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<document id="EA386F13D2BEF134208B5C41B2D27151" ID="10.11646/zootaxa.4179.2.3" ID-DOI="10.11646/zootaxa.4179.2.3" ID-GBIF-Dataset="6fc39bea-49f6-4bcb-a07a-6027a4e829d5" ID-ISSN="1175-5326" ID-Zenodo-Dep="261616" ID-ZooBank="A456F585-0F3A-4F98-99CC-795A1A3603C4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1477898728895" checkinUser="plazi" docAuthor="Delventhal, Naomi R., Mooi, Randall D., Bogorodsky, Sergey V. &amp; Mal, Ahmad O." docDate="2016" docId="03B187A8CD13FE0C4C9FFF5E58D3FA25" docLanguage="en" docName="zootaxa.4179.2.3.pdf" docOrigin="Zootaxa 4179 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Callogobius pilosimentum Delventhal, Mooi, Bogorodsky &amp; Mal, 2016, sp. nov." docType="treatment" docVersion="8" lastPageNumber="232" masterDocId="FF88FFD0CD11FE0B4C08FFC9593BFFC1" masterDocTitle="A review of the Callogobius (Teleostei: Gobiidae) from the Red Sea with the description of a new species" masterLastPageNumber="243" masterPageNumber="225" pageNumber="227" updateTime="1698391306123" updateUser="plazi">
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<mods:title id="FD3986C34EA861D1587E517F4330AB83">A review of the Callogobius (Teleostei: Gobiidae) from the Red Sea with the description of a new species</mods:title>
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<mods:namePart id="9BA0E85920BD2E5269156726D1D4F12A">Delventhal, Naomi R.</mods:namePart>
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<mods:namePart id="4F27BFEA804032A65B4CF7B031D2D490">Mooi, Randall D.</mods:namePart>
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<mods:namePart id="AC1FE385A0547207734322EA27DFBEC8">Bogorodsky, Sergey V.</mods:namePart>
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<mods:namePart id="C60E6C575C3910EFB424AAC0CD6C33BC">Mal, Ahmad O.</mods:namePart>
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<mods:date id="7AEB2AE9FB83FD2A785C8A9E16C75B98">2016</mods:date>
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<treatment id="03B187A8CD13FE0C4C9FFF5E58D3FA25" ID-DOI="http://doi.org/10.5281/zenodo.5679403" ID-GBIF-Taxon="124560751" ID-Zenodo-Dep="5679403" LSID="urn:lsid:plazi:treatment:03B187A8CD13FE0C4C9FFF5E58D3FA25" httpUri="http://treatment.plazi.org/id/03B187A8CD13FE0C4C9FFF5E58D3FA25" lastPageId="7" lastPageNumber="232" pageId="2" pageNumber="227">
<subSubSection id="C3026535CD13FE094C9FFF5E58B4FF36" pageId="2" pageNumber="227" type="nomenclature">
<paragraph id="8BA736BECD13FE094C9FFF5E5B03FF73" blockId="2.[151,568,151,247]" box="[151,568,151,178]" pageId="2" pageNumber="227">
<heading id="D0EF81D2CD13FE094C9FFF5E5B03FF73" bold="true" box="[151,568,151,178]" fontSize="11" level="1" pageId="2" pageNumber="227" reason="1">
<emphasis id="B96CEAACCD13FE094C9FFF5E5B03FF73" bold="true" box="[151,568,151,178]" pageId="2" pageNumber="227">
<taxonomicName id="4C184D3DCD13FE094C9FFF5E58EFFF70" ID-CoL="5X7R7" box="[151,468,151,177]" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="227" phylum="Chordata" rank="species" species="pilosimentum" status="sp. nov.">
<emphasis id="B96CEAACCD13FE094C9FFF5E58EFFF70" bold="true" box="[151,468,151,177]" italics="true" pageId="2" pageNumber="227">Callogobius pilosimentum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F57D7CD13FE094DD3FF515B03FF73" box="[475,568,152,178]" pageId="2" pageNumber="227" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA736BECD13FE094C9FFF7358B4FF36" blockId="2.[151,568,151,247]" pageId="2" pageNumber="227">
Hairy-chinned Flapheaded Goby (
<figureCitation id="13232A3BCD13FE094C96FF17581CFF36" box="[158,295,222,247]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="3.[151,250,1751,1773]" captionStartId-1="4.[151,250,1952,1974]" captionTargetBox-0="[247,1331,204,1700]" captionTargetBox-1="[187,1400,203,1903]" captionTargetId-0="figure@3.[237,1347,193,1715]" captionTargetId-1="figure@4.[187,1400,203,1904]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURE 1. Sensory pore and papillae pattern on the head of Callogobius pilosimentum sp. nov., SMF 35756, holotype, female, 36.4 mm SL. A. Lateral view; B. Ventral view. Letter abbreviations of sensory pores follow Akihito &amp; Meguro (1977) and descriptive names are modified from Takagi (1957). Papillae row numbering follows Akihito and Meguro and descriptive names are from Delventhal &amp; Mooi (2013). Sensory pores: B = posterior nasal; C = anterior interorbital; D = posterior interorbital; E = supraotic; F = anterior otic; G = posterior otic; H = intertemporal; M, N, O = preopercular; K = anterior temporal; L = posterior temporal. Papillae rows: 2 = postnasal; 9 = anterior suborbital; 10 = mid suborbital; 11 = posterior suborbital; 12 = longitudinal cheek; 13 = transverse cheek; 14 = longitudinal maxillary; 15 = longitudinal mandibular; 16 = transverse mandibular; 17 = postorbital; 20 = preopercular; 21 = transverse opercular (rows 20 and 21 continuous). Scale bar = 2 mm. Illustration by RDM and NRD." captionText-1="FIGURE 2. Callogobius pilosimentum sp. nov., live coloration: A. SMF 35756, holotype, female, 36.4 mm SL, Farasan Island, Saudi Arabia; B. Fresh coloration, uncatalogued, Farasan Island, Saudi Arabia, illustrating pattern frequently induced by stress; C. SMF 35760, paratype, female, 55.8 mm SL, Obhur, Jeddah, Saudi Arabia. Photos by SVB." httpUri-0="https://zenodo.org/record/261617/files/figure.png" httpUri-1="https://zenodo.org/record/261618/files/figure.png" pageId="2" pageNumber="227">Figures 12</figureCitation>
,
<tableCitation id="C69A0305CD13FE094D3BFF1758BFFF36" box="[307,388,222,247]" captionStart="TABLE 1" captionStartId="5.[151,239,152,175]" captionTargetBox="[159,1268,298,858]" captionTargetPageId="5" captionText="TABLE 1. Selected measurements of the holotype and paratypes of C. pilosimentum sp. nov.; standard length is provided in mm, values for other morphometrics are as percentage of standard length (SL) or head length (HL), as indicated. For paratypes, the range is followed by an average value for all types in parentheses. The two largest specimens (63 and 67 mm SL) were males (M), the next 3 largest specimens (55.8, 50.3, 49.3 mm SL) were females (F)." httpUri="http://table.plazi.org/id/DF676636CD14FE0E4C9FFF515CA7FECE" pageId="2" pageNumber="227" tableUuid="DF676636CD14FE0E4C9FFF515CA7FECE">Table 1</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3026535CD13FE094C9FFEEF5A76FE9A" pageId="2" pageNumber="227" type="description">
<paragraph id="8BA736BECD13FE094C9FFEEF5BBAFEFD" blockId="2.[151,1389,294,378]" box="[151,641,294,317]" pageId="2" pageNumber="227">
<taxonomicName id="4C184D3DCD13FE094C9FFEEF5879FEFC" box="[151,322,294,317]" class="Actinopterygii" family="Gobiidae" genus="Drombus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="227" phylum="Chordata" rank="species" species="irrasus">
<emphasis id="B96CEAACCD13FE094C9FFEEF5879FEFC" box="[151,322,294,317]" italics="true" pageId="2" pageNumber="227">Drombus irrasus</emphasis>
</taxonomicName>
(non Smith)—Goren 1979: 36.
</paragraph>
<paragraph id="8BA736BECD13FE094C9FFE8C5A76FE9A" blockId="2.[151,1389,294,378]" box="[151,845,325,347]" pageId="2" pageNumber="227">
<treatmentCitationGroup id="AB081190CD13FE094C9FFE8C5A76FE9A" box="[151,845,325,347]" pageId="2" pageNumber="227">
<taxonomicName id="4C184D3DCD13FE094C9FFE8C585AFE9A" ID-CoL="37PLZ" box="[151,353,325,347]" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="227" phylum="Chordata" rank="species" species="irrasus">
<emphasis id="B96CEAACCD13FE094C9FFE8C585AFE9A" box="[151,353,325,347]" italics="true" pageId="2" pageNumber="227">Callogobius irrasus</emphasis>
</taxonomicName>
(non Smith)—
<treatmentCitation id="0AB910AFCD13FE094DF1FE8C5B93FE9A" author="Goren" box="[505,680,325,347]" page="213" pageId="2" pageNumber="227" year="1980">
<bibRefCitation id="EF894B4FCD13FE094DF1FE8C5B4DFE9A" author="Goren" box="[505,630,325,347]" pageId="2" pageNumber="227" refString="Goren, M. (1980) Red Sea fishes assigned to the genus Callogobius Bleeker with a description of a new species (Teleostei: Gobiidae). Israel Journal of Zoology, 28, 209 - 217. [for 1979]" type="journal article" year="1980">Goren 1980</bibRefCitation>
: 213
</treatmentCitation>
; Dor 1984: 241.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3026535CD13FE094C9FFEAD5C56FEBB" box="[151,1389,356,378]" pageId="2" pageNumber="227" type="reference_group">
<paragraph id="8BA736BECD13FE094C9FFEAD5C56FEBB" blockId="2.[151,1389,294,378]" box="[151,1389,356,378]" pageId="2" pageNumber="227">
<treatmentCitationGroup id="AB081190CD13FE094C9FFEAD5C56FEBB" box="[151,1389,356,378]" pageId="2" pageNumber="227">
<taxonomicName id="4C184D3DCD13FE094C9FFEAD58A7FEBB" ID-CoL="PV9Y" box="[151,412,356,378]" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="227" phylum="Chordata" rank="species" species="maculipinnis">
<emphasis id="B96CEAACCD13FE094C9FFEAD58A7FEBB" box="[151,412,356,378]" italics="true" pageId="2" pageNumber="227">Callogobius maculipinnis</emphasis>
</taxonomicName>
(non Fowler)—
<treatmentCitation id="0AB910AFCD13FE094E48FEAD5A13FEBB" author="Goren" box="[576,808,356,378]" page="299" pageId="2" pageNumber="227" year="1991">
<bibRefCitation id="EF894B4FCD13FE094E48FEAD5BCEFEBB" author="Goren" box="[576,757,356,378]" pageId="2" pageNumber="227" refString="Goren, M., Miroz, A. &amp; Baranes, A. (1991) Callogobius amikami, a new species of goby (Gobiidae) from the Red Sea. Cybium, 15 (4), 299 - 302." type="journal article" year="1991">
Goren
<emphasis id="B96CEAACCD13FE094E8EFEAC5B89FEBB" box="[646,690,356,378]" italics="true" pageId="2" pageNumber="227">et al</emphasis>
. 1991
</bibRefCitation>
: 299
</treatmentCitation>
; Goren &amp; Dor 1994: 53;
<treatmentCitation id="0AB910AFCD13FE09482EFEAD5C52FEBB" author="Golani" box="[1062,1385,356,378]" page="46" pageId="2" pageNumber="227" year="2010">
<bibRefCitation id="EF894B4FCD13FE09482EFEAD5C7FFEBB" author="Golani" box="[1062,1348,356,378]" pageId="2" pageNumber="227" refString="Golani, D. &amp; Bogorodsky, S. V. (2010) The fishes of the Red Sea-reappraisal and updated checklist. Zootaxa, 2463, 1 - 135." type="journal article" year="2010">Golani &amp; Bogorodsky 2010</bibRefCitation>
: 46
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3026535CD13FE094C9FFE6F58E7FB3A" pageId="2" pageNumber="227" type="materials_examined">
<paragraph id="8BA736BECD13FE094C9FFE6F5A5FFE7E" blockId="2.[151,1438,422,1923]" box="[151,868,422,447]" pageId="2" pageNumber="227">
<emphasis id="B96CEAACCD13FE094C9FFE6F58BBFE7E" bold="true" box="[151,384,422,447]" pageId="2" pageNumber="227">Material examined.</emphasis>
All specimens collected from the Red Sea.
</paragraph>
<paragraph id="8BA736BECD13FE094CCFFE0359F1FDEA" blockId="2.[151,1438,422,1923]" pageId="2" pageNumber="227">
<materialsCitation id="3B703CE3CD13FE094CCFFE0359F1FDEA" ID-GBIF-Occurrence="1324332036" collectingDate="2012-02-20" collectionCode="SMF" collectorName="S. V. Bogorodsky &amp; T. J. Alpermann" country="Saudi Arabia" elevation="4" latitude="16.71805" location="Farasan Island" longLatPrecision="1" longitude="42.065567" pageId="2" pageNumber="227" specimenCount="1" typeStatus="holotype">
<emphasis id="B96CEAACCD13FE094CCFFE035800FE22" bold="true" box="[199,315,458,483]" pageId="2" pageNumber="227">
<typeStatus id="54A3881CCD13FE094CCFFE03580DFE22" box="[199,310,458,483]" pageId="2" pageNumber="227" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED09AE7BCD13FE094D4FFE025844FE22" box="[327,383,459,483]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="2" pageNumber="227">SMF</collectionCode>
35756 (KAU12-0224), female,
<quantity id="4CE09B5BCD13FE094EF6FE035A5CFE23" box="[766,871,458,483]" metricMagnitude="-2" metricUnit="m" metricValue="3.6399999999999997" pageId="2" pageNumber="227" unit="mm" value="36.4">36.4 mm</quantity>
SL,
<collectingCountry id="F30F762ECD13FE094FAFFE035D78FE22" box="[935,1091,458,483]" name="Saudi Arabia" pageId="2" pageNumber="227">Saudi Arabia</collectingCountry>
,
<location id="8EC76065CD13FE09485BFE025DC5FE22" LSID="urn:lsid:plazi:treatment:03B187A8CD13FE0C4C9FFF5E58D3FA25:8EC76065CD13FE09485BFE025DC5FE22" box="[1107,1278,458,483]" country="Saudi Arabia" latitude="16.71805" longLatPrecision="1" longitude="42.065567" name="Farasan Island" pageId="2" pageNumber="227">Farasan Island</location>
,
<geoCoordinate id="EE2C5079CD13FE094907FE035CA6FE22" box="[1295,1437,458,483]" direction="north" orientation="latitude" pageId="2" pageNumber="227" precision="1" value="16.71805">N16°43.083'</geoCoordinate>
<geoCoordinate id="EE2C5079CD13FE094C9FFE26581EFDC6" box="[151,293,495,519]" direction="east" orientation="longitude" pageId="2" pageNumber="227" precision="1" value="42.065567">E42°03.934'</geoCoordinate>
, isolated coral patch of lagoon,
<elevation id="0035D18DCD13FE094E8EFE275BF5FDC7" box="[646,718,494,519]" metricMagnitude="0" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="2" pageNumber="227" unit="m" value="4.0" valueMax="5.0" valueMin="3.0">
<quantity id="4CE09B5BCD13FE094E8EFE275BF5FDC7" box="[646,718,494,519]" metricMagnitude="0" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="2" pageNumber="227" unit="m" value="4.0" valueMax="5.0" valueMin="3.0">35 m</quantity>
</elevation>
,
<collectorName id="26ED5368CD13FE094ED1FE265AACFDC6" box="[729,919,494,519]" pageId="2" pageNumber="227">S.V. Bogorodsky</collectorName>
&amp;
<collectorName id="26ED5368CD13FE094FB1FE265D52FDC6" box="[953,1129,494,519]" pageId="2" pageNumber="227">T.J. Alpermann</collectorName>
,
<collectingDate id="EFE2E996CD13FE09487BFE265C01FDC6" box="[1139,1338,494,519]" pageId="2" pageNumber="227" value="2012-02-20">
<date id="FFA6107ECD13FE09487BFE265C01FDC6" box="[1139,1338,494,519]" pageId="2" pageNumber="227" value="2012-02-20">20 February 2012</date>
</collectingDate>
(
<figureCitation id="13232A3BCD13FE094940FE275CAEFDC6" box="[1352,1429,494,519]" captionStart="FIGURE 1" captionStartId="3.[151,250,1751,1773]" captionTargetBox="[247,1331,204,1700]" captionTargetId="figure@3.[237,1347,193,1715]" captionTargetPageId="3" captionText="FIGURE 1. Sensory pore and papillae pattern on the head of Callogobius pilosimentum sp. nov., SMF 35756, holotype, female, 36.4 mm SL. A. Lateral view; B. Ventral view. Letter abbreviations of sensory pores follow Akihito &amp; Meguro (1977) and descriptive names are modified from Takagi (1957). Papillae row numbering follows Akihito and Meguro and descriptive names are from Delventhal &amp; Mooi (2013). Sensory pores: B = posterior nasal; C = anterior interorbital; D = posterior interorbital; E = supraotic; F = anterior otic; G = posterior otic; H = intertemporal; M, N, O = preopercular; K = anterior temporal; L = posterior temporal. Papillae rows: 2 = postnasal; 9 = anterior suborbital; 10 = mid suborbital; 11 = posterior suborbital; 12 = longitudinal cheek; 13 = transverse cheek; 14 = longitudinal maxillary; 15 = longitudinal mandibular; 16 = transverse mandibular; 17 = postorbital; 20 = preopercular; 21 = transverse opercular (rows 20 and 21 continuous). Scale bar = 2 mm. Illustration by RDM and NRD." httpUri="https://zenodo.org/record/261617/files/figure.png" pageId="2" pageNumber="227">Figs. 1</figureCitation>
,
<figureCitation id="13232A3BCD13FE094C9FFDDA599DFDEA" box="[151,166,531,555]" captionStart="FIGURE 2" captionStartId="4.[151,250,1952,1974]" captionTargetBox="[187,1400,203,1903]" captionTargetId="figure@4.[187,1400,203,1904]" captionTargetPageId="4" captionText="FIGURE 2. Callogobius pilosimentum sp. nov., live coloration: A. SMF 35756, holotype, female, 36.4 mm SL, Farasan Island, Saudi Arabia; B. Fresh coloration, uncatalogued, Farasan Island, Saudi Arabia, illustrating pattern frequently induced by stress; C. SMF 35760, paratype, female, 55.8 mm SL, Obhur, Jeddah, Saudi Arabia. Photos by SVB." httpUri="https://zenodo.org/record/261618/files/figure.png" pageId="2" pageNumber="227">2</figureCitation>
A).
