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<document id="389D0654D423378CC4F931157499D2EB" ID-CLB-Dataset="6194" ID-DOI="10.11646/zootaxa.4975.1.3" ID-GBIF-Dataset="529d48db-4299-4e2d-9aaf-6c4178c08b18" ID-ISSN="1175-5326" ID-Zenodo-Dep="4804909" ID-ZooBank="DB7C9028-3EDF-454F-88D0-336624AD1DC4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1622016306479" checkinUser="plazi" docAuthor="Shear, William A. &amp; Marek, Paul E." docDate="2021" docId="03B18797FFE59D054BA507C3DAAC27EE" docLanguage="en" docName="zootaxa.4975.1.3.pdf" docOrigin="Zootaxa 4975 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Retrorsia Shelley 2003" docType="treatment" docVersion="8" lastPageNumber="93" masterDocId="FF88FFEFFFE99D094B32053AD943204D" masterDocTitle="Three new genera and eighteen new species of miniature polydesmid millipedes from the northwestern United States (Diplopoda, Polydesmida, Polydesmidae)" masterLastPageNumber="126" masterPageNumber="81" pageNumber="93" updateTime="1698991024886" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="23AC04CAF06308D1D13D7EDA3860214F">Three new genera and eighteen new species of miniature polydesmid millipedes from the northwestern United States (Diplopoda, Polydesmida, Polydesmidae)</mods:title>
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<mods:name id="88B5E3BB7D64EBEF743E0012C8585800" type="personal">
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<mods:namePart id="140501AFD58583E954E5CA1B64461FFB">Shear, William A.</mods:namePart>
<mods:affiliation id="9560936AB6DCED86D55C6EC7A9346B1B">Professor Emeritus, Department of Biology, Hampden-Sydney College, Hampden-Sydney VA 23943; present address 1950 Price Drive, Farmville VA 23901.</mods:affiliation>
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<mods:namePart id="764105702DB9095DE4AD69679C7FCFB2">Marek, Paul E.</mods:namePart>
<mods:nameIdentifier id="5DEA3BEA77E28ED929D612E117B43349" type="ORCID">0000-0002-7048-2514</mods:nameIdentifier>
<mods:affiliation id="8B9B4991093F0E8C8AB45AF58DF2EBE9">Associate Professor, Department of Entomology, Virginia Polytechnic Institute and State University, Blacksburg, VA 24061. pmarek @ vt. edu; https: // orcid. org / 0000 - 0002 - 7048 - 2514</mods:affiliation>
<mods:nameIdentifier id="89E6F433FD12337495662DD919A3AF1B" type="email">pmarek@vt.edu</mods:nameIdentifier>
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<mods:date id="A26A4448EF9AA2D26466851F03768B53">2021</mods:date>
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<mods:number id="C36BF238A3CEDE926B31A93DA8700DDD">2021-05-24</mods:number>
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<mods:number id="710D4023451FA243911AA0DA644E95E8">4975</mods:number>
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<treatment id="03B18797FFE59D054BA507C3DAAC27EE" ID-DOI="http://doi.org/10.5281/zenodo.4804988" ID-GBIF-Taxon="182644906" ID-Zenodo-Dep="4804988" LSID="urn:lsid:plazi:treatment:03B18797FFE59D054BA507C3DAAC27EE" httpUri="http://treatment.plazi.org/id/03B18797FFE59D054BA507C3DAAC27EE" lastPageNumber="93" pageId="12" pageNumber="93">
<subSubSection id="C302650AFFE59D054BA507C3DB4B2359" box="[151,520,761,788]" pageId="12" pageNumber="93" type="nomenclature">
<paragraph id="8BA73681FFE59D054BA507C3DB4B2359" blockId="12.[151,520,761,788]" box="[151,520,761,788]" pageId="12" pageNumber="93">
