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<document id="2322D801F0135C7873CB87F3BE00B618" ID-CLB-Dataset="6194" ID-DOI="10.11646/zootaxa.4975.1.3" ID-GBIF-Dataset="529d48db-4299-4e2d-9aaf-6c4178c08b18" ID-ISSN="1175-5326" ID-Zenodo-Dep="4804909" ID-ZooBank="DB7C9028-3EDF-454F-88D0-336624AD1DC4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1622016306479" checkinUser="plazi" docAuthor="Shear, William A. &amp; Marek, Paul E." docDate="2021" docId="03B18797FFF29D144BA500F0DBB22128" docLanguage="en" docName="zootaxa.4975.1.3.pdf" docOrigin="Zootaxa 4975 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Retrorsioides arboramagna Shear &amp; Marek 2021, n. sp." docType="treatment" docVersion="6" lastPageNumber="110" masterDocId="FF88FFEFFFE99D094B32053AD943204D" masterDocTitle="Three new genera and eighteen new species of miniature polydesmid millipedes from the northwestern United States (Diplopoda, Polydesmida, Polydesmidae)" masterLastPageNumber="126" masterPageNumber="81" pageNumber="108" updateTime="1698991024886" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="5426361CF3555B43CA4D38A33A360D0D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3207FC6B58BB2F8A33FF2A62CDEBBB71">
<mods:title id="9A1B521F5F0E195B91430B8CD49E43CC">Three new genera and eighteen new species of miniature polydesmid millipedes from the northwestern United States (Diplopoda, Polydesmida, Polydesmidae)</mods:title>
</mods:titleInfo>
<mods:name id="A0AAF61861CF374B06AB652BEF332102" type="personal">
<mods:role id="1B870DDF89881523DDBEF35CA6BA52BE">
<mods:roleTerm id="6162CBD67DAA3361B7707AB9D67F1586">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2AA13AB81ADF9A37BFBF06DFF3FBF459">Shear, William A.</mods:namePart>
<mods:affiliation id="8FD17AA6F1D9533455202071E0406E17">Professor Emeritus, Department of Biology, Hampden-Sydney College, Hampden-Sydney VA 23943; present address 1950 Price Drive, Farmville VA 23901.</mods:affiliation>
</mods:name>
<mods:name id="D8EB422CFC0434EA81A43349EBF1BD1E" type="personal">
<mods:role id="2FF9842C84AEC40CB6FB0FA61BF8EBCA">
<mods:roleTerm id="50F9B17FBE890DEE79C52B858A7444D8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FFEC2BB9DE9448740BDE0B81F4965ACB">Marek, Paul E.</mods:namePart>
<mods:nameIdentifier id="65692476DECEB3B3B07D04E2236F55D8" type="ORCID">0000-0002-7048-2514</mods:nameIdentifier>
<mods:affiliation id="714552FF3C47B1EAD86B5CBAE2E1F3FD">Associate Professor, Department of Entomology, Virginia Polytechnic Institute and State University, Blacksburg, VA 24061. pmarek @ vt. edu; https: // orcid. org / 0000 - 0002 - 7048 - 2514</mods:affiliation>
<mods:nameIdentifier id="0A07A0411D754A975FA7A22CA732AB3C" type="email">pmarek@vt.edu</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="1F4827654ED355C30923044AD6FBAA26">text</mods:typeOfResource>
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<mods:title id="9F65D0593206C3255D7626578C7BAA4D">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="8475943E4533D9A5B0A46B910884A13B">
<mods:date id="130C0600CE7E5F5C9B30376D1FD674C4">2021</mods:date>
<mods:detail id="9F99170F368085AA65FA62B677A7F491" type="pubDate">
<mods:number id="46709DF0BCB5A17EF0B26BBFE6623AD7">2021-05-24</mods:number>
</mods:detail>
<mods:detail id="E491BAE14872DCBE7C741494748FA496" type="volume">
<mods:number id="EFEC7AA0AF5B10CCA0B6AD6A29D9151D">4975</mods:number>
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<mods:detail id="75AEBC23C3F651A74FBB5A3F487DB3A6" type="issue">
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<mods:start id="8B74A080B0A9608229D97BEE87CB0C81">81</mods:start>
<mods:end id="7723FC65F2619DB0927C787772AACEF8">126</mods:end>
</mods:extent>
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<mods:classification id="8F0492C26253FD4B9196318CC4DEBFD6">journal article</mods:classification>
<mods:identifier id="437F7A150A5D8AA3E9B121DE57E5975E" type="CLB-Dataset">6194</mods:identifier>
<mods:identifier id="307FE5182AF77C5FB5C281D9EB8D4896" type="DOI">10.11646/zootaxa.4975.1.3</mods:identifier>
<mods:identifier id="6D48C9264E34836BB9905C392A762E27" type="GBIF-Dataset">529d48db-4299-4e2d-9aaf-6c4178c08b18</mods:identifier>
