, ventral view. B. Second leg pair, posterior view. C. First leg pair, posterior view. D. Left midbody leg pair, posterior view. E. Anterior gonopod, anterior view. F. Anterior gonopod, posterior view. Abbreviations: cp = coxite process; cP = central pads; Cx = coxite; eP = lateral palpus; Fe = femur; iP = inner palpus; LL = lamellae lingualis; Me = mentum; Po = postfemur; Pre = prefemur; St = sternite; St-Cx = sterno-coxite; T = telopodite; Ta = tarsus; Ti = tibia; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4085212" httpUri="https://zenodo.org/record/4085212/files/figure.png" pageId="8" pageNumber="115">Fig. 4E</figureCitation>
|
||
), lobe lower than coxite. Sternite in posterior view well-visible, without discernable apodemes. Coxite mesally in anterior view with wide, well-rounded process, apical part of process recessed (
|
||
<figureCitation id="13232A22B71CFFAB234F3782FB7220CF" box="[1135,1255,449,476]" captionStart="Fig" captionStartId="7.[189,232,1784,1810]" captionTargetBox="[253,1335,268,1747]" captionTargetId="figure-11@7.[253,1335,265,1747]" captionTargetPageId="7" captionText="Fig. 4. Dichromatobolus elephantulus gen. et sp. nov., holotype, ♂ (CAS MG-54C-78), drawings. A. Gnathochilarium, ventral view. B. Second leg pair, posterior view. C. First leg pair, posterior view. D. Left midbody leg pair, posterior view. E. Anterior gonopod, anterior view. F. Anterior gonopod, posterior view. Abbreviations: cp = coxite process; cP = central pads; Cx = coxite; eP = lateral palpus; Fe = femur; iP = inner palpus; LL = lamellae lingualis; Me = mentum; Po = postfemur; Pre = prefemur; St = sternite; St-Cx = sterno-coxite; T = telopodite; Ta = tarsus; Ti = tibia; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4085212" httpUri="https://zenodo.org/record/4085212/files/figure.png" pageId="8" pageNumber="115">Fig. 4E, F</figureCitation>
|
||
). Telopodite with slender process arising mesally (
|
||
<figureCitation id="13232A22B71CFFAB25A637ABFD772311" box="[646,738,488,514]" captionStart="Fig" captionStartId="7.[189,232,1784,1810]" captionTargetBox="[253,1335,268,1747]" captionTargetId="figure-11@7.[253,1335,265,1747]" captionTargetPageId="7" captionText="Fig. 4. Dichromatobolus elephantulus gen. et sp. nov., holotype, ♂ (CAS MG-54C-78), drawings. A. Gnathochilarium, ventral view. B. Second leg pair, posterior view. C. First leg pair, posterior view. D. Left midbody leg pair, posterior view. E. Anterior gonopod, anterior view. F. Anterior gonopod, posterior view. Abbreviations: cp = coxite process; cP = central pads; Cx = coxite; eP = lateral palpus; Fe = femur; iP = inner palpus; LL = lamellae lingualis; Me = mentum; Po = postfemur; Pre = prefemur; St = sternite; St-Cx = sterno-coxite; T = telopodite; Ta = tarsus; Ti = tibia; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4085212" httpUri="https://zenodo.org/record/4085212/files/figure.png" pageId="8" pageNumber="115">Fig. 4F</figureCitation>
|
||
), process conspicuously curved laterad (
|
||
<figureCitation id="13232A22B71CFFAB23EC37ABFABF2311" box="[1228,1322,488,514]" captionStart="Fig" captionStartId="7.[189,232,1784,1810]" captionTargetBox="[253,1335,268,1747]" captionTargetId="figure-11@7.[253,1335,265,1747]" captionTargetPageId="7" captionText="Fig. 4. Dichromatobolus elephantulus gen. et sp. nov., holotype, ♂ (CAS MG-54C-78), drawings. A. Gnathochilarium, ventral view. B. Second leg pair, posterior view. C. First leg pair, posterior view. D. Left midbody leg pair, posterior view. E. Anterior gonopod, anterior view. F. Anterior gonopod, posterior view. Abbreviations: cp = coxite process; cP = central pads; Cx = coxite; eP = lateral palpus; Fe = femur; iP = inner palpus; LL = lamellae lingualis; Me = mentum; Po = postfemur; Pre = prefemur; St = sternite; St-Cx = sterno-coxite; T = telopodite; Ta = tarsus; Ti = tibia; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4085212" httpUri="https://zenodo.org/record/4085212/files/figure.png" pageId="8" pageNumber="115">Fig. 4E</figureCitation>
