254 lines
40 KiB
XML
254 lines
40 KiB
XML
<document id="F108DE2EBAC72D9682563D8775C26077" ID-DOI="10.5281/zenodo.169460" ID-GBIF-Dataset="c6b6aaea-4e91-4f23-b4bd-c80a4f676e13" ID-ISSN="11755326" ID-Zenodo-Dep="169460" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461032056718" checkinUser="plazi" docAuthor="Johanson, Kjell Arne & Holzenthal, Ralph W." docDate="2004" docId="03B1879DFFA5DE5EFE98F9947E8DF983" docLanguage="en" docName="zt00711.pdf" docOrigin="Zootaxa 711" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Helicopsyche (Feropsyche) lara Johanson & Holzenthal, 2004, new species" docType="treatment" docVersion="9" lastPageNumber="21" masterDocId="FF88FFE5FFB7DE4AFF90FFD07F67FFB5" masterDocTitle="Thirteen new species and new distribution records of Helicopsyche (Feropsyche) Johanson from Venezuela (Trichoptera: Helicopsychidae)" masterLastPageNumber="40" masterPageNumber="1" pageNumber="19" updateTime="1698573568728" updateUser="plazi">
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<mods:title id="FBAA2381D204E11541D0DAB3996B41CC">Thirteen new species and new distribution records of Helicopsyche (Feropsyche) Johanson from Venezuela (Trichoptera: Helicopsychidae)</mods:title>
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<mods:name id="BFA88D1F998DF20EDA4727CFC65F59A3" type="personal">
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<mods:namePart id="10D588EDB6749A6E3153BFF3B2B09D34">Johanson, Kjell Arne</mods:namePart>
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<mods:namePart id="DA4267D27970541266368DA2DD6BE327">Holzenthal, Ralph W.</mods:namePart>
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<mods:typeOfResource id="58BA5CF319FCAB209BE778A53D36525C">text</mods:typeOfResource>
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<mods:part id="4CFC07A4A4C679D4AE14026CF9329BDF">
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<mods:date id="D3610C559CEA6F2AAA0A6A7EC9C5A309">2004</mods:date>
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<mods:identifier id="D78DE48C0953F2F80612E17AA2219914" type="DOI">10.5281/zenodo.169460</mods:identifier>
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<treatment id="03B1879DFFA5DE5EFE98F9947E8DF983" ID-DOI="http://doi.org/10.5281/zenodo.6271568" ID-GBIF-Taxon="119608257" ID-Zenodo-Dep="6271568" LSID="urn:lsid:plazi:treatment:03B1879DFFA5DE5EFE98F9947E8DF983" httpUri="http://treatment.plazi.org/id/03B1879DFFA5DE5EFE98F9947E8DF983" lastPageId="20" lastPageNumber="21" pageId="18" pageNumber="19">
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<subSubSection id="C3026500FFA5DE58FE98F9947EDEF934" pageId="18" pageNumber="19" type="nomenclature">
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<paragraph id="8BA7368BFFA5DE58FE98F9947C40F9EB" blockId="18.[264,807,1604,1665]" box="[264,807,1604,1630]" pageId="18" pageNumber="19">
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<heading id="D0EF81E7FFA5DE58FE98F9947C40F9EB" bold="true" box="[264,807,1604,1630]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
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<emphasis id="B96CEA99FFA5DE58FE98F9947C40F9EB" bold="true" box="[264,807,1604,1630]" pageId="18" pageNumber="19">
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<taxonomicName id="4C184D08FFA5DE58FE98F9947DEFF9EB" box="[264,648,1604,1630]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="lara" status="sp. nov." subGenus="Feropsyche">
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<emphasis id="B96CEA99FFA5DE58FE98F9947ECEF9EB" bold="true" box="[264,425,1604,1630]" italics="true" pageId="18" pageNumber="19">Helicopsyche</emphasis>
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(
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<emphasis id="B96CEA99FFA5DE58FE2AF9947D21F9EB" bold="true" box="[442,582,1604,1630]" italics="true" pageId="18" pageNumber="19">Feropsyche</emphasis>
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)
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<emphasis id="B96CEA99FFA5DE58FDC7F9947DEFF9EB" bold="true" box="[599,648,1604,1630]" italics="true" pageId="18" pageNumber="19">lara</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A25F57E2FFA5DE58FD07F9947C40F9EB" box="[663,807,1604,1630]" pageId="18" pageNumber="19" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BA7368BFFA5DE58FE98F9B77EDEF934" blockId="18.[264,807,1604,1665]" box="[264,441,1639,1665]" pageId="18" pageNumber="19">
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<figureCitation id="13232A0EFFA5DE58FE98F9B77EEFF934" box="[264,392,1639,1665]" captionStart="FIGURES 55 – 61" captionStartId="19.[264,382,1643,1667]" captionTargetBox="[293,1285,597,1611]" captionTargetId="figure@19.[270,1317,587,1634]" captionTargetPageId="19" captionText="FIGURES 55 – 61. Helicopsyche lara, new species, holotype. 55 — sternum VI process, lateral; 56 — sternum VI process, ventral; 57 — male genitalia, lateral; 58 — male genitalia, dorsal; 59 — male genitalia, ventral; 60 — phallus, lateral; 61 — phallus, ventral." httpUri="https://zenodo.org/record/169469/files/figure.png" pageId="18" pageNumber="19">Fig. 55–61</figureCitation>
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,
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<figureCitation id="13232A0EFFA5DE58FE05F9B77ED1F934" box="[405,438,1639,1665]" captionStart="FIGURE 86" captionStartId="35.[264,367,1583,1607]" captionTargetBox="[268,1314,656,1548]" captionTargetId="figure@35.[253,1333,639,1559]" captionTargetPageId="35" captionText="FIGURE 86. Map of Venezuela showing distributions of H. vergelana (circle), H. extensa (star), and H. lara (rectangle)." httpUri="https://zenodo.org/record/169474/files/figure.png" pageId="18" pageNumber="19">86</figureCitation>
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.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3026500FFA5DE5EFE98F9677BE7FB13" lastPageId="20" lastPageNumber="21" pageId="18" pageNumber="19" type="description">
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<paragraph id="8BA7368BFFA5DE58FE98F9677C1FF8FC" blockId="18.[264,1323,1719,1865]" pageId="18" pageNumber="19">
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Together with
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<taxonomicName id="4C184D08FFA5DE58FE21F9677D3FF965" box="[433,600,1719,1745]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="neblinensis">
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<emphasis id="B96CEA99FFA5DE58FE21F9677EA1F965" box="[433,454,1719,1744]" italics="true" pageId="18" pageNumber="19">H</emphasis>
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.
