187 lines
19 KiB
XML
187 lines
19 KiB
XML
<document id="B5711F74075876E73DEAF3CC70B51C72" ID-CLB-Dataset="6194" ID-DOI="10.11646/zootaxa.4975.1.3" ID-GBIF-Dataset="529d48db-4299-4e2d-9aaf-6c4178c08b18" ID-ISSN="1175-5326" ID-Zenodo-Dep="4804909" ID-ZooBank="DB7C9028-3EDF-454F-88D0-336624AD1DC4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1622016306479" checkinUser="plazi" docAuthor="Shear, William A. & Marek, Paul E." docDate="2021" docId="03B18797FFFF9D1F4BA50788D9B025B9" docLanguage="en" docName="zootaxa.4975.1.3.pdf" docOrigin="Zootaxa 4975 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Retrorsioides Shear & Marek 2021, new genus" docType="treatment" docVersion="6" lastPageNumber="103" masterDocId="FF88FFEFFFE99D094B32053AD943204D" masterDocTitle="Three new genera and eighteen new species of miniature polydesmid millipedes from the northwestern United States (Diplopoda, Polydesmida, Polydesmidae)" masterLastPageNumber="126" masterPageNumber="81" pageNumber="103" updateTime="1698991024886" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="FD6FD940C4997F2627B23FEEE07A2F78" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="ACD72054AC824F05D86705C0E062B36F">
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<mods:title id="2EC1DDD6F41A529146F1DE35BD5CD1D0">Three new genera and eighteen new species of miniature polydesmid millipedes from the northwestern United States (Diplopoda, Polydesmida, Polydesmidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="B7B63BD1665A550CEE0D4CB9BE9359CF" type="personal">
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<mods:role id="212603398B439913FE6EAA4BF0656935">
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<mods:roleTerm id="F760F6305E14C337E5F00A0985BFFE0C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="89F61B8FB09028DCEE491A5FB1610E1F">Shear, William A.</mods:namePart>
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<mods:affiliation id="3D85CEE07A56BD7AD8D88825CDCBAD66">Professor Emeritus, Department of Biology, Hampden-Sydney College, Hampden-Sydney VA 23943; present address 1950 Price Drive, Farmville VA 23901.</mods:affiliation>
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</mods:name>
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<mods:name id="752ECE715CB84884A891F35807C200F3" type="personal">
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<mods:role id="29526F17949CC6953E14175D84AA37EF">
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<mods:roleTerm id="A84D93C46D54D8A7057562D32B3188B4">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0D49051D086BEC74B848DD74CA093C25">Marek, Paul E.</mods:namePart>
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<mods:nameIdentifier id="B0654BA7F7E1594421E7E49155A025BE" type="ORCID">0000-0002-7048-2514</mods:nameIdentifier>
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<mods:affiliation id="32C73D33B95804CCE723F9185B1B830A">Associate Professor, Department of Entomology, Virginia Polytechnic Institute and State University, Blacksburg, VA 24061. pmarek @ vt. edu; https: // orcid. org / 0000 - 0002 - 7048 - 2514</mods:affiliation>
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<mods:nameIdentifier id="D059049D9A1443DD8DB194419404217A" type="email">pmarek@vt.edu</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource id="503F8921ECEB93036319BA41BB70D5BF">text</mods:typeOfResource>
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<mods:titleInfo id="472EA76D954730584802F07887E3B5AC">
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<mods:title id="8F794B2B3C8192D5996C35692192B99A">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="7EBB9B28E9B98F4DF3B58FF9A09CA01D">
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<mods:date id="C8B86E8652B2CBC2F25D9F9BAFDD6E01">2021</mods:date>
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<mods:detail id="701385726A06735ADD972EC01724C674" type="pubDate">
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<mods:number id="1BE81DAA8A672D6884FE5385B7CA6BB3">2021-05-24</mods:number>
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</mods:detail>
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<mods:detail id="39731191A9A135E2AC1A1A7C0D9463D0" type="volume">
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<mods:number id="F4095F76AFB785B42D92AF347BCA0303">4975</mods:number>
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</mods:detail>
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<mods:detail id="4CEA64B57FA8B167ED35085BF3CC1942" type="issue">
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<mods:number id="83C00EC298C6A4FDCC4837A32F8FC50A">1</mods:number>
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<mods:start id="A106BF770F60457F4B797D4B2AC89A87">81</mods:start>