|
||
), with well-rounded tip, not reaching lateral margin of telopodite (
|
||
<figureCitation id="13232A22B71CFFAB2457344DFC44233A" box="[887,977,526,553]" captionStart="Fig" captionStartId="7.[189,232,1784,1810]" captionTargetBox="[253,1335,268,1747]" captionTargetId="figure-11@7.[253,1335,265,1747]" captionTargetPageId="7" captionText="Fig. 4. Dichromatobolus elephantulus gen. et sp. nov., holotype, ♂ (CAS MG-54C-78), drawings. A. Gnathochilarium, ventral view. B. Second leg pair, posterior view. C. First leg pair, posterior view. D. Left midbody leg pair, posterior view. E. Anterior gonopod, anterior view. F. Anterior gonopod, posterior view. Abbreviations: cp = coxite process; cP = central pads; Cx = coxite; eP = lateral palpus; Fe = femur; iP = inner palpus; LL = lamellae lingualis; Me = mentum; Po = postfemur; Pre = prefemur; St = sternite; St-Cx = sterno-coxite; T = telopodite; Ta = tarsus; Ti = tibia; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4085212" httpUri="https://zenodo.org/record/4085212/files/figure.png" pageId="8" pageNumber="115">Fig. 4F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA736A7B71CFFAB279D3410FB8B22D9" blockId="8.[189,1399,595,970]" pageId="8" pageNumber="115">
|
||
Posterior gonopods each consisting of two parts, separated by sutures (
|
||
<figureCitation id="13232A22B71CFFAB23323410FBCC237D" box="[1042,1113,595,622]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[189,1398,1021,1852]" captionTargetId="figure-239@8.[189,1398,1017,1852]" captionTargetPageId="8" captionText="Fig. 5. Dichromatobolus elephantulus gen. et sp. nov., holotype, ♂ (CAS MG-54C-78), left posterior gonopod, drawings. A. Anterior view. B. Posterior view. C. Detail of telopodite, posterior view. Abbreviations: Apo = apodeme; Cx = coxite; sg =efferent dUct; scl = small mesal sclerite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4085214" httpUri="https://zenodo.org/record/4085214/files/figure.png" pageId="8" pageNumber="115">Fig. 5</figureCitation>
|
||
A–C): (1) basally large, bulky coxite with slender, elongated process of coxite, and (2) apically almost rectangular telopodite with slight membranoUs folds extending meso-apicad and apicad, efferent dUct discharging apically (
|
||
<figureCitation id="13232A22B71CFFAB27E63484FE9C23F1" box="[198,265,711,738]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[189,1398,1021,1852]" captionTargetId="figure-239@8.[189,1398,1017,1852]" captionTargetPageId="8" captionText="Fig. 5. Dichromatobolus elephantulus gen. et sp. nov., holotype, ♂ (CAS MG-54C-78), left posterior gonopod, drawings. A. Anterior view. B. Posterior view. C. Detail of telopodite, posterior view. Abbreviations: Apo = apodeme; Cx = coxite; sg =efferent dUct; scl = small mesal sclerite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4085214" httpUri="https://zenodo.org/record/4085214/files/figure.png" pageId="8" pageNumber="115">Fig. 5</figureCitation>
|
||
A–C). Coxal process and telopodite placed in same axis (
|
||
<figureCitation id="13232A22B71CFFAB248A3484FC7B23F1" box="[938,1006,711,738]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[189,1398,1021,1852]" captionTargetId="figure-239@8.[189,1398,1017,1852]" captionTargetPageId="8" captionText="Fig. 5. Dichromatobolus elephantulus gen. et sp. nov., holotype, ♂ (CAS MG-54C-78), left posterior gonopod, drawings. A. Anterior view. B. Posterior view. C. Detail of telopodite, posterior view. Abbreviations: Apo = apodeme; Cx = coxite; sg =efferent dUct; scl = small mesal sclerite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4085214" httpUri="https://zenodo.org/record/4085214/files/figure.png" pageId="8" pageNumber="115">Fig. 5</figureCitation>
|
||
A–B). Pair of posterior gonopods located parallel to each other, connected by small, sclerotized and visible sternite. Basal part of coxite wide, mesally with small sclerite located on lower level than remaining part (
|
||