<heading id="D0EF81EDFFE59D054BA507C3DB4B2359" bold="true" box="[151,520,761,788]" fontSize="11" level="1" pageId="12" pageNumber="93" reason="1">
<emphasis id="B96CEA93FFE59D054BA507C3DB4B2359" bold="true" box="[151,520,761,788]" pageId="12" pageNumber="93">
Genus
<taxonomicName id="4C184D02FFE59D054BDC07C0DB4B2359" ID-CoL="7PXJN" authority="Shelley, 2003" authorityName="Shelley" authorityYear="2003" box="[238,520,761,788]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="B96CEA93FFE59D054BDC07C0D81D2359" bold="true" box="[238,350,762,788]" italics="true" pageId="12" pageNumber="93">Retrorsia</emphasis>
<bibRefCitation id="EF894B70FFE59D054A5407C3DB4B2359" author="Shelley, R. M." box="[358,520,761,788]" pageId="12" pageNumber="93" pagination="1 - 12" refId="ref20733" refString="Shelley, R. M. (2003) A new polydesmid genus and two new species from Oregon and Washington, U. S. A., with a review of Bidentogon Buckett &amp; Gardner, 1968, and a summary of the family in western North America (Polydesmida: Polydesmidae). Zootaxa, 296, 1 - 12. https: // doi. org / 10.11646 / zootaxa. 296.1.1" type="journal article" year="2003">Shelley, 2003</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C302650AFFE59D054BA50605DB8623FF" pageId="12" pageNumber="93" type="reference_group">
<paragraph id="8BA73681FFE59D054BA50605DAC5231B" blockId="12.[151,902,830,855]" box="[151,902,830,855]" pageId="12" pageNumber="93">
<treatmentCitationGroup id="AB0811AFFFE59D054BA50605DAC5231B" box="[151,902,830,855]" pageId="12" pageNumber="93">
<treatmentCitation id="0AB91090FFE59D054BA50605D8DE231B" ID-DOI="http://doi.org/10.5281/zenodo.5014401" ID-Zenodo-Dep="5014401" author="Shelley, R. M." box="[151,413,830,854]" httpUri="http://treatment.plazi.org/id/03A3096E3A17FFF6F462FA3E4E25C1DB" page="3" pageId="12" pageNumber="93" year="2003">
<taxonomicName id="4C184D02FFE59D054BA50605D8DE231B" ID-CoL="7PXJN" authority="Shelley, 2003: 3" authorityName="Shelley" authorityPageNumber="3" authorityYear="2003" box="[151,413,830,854]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="B96CEA93FFE59D054BA50605D9B5231B" box="[151,246,831,854]" italics="true" pageId="12" pageNumber="93">Retrorsia</emphasis>
<bibRefCitation id="EF894B70FFE59D054BCE0604D8DE231B" author="Shelley, R. M." box="[252,413,830,854]" pageId="12" pageNumber="93" pagination="1 - 12" refId="ref20733" refString="Shelley, R. M. (2003) A new polydesmid genus and two new species from Oregon and Washington, U. S. A., with a review of Bidentogon Buckett &amp; Gardner, 1968, and a summary of the family in western North America (Polydesmida: Polydesmidae). Zootaxa, 296, 1 - 12. https: // doi. org / 10.11646 / zootaxa. 296.1.1" type="journal article" year="2003">Shelley, 2003:3</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="54A38823FFE59D054A940605D89B231A" box="[422,472,831,855]" pageId="12" pageNumber="93">Type</typeStatus>
species,
<taxonomicName id="4C184D02FFE59D0549060605DAC1231B" ID-CoL="78RVF" authority="Shelley, 2003" authorityName="Shelley" authorityYear="2003" box="[564,898,830,854]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="93" phylum="Arthropoda" rank="species" species="leonardi">
<emphasis id="B96CEA93FFE59D0549060605DBAD231B" box="[564,750,830,854]" italics="true" pageId="12" pageNumber="93">Retrorsia leonardi</emphasis>
<bibRefCitation id="EF894B70FFE59D0549C70604DAC1231B" author="Shelley, R. M." box="[757,898,830,854]" pageId="12" pageNumber="93" pagination="1 - 12" refId="ref20733" refString="Shelley, R. M. (2003) A new polydesmid genus and two new species from Oregon and Washington, U. S. A., with a review of Bidentogon Buckett &amp; Gardner, 1968, and a summary of the family in western North America (Polydesmida: Polydesmidae). Zootaxa, 296, 1 - 12. https: // doi. org / 10.11646 / zootaxa. 296.1.1" type="journal article" year="2003">Shelley, 2003</bibRefCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA73681FFE59D054BA50646DB8623FF" blockId="12.[151,1436,891,946]" pageId="12" pageNumber="93">
<emphasis id="B96CEA93FFE59D054BA50646D80823D9" box="[151,331,892,916]" italics="true" pageId="12" pageNumber="93">Species included:</emphasis>
<taxonomicName id="4C184D02FFE59D054A610646DBE523D9" ID-CoL="78RVF" authority="Shelley, 2003" authorityName="Shelley" authorityYear="2003" box="[339,678,892,916]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="93" phylum="Arthropoda" rank="species" species="leonardi">
<emphasis id="B96CEA93FFE59D054A610646DB4C23D9" box="[339,527,892,916]" italics="true" pageId="12" pageNumber="93">Retrorsia leonardi</emphasis>
<bibRefCitation id="EF894B70FFE59D0549250646DBE523D9" author="Shelley, R. M." box="[535,678,892,916]" pageId="12" pageNumber="93" pagination="1 - 12" refId="ref20733" refString="Shelley, R. M. (2003) A new polydesmid genus and two new species from Oregon and Washington, U. S. A., with a review of Bidentogon Buckett &amp; Gardner, 1968, and a summary of the family in western North America (Polydesmida: Polydesmidae). Zootaxa, 296, 1 - 12. https: // doi. org / 10.11646 / zootaxa. 296.1.1" type="journal article" year="2003">Shelley, 2003</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C184D02FFE59D0549800646DD5E23D9" ID-CoL="6X47F" authority="Shelley, 2003" authorityName="Shelley" authorityYear="2003" box="[690,1053,892,916]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="93" phylum="Arthropoda" rank="species" species="benedictae">
<emphasis id="B96CEA93FFE59D0549800646DAC423D9" box="[690,903,892,916]" italics="true" pageId="12" pageNumber="93">Retrorsia benedictae</emphasis>
<bibRefCitation id="EF894B70FFE59D0548BC0646DD5E23D9" author="Shelley, R. M." box="[910,1053,892,916]" pageId="12" pageNumber="93" pagination="1 - 12" refId="ref20733" refString="Shelley, R. M. (2003) A new polydesmid genus and two new species from Oregon and Washington, U. S. A., with a review of Bidentogon Buckett &amp; Gardner, 1968, and a summary of the family in western North America (Polydesmida: Polydesmidae). Zootaxa, 296, 1 - 12. https: // doi. org / 10.11646 / zootaxa. 296.1.1" type="journal article" year="2003">Shelley, 2003</bibRefCitation>
</taxonomicName>
;
<emphasis id="B96CEA93FFE59D054F1B0641D8D923FF" bold="true" pageId="12" pageNumber="93">
<taxonomicName id="4C184D02FFE59D054F1B0641DDA223DE" authority="Shear &amp; Marek, 2021" authorityName="Shear &amp; Marek" authorityYear="2021" box="[1065,1249,891,915]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="93" phylum="Arthropoda" rank="species" species="richarti" status="sp. nov.">
<emphasis id="B96CEA93FFE59D054F1B0641DDA223DE" bold="true" box="[1065,1249,891,915]" italics="true" pageId="12" pageNumber="93">Retrorsia richarti</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25F57E8FFE59D054FC20646DC6E23D9" box="[1264,1325,892,916]" pageId="12" pageNumber="93" rank="species">n. sp.</taxonomicNameLabel>
;
<taxonomicName id="4C184D02FFE59D054E090641D81723FF" authority="Shear &amp; Marek, 2021" authorityName="Shear &amp; Marek" authorityYear="2021" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="93" phylum="Arthropoda" rank="species" species="simplicissima" status="sp. nov.">
<emphasis id="B96CEA93FFE59D054E090641D81723FF" bold="true" italics="true" pageId="12" pageNumber="93">Retrorsia simplicissima</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25F57E8FFE59D054A5306A0D8D923FF" box="[353,410,922,946]" pageId="12" pageNumber="93" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and
<emphasis id="B96CEA93FFE59D054AF906A0DBC523FF" bold="true" box="[459,646,922,946]" pageId="12" pageNumber="93">
<taxonomicName id="4C184D02FFE59D054AF906A0DB3C23FF" authority="Shear &amp; Marek, 2021" authorityName="Shear &amp; Marek" authorityYear="2021" box="[459,639,922,946]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="93" phylum="Arthropoda" rank="species" species="gracilis" status="sp. nov.">
<emphasis id="B96CEA93FFE59D054AF906A0DB3C23FF" bold="true" box="[459,639,922,946]" italics="true" pageId="12" pageNumber="93">Retrorsia gracilis</emphasis>
</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A25F57E8FFE59D0549BE06A0DB8623FF" box="[652,709,922,946]" pageId="12" pageNumber="93" rank="species">
<emphasis id="B96CEA93FFE59D0549BE06A0DBFC23FF" bold="true" box="[652,703,922,946]" pageId="12" pageNumber="93">n. sp</emphasis>
.
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C302650AFFE59D054BA506DADC46242B" pageId="12" pageNumber="93" type="diagnosis">
<paragraph id="8BA73681FFE59D054BA506DADC46242B" blockId="12.[151,1437,992,1955]" pageId="12" pageNumber="93">
<emphasis id="B96CEA93FFE59D054BA506DAD85123B7" bold="true" box="[151,274,992,1018]" pageId="12" pageNumber="93">Diagnosis.</emphasis>
The diagnosis provided by
<bibRefCitation id="EF894B70FFE59D05496206DADBB423B6" author="Shelley, R. M." box="[592,759,992,1019]" pageId="12" pageNumber="93" pagination="1 - 12" refId="ref20733" refString="Shelley, R. M. (2003) A new polydesmid genus and two new species from Oregon and Washington, U. S. A., with a review of Bidentogon Buckett &amp; Gardner, 1968, and a summary of the family in western North America (Polydesmida: Polydesmidae). Zootaxa, 296, 1 - 12. https: // doi. org / 10.11646 / zootaxa. 296.1.1" type="journal article" year="2003">Shelley (2003)</bibRefCitation>
is actually a description and does not compare
<taxonomicName id="4C184D02FFE59D054E2706DBDC3D23B7" authorityName="Shelley" authorityYear="2003" box="[1301,1406,993,1018]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="B96CEA93FFE59D054E2706DBDC3D23B7" box="[1301,1406,993,1018]" italics="true" pageId="12" pageNumber="93">Retrorsia</emphasis>
</taxonomicName>
to other polydesmid genera. However, he implied that the retrorse branch of the terminal zone of the gonopod distinguished
<taxonomicName id="4C184D02FFE59D054BC70113D8F6240F" box="[245,437,1064,1090]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="93" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B96CEA93FFE59D054BC70113D81D240F" box="[245,350,1065,1090]" italics="true" pageId="12" pageNumber="93">Retrorsia</emphasis>
species
</taxonomicName>
from all others. This branch is very obvious in the two species he described, but less so in the three new species described below. Nevertheless it is present and serves as a diagnostic character.
</paragraph>
</subSubSection>
<subSubSection id="C302650AFFE59D054BF5014ADAAC27EE" pageId="12" pageNumber="93" type="distribution">
<paragraph id="8BA73681FFE59D054BF5014ADDBE24C6" blockId="12.[151,1437,992,1955]" box="[199,1277,1136,1163]" pageId="12" pageNumber="93">
<emphasis id="B96CEA93FFE59D054BF5014AD82224C7" bold="true" box="[199,353,1136,1162]" pageId="12" pageNumber="93">Distribution.</emphasis>
Northwestern
<collectingRegion id="49DCF863FFE59D054935014BDB1824C6" box="[519,603,1137,1163]" country="United States of America" name="Oregon" pageId="12" pageNumber="93">Oregon</collectingRegion>
and western
<collectingRegion id="49DCF863FFE59D0549DD014ADA3424C7" box="[751,887,1136,1162]" country="United States of America" name="Washington" pageId="12" pageNumber="93">Washington</collectingRegion>
, in or west of the Cascade Ranges.
</paragraph>
<paragraph id="8BA73681FFE59D054BF501AFDB462687" blockId="12.[151,1437,992,1955]" pageId="12" pageNumber="93">
<emphasis id="B96CEA93FFE59D054BF501AFD85224E2" bold="true" box="[199,273,1173,1199]" pageId="12" pageNumber="93">Notes.</emphasis>
<taxonomicName id="4C184D02FFE59D054A2801AFD89F24E3" box="[282,476,1172,1198]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="93" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B96CEA93FFE59D054A2801AFD8C024E3" box="[282,387,1173,1198]" italics="true" pageId="12" pageNumber="93">Retrorsia</emphasis>
species
</taxonomicName>
made up the first component of the diverse minute polydesmid fauna of the Pacific Northwest to be recently described. The general appearance of the two previously described species is typical of the tiny polydesmids, with large, setose heads, relatively small collums with scattered setae, and metazonites with toothed margins and with rows of setae on prominent sockets. Alveolate cuticle is confined to the head, epiproct and sometimes the anterior margin of the collum. The epiproct is short and not swollen. The gonopods are somewhat variable from species to species; the two described by
<bibRefCitation id="EF894B70FFE59D0548720072DAAB252E" author="Shelley, R. M." box="[832,1000,1352,1379]" pageId="12" pageNumber="93" pagination="1 - 12" refId="ref20733" refString="Shelley, R. M. (2003) A new polydesmid genus and two new species from Oregon and Washington, U. S. A., with a review of Bidentogon Buckett &amp; Gardner, 1968, and a summary of the family in western North America (Polydesmida: Polydesmidae). Zootaxa, 296, 1 - 12. https: // doi. org / 10.11646 / zootaxa. 296.1.1" type="journal article" year="2003">Shelley (2003)</bibRefCitation>
,
<taxonomicName id="4C184D02FFE59D0548C70073DD30252F" box="[1013,1139,1352,1378]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="93" phylum="Arthropoda" rank="species" species="leonardi">
<emphasis id="B96CEA93FFE59D0548C70073DD30252F" box="[1013,1139,1352,1378]" italics="true" pageId="12" pageNumber="93">R. leonardi</emphasis>
</taxonomicName>
and
<emphasis id="B96CEA93FFE59D054F990073DC08252F" box="[1195,1355,1352,1378]" italics="true" pageId="12" pageNumber="93">
<taxonomicName id="4C184D02FFE59D054F990073DC04252F" box="[1195,1351,1352,1378]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="93" phylum="Arthropoda" rank="species" species="benedictae">R. benedictae</taxonomicName>
,
</emphasis>
have a prominent subterminal branch in the terminal zone, the lateral one of which Shelley termed the endomerite and the mesal one the tibiotarsus. In the terminology used here, the gonopod consists of a simple acropodite (
<emphasis id="B96CEA93FFE59D054E2D00ABDC0225E6" bold="true" box="[1311,1345,1425,1451]" pageId="12" pageNumber="93">ac,</emphasis>
<figureCitation id="13232A04FFE59D054E7B00AADCD725E6" box="[1353,1428,1424,1451]" captionStart="FIGURES 27, 28" captionStartId="14.[151,264,1981,2005]" captionTargetBox="[153,1436,1066,1957]" captionTargetId="figure-465@14.[151,1436,1064,1957]" captionTargetPageId="14" captionText="FIGURES 27, 28. Retrorsia leonardi Shelley gonopods. 27. Gonopods in situ, posterior view. 28. Gonopod tips. SEM stub WS32-15. Abbreviations: ac, acropodite; rp, retrorse process; sm, solenomere; tz, terminal zone." figureDoi="http://doi.org/10.5281/zenodo.4804941" httpUri="https://zenodo.org/record/4804941/files/figure.png" pageId="12" pageNumber="93">Fig 27</figureCitation>
) bearing the pulvillus, and a terminal zone with two processes, one of which recurves anteriorly (
<emphasis id="B96CEA93FFE59D054FE2008FDDB02582" bold="true" box="[1232,1267,1461,1487]" pageId="12" pageNumber="93">rp,</emphasis>
<figureCitation id="13232A04FFE59D054FC8008EDC092582" box="[1274,1354,1460,1487]" captionStart="FIGURES 27, 28" captionStartId="14.[151,264,1981,2005]" captionTargetBox="[153,1436,1066,1957]" captionTargetId="figure-465@14.[151,1436,1064,1957]" captionTargetPageId="14" captionText="FIGURES 27, 28. Retrorsia leonardi Shelley gonopods. 27. Gonopods in situ, posterior view. 28. Gonopod tips. SEM stub WS32-15. Abbreviations: ac, acropodite; rp, retrorse process; sm, solenomere; tz, terminal zone." figureDoi="http://doi.org/10.5281/zenodo.4804941" httpUri="https://zenodo.org/record/4804941/files/figure.png" pageId="12" pageNumber="93">Fig. 28</figureCitation>
) and is the source of the generic name. A further uniting character is a rounded notch just distal to the pulvillus, although this is not obvious in
<emphasis id="B96CEA93FFE59D054AA200C7DB55265B" box="[400,534,1532,1558]" italics="true" pageId="12" pageNumber="93">
<taxonomicName id="4C184D02FFE59D054AA200C7DB51265B" box="[400,530,1532,1558]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="93" phylum="Arthropoda" rank="species" species="leonardi">R. leonardi</taxonomicName>
,
</emphasis>
the
<typeStatus id="54A38823FFE59D05497800C7DB39265A" box="[586,634,1533,1559]" pageId="12" pageNumber="93">type</typeStatus>
species of the genus. Shelley’s illustrations of the gonopods show the course of the seminal canal as lacking a loop and a seminal vesicle. Close examination reveals that a loop is indeed lacking, but that the seminal canal at the very least expands just before the pore (
<figureCitation id="13232A04FFE59D054F29037EDDFB2612" box="[1051,1208,1604,1631]" captionStart="FIGURES 121–126" captionStartId="40.[151,264,1408,1432]" captionTargetBox="[160,1431,185,1372]" captionTargetId="figure-24@40.[151,1436,181,1384]" captionTargetPageId="40" captionText="FIGURES 121–126. Drawings of gonopods of polydesmid species. 121. Left gonopod of Retrorisia leonardi, mesal view. 122. Left gonopod of Retrorsia benedictae, mesal view. 123. Left gonopod of Retrorsia richarti,n. sp., mesal view. 124. Right gonopod of Retrorsia gracilis, n. sp., lateral view. 125, Gonopod tip of Retrorsia gracilis, mesal view. 126. Right gonopod of Retrorsia simplicissima, n. sp., lateral view." figureDoi="http://doi.org/10.5281/zenodo.4805071" httpUri="https://zenodo.org/record/4805071/files/figure.png" pageId="12" pageNumber="93">Figs 121, 122</figureCitation>
), so it can be argued that a vesicle is indeed present and perhaps has been reduced, or is in the process of developing. The three new species described below have simpler gonopods than the two described by
<bibRefCitation id="EF894B70FFE59D0548F103B6DD2A26EA" author="Shelley, R. M." box="[963,1129,1676,1703]" pageId="12" pageNumber="93" pagination="1 - 12" refId="ref20733" refString="Shelley, R. M. (2003) A new polydesmid genus and two new species from Oregon and Washington, U. S. A., with a review of Bidentogon Buckett &amp; Gardner, 1968, and a summary of the family in western North America (Polydesmida: Polydesmidae). Zootaxa, 296, 1 - 12. https: // doi. org / 10.11646 / zootaxa. 296.1.1" type="journal article" year="2003">Shelley (2003)</bibRefCitation>
and the retrorse process of the terminal zone is less distinct.
</paragraph>
<paragraph id="8BA73681FFE59D054BF503EEDC272717" blockId="12.[151,1437,992,1955]" pageId="12" pageNumber="93">
The detailed descriptions of the nonsexual characters of the two species named by
<bibRefCitation id="EF894B70FFE59D054F5A03EEDC4E26A2" author="Shelley, R. M." box="[1128,1293,1748,1775]" pageId="12" pageNumber="93" pagination="1 - 12" refId="ref20733" refString="Shelley, R. M. (2003) A new polydesmid genus and two new species from Oregon and Washington, U. S. A., with a review of Bidentogon Buckett &amp; Gardner, 1968, and a summary of the family in western North America (Polydesmida: Polydesmidae). Zootaxa, 296, 1 - 12. https: // doi. org / 10.11646 / zootaxa. 296.1.1" type="journal article" year="2003">Shelley (2003)</bibRefCitation>
can serve as well for the three additional species described here. The anterior setal row on the collum consists of 16–20 setae, varying according to species. The anterior metazonites have four rows of 14–18 setae set on stout, discrete tubercles; the number of rows increases to five midbody and may or may not decrease again to four at the posterior end.
</paragraph>
<paragraph id="8BA73681FFE59D054BF5025EDAAC27EE" blockId="12.[151,1437,992,1955]" pageId="12" pageNumber="93">
<bibRefCitation id="EF894B70FFE59D054BF5025ED8332732" author="Shelley, R. M." box="[199,368,1892,1919]" pageId="12" pageNumber="93" pagination="1 - 12" refId="ref20733" refString="Shelley, R. M. (2003) A new polydesmid genus and two new species from Oregon and Washington, U. S. A., with a review of Bidentogon Buckett &amp; Gardner, 1968, and a summary of the family in western North America (Polydesmida: Polydesmidae). Zootaxa, 296, 1 - 12. https: // doi. org / 10.11646 / zootaxa. 296.1.1" type="journal article" year="2003">Shelley (2003)</bibRefCitation>
did not mention
<taxonomicName id="4C184D02FFE59D054908025EDBF52733" authorityName="Chamberlin &amp; Hoffman" authorityYear="1950" box="[570,694,1892,1918]" class="Diplopoda" family="Polydesmidae" genus="Utadesmus" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="B96CEA93FFE59D054908025EDBF52733" box="[570,694,1892,1918]" italics="true" pageId="12" pageNumber="93">Utadesmus</emphasis>
</taxonomicName>
as possibly related to
<emphasis id="B96CEA93FFE59D05488D025FDD6D2733" box="[959,1070,1893,1918]" italics="true" pageId="12" pageNumber="93">
<taxonomicName id="4C184D02FFE59D05488D025FDD682733" authorityName="Shelley" authorityYear="2003" box="[959,1067,1893,1918]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="93" phylum="Arthropoda" rank="genus">Retrorsia</taxonomicName>
,
</emphasis>
but there are similarities in the gonopods, especially in the division of the terminal zone into two processes.
</paragraph>
</subSubSection>
</treatment>
</document>
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