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<emphasis id="B96CEA99FFA5DE58FE44F9677D3FF965" box="[468,600,1719,1744]" italics="true" pageId="18" pageNumber="19">neblinensis</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C184D08FFA5DE58FDF6F9677DDCF965" box="[614,699,1719,1745]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="lara">
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<emphasis id="B96CEA99FFA5DE58FDF6F9677D1CF965" box="[614,635,1719,1744]" italics="true" pageId="18" pageNumber="19">H</emphasis>
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.
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<emphasis id="B96CEA99FFA5DE58FD1AF9677DDCF965" box="[650,699,1719,1744]" italics="true" pageId="18" pageNumber="19">lara</emphasis>
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</taxonomicName>
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has a rounded, triangular gonocoxite which is unique among American
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<taxonomicName id="4C184D08FFA5DE58FE4BF90F7D1EF94D" box="[475,633,1759,1784]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
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<emphasis id="B96CEA99FFA5DE58FE4BF90F7D1EF94D" box="[475,633,1759,1784]" italics="true" pageId="18" pageNumber="19">Helicopsyche</emphasis>
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</taxonomicName>
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.
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<taxonomicName id="4C184D08FFA5DE58FD00F90F7C01F94D" box="[656,870,1759,1784]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="lara">
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<emphasis id="B96CEA99FFA5DE58FD00F90F7C01F94D" box="[656,870,1759,1784]" italics="true" pageId="18" pageNumber="19">Helicopsyche lara</emphasis>
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</taxonomicName>
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can be separated from
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<taxonomicName id="4C184D08FFA5DE58FBEBF90F7A44F94D" box="[1147,1315,1759,1785]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="neblinensis">
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<emphasis id="B96CEA99FFA5DE58FBEBF90F7BF7F94D" box="[1147,1168,1759,1784]" italics="true" pageId="18" pageNumber="19">H</emphasis>
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.
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<emphasis id="B96CEA99FFA5DE58FB0FF90F7A44F94D" box="[1183,1315,1759,1784]" italics="true" pageId="18" pageNumber="19">neblinensis</emphasis>
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</taxonomicName>
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, in dorsal view, by a slightly wider tergum X having about 16 pairs of dorsal setae, and in lateral view, by the slightly more slender gonocoxite.
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</paragraph>
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<paragraph id="8BA7368BFFA4DE59FEA8FECC7B1AFD93" blockId="19.[264,1324,284,550]" pageId="19" pageNumber="20">
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Male. Head: Antennal scape as long as eye diameter, with pale yellowish setae. Maxillary palp segments equally long, with dark brown and grey setae. Cephalic warts oval, about half as long as eye diameter; with yellowish setae, some dark brown near lateral margins. Forewing golden brown, length
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<quantity id="4CE09B6EFFA4DE59FD7DFE447C1CFE1B" box="[749,891,404,430]" metricMagnitude="-3" metricUnit="m" metricValue="5.25" metricValueMax="5.4" metricValueMin="5.1" pageId="19" pageNumber="20" unit="mm" value="5.25" valueMax="5.4" valueMin="5.1">5.1–5.4 mm</quantity>
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. Sternum VI process (
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<figureCitation id="13232A0EFFA4DE59FB14FE447A62FE1B" box="[1156,1285,404,430]" captionStart="FIGURES 55 – 61" captionStartId="19.[264,382,1643,1667]" captionTargetBox="[293,1285,597,1611]" captionTargetId="figure@19.[270,1317,587,1634]" captionTargetPageId="19" captionText="FIGURES 55 – 61. Helicopsyche lara, new species, holotype. 55 — sternum VI process, lateral; 56 — sternum VI process, ventral; 57 — male genitalia, lateral; 58 — male genitalia, dorsal; 59 — male genitalia, ventral; 60 — phallus, lateral; 61 — phallus, ventral." httpUri="https://zenodo.org/record/169469/files/figure.png" pageId="19" pageNumber="20">Fig. 55, 56</figureCitation>
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) 1/ 3 its segment length, covered by microtrichiae; in lateral view (