<figureCitation id="13232A22B71CFFAB231F3556FB0E223C" box="[1087,1179,788,815]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[189,1398,1021,1852]" captionTargetId="figure-239@8.[189,1398,1017,1852]" captionTargetPageId="8" captionText="Fig. 5. Dichromatobolus elephantulus gen. et sp. nov., holotype, ♂ (CAS MG-54C-78), left posterior gonopod, drawings. A. Anterior view. B. Posterior view. C. Detail of telopodite, posterior view. Abbreviations: Apo = apodeme; Cx = coxite; sg =efferent dUct; scl = small mesal sclerite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4085214" httpUri="https://zenodo.org/record/4085214/files/figure.png" pageId="8" pageNumber="115">Fig. 5B</figureCitation>
|
||
). Coxite elongated into long coxal process (
|
||
<figureCitation id="13232A22B71CFFAB26FC3578FDAF2245" box="[476,570,827,854]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[189,1398,1021,1852]" captionTargetId="figure-239@8.[189,1398,1017,1852]" captionTargetPageId="8" captionText="Fig. 5. Dichromatobolus elephantulus gen. et sp. nov., holotype, ♂ (CAS MG-54C-78), left posterior gonopod, drawings. A. Anterior view. B. Posterior view. C. Detail of telopodite, posterior view. Abbreviations: Apo = apodeme; Cx = coxite; sg =efferent dUct; scl = small mesal sclerite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4085214" httpUri="https://zenodo.org/record/4085214/files/figure.png" pageId="8" pageNumber="115">Fig. 5A</figureCitation>
|
||
). Efferent dUct rUnning at mesal margin of coxite (
|
||
<figureCitation id="13232A22B71CFFAB23A93578FB702245" box="[1161,1253,827,854]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[189,1398,1021,1852]" captionTargetId="figure-239@8.[189,1398,1017,1852]" captionTargetPageId="8" captionText="Fig. 5. Dichromatobolus elephantulus gen. et sp. nov., holotype, ♂ (CAS MG-54C-78), left posterior gonopod, drawings. A. Anterior view. B. Posterior view. C. Detail of telopodite, posterior view. Abbreviations: Apo = apodeme; Cx = coxite; sg =efferent dUct; scl = small mesal sclerite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4085214" httpUri="https://zenodo.org/record/4085214/files/figure.png" pageId="8" pageNumber="115">Fig. 5B</figureCitation>
|
||
). Telopodite straight, as wide as coxite process, discharge opening of efferent dUct apically. Telopodite shorter than coxite process (
|
||
<figureCitation id="13232A22B71CFFAB265735CBFE4322B0" box="[375,470,904,931]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[189,1398,1021,1852]" captionTargetId="figure-239@8.[189,1398,1017,1852]" captionTargetPageId="8" captionText="Fig. 5. Dichromatobolus elephantulus gen. et sp. nov., holotype, ♂ (CAS MG-54C-78), left posterior gonopod, drawings. A. Anterior view. B. Posterior view. C. Detail of telopodite, posterior view. Abbreviations: Apo = apodeme; Cx = coxite; sg =efferent dUct; scl = small mesal sclerite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4085214" httpUri="https://zenodo.org/record/4085214/files/figure.png" pageId="8" pageNumber="115">Fig. 5A</figureCitation>
|
||
), meso-apically with well-rounded, triangular, membranous process (
|
||
<figureCitation id="13232A22B71CFFAB222D35CBFAC622B0" box="[1293,1363,904,931]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[189,1398,1021,1852]" captionTargetId="figure-239@8.[189,1398,1017,1852]" captionTargetPageId="8" captionText="Fig. 5. Dichromatobolus elephantulus gen. et sp. nov., holotype, ♂ (CAS MG-54C-78), left posterior gonopod, drawings. A. Anterior view. B. Posterior view. C. Detail of telopodite, posterior view. Abbreviations: Apo = apodeme; Cx = coxite; sg =efferent dUct; scl = small mesal sclerite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4085214" httpUri="https://zenodo.org/record/4085214/files/figure.png" pageId="8" pageNumber="115">Fig. 5</figureCitation>
|
||
B– C), apically with slightly projecting opening of efferent dUct (
|
||