<mods:identifier id="036A9B9F9A8679E57A5960E92543B4C3" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="A3E4F1893C65642C5D5E73A633214449" type="Zenodo-Dep">4804909</mods:identifier>
<mods:identifier id="C8B232F4C7242652553A7DFA83BE5D18" type="ZooBank">DB7C9028-3EDF-454F-88D0-336624AD1DC4</mods:identifier>
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<treatment id="03B18797FFF29D144BA500F0DBB22128" ID-DOI="http://doi.org/10.5281/zenodo.4805023" ID-GBIF-Taxon="182644896" ID-Zenodo-Dep="4805023" LSID="urn:lsid:plazi:treatment:03B18797FFF29D144BA500F0DBB22128" httpUri="http://treatment.plazi.org/id/03B18797FFF29D144BA500F0DBB22128" lastPageId="29" lastPageNumber="110" pageId="27" pageNumber="108">
<subSubSection id="C302650AFFF29D124BA500F0DBBC25A8" box="[151,767,1482,1509]" pageId="27" pageNumber="108" type="nomenclature">
<paragraph id="8BA73681FFF29D124BA500F0DBBC25A8" blockId="27.[151,767,1482,1545]" box="[151,767,1482,1509]" pageId="27" pageNumber="108">
<heading id="D0EF81EDFFF29D124BA500F0DBBC25A8" bold="true" box="[151,767,1482,1509]" fontSize="11" level="1" pageId="27" pageNumber="108" reason="1">
<emphasis id="B96CEA93FFF29D124BA500F0DBBC25A8" bold="true" box="[151,767,1482,1509]" pageId="27" pageNumber="108">
<taxonomicName id="4C184D02FFF29D124BA500F0DBF325A8" authority="Shear &amp; Marek" authorityName="Shear &amp; Marek" authorityYear="2021" box="[151,688,1482,1509]" class="Diplopoda" family="Polydesmidae" genus="Retrorsioides" kingdom="Animalia" order="Polydesmida" pageId="27" pageNumber="108" phylum="Arthropoda" rank="species" species="arboramagna" status="sp. nov.">
<emphasis id="B96CEA93FFF29D124BA500F0D8A025A8" bold="true" box="[151,483,1482,1509]" italics="true" pageId="27" pageNumber="108">Retrorsioides arboramagna</emphasis>
Shear &amp; Marek
</taxonomicName>
,
<taxonomicNameLabel id="A25F57E8FFF29D12498F00F1DBBC25A8" box="[701,767,1483,1509]" pageId="27" pageNumber="108" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C302650AFFF29D124BA500D5D8572644" box="[151,276,1519,1545]" pageId="27" pageNumber="108" type="description">
<paragraph id="8BA73681FFF29D124BA500D5D8572644" blockId="27.[151,767,1482,1545]" box="[151,276,1519,1545]" pageId="27" pageNumber="108">
<figureCitation id="13232A04FFF29D124BA500D5D8572644" box="[151,276,1519,1545]" captionStart="FIGURES 79–84" captionStartId="29.[151,264,1877,1901]" captionTargetBox="[151,1436,381,1854]" captionTargetId="figure-101@29.[151,1436,381,1854]" captionTargetPageId="29" captionText="FIGURES 79–84. Retrorsioides arboramagna, n. sp. 79. Head and anterior rings, dorsal view. 80. Midbody ring, dorsal view. 81. Pygidium, lateral view. 82. Gonopods in situ, lateroposterior view. 83. Gonopod tip, mesal view. 84. Pulvillus of gonopod. SEM stub WS33-16. Abbreviations: adp, anteriorly directed process; p, pulvillus; pp, pulvillar process; tz, terminal zone, x, unnamed gonopod process." figureDoi="http://doi.org/10.5281/zenodo.4805015" httpUri="https://zenodo.org/record/4805015/files/figure.png" pageId="27" pageNumber="108">Figs 79–84</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C302650AFFF29D124BA5030DD82726D4" pageId="27" pageNumber="108" type="materials_examined">
<paragraph id="8BA73681FFF29D124BA5030DD82726D4" blockId="27.[151,1437,1590,2013]" pageId="27" pageNumber="108">
<emphasis id="B96CEA93FFF29D124BA5030DD99A261C" bold="true" box="[151,217,1591,1617]" pageId="27" pageNumber="108">Type.</emphasis>
<materialsCitation id="3B703CDCFFF29D124BED030DD82326D4" ID-GBIF-Occurrence="3128519333" collectingDate="2006-12-02" collectionCode="CAS" collectorName="C. Richart. The" county="Humboldt Co." elevation="77" location="Prairie Creek Redwoods State Park" municipality="Male" pageId="27" pageNumber="108" specimenCount="2" stateProvince="California" typeStatus="holotype">
<collectingMunicipality id="6BC3ACFBFFF29D124BED030DD85A261C" box="[223,281,1591,1617]" pageId="27" pageNumber="108">Male</collectingMunicipality>
<typeStatus id="54A38823FFF29D124A2D030DD8C2261C" box="[287,385,1591,1617]" pageId="27" pageNumber="108" type="holotype">holotype</typeStatus>
from