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<figureCitation id="13232A0EFFA4DE59FB97FE6C7B07FE63" box="[1031,1120,444,470]" captionStart="FIGURES 55 – 61" captionStartId="19.[264,382,1643,1667]" captionTargetBox="[293,1285,597,1611]" captionTargetId="figure@19.[270,1317,587,1634]" captionTargetPageId="19" captionText="FIGURES 55 – 61. Helicopsyche lara, new species, holotype. 55 — sternum VI process, lateral; 56 — sternum VI process, ventral; 57 — male genitalia, lateral; 58 — male genitalia, dorsal; 59 — male genitalia, ventral; 60 — phallus, lateral; 61 — phallus, ventral." httpUri="https://zenodo.org/record/169469/files/figure.png" pageId="19" pageNumber="20">Fig. 55</figureCitation>
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), nearly straight, slender along its length; in ventral view (
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<figureCitation id="13232A0EFFA4DE59FD60FE347C2EFE4B" box="[752,841,484,510]" captionStart="FIGURES 48 – 54" captionStartId="17.[264,382,1504,1528]" captionTargetBox="[280,1312,417,1475]" captionTargetId="figure@17.[270,1317,405,1491]" captionTargetPageId="17" captionText="FIGURES 48 – 54. Helicopsyche venezuelensis, new species, holotype. 48 — sternum VI process, lateral; 49 — sternum VI process, ventral; 50 — male genitalia, lateral; 51 — male genitalia, dorsal; 52 — male genitalia, ventral; 53 — phallus, lateral; 54 — phallus, ventral." httpUri="https://zenodo.org/record/169468/files/figure.png" pageId="19" pageNumber="20">Fig. 49</figureCitation>
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), with proximal half slightly tapering, in distal half parallelsided; apex bearing numerous ventral lamellae (
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<figureCitation id="13232A0EFFA4DE59FB87FDDC7B08FD93" box="[1047,1135,524,550]" captionStart="FIGURES 55 – 61" captionStartId="19.[264,382,1643,1667]" captionTargetBox="[293,1285,597,1611]" captionTargetId="figure@19.[270,1317,587,1634]" captionTargetPageId="19" captionText="FIGURES 55 – 61. Helicopsyche lara, new species, holotype. 55 — sternum VI process, lateral; 56 — sternum VI process, ventral; 57 — male genitalia, lateral; 58 — male genitalia, dorsal; 59 — male genitalia, ventral; 60 — phallus, lateral; 61 — phallus, ventral." httpUri="https://zenodo.org/record/169469/files/figure.png" pageId="19" pageNumber="20">Fig. 56</figureCitation>
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).
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</paragraph>
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<caption id="DF676603FFA4DE59FE98F9BB7CEDF97A" httpUri="https://zenodo.org/record/169469/files/figure.png" pageId="19" pageNumber="20" targetBox="[293,1285,597,1611]" targetPageId="19">
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<paragraph id="8BA7368BFFA4DE59FE98F9BB7CEDF97A" blockId="19.[264,1323,1643,1743]" pageId="19" pageNumber="20">
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<emphasis id="B96CEA99FFA4DE59FE98F9BB7EB6F936" bold="true" box="[264,465,1643,1667]" pageId="19" pageNumber="20">FIGURES 55–61.</emphasis>
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<taxonomicName id="4C184D08FFA4DE59FE74F9BB7DCCF937" box="[484,683,1643,1666]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="lara" status="sp. nov.">
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<emphasis id="B96CEA99FFA4DE59FE74F9BB7DCCF937" box="[484,683,1643,1666]" italics="true" pageId="19" pageNumber="20">Helicopsyche lara</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A25F57E2FFA4DE59FD2BF9BB7C26F937" box="[699,833,1643,1666]" pageId="19" pageNumber="20" rank="species">new species</taxonomicNameLabel>
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, holotype. 55—sternum VI process, lateral; 56—sternum VI process, ventral; 57—male genitalia, lateral; 58—male genitalia, dorsal; 59—male genitalia, ventral; 60—phallus, lateral; 61—phallus, ventral.
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</paragraph>
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</caption>
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<paragraph id="8BA7368BFFA4DE5EFEA8F8C37BE7FB13" blockId="19.[264,1322,1811,1877]" lastBlockId="20.[264,1323,284,1590]" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
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Male genitalia (
|
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<figureCitation id="13232A0EFFA4DE59FE62F8C37D12F898" box="[498,629,1811,1837]" captionStart="FIGURES 55 – 61" captionStartId="19.[264,382,1643,1667]" captionTargetBox="[293,1285,597,1611]" captionTargetId="figure@19.[270,1317,587,1634]" captionTargetPageId="19" captionText="FIGURES 55 – 61. Helicopsyche lara, new species, holotype. 55 — sternum VI process, lateral; 56 — sternum VI process, ventral; 57 — male genitalia, lateral; 58 — male genitalia, dorsal; 59 — male genitalia, ventral; 60 — phallus, lateral; 61 — phallus, ventral." httpUri="https://zenodo.org/record/169469/files/figure.png" pageId="19" pageNumber="20">Fig. 57–61</figureCitation>
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). Segment IX, in lateral view (
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<figureCitation id="13232A0EFFA4DE59FC7AF8C37B25F898" box="[1002,1090,1811,1837]" captionStart="FIGURES 55 – 61" captionStartId="19.[264,382,1643,1667]" captionTargetBox="[293,1285,597,1611]" captionTargetId="figure@19.[270,1317,587,1634]" captionTargetPageId="19" captionText="FIGURES 55 – 61. Helicopsyche lara, new species, holotype. 55 — sternum VI process, lateral; 56 — sternum VI process, ventral; 57 — male genitalia, lateral; 58 — male genitalia, dorsal; 59 — male genitalia, ventral; 60 — phallus, lateral; 61 — phallus, ventral." httpUri="https://zenodo.org/record/169469/files/figure.png" pageId="19" pageNumber="20">Fig. 57</figureCitation>
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), with anterior lobe triangular, oriented anteriad and located at midlaterally on segment; anterodorsal and anteroventral margins slightly concave; in dorsal view (
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<figureCitation id="13232A0EFFA3DE5EFC05FECC7C8BFE83" box="[917,1004,284,310]" captionStart="FIGURES 55 – 61" captionStartId="19.[264,382,1643,1667]" captionTargetBox="[293,1285,597,1611]" captionTargetId="figure@19.[270,1317,587,1634]" captionTargetPageId="19" captionText="FIGURES 55 – 61. Helicopsyche lara, new species, holotype. 55 — sternum VI process, lateral; 56 — sternum VI process, ventral; 57 — male genitalia, lateral; 58 — male genitalia, dorsal; 59 — male genitalia, ventral; 60 — phallus, lateral; 61 — phallus, ventral." httpUri="https://zenodo.org/record/169469/files/figure.png" pageId="20" pageNumber="21">Fig. 58</figureCitation>
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), with inner margin widely ellipsoid; in ventral view (
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<figureCitation id="13232A0EFFA3DE5EFDACFE947DF3FEEB" box="[572,660,324,350]" captionStart="FIGURES 55 – 61" captionStartId="19.[264,382,1643,1667]" captionTargetBox="[293,1285,597,1611]" captionTargetId="figure@19.[270,1317,587,1634]" captionTargetPageId="19" captionText="FIGURES 55 – 61. Helicopsyche lara, new species, holotype. 55 — sternum VI process, lateral; 56 — sternum VI process, ventral; 57 — male genitalia, lateral; 58 — male genitalia, dorsal; 59 — male genitalia, ventral; 60 — phallus, lateral; 61 — phallus, ventral." httpUri="https://zenodo.org/record/169469/files/figure.png" pageId="20" pageNumber="21">Fig. 59</figureCitation>
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), with minute posterior process; lateral apodeme present as anteriorly oriented, nearly straight line (
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<figureCitation id="13232A0EFFA3DE5EFD6BFEBC7C36FE33" box="[763,849,364,390]" captionStart="FIGURES 55 – 61" captionStartId="19.[264,382,1643,1667]" captionTargetBox="[293,1285,597,1611]" captionTargetId="figure@19.[270,1317,587,1634]" captionTargetPageId="19" captionText="FIGURES 55 – 61. Helicopsyche lara, new species, holotype. 55 — sternum VI process, lateral; 56 — sternum VI process, ventral; 57 — male genitalia, lateral; 58 — male genitalia, dorsal; 59 — male genitalia, ventral; 60 — phallus, lateral; 61 — phallus, ventral." httpUri="https://zenodo.org/record/169469/files/figure.png" pageId="20" pageNumber="21">Fig. 57</figureCitation>
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) reaching anterior margin; submarginal line absent; tergal transverse apodeme narrow; sternal transverse apodeme well developed. Segment X, in lateral view (
|
||
<figureCitation id="13232A0EFFA3DE5EFDF3FE6C7DDAFE63" box="[611,701,444,470]" captionStart="FIGURES 55 – 61" captionStartId="19.[264,382,1643,1667]" captionTargetBox="[293,1285,597,1611]" captionTargetId="figure@19.[270,1317,587,1634]" captionTargetPageId="19" captionText="FIGURES 55 – 61. Helicopsyche lara, new species, holotype. 55 — sternum VI process, lateral; 56 — sternum VI process, ventral; 57 — male genitalia, lateral; 58 — male genitalia, dorsal; 59 — male genitalia, ventral; 60 — phallus, lateral; 61 — phallus, ventral." httpUri="https://zenodo.org/record/169469/files/figure.png" pageId="20" pageNumber="21">Fig. 57</figureCitation>
|
||
), oriented posteroventrad, nearly straight; proximal half with dorsal and ventral margins parallel, distal half tapering toward rounded apex; in dorsal view (
|
||
<figureCitation id="13232A0EFFA3DE5EFE30FDDC7E90FD93" box="[416,503,524,550]" captionStart="FIGURES 55 – 61" captionStartId="19.[264,382,1643,1667]" captionTargetBox="[293,1285,597,1611]" captionTargetId="figure@19.[270,1317,587,1634]" captionTargetPageId="19" captionText="FIGURES 55 – 61. Helicopsyche lara, new species, holotype. 55 — sternum VI process, lateral; 56 — sternum VI process, ventral; 57 — male genitalia, lateral; 58 — male genitalia, dorsal; 59 — male genitalia, ventral; 60 — phallus, lateral; 61 — phallus, ventral." httpUri="https://zenodo.org/record/169469/files/figure.png" pageId="20" pageNumber="21">Fig. 58</figureCitation>