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<mods:end id="7FD7566CA7901A5AB40060FA7E0F2D2A">126</mods:end>
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<mods:classification id="290859CC9FEF0FA2FD64750B826F7FB9">journal article</mods:classification>
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<mods:identifier id="331BD5BF31A2D331E4A5A8891714586E" type="CLB-Dataset">6194</mods:identifier>
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<mods:identifier id="61403CA24ABE73F91657267AD5B02E3A" type="DOI">10.11646/zootaxa.4975.1.3</mods:identifier>
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<mods:identifier id="477BE698344C8E28935FD0BAE6E02B64" type="GBIF-Dataset">529d48db-4299-4e2d-9aaf-6c4178c08b18</mods:identifier>
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<mods:identifier id="0ABDF80F549A5418484ED09F5B5E3DE7" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="16FD8186E00C450C07EFB1D4A3A2C5C7" type="Zenodo-Dep">4804909</mods:identifier>
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</mods:mods>
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<treatment id="03B18797FFFF9D1F4BA50788D9B025B9" ID-DOI="http://doi.org/10.5281/zenodo.4805017" ID-GBIF-Taxon="182644887" ID-Zenodo-Dep="4805017" LSID="urn:lsid:plazi:treatment:03B18797FFFF9D1F4BA50788D9B025B9" httpUri="http://treatment.plazi.org/id/03B18797FFFF9D1F4BA50788D9B025B9" lastPageNumber="103" pageId="22" pageNumber="103">
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<subSubSection id="C302650AFFFF9D1F4BA50788DBD02280" box="[151,659,690,717]" pageId="22" pageNumber="103" type="nomenclature">
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<paragraph id="8BA73681FFFF9D1F4BA50788DBD02280" blockId="22.[151,659,690,717]" box="[151,659,690,717]" pageId="22" pageNumber="103">
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<heading id="D0EF81EDFFFF9D1F4BA50788DBD02280" bold="true" box="[151,659,690,717]" fontSize="11" level="1" pageId="22" pageNumber="103" reason="1">
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<emphasis id="B96CEA93FFFF9D1F4BA50788DBD02280" bold="true" box="[151,659,690,717]" pageId="22" pageNumber="103">
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<taxonomicName id="4C184D02FFFF9D1F4BA50788DB412280" authority="Shear & Marek" authorityName="Shear & Marek" authorityYear="2021" box="[151,514,690,717]" class="Diplopoda" family="Polydesmidae" genus="Retrorsioides" kingdom="Animalia" order="Polydesmida" pageId="22" pageNumber="103" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B96CEA93FFFF9D1F4BA50788D8742280" bold="true" box="[151,311,690,717]" italics="true" pageId="22" pageNumber="103">Retrorsioides</emphasis>
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Shear & Marek
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</taxonomicName>
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,
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<taxonomicNameLabel id="A25F57E8FFFF9D1F49230789DBD02280" box="[529,659,691,717]" pageId="22" pageNumber="103" rank="genus">new genus</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C302650AFFFF9D1F4BA507C2D8162326" pageId="22" pageNumber="103" type="materials_examined">
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<paragraph id="8BA73681FFFF9D1F4BA507C2DB152342" blockId="22.[151,598,759,783]" box="[151,598,759,783]" pageId="22" pageNumber="103">
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<emphasis id="B96CEA93FFFF9D1F4BA507C2D8552342" box="[151,278,760,783]" italics="true" pageId="22" pageNumber="103">
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<typeStatus id="54A38823FFFF9D1F4BA507C2D9852342" box="[151,198,760,783]" pageId="22" pageNumber="103">Type</typeStatus>
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species
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</emphasis>
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:
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<emphasis id="B96CEA93FFFF9D1F4A1107CDDB5A2342" box="[291,537,759,783]" italics="true" pageId="22" pageNumber="103">
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<taxonomicName id="4C184D02FFFF9D1F4A1107CDDB562342" authority="Shear & Marek, 2021" authorityName="Shear & Marek" authorityYear="2021" box="[291,533,759,783]" class="Diplopoda" family="Polydesmidae" genus="Retrorsioides" kingdom="Animalia" order="Polydesmida" pageId="22" pageNumber="103" phylum="Arthropoda" rank="species" species="castellum" status="sp. nov.">Retrorsioides castellum</taxonomicName>
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,
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</emphasis>