<location id="8EC7605AFFF29D124AF1030DDA0D261C" LSID="urn:lsid:plazi:treatment:03B18797FFF29D144BA500F0DBB22128:8EC7605AFFF29D124AF1030DDA0D261C" box="[451,846,1591,1617]" county="Humboldt Co." municipality="Male" name="Prairie Creek Redwoods State Park" pageId="27" pageNumber="108" stateProvince="California">Prairie Creek Redwoods State Park</location>
, “Big Trees,”
<elevation id="0035D1B2FFF29D1248DB030CDD10261C" box="[1001,1107,1590,1617]" metricMagnitude="1" metricUnit="m" metricValue="7.68096" pageId="27" pageNumber="108" unit="ft" value="252.0">
<quantity id="4CE09B64FFF29D1248DB030CDD69261C" box="[1001,1066,1590,1617]" metricMagnitude="1" metricUnit="m" metricValue="7.68096" pageId="27" pageNumber="108" unit="ft" value="252.0">252 ft</quantity>
asl
</elevation>
, 41.4006°, -124.0003°.
<collectingCounty id="62C64E0DFFF29D124E69030DD8422638" pageId="27" pageNumber="108">Humboldt Co.</collectingCounty>
<collectingRegion id="49DCF863FFF29D124A3A0361D83F2638" box="[264,380,1627,1653]" country="United States of America" name="California" pageId="27" pageNumber="108">California</collectingRegion>
, collected
<date id="FFA61041FFF29D124AC40361DBFD2638" box="[502,702,1627,1653]" pageId="27" pageNumber="108" value="2006-12-02">
<collectingDate id="EFE2E9A9FFF29D124AC40361DBFD2638" box="[502,702,1627,1653]" pageId="27" pageNumber="108" value="2006-12-02">2 December 2006</collectingDate>
</date>
by
<collectorName id="26ED5357FFF29D1249D80361DADA2638" box="[746,921,1627,1653]" pageId="27" pageNumber="108">C. Richart. The</collectorName>
<typeStatus id="54A38823FFF29D1248920361DD412638" box="[928,1026,1627,1653]" pageId="27" pageNumber="108" type="holotype">holotype</typeStatus>
is mounted on SEM stub WS33-16, deposited in
<collectionCode id="ED09AE44FFF29D124A140345D82326D4" box="[294,352,1663,1689]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="27" pageNumber="108" type="Museum">CAS</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C302650AFFF29D124BF50398DBAF26AC" pageId="27" pageNumber="108" type="diagnosis">
<paragraph id="8BA73681FFF29D124BF50398DBAF26AC" blockId="27.[151,1437,1590,2013]" pageId="27" pageNumber="108">
<emphasis id="B96CEA93FFF29D124BF50398D80126F1" bold="true" box="[199,322,1698,1724]" pageId="27" pageNumber="108">Diagnosis.</emphasis>
Distinct from other species of the genus in the extended pulvillus (
<emphasis id="B96CEA93FFF29D124F1E0399DD0026F0" bold="true" box="[1068,1091,1699,1725]" pageId="27" pageNumber="108">p,</emphasis>
<figureCitation id="13232A04FFF29D124F780399DDD826F0" box="[1098,1179,1699,1725]" captionStart="FIGURES 79–84" captionStartId="29.[151,264,1877,1901]" captionTargetBox="[151,1436,381,1854]" captionTargetId="figure-101@29.[151,1436,381,1854]" captionTargetPageId="29" captionText="FIGURES 79–84. Retrorsioides arboramagna, n. sp. 79. Head and anterior rings, dorsal view. 80. Midbody ring, dorsal view. 81. Pygidium, lateral view. 82. Gonopods in situ, lateroposterior view. 83. Gonopod tip, mesal view. 84. Pulvillus of gonopod. SEM stub WS33-16. Abbreviations: adp, anteriorly directed process; p, pulvillus; pp, pulvillar process; tz, terminal zone, x, unnamed gonopod process." figureDoi="http://doi.org/10.5281/zenodo.4805015" httpUri="https://zenodo.org/record/4805015/files/figure.png" pageId="27" pageNumber="108">Fig. 83</figureCitation>
), and a strong tooth (
<emphasis id="B96CEA93FFF29D124EB50399DCDF26F0" bold="true" box="[1415,1436,1699,1725]" pageId="27" pageNumber="108">x,</emphasis>
<figureCitation id="13232A04FFF29D124BA503FDD9AB26AC" box="[151,232,1735,1761]" captionStart="FIGURES 79–84" captionStartId="29.[151,264,1877,1901]" captionTargetBox="[151,1436,381,1854]" captionTargetId="figure-101@29.[151,1436,381,1854]" captionTargetPageId="29" captionText="FIGURES 79–84. Retrorsioides arboramagna, n. sp. 79. Head and anterior rings, dorsal view. 80. Midbody ring, dorsal view. 81. Pygidium, lateral view. 82. Gonopods in situ, lateroposterior view. 83. Gonopod tip, mesal view. 84. Pulvillus of gonopod. SEM stub WS33-16. Abbreviations: adp, anteriorly directed process; p, pulvillus; pp, pulvillar process; tz, terminal zone, x, unnamed gonopod process." figureDoi="http://doi.org/10.5281/zenodo.4805015" httpUri="https://zenodo.org/record/4805015/files/figure.png" pageId="27" pageNumber="108">Fig. 83</figureCitation>
) at the base of the anteriorly directed process.
</paragraph>
</subSubSection>
<subSubSection id="C302650AFFF29D124BF503D1DB232764" pageId="27" pageNumber="108" type="etymology">
<paragraph id="8BA73681FFF29D124BF503D1DB232764" blockId="27.[151,1437,1590,2013]" pageId="27" pageNumber="108">
<emphasis id="B96CEA93FFF29D124BF503D1D80D2748" bold="true" box="[199,334,1771,1797]" pageId="27" pageNumber="108">Etymology.</emphasis>
The species epithet (Latin) is a noun in appostion meaning “big tree” and refers to the
<typeStatus id="54A38823FFF29D124E3F03D1DC7E2748" box="[1293,1341,1771,1797]" pageId="27" pageNumber="108">type</typeStatus>
locality, famous for its gigantic Coast Redwoods.
</paragraph>
</subSubSection>
<subSubSection id="C302650AFFF29D144BF50208D828210C" lastPageId="29" lastPageNumber="110" pageId="27" pageNumber="108" type="description">
<paragraph id="8BA73681FFF29D144BF50208D828210C" blockId="27.[151,1437,1590,2013]" lastBlockId="29.[151,1437,151,357]" lastPageId="29" lastPageNumber="110" pageId="27" pageNumber="108">
<emphasis id="B96CEA93FFF29D124BF50208D81A2701" bold="true" box="[199,345,1842,1868]" pageId="27" pageNumber="108">Description.</emphasis>
Male
<typeStatus id="54A38823FFF29D124A910209DB4A2700" box="[419,521,1843,1869]" pageId="27" pageNumber="108" type="holotype">holotype</typeStatus>
. Length about 3.0 mm, greatest width
<quantity id="4CE09B64FFF29D1248F30208DD692700" box="[961,1066,1842,1869]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="27" pageNumber="108" unit="mm" value="0.45">0.45 mm</quantity>
. Head densely setose, cuticle alveolate (
<figureCitation id="13232A04FFF29D124BCB026DD80C273C" box="[249,335,1879,1905]" captionStart="FIGURES 79–84" captionStartId="29.[151,264,1877,1901]" captionTargetBox="[151,1436,381,1854]" captionTargetId="figure-101@29.[151,1436,381,1854]" captionTargetPageId="29" captionText="FIGURES 79–84. Retrorsioides arboramagna, n. sp. 79. Head and anterior rings, dorsal view. 80. Midbody ring, dorsal view. 81. Pygidium, lateral view. 82. Gonopods in situ, lateroposterior view. 83. Gonopod tip, mesal view. 84. Pulvillus of gonopod. SEM stub WS33-16. Abbreviations: adp, anteriorly directed process; p, pulvillus; pp, pulvillar process; tz, terminal zone, x, unnamed gonopod process." figureDoi="http://doi.org/10.5281/zenodo.4805015" httpUri="https://zenodo.org/record/4805015/files/figure.png" pageId="27" pageNumber="108">Fig. 79</figureCitation>
). Collum (
<figureCitation id="13232A04FFF29D124AF8026DDB5E273C" box="[458,541,1879,1905]" captionStart="FIGURES 79–84" captionStartId="29.[151,264,1877,1901]" captionTargetBox="[151,1436,381,1854]" captionTargetId="figure-101@29.[151,1436,381,1854]" captionTargetPageId="29" captionText="FIGURES 79–84. Retrorsioides arboramagna, n. sp. 79. Head and anterior rings, dorsal view. 80. Midbody ring, dorsal view. 81. Pygidium, lateral view. 82. Gonopods in situ, lateroposterior view. 83. Gonopod tip, mesal view. 84. Pulvillus of gonopod. SEM stub WS33-16. Abbreviations: adp, anteriorly directed process; p, pulvillus; pp, pulvillar process; tz, terminal zone, x, unnamed gonopod process." figureDoi="http://doi.org/10.5281/zenodo.4805015" httpUri="https://zenodo.org/record/4805015/files/figure.png" pageId="27" pageNumber="108">Fig. 79</figureCitation>
) with anterior marginal row of 16 setae. Anterior metazonites with three rows of setae, transitioning to four rows about ring 5, setal tubercles becoming almost obsolete on midbody to posterior rings (
<figureCitation id="13232A04FFF29D124A2302A5D82427F4" box="[273,359,1951,1977]" captionStart="FIGURES 79–84" captionStartId="29.[151,264,1877,1901]" captionTargetBox="[151,1436,381,1854]" captionTargetId="figure-101@29.[151,1436,381,1854]" captionTargetPageId="29" captionText="FIGURES 79–84. Retrorsioides arboramagna, n. sp. 79. Head and anterior rings, dorsal view. 80. Midbody ring, dorsal view. 81. Pygidium, lateral view. 82. Gonopods in situ, lateroposterior view. 83. Gonopod tip, mesal view. 84. Pulvillus of gonopod. SEM stub WS33-16. Abbreviations: adp, anteriorly directed process; p, pulvillus; pp, pulvillar process; tz, terminal zone, x, unnamed gonopod process." figureDoi="http://doi.org/10.5281/zenodo.4805015" httpUri="https://zenodo.org/record/4805015/files/figure.png" pageId="27" pageNumber="108">Fig. 80</figureCitation>
). Alveolate cuticle absent from metazonite posterior to collum. Epiproct (
<figureCitation id="13232A04FFF29D124F9C02A5DC4227F4" box="[1198,1281,1951,1977]" captionStart="FIGURES 79–84" captionStartId="29.[151,264,1877,1901]" captionTargetBox="[151,1436,381,1854]" captionTargetId="figure-101@29.[151,1436,381,1854]" captionTargetPageId="29" captionText="FIGURES 79–84. Retrorsioides arboramagna, n. sp. 79. Head and anterior rings, dorsal view. 80. Midbody ring, dorsal view. 81. Pygidium, lateral view. 82. Gonopods in situ, lateroposterior view. 83. Gonopod tip, mesal view. 84. Pulvillus of gonopod. SEM stub WS33-16. Abbreviations: adp, anteriorly directed process; p, pulvillus; pp, pulvillar process; tz, terminal zone, x, unnamed gonopod process." figureDoi="http://doi.org/10.5281/zenodo.4805015" httpUri="https://zenodo.org/record/4805015/files/figure.png" pageId="27" pageNumber="108">Fig. 81</figureCitation>
) not swollen, short, straight, with alveolate cuticle. Anterior legs crassate, tarsi with sphaerotrichomes. Gonopod (
<figureCitation id="13232A04FFF29D124E2002F9DCD72790" box="[1298,1428,1987,2013]" captionStart="FIGURES 79–84" captionStartId="29.[151,264,1877,1901]" captionTargetBox="[151,1436,381,1854]" captionTargetId="figure-101@29.[151,1436,381,1854]" captionTargetPageId="29" captionText="FIGURES 79–84. Retrorsioides arboramagna, n. sp. 79. Head and anterior rings, dorsal view. 80. Midbody ring, dorsal view. 81. Pygidium, lateral view. 82. Gonopods in situ, lateroposterior view. 83. Gonopod tip, mesal view. 84. Pulvillus of gonopod. SEM stub WS33-16. Abbreviations: adp, anteriorly directed process; p, pulvillus; pp, pulvillar process; tz, terminal zone, x, unnamed gonopod process." figureDoi="http://doi.org/10.5281/zenodo.4805015" httpUri="https://zenodo.org/record/4805015/files/figure.png" pageId="27" pageNumber="108">Figs 82–84</figureCitation>
) with moderately inflated prefemorite. Acropodite long, robust, basally thickened. Pulvillus (
<emphasis id="B96CEA93FFF49D144F9505ADDDFE20FC" bold="true" box="[1191,1213,151,177]" pageId="29" pageNumber="110">p,</emphasis>
<figureCitation id="13232A04FFF49D144FF705ADDC5B20FC" box="[1221,1304,151,177]" captionStart="FIGURES 79–84" captionStartId="29.[151,264,1877,1901]" captionTargetBox="[151,1436,381,1854]" captionTargetId="figure-101@29.[151,1436,381,1854]" captionTargetPageId="29" captionText="FIGURES 79–84. Retrorsioides arboramagna, n. sp. 79. Head and anterior rings, dorsal view. 80. Midbody ring, dorsal view. 81. Pygidium, lateral view. 82. Gonopods in situ, lateroposterior view. 83. Gonopod tip, mesal view. 84. Pulvillus of gonopod. SEM stub WS33-16. Abbreviations: adp, anteriorly directed process; p, pulvillus; pp, pulvillar process; tz, terminal zone, x, unnamed gonopod process." figureDoi="http://doi.org/10.5281/zenodo.4805015" httpUri="https://zenodo.org/record/4805015/files/figure.png" pageId="29" pageNumber="110">Fig. 83</figureCitation>
;
<figureCitation id="13232A04FFF49D144E1705ADDC3520FC" box="[1317,1398,151,177]" captionStart="FIGURES 79–84" captionStartId="29.[151,264,1877,1901]" captionTargetBox="[151,1436,381,1854]" captionTargetId="figure-101@29.[151,1436,381,1854]" captionTargetPageId="29" captionText="FIGURES 79–84. Retrorsioides arboramagna, n. sp. 79. Head and anterior rings, dorsal view. 80. Midbody ring, dorsal view. 81. Pygidium, lateral view. 82. Gonopods in situ, lateroposterior view. 83. Gonopod tip, mesal view. 84. Pulvillus of gonopod. SEM stub WS33-16. Abbreviations: adp, anteriorly directed process; p, pulvillus; pp, pulvillar process; tz, terminal zone, x, unnamed gonopod process." figureDoi="http://doi.org/10.5281/zenodo.4805015" httpUri="https://zenodo.org/record/4805015/files/figure.png" pageId="29" pageNumber="110">Fig. 84</figureCitation>
) in distal third of acropodite, its dorsal side prominently extended. Pulvillar process (
<emphasis id="B96CEA93FFF49D144F000581DD1B2098" bold="true" box="[1074,1112,187,213]" pageId="29" pageNumber="110">pp,</emphasis>
<figureCitation id="13232A04FFF49D144F520581DDF22098" box="[1120,1201,187,213]" captionStart="FIGURES 79–84" captionStartId="29.[151,264,1877,1901]" captionTargetBox="[151,1436,381,1854]" captionTargetId="figure-101@29.[151,1436,381,1854]" captionTargetPageId="29" captionText="FIGURES 79–84. Retrorsioides arboramagna, n. sp. 79. Head and anterior rings, dorsal view. 80. Midbody ring, dorsal view. 81. Pygidium, lateral view. 82. Gonopods in situ, lateroposterior view. 83. Gonopod tip, mesal view. 84. Pulvillus of gonopod. SEM stub WS33-16. Abbreviations: adp, anteriorly directed process; p, pulvillus; pp, pulvillar process; tz, terminal zone, x, unnamed gonopod process." figureDoi="http://doi.org/10.5281/zenodo.4805015" httpUri="https://zenodo.org/record/4805015/files/figure.png" pageId="29" pageNumber="110">Fig. 83</figureCitation>
) short, thin, directed posteriorly. Anteriorly directed process (
<emphasis id="B96CEA93FFF49D14496B05E5DBCD20B4" bold="true" box="[601,654,223,249]" pageId="29" pageNumber="110">adp,</emphasis>
<figureCitation id="13232A04FFF49D1449A105E5DBA020B4" box="[659,739,223,249]" captionStart="FIGURES 79–84" captionStartId="29.[151,264,1877,1901]" captionTargetBox="[151,1436,381,1854]" captionTargetId="figure-101@29.[151,1436,381,1854]" captionTargetPageId="29" captionText="FIGURES 79–84. Retrorsioides arboramagna, n. sp. 79. Head and anterior rings, dorsal view. 80. Midbody ring, dorsal view. 81. Pygidium, lateral view. 82. Gonopods in situ, lateroposterior view. 83. Gonopod tip, mesal view. 84. Pulvillus of gonopod. SEM stub WS33-16. Abbreviations: adp, anteriorly directed process; p, pulvillus; pp, pulvillar process; tz, terminal zone, x, unnamed gonopod process." figureDoi="http://doi.org/10.5281/zenodo.4805015" httpUri="https://zenodo.org/record/4805015/files/figure.png" pageId="29" pageNumber="110">Fig. 83</figureCitation>
) long, tapering to acute point, without small teeth along length but with single, sharp, slightly curved tooth (
<emphasis id="B96CEA93FFF49D1449BC0439DBE02150" bold="true" box="[654,675,259,285]" pageId="29" pageNumber="110">x,</emphasis>
<figureCitation id="13232A04FFF49D1449980439DBB92150" box="[682,762,259,285]" captionStart="FIGURES 79–84" captionStartId="29.[151,264,1877,1901]" captionTargetBox="[151,1436,381,1854]" captionTargetId="figure-101@29.[151,1436,381,1854]" captionTargetPageId="29" captionText="FIGURES 79–84. Retrorsioides arboramagna, n. sp. 79. Head and anterior rings, dorsal view. 80. Midbody ring, dorsal view. 81. Pygidium, lateral view. 82. Gonopods in situ, lateroposterior view. 83. Gonopod tip, mesal view. 84. Pulvillus of gonopod. SEM stub WS33-16. Abbreviations: adp, anteriorly directed process; p, pulvillus; pp, pulvillar process; tz, terminal zone, x, unnamed gonopod process." figureDoi="http://doi.org/10.5281/zenodo.4805015" httpUri="https://zenodo.org/record/4805015/files/figure.png" pageId="29" pageNumber="110">Fig. 83</figureCitation>
) at base. Terminal zone (
<emphasis id="B96CEA93FFF49D144F260439DD722150" bold="true" box="[1044,1073,259,285]" pageId="29" pageNumber="110">tz,</emphasis>
<figureCitation id="13232A04FFF49D144F0A0439DDCB2150" box="[1080,1160,259,285]" captionStart="FIGURES 79–84" captionStartId="29.[151,264,1877,1901]" captionTargetBox="[151,1436,381,1854]" captionTargetId="figure-101@29.[151,1436,381,1854]" captionTargetPageId="29" captionText="FIGURES 79–84. Retrorsioides arboramagna, n. sp. 79. Head and anterior rings, dorsal view. 80. Midbody ring, dorsal view. 81. Pygidium, lateral view. 82. Gonopods in situ, lateroposterior view. 83. Gonopod tip, mesal view. 84. Pulvillus of gonopod. SEM stub WS33-16. Abbreviations: adp, anteriorly directed process; p, pulvillus; pp, pulvillar process; tz, terminal zone, x, unnamed gonopod process." figureDoi="http://doi.org/10.5281/zenodo.4805015" httpUri="https://zenodo.org/record/4805015/files/figure.png" pageId="29" pageNumber="110">Fig. 83</figureCitation>
) not bifurcate, flattened. Females unknown.
</paragraph>
</subSubSection>
<caption id="DF676609FFF59D154BA5022FDD8127D4" ID-DOI="http://doi.org/10.5281/zenodo.4805007" ID-Zenodo-Dep="4805007" httpUri="https://zenodo.org/record/4805007/files/figure.png" pageId="28" pageNumber="109" startId="28.[151,264,1813,1837]" targetBox="[151,1436,182,1786]" targetPageId="28">
<paragraph id="8BA73681FFF59D154BA5022FDD8127D4" blockId="28.[151,1437,1813,1945]" pageId="28" pageNumber="109">
<emphasis id="B96CEA93FFF59D154BA5022FD80C2760" bold="true" box="[151,335,1813,1837]" pageId="28" pageNumber="109">FIGURES 73–78</emphasis>
. Species of
<taxonomicName id="4C184D02FFF59D154AFD022FDB192760" authority="Shear &amp; Marek, 2021" authorityName="Shear &amp; Marek" authorityYear="2021" box="[463,602,1813,1837]" class="Diplopoda" family="Polydesmidae" genus="Retrorsioides" kingdom="Animalia" order="Polydesmida" pageId="28" pageNumber="109" phylum="Arthropoda" rank="genus" status="gen. nov.">Retrorsioides</taxonomicName>
,
<taxonomicNameLabel id="A25F57E8FFF59D154955022FDBEB2760" box="[615,680,1813,1837]" pageId="28" pageNumber="109" rank="genus">
<emphasis id="B96CEA93FFF59D154955022FDBE12760" bold="true" box="[615,674,1813,1837]" pageId="28" pageNumber="109">n.gen</emphasis>
.
</taxonomicNameLabel>
<emphasis id="B96CEA93FFF59D154982022FDBAC2760" bold="true" box="[688,751,1813,1837]" pageId="28" pageNumber="109">73–74</emphasis>
.
<emphasis id="B96CEA93FFF59D1549CF022FDD7C2760" bold="true" box="[765,1087,1813,1837]" pageId="28" pageNumber="109">
<taxonomicName id="4C184D02FFF59D1549CF022FDA962760" authority="Shear &amp; Marek, 2021" authorityName="Shear &amp; Marek" authorityYear="2021" box="[765,981,1813,1837]" class="Diplopoda" family="Polydesmidae" genus="Retrorsioides" kingdom="Animalia" order="Polydesmida" pageId="28" pageNumber="109" phylum="Arthropoda" rank="species" species="kittitas" status="sp. nov.">
<emphasis id="B96CEA93FFF59D1549CF022FDA962760" bold="true" box="[765,981,1813,1837]" italics="true" pageId="28" pageNumber="109">Retrorsioides kittitas</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25F57E8FFF59D1548D1022FDD5D2760" box="[995,1054,1813,1837]" pageId="28" pageNumber="109" rank="species">n. sp.</taxonomicNameLabel>
73
</emphasis>
. Gonopods
<emphasis id="B96CEA93FFF59D154F8F022CDDBD2760" box="[1213,1278,1814,1837]" italics="true" pageId="28" pageNumber="109">in situ</emphasis>
, posteriolateral view.
<emphasis id="B96CEA93FFF59D154BE60203D9AE271C" bold="true" box="[212,237,1849,1873]" pageId="28" pageNumber="109">74</emphasis>
. Same, posterior view.
<emphasis id="B96CEA93FFF59D154AEC0203D8B4271C" bold="true" box="[478,503,1849,1873]" pageId="28" pageNumber="109">75</emphasis>
. Gonopod tip, lateral view. SEM stub WS32-17.
<emphasis id="B96CEA93FFF59D1548DC0203DD6E271C" bold="true" box="[1006,1069,1849,1873]" pageId="28" pageNumber="109">76–78</emphasis>
.
<emphasis id="B96CEA93FFF59D154F080203DCD5271C" bold="true" box="[1082,1430,1849,1873]" pageId="28" pageNumber="109">
<taxonomicName id="4C184D02FFF59D154F080203DC6C271C" authority="Shear &amp; Marek, 2021" authorityName="Shear &amp; Marek" authorityYear="2021" box="[1082,1327,1849,1873]" class="Diplopoda" family="Polydesmidae" genus="Retrorsioides" kingdom="Animalia" order="Polydesmida" pageId="28" pageNumber="109" phylum="Arthropoda" rank="species" species="bammerti" status="sp. nov.">
<emphasis id="B96CEA93FFF59D154F080203DC6C271C" bold="true" box="[1082,1327,1849,1873]" italics="true" pageId="28" pageNumber="109">Retrorsioides bammerti</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25F57E8FFF59D154E0E0203DC35271C" box="[1340,1398,1849,1873]" pageId="28" pageNumber="109" rank="species">n. sp.</taxonomicNameLabel>
76
</emphasis>
. Head and anterior rings, dorsal view.
<emphasis id="B96CEA93FFF59D1549230267DB692738" bold="true" box="[529,554,1885,1909]" pageId="28" pageNumber="109">77</emphasis>
. Midbody ring, dorsal view.
<emphasis id="B96CEA93FFF59D15487D0267DA2C2738" bold="true" box="[847,879,1885,1909]" pageId="28" pageNumber="109">78.</emphasis>
Pygidium, ventral view. SEM stub WS33-1. Abbreviations:
<emphasis id="B96CEA93FFF59D154BE402BBD9AD27D4" bold="true" box="[214,238,1921,1945]" pageId="28" pageNumber="109">ac</emphasis>
, acropodite;
<emphasis id="B96CEA93FFF59D154A4102BBD8DF27D4" bold="true" box="[371,412,1921,1945]" pageId="28" pageNumber="109">adp</emphasis>
, anteriorly directed process;
<emphasis id="B96CEA93FFF59D1549F002BBDB9327D4" bold="true" box="[706,720,1921,1945]" pageId="28" pageNumber="109">p</emphasis>
, pulvillus;
<emphasis id="B96CEA93FFF59D15487102BBDA1927D4" bold="true" box="[835,858,1921,1945]" pageId="28" pageNumber="109">pf</emphasis>
, prefemorite;
<emphasis id="B96CEA93FFF59D1548D802BBDD4527D4" bold="true" box="[1002,1030,1921,1945]" pageId="28" pageNumber="109">pp</emphasis>
, pulvillar process.
</paragraph>
</caption>
<subSubSection id="C302650AFFF49D144BF50470DBB22128" box="[199,753,330,357]" pageId="29" pageNumber="110" type="distribution">
<paragraph id="8BA73681FFF49D144BF50470DBB22128" blockId="29.[151,1437,151,357]" box="[199,753,330,357]" pageId="29" pageNumber="110">
<emphasis id="B96CEA93FFF49D144BF50470D8222129" bold="true" box="[199,353,330,356]" pageId="29" pageNumber="110">Distribution.</emphasis>
Known only from the
<typeStatus id="54A38823FFF49D1449520471DBD32128" box="[608,656,331,357]" pageId="29" pageNumber="110">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>
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