</materialsCitation>
</paragraph>
<paragraph id="8BA736BECD13FE094CCFFDFF5A7AFBAA" blockId="2.[151,1438,422,1923]" pageId="2" pageNumber="227">
<materialsCitation id="3B703CE3CD13FE094CCFFDFF5BCDFDB2" ID-GBIF-Occurrence="1324332045" collectingDate="1977-04-25" collectionCode="BPBM" collectorName="El Hameira &amp; J. E. Randall &amp; O. Gon" country="Egypt" elevation="12" location="El Hameira" pageId="2" pageNumber="227" specimenCode="BPBM 21518" specimenCount="17" typeStatus="paratype">
<typeStatus id="54A3881CCD13FE094CCFFDFF5804FD8E" box="[199,319,566,591]" pageId="2" pageNumber="227" type="paratype">
<emphasis id="B96CEAACCD13FE094CCFFDFF5804FD8E" bold="true" box="[199,319,566,591]" pageId="2" pageNumber="227">Paratypes</emphasis>
</typeStatus>
(
<specimenCount id="9D1EFD37CD13FE094D58FDFE58D7FD8E" box="[336,492,566,591]" pageId="2" pageNumber="227" type="generic">17 specimens</specimenCount>
, 25.567.0 mm SL).
<collectingCountry id="F30F762ECD13FE094ED7FDFF5A1DFD8E" box="[735,806,566,591]" name="Egypt" pageId="2" pageNumber="227">
<emphasis id="B96CEAACCD13FE094ED7FDFF5A1DFD8E" bold="true" box="[735,806,566,591]" pageId="2" pageNumber="227">Egypt</emphasis>
</collectingCountry>
:
<specimenCode id="DBBE9EC5CD13FE094F3FFDFE5AEDFD8E" box="[823,982,566,591]" collectionCode="BPBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="2" pageNumber="227">
<collectionCode id="ED09AE7BCD13FE094F3FFDFE5ABEFD8F" box="[823,901,567,590]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="2" pageNumber="227">BPBM</collectionCode>
21518
</specimenCode>
, male?,
<quantity id="4CE09B5BCD13FE09483DFDFF5DA1FD8F" box="[1077,1178,566,591]" metricMagnitude="-2" metricUnit="m" metricValue="4.15" pageId="2" pageNumber="227" unit="mm" value="41.5">41.5 mm</quantity>
SL,
<collectorName id="26ED5368CD13FE0948DBFDFF5C6CFD8E" box="[1235,1367,566,591]" pageId="2" pageNumber="227">
<location id="8EC76065CD13FE0948DBFDFF5C6CFD8E" LSID="urn:lsid:plazi:treatment:03B187A8CD13FE0C4C9FFF5E58D3FA25:8EC76065CD13FE0948DBFDFF5C6CFD8E" box="[1235,1367,566,591]" country="Egypt" name="El Hameira" pageId="2" pageNumber="227">El Hameira</location>
</collectorName>
, coral knoll in
<elevation id="0035D18DCD13FE094CFFFD925808FDB3" box="[247,307,603,627]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="227" unit="m" value="12.0">
<quantity id="4CE09B5BCD13FE094CFFFD925808FDB3" box="[247,307,603,627]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="227" unit="m" value="12.0">12 m</quantity>
</elevation>
,
<collectorName id="26ED5368CD13FE094D48FD9258F1FDB2" box="[320,458,602,627]" pageId="2" pageNumber="227">J.E. Randall</collectorName>
&amp;
<collectorName id="26ED5368CD13FE094DF8FD925B7CFDB2" box="[496,583,603,627]" pageId="2" pageNumber="227">O. Gon</collectorName>
,
<collectingDate id="EFE2E996CD13FE094E5BFD935BCDFDB2" box="[595,758,602,627]" pageId="2" pageNumber="227" value="1977-04-25">
<date id="FFA6107ECD13FE094E5BFD935BCDFDB2" box="[595,758,602,627]" pageId="2" pageNumber="227" value="1977-04-25">25 April 1977</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B703CE3CD13FE094F0BFD935D77FD56" ID-GBIF-Occurrence="1324332040" collectingDate="2011-04-13" collectionCode="KAUMM" collectorName="S. V. Bogorodsky" country="Saudi Arabia" elevation="7" latitude="28.437056" location="Maqna" longLatPrecision="1" longitude="34.76328" pageId="2" pageNumber="227" specimenCode="KAUMM 373" specimenCount="1" typeStatus="paratype">
<collectingCountry id="F30F762ECD13FE094F0BFD935A9FFDB2" box="[771,932,602,627]" name="Saudi Arabia" pageId="2" pageNumber="227">
<emphasis id="B96CEAACCD13FE094F0BFD935A9FFDB2" bold="true" box="[771,932,602,627]" pageId="2" pageNumber="227">Saudi Arabia</emphasis>
</collectingCountry>
:
<specimenCode id="DBBE9EC5CD13FE094FBCFD925D63FDB2" box="[948,1112,603,627]" collectionCode="KAUMM" pageId="2" pageNumber="227">
<collectionCode id="ED09AE7BCD13FE094FBCFD925D19FDB2" box="[948,1058,603,627]" pageId="2" pageNumber="227">KAUMM</collectionCode>
373
</specimenCode>
, male,
<quantity id="4CE09B5BCD13FE0948A2FD925C34FDB3" box="[1194,1295,603,627]" metricMagnitude="-2" metricUnit="m" metricValue="4.71" pageId="2" pageNumber="227" unit="mm" value="47.1">47.1 mm</quantity>
SL,
<location id="8EC76065CD13FE09494FFD925CA3FDB3" LSID="urn:lsid:plazi:treatment:03B187A8CD13FE0C4C9FFF5E58D3FA25:8EC76065CD13FE09494FFD925CA3FDB3" box="[1351,1432,603,626]" country="Saudi Arabia" latitude="28.437056" longLatPrecision="1" longitude="34.76328" name="Maqna" pageId="2" pageNumber="227">Maqna</location>
,
<geoCoordinate id="EE2C5079CD13FE094C9FFDB75805FD56" box="[151,318,638,663]" direction="north" orientation="latitude" pageId="2" pageNumber="227" precision="1" value="28.437056">N28°26'13.40&quot;</geoCoordinate>
<geoCoordinate id="EE2C5079CD13FE094D4FFDB758C7FD56" box="[327,508,638,663]" direction="east" orientation="longitude" pageId="2" pageNumber="227" precision="1" value="34.76328">E034°45'47.80&quot;</geoCoordinate>
, steep slope,
<elevation id="0035D18DCD13FE094E91FDB65BF2FD57" box="[665,713,639,662]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="227" unit="m" value="7.0">
<quantity id="4CE09B5BCD13FE094E91FDB65BF2FD57" box="[665,713,639,662]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="227" unit="m" value="7.0">7 m</quantity>
</elevation>
,
<collectorName id="26ED5368CD13FE094ED0FDB65AA1FD56" box="[728,922,638,663]" pageId="2" pageNumber="227">S.V. Bogorodsky</collectorName>
,
<collectingDate id="EFE2E996CD13FE094FA0FDB65D77FD56" box="[936,1100,638,663]" pageId="2" pageNumber="227" value="2011-04-13">
<date id="FFA6107ECD13FE094FA0FDB65D77FD56" box="[936,1100,638,663]" pageId="2" pageNumber="227" value="2011-04-13">13 April 2011</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B703CE3CD13FE094853FDB65806FD1E" ID-GBIF-Occurrence="1324332050" collectingDate="2011-04-19" collectionCode="KAUMM" collectorName="S. V. Bogorodsky" country="Saudi Arabia" elevation="2" latitude="21.7092" location="Jeddah" longLatPrecision="1" longitude="39.096737" municipality="Obhur" pageId="2" pageNumber="227" specimenCode="KAUMM 374" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBBE9EC5CD13FE094853FDB65C3BFD56" box="[1115,1280,639,663]" collectionCode="KAUMM" pageId="2" pageNumber="227">
<collectionCode id="ED09AE7BCD13FE094853FDB65DF2FD56" box="[1115,1225,639,663]" pageId="2" pageNumber="227">KAUMM</collectionCode>
374
</specimenCode>
, female,
<quantity id="4CE09B5BCD13FE094963FDB659F8FD7B" metricMagnitude="-2" metricUnit="m" metricValue="2.83" pageId="2" pageNumber="227" unit="mm" value="28.3">28.3 mm</quantity>
SL,
<location id="8EC76065CD13FE094CF2FD6B5870FD7A" LSID="urn:lsid:plazi:treatment:03B187A8CD13FE0C4C9FFF5E58D3FA25:8EC76065CD13FE094CF2FD6B5870FD7A" box="[250,331,674,699]" country="Saudi Arabia" latitude="21.7092" longLatPrecision="1" longitude="39.096737" municipality="Obhur" name="Jeddah" pageId="2" pageNumber="227">Jeddah</location>
,
<collectingMunicipality id="6BC3ACC4CD13FE094D5FFD6B58A5FD7A" box="[343,414,674,699]" pageId="2" pageNumber="227">Obhur</collectingMunicipality>
(Sharm Obhur),
<geoCoordinate id="EE2C5079CD13FE094E68FD6A5A3CFD7A" box="[608,775,675,699]" direction="north" orientation="latitude" pageId="2" pageNumber="227" precision="1" value="21.7092">N21°42'33.12&quot;</geoCoordinate>
<geoCoordinate id="EE2C5079CD13FE094F07FD6B5A8DFD7A" box="[783,950,674,699]" direction="east" orientation="longitude" pageId="2" pageNumber="227" precision="1" value="39.096737">E39°05'48.26&quot;</geoCoordinate>
, base of rocky reef,
<elevation id="0035D18DCD13FE094895FD6A5DF0FD7B" box="[1181,1227,675,699]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="227" unit="m" value="2.0">
<quantity id="4CE09B5BCD13FE094895FD6A5DF0FD7B" box="[1181,1227,675,699]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="227" unit="m" value="2.0">2 m</quantity>
</elevation>
,
<collectorName id="26ED5368CD13FE0948D0FD6A5CA3FD7A" box="[1240,1432,674,699]" pageId="2" pageNumber="227">S.V. Bogorodsky</collectorName>
,
<collectingDate id="EFE2E996CD13FE094C9FFD0E5806FD1E" box="[151,317,710,735]" pageId="2" pageNumber="227" value="2011-04-19">
<date id="FFA6107ECD13FE094C9FFD0E5806FD1E" box="[151,317,710,735]" pageId="2" pageNumber="227" value="2011-04-19">19 April 2011</date>
</collectingDate>
</materialsCitation>
; KAUMM 375 (KAU12-0220), male,
<quantity id="4CE09B5BCD13FE094F08FD0F5A53FD1F" box="[768,872,710,735]" metricMagnitude="-2" metricUnit="m" metricValue="2.55" pageId="2" pageNumber="227" unit="mm" value="25.5">25.5 mm</quantity>
SL, collected with the holotype;
<materialsCitation id="3B703CE3CD13FE0948FAFD0E5BF6FCE6" ID-GBIF-Occurrence="1324332031" collectingDate="2012-02-20" collectionCode="KAUMM" collectorName="S. V. Bogorodsky &amp; T. J. Alpermann" country="Iceland" elevation="4" latitude="16.71805" location="Island" longLatPrecision="1" longitude="42.065567" pageId="2" pageNumber="227" specimenCode="KAUMM 376" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBBE9EC5CD13FE0948FAFD0E5CA3FD1E" box="[1266,1432,710,735]" collectionCode="KAUMM" pageId="2" pageNumber="227">
<collectionCode id="ED09AE7BCD13FE0948FAFD0E5C5BFD1E" box="[1266,1376,711,735]" pageId="2" pageNumber="227">KAUMM</collectionCode>
376
</specimenCode>
, male, 37.0 mm SL, Farasan
<collectingCountry id="F30F762ECD13FE094DFFFD235B05FCC2" box="[503,574,746,771]" name="Iceland" pageId="2" pageNumber="227">Island</collectingCountry>
,
<geoCoordinate id="EE2C5079CD13FE094E59FD235BDBFCC2" box="[593,736,746,771]" direction="north" orientation="latitude" pageId="2" pageNumber="227" precision="1" value="16.71805">N16°43.083'</geoCoordinate>
<geoCoordinate id="EE2C5079CD13FE094EE5FD225A40FCC2" box="[749,891,747,771]" direction="east" orientation="longitude" pageId="2" pageNumber="227" precision="1" value="42.065567">E42°03.934'</geoCoordinate>
, isolated coral patch of lagoon,
<elevation id="0035D18DCD13FE094903FD235C60FCC3" box="[1291,1371,746,771]" metricMagnitude="0" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="2" pageNumber="227" unit="m" value="4.0" valueMax="5.0" valueMin="3.0">
<quantity id="4CE09B5BCD13FE094903FD235C60FCC3" box="[1291,1371,746,771]" metricMagnitude="0" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="2" pageNumber="227" unit="m" value="4.0" valueMax="5.0" valueMin="3.0">35 m</quantity>
</elevation>
,
<collectorName id="26ED5368CD13FE094966FD22581BFCE6" pageId="2" pageNumber="227">S.V. Bogorodsky</collectorName>
&amp;
<collectorName id="26ED5368CD13FE094D4DFCC658CDFCE6" box="[325,502,782,807]" pageId="2" pageNumber="227">T.J. Alpermann</collectorName>
,
<collectingDate id="EFE2E996CD13FE094E0AFCC65BF6FCE6" box="[514,717,782,807]" pageId="2" pageNumber="227" value="2012-02-20">
<date id="FFA6107ECD13FE094E0AFCC65BF6FCE6" box="[514,717,782,807]" pageId="2" pageNumber="227" value="2012-02-20">20 February 2012</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B703CE3CD13FE094ED2FCC65CACFC8A" ID-GBIF-Occurrence="1324332042" collectingDate="2013-06-14" collectionCode="KAUMM" collectorName="S. V. Bogorodsky &amp; T. J. Alpermann" country="Iceland" elevation="8" latitude="26.058432" location="Al Wajh" longLatPrecision="1" longitude="36.64305" pageId="2" pageNumber="227" specimenCode="KAUMM 377" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBBE9EC5CD13FE094ED2FCC65A41FCE6" box="[730,890,783,807]" collectionCode="KAUMM" pageId="2" pageNumber="227">
<collectionCode id="ED09AE7BCD13FE094ED2FCC65A73FCE6" box="[730,840,783,807]" pageId="2" pageNumber="227">KAUMM</collectionCode>
377
</specimenCode>
(KAU13-378), male,
<quantity id="4CE09B5BCD13FE09487CFCC75DE2FCE7" box="[1140,1241,782,807]" metricMagnitude="-2" metricUnit="m" metricValue="2.85" pageId="2" pageNumber="227" unit="mm" value="28.5">28.5 mm</quantity>
SL,
<quantity id="4CE09B5BCD13FE094918FCC65C63FCE6" box="[1296,1368,782,807]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="227" unit="km" value="30.0">30 km</quantity>
south of
<location id="8EC76065CD13FE094CBEFCFB582CFC8A" LSID="urn:lsid:plazi:treatment:03B187A8CD13FE0C4C9FFF5E58D3FA25:8EC76065CD13FE094CBEFCFB582CFC8A" box="[182,279,818,843]" country="Iceland" latitude="26.058432" longLatPrecision="1" longitude="36.64305" name="Al Wajh" pageId="2" pageNumber="227">Al Wajh</location>
,
<geoCoordinate id="EE2C5079CD13FE094D2BFCFB58F1FC8A" box="[291,458,818,843]" direction="north" orientation="latitude" pageId="2" pageNumber="227" precision="1" value="26.058432">N26°03'30.36&quot;</geoCoordinate>
<geoCoordinate id="EE2C5079CD13FE094DDAFCFB5B43FC8A" box="[466,632,818,843]" direction="east" orientation="longitude" pageId="2" pageNumber="227" precision="1" value="36.64305">E36°38'34.98&quot;</geoCoordinate>
, fringing reef,
<elevation id="0035D18DCD13FE094F29FCFA5A74FC8B" box="[801,847,819,843]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="227" unit="m" value="8.0">
<quantity id="4CE09B5BCD13FE094F29FCFA5A74FC8B" box="[801,847,819,843]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="227" unit="m" value="8.0">8 m</quantity>
</elevation>
,
<collectorName id="26ED5368CD13FE094F54FCFA5D20FC8A" box="[860,1051,818,843]" pageId="2" pageNumber="227">S.V. Bogorodsky</collectorName>
&amp;
<collectorName id="26ED5368CD13FE094848FCFA5DCAFC8A" box="[1088,1265,818,843]" pageId="2" pageNumber="227">T.J. Alpermann</collectorName>
,
<collectingDate id="EFE2E996CD13FE0948F5FCFA5CACFC8A" box="[1277,1431,819,843]" pageId="2" pageNumber="227" value="2013-06-14">
<date id="FFA6107ECD13FE0948F5FCFA5CACFC8A" box="[1277,1431,819,843]" pageId="2" pageNumber="227" value="2013-06-14">14 June 2013</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B703CE3CD13FE094C9FFC9E5A56FC52" ID-GBIF-Occurrence="1324332035" collectingDate="2011-04-01" collectionCode="SMF" collectorName="S. V. Bogorodsky" country="Iceland" elevation="1" latitude="16.925917" location="Island" longLatPrecision="1" longitude="41.85447" pageId="2" pageNumber="227" specimenCode="SMF 35757, 2" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBBE9EC5CD13FE094C9FFC9E5802FCAE" box="[151,313,854,879]" pageId="2" pageNumber="227">
<collectionCode id="ED09AE7BCD13FE094C9FFC9E59F4FCAE" box="[151,207,855,879]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="2" pageNumber="227">SMF</collectionCode>
35757, 2
</specimenCode>
males,
<quantity id="4CE09B5BCD13FE094D98FC9E58CEFCAF" box="[400,501,855,879]" metricMagnitude="-2" metricUnit="m" metricValue="2.9699999999999998" pageId="2" pageNumber="227" unit="mm" value="29.7">29.7 mm</quantity>
SL (C&amp;S) &amp;
<quantity id="4CE09B5BCD13FE094E9CFC9E5BC2FCAF" box="[660,761,855,879]" metricMagnitude="-2" metricUnit="m" metricValue="4.7299999999999995" pageId="2" pageNumber="227" unit="mm" value="47.3">47.3 mm</quantity>
SL, Farasan
<collectingCountry id="F30F762ECD13FE094F86FC9F5AEEFCAE" box="[910,981,854,879]" name="Iceland" pageId="2" pageNumber="227">Island</collectingCountry>
,
<geoCoordinate id="EE2C5079CD13FE094FE9FC9F5DB4FCAE" box="[993,1167,854,879]" direction="north" orientation="latitude" pageId="2" pageNumber="227" precision="1" value="16.925917">N16°54' 93.30&quot;</geoCoordinate>
<geoCoordinate id="EE2C5079CD13FE09489FFC9F5C06FCAE" box="[1175,1341,854,879]" direction="east" orientation="longitude" pageId="2" pageNumber="227" precision="1" value="41.85447">E41°50'76.10&quot;</geoCoordinate>
, lagoon, isolated small coral patch,
<elevation id="0035D18DCD13FE094DCEFCB258CFFC53" box="[454,500,891,915]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="227" unit="m" value="1.0">
<quantity id="4CE09B5BCD13FE094DCEFCB258CFFC53" box="[454,500,891,915]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="227" unit="m" value="1.0">1 m</quantity>
</elevation>
,
<collectorName id="26ED5368CD13FE094E08FCB25BFBFC52" box="[512,704,890,915]" pageId="2" pageNumber="227">S.V. Bogorodsky</collectorName>
, 0
<collectingDate id="EFE2E996CD13FE094ED5FCB25A56FC52" box="[733,877,890,915]" pageId="2" pageNumber="227" value="2011-04-01">
<date id="FFA6107ECD13FE094ED5FCB25A56FC52" box="[733,877,890,915]" pageId="2" pageNumber="227" value="2011-04-01">1 April 2011</date>
</collectingDate>
</materialsCitation>
; SMF 35758 (KAU12-0218), male,
<quantity id="4CE09B5BCD13FE094900FCB25C56FC53" box="[1288,1389,891,915]" metricMagnitude="-2" metricUnit="m" metricValue="3.84" pageId="2" pageNumber="227" unit="mm" value="38.4">38.4 mm</quantity>
SL, collected with the holotype;
<materialsCitation id="3B703CE3CD13FE094DE6FC565C6CFC1A" ID-GBIF-Occurrence="1324332034" collectingDate="2012-02-28" collectionCode="SMF" collectorName="S. V. Bogorodsky &amp; T. J. Alpermann" country="Iceland" elevation="4" latitude="16.71805" location="Island" longLatPrecision="1" longitude="42.065567" pageId="2" pageNumber="227" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED09AE7BCD13FE094DE6FC565B1DFC76" box="[494,550,927,951]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="2" pageNumber="227">SMF</collectionCode>
35759 (KAU12-0538), female,
<quantity id="4CE09B5BCD13FE094FAFFC575D2AFC77" box="[935,1041,926,951]" metricMagnitude="-2" metricUnit="m" metricValue="3.5799999999999996" pageId="2" pageNumber="227" unit="mm" value="35.8">35.8 mm</quantity>
SL, Farasan
<collectingCountry id="F30F762ECD13FE0948BEFC575DC6FC76" box="[1206,1277,926,951]" name="Iceland" pageId="2" pageNumber="227">Island</collectingCountry>
,
<geoCoordinate id="EE2C5079CD13FE094907FC575CA5FC76" box="[1295,1438,926,951]" direction="north" orientation="latitude" pageId="2" pageNumber="227" precision="1" value="16.71805">N16°43.083'</geoCoordinate>
<geoCoordinate id="EE2C5079CD13FE094C9FFC0A581EFC1A" box="[151,293,963,987]" direction="east" orientation="longitude" pageId="2" pageNumber="227" precision="1" value="42.065567">E42°03.934'</geoCoordinate>
, isolated coral patch of lagoon,
<elevation id="0035D18DCD13FE094E98FC0B5BE1FC1B" box="[656,730,962,987]" metricMagnitude="0" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="2" pageNumber="227" unit="m" value="4.0" valueMax="5.0" valueMin="3.0">
<quantity id="4CE09B5BCD13FE094E98FC0B5BE1FC1B" box="[656,730,962,987]" metricMagnitude="0" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="2" pageNumber="227" unit="m" value="4.0" valueMax="5.0" valueMin="3.0">35 m</quantity>
</elevation>
,
<collectorName id="26ED5368CD13FE094EE0FC0A5A9CFC1A" box="[744,935,962,987]" pageId="2" pageNumber="227">S.V. Bogorodsky</collectorName>
&amp;
<collectorName id="26ED5368CD13FE094FC5FC0A5D45FC1A" box="[973,1150,962,987]" pageId="2" pageNumber="227">T.J. Alpermann</collectorName>
,
<collectingDate id="EFE2E996CD13FE094882FC0A5C6CFC1A" box="[1162,1367,962,987]" pageId="2" pageNumber="227" value="2012-02-28">
<date id="FFA6107ECD13FE094882FC0A5C6CFC1A" box="[1162,1367,962,987]" pageId="2" pageNumber="227" value="2012-02-28">28 February 2012</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B703CE3CD13FE09496CFC0A5CACFBE2" ID-GBIF-Occurrence="1324332049" collectingDate="2013-07-01" collectionCode="SMF" collectorName="S. V. Bogorodsky" country="Iceland" elevation="15" latitude="21.708967" location="Jeddah" longLatPrecision="1" longitude="39.09643" municipality="Obhur" pageId="2" pageNumber="227" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED09AE7BCD13FE09496CFC0A5CA7FC1A" box="[1380,1436,963,987]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="2" pageNumber="227">SMF</collectionCode>
35760, male, 67.0 mm SL and female,
<quantity id="4CE09B5BCD13FE094E6FFC2F5BF4FC3F" box="[615,719,998,1023]" metricMagnitude="-2" metricUnit="m" metricValue="5.58" pageId="2" pageNumber="227" unit="mm" value="55.8">55.8 mm</quantity>
SL (
<figureCitation id="13232A3BCD13FE094F06FC2F5A6DFC3E" box="[782,854,998,1023]" captionStart="FIGURE 2" captionStartId="4.[151,250,1952,1974]" captionTargetBox="[187,1400,203,1903]" captionTargetId="figure@4.[187,1400,203,1904]" captionTargetPageId="4" captionText="FIGURE 2. Callogobius pilosimentum sp. nov., live coloration: A. SMF 35756, holotype, female, 36.4 mm SL, Farasan Island, Saudi Arabia; B. Fresh coloration, uncatalogued, Farasan Island, Saudi Arabia, illustrating pattern frequently induced by stress; C. SMF 35760, paratype, female, 55.8 mm SL, Obhur, Jeddah, Saudi Arabia. Photos by SVB." httpUri="https://zenodo.org/record/261618/files/figure.png" pageId="2" pageNumber="227">Fig. 2</figureCitation>
C),
<location id="8EC76065CD13FE094F8AFC2F5AE8FC3E" LSID="urn:lsid:plazi:treatment:03B187A8CD13FE0C4C9FFF5E58D3FA25:8EC76065CD13FE094F8AFC2F5AE8FC3E" box="[898,979,998,1023]" country="Iceland" latitude="21.708967" longLatPrecision="1" longitude="39.09643" municipality="Obhur" name="Jeddah" pageId="2" pageNumber="227">Jeddah</location>
,
<collectingMunicipality id="6BC3ACC4CD13FE094FEAFC2F5D12FC3E" box="[994,1065,998,1023]" pageId="2" pageNumber="227">Obhur</collectingMunicipality>
(Sharm Obhur),
<geoCoordinate id="EE2C5079CD13FE0948FDFC2E5CA7FC3E" box="[1269,1436,999,1023]" direction="north" orientation="latitude" pageId="2" pageNumber="227" precision="1" value="21.708967">N21°42'32.28&quot;</geoCoordinate>
<geoCoordinate id="EE2C5079CD13FE094C9FFBC35806FBE2" box="[151,317,1034,1059]" direction="east" orientation="longitude" pageId="2" pageNumber="227" precision="1" value="39.09643">E39°05'47.16&quot;</geoCoordinate>
, steep slope with patches of corals and small sandy flats,
<elevation id="0035D18DCD13FE094FCEFBC35D10FBE3" box="[966,1067,1034,1059]" metricMagnitude="1" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="2" pageNumber="227" unit="m" value="15.0" valueMax="16.0" valueMin="14.0">
<quantity id="4CE09B5BCD13FE094FCEFBC35D10FBE3" box="[966,1067,1034,1059]" metricMagnitude="1" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="2" pageNumber="227" unit="m" value="15.0" valueMax="16.0" valueMin="14.0">1416 m</quantity>
</elevation>
,
<collectorName id="26ED5368CD13FE094830FBC25DCCFBE2" box="[1080,1271,1034,1059]" pageId="2" pageNumber="227">S.V. Bogorodsky</collectorName>
, 0
<collectingDate id="EFE2E996CD13FE09491CFBC25CACFBE2" box="[1300,1431,1034,1059]" pageId="2" pageNumber="227" value="2013-07-01">
<date id="FFA6107ECD13FE09491CFBC25CACFBE2" box="[1300,1431,1034,1059]" pageId="2" pageNumber="227" value="2013-07-01">1 July 2013</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B703CE3CD13FE094C9FFBE75A06FBAA" ID-GBIF-Occurrence="1324332041" collectingDate="1950-12-05" collectionCode="BMNH" collectorName="Suakin &amp; Manihine" country="Sudan" location="Sudan" pageId="2" pageNumber="227" specimenCount="10" specimenCount-female="4" specimenCount-male="1" typeStatus="paratype">
<collectingCountry id="F30F762ECD13FE094C9FFBE759D8FB86" box="[151,227,1070,1095]" name="Sudan" pageId="2" pageNumber="227">
<emphasis id="B96CEAACCD13FE094C9FFBE759D8FB86" bold="true" box="[151,227,1070,1095]" pageId="2" pageNumber="227">Sudan</emphasis>
</collectingCountry>
:
<collectionCode id="ED09AE7BCD13FE094CFCFBE65873FB87" box="[244,328,1071,1094]" pageId="2" pageNumber="227">BMNH</collectionCode>
1978.9.8.12-16,
<specimenCount id="9D1EFD37CD13FE094E04FBE75BA2FB86" box="[524,665,1070,1095]" pageId="2" pageNumber="227" type="generic">5 specimens</specimenCount>
(
<specimenCount id="9D1EFD37CD13FE094EA2FBE65A21FB86" box="[682,794,1070,1095]" pageId="2" pageNumber="227" type="female">4 females</specimenCount>
with 1 C&amp;S,
<specimenCount id="9D1EFD37CD13FE094FB6FBE65D34FB86" box="[958,1039,1070,1095]" pageId="2" pageNumber="227" type="male">1 male</specimenCount>
,
<quantity id="4CE09B5BCD13FE094815FBE75DF9FB87" box="[1053,1218,1070,1095]" metricMagnitude="-2" metricUnit="m" metricValue="5.32" metricValueMax="6.1899999999999995" metricValueMin="4.45" pageId="2" pageNumber="227" unit="mm" value="53.2" valueMax="61.9" valueMin="44.5">44.561.9 mm</quantity>
SL),
<collectorName id="26ED5368CD13FE09490EFBE75C6CFB86" box="[1286,1367,1070,1095]" pageId="2" pageNumber="227">Suakin</collectorName>
, coral crevices in seawall,
<collectorName id="26ED5368CD13FE094D70FB9B58D8FBAA" box="[376,483,1106,1131]" pageId="2" pageNumber="227">Manihine</collectorName>
collections,
<collectingDate id="EFE2E996CD13FE094E7AFB9B5A06FBAA" box="[626,829,1106,1131]" pageId="2" pageNumber="227" value="1950-12-05">
<date id="FFA6107ECD13FE094E7AFB9B5A06FBAA" box="[626,829,1106,1131]" pageId="2" pageNumber="227" value="1950-12-05">5 December 1950</date>
</collectingDate>
</materialsCitation>
.
</paragraph>
<paragraph id="8BA736BECD13FE094CCFFBBF5D4BFB72" blockId="2.[151,1438,422,1923]" pageId="2" pageNumber="227">
<materialsCitation id="3B703CE3CD13FE094CCFFBBF5D4BFB72" ID-GBIF-Occurrence="1324332052" collectingDate="1951-01-12" collectionCode="BMNH" country="Sudan" location="Suakin Archipelago" municipality="Manihine" pageId="2" pageNumber="227" specimenCount="20" specimenCount-female="6" specimenCount-male="4">
<emphasis id="B96CEAACCD13FE094CCFFBBF58BAFB4E" bold="true" box="[199,385,1142,1167]" pageId="2" pageNumber="227">Other material.</emphasis>
<collectionCode id="ED09AE7BCD13FE094D8FFBBE58E0FB4F" box="[391,475,1143,1166]" pageId="2" pageNumber="227">BMNH</collectionCode>
1978.9.8.17-26,
<specimenCount id="9D1EFD37CD13FE094E91FBBE5A0BFB4E" box="[665,816,1142,1167]" pageId="2" pageNumber="227" type="generic">10 specimens</specimenCount>
(
<specimenCount id="9D1EFD37CD13FE094F37FBBF5A95FB4E" box="[831,942,1142,1167]" pageId="2" pageNumber="227" type="female">6 females</specimenCount>
,
<specimenCount id="9D1EFD37CD13FE094FB1FBBE5D2AFB4E" box="[953,1041,1142,1167]" pageId="2" pageNumber="227" type="male">4 males</specimenCount>
,
<quantity id="4CE09B5BCD13FE094813FBBF5D85FB4F" box="[1051,1214,1142,1167]" metricMagnitude="-2" metricUnit="m" metricValue="5.38" metricValueMax="7.07" metricValueMin="3.69" pageId="2" pageNumber="227" unit="mm" value="53.8" valueMax="70.7" valueMin="36.9">36.970.7 mm</quantity>
SL),
<collectingCountry id="F30F762ECD13FE0948F3FBBF5C7FFB4E" box="[1275,1348,1142,1167]" name="Sudan" pageId="2" pageNumber="227">Sudan</collectingCountry>
,
<location id="8EC76065CD13FE094946FBBF581EFB72" LSID="urn:lsid:plazi:treatment:03B187A8CD13FE0C4C9FFF5E58D3FA25:8EC76065CD13FE094946FBBF581EFB72" country="Sudan" municipality="Manihine" name="Suakin Archipelago" pageId="2" pageNumber="227">Suakin Archipelago</location>
, around seawall and from crevices,
<collectingMunicipality id="6BC3ACC4CD13FE094EBEFB535A1AFB72" box="[694,801,1178,1203]" pageId="2" pageNumber="227">Manihine</collectingMunicipality>
collections,
<collectingDate id="EFE2E996CD13FE094FA7FB525D50FB72" box="[943,1131,1178,1203]" pageId="2" pageNumber="227" value="1951-01-12">
<date id="FFA6107ECD13FE094FA7FB525D50FB72" box="[943,1131,1178,1203]" pageId="2" pageNumber="227" value="1951-01-12">12 January 1951</date>
</collectingDate>
.
</materialsCitation>
</paragraph>
<paragraph id="8BA736BECD13FE094CCFFB7758E7FB3A" blockId="2.[151,1438,422,1923]" pageId="2" pageNumber="227">
<materialsCitation id="3B703CE3CD13FE094CCFFB7758E7FB3A" ID-GBIF-Occurrence="1324332054" collectingDate="1969-07-23" collectionCode="USNM" collectorName="Ras Burqa, V. G." country="Egypt" location="Ras Burqa" pageId="2" pageNumber="227" specimenCode="USNM 296956" specimenCount="1" stateProvince="Red Sea">
<emphasis id="B96CEAACCD13FE094CCFFB7758DEFB16" bold="true" box="[199,485,1214,1239]" pageId="2" pageNumber="227">Tentative identification:</emphasis>
<specimenCode id="DBBE9EC5CD13FE094DE4FB765BA7FB16" box="[492,668,1214,1239]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="227">
<collectionCode id="ED09AE7BCD13FE094DE4FB765B06FB16" box="[492,573,1215,1239]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="227">USNM</collectionCode>
296956
</specimenCode>
, female,
<quantity id="4CE09B5BCD13FE094F09FB765A5DFB17" box="[769,870,1215,1239]" metricMagnitude="-2" metricUnit="m" metricValue="4.43" pageId="2" pageNumber="227" unit="mm" value="44.3">44.3 mm</quantity>
SL,
<collectingRegion id="49DCF85CCD13FE094F93FB775AC0FB16" box="[923,1019,1214,1239]" country="Sudan" name="Al Bahr al Ahmar" pageId="2" pageNumber="227">Red Sea</collectingRegion>
,
<collectingCountry id="F30F762ECD13FE09480FFB765D76FB16" box="[1031,1101,1215,1239]" name="Egypt" pageId="2" pageNumber="227">Egypt</collectingCountry>
, just north of
<collectorName id="26ED5368CD13FE0948E1FB765CA6FB16" box="[1257,1437,1215,1239]" pageId="2" pageNumber="227">
<location id="8EC76065CD13FE0948E1FB765C5AFB17" LSID="urn:lsid:plazi:treatment:03B187A8CD13FE0C4C9FFF5E58D3FA25:8EC76065CD13FE0948E1FB765C5AFB17" box="[1257,1377,1215,1238]" country="Egypt" name="Ras Burqa" pageId="2" pageNumber="227" stateProvince="Red Sea">Ras Burqa</location>
, V.G.
</collectorName>
Springer
<emphasis id="B96CEAACCD13FE094CF6FB2A5814FB3A" box="[254,303,1251,1275]" italics="true" pageId="2" pageNumber="227">et al</emphasis>
.,
<collectingDate id="EFE2E996CD13FE094D4CFB2A58E3FB3A" box="[324,472,1250,1275]" pageId="2" pageNumber="227" value="1969-07-23">
<date id="FFA6107ECD13FE094D4CFB2A58E3FB3A" box="[324,472,1250,1275]" pageId="2" pageNumber="227" value="1969-07-23">23 July 1969</date>
</collectingDate>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3026535CD13FE094CCFFACF5C3AFA4A" pageId="2" pageNumber="227" type="diagnosis">
<paragraph id="8BA736BECD13FE094CCFFACF5C3AFA4A" blockId="2.[151,1438,422,1923]" pageId="2" pageNumber="227">
<emphasis id="B96CEAACCD13FE094CCFFACF587AFADE" bold="true" box="[199,321,1286,1311]" pageId="2" pageNumber="227">Diagnosis.</emphasis>
<taxonomicName id="4C184D3DCD13FE094D46FACF5B47FADF" box="[334,636,1286,1310]" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="227" phylum="Chordata" rank="species" species="pilosimentum">
<emphasis id="B96CEAACCD13FE094D46FACF5B47FADF" box="[334,636,1286,1310]" italics="true" pageId="2" pageNumber="227">Callogobius pilosimentum</emphasis>
</taxonomicName>
is distinguished from all other known
<taxonomicName id="4C184D3DCD13FE094853FACF5DDFFADF" box="[1115,1252,1286,1310]" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="227" phylum="Chordata" rank="genus">
<emphasis id="B96CEAACCD13FE094853FACF5DDFFADF" box="[1115,1252,1286,1310]" italics="true" pageId="2" pageNumber="227">Callogobius</emphasis>
</taxonomicName>
species by the following combination of characters: interorbital canal normally containing pores B, C, D, E, F, G and H; preopercular canal containing pores M, N, and O; temporal canal containing pores K and L; scales in lateral series 2125 (usually 24); normally four transverse mandibular papillae rows (Row 16) on each side.
</paragraph>
</subSubSection>
<subSubSection id="C3026535CD13FE0D4CCFFA5F5A44FCBA" lastPageId="6" lastPageNumber="231" pageId="2" pageNumber="227" type="description">
<paragraph id="8BA736BECD13FE094CCFFA5F5C53F9FE" blockId="2.[151,1438,422,1923]" pageId="2" pageNumber="227">
<emphasis id="B96CEAACCD13FE094CCFFA5F5862FA6E" bold="true" box="[199,345,1430,1455]" pageId="2" pageNumber="227">Description.</emphasis>
<typeStatus id="54A3881CCD13FE094D57FA5F58FCFA6E" box="[351,455,1430,1455]" pageId="2" pageNumber="227" type="holotype">Holotype</typeStatus>
values indicated by an asterisk. Parentheses enclose number of
<typeStatus id="54A3881CCD13FE094898FA5E5DFBFA6E" box="[1168,1216,1431,1455]" pageId="2" pageNumber="227">type</typeStatus>
specimens with the particular value, counts made on both sides when applicable or possible. Dorsal fin VI + I,9(17*), VI + I,10(1); anal fin I,6(1), I,7(17*); pectoral-fin rays 15(2), 16(11), 17(15*), 18(5); pelvic-fin rays I,5(36*); segmented caudalfin rays 9 + 8(1),
<date id="FFA6107ECD13FE094D56F9CB584FF9DA" box="[350,372,1538,1563]" pageId="2" pageNumber="227">i8</date>
+ 8(4*),
<date id="FFA6107ECD13FE094DD6F9CB58CFF9DA" box="[478,500,1538,1563]" pageId="2" pageNumber="227">i8</date>
+
<date id="FFA6107ECD13FE094E1BF9CB5B12F9DA" box="[531,553,1538,1563]" pageId="2" pageNumber="227">7i</date>
(12),
<date id="FFA6107ECD13FE094E6EF9CB5BB8F9DA" box="[614,643,1538,1563]" pageId="2" pageNumber="227">ii7</date>
+ 8(1); procurrent rays 4 + 4(2), 5 + 4(15*), 5 + 5(1); scales in lateral series 21(1), 22(3*), 23(4), 24(20*), 25(5); predorsal scales 6(5), 7(13*); transverse scales 8(3), 9(13*), 10(2).
</paragraph>
<paragraph id="8BA736BECD13FE094CCFF9825BB1F9A2" blockId="2.[151,1438,422,1923]" box="[199,650,1610,1635]" pageId="2" pageNumber="227">
See
<tableCitation id="C69A0305CD13FE094CFEF9835872F9A2" box="[246,329,1610,1635]" captionStart="TABLE 1" captionStartId="5.[151,239,152,175]" captionTargetBox="[159,1268,298,858]" captionTargetPageId="5" captionText="TABLE 1. Selected measurements of the holotype and paratypes of C. pilosimentum sp. nov.; standard length is provided in mm, values for other morphometrics are as percentage of standard length (SL) or head length (HL), as indicated. For paratypes, the range is followed by an average value for all types in parentheses. The two largest specimens (63 and 67 mm SL) were males (M), the next 3 largest specimens (55.8, 50.3, 49.3 mm SL) were females (F)." httpUri="http://table.plazi.org/id/DF676636CD14FE0E4C9FFF515CA7FECE" pageId="2" pageNumber="227" tableUuid="DF676636CD14FE0E4C9FFF515CA7FECE">Table 1</tableCitation>
for selected morphometrics.
</paragraph>
<paragraph id="8BA736BECD13FE094CCFF9A75A84F842" blockId="2.[151,1438,422,1923]" pageId="2" pageNumber="227">
Body moderately robust for the genus (
<figureCitation id="13232A3BCD13FE094E92F9A75BDFF946" box="[666,740,1646,1671]" captionStart="FIGURE 2" captionStartId="4.[151,250,1952,1974]" captionTargetBox="[187,1400,203,1903]" captionTargetId="figure@4.[187,1400,203,1904]" captionTargetPageId="4" captionText="FIGURE 2. Callogobius pilosimentum sp. nov., live coloration: A. SMF 35756, holotype, female, 36.4 mm SL, Farasan Island, Saudi Arabia; B. Fresh coloration, uncatalogued, Farasan Island, Saudi Arabia, illustrating pattern frequently induced by stress; C. SMF 35760, paratype, female, 55.8 mm SL, Obhur, Jeddah, Saudi Arabia. Photos by SVB." httpUri="https://zenodo.org/record/261618/files/figure.png" pageId="2" pageNumber="227">Fig. 2</figureCitation>
). Head depressed, broader than deep. Snout obtuse. Mouth slightly oblique, forming an angle of about 45º with body axis; lower jaw slightly beyond tip of upper jaw, posterior end of jaws at or just before vertical through anterior margin of orbit. Anterior nostril moderately long, slender tube, reaching halfway to anterior outer edge of upper lip; posterior nostril very short, upright tube. Eye moderately large; upper margin of orbit slightly elevated above profile of head. Interorbital narrow. Gill opening ending ventrally at lower edge of pectoral-fin base. Tongue broad, tip slightly bilobed. Urogenital papilla long, slender and darkly pigmented in males; broad and darkly pigmented in females, no lateral flaps. Vertebral count (based on two cleared and stained specimens) 10 precaudal + 16 caudal.
</paragraph>
<caption id="DF676636CD12FE084C9FF91E5B21F823" httpUri="https://zenodo.org/record/261617/files/figure.png" pageId="3" pageNumber="228" targetBox="[247,1331,204,1700]" targetPageId="3">
<paragraph id="8BA736BECD12FE084C9FF91E5B21F823" blockId="3.[151,1436,1751,2018]" pageId="3" pageNumber="228">
<emphasis id="B96CEAACCD12FE084C9FF91E582CF92D" bold="true" box="[151,279,1751,1773]" pageId="3" pageNumber="228">FIGURE 1.</emphasis>
Sensory pore and papillae pattern on the head of
<taxonomicName id="4C184D3DCD12FE084F3BF91E5D7AF92C" box="[819,1089,1751,1773]" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="228" phylum="Chordata" rank="species" species="pilosimentum" status="sp. nov.">
<emphasis id="B96CEAACCD12FE084F3BF91E5D7AF92C" box="[819,1089,1751,1773]" italics="true" pageId="3" pageNumber="228">Callogobius pilosimentum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F57D7CD12FE084843F91E5DA4F92D" box="[1099,1183,1751,1772]" pageId="3" pageNumber="228" rank="species">
<emphasis id="B96CEAACCD12FE084843F91E5DA4F92D" bold="true" box="[1099,1183,1751,1772]" pageId="3" pageNumber="228">sp. nov.</emphasis>
</taxonomicNameLabel>
, SMF 35756, holotype, female, 36.4 mm SL.
<emphasis id="B96CEAACCD12FE084D7CF93C58B6F8CB" bold="true" box="[372,397,1781,1802]" pageId="3" pageNumber="228">A.</emphasis>
Lateral view;
<emphasis id="B96CEAACCD12FE084E29F93F5B03F8CA" bold="true" box="[545,568,1782,1803]" pageId="3" pageNumber="228">B.</emphasis>
Ventral view. Letter abbreviations of sensory pores follow
<taxonomicName id="4C184D3DCD12FE08489BF93C5DDBF8CA" authority="Meguro" authorityName="Meguro" authorityYear="1977" box="[1171,1248,1781,1803]" class="Actinopterygii" family="Gobiidae" genus="Akihito" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="228" phylum="Chordata" rank="genus">Akihito</taxonomicName>
&amp; Meguro (1977) and descriptive names are modified from Takagi (1957). Papillae row numbering follows
<taxonomicName id="4C184D3DCD12FE084822F8DD5D4CF8EB" authority="Meguro" authorityName="Meguro" authorityYear="1977" box="[1066,1143,1812,1834]" class="Actinopterygii" family="Gobiidae" genus="Akihito" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="228" phylum="Chordata" rank="genus">Akihito</taxonomicName>
and Meguro and descriptive names are from Delventhal &amp; Mooi (2013). Sensory pores: B = posterior nasal; C = anterior interorbital; D = posterior interorbital; E = supraotic; F = anterior otic; G = posterior otic; H = intertemporal; M,N,O = preopercular; K = anterior temporal; L = posterior temporal. Papillae rows: 2 = postnasal; 9 = anterior suborbital; 10 = mid suborbital; 11 = posterior suborbital; 12 = longitudinal cheek; 13 = transverse cheek; 14 = longitudinal maxillary; 15 = longitudinal mandibular; 16 = transverse mandibular; 17 = postorbital; 20 = preopercular; 21 = transverse opercular (rows 20 and 21 continuous). Scale bar = 2 mm. Illustration by RDM and NRD.
</paragraph>
</caption>
<caption id="DF676636CD15FE0F4C9FF8695D75F832" httpUri="https://zenodo.org/record/261618/files/figure.png" pageId="4" pageNumber="229" targetBox="[187,1400,203,1903]" targetPageId="4">
<paragraph id="8BA736BECD15FE0F4C9FF8695D75F832" blockId="4.[151,1436,1952,2035]" pageId="4" pageNumber="229">
<emphasis id="B96CEAACCD15FE0F4C9FF8695829F874" bold="true" box="[151,274,1952,1974]" pageId="4" pageNumber="229">FIGURE 2.</emphasis>
<taxonomicName id="4C184D3DCD15FE0F4D10F8695B1AF877" box="[280,545,1952,1974]" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="229" phylum="Chordata" rank="species" species="pilosimentum" status="sp. nov.">
<emphasis id="B96CEAACCD15FE0F4D10F8695B1AF877" box="[280,545,1952,1974]" italics="true" pageId="4" pageNumber="229">Callogobius pilosimentum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F57D7CD15FE0F4E2FF8695B4CF874" box="[551,631,1952,1973]" pageId="4" pageNumber="229" rank="species">
<emphasis id="B96CEAACCD15FE0F4E2FF8695B4CF874" bold="true" box="[551,631,1952,1973]" pageId="4" pageNumber="229">sp. nov.</emphasis>
</taxonomicNameLabel>
, live coloration:
<emphasis id="B96CEAACCD15FE0F4F2BF8695A07F874" bold="true" box="[803,828,1952,1973]" pageId="4" pageNumber="229">A.</emphasis>
SMF 35756, holotype, female, 36.4 mm SL, Farasan Island, Saudi Arabia;
<emphasis id="B96CEAACCD15FE0F4D20F876587BF815" bold="true" box="[296,320,1983,2004]" pageId="4" pageNumber="229">B.</emphasis>
Fresh coloration, uncatalogued, Farasan Island, Saudi Arabia, illustrating pattern frequently induced by stress;
<emphasis id="B96CEAACCD15FE0F4C9FF8145994F832" bold="true" box="[151,175,2013,2035]" pageId="4" pageNumber="229">C.</emphasis>
SMF 35760, paratype, female, 55.8 mm SL, Obhur, Jeddah, Saudi Arabia. Photos by SVB.
</paragraph>
</caption>
<caption id="DF676636CD14FE0E4C9FFF515CA7FECE" ID-Table-UUID="DF676636CD14FE0E4C9FFF515CA7FECE" httpUri="http://table.plazi.org/id/DF676636CD14FE0E4C9FFF515CA7FECE" pageId="5" pageNumber="230" targetBox="[159,1268,298,858]" targetIsTable="true" targetPageId="5">
<paragraph id="8BA736BECD14FE0E4C9FFF515CA7FECE" blockId="5.[151,1436,151,271]" pageId="5" pageNumber="230">
<emphasis id="B96CEAACCD14FE0E4C9FFF515834FF6E" bold="true" box="[151,271,152,175]" pageId="5" pageNumber="230">TABLE 1.</emphasis>
Selected measurements of the holotype and paratypes of
<taxonomicName id="4C184D3DCD14FE0E4FAEFF515D60FF6E" box="[934,1115,151,175]" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="230" phylum="Chordata" rank="species" species="pilosimentum" status="sp. nov.">
<emphasis id="B96CEAACCD14FE0E4FAEFF515D60FF6E" box="[934,1115,151,175]" italics="true" pageId="5" pageNumber="230">C. pilosimentum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F57D7CD14FE0E486FFF515DF9FF6E" box="[1127,1218,152,175]" pageId="5" pageNumber="230" rank="species">
<emphasis id="B96CEAACCD14FE0E486FFF515DF9FF6E" bold="true" box="[1127,1218,152,175]" pageId="5" pageNumber="230">sp. nov.</emphasis>
</taxonomicNameLabel>
; standard length is provided in mm, values for other morphometrics are as percentage of standard length (SL) or head length (HL), as indicated. For paratypes, the range is followed by an average value for all types in parentheses. The two largest specimens (63 and 67 mm SL) were males (M), the next 3 largest specimens (55.8, 50.3, 49.3 mm SL) were females (F).
</paragraph>
</caption>
<paragraph id="8BA736BECD14FE0E4EF4FEE35DC8FC9B" pageId="5" pageNumber="230">
<table id="F918C41ECD1401F44C97FEE35DCFFC9B" box="[159,1268,298,858]" gridcols="3" gridrows="14" pageId="5" pageNumber="230">
<tr id="352834FCCD1401F44C97FEE35DCFFE81" box="[159,1268,298,320]" gridrow="0" pageId="5" pageNumber="230" rowspan-0="1">
<th id="76F95D80CD1401F44EF4FEE35A61FE81" box="[764,858,298,320]" gridcol="1" gridrow="0" pageId="5" pageNumber="230">Holotype</th>
<th id="76F95D80CD1401F44843FEE35DCFFE81" box="[1099,1268,298,320]" gridcol="2" gridrow="0" pageId="5" pageNumber="230">Paratypes</th>
</tr>
<tr id="352834FCCD1401F44C97FE9D5DCFFEAB" box="[159,1268,340,362]" gridrow="1" pageId="5" pageNumber="230">
<th id="76F95D80CD1401F44C97FE9D58C9FEAB" box="[159,498,340,362]" gridcol="0" gridrow="1" pageId="5" pageNumber="230">Gender</th>
<td id="76F95D80CD1401F44EF4FE9D5A61FEAB" box="[764,858,340,362]" gridcol="1" gridrow="1" pageId="5" pageNumber="230">F</td>
<td id="76F95D80CD1401F44843FE9D5DCFFEAB" box="[1099,1268,340,362]" gridcol="2" gridrow="1" pageId="5" pageNumber="230">9M,8F</td>
</tr>
<tr id="352834FCCD1401F44C97FEB45DCFFE52" box="[159,1268,381,403]" gridrow="2" pageId="5" pageNumber="230">
<th id="76F95D80CD1401F44C97FEB458C9FE52" box="[159,498,381,403]" gridcol="0" gridrow="2" pageId="5" pageNumber="230">Standard Length (mm)</th>
<td id="76F95D80CD1401F44EF4FEB45A61FE52" box="[764,858,381,403]" gridcol="1" gridrow="2" pageId="5" pageNumber="230">36.4</td>
<td id="76F95D80CD1401F44843FEB45DCFFE52" box="[1099,1268,381,403]" gridcol="2" gridrow="2" pageId="5" pageNumber="230">25.567.0 (42.8)</td>
</tr>
<tr id="352834FCCD1401F44C97FE6F5DCFFE7D" box="[159,1268,422,444]" gridrow="3" pageId="5" pageNumber="230">
<th id="76F95D80CD1401F44C97FE6F58C9FE7D" box="[159,498,422,444]" gridcol="0" gridrow="3" pageId="5" pageNumber="230">Head length (% of SL)</th>
<td id="76F95D80CD1401F44EF4FE6F5A61FE7D" box="[764,858,422,444]" gridcol="1" gridrow="3" pageId="5" pageNumber="230">33.0</td>
<td id="76F95D80CD1401F44843FE6F5DCFFE7D" box="[1099,1268,422,444]" gridcol="2" gridrow="3" pageId="5" pageNumber="230">29.134.1 (32.3)</td>
</tr>
<tr id="352834FCCD1401F44C97FE195DCFFE27" box="[159,1268,464,486]" gridrow="4" pageId="5" pageNumber="230">
<th id="76F95D80CD1401F44C97FE1958C9FE27" box="[159,498,464,486]" gridcol="0" gridrow="4" pageId="5" pageNumber="230">Head depth (% of HL)</th>
<td id="76F95D80CD1401F44EF4FE195A61FE27" box="[764,858,464,486]" gridcol="1" gridrow="4" pageId="5" pageNumber="230">59.2</td>
<td id="76F95D80CD1401F44843FE195DCFFE27" box="[1099,1268,464,486]" gridcol="2" gridrow="4" pageId="5" pageNumber="230">50.664.6 (58.8)</td>
</tr>
<tr id="352834FCCD1401F44C97FE305DCFFDCE" box="[159,1268,505,527]" gridrow="5" pageId="5" pageNumber="230">
<th id="76F95D80CD1401F44C97FE3058C9FDCE" box="[159,498,505,527]" gridcol="0" gridrow="5" pageId="5" pageNumber="230">Head width (% of HL)</th>
<td id="76F95D80CD1401F44EF4FE305A61FDCE" box="[764,858,505,527]" gridcol="1" gridrow="5" pageId="5" pageNumber="230">73.3</td>
<td id="76F95D80CD1401F44843FE305DCFFDCE" box="[1099,1268,505,527]" gridcol="2" gridrow="5" pageId="5" pageNumber="230">64.675.8 (70.2)</td>
</tr>
<tr id="352834FCCD1401F44C97FDEB5DCFFDF9" box="[159,1268,546,568]" gridrow="6" pageId="5" pageNumber="230">
<th id="76F95D80CD1401F44C97FDEB58C9FDF9" box="[159,498,546,568]" gridcol="0" gridrow="6" pageId="5" pageNumber="230">Interorbital width (% of HL)</th>
<td id="76F95D80CD1401F44EF4FDEB5A61FDF9" box="[764,858,546,568]" gridcol="1" gridrow="6" pageId="5" pageNumber="230">5.4</td>
<td id="76F95D80CD1401F44843FDEB5DCFFDF9" box="[1099,1268,546,568]" gridcol="2" gridrow="6" pageId="5" pageNumber="230">4.17.8 (5.7)</td>
</tr>
<tr id="352834FCCD1401F44C97FD855DCFFDA3" box="[159,1268,588,610]" gridrow="7" pageId="5" pageNumber="230">
<th id="76F95D80CD1401F44C97FD8558C9FDA3" box="[159,498,588,610]" gridcol="0" gridrow="7" pageId="5" pageNumber="230">Predorsal fin distance (% of SL)</th>
<td id="76F95D80CD1401F44EF4FD855A61FDA3" box="[764,858,588,610]" gridcol="1" gridrow="7" pageId="5" pageNumber="230">38.5</td>
<td id="76F95D80CD1401F44843FD855DCFFDA3" box="[1099,1268,588,610]" gridcol="2" gridrow="7" pageId="5" pageNumber="230">34.340.4 (37.0)</td>
</tr>
<tr id="352834FCCD1401F44C97FDBC5DCFFD4A" box="[159,1268,629,651]" gridrow="8" pageId="5" pageNumber="230">
<th id="76F95D80CD1401F44C97FDBC58C9FD4A" box="[159,498,629,651]" gridcol="0" gridrow="8" pageId="5" pageNumber="230">Preanal fin distance (% of SL)</th>
<td id="76F95D80CD1401F44EF4FDBC5A61FD4A" box="[764,858,629,651]" gridcol="1" gridrow="8" pageId="5" pageNumber="230">59.6</td>
<td id="76F95D80CD1401F44843FDBC5DCFFD4A" box="[1099,1268,629,651]" gridcol="2" gridrow="8" pageId="5" pageNumber="230">57.862.8 (60.3)</td>
</tr>
<tr id="352834FCCD1401F44C97FD575DCFFD75" box="[159,1268,670,692]" gridrow="9" pageId="5" pageNumber="230">
<th id="76F95D80CD1401F44C97FD5758C9FD75" box="[159,498,670,692]" gridcol="0" gridrow="9" pageId="5" pageNumber="230">Prepelvic fin distance (% of SL)</th>
<td id="76F95D80CD1401F44EF4FD575A61FD75" box="[764,858,670,692]" gridcol="1" gridrow="9" pageId="5" pageNumber="230">33.0</td>
<td id="76F95D80CD1401F44843FD575DCFFD75" box="[1099,1268,670,692]" gridcol="2" gridrow="9" pageId="5" pageNumber="230">29.635.8 (32.8)</td>
</tr>
<tr id="352834FCCD1401F44C97FD015DCFFD1F" box="[159,1268,712,734]" gridrow="10" pageId="5" pageNumber="230">
<th id="76F95D80CD1401F44C97FD0158C9FD1F" box="[159,498,712,734]" gridcol="0" gridrow="10" pageId="5" pageNumber="230">Pectoral-fin length (% of SL)</th>
<td id="76F95D80CD1401F44EF4FD015A61FD1F" box="[764,858,712,734]" gridcol="1" gridrow="10" pageId="5" pageNumber="230">31.3</td>
<td id="76F95D80CD1401F44843FD015DCFFD1F" box="[1099,1268,712,734]" gridcol="2" gridrow="10" pageId="5" pageNumber="230">29.735.4 (32.6)</td>
</tr>
<tr id="352834FCCD1401F44C97FD385DCFFCC6" box="[159,1268,753,775]" gridrow="11" pageId="5" pageNumber="230">
<th id="76F95D80CD1401F44C97FD3858C9FCC6" box="[159,498,753,775]" gridcol="0" gridrow="11" pageId="5" pageNumber="230">Pelvic-fin length (% of SL)</th>
<td id="76F95D80CD1401F44EF4FD385A61FCC6" box="[764,858,753,775]" gridcol="1" gridrow="11" pageId="5" pageNumber="230">23.9</td>
<td id="76F95D80CD1401F44843FD385DCFFCC6" box="[1099,1268,753,775]" gridcol="2" gridrow="11" pageId="5" pageNumber="230">22.028.1 (24.9)</td>
</tr>
<tr id="352834FCCD1401F44C97FCD35DCFFCF1" box="[159,1268,794,816]" gridrow="12" pageId="5" pageNumber="230">
<th id="76F95D80CD1401F44C97FCD358C9FCF1" box="[159,498,794,816]" gridcol="0" gridrow="12" pageId="5" pageNumber="230">Caudal-fin length (% of SL)</th>
<td id="76F95D80CD1401F44EF4FCD35A61FCF1" box="[764,858,794,816]" gridcol="1" gridrow="12" pageId="5" pageNumber="230">32.4</td>
<td id="76F95D80CD1401F44843FCD35DCFFCF1" box="[1099,1268,794,816]" gridcol="2" gridrow="12" pageId="5" pageNumber="230">26.437.3 (31.9)</td>
</tr>
<tr id="352834FCCD1401F44C97FC8D5DCFFC9B" box="[159,1268,836,858]" gridrow="13" pageId="5" pageNumber="230">
<th id="76F95D80CD1401F44C97FC8D58C9FC9B" box="[159,498,836,858]" gridcol="0" gridrow="13" pageId="5" pageNumber="230">Caudal peduncle depth (% of SL)</th>
<td id="76F95D80CD1401F44EF4FC8D5A61FC9B" box="[764,858,836,858]" gridcol="1" gridrow="13" pageId="5" pageNumber="230">13.7</td>
<td id="76F95D80CD1401F44843FC8D5DCFFC9B" box="[1099,1268,836,858]" gridcol="2" gridrow="13" pageId="5" pageNumber="230">12.315.5 (13.8)</td>
</tr>
</table>
</paragraph>
<paragraph id="8BA736BECD14FE0E4CCFFC575C53FBE3" blockId="5.[151,1436,925,2028]" pageId="5" pageNumber="230">
<emphasis id="B96CEAACCD14FE0E4CCFFC575803FC74" box="[199,312,926,949]" italics="true" pageId="5" pageNumber="230">Dentition.</emphasis>
Teeth in jaws conical and slender in rows; outer teeth slightly larger than inner teeth. Outer teeth present on anterior two-fifths of toothed portion of lower jaw; inner teeth of lower jaw tightly packed anteriorly, about 6 irregular rows merging to single row posteriorly. Outer teeth of upper jaw extend over entire toothed portion of upper jaw, about 5 irregular rows of inner teeth merging to single row parallel to row of outer teeth.
</paragraph>
<paragraph id="8BA736BECD14FE0E4CD8FBE75BFFFAA7" blockId="5.[151,1436,925,2028]" pageId="5" pageNumber="230">
<emphasis id="B96CEAACCD14FE0E4CD8FBE75839FB84" box="[208,258,1070,1093]" italics="true" pageId="5" pageNumber="230">Fins</emphasis>
. First dorsal-fin base short; anterior fin membranes incised, first three interspinous membranes incised with spine tips free up to one-fourth their length, occasionally more for second and third spine (particularly in males that have longer, filamentous fin spines), posterior interspinous membrane only slightly incised; second spine longest. Second dorsal-fin base about twice that of first dorsal; segmented rays branched, penultimate ray longest. Anal-fin base short; segmented rays branched, penultimate ray longest. Pectoral-fin rays branched except for dorsalmost one or two rays; fin reaching to level of second or third anal-fin ray. Pelvic fins fully united with membrane over entire length of medial rays; fourth segmented pelvic-fin ray barely shorter or subequal to fifth ray, fifth ray almost reaching to anus; all segmented pelvic-fin rays branched. Pelvic frenum present and well developed. Caudal fin rounded, length &lt;40% SL.
</paragraph>
<paragraph id="8BA736BECD14FE0E4CCFFABB5ABEF9FF" blockId="5.[151,1436,925,2028]" pageId="5" pageNumber="230">
<emphasis id="B96CEAACCD14FE0E4CCFFABB586FFA48" box="[199,340,1394,1417]" italics="true" pageId="5" pageNumber="230">Squamation.</emphasis>
All scales large and deciduous, with distinctly outlined centres. Scales cycloid on head and anterior half of body; cycloid scales present on nape to interorbital, in spaces between papillae rows on cheeks, preoperculum and operculum, on lateral side of pectoral-fin bases, prepelvic and predorsal areas, and belly. Scales ctenoid on posterior half of body from mid-flank region below first to third spines of first dorsal fin. Ctenii numerous, long, slender and pointed; no elongate ctenii on caudal peduncle. Most specimens with single large cycloid scale in centre of pelvic disc (visible under pelvic frenum).
</paragraph>
<paragraph id="8BA736BECD14FE0E4CCFF9805BCFF933" blockId="5.[151,1436,925,2028]" pageId="5" pageNumber="230">
<emphasis id="B96CEAACCD14FE0E4CCFF98058D6F9A0" box="[199,493,1609,1633]" italics="true" pageId="5" pageNumber="230">Cephalic sensory systems.</emphasis>
Pores present (
<figureCitation id="13232A3BCD14FE0E4E92F9805BE7F9A3" box="[666,732,1609,1634]" captionStart="FIGURE 1" captionStartId="3.[151,250,1751,1773]" captionTargetBox="[247,1331,204,1700]" captionTargetId="figure@3.[237,1347,193,1715]" captionTargetPageId="3" captionText="FIGURE 1. Sensory pore and papillae pattern on the head of Callogobius pilosimentum sp. nov., SMF 35756, holotype, female, 36.4 mm SL. A. Lateral view; B. Ventral view. Letter abbreviations of sensory pores follow Akihito &amp; Meguro (1977) and descriptive names are modified from Takagi (1957). Papillae row numbering follows Akihito and Meguro and descriptive names are from Delventhal &amp; Mooi (2013). Sensory pores: B = posterior nasal; C = anterior interorbital; D = posterior interorbital; E = supraotic; F = anterior otic; G = posterior otic; H = intertemporal; M, N, O = preopercular; K = anterior temporal; L = posterior temporal. Papillae rows: 2 = postnasal; 9 = anterior suborbital; 10 = mid suborbital; 11 = posterior suborbital; 12 = longitudinal cheek; 13 = transverse cheek; 14 = longitudinal maxillary; 15 = longitudinal mandibular; 16 = transverse mandibular; 17 = postorbital; 20 = preopercular; 21 = transverse opercular (rows 20 and 21 continuous). Scale bar = 2 mm. Illustration by RDM and NRD." httpUri="https://zenodo.org/record/261617/files/figure.png" pageId="5" pageNumber="230">Fig. 1</figureCitation>
). Interorbital canal with pair of posterior nasal pores (pore B), anterior interorbital pore (pore C), posterior interorbital pore (pore D), pair of supraotic pores (pore E), pair of anterior otic pores (pore F), pair of posterior otic pores (pore G), and pair of intertemporal pores (pore H); preopercular canal with three preopercular pores (pores M, N and O); each temporal canal with anterior and posterior temporal pore (pores K and L, respectively).
</paragraph>
<paragraph id="8BA736BECD14FE0D4CCFF9345DA3FEA2" blockId="5.[151,1436,925,2028]" lastBlockId="6.[151,1437,151,2035]" lastPageId="6" lastPageNumber="231" pageId="5" pageNumber="230">
<emphasis id="B96CEAACCD14FE0E4CCFF9345B3DF8D4" box="[199,518,1789,1813]" italics="true" pageId="5" pageNumber="230">Papillae row configuration</emphasis>
(
<figureCitation id="13232A3BCD14FE0E4E14F9345B5DF8D7" box="[540,614,1789,1814]" captionStart="FIGURE 1" captionStartId="3.[151,250,1751,1773]" captionTargetBox="[247,1331,204,1700]" captionTargetId="figure@3.[237,1347,193,1715]" captionTargetPageId="3" captionText="FIGURE 1. Sensory pore and papillae pattern on the head of Callogobius pilosimentum sp. nov., SMF 35756, holotype, female, 36.4 mm SL. A. Lateral view; B. Ventral view. Letter abbreviations of sensory pores follow Akihito &amp; Meguro (1977) and descriptive names are modified from Takagi (1957). Papillae row numbering follows Akihito and Meguro and descriptive names are from Delventhal &amp; Mooi (2013). Sensory pores: B = posterior nasal; C = anterior interorbital; D = posterior interorbital; E = supraotic; F = anterior otic; G = posterior otic; H = intertemporal; M, N, O = preopercular; K = anterior temporal; L = posterior temporal. Papillae rows: 2 = postnasal; 9 = anterior suborbital; 10 = mid suborbital; 11 = posterior suborbital; 12 = longitudinal cheek; 13 = transverse cheek; 14 = longitudinal maxillary; 15 = longitudinal mandibular; 16 = transverse mandibular; 17 = postorbital; 20 = preopercular; 21 = transverse opercular (rows 20 and 21 continuous). Scale bar = 2 mm. Illustration by RDM and NRD." httpUri="https://zenodo.org/record/261617/files/figure.png" pageId="5" pageNumber="230">Fig. 1</figureCitation>
): Postnasal rows (Row 2) long and joined across midline (16*) or overlapping (1). Anterior suborbital row (Row 9) short to moderately long, not reaching eye (34*). Mid suborbital row (Row 10) short to moderately long, not reaching eye (34*). Posterior suborbital rows (Row 11) long and overlapping (32*) or rarely short or irregular (2). Longitudinal maxillary row (Row 14) rarely irregular (1), normally continuous and extending posteriorly to below and beyond (33*) transverse cheek row (Row 13), latter short (31*) or rarely irregular (1). Longitudinal mandibular row (Row 15) continuous (21*) or irregular (3). With 4(31*), 5(2), or 2(1) transverse mandibular rows (Row 16) on each side of lower jaw, all but one specimen with 4 on at least one side; 15 of
<specimenCount id="9D1EFD37CD17FE0D4DCAFF515B2DFF71" box="[450,534,152,176]" pageId="6" pageNumber="231" type="generic" typeStatus="type">17 type</specimenCount>
specimens with 4 on both sides. One specimen with 5 on one side and 4 on the other (SMF 35760, 67 mm SL), and one with 5 on one side and 2 on the other (SMF 35759,
<quantity id="4CE09B5BCD17FE0D48B3FF725C1AFF12" box="[1211,1313,187,212]" metricMagnitude="-2" metricUnit="m" metricValue="3.5799999999999996" pageId="6" pageNumber="231" unit="mm" value="35.8">35.8 mm</quantity>
SL); latter exhibits irregular patterns in several other papillae rows (Rows 13, 14, 15, 17, 20, 21). Postorbital rows (Row 17) short (less than two-thirds of distance from the dorsal mid-line to bony edge of cranium) (8) or medium (more than two-thirds of distance from dorsal mid-line to bony edge of the cranium) (24*), rarely irregular (1). Preopercular row (Row 20) continuous with the transverse opercular row (Row 21) (25*) or separate (5).
</paragraph>
<paragraph id="8BA736BECD17FE0D4CCFFEA75AFCFCD1" blockId="6.[151,1437,151,2035]" pageId="6" pageNumber="231">
<emphasis id="B96CEAACCD17FE0D4CCFFEA7586BFE47" box="[199,336,366,390]" italics="true" pageId="6" pageNumber="231">Color in life</emphasis>
(
<figureCitation id="13232A3BCD17FE0D4D57FEA7589FFE46" box="[351,420,366,391]" captionStart="FIGURE 2" captionStartId="4.[151,250,1952,1974]" captionTargetBox="[187,1400,203,1903]" captionTargetId="figure@4.[187,1400,203,1904]" captionTargetPageId="4" captionText="FIGURE 2. Callogobius pilosimentum sp. nov., live coloration: A. SMF 35756, holotype, female, 36.4 mm SL, Farasan Island, Saudi Arabia; B. Fresh coloration, uncatalogued, Farasan Island, Saudi Arabia, illustrating pattern frequently induced by stress; C. SMF 35760, paratype, female, 55.8 mm SL, Obhur, Jeddah, Saudi Arabia. Photos by SVB." httpUri="https://zenodo.org/record/261618/files/figure.png" pageId="6" pageNumber="231">Fig. 2</figureCitation>
). General color pattern brown to dark brown with paler patches (
<figureCitation id="13232A3BCD17FE0D4876FEA75DFAFE46" box="[1150,1217,366,391]" captionStart="FIGURE 2" captionStartId="4.[151,250,1952,1974]" captionTargetBox="[187,1400,203,1903]" captionTargetId="figure@4.[187,1400,203,1904]" captionTargetPageId="4" captionText="FIGURE 2. Callogobius pilosimentum sp. nov., live coloration: A. SMF 35756, holotype, female, 36.4 mm SL, Farasan Island, Saudi Arabia; B. Fresh coloration, uncatalogued, Farasan Island, Saudi Arabia, illustrating pattern frequently induced by stress; C. SMF 35760, paratype, female, 55.8 mm SL, Obhur, Jeddah, Saudi Arabia. Photos by SVB." httpUri="https://zenodo.org/record/261618/files/figure.png" pageId="6" pageNumber="231">Fig. 2</figureCitation>
A), or pale grayishbrown with up to five dark brown mottled vertical bars on body and dark brown blotchy patterns with varying contrast on head. Body bars, when visible (e.g.
<figureCitation id="13232A3BCD17FE0D4EAFFE7C5BC8FE0F" box="[679,755,437,462]" captionStart="FIGURE 2" captionStartId="4.[151,250,1952,1974]" captionTargetBox="[187,1400,203,1903]" captionTargetId="figure@4.[187,1400,203,1904]" captionTargetPageId="4" captionText="FIGURE 2. Callogobius pilosimentum sp. nov., live coloration: A. SMF 35756, holotype, female, 36.4 mm SL, Farasan Island, Saudi Arabia; B. Fresh coloration, uncatalogued, Farasan Island, Saudi Arabia, illustrating pattern frequently induced by stress; C. SMF 35760, paratype, female, 55.8 mm SL, Obhur, Jeddah, Saudi Arabia. Photos by SVB." httpUri="https://zenodo.org/record/261618/files/figure.png" pageId="6" pageNumber="231">Figs. 2</figureCitation>
B &amp; C, but not
<figureCitation id="13232A3BCD17FE0D4F95FE7C5AE4FE0F" box="[925,991,437,462]" captionStart="FIGURE 2" captionStartId="4.[151,250,1952,1974]" captionTargetBox="[187,1400,203,1903]" captionTargetId="figure@4.[187,1400,203,1904]" captionTargetPageId="4" captionText="FIGURE 2. Callogobius pilosimentum sp. nov., live coloration: A. SMF 35756, holotype, female, 36.4 mm SL, Farasan Island, Saudi Arabia; B. Fresh coloration, uncatalogued, Farasan Island, Saudi Arabia, illustrating pattern frequently induced by stress; C. SMF 35760, paratype, female, 55.8 mm SL, Obhur, Jeddah, Saudi Arabia. Photos by SVB." httpUri="https://zenodo.org/record/261618/files/figure.png" pageId="6" pageNumber="231">Fig. 2</figureCitation>
A), appear as follows: irregular bar may extend dorsally over operculum and pectoral-fin base, slightly indented at first dorsal-fin spine; wide bar extends below first dorsal fin; two wide, slightly oblique bars extend below mid and posterior portions of second dorsal fin; narrow bar encircles posterior caudal peduncle, edge of hypural plate and proximal regions of the caudal fin. Underside of head light brown or dusky, papillae rows may show great color contrast. First dorsal fin with dark brown basal spot centered on fourth ray and continuous with dark body bar; may exhibit yellow markings on distal margin. First and second dorsal fins dark brown with irregular, oblique, pale bands or rows of spots, some individuals with irregular dark and pale pattern; caudal fin dark brown usually with pale spots forming irregular bars. Pelvic and anal fins dusky or dark brown with or without paler blotches. Pectoral fin with dark irregular bands or rows of spots and a dark medial mark at base. Urogenital papilla dark.
</paragraph>
<paragraph id="8BA736BECD17FE0D4CCFFCD25A44FCBA" blockId="6.[151,1437,151,2035]" pageId="6" pageNumber="231">
<emphasis id="B96CEAACCD17FE0D4CCFFCD25881FCF2" box="[199,442,795,819]" italics="true" pageId="6" pageNumber="231">Color in preservative</emphasis>
. As in life, although older specimens can be orange-brown with little pattern on body. Most museum specimens damaged (species with fragile skin and deciduous scales) and body color pattern often reduced to light brown or grey with dark outlines of scale pockets.
</paragraph>
</subSubSection>
<subSubSection id="C3026535CD17FE0D4CCFFC4F5B95FC27" pageId="6" pageNumber="231" type="etymology">
<paragraph id="8BA736BECD17FE0D4CCFFC4F5B95FC27" blockId="6.[151,1437,151,2035]" pageId="6" pageNumber="231">
<emphasis id="B96CEAACCD17FE0D4CCFFC4F5875FC5E" bold="true" box="[199,334,902,927]" pageId="6" pageNumber="231">Etymology.</emphasis>
The species name is derived from the Latin
<emphasis id="B96CEAACCD17FE0D4F37FC4F5AB5FC5F" box="[831,910,902,926]" italics="true" pageId="6" pageNumber="231">pilosus</emphasis>
meaning “hairy” and
<emphasis id="B96CEAACCD17FE0D488DFC4E5DE6FC5F" box="[1157,1245,903,926]" italics="true" pageId="6" pageNumber="231">mentum</emphasis>
meaning “chin”, referring to the extra rows of papillae on the chin. Specific epithet to be treated as a noun in apposition. Suggested common name: hairy-chinned flapheaded goby.
</paragraph>
</subSubSection>
<subSubSection id="C3026535CD17FE0D4CCFFC385B2AFB90" pageId="6" pageNumber="231" type="distribution">
<paragraph id="8BA736BECD17FE0D4CCFFC385B2AFB90" blockId="6.[151,1437,151,2035]" pageId="6" pageNumber="231">
<emphasis id="B96CEAACCD17FE0D4CCFFC3858CFFBCB" bold="true" box="[199,500,1009,1034]" pageId="6" pageNumber="231">Distribution and habitat.</emphasis>
Restricted to the
<collectingRegion id="49DCF85CCD17FE0D4EB6FC385A1BFBCB" box="[702,800,1009,1034]" country="Sudan" name="Al Bahr al Ahmar" pageId="6" pageNumber="231">Red Sea</collectingRegion>
. It is usually collected in shelters or close to shelters on small sand flats (about
<quantity id="4CE09B5BCD17FE0D4D92FBDC58CDFBEC" box="[410,502,1045,1070]" metricMagnitude="-1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="6" pageNumber="231" unit="m" value="0.75" valueMax="1.0" valueMin="0.5">0.51 m</quantity>
across) of steep slopes, sometimes at the base of coral reefs, in closed areas in bays and lagoons, at depths of
<quantity id="4CE09B5BCD17FE0D4DBDFBF05B36FB90" box="[437,525,1081,1105]" metricMagnitude="1" metricUnit="m" metricValue="1.05" metricValueMax="2.0" metricValueMin="0.1" pageId="6" pageNumber="231" unit="m" value="10.5" valueMax="20.0" valueMin="1.0">
1
<specimenCount id="9D1EFD37CD17FE0D4DC6FBF05B36FB90" box="[462,525,1081,1105]" pageId="6" pageNumber="231" type="generic">20 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C3026535CD17FE0D4CCFFB9558A5FA8A" pageId="6" pageNumber="231" type="discussion">
<paragraph id="8BA736BECD17FE0D4CCFFB9558A5FA8A" blockId="6.[151,1437,151,2035]" pageId="6" pageNumber="231">
<emphasis id="B96CEAACCD17FE0D4CCFFB955800FBB4" bold="true" box="[199,315,1116,1141]" pageId="6" pageNumber="231">Remarks.</emphasis>
<bibRefCitation id="EF894B4FCD17FE0D4D4FFB9458DEFBB4" author="Goren" box="[327,485,1117,1141]" pageId="6" pageNumber="231" refString="Goren, M. (1980) Red Sea fishes assigned to the genus Callogobius Bleeker with a description of a new species (Teleostei: Gobiidae). Israel Journal of Zoology, 28, 209 - 217. [for 1979]" type="journal article" year="1980">Goren (1980)</bibRefCitation>
listed
<taxonomicName id="4C184D3DCD17FE0D4E33FB945A75FBB4" authority="Smith" authorityName="Smith" box="[571,846,1116,1141]" class="Actinopterygii" family="Gobiidae" genus="Drombus" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="231" phylum="Chordata" rank="species" species="irrasus">
<emphasis id="B96CEAACCD17FE0D4E33FB945BC5FBB5" box="[571,766,1117,1141]" italics="true" pageId="6" pageNumber="231">Drombus irrasus</emphasis>
Smith
</taxonomicName>
in the Red Sea as a first record. This species is considered a synonym of
<taxonomicName id="4C184D3DCD17FE0D4DBFFB495BE3FB59" box="[439,728,1152,1176]" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="231" phylum="Chordata" rank="species" species="maculipinnis">
<emphasis id="B96CEAACCD17FE0D4DBFFB495BE3FB59" box="[439,728,1152,1176]" italics="true" pageId="6" pageNumber="231">Callogobius maculipinnis</emphasis>
</taxonomicName>
by most authors (e.g.
<bibRefCitation id="EF894B4FCD17FE0D4FDAFB495C37FB58" author="McKinney" box="[978,1292,1152,1177]" pageId="6" pageNumber="231" refString="McKinney, J. F. &amp; Lachner, E. A. (1984) Callogobius crassus, a new fish from the Indo-Pacific region (Teleostei: Gobiidae). Proceedings of the Biological Society of Washington, 97 (3), 627 - 631." type="journal article" year="1984">McKinney &amp; Lachner 1984</bibRefCitation>
,
<bibRefCitation id="EF894B4FCD17FE0D491FFB4859E8FB7C" author="Goren" pageId="6" pageNumber="231" refString="Goren, M., Miroz, A. &amp; Baranes, A. (1991) Callogobius amikami, a new species of goby (Gobiidae) from the Red Sea. Cybium, 15 (4), 299 - 302." type="journal article" year="1991">
Goren
<emphasis id="B96CEAACCD17FE0D496CFB485CAEFB59" box="[1380,1429,1152,1176]" italics="true" pageId="6" pageNumber="231">et al</emphasis>
. 1991
</bibRefCitation>
);
<bibRefCitation id="EF894B4FCD17FE0D4CEEFB6D5B25FB7C" author="Eschmeyer" box="[230,542,1188,1213]" pageId="6" pageNumber="231" refString="Eschmeyer, W. N. &amp; Fricke, R. (Eds.) (2016) Catalog of Fishes: Genera, Species, References. California Academy of Sciences, San Francisco. Available from: http: // researcharchive. calacademy. org / research / ichthyology / catalog / fishcatmain. asp (accessed 5 January 2016)" type="journal article" year="2016">Eschmeyer &amp; Fricke (2016)</bibRefCitation>
incorrectly attributed
<bibRefCitation id="EF894B4FCD17FE0D4F1DFB6D5D7BFB7C" author="Delventhal" box="[789,1088,1188,1213]" pageId="6" pageNumber="231" refString="Delventhal, N. R. &amp; Mooi, R. D. (2013) Callogobius winterbottomi, a new species of goby (Teleostei: Gobiidae) from the Western Indian Ocean. Zootaxa, 3630 (1), 155 - 164. http: // dx. doi. org / 10.11646 / zootaxa. 3630.1.6" type="journal article" year="2013">Delventhal &amp; Mooi (2013)</bibRefCitation>
as recognizing Smiths species when the type was listed as comparative material. Although we have not examined Gorens (1980)
<taxonomicName id="4C184D3DCD17FE0D4920FB015CA0FB1E" box="[1320,1435,1224,1247]" class="Actinopterygii" family="Gobiidae" genus="Drombus" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="231" phylum="Chordata" rank="species" species="irrasus">
<emphasis id="B96CEAACCD17FE0D4920FB015CA0FB1E" box="[1320,1435,1224,1247]" italics="true" pageId="6" pageNumber="231">C. irrasus</emphasis>
</taxonomicName>
material, we suspect that these specimens will belong to
<taxonomicName id="4C184D3DCD17FE0D4F04FB255AF8FAC5" box="[780,963,1260,1284]" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="231" phylum="Chordata" rank="species" species="pilosimentum">
<emphasis id="B96CEAACCD17FE0D4F04FB255AF8FAC5" box="[780,963,1260,1284]" italics="true" pageId="6" pageNumber="231">C. pilosimentum</emphasis>
</taxonomicName>
. Two lots of specimens that we identify as
<taxonomicName id="4C184D3DCD17FE0D4C9FFAD9586EFAE6" box="[151,341,1295,1319]" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="231" phylum="Chordata" rank="species" species="pilosimentum">
<emphasis id="B96CEAACCD17FE0D4C9FFAD9586EFAE6" box="[151,341,1295,1319]" italics="true" pageId="6" pageNumber="231">C. pilosimentum</emphasis>
</taxonomicName>
(BMNH 1978.9.8.12-16 and BMNH 1978.9.8.17-26) were examined by
<bibRefCitation id="EF894B4FCD17FE0D48C0FAD95C5DFAE9" author="Goren" box="[1224,1382,1296,1320]" pageId="6" pageNumber="231" refString="Goren, M. (1980) Red Sea fishes assigned to the genus Callogobius Bleeker with a description of a new species (Teleostei: Gobiidae). Israel Journal of Zoology, 28, 209 - 217. [for 1979]" type="journal article" year="1980">Goren (1980)</bibRefCitation>
and identified as
<taxonomicName id="4C184D3DCD17FE0D4D2FFAFA58ACFA8A" box="[295,407,1331,1355]" class="Actinopterygii" family="Gobiidae" genus="Drombus" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="231" phylum="Chordata" rank="species" species="irrasus">
<emphasis id="B96CEAACCD17FE0D4D2FFAFA58ACFA8A" box="[295,407,1331,1355]" italics="true" pageId="6" pageNumber="231">C. irrasus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3026535CD17FE0D4CCFFA9E5BF7F833" pageId="6" pageNumber="231" type="reference_group">
<paragraph id="8BA736BECD17FE0D4CCFFA9E5BF7F833" blockId="6.[151,1437,151,2035]" pageId="6" pageNumber="231">
<taxonomicName id="4C184D3DCD17FE0D4CCFFA9E58CFFAAE" ID-CoL="5X7R7" box="[199,500,1367,1391]" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="231" phylum="Chordata" rank="species" species="pilosimentum">
<emphasis id="B96CEAACCD17FE0D4CCFFA9E58CFFAAE" box="[199,500,1367,1391]" italics="true" pageId="6" pageNumber="231">Callogobius pilosimentum</emphasis>
</taxonomicName>
belongs to a “
<taxonomicName id="4C184D3DCD17FE0D4EA4FA9E5A06FAAE" ID-CoL="PV9Y" box="[684,829,1367,1391]" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="231" phylum="Chordata" rank="species" species="maculipinnis">
<emphasis id="B96CEAACCD17FE0D4EA4FA9E5A06FAAE" box="[684,829,1367,1391]" italics="true" pageId="6" pageNumber="231">maculipinnis</emphasis>
</taxonomicName>
species complex” including the following nominal species:
<taxonomicName id="4C184D3DCD17FE0D4CF3FAB25B10FA52" ID-CoL="PV9H" authority="Goren 1979" authorityName="Goren" authorityYear="1979" box="[251,555,1403,1427]" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="231" phylum="Chordata" rank="species" species="bauchotae">
<emphasis id="B96CEAACCD17FE0D4CF3FAB258AFFA52" box="[251,404,1403,1427]" italics="true" pageId="6" pageNumber="231">C. bauchotae</emphasis>
Goren 1979
</taxonomicName>
,
<taxonomicName id="4C184D3DCD17FE0D4E32FAB25A6FFA52" ID-CoL="37PLZ" authority="Smith 1959" authorityName="Smith" authorityYear="1959" box="[570,852,1402,1427]" class="Actinopterygii" family="Gobiidae" genus="Drombus" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="231" phylum="Chordata" rank="species" species="irrasus">
<emphasis id="B96CEAACCD17FE0D4E32FAB25B95FA53" box="[570,686,1403,1426]" italics="true" pageId="6" pageNumber="231">C. irrasus</emphasis>
(
<bibRefCitation id="EF894B4FCD17FE0D4EC9FAB35A76FA52" author="Smith" box="[705,845,1402,1427]" pageId="6" pageNumber="231" refString="Smith, J. L. B. (1959) Gobioid fishes of the families Gobiidae, Periophthalmidae, Trypauchenidae, Taenioididae and Kraemeriidae of the western Indian Ocean. Ichthyological Bulletin, Department of Ichthyology, Rhodes University, 13, 185 - 225, pls. 9 - 13." type="journal article" year="1959">Smith 1959</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C184D3DCD17FE0D4F6CFAB25D49FA52" ID-CoL="3PSW8" authority="Herre 1943" authorityName="Herre" authorityYear="1943" box="[868,1138,1403,1427]" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="231" phylum="Chordata" rank="species" species="kuderi">
<emphasis id="B96CEAACCD17FE0D4F6CFAB25AEAFA52" box="[868,977,1403,1427]" italics="true" pageId="6" pageNumber="231">C. kuderi</emphasis>
(
<bibRefCitation id="EF894B4FCD17FE0D4FEAFAB25D51FA52" author="Herre" box="[994,1130,1403,1427]" pageId="6" pageNumber="231" refString="Herre, A. W. C. T. (1943) Notes on fishes in the Zoological Museum of Stanford University. XI. Two new genera and species. With key to the genera of gobies with vomerine teeth. Proceedings of the Biological Society of Washington, 56, 91 - 95." type="journal article" year="1943">Herre 1943</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C184D3DCD17FE0D488BFAB259E1FA76" ID-CoL="PV9Y" authority="Fowler 1918" authorityName="Fowler" authorityYear="1918" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="231" phylum="Chordata" rank="species" species="maculipinnis">
<emphasis id="B96CEAACCD17FE0D488BFAB25C03FA52" box="[1155,1336,1403,1427]" italics="true" pageId="6" pageNumber="231">C. maculipinnis</emphasis>
(
<bibRefCitation id="EF894B4FCD17FE0D4942FAB359E9FA76" author="Fowler" pageId="6" pageNumber="231" refString="Fowler, H. W. (1918) New and little-known fishes from the Philippine Islands. Proceedings of the Academy of Natural Sciences of Philadelphia, 70, 2 - 71." type="journal article" year="1918">Fowler 1918</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C184D3DCD17FE0D4CE3FA565B86FA76" ID-CoL="5WW2V" authority="Chen &amp; Shao 2000" authorityName="Chen &amp; Shao" authorityYear="2000" box="[235,701,1438,1463]" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="231" phylum="Chordata" rank="species" species="nigromarginatus">
<emphasis id="B96CEAACCD17FE0D4CE3FA5658F1FA77" box="[235,458,1439,1462]" italics="true" pageId="6" pageNumber="231">C. nigromarginatus</emphasis>
<bibRefCitation id="EF894B4FCD17FE0D4DDEFA575B86FA76" author="Chen" box="[470,701,1438,1463]" pageId="6" pageNumber="231" refString="Chen, J. - P. &amp; Shao, K. - T. (2000) Callogobius nigromarginatus, a new species of goby (Pisces: Gobiidae) from Taiwan. Bulletin of Marine Science, 66, 457 - 466." type="journal article" year="2000">Chen &amp; Shao 2000</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C184D3DCD17FE0D4EC5FA565AD5FA76" authority="Takagi 1957" authorityName="Takagi" authorityYear="1957" box="[717,1006,1438,1463]" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="231" phylum="Chordata" rank="species" species="shunkan">
<emphasis id="B96CEAACCD17FE0D4EC5FA565A74FA77" box="[717,847,1438,1462]" italics="true" pageId="6" pageNumber="231">C. shunkan</emphasis>
<bibRefCitation id="EF894B4FCD17FE0D4F54FA575AD5FA76" author="Takagi" box="[860,1006,1438,1463]" pageId="6" pageNumber="231" refString="Takagi, K. (1957) Descriptions of some new gobioid fishes of Japan, with a proposition on the sensory line system as a taxonomic appliance. Journal of the Tokyo University of Fisheries, 43 (1), 97 - 126." type="journal article" year="1957">Takagi 1957</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C184D3DCD17FE0D4FF6FA565C05FA76" ID-CoL="6982V" authority="Koumans 1953" authorityName="Koumans" authorityYear="1953" box="[1022,1342,1438,1463]" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="231" phylum="Chordata" rank="species" species="snelliusi">
<emphasis id="B96CEAACCD17FE0D4FF6FA565DBBFA77" box="[1022,1152,1438,1462]" italics="true" pageId="6" pageNumber="231">C. snelliusi</emphasis>
<bibRefCitation id="EF894B4FCD17FE0D4884FA565C05FA76" author="Koumans" box="[1164,1342,1438,1463]" pageId="6" pageNumber="231" refString="Koumans, F. P. (1953) Biological results of the Snellius expedition. XVI. The Pisces and Leptocardii of the Snellius expedition. Temminckia, 9, 177 - 275." type="journal article" year="1953">Koumans 1953</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C184D3DCD17FE0D498AFA5658A0FA1A" ID-CoL="3PSW9" authority="Herre 1950" authorityName="Herre" authorityYear="1950" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="231" phylum="Chordata" rank="species" species="vanclevei">
<emphasis id="B96CEAACCD17FE0D498AFA56583AFA1B" italics="true" pageId="6" pageNumber="231">C. vanclevei</emphasis>
(
<bibRefCitation id="EF894B4FCD17FE0D4D06FA0B58A9FA1A" author="Herre" box="[270,402,1474,1499]" pageId="6" pageNumber="231" refString="Herre, A. W. C. T. (1950) Two new gobies from the Philippines with notes on a third rare goby. Proceedings of the Biological Society of Washington, 63, 73 - 76." type="journal article" year="1950">Herre 1950</bibRefCitation>
)
</taxonomicName>
. Species of this complex are characterized by having the temporal canal with pores K and L (absent in all other
<taxonomicName id="4C184D3DCD17FE0D4D71FA2F5B39FA3F" ID-CoL="62L29" box="[377,514,1510,1534]" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="231" phylum="Chordata" rank="genus">
<emphasis id="B96CEAACCD17FE0D4D71FA2F5B39FA3F" box="[377,514,1510,1534]" italics="true" pageId="6" pageNumber="231">Callogobius</emphasis>
</taxonomicName>
) and normally four or fewer transverse mandibular rows on each side (all other
<taxonomicName id="4C184D3DCD17FE0D4C9FF9C0581BF9E0" ID-CoL="62L29" box="[151,288,1545,1569]" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="231" phylum="Chordata" rank="genus">
<emphasis id="B96CEAACCD17FE0D4C9FF9C0581BF9E0" box="[151,288,1545,1569]" italics="true" pageId="6" pageNumber="231">Callogobius</emphasis>
</taxonomicName>
normally have more than 10).
<bibRefCitation id="EF894B4FCD17FE0D4E82F9C05AE7F9E3" author="McKinney" box="[650,988,1545,1570]" pageId="6" pageNumber="231" refString="McKinney, J. F. &amp; Lachner, E. A. (1984) Callogobius crassus, a new fish from the Indo-Pacific region (Teleostei: Gobiidae). Proceedings of the Biological Society of Washington, 97 (3), 627 - 631." type="journal article" year="1984">McKinney &amp; Lachner (1984)</bibRefCitation>
synonymised
<taxonomicName id="4C184D3DCD17FE0D488BF9C35DCEF9E0" ID-CoL="37PLZ" box="[1155,1269,1546,1569]" class="Actinopterygii" family="Gobiidae" genus="Drombus" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="231" phylum="Chordata" rank="species" species="irrasus">
<emphasis id="B96CEAACCD17FE0D488BF9C35DCEF9E0" box="[1155,1269,1546,1569]" italics="true" pageId="6" pageNumber="231">C. irrasus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C184D3DCD17FE0D490EF9C35C4AF9E0" ID-CoL="3PSW8" box="[1286,1393,1545,1569]" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="231" phylum="Chordata" rank="species" species="kuderi">
<emphasis id="B96CEAACCD17FE0D490EF9C35C4AF9E0" box="[1286,1393,1545,1569]" italics="true" pageId="6" pageNumber="231">C. kuderi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C184D3DCD17FE0D498AF9C359CFF984" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="231" phylum="Chordata" rank="species" species="shunkan">
<emphasis id="B96CEAACCD17FE0D498AF9C359CFF984" italics="true" pageId="6" pageNumber="231">C. shunkan</emphasis>
</taxonomicName>
,
<taxonomicName id="4C184D3DCD17FE0D4D02F9E758ABF984" ID-CoL="6982V" box="[266,400,1581,1605]" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="231" phylum="Chordata" rank="species" species="snelliusi">
<emphasis id="B96CEAACCD17FE0D4D02F9E758ABF984" box="[266,400,1581,1605]" italics="true" pageId="6" pageNumber="231">C. snelliusi</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C184D3DCD17FE0D4DD6F9E75B4AF984" ID-CoL="3PSW9" box="[478,625,1581,1605]" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="231" phylum="Chordata" rank="species" species="vanclevei">
<emphasis id="B96CEAACCD17FE0D4DD6F9E75B4AF984" box="[478,625,1581,1605]" italics="true" pageId="6" pageNumber="231">C. vanclevei</emphasis>
</taxonomicName>
with
<taxonomicName id="4C184D3DCD17FE0D4EC9F9E75A40F984" ID-CoL="PV9Y" box="[705,891,1581,1605]" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="231" phylum="Chordata" rank="species" species="maculipinnis">
<emphasis id="B96CEAACCD17FE0D4EC9F9E75A40F984" box="[705,891,1581,1605]" italics="true" pageId="6" pageNumber="231">C. maculipinnis</emphasis>
</taxonomicName>
; they did not examine
<taxonomicName id="4C184D3DCD17FE0D48ADF9E75C78F984" ID-CoL="PV9H" box="[1189,1347,1581,1605]" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="231" phylum="Chordata" rank="species" species="bauchotae">
<emphasis id="B96CEAACCD17FE0D48ADF9E75C78F984" box="[1189,1347,1581,1605]" italics="true" pageId="6" pageNumber="231">C. bauchotae</emphasis>
</taxonomicName>
(or
<taxonomicName id="4C184D3DCD17FE0D498AF9E7586AF9A8" ID-CoL="5WW2V" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="231" phylum="Chordata" rank="species" species="nigromarginatus">
<emphasis id="B96CEAACCD17FE0D498AF9E7586AF9A8" italics="true" pageId="6" pageNumber="231">C. nigromarginatus</emphasis>
</taxonomicName>
as it was described after their work). However, their synonymies should be considered questionable because all included species are poorly defined. Based on the first authors examination of the
<typeStatus id="54A3881CCD17FE0D4952F9BC5CA3F94C" box="[1370,1432,1653,1677]" pageId="6" pageNumber="231">types</typeStatus>
, we are confident that several, or even most, of these species will prove to be valid after complete investigation. Despite this,
<taxonomicName id="4C184D3DCD17FE0D4D25F97458DDF915" ID-CoL="5X7R7" box="[301,486,1724,1748]" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="231" phylum="Chordata" rank="species" species="pilosimentum">
<emphasis id="B96CEAACCD17FE0D4D25F97458DDF915" box="[301,486,1724,1748]" italics="true" pageId="6" pageNumber="231">C. pilosimentum</emphasis>
</taxonomicName>
can readily be distinguished from the other members of this complex, all of which normally have three transverse mandibular rows on each side. We previously examined the
<typeStatus id="54A3881CCD17FE0D48A0F9285DD8F938" box="[1192,1251,1761,1785]" pageId="6" pageNumber="231">types</typeStatus>
of all the above species in the
<taxonomicName id="4C184D3DCD17FE0D4D3EF8CD58FDF8DD" ID-CoL="PV9Y" box="[310,454,1796,1820]" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="231" phylum="Chordata" rank="species" species="maculipinnis">
<emphasis id="B96CEAACCD17FE0D4D3EF8CD58FDF8DD" box="[310,454,1796,1820]" italics="true" pageId="6" pageNumber="231">maculipinnis</emphasis>
</taxonomicName>
complex except
<emphasis id="B96CEAACCD17FE0D4E80F8CD5A50F8DA" box="[648,875,1796,1819]" italics="true" pageId="6" pageNumber="231">
<taxonomicName id="4C184D3DCD17FE0D4E80F8CD5A5CF8DA" ID-CoL="5WW2V" box="[648,871,1796,1819]" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="231" phylum="Chordata" rank="species" species="nigromarginatus">C. nigromarginatus</taxonomicName>
.
</emphasis>
They all have three sets of transverse mandibular rows, except
<taxonomicName id="4C184D3DCD17FE0D4D27F8E15894F8FE" box="[303,431,1831,1855]" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="231" phylum="Chordata" rank="species" species="shunkan">
<emphasis id="B96CEAACCD17FE0D4D27F8E15894F8FE" box="[303,431,1831,1855]" italics="true" pageId="6" pageNumber="231">C. shunkan</emphasis>
</taxonomicName>
which has extra irregular rows and is easily differentiated by other characters (Y. Ikeda pers. comm.). These observations of the
<typeStatus id="54A3881CCD17FE0D4E54F8855BACF8A5" box="[604,663,1868,1892]" pageId="6" pageNumber="231">types</typeStatus>
were recently confirmed by Y. Ikeda (
<taxonomicName id="4C184D3DCD17FE0D484DF8825DE1F8A2" ID-CoL="PV9H" box="[1093,1242,1867,1891]" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="231" phylum="Chordata" rank="species" species="bauchotae">
<emphasis id="B96CEAACCD17FE0D484DF8825DE1F8A2" box="[1093,1242,1867,1891]" italics="true" pageId="6" pageNumber="231">C. bauchotae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C184D3DCD17FE0D48E0F8825C5EF8A2" ID-CoL="6982V" box="[1256,1381,1867,1891]" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="231" phylum="Chordata" rank="species" species="snelliusi">
<emphasis id="B96CEAACCD17FE0D48E0F8825C5EF8A2" box="[1256,1381,1867,1891]" italics="true" pageId="6" pageNumber="231">C. snelliusi</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C184D3DCD17FE0D4C9FF8A6582EF846" box="[151,277,1903,1927]" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="231" phylum="Chordata" rank="species" species="shunkan">
<emphasis id="B96CEAACCD17FE0D4C9FF8A6582EF846" box="[151,277,1903,1927]" italics="true" pageId="6" pageNumber="231">C. shunkan</emphasis>
</taxonomicName>
), M. Sabaj Pérez (
<taxonomicName id="4C184D3DCD17FE0D4DE2F8A65BA7F846" ID-CoL="PV9Y" box="[490,668,1903,1927]" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="231" phylum="Chordata" rank="species" species="maculipinnis">
<emphasis id="B96CEAACCD17FE0D4DE2F8A65BA7F846" box="[490,668,1903,1927]" italics="true" pageId="6" pageNumber="231">C. maculipinnis</emphasis>
</taxonomicName>
) and D. Catania (
<taxonomicName id="4C184D3DCD17FE0D4F6EF8A65AF5F846" ID-CoL="3PSW8" box="[870,974,1903,1927]" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="231" phylum="Chordata" rank="species" species="kuderi">
<emphasis id="B96CEAACCD17FE0D4F6EF8A65AF5F846" box="[870,974,1903,1927]" italics="true" pageId="6" pageNumber="231">C. kuderi</emphasis>
</taxonomicName>
). L. Parenti re-examined the
<typeStatus id="54A3881CCD17FE0D4913F8A75C46F846" box="[1307,1405,1902,1927]" pageId="6" pageNumber="231" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C184D3DCD17FE0D4C9FF85A581FF86A" ID-CoL="3PSW9" box="[151,292,1939,1963]" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="231" phylum="Chordata" rank="species" species="vanclevei">
<emphasis id="B96CEAACCD17FE0D4C9FF85A581FF86A" box="[151,292,1939,1963]" italics="true" pageId="6" pageNumber="231">C. vanclevei</emphasis>
</taxonomicName>
, a distorted specimen that appears to have no fourth transverse mandibular row on either side. The
<typeStatus id="54A3881CCD17FE0D4C9FF87F59C2F80E" box="[151,249,1974,1999]" pageId="6" pageNumber="231" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C184D3DCD17FE0D4D24F87E5B35F80F" ID-CoL="5WW2V" box="[300,526,1975,1998]" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="231" phylum="Chordata" rank="species" species="nigromarginatus">
<emphasis id="B96CEAACCD17FE0D4D24F87E5B35F80F" box="[300,526,1975,1998]" italics="true" pageId="6" pageNumber="231">C. nigromarginatus</emphasis>
</taxonomicName>
was unavailable, but the illustration and text indicate that three transverse mandibular rows are present (
<bibRefCitation id="EF894B4FCD17FE0D4DEBF8135BFBF833" author="Chen" box="[483,704,2010,2035]" pageId="6" pageNumber="231" refString="Chen, J. - P. &amp; Shao, K. - T. (2000) Callogobius nigromarginatus, a new species of goby (Pisces: Gobiidae) from Taiwan. Bulletin of Marine Science, 66, 457 - 466." type="journal article" year="2000">Chen &amp; Shao 2000</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3026535CD16FE0C4CCFFF5158D3FA25" pageId="7" pageNumber="232" type="materials_examined">
<paragraph id="8BA736BECD16FE0C4CCFFF5159D7FE49" blockId="7.[151,1437,151,1509]" pageId="7" pageNumber="232">
We found only a single Red Sea specimen (USNM 296956, female,
<quantity id="4CE09B5BCD16FE0C4FEAFF515D72FF6E" box="[994,1097,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="4.43" pageId="7" pageNumber="232" unit="mm" value="44.3">44.3 mm</quantity>
SL) that has three transverse mandibular rows on each side. We tentatively identify it as
<taxonomicName id="4C184D3DCD16FE0C4F3CFF755D60FF15" box="[820,1115,188,212]" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="232" phylum="Chordata" rank="species" species="pilosimentum">
<emphasis id="B96CEAACCD16FE0C4F3CFF755D60FF15" box="[820,1115,188,212]" italics="true" pageId="7" pageNumber="232">Callogobius pilosimentum</emphasis>
</taxonomicName>
, but we have not designated it as a paratype. We presume that this is simply unusual variation in a species that normally has four on each side. To evaluate variation in transverse mandibular papillae rows (Row 16,
<figureCitation id="13232A3BCD16FE0C4FB8FECD5AD4FEDD" box="[944,1007,260,285]" captionStart="FIGURE 1" captionStartId="3.[151,250,1751,1773]" captionTargetBox="[247,1331,204,1700]" captionTargetId="figure@3.[237,1347,193,1715]" captionTargetPageId="3" captionText="FIGURE 1. Sensory pore and papillae pattern on the head of Callogobius pilosimentum sp. nov., SMF 35756, holotype, female, 36.4 mm SL. A. Lateral view; B. Ventral view. Letter abbreviations of sensory pores follow Akihito &amp; Meguro (1977) and descriptive names are modified from Takagi (1957). Papillae row numbering follows Akihito and Meguro and descriptive names are from Delventhal &amp; Mooi (2013). Sensory pores: B = posterior nasal; C = anterior interorbital; D = posterior interorbital; E = supraotic; F = anterior otic; G = posterior otic; H = intertemporal; M, N, O = preopercular; K = anterior temporal; L = posterior temporal. Papillae rows: 2 = postnasal; 9 = anterior suborbital; 10 = mid suborbital; 11 = posterior suborbital; 12 = longitudinal cheek; 13 = transverse cheek; 14 = longitudinal maxillary; 15 = longitudinal mandibular; 16 = transverse mandibular; 17 = postorbital; 20 = preopercular; 21 = transverse opercular (rows 20 and 21 continuous). Scale bar = 2 mm. Illustration by RDM and NRD." httpUri="https://zenodo.org/record/261617/files/figure.png" pageId="7" pageNumber="232">Fig. 1</figureCitation>
) in specimens outside of the Red Sea, we examined 174 adult specimens identified in collections as
<taxonomicName id="4C184D3DCD16FE0C4F60FEE15D26FE81" box="[872,1053,296,320]" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="232" phylum="Chordata" rank="species" species="maculipinnis">
<emphasis id="B96CEAACCD16FE0C4F60FEE15D26FE81" box="[872,1053,296,320]" italics="true" pageId="7" pageNumber="232">C. maculipinnis</emphasis>
</taxonomicName>
ranging across the Indian Ocean eastward to Mangareva (Gambier Islands, French Polynesia) in the Pacific Ocean (listed in Comparative material below).
</paragraph>
<paragraph id="8BA736BECD16FE0C4CCFFE5D5C42FCF9" blockId="7.[151,1437,151,1509]" pageId="7" pageNumber="232">
Of the 174 specimens examined from outside of the Red Sea, only three (&lt;1.7%) exhibited the condition of
<taxonomicName id="4C184D3DCD16FE0C498AFE5D5815FE11" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="232" phylum="Chordata" rank="species" species="pilosimentum">
<emphasis id="B96CEAACCD16FE0C498AFE5D5815FE11" italics="true" pageId="7" pageNumber="232">C. pilosimentum</emphasis>
</taxonomicName>
of having four transverse mandibular papillae rows on both sides of the jaw. Even among these, only one (WAM P27935-024) exhibited a normal, sequential four-row anatomy, and this specimen belongs to an Australian population that has distinctly higher meristic counts than other members of the complex (D. Hoese pers. comm.). The other two specimens (USNM 332226 and ZRC 40669) have unusual morphologies where the fourth transverse mandibular row on one side follows a free papilla and/or the first row is a branch of the intermandibular row. An additional 11 specimens had four rows on only one side and three or fewer on the other (6.3% of all specimens). From localities with the largest representation, an Indonesian lot (USNM 241882) had three of 32 specimens having four transverse mandibular papillae rows on one side only and of 42 specimens in 11 lots from Fiji only two specimens had four such rows on one side. This is in contrast to the condition in
<taxonomicName id="4C184D3DCD16FE0C498AFD115815FCD5" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="232" phylum="Chordata" rank="species" species="pilosimentum">
<emphasis id="B96CEAACCD16FE0C498AFD115815FCD5" italics="true" pageId="7" pageNumber="232">C. pilosimentum</emphasis>
</taxonomicName>
in the Red Sea where all specimens exhibited four or more transverse mandibular rows on each side except one that had five on one side and two on the other, and one that had three on each side (USNM 296956).
</paragraph>
<paragraph id="8BA736BECD16FE0C4CCFFC8C5BE1FC2C" blockId="7.[151,1437,151,1509]" pageId="7" pageNumber="232">
In
<taxonomicName id="4C184D3DCD16FE0C4CEDFC8D5B30FC9D" box="[229,523,836,860]" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="232" phylum="Chordata" rank="species" species="pilosimentum">
<emphasis id="B96CEAACCD16FE0C4CEDFC8D5B30FC9D" box="[229,523,836,860]" italics="true" pageId="7" pageNumber="232">Callogobius pilosimentum</emphasis>
</taxonomicName>
, the anterior first dorsal-fin spines are usually free of membrane for one-fourth or more of the spine length, particularly the second spine. Although clearly evident on some live specimens (
<figureCitation id="13232A3BCD16FE0C4941FCAE5CB5FC41" box="[1353,1422,871,896]" captionStart="FIGURE 2" captionStartId="4.[151,250,1952,1974]" captionTargetBox="[187,1400,203,1903]" captionTargetId="figure@4.[187,1400,203,1904]" captionTargetPageId="4" captionText="FIGURE 2. Callogobius pilosimentum sp. nov., live coloration: A. SMF 35756, holotype, female, 36.4 mm SL, Farasan Island, Saudi Arabia; B. Fresh coloration, uncatalogued, Farasan Island, Saudi Arabia, illustrating pattern frequently induced by stress; C. SMF 35760, paratype, female, 55.8 mm SL, Obhur, Jeddah, Saudi Arabia. Photos by SVB." httpUri="https://zenodo.org/record/261618/files/figure.png" pageId="7" pageNumber="232">Fig. 2</figureCitation>
), this can be difficult to assess on preserved specimens. This feature is common to most, if not all, members of the
<taxonomicName id="4C184D3DCD16FE0C4C9FFC795813FC09" box="[151,296,944,968]" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="232" phylum="Chordata" rank="species" species="maculipinnis">
<emphasis id="B96CEAACCD16FE0C4C9FFC795813FC09" box="[151,296,944,968]" italics="true" pageId="7" pageNumber="232">maculipinnis</emphasis>
</taxonomicName>
species complex so is not diagnostic, but is potentially of value to distinguish live or particularly well-preserved specimens from
<collectingRegion id="49DCF85CCD16FE0C4DF5FC1D5B62FC2D" box="[509,601,980,1005]" country="Sudan" name="Al Bahr al Ahmar" pageId="7" pageNumber="232">Red Sea</collectingRegion>
congeners.
</paragraph>
<paragraph id="8BA736BECD16FE0C4CCFFC3E58C1FA5D" blockId="7.[151,1437,151,1509]" pageId="7" pageNumber="232">
<emphasis id="B96CEAACCD16FE0C4CCFFC3E58E3FBD1" bold="true" box="[199,472,1015,1040]" pageId="7" pageNumber="232">Comparative material.</emphasis>
<taxonomicName id="4C184D3DCD16FE0C4DD7FC315A3AFBD1" box="[479,769,1016,1040]" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="232" phylum="Chordata" rank="species" species="maculipinnis">
<emphasis id="B96CEAACCD16FE0C4DD7FC315A3AFBD1" box="[479,769,1016,1040]" italics="true" pageId="7" pageNumber="232">Callogobius maculipinnis</emphasis>
</taxonomicName>
(listed west to east): Comoros—ROM 92691 (4); Oman— ROM 39895 (6); Chagos—ROM 55107 (4); Sri Lanka—USNM 220035 (5); Cocos-Keeling—WAM P29928-022 (4); Western Australia—WAM 27935-024 (3); Thailand—ROM 58036 (1), 68041 (4); Indonesia—USNM 241882 (32), WAM P33093-002 (2); Singapore—ZRC 40669 (1); Philippines—ROM 53339 (3), USNM 297102 (2); Taiwan—BPBM 23242 (1), USNM 298439 (8); Palau—BPBM 37767 (2), ROM 75955 (1); 76134 (1); Papua New Guinea—BPBM 32638 (1), 32674 (2), USNM 297051 (2); Australia—MPM 48365 (1), ROM 38903 (1), USNM 297048 (3); One Tree Island—BPBM 14432 (4); Lord Howe Island—BPBM 14862 (1); Coral Sea—BPBM 33517 (1), 33629 (3); New Caledonia—BPBM 34270 (1), ROM 64155 (1), 64160 (2), 64416 (3); Vanuatu—BPBM 5765 (1), MPM 32132 (1), 46721 (1); Marshall Islands—BPBM 8296 (1), 17739 (1), 22349 (1); Fiji—BPBM 39866 (1), 40082 (1), ROM 57700 (5), 57701 (1), 57703 (3), 57705 (6), 57707 (2), 57708 (1), 57709 (2), 57710 (1), USNM 332226 (19); Tonga—BPBM 38115 (3), USNM 339828 (7); Tahiti—BPBM 8315 (2); Rapa—BPBM 17317 (1); Mangareva—BPBM 13595 (3).
</paragraph>
<paragraph id="8BA736BECD16FE0C4CCFFA6E58D3FA25" blockId="7.[151,1437,151,1509]" pageId="7" pageNumber="232">
Comparisons to other described
<taxonomicName id="4C184D3DCD16FE0C4E3CFA615B86FA01" box="[564,701,1448,1472]" class="Actinopterygii" family="Gobiidae" genus="Callogobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="232" phylum="Chordata" rank="genus">
<emphasis id="B96CEAACCD16FE0C4E3CFA615B86FA01" box="[564,701,1448,1472]" italics="true" pageId="7" pageNumber="232">Callogobius</emphasis>
</taxonomicName>
species are based on our data from the available
<typeStatus id="54A3881CCD16FE0C48E2FA6E5C6CFA01" box="[1258,1367,1447,1472]" pageId="7" pageNumber="232" type="holotype">holotypes</typeStatus>
listed in
<bibRefCitation id="EF894B4FCD16FE0C4CBCFA0558DFFA25" author="Delventhal" box="[180,484,1484,1509]" pageId="7" pageNumber="232" refString="Delventhal, N. R. &amp; Mooi, R. D. (2013) Callogobius winterbottomi, a new species of goby (Teleostei: Gobiidae) from the Western Indian Ocean. Zootaxa, 3630 (1), 155 - 164. http: // dx. doi. org / 10.11646 / zootaxa. 3630.1.6" type="journal article" year="2013">Delventhal &amp; Mooi (2013)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>