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<taxonomicNameLabel id="A25F57E8FFFF9D1F492D07C2DB152342" box="[543,598,760,783]" pageId="22" pageNumber="103" rank="species">n. sp.</taxonomicNameLabel>
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</paragraph>
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<paragraph id="8BA73681FFFF9D1F4BA5060FD8162326" blockId="22.[151,1436,820,875]" pageId="22" pageNumber="103">
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<emphasis id="B96CEA93FFFF9D1F4BA5060FDB0E2301" box="[151,589,820,844]" italics="true" pageId="22" pageNumber="103">
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Species included:
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<emphasis id="B96CEA93FFFF9D1F4A60060EDB0E2301" bold="true" box="[338,589,820,844]" pageId="22" pageNumber="103">
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<taxonomicName id="4C184D02FFFF9D1F4A60060EDB0B2301" authority="Shear & Marek, 2021" authorityName="Shear & Marek" authorityYear="2021" box="[338,584,820,844]" class="Diplopoda" family="Polydesmidae" genus="Retrorsioides" kingdom="Animalia" order="Polydesmida" pageId="22" pageNumber="103" phylum="Arthropoda" rank="species" species="castellum" status="sp. nov.">Retrorsioides castellum</taxonomicName>
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,
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</emphasis>
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</emphasis>
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<taxonomicNameLabel id="A25F57E8FFFF9D1F4967060FDBD32300" box="[597,656,821,845]" pageId="22" pageNumber="103" rank="species">n. sp.</taxonomicNameLabel>
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,
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<emphasis id="B96CEA93FFFF9D1F49AF060FDA632301" bold="true" box="[669,800,820,844]" italics="true" pageId="22" pageNumber="103">
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<taxonomicName id="4C184D02FFFF9D1F49AF060FDA592301" authority="Shear & Marek, 2021" authorityName="Shear & Marek" authorityYear="2021" box="[669,794,820,844]" class="Diplopoda" family="Polydesmidae" genus="Retrorsioides" kingdom="Animalia" order="Polydesmida" pageId="22" pageNumber="103" phylum="Arthropoda" rank="species" species="linnensis" status="sp. nov.">R. linnensis</taxonomicName>
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,
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</emphasis>
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<taxonomicNameLabel id="A25F57E8FFFF9D1F481A060FDA202300" box="[808,867,821,845]" pageId="22" pageNumber="103" rank="species">n. sp.</taxonomicNameLabel>
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,
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<emphasis id="B96CEA93FFFF9D1F4842060FDA982301" bold="true" box="[880,987,820,844]" italics="true" pageId="22" pageNumber="103">
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<taxonomicName id="4C184D02FFFF9D1F4842060FDA952301" authority="Shear & Marek, 2021" authorityName="Shear & Marek" authorityYear="2021" box="[880,982,820,844]" class="Diplopoda" family="Polydesmidae" genus="Retrorsioides" kingdom="Animalia" order="Polydesmida" pageId="22" pageNumber="103" phylum="Arthropoda" rank="species" species="kittitas" status="sp. nov.">R. kittitas</taxonomicName>
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,
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</emphasis>
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<taxonomicNameLabel id="A25F57E8FFFF9D1F48D0060FDD5E2300" box="[994,1053,821,845]" pageId="22" pageNumber="103" rank="species">n. sp.</taxonomicNameLabel>
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,
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<emphasis id="B96CEA93FFFF9D1F4F18060FDDF02301" bold="true" box="[1066,1203,820,844]" italics="true" pageId="22" pageNumber="103">
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<taxonomicName id="4C184D02FFFF9D1F4F18060FDDED2301" authority="Shear & Marek, 2021" authorityName="Shear & Marek" authorityYear="2021" box="[1066,1198,820,844]" class="Diplopoda" family="Polydesmidae" genus="Retrorsioides" kingdom="Animalia" order="Polydesmida" pageId="22" pageNumber="103" phylum="Arthropoda" rank="species" species="bammerti" status="sp. nov.">R. bammerti</taxonomicName>
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,
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</emphasis>
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<taxonomicNameLabel id="A25F57E8FFFF9D1F4F89060FDDB52300" box="[1211,1270,821,845]" pageId="22" pageNumber="103" rank="species">n. sp.</taxonomicNameLabel>
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, and
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<emphasis id="B96CEA93FFFF9D1F4E1D060FD8162326" bold="true" pageId="22" pageNumber="103">
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<emphasis id="B96CEA93FFFF9D1F4E1D060FD8562327" bold="true" italics="true" pageId="22" pageNumber="103">
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<taxonomicName id="4C184D02FFFF9D1F4E1D060FD8532327" authority="Shear & Marek, 2021" authorityName="Shear & Marek" authorityYear="2021" class="Diplopoda" family="Polydesmidae" genus="Retrorsioides" kingdom="Animalia" order="Polydesmida" pageId="22" pageNumber="103" phylum="Arthropoda" rank="species" species="arboramagna" status="sp. nov.">R. arboramagna</taxonomicName>
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,
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</emphasis>
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<taxonomicNameLabel id="A25F57E8FFFF9D1F4A2E0669D8162326" box="[284,341,851,875]" pageId="22" pageNumber="103" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C302650AFFFF9D1F4BA506A3DAC323B1" pageId="22" pageNumber="103" type="diagnosis">
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<paragraph id="8BA73681FFFF9D1F4BA506A3DAC323B1" blockId="22.[151,1437,921,1524]" pageId="22" pageNumber="103">
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<emphasis id="B96CEA93FFFF9D1F4BA506A3D85123FE" bold="true" box="[151,274,921,947]" pageId="22" pageNumber="103">Diagnosis.</emphasis>
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Clearly related to
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<emphasis id="B96CEA93FFFF9D1F4AD106A0DB1123FE" box="[483,594,922,947]" italics="true" pageId="22" pageNumber="103">
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<taxonomicName id="4C184D02FFFF9D1F4AD106A0DB0C23FE" authorityName="Shelley" authorityYear="2003" box="[483,591,922,947]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="22" pageNumber="103" phylum="Arthropoda" rank="genus">Retrorsia</taxonomicName>
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,
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</emphasis>
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species of this genus are distinct in the gonopod having a prominent, anteriodorsally directed process at about the level of the pulvillus (absent in
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<taxonomicName id="4C184D02FFFF9D1F48850684DD382395" box="[951,1147,957,984]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="22" pageNumber="103" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis id="B96CEA93FFFF9D1F48850684DD63239A" box="[951,1056,958,983]" italics="true" pageId="22" pageNumber="103">Retrorsia</emphasis>
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species
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</taxonomicName>
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), and in having a pulvillar process rather than a pulvillar notch (present in
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<taxonomicName id="4C184D02FFFF9D1F499C06D8DA3723B1" box="[686,884,993,1020]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="22" pageNumber="103" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis id="B96CEA93FFFF9D1F499C06D8DA5423B6" box="[686,791,994,1019]" italics="true" pageId="22" pageNumber="103">Retrorsia</emphasis>
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species
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</taxonomicName>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C302650AFFFF9D1F4BF5013FDCC72452" box="[199,1412,1029,1056]" pageId="22" pageNumber="103" type="etymology">
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<paragraph id="8BA73681FFFF9D1F4BF5013FDCC72452" blockId="22.[151,1437,921,1524]" box="[199,1412,1029,1056]" pageId="22" pageNumber="103">
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<emphasis id="B96CEA93FFFF9D1F4BF5013FD80D2452" bold="true" box="[199,334,1029,1055]" pageId="22" pageNumber="103">Etymology.</emphasis>
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The name of the genus is based on the similarity of the included species to species of
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<emphasis id="B96CEA93FFFF9D1F4E21013CDCC72452" box="[1299,1412,1030,1055]" italics="true" pageId="22" pageNumber="103">
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<taxonomicName id="4C184D02FFFF9D1F4E21013CDC3C2452" authorityName="Shelley" authorityYear="2003" box="[1299,1407,1030,1055]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="22" pageNumber="103" phylum="Arthropoda" rank="genus">Retrorsia</taxonomicName>
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.
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C302650AFFFF9D1F4BF50113D9B025B9" pageId="22" pageNumber="103" type="distribution">
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<paragraph id="8BA73681FFFF9D1F4BF50113DA27240E" blockId="22.[151,1437,921,1524]" box="[199,868,1065,1091]" pageId="22" pageNumber="103">
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<emphasis id="B96CEA93FFFF9D1F4BF50113D822240E" bold="true" box="[199,353,1065,1091]" pageId="22" pageNumber="103">Distribution.</emphasis>
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<collectingRegion id="49DCF863FFFF9D1F4A550113D8AC240E" box="[359,495,1065,1091]" country="United States of America" name="Washington" pageId="22" pageNumber="103">Washington</collectingRegion>
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,
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<collectingRegion id="49DCF863FFFF9D1F4AC90113DB0C240E" box="[507,591,1065,1091]" country="United States of America" name="Oregon" pageId="22" pageNumber="103">Oregon</collectingRegion>
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and northern
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<collectingRegion id="49DCF863FFFF9D1F49D90113DA23240E" box="[747,864,1065,1091]" country="United States of America" name="California" pageId="22" pageNumber="103">California</collectingRegion>
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.
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</paragraph>
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<paragraph id="8BA73681FFFF9D1F4BF50177D9B025B9" blockId="22.[151,1437,921,1524]" pageId="22" pageNumber="103">
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<emphasis id="B96CEA93FFFF9D1F4BF50177D852242A" bold="true" box="[199,273,1101,1127]" pageId="22" pageNumber="103">Notes.</emphasis>
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<taxonomicName id="4C184D02FFFF9D1F4A2B0177DB4A242A" box="[281,521,1101,1127]" class="Diplopoda" family="Polydesmidae" genus="Retrorsioides" kingdom="Animalia" order="Polydesmida" pageId="22" pageNumber="103" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis id="B96CEA93FFFF9D1F4A2B0177D8EC242A" box="[281,431,1101,1127]" italics="true" pageId="22" pageNumber="103">Retrorsioides</emphasis>
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species
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</taxonomicName>
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encompass a wide distribution, from Humboldt Co. in northern
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<collectingRegion id="49DCF863FFFF9D1F4FD50177DC1A242A" box="[1255,1369,1101,1127]" country="United States of America" name="California" pageId="22" pageNumber="103">California</collectingRegion>
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north to Thurston Co.,
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<collectingRegion id="49DCF863FFFF9D1F4A64014BD89C24C6" box="[342,479,1137,1163]" country="United States of America" name="Washington" pageId="22" pageNumber="103">Washington</collectingRegion>
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. The general appearance of the species is much like that of
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<taxonomicName id="4C184D02FFFF9D1F4F490148DC7C24C6" box="[1147,1343,1137,1163]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="22" pageNumber="103" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis id="B96CEA93FFFF9D1F4F490148DDA724C6" box="[1147,1252,1138,1163]" italics="true" pageId="22" pageNumber="103">Retrorsia</emphasis>
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species
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</taxonomicName>
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, and the other polydesmids described herein. In addition to the accessory sensory area on the sixth antennal segment, there may also be a small cluster of sensilla near the distal end of the seventh segment. There may be up to 24 setae in the anterior marginal row on the collum, and alveolate cuticle, while present on the anterior part of the collum, is not seen on the dorsa of any of the anterior metazonites. However, the epiprocts of all the species have alveolate cuticle posteriorly. Males have crassate legs and typical (for this group of genera) sphaerotrichomes on the anterior tarsi. In two of the species, the pygidium or epiproct is swollen and curved ventrally, as it is in
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<taxonomicName id="4C184D02FFFF9D1F4F8A0073DCDE252E" box="[1208,1437,1353,1379]" class="Diplopoda" family="Polydesmidae" genus="Snoqualmia" kingdom="Animalia" order="Polydesmida" pageId="22" pageNumber="103" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis id="B96CEA93FFFF9D1F4F8A0073DC7D252E" box="[1208,1342,1353,1379]" italics="true" pageId="22" pageNumber="103">Snoqualmia</emphasis>
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species.
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</taxonomicName>
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The gonopod prefemora may be bulbous or small and flattened. The gonopod acropodites are shorter and stouter than in
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<taxonomicName id="4C184D02FFFF9D1F4BDE00A8D8EE25E6" box="[236,429,1425,1451]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="22" pageNumber="103" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis id="B96CEA93FFFF9D1F4BDE00A8D81625E6" box="[236,341,1426,1451]" italics="true" pageId="22" pageNumber="103">Retrorsia</emphasis>
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species
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</taxonomicName>
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|
and are not apically recurved, nor are they distally divided. Instead, a long, acute process arises at about the level of the pulvillus, which is not associated with a semicircular notch, but with a short pulvillar process.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |