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<document id="46AE3C04C156E79C3AA9253D61551CC1" ID-CLB-Dataset="7631" ID-DOI="10.1206/0003-0090-423.1.1" ID-GBIF-Dataset="2e886aea-b59a-45a6-aeaa-2427d584a894" ID-ISSN="0003-0090" ID-Zenodo-Dep="4613008" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615995240537" checkinUser="felipe" docAuthor="Grimaldi, David A." docDate="2018" docId="03B187A8FFAAFFD2FF5F38A97B7DFE37" docLanguage="en" docName="BulAmeMusNatHis.2018.423.1-97.pdf.imf" docOrigin="Bulletin of the American Museum of Natural History 2018 (423)" docSource="http://www.bioone.org/doi/10.1206/0003-0090-423.1.1" docStyle="DocumentStyle:C5E2DA72A22EF33813C92A197453A310.5:BulAmeMusNatHis.2011-.journal_article.0cover" docStyleId="C5E2DA72A22EF33813C92A197453A310" docStyleName="BulAmeMusNatHis.2011-.journal_article.0cover" docStyleVersion="5" docTitle="Lebanopeza Grimaldi 2018, new genus" docType="treatment" docVersion="7" lastPageNumber="97" masterDocId="FF88FFD0FF86FFFEFFAA39537F49FFF7" masterDocTitle="Basal Cyclorrhapha In Amber From The Cretaceous And Tertiary (Insecta: Diptera), And Their Relationships: Brachycera In Cretaceous Amber Part Ix David A. Grimaldi" masterLastPageNumber="97" masterPageNumber="1" pageNumber="1" updateTime="1698933720305" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="4887BCB65D45FE05E03A2A40AF1A1B6A">Basal Cyclorrhapha In Amber From The Cretaceous And Tertiary (Insecta: Diptera), And Their Relationships: Brachycera In Cretaceous Amber Part Ix David A. Grimaldi</mods:title>
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<mods:namePart id="BF1DA460F8FEBD2F081E266F1340D69D">Grimaldi, David A.</mods:namePart>
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<treatment id="03B187A8FFAAFFD2FF5F38A97B7DFE37" ID-DOI="http://doi.org/10.5281/zenodo.4612803" ID-GBIF-Taxon="180670809" ID-Zenodo-Dep="4612803" LSID="urn:lsid:plazi:treatment:03B187A8FFAAFFD2FF5F38A97B7DFE37" httpUri="http://treatment.plazi.org/id/03B187A8FFAAFFD2FF5F38A97B7DFE37" pageId="44">
<subSubSection id="C3026535FFAAFFD2FF5F38A97EB2FDE4" box="[245,507,506,531]" pageId="44" type="nomenclature">
<paragraph id="8BA736BEFFAAFFD2FF5F38A97EB2FDE4" blockId="44.[245,507,506,531]" box="[245,507,506,531]" pageId="44">
<heading id="D0EF81D2FFAAFFD2FF5F38A97EB2FDE4" box="[245,507,506,531]" centered="true" fontSize="9" level="2" pageId="44" reason="2">
<taxonomicName id="4C184D3DFFAAFFD2FF5F38A97E33FDE5" ID-CoL="89QJV" authority="Grimaldi, 2018" authorityName="Grimaldi" authorityYear="2018" box="[245,378,506,530]" class="Insecta" family="Platypezidae" genus="Lebanopeza" kingdom="Animalia" order="Diptera" pageId="44" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96CEAACFFAAFFD2FF5F38A97E33FDE5" bold="true" box="[245,378,506,530]" italics="true" pageId="44">Lebanopeza</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25F57D7FFAAFFD2FE2D38A87EB2FDE4" box="[391,507,507,531]" pageId="44" rank="genus">new genus</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3026535FFAAFFD2FF263B7F7FB5FC47" pageId="44" type="diagnosis">
<paragraph id="8BA736BEFFAAFFD2FF263B7F7FB5FC47" blockId="44.[108,638,556,1742]" pageId="44">
DIAGNOSIS: (male only). Eyes holoptic, dorsal facets larger; small prescutellum present; acrostichals in short uniserial row; wing with microtrichia over entire surface; C ends at apex of R
<subScript id="179C34FBFFAAFFD2FDCA3BC87D35FD5B" attach="left" box="[608,636,667,684]" fontSize="7" pageId="44">4+5</subScript>
(vs. M
<subScript id="179C34FBFFAAFFD2FF183BEF7FF2FD3A" attach="left" box="[178,187,700,717]" fontSize="7" pageId="44">1</subScript>
); crossvein dm-cu (cell dm) absent, CuP curved; M entirely spectral, branches of M
<subScript id="179C34FBFFAAFFD2FDE33B8E7D1BFD19" attach="left" box="[585,594,733,750]" fontSize="7" pageId="44">1</subScript>
-M
<subScript id="179C34FBFFAAFFD2FDD93B8E7D35FD19" attach="left" box="[627,636,733,750]" fontSize="7" pageId="44">2</subScript>
fork virtually symmetrical; dark bifid scales lacking on legs; metatarsomeres not expanded; male terminalia lateroflexed to right. Except for the lack of crossvein dm-cu and the faint M vein, all these features appear plesiomorphic for the
<taxonomicName id="4C184D3DFFAAFFD2FFC63ACB7FB1FC47" box="[108,248,920,944]" class="Insecta" family="Platypezidae" kingdom="Animalia" order="Diptera" pageId="44" phylum="Arthropoda" rank="family">Platypezidae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3026535FFAAFFD2FF263AE97D5EFC26" box="[140,535,953,977]" pageId="44" type="materials_examined">
<paragraph id="8BA736BEFFAAFFD2FF263AE97D5EFC26" blockId="44.[108,638,556,1742]" box="[140,535,953,977]" pageId="44">
<collectionCode id="ED09AE7BFFAAFFD2FF263AE97E69FC26" box="[140,288,953,977]" pageId="44">
<typeStatus id="54A3881CFFAAFFD2FF263AE97F8AFC26" box="[140,195,954,977]" pageId="44">TYPE</typeStatus>
SPECIES
</collectionCode>
:
<taxonomicName id="4C184D3DFFAAFFD2FE843AE97EC8FC26" authority="Grimaldi, 2018" authorityName="Grimaldi" authorityYear="2018" box="[302,385,954,977]" class="Insecta" family="Platypezidae" genus="Lebanopeza" kingdom="Animalia" order="Diptera" pageId="44" phylum="Arthropoda" rank="species" species="azari" status="sp. nov.">
<emphasis id="B96CEAACFFAAFFD2FE843AE97EC8FC26" box="[302,385,954,977]" italics="true" pageId="44">L. azari</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25F57D7FFAAFFD2FE253AE97D5BFC26" box="[399,530,953,977]" pageId="44" rank="species">new species</taxonomicNameLabel>
.
</paragraph>
</subSubSection>
<subSubSection id="C3026535FFAAFFD2FF263A897EA1FBC3" pageId="44" type="etymology">
<paragraph id="8BA736BEFFAAFFD2FF263A897EA1FBC3" blockId="44.[108,638,556,1742]" pageId="44">
ETYMOLOGY: From
<collectingCountry id="F30F762EFFAAFFD2FEC03A897E85FC05" box="[362,460,986,1010]" name="Lebanon" pageId="44">Lebanon</collectingCountry>
, the country of origin, and -
<emphasis id="B96CEAACFFAAFFD2FEAA3AAF7E78FBE4" box="[256,305,1020,1043]" italics="true" pageId="44">peza</emphasis>
(foot), a common suffix for generic names in the
<taxonomicName id="4C184D3DFFAAFFD2FEF23D4F7EADFBC3" box="[344,484,1052,1076]" class="Insecta" family="Platypezidae" kingdom="Animalia" order="Diptera" pageId="44" phylum="Arthropoda" rank="family">Platypezidae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3026535FFAAFFD2FF263D6E7B7DFE37" pageId="44" type="discussion">
<paragraph id="8BA736BEFFAAFFD2FF263D6E7EA9FA57" blockId="44.[108,638,556,1742]" pageId="44">
COMMENTS:
<taxonomicName id="4C184D3DFFAAFFD2FE883D6E7EEDFBA2" authorityName="Grimaldi" authorityYear="2018" box="[290,420,1085,1109]" class="Insecta" family="Platypezidae" genus="Lebanopeza" kingdom="Animalia" order="Diptera" pageId="44" phylum="Arthropoda" rank="genus">
<emphasis id="B96CEAACFFAAFFD2FE883D6E7EEDFBA2" box="[290,420,1085,1109]" italics="true" pageId="44">Lebanopeza</emphasis>
</taxonomicName>
appears to be an extinct stem group to
<taxonomicName id="4C184D3DFFAAFFD2FECD3D0C7E96FB81" authorityName="Zetterstedt" authorityYear="1837" box="[359,479,1119,1142]" class="Insecta" family="Platypezidae" genus="Microsania" kingdom="Animalia" order="Diptera" pageId="44" phylum="Arthropoda" rank="genus">
<emphasis id="B96CEAACFFAAFFD2FECD3D0C7E96FB81" box="[359,479,1119,1142]" italics="true" pageId="44">Microsania</emphasis>
</taxonomicName>
+
<taxonomicName id="4C184D3DFFAAFFD2FDAB3D0D7FEBFB60" authorityName="Kessel" authorityYear="1960" class="Insecta" family="Platypezidae" genus="Melanderomyia" kingdom="Animalia" order="Diptera" pageId="44" phylum="Arthropoda" rank="genus">
<emphasis id="B96CEAACFFAAFFD2FDAB3D0D7FEBFB60" italics="true" pageId="44">Melanderomyia</emphasis>
</taxonomicName>
(fig. 27). It lacks at least four features that define these two extant genera (chars. 3, 16, 17, 18; table 2), but shares with them the loss of crossvein dm-cu (cell dm), a distinctly shortened Sc vein and somewhat shortened R
<subScript id="179C34FBFFAAFFD2FE443C5C7EBEFAD7" attach="left" box="[494,503,1295,1312]" fontSize="7" pageId="44">1</subScript>
, and reduction in the medial veins. In
<taxonomicName id="4C184D3DFFAAFFD2FE3C3C767D77FACA" authorityName="Kessel" authorityYear="1960" box="[406,574,1317,1341]" class="Insecta" family="Platypezidae" genus="Melanderomyia" kingdom="Animalia" order="Diptera" pageId="44" phylum="Arthropoda" rank="genus">
<emphasis id="B96CEAACFFAAFFD2FE3C3C767D77FACA" box="[406,574,1317,1341]" italics="true" pageId="44">Melanderomyia</emphasis>
</taxonomicName>
M
<subScript id="179C34FBFFAAFFD2FDF63C637D2CFAB6" attach="left" box="[604,613,1328,1345]" fontSize="7" pageId="44">2</subScript>
is lost; in
<taxonomicName id="4C184D3DFFAAFFD2FF153C147E7EFAA9" authorityName="Zetterstedt" authorityYear="1837" box="[191,311,1351,1374]" class="Insecta" family="Platypezidae" genus="Microsania" kingdom="Animalia" order="Diptera" pageId="44" phylum="Arthropoda" rank="genus">
<emphasis id="B96CEAACFFAAFFD2FF153C147E7EFAA9" box="[191,311,1351,1374]" italics="true" pageId="44">Microsania</emphasis>
</taxonomicName>
the base of M
<subScript id="179C34FBFFAAFFD2FE793C027E95FA95" attach="left" box="[467,476,1361,1378]" fontSize="7" pageId="44">1</subScript>
is lost; in
<taxonomicName id="4C184D3DFFAAFFD2FDE43C157F8CFA88" authorityName="Grimaldi" authorityYear="2018" class="Insecta" family="Platypezidae" genus="Lebanopeza" kingdom="Animalia" order="Diptera" pageId="44" phylum="Arthropoda" rank="genus">
<emphasis id="B96CEAACFFAAFFD2FDE43C157F8CFA88" italics="true" pageId="44">Lebanopeza</emphasis>
</taxonomicName>
the entire stem and fork of M
<subScript id="179C34FBFFAAFFD2FD863C217D7CFA74" attach="left" box="[556,565,1394,1411]" fontSize="7" pageId="44">1</subScript>
-M
<subScript id="179C34FBFFAAFFD2FDFD3C217D29FA74" attach="left" box="[599,608,1394,1411]" fontSize="7" pageId="44">2</subScript>
is extremely faint and unsclerotized.
</paragraph>
<paragraph id="8BA736BEFFAAFFD2FF263CFA7B7DFE37" blockId="44.[108,638,556,1742]" lastBlockId="44.[684,1213,226,448]" pageId="44">
The wing of
<taxonomicName id="4C184D3DFFAAFFD2FEBB3CFA7EC6FA36" authorityName="Grimaldi" authorityYear="2018" box="[273,399,1449,1473]" class="Insecta" family="Platypezidae" genus="Lebanopeza" kingdom="Animalia" order="Diptera" pageId="44" phylum="Arthropoda" rank="genus">
<emphasis id="B96CEAACFFAAFFD2FEBB3CFA7EC6FA36" box="[273,399,1449,1473]" italics="true" pageId="44">Lebanopeza</emphasis>
</taxonomicName>
is similar to
<emphasis id="B96CEAACFFAAFFD2FDB33CF97F98FA15" italics="true" pageId="44">Mauritulus sospes</emphasis>
Mostovski, preserved as a compression (along with portions of the body) in shale from the Early Cretaceous of Eurasia (
<bibRefCitation id="EF894B4FFFAAFFD2FDA83F5F7FF9F9B2" author="Mostovski, M." pageId="44" pagination="106 - 118" refId="ref40396" refString="Mostovski, M., 1995 a. New representatives of Platypezidae (Diptera) from the Mesozoic, and main directions in the evolution of the family. Paleontologischeskii Zhurnal 2: 106 - 118." type="journal article" year="1995">Mostovski, 1995a</bibRefCitation>
). Both genera have a slender wing; membrane densely covered with microtrichia; R
<subScript id="179C34FBFFAAFFD2FDD93F097D35F99C" attach="left" box="[627,636,1626,1643]" fontSize="7" pageId="44">1</subScript>
straight, relatively short; C definitely (
<taxonomicName id="4C184D3DFFAAFFD2FD8E3F3C7FD4F95F" authorityName="Grimaldi" authorityYear="2018" class="Insecta" family="Platypezidae" genus="Lebanopeza" kingdom="Animalia" order="Diptera" pageId="44" phylum="Arthropoda" rank="genus">
<emphasis id="B96CEAACFFAAFFD2FD8E3F3C7FD4F95F" italics="true" pageId="44">Lebanopeza</emphasis>
</taxonomicName>
) or apparently (
<emphasis id="B96CEAACFFAAFFD2FEF33FC37E95F95F" box="[345,476,1680,1704]" italics="true" pageId="44">Mauritulus)</emphasis>
ending at the apex of R
<subScript id="179C34FBFFAAFFD2FF773FEE7FB0F939" attach="left" box="[221,249,1725,1742]" fontSize="7" pageId="44">4+5</subScript>
; M
<subScript id="179C34FBFFAAFFD2FE8E3FEE7E64F939" attach="right" box="[292,301,1725,1742]" fontSize="7" pageId="44">1</subScript>
-M
<subScript id="179C34FBFFAAFFD2FEE53FEE7E11F939" attach="left" box="[335,344,1725,1742]" fontSize="7" pageId="44">2</subScript>
fork nearly symmetrical; and crossvein dm-cu (cell dm) absent.
<taxonomicName id="4C184D3DFFAAFFD2FBCF39B17D94FEED" authorityName="Grimaldi" authorityYear="2018" class="Insecta" family="Platypezidae" genus="Lebanopeza" kingdom="Animalia" order="Diptera" pageId="44" phylum="Arthropoda" rank="genus">
<emphasis id="B96CEAACFFAAFFD2FBCF39B17D94FEED" italics="true" pageId="44">Lebanopeza</emphasis>
</taxonomicName>
differs from
<emphasis id="B96CEAACFFAAFFD2FCC638507CADFEEC" box="[876,996,259,283]" italics="true" pageId="44">Mauritulus</emphasis>
by having a shorter Sc cell (0.3× wing length, vs. 0.45×), cell cup with apical stem of CuP+A
<subScript id="179C34FBFFAAFFD2FC3D38037CE9FE96" attach="left" box="[919,928,336,353]" fontSize="7" pageId="44">1</subScript>
(in
<emphasis id="B96CEAACFFAAFFD2FC6038167B08FEAA" box="[970,1089,325,349]" italics="true" pageId="44">Mauritulus</emphasis>
these veins join virtually at the wing margin), and in having no metatarsomeres expanded (in
<emphasis id="B96CEAACFFAAFFD2FBB138D47BDAFE68" box="[1051,1171,391,415]" italics="true" pageId="44">Mauritulus</emphasis>
the basitarsomere is slightly expanded).
</paragraph>
</subSubSection>
</treatment>
</document>