|
||
), slightly narrowing distally; apex without notch (
|
||
<figureCitation id="13232A0EFFA3DE5EFBD1FDDC7BFEFD93" box="[1089,1177,524,550]" captionStart="FIGURES 55 – 61" captionStartId="19.[264,382,1643,1667]" captionTargetBox="[293,1285,597,1611]" captionTargetId="figure@19.[270,1317,587,1634]" captionTargetPageId="19" captionText="FIGURES 55 – 61. Helicopsyche lara, new species, holotype. 55 — sternum VI process, lateral; 56 — sternum VI process, ventral; 57 — male genitalia, lateral; 58 — male genitalia, dorsal; 59 — male genitalia, ventral; 60 — phallus, lateral; 61 — phallus, ventral." httpUri="https://zenodo.org/record/169469/files/figure.png" pageId="20" pageNumber="21">Fig. 58</figureCitation>
|
||
); with about 16 pairs of about equally long megasetae in longitudinal group starting at midway on segment. Superior appendage (
|
||
<figureCitation id="13232A0EFFA3DE5EFDC5FD8C7DCCFDC3" box="[597,683,604,630]" captionStart="FIGURES 55 – 61" captionStartId="19.[264,382,1643,1667]" captionTargetBox="[293,1285,597,1611]" captionTargetId="figure@19.[270,1317,587,1634]" captionTargetPageId="19" captionText="FIGURES 55 – 61. Helicopsyche lara, new species, holotype. 55 — sternum VI process, lateral; 56 — sternum VI process, ventral; 57 — male genitalia, lateral; 58 — male genitalia, dorsal; 59 — male genitalia, ventral; 60 — phallus, lateral; 61 — phallus, ventral." httpUri="https://zenodo.org/record/169469/files/figure.png" pageId="20" pageNumber="21">Fig. 57</figureCitation>
|
||
) clubshaped in lateral and dorsal views (
|
||
<figureCitation id="13232A0EFFA3DE5EFB07FD8C7A7BFDC3" box="[1175,1308,604,630]" captionStart="FIGURES 55 – 61" captionStartId="19.[264,382,1643,1667]" captionTargetBox="[293,1285,597,1611]" captionTargetId="figure@19.[270,1317,587,1634]" captionTargetPageId="19" captionText="FIGURES 55 – 61. Helicopsyche lara, new species, holotype. 55 — sternum VI process, lateral; 56 — sternum VI process, ventral; 57 — male genitalia, lateral; 58 — male genitalia, dorsal; 59 — male genitalia, ventral; 60 — phallus, lateral; 61 — phallus, ventral." httpUri="https://zenodo.org/record/169469/files/figure.png" pageId="20" pageNumber="21">Fig. 57, 58</figureCitation>
|
||
); curving posteriorly in dorsal view (
|
||
<figureCitation id="13232A0EFFA3DE5EFD34FD547D9CFD2B" box="[676,763,644,670]" captionStart="FIGURES 55 – 61" captionStartId="19.[264,382,1643,1667]" captionTargetBox="[293,1285,597,1611]" captionTargetId="figure@19.[270,1317,587,1634]" captionTargetPageId="19" captionText="FIGURES 55 – 61. Helicopsyche lara, new species, holotype. 55 — sternum VI process, lateral; 56 — sternum VI process, ventral; 57 — male genitalia, lateral; 58 — male genitalia, dorsal; 59 — male genitalia, ventral; 60 — phallus, lateral; 61 — phallus, ventral." httpUri="https://zenodo.org/record/169469/files/figure.png" pageId="20" pageNumber="21">Fig. 58</figureCitation>
|
||
). Primary branch of gonocoxite, in lateral view (
|
||
<figureCitation id="13232A0EFFA3DE5EFE81FD7C7E0DFD73" box="[273,362,684,710]" captionStart="FIGURES 55 – 61" captionStartId="19.[264,382,1643,1667]" captionTargetBox="[293,1285,597,1611]" captionTargetId="figure@19.[270,1317,587,1634]" captionTargetPageId="19" captionText="FIGURES 55 – 61. Helicopsyche lara, new species, holotype. 55 — sternum VI process, lateral; 56 — sternum VI process, ventral; 57 — male genitalia, lateral; 58 — male genitalia, dorsal; 59 — male genitalia, ventral; 60 — phallus, lateral; 61 — phallus, ventral." httpUri="https://zenodo.org/record/169469/files/figure.png" pageId="20" pageNumber="21">Fig. 57</figureCitation>
|
||
), generally triangular, with undulate dorsal margin; apex rounded in lateral and dorsal views (
|
||
<figureCitation id="13232A0EFFA3DE5EFE3AFD047D4AFD5B" box="[426,557,724,750]" captionStart="FIGURES 55 – 61" captionStartId="19.[264,382,1643,1667]" captionTargetBox="[293,1285,597,1611]" captionTargetId="figure@19.[270,1317,587,1634]" captionTargetPageId="19" captionText="FIGURES 55 – 61. Helicopsyche lara, new species, holotype. 55 — sternum VI process, lateral; 56 — sternum VI process, ventral; 57 — male genitalia, lateral; 58 — male genitalia, dorsal; 59 — male genitalia, ventral; 60 — phallus, lateral; 61 — phallus, ventral." httpUri="https://zenodo.org/record/169469/files/figure.png" pageId="20" pageNumber="21">Fig. 57, 58</figureCitation>
|
||
); central part of primary branch about as wide as height of central part of tergum X (
|
||
<figureCitation id="13232A0EFFA3DE5EFE7BFD2C7D20FCA3" box="[491,583,764,790]" captionStart="FIGURES 55 – 61" captionStartId="19.[264,382,1643,1667]" captionTargetBox="[293,1285,597,1611]" captionTargetId="figure@19.[270,1317,587,1634]" captionTargetPageId="19" captionText="FIGURES 55 – 61. Helicopsyche lara, new species, holotype. 55 — sternum VI process, lateral; 56 — sternum VI process, ventral; 57 — male genitalia, lateral; 58 — male genitalia, dorsal; 59 — male genitalia, ventral; 60 — phallus, lateral; 61 — phallus, ventral." httpUri="https://zenodo.org/record/169469/files/figure.png" pageId="20" pageNumber="21">Fig. 57</figureCitation>
|
||
); anterodorsal margin nearly straight, smooth; posteroventral margin nearly straight, undulated; basimesal lobe large, rectangular and protruding primary branch in lateral view (
|
||
<figureCitation id="13232A0EFFA3DE5EFDCBFC9C7DD4FCD3" box="[603,691,844,870]" captionStart="FIGURES 55 – 61" captionStartId="19.[264,382,1643,1667]" captionTargetBox="[293,1285,597,1611]" captionTargetId="figure@19.[270,1317,587,1634]" captionTargetPageId="19" captionText="FIGURES 55 – 61. Helicopsyche lara, new species, holotype. 55 — sternum VI process, lateral; 56 — sternum VI process, ventral; 57 — male genitalia, lateral; 58 — male genitalia, dorsal; 59 — male genitalia, ventral; 60 — phallus, lateral; 61 — phallus, ventral." httpUri="https://zenodo.org/record/169469/files/figure.png" pageId="20" pageNumber="21">Fig. 57</figureCitation>
|
||
); in ventral view, coneshaped (
|
||
<figureCitation id="13232A0EFFA3DE5EFBB9FC9C7BE6FCD3" box="[1065,1153,844,870]" captionStart="FIGURES 55 – 61" captionStartId="19.[264,382,1643,1667]" captionTargetBox="[293,1285,597,1611]" captionTargetId="figure@19.[270,1317,587,1634]" captionTargetPageId="19" captionText="FIGURES 55 – 61. Helicopsyche lara, new species, holotype. 55 — sternum VI process, lateral; 56 — sternum VI process, ventral; 57 — male genitalia, lateral; 58 — male genitalia, dorsal; 59 — male genitalia, ventral; 60 — phallus, lateral; 61 — phallus, ventral." httpUri="https://zenodo.org/record/169469/files/figure.png" pageId="20" pageNumber="21">Fig. 59</figureCitation>
|
||
); with slightly undulating and diverging median margins armed with a strong seta; with about 8 short megasetae on apical margin; basal plate, in lateral view (
|
||
<figureCitation id="13232A0EFFA3DE5EFC51FC4C7B7BFC03" box="[961,1052,924,950]" captionStart="FIGURES 55 – 61" captionStartId="19.[264,382,1643,1667]" captionTargetBox="[293,1285,597,1611]" captionTargetId="figure@19.[270,1317,587,1634]" captionTargetPageId="19" captionText="FIGURES 55 – 61. Helicopsyche lara, new species, holotype. 55 — sternum VI process, lateral; 56 — sternum VI process, ventral; 57 — male genitalia, lateral; 58 — male genitalia, dorsal; 59 — male genitalia, ventral; 60 — phallus, lateral; 61 — phallus, ventral." httpUri="https://zenodo.org/record/169469/files/figure.png" pageId="20" pageNumber="21">Fig. 57</figureCitation>
|
||
), nearly straight, apex pointed; in ventral view (
|
||
<figureCitation id="13232A0EFFA3DE5EFDADFC147DFFFC6B" box="[573,664,964,990]" captionStart="FIGURES 55 – 61" captionStartId="19.[264,382,1643,1667]" captionTargetBox="[293,1285,597,1611]" captionTargetId="figure@19.[270,1317,587,1634]" captionTargetPageId="19" captionText="FIGURES 55 – 61. Helicopsyche lara, new species, holotype. 55 — sternum VI process, lateral; 56 — sternum VI process, ventral; 57 — male genitalia, lateral; 58 — male genitalia, dorsal; 59 — male genitalia, ventral; 60 — phallus, lateral; 61 — phallus, ventral." httpUri="https://zenodo.org/record/169469/files/figure.png" pageId="20" pageNumber="21">Fig. 59</figureCitation>
|
||
), strongly pointing anteriorly. Phallus, in lateral view (
|
||
<figureCitation id="13232A0EFFA3DE5EFE81FC3C7E0DFBB3" box="[273,362,1004,1030]" captionStart="FIGURES 55 – 61" captionStartId="19.[264,382,1643,1667]" captionTargetBox="[293,1285,597,1611]" captionTargetId="figure@19.[270,1317,587,1634]" captionTargetPageId="19" captionText="FIGURES 55 – 61. Helicopsyche lara, new species, holotype. 55 — sternum VI process, lateral; 56 — sternum VI process, ventral; 57 — male genitalia, lateral; 58 — male genitalia, dorsal; 59 — male genitalia, ventral; 60 — phallus, lateral; 61 — phallus, ventral." httpUri="https://zenodo.org/record/169469/files/figure.png" pageId="20" pageNumber="21">Fig. 60</figureCitation>
|
||
), gently bent ventrad at midlenth, dorsal margin nearly straight along its length; ventral margin gently concave; anterior 1/5 nearly 1.5x broader than its central part (
|
||
<figureCitation id="13232A0EFFA3DE5EFB6BFBC47E4EFBE3" captionStart="FIGURES 55 – 61" captionStartId="19.[264,382,1643,1667]" captionTargetBox="[293,1285,597,1611]" captionTargetId="figure@19.[270,1317,587,1634]" captionTargetPageId="19" captionText="FIGURES 55 – 61. Helicopsyche lara, new species, holotype. 55 — sternum VI process, lateral; 56 — sternum VI process, ventral; 57 — male genitalia, lateral; 58 — male genitalia, dorsal; 59 — male genitalia, ventral; 60 — phallus, lateral; 61 — phallus, ventral." httpUri="https://zenodo.org/record/169469/files/figure.png" pageId="20" pageNumber="21">Fig. 60</figureCitation>
|
||
); in ventral view (
|
||
<figureCitation id="13232A0EFFA3DE5EFD99FBEC7D04FBE3" box="[521,611,1084,1110]" captionStart="FIGURES 55 – 61" captionStartId="19.[264,382,1643,1667]" captionTargetBox="[293,1285,597,1611]" captionTargetId="figure@19.[270,1317,587,1634]" captionTargetPageId="19" captionText="FIGURES 55 – 61. Helicopsyche lara, new species, holotype. 55 — sternum VI process, lateral; 56 — sternum VI process, ventral; 57 — male genitalia, lateral; 58 — male genitalia, dorsal; 59 — male genitalia, ventral; 60 — phallus, lateral; 61 — phallus, ventral." httpUri="https://zenodo.org/record/169469/files/figure.png" pageId="20" pageNumber="21">Fig. 61</figureCitation>
|
||
), tapering toward midlength; phallobase apparently absent; endotheca strongly produced (
|
||
<figureCitation id="13232A0EFFA3DE5EFDFFFBB47DAFFBCB" box="[623,712,1124,1150]" captionStart="FIGURES 55 – 61" captionStartId="19.[264,382,1643,1667]" captionTargetBox="[293,1285,597,1611]" captionTargetId="figure@19.[270,1317,587,1634]" captionTargetPageId="19" captionText="FIGURES 55 – 61. Helicopsyche lara, new species, holotype. 55 — sternum VI process, lateral; 56 — sternum VI process, ventral; 57 — male genitalia, lateral; 58 — male genitalia, dorsal; 59 — male genitalia, ventral; 60 — phallus, lateral; 61 — phallus, ventral." httpUri="https://zenodo.org/record/169469/files/figure.png" pageId="20" pageNumber="21">Fig. 60</figureCitation>
|
||
); sperm channel undivided; sclerotized posteroventral part narrow (
|
||
<figureCitation id="13232A0EFFA3DE5EFE5EFB5C7D41FB13" box="[462,550,1164,1190]" captionStart="FIGURES 55 – 61" captionStartId="19.[264,382,1643,1667]" captionTargetBox="[293,1285,597,1611]" captionTargetId="figure@19.[270,1317,587,1634]" captionTargetPageId="19" captionText="FIGURES 55 – 61. Helicopsyche lara, new species, holotype. 55 — sternum VI process, lateral; 56 — sternum VI process, ventral; 57 — male genitalia, lateral; 58 — male genitalia, dorsal; 59 — male genitalia, ventral; 60 — phallus, lateral; 61 — phallus, ventral." httpUri="https://zenodo.org/record/169469/files/figure.png" pageId="20" pageNumber="21">Fig. 60</figureCitation>
|
||
), forming a truncate plate in ventral view (
|
||
<figureCitation id="13232A0EFFA3DE5EFB8AFB5C7B15FB13" box="[1050,1138,1164,1190]" captionStart="FIGURES 55 – 61" captionStartId="19.[264,382,1643,1667]" captionTargetBox="[293,1285,597,1611]" captionTargetId="figure@19.[270,1317,587,1634]" captionTargetPageId="19" captionText="FIGURES 55 – 61. Helicopsyche lara, new species, holotype. 55 — sternum VI process, lateral; 56 — sternum VI process, ventral; 57 — male genitalia, lateral; 58 — male genitalia, dorsal; 59 — male genitalia, ventral; 60 — phallus, lateral; 61 — phallus, ventral." httpUri="https://zenodo.org/record/169469/files/figure.png" pageId="20" pageNumber="21">Fig. 61</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3026500FFA3DE5EFEA8FB647E8DF983" pageId="20" pageNumber="21" type="materials_examined">
|
||
<paragraph id="8BA7368BFFA3DE5EFEA8FB647C50FAAB" blockId="20.[264,1323,284,1590]" pageId="20" pageNumber="21">
|
||
<emphasis id="B96CEA99FFA3DE5EFEA8FB647C43FB7B" bold="true" box="[312,804,1204,1230]" pageId="20" pageNumber="21">
|
||
<typeStatus id="54A38829FFA3DE5EFEA8FB647ECDFB7B" box="[312,426,1204,1230]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
|
||
male:
|
||
<collectingCountry id="F30F761BFFA3DE5EFD97FB647DD8FB7B" box="[519,703,1204,1230]" name="Venezuela" pageId="20" pageNumber="21">VENEZUELA</collectingCountry>
|
||
:
|
||
<taxonomicName id="4C184D08FFA3DE5EFD4CFB647C79FB7B" box="[732,798,1204,1230]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="20" pageNumber="36" phylum="Arthropoda" rank="species" species="lara">Lara</taxonomicName>
|
||
:
|
||
</emphasis>
|
||
Parque Nacional Dinira, Quebrada Buenos Aires,
|
||
<geoCoordinate id="EE2C504CFFA3DE5EFEC8FB0C7E86FB43" box="[344,481,1244,1270]" direction="north" orientation="latitude" pageId="20" pageNumber="21" precision="1" value="9.606783">9°36.407'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE2C504CFFA3DE5EFE7FFB0C7DEEFB43" box="[495,649,1244,1270]" direction="west" orientation="longitude" pageId="20" pageNumber="21" precision="1" value="-70.06963">70°04.178'W</geoCoordinate>
|
||
,
|
||
<quantity id="4CE09B6EFFA3DE5EFD07FB0C7D92FB43" box="[663,757,1244,1270]" metricMagnitude="3" metricUnit="m" metricValue="1.85" pageId="20" pageNumber="21" unit="m" value="1850.0">1850 m</quantity>
|
||
,
|
||
<date id="FFA6104BFFA3DE5EFC93FB0C7CCAFB43" box="[771,941,1244,1270]" pageId="20" pageNumber="21" value="2001-06-18" valueMax="2001-06-19" valueMin="2001-06-18">18–19.vi.2001</date>
|
||
, Holzenthal, Blahnik, Paprocki, & Cressa, (
|
||
<collectionCode id="ED09AE4EFFA3DE5EFE1BFAD47EBBFAAB" box="[395,476,1284,1310]" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="20" pageNumber="21">UMSP</collectionCode>
|
||
000074735) (
|
||
<collectionCode id="ED09AE4EFFA3DE5EFDE9FAD47DADFAAB" box="[633,714,1284,1310]" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="20" pageNumber="21">UMSP</collectionCode>
|
||
, pinned).
|
||
</paragraph>
|
||
<paragraph id="8BA7368BFFA3DE5EFEA8FAFC7C59FA0B" blockId="20.[264,1323,284,1590]" pageId="20" pageNumber="21">
|
||
<emphasis id="B96CEA99FFA3DE5EFEA8FAFC7ED9FAF3" bold="true" box="[312,446,1324,1350]" pageId="20" pageNumber="21">
|
||
<typeStatus id="54A38829FFA3DE5EFEA8FAFC7EDDFAF3" box="[312,442,1324,1350]" pageId="20" pageNumber="21" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
</emphasis>
|
||
same data as
|
||
<typeStatus id="54A38829FFA3DE5EFDE7FAFC7DB9FAF3" box="[631,734,1324,1350]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
|
||
—
|
||
<specimenCount id="9D1EFD02FFA3DE5EFC9FFAFC7C19FAF3" box="[783,894,1324,1350]" pageId="20" pageNumber="21" type="male">19 males</specimenCount>
|
||
,
|
||
<specimenCount id="9D1EFD02FFA3DE5EFC1DFAFC7B68FAF3" box="[909,1039,1324,1350]" pageId="20" pageNumber="21" type="female">19 females</specimenCount>
|
||
(
|
||
<collectionCode id="ED09AE4EFFA3DE5EFBB1FAFC7B15FAF3" box="[1057,1138,1324,1350]" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="20" pageNumber="21">UMSP</collectionCode>
|
||
, pinned); same data as
|
||
<typeStatus id="54A38829FFA3DE5EFEE4FA847EBAFADB" box="[372,477,1364,1390]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
|
||
, except Quebrada Las Minas,
|
||
<geoCoordinate id="EE2C504CFFA3DE5EFCE1FA847C9DFADB" box="[881,1018,1364,1390]" direction="north" orientation="latitude" pageId="20" pageNumber="21" precision="1" value="9.604383">9°36.263'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE2C504CFFA3DE5EFB9FFA847BCDFADB" box="[1039,1194,1364,1390]" direction="west" orientation="longitude" pageId="20" pageNumber="21" precision="1" value="-70.06945">70°04.167'W</geoCoordinate>
|
||
,
|
||
<quantity id="4CE09B6EFFA3DE5EFB50FA847A41FADB" box="[1216,1318,1364,1390]" metricMagnitude="3" metricUnit="m" metricValue="1.825" pageId="20" pageNumber="21" unit="m" value="1825.0">1825 m</quantity>
|
||
,
|
||
<date id="FFA6104BFFA3DE5EFE98FAAC7EEFFA23" box="[264,392,1404,1430]" pageId="20" pageNumber="21" value="2001-06-20">20.vi.2001</date>
|
||
, Holzenthal, Blahnik, Paprocki, & Cressa —
|
||
<specimenCount id="9D1EFD02FFA3DE5EFC0CFAAC7C89FA23" box="[924,1006,1404,1430]" pageId="20" pageNumber="21" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount id="9D1EFD02FFA3DE5EFC6AFAAC7B38FA23" box="[1018,1119,1404,1430]" pageId="20" pageNumber="21" type="female">1 female</specimenCount>
|
||
(
|
||
<collectionCode id="ED09AE4EFFA3DE5EFBFFFAAC7BD9FA23" box="[1135,1214,1404,1430]" pageId="20" pageNumber="21">IZAM</collectionCode>
|
||
, pinned),
|
||
<specimenCount id="9D1EFD02FFA3DE5EFE98FA747E30FA0B" box="[264,343,1444,1470]" pageId="20" pageNumber="21" type="male">1 male</specimenCount>
|
||
(
|
||
<collectionCode id="ED09AE4EFFA3DE5EFEF6FA747EDFFA0B" box="[358,440,1444,1470]" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="20" pageNumber="21">UMSP</collectionCode>
|
||
, pinned),
|
||
<specimenCount id="9D1EFD02FFA3DE5EFDBBFA747D1DFA0B" box="[555,634,1444,1470]" pageId="20" pageNumber="21" type="male">1 male</specimenCount>
|
||
(
|
||
<collectionCode id="ED09AE4EFFA3DE5EFD1AFA747DB6FA0B" box="[650,721,1444,1470]" httpUri="http://grbio.org/cool/4m0z-xq14" name="Swedish Museum of Natural History - Zoological Collections" pageId="20" pageNumber="21">NRM</collectionCode>
|
||
, pinned).
|
||
</paragraph>
|
||
<paragraph id="8BA7368BFFA3DE5EFEA8FA1C7DCCFA53" blockId="20.[264,1323,284,1590]" box="[312,683,1484,1510]" pageId="20" pageNumber="21">
|
||
Distribution.
|
||
<collectingCountry id="F30F761BFFA3DE5EFE4BFA1C7D34FA53" box="[475,595,1484,1510]" name="Venezuela" pageId="20" pageNumber="21">Venezuela</collectingCountry>
|
||
(
|
||
<taxonomicName id="4C184D08FFA3DE5EFDF3FA1C7DF8FA53" box="[611,671,1484,1510]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="20" pageNumber="36" phylum="Arthropoda" rank="species" species="lara">Lara</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA7368BFFA3DE5EFEA8FA247E8DF983" blockId="20.[264,1323,284,1590]" pageId="20" pageNumber="21">
|
||
Etymology.
|
||
<taxonomicName id="4C184D08FFA3DE5EFE40FA247D66F9B8" box="[464,513,1524,1549]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="lara">
|
||
<emphasis id="B96CEA99FFA3DE5EFE40FA247D66F9B8" box="[464,513,1524,1549]" italics="true" pageId="20" pageNumber="21">lara</emphasis>
|
||
</taxonomicName>
|
||
, derived from the
|
||
<typeStatus id="54A38829FFA3DE5EFD70FA247C75F9BB" box="[736,786,1524,1550]" pageId="20" pageNumber="21">type</typeStatus>
|
||
locality,
|
||
<taxonomicName id="4C184D08FFA3DE5EFC11FA247CDDF9BB" box="[897,954,1524,1550]" class="Insecta" family="Helicopsychidae" genus="Helicopsyche" kingdom="Animalia" order="Trichoptera" pageId="20" pageNumber="36" phylum="Arthropoda" rank="species" species="lara">Lara</taxonomicName>
|
||
. The name is to be treated as a noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |