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<document id="902512A4428561C01D5DFA98C560EFA0" ID-CLB-Dataset="7631" ID-DOI="10.1206/0003-0090-423.1.1" ID-GBIF-Dataset="2e886aea-b59a-45a6-aeaa-2427d584a894" ID-ISSN="0003-0090" ID-Zenodo-Dep="4613008" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615995240537" checkinUser="felipe" docAuthor="Grimaldi, David A." docDate="2018" docId="03B187A8FFB4FFCDFCB63D587EA6FB6E" docLanguage="en" docName="BulAmeMusNatHis.2018.423.1-97.pdf.imf" docOrigin="Bulletin of the American Museum of Natural History 2018 (423)" docSource="http://www.bioone.org/doi/10.1206/0003-0090-423.1.1" docStyle="DocumentStyle:C5E2DA72A22EF33813C92A197453A310.5:BulAmeMusNatHis.2011-.journal_article.0cover" docStyleId="C5E2DA72A22EF33813C92A197453A310" docStyleName="BulAmeMusNatHis.2011-.journal_article.0cover" docStyleVersion="5" docTitle="Chandleromyia Grimaldi 2018, new genus" docType="treatment" docVersion="7" lastPageNumber="97" masterDocId="FF88FFD0FF86FFFEFFAA39537F49FFF7" masterDocTitle="Basal Cyclorrhapha In Amber From The Cretaceous And Tertiary (Insecta: Diptera), And Their Relationships: Brachycera In Cretaceous Amber Part Ix David A. Grimaldi" masterLastPageNumber="97" masterPageNumber="1" pageNumber="1" updateTime="1698933720305" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="AFCDE6DA9FE840FE17ABCEBF89B93B1B">Basal Cyclorrhapha In Amber From The Cretaceous And Tertiary (Insecta: Diptera), And Their Relationships: Brachycera In Cretaceous Amber Part Ix David A. Grimaldi</mods:title>
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<mods:name id="935B5F1EF3E7884B60C6E3DC460102A5" type="personal">
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<mods:namePart id="DAB542F08446222D4E82E341EF969762">Grimaldi, David A.</mods:namePart>
<mods:affiliation id="2376179034E90AE1E1AF1A1B7FD4A1A0">Division of Invertebrate Zoology American Museum of Natural History, New York</mods:affiliation>
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<mods:title id="D5185069AA220AE6056679B87407B808">Bulletin of the American Museum of Natural History</mods:title>
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<mods:date id="394D019317DC39E16153722BDE95E422">2018</mods:date>
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<mods:number id="3FDC70856F0348F1E5DA7F86D712E5EF">2018-10-24</mods:number>
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<treatment id="03B187A8FFB4FFCDFCB63D587EA6FB6E" ID-DOI="http://doi.org/10.5281/zenodo.4631121" ID-GBIF-Taxon="180670798" ID-Zenodo-Dep="4631121" LSID="urn:lsid:plazi:treatment:03B187A8FFB4FFCDFCB63D587EA6FB6E" httpUri="http://treatment.plazi.org/id/03B187A8FFB4FFCDFCB63D587EA6FB6E" lastPageId="51" pageId="50">
<subSubSection id="C3026535FFB4FFCCFCB63D587B05FBD3" box="[796,1100,1035,1060]" pageId="50" type="nomenclature">
<paragraph id="8BA736BEFFB4FFCCFCB63D587B05FBD3" blockId="50.[796,1100,1035,1060]" box="[796,1100,1035,1060]" pageId="50">
<heading id="D0EF81D2FFB4FFCCFCB63D587B05FBD3" box="[796,1100,1035,1060]" centered="true" fontSize="9" level="2" pageId="50" reason="2">
<taxonomicName id="4C184D3DFFB4FFCCFCB63D587C82FBD4" ID-CoL="89PW5" authority="Grimaldi, 2018" authorityName="Grimaldi" authorityYear="2018" box="[796,971,1035,1059]" class="Insecta" family="Platypezidae" genus="Chandleromyia" kingdom="Animalia" order="Diptera" pageId="50" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96CEAACFFB4FFCCFCB63D587C82FBD4" bold="true" box="[796,971,1035,1059]" italics="true" pageId="50">Chandleromyia</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25F57D7FFB4FFCCFC723D5F7B05FBD3" box="[984,1100,1036,1060]" pageId="50" rank="genus">new genus</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3026535FFB4FFCCFD663D6E7CF4FA15" pageId="50" type="diagnosis">
<paragraph id="8BA736BEFFB4FFCCFD663D6E7CF4FA15" blockId="50.[684,1215,1085,1738]" pageId="50">
DIAGNOSIS: (male only). Eyes extensively holoptic; gena bare, occiput with very fine, scattered short setae; clypeus a broad triangle; palp minute; arista bare; acrostichals entirely absent, only 1 pair of (short, prescutellar) dorsocentrals; 1 short pair of scutellars; 45 notopleurals. Wing entirely covered with dense microtrichia; M not forked (M
<subScript id="179C34FBFFB4FFCCFCB53C637C61FAB6" attach="left" box="[799,808,1328,1345]" fontSize="7" pageId="50">2</subScript>
lost), dm-cu close to wing margin; CuP slightly curved. Tibiae and tarsi with bifid scales. Metatarsomeres 14 flattened, expanded; tarsomere 2 shortest, with blunt spine; tarsomere 3 with clavate seta. Abdomen short, broad; male tergite 7 narrow, tubular.
</paragraph>
</subSubSection>
<subSubSection id="C3026535FFB4FFCCFD663CBF7B7AF9F4" box="[716,1075,1515,1539]" pageId="50" type="materials_examined">
<paragraph id="8BA736BEFFB4FFCCFD663CBF7B7AF9F4" blockId="50.[684,1215,1085,1738]" box="[716,1075,1515,1539]" pageId="50">
<collectionCode id="ED09AE7BFFB4FFCCFD663CBF7C29F9F4" box="[716,864,1515,1539]" pageId="50">
<typeStatus id="54A3881CFFB4FFCCFD663CBF7C4AF9F4" box="[716,771,1516,1539]" pageId="50">TYPE</typeStatus>
SPECIES
</collectionCode>
:
<taxonomicName id="4C184D3DFFB4FFCCFCC43CBF7CA2F9F4" authority="Grimaldi, 2018" authorityName="Grimaldi" authorityYear="2018" box="[878,1003,1515,1539]" class="Insecta" family="Platypezidae" genus="Chandleromyia" kingdom="Animalia" order="Diptera" pageId="50" phylum="Arthropoda" rank="species" species="anomala" status="sp. nov.">
<emphasis id="B96CEAACFFB4FFCCFCC43CBF7CA2F9F4" box="[878,1003,1515,1539]" italics="true" pageId="50">C. anomala</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25F57D7FFB4FFCCFC523CBF7B7AF9F4" box="[1016,1075,1516,1539]" pageId="50" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C3026535FFB4FFCCFD663F5F7BF2F991" pageId="50" type="etymology">
<paragraph id="8BA736BEFFB4FFCCFD663F5F7BF2F991" blockId="50.[684,1215,1085,1738]" pageId="50">
ETYMOLOGY: For Peter J. Chandler, in recognition of his work on global
<taxonomicName id="4C184D3DFFB4FFCCFC763F7E7B2EF9B2" box="[988,1127,1581,1605]" class="Insecta" family="Platypezidae" kingdom="Animalia" order="Diptera" pageId="50" phylum="Arthropoda" rank="family">Platypezidae</taxonomicName>
, and on
<taxonomicName id="4C184D3DFFB4FFCCFD063F1D7DB6F991" box="[684,767,1614,1638]" kingdom="Plantae" order="Diptera" pageId="50" rank="order">Diptera</taxonomicName>
in general. The genus name is feminine.
</paragraph>
</subSubSection>
<subSubSection id="C3026535FFB4FFCDFD663F237EA6FB6E" lastPageId="51" pageId="50" type="discussion">
<paragraph id="8BA736BEFFB4FFCDFD663F237ECDFC98" blockId="50.[684,1215,1085,1738]" lastBlockId="51.[108,637,226,1178]" lastPageId="51" pageId="50">
COMMENTS: This genus is extremely similar phenetically, and apparently close phylogenetically, to
<taxonomicName id="4C184D3DFFB4FFCCFCBF3FE17B72F93D" authority="Grimaldi, 2018" authorityName="Grimaldi" authorityYear="2018" box="[789,1083,1714,1738]" class="Insecta" family="Platypezidae" genus="Lindneromyia" kingdom="Animalia" order="Diptera" pageId="50" phylum="Arthropoda" rank="species" species="neomedialis" status="sp. nov.">
<emphasis id="B96CEAACFFB4FFCCFCBF3FE17B72F93D" box="[789,1083,1714,1738]" italics="true" pageId="50">Lindneromyia neomedialis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25F57D7FFB4FFCCFBFA3FE17BDAF93E" box="[1104,1171,1714,1737]" pageId="50" rank="species">n. sp.</taxonomicNameLabel>
, in Dominican amber, described below. This is based on the squared oral margin, minute palps; face, gena, arista bare; complete lack of acrostichals, one (short, prescutellar) pair of dorsocentral setae; one short pair of scutellars; no supraalars, one short postalar; loss of vein M
<subScript id="179C34FBFFB5FFCDFE0A38C07EE1FE53" attach="left" box="[416,424,403,420]" fontSize="7" pageId="51">2</subScript>
; r-m crossvein near level of cell Sc apex; and thick, stiff, spinelike setae on posterior margin of tergite 6 lacking.
<taxonomicName id="4C184D3DFFB5FFCDFD93389A7F9DFDF4" authorityName="Grimaldi" authorityYear="2018" class="Insecta" family="Platypezidae" genus="Chandleromyia" kingdom="Animalia" order="Diptera" pageId="51" phylum="Arthropoda" rank="genus">
<emphasis id="B96CEAACFFB5FFCDFD93389A7F9DFDF4" italics="true" pageId="51">Chandleromyia</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C184D3DFFB5FFCDFEF638B87EB8FDF4" authorityName="Grimaldi" authorityYear="2018" box="[348,497,491,515]" class="Insecta" family="Platypezidae" genus="Lindneromyia" kingdom="Animalia" order="Diptera" pageId="51" phylum="Arthropoda" rank="species" species="neomedialis">
<emphasis id="B96CEAACFFB5FFCDFEF638B87EB8FDF4" box="[348,497,491,515]" italics="true" pageId="51">L. neomedialis</emphasis>
</taxonomicName>
based on the setose occiput (completely bare in
<taxonomicName id="4C184D3DFFB5FFCDFE713B5F7D24FDD3" authorityName="Kessel" authorityYear="1965" box="[475,621,524,548]" class="Insecta" family="Platypezidae" genus="Lindneromyia" kingdom="Animalia" order="Diptera" pageId="51" phylum="Arthropoda" rank="genus">
<emphasis id="B96CEAACFFB5FFCDFE713B5F7D24FDD3" box="[475,621,524,548]" italics="true" pageId="51">Lindneromyia</emphasis>
</taxonomicName>
); 45 notopleurals (vs. 2); wing membrane entirely with dense microtrichia (vs. partially covered with minute, faint ones); cell dm long, with dm-cu close to wing margin (vs. cell short, vein near middle of wing); vein M
<subScript id="179C34FBFFB5FFCDFE2D3BEE7ED9FD39" attach="left" box="[391,400,701,718]" fontSize="7" pageId="51">1</subScript>
deflexed (vs. almost straight); vein CuP curved (vs. straight); spine and clavate seta on metarsomeres 2 and 3, respectively (vs. without); male tergite 7 narrow, tubular (vs. short, much broader than long, inserted into emargination of tergite 6).
</paragraph>
<paragraph id="8BA736BEFFB5FFCDFF263A2B7EA6FB6E" blockId="51.[108,637,226,1178]" pageId="51">
Though separated by approximately 80 million years, the two flies appear closely related. There is no question about the provenance of the two specimens; AMNH DR14-35 was acquired in the 1990s in
<collectingCountry id="F30F762EFFB5FFCDFEB03AAE7E88FBE2" box="[282,449,1021,1045]" name="Dominican Republic" pageId="51">the Dominican</collectingCountry>
Republic by me; AMNH Bu-KL30-27 was acquired from
<collectingCountry id="F30F762EFFB5FFCDFD9C3D4D7FD0FBA1" name="Myanmar" pageId="51">Myanmar</collectingCountry>
. With a derived platypezine in the mid-Cretaceous, this greatly extends the age of crown-group
<taxonomicName id="4C184D3DFFB5FFCDFEA83DD27EC2FB6E" box="[258,395,1153,1177]" class="Insecta" family="Platypezidae" kingdom="Animalia" order="Diptera" pageId="51" phylum="Arthropoda" rank="family">Platypezidae</taxonomicName>
(fig. 27).
</paragraph>
</subSubSection>
</treatment>
</document>