<figureCitation id="13232A22B71CFFAB24AF35ECFC4622D9" box="[911,979,943,970]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[189,1398,1021,1852]" captionTargetId="figure-239@8.[189,1398,1017,1852]" captionTargetPageId="8" captionText="Fig. 5. Dichromatobolus elephantulus gen. et sp. nov., holotype, ♂ (CAS MG-54C-78), left posterior gonopod, drawings. A. Anterior view. B. Posterior view. C. Detail of telopodite, posterior view. Abbreviations: Apo = apodeme; Cx = coxite; sg =efferent dUct; scl = small mesal sclerite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4085214" httpUri="https://zenodo.org/record/4085214/files/figure.png" pageId="8" pageNumber="115">Fig. 5</figureCitation>
|
||
A–C).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF67662FB71CFFAB279D3127FE0A26FB" ID-DOI="http://doi.org/10.5281/zenodo.4085214" ID-Zenodo-Dep="4085214" httpUri="https://zenodo.org/record/4085214/files/figure.png" pageId="8" pageNumber="115" startId="8.[189,232,1892,1918]" targetBox="[189,1398,1021,1852]" targetPageId="8">
|
||
<paragraph id="8BA736A7B71CFFAB279D3127FE0A26FB" blockId="8.[189,1399,1892,2024]" pageId="8" pageNumber="115">
|
||
<emphasis id="B96CEAB5B71CFFAB279D3127FE99266D" bold="true" box="[189,268,1892,1918]" pageId="8" pageNumber="115">Fig. 5.</emphasis>
|
||
<taxonomicName id="4C184D24B71CFFAB26353127FD14266D" authority="Wesener, 2020" authorityName="Wesener" authorityYear="2020" box="[277,641,1892,1918]" class="Diplopoda" family="Pachybolidae" genus="Dichromatobolus" kingdom="Animalia" order="Spirobolida" pageId="8" pageNumber="115" phylum="Arthropoda" rank="species" species="elephantulus" status="gen. et sp. nov.">
|
||
<emphasis id="B96CEAB5B71CFFAB26353127FD14266D" box="[277,641,1892,1918]" italics="true" pageId="8" pageNumber="115">Dichromatobolus elephantulus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25F57CEB71CFFAB25AA3126FCD7266C" box="[650,834,1893,1919]" pageId="8" pageNumber="115" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
, holotype, ♂ (CAS MG-54C-78), left posterior gonopod, drawings.
|
||
<emphasis id="B96CEAB5B71CFFAB269A31C4FE5A26B2" bold="true" box="[442,463,1927,1953]" pageId="8" pageNumber="115">A</emphasis>
|
||
. Anterior view.
|
||
<emphasis id="B96CEAB5B71CFFAB258231C4FD2326B2" bold="true" box="[674,694,1927,1953]" pageId="8" pageNumber="115">B</emphasis>
|
||
. Posterior view.
|
||
<emphasis id="B96CEAB5B71CFFAB24AF31C4FC3126B2" bold="true" box="[911,932,1927,1953]" pageId="8" pageNumber="115">C</emphasis>
|
||
. Detail of telopodite, posterior view. Abbreviations: Apo = apodeme; Cx = coxite; sg =efferent dUct; scl = small mesal sclerite; T = telopodite. Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF67662FB71DFFAA279D31E8FD9126FA" ID-DOI="http://doi.org/10.5281/zenodo.4085216" ID-Zenodo-Dep="4085216" httpUri="https://zenodo.org/record/4085216/files/figure.png" pageId="9" pageNumber="116" startId="9.[189,232,1963,1989]" targetBox="[204,1383,279,1926]" targetPageId="9">
|
||
<paragraph id="8BA736A7B71DFFAA279D31E8FD9126FA" blockId="9.[189,1398,1963,2025]" pageId="9" pageNumber="116">
|
||
<emphasis id="B96CEAB5B71DFFAA279D31E8FE9E26D6" bold="true" box="[189,267,1963,1989]" pageId="9" pageNumber="116">Fig. 6.</emphasis>
|
||
Distribution of
|
||
<taxonomicName id="4C184D24B71DFFAA26E931EFFCA126D5" authority="Wesener, 2020" authorityName="Wesener" authorityYear="2020" box="[457,820,1964,1990]" class="Diplopoda" family="Pachybolidae" genus="Dichromatobolus" kingdom="Animalia" order="Spirobolida" pageId="9" pageNumber="116" phylum="Arthropoda" rank="species" species="elephantulus" status="gen. et sp. nov.">
|
||
<emphasis id="B96CEAB5B71DFFAA26E931EFFCA126D5" box="[457,820,1964,1990]" italics="true" pageId="9" pageNumber="116">Dichromatobolus elephantulus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25F57CEB71DFFAA241B31EFFC7B26D5" box="[827,1006,1964,1990]" pageId="9" pageNumber="116" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
Vegetation color marks modified from
|
||
<bibRefCitation id="EF894B56B71DFFAA27DD318CFD9526FA" author="Moat J. & Smith P." box="[253,512,1999,2025]" pageId="9" pageNumber="116" refId="ref5211" refString="Moat J. & Smith P. 2007. Atlas of the Vegetation of Madagscar. Royal botanic gardens, Kew, Richmond." type="book" year="2007">Moat & Smith (2007)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C302652CB71EFFA9279D374BFC312041" box="[189,932,264,338]" pageId="10" pageNumber="117" type="distribution">
|
||
<paragraph id="8BA736A7B71EFFA9279D374BFEC32031" blockId="10.[189,342,264,290]" box="[189,342,264,290]" pageId="10" pageNumber="117">
|
||
<heading id="D0EF81CBB71EFFA9279D374BFEC32031" bold="true" box="[189,342,264,290]" fontSize="11" level="3" pageId="10" pageNumber="117" reason="3">
|
||
<emphasis id="B96CEAB5B71EFFA9279D374BFEC32031" bold="true" box="[189,342,264,290]" pageId="10" pageNumber="117">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA736A7B71EFFA9279D3774FC312041" blockId="10.[189,932,311,338]" box="[189,932,311,338]" pageId="10" pageNumber="117">
|
||
Spiny forests around
|
||
<collectingRegion id="49DCF845B71EFFA926973774FD9F2042" box="[439,522,311,337]" country="Madagascar" name="Toliara" pageId="10" pageNumber="117">Toliara</collectingRegion>
|
||
in Southwest
|
||
<collectingCountry id="F30F7637B71EFFA925903774FCAB2042" box="[688,830,311,337]" name="Madagascar" pageId="10" pageNumber="117">Madagascar</collectingCountry>
|
||
(
|
||
<figureCitation id="13232A22B71EFFA9246E3774FC032041" box="[846,918,311,338]" captionStart="Fig" captionStartId="9.[189,232,1963,1989]" captionTargetBox="[204,1383,279,1926]" captionTargetId="figure-12@9.[204,1383,265,1926]" captionTargetPageId="9" captionText="Fig. 6. Distribution of Dichromatobolus elephantulus gen. et sp. nov. Vegetation color marks modified from Moat & Smith (2007)." figureDoi="http://doi.org/10.5281/zenodo.4085216" httpUri="https://zenodo.org/record/4085216/files/figure.png" pageId="10" pageNumber="117">Fig. 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C302652CB71EFFA9279D373BFBF2231B" pageId="10" pageNumber="117" type="description">
|
||
<paragraph id="8BA736A7B71EFFA9279D373BFE462081" blockId="10.[189,467,376,402]" box="[189,467,376,402]" pageId="10" pageNumber="117">
|
||
<heading id="D0EF81CBB71EFFA9279D373BFE462081" bold="true" box="[189,467,376,402]" fontSize="11" level="3" pageId="10" pageNumber="117" reason="3">
|
||
<emphasis id="B96CEAB5B71EFFA9279D373BFE462081" bold="true" box="[189,467,376,402]" pageId="10" pageNumber="117">Intraspecific variation</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA736A7B71EFFA9279D37E4FBF2231B" blockId="10.[189,1399,423,520]" pageId="10" pageNumber="117">
|
||
The three known populations from southwestern
|
||
<collectingCountry id="F30F7637B71EFFA9242F37E4FC0A20D2" box="[783,927,423,449]" name="Madagascar" pageId="10" pageNumber="117">Madagascar</collectingCountry>
|
||
, as well as the specimens from the pet trade, coming from at least two different imports
|
||
<date id="FFA61067B71EFFA924013788FC4D20F6" box="[801,984,459,485]" pageId="10" pageNumber="117" value="2007" valueMax="2019">2007 and 2019</date>
|
||
look very similar to one another, showing only a slight variation in body ring counts, varying between 47 and 48.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C302652CB71EFFA9279D3473FF6C23AC" pageId="10" pageNumber="117" type="discussion">
|
||
<paragraph id="8BA736A7B71EFFA9279D3473FEBA2359" blockId="10.[189,303,560,586]" box="[189,303,560,586]" pageId="10" pageNumber="117">
|
||
<heading id="D0EF81CBB71EFFA9279D3473FEBA2359" bold="true" box="[189,303,560,586]" fontSize="11" level="3" pageId="10" pageNumber="117" reason="3">
|
||
<emphasis id="B96CEAB5B71EFFA9279D3473FEBA2359" bold="true" box="[189,303,560,586]" pageId="10" pageNumber="117">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA736A7B71EFFA9279D341DFF6C23AC" blockId="10.[189,1399,606,703]" pageId="10" pageNumber="117">Some element associated with the species, or the surroundings they live in, reacts to ethanol. White crystallizations appear around the specimens and appendages when conserved in ethanol for a longer time.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |