244 lines
32 KiB
XML
244 lines
32 KiB
XML
<document id="27CF3FD179142D19AFA12632E2D7E7D0" ID-CLB-Dataset="6194" ID-DOI="10.11646/zootaxa.4975.1.3" ID-GBIF-Dataset="529d48db-4299-4e2d-9aaf-6c4178c08b18" ID-ISSN="1175-5326" ID-Zenodo-Dep="4804909" ID-ZooBank="DB7C9028-3EDF-454F-88D0-336624AD1DC4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1622016306479" checkinUser="plazi" docAuthor="Shear, William A. & Marek, Paul E." docDate="2021" docId="03B18797FFE49D074BA505ADDDCD245D" docLanguage="en" docName="zootaxa.4975.1.3.pdf" docOrigin="Zootaxa 4975 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Retrorsia leonardi Shelley 2003" docType="treatment" docVersion="8" lastPageNumber="95" masterDocId="FF88FFEFFFE99D094B32053AD943204D" masterDocTitle="Three new genera and eighteen new species of miniature polydesmid millipedes from the northwestern United States (Diplopoda, Polydesmida, Polydesmidae)" masterLastPageNumber="126" masterPageNumber="81" pageNumber="94" updateTime="1698991024886" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="2323A79B686B30C6278304A35C207706">Three new genera and eighteen new species of miniature polydesmid millipedes from the northwestern United States (Diplopoda, Polydesmida, Polydesmidae)</mods:title>
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<mods:namePart id="E04222C72D17834414536048693E09B5">Shear, William A.</mods:namePart>
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<mods:affiliation id="95F681DCAAB201B86552F831A6276246">Professor Emeritus, Department of Biology, Hampden-Sydney College, Hampden-Sydney VA 23943; present address 1950 Price Drive, Farmville VA 23901.</mods:affiliation>
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<mods:namePart id="FF628622AB1D91F67F7E146F1E4E4428">Marek, Paul E.</mods:namePart>
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<mods:nameIdentifier id="00FC159F51D756130A8B36E83674A515" type="ORCID">0000-0002-7048-2514</mods:nameIdentifier>
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<mods:affiliation id="521C7F351D83F631AA5CDDDDAC59B222">Associate Professor, Department of Entomology, Virginia Polytechnic Institute and State University, Blacksburg, VA 24061. pmarek @ vt. edu; https: // orcid. org / 0000 - 0002 - 7048 - 2514</mods:affiliation>
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<mods:nameIdentifier id="F2859251C205C9D9A6AD9264260E79E4" type="email">pmarek@vt.edu</mods:nameIdentifier>
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<mods:title id="A10C59EDC65DAF20829A88D6A14397A8">Zootaxa</mods:title>
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<mods:date id="995C2700972E0C0A70146B38AF085636">2021</mods:date>
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<treatment id="03B18797FFE49D074BA505ADDDCD245D" ID-DOI="http://doi.org/10.5281/zenodo.4804995" ID-GBIF-Taxon="182644911" ID-Zenodo-Dep="4804995" LSID="urn:lsid:plazi:treatment:03B18797FFE49D074BA505ADDDCD245D" httpUri="http://treatment.plazi.org/id/03B18797FFE49D074BA505ADDDCD245D" lastPageId="14" lastPageNumber="95" pageId="13" pageNumber="94">
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<subSubSection id="C302650AFFE49D044BA505ADDB5E20FC" box="[151,541,151,178]" pageId="13" pageNumber="94" type="nomenclature">
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<paragraph id="8BA73681FFE49D044BA505ADDB5E20FC" blockId="13.[151,541,151,213]" box="[151,541,151,178]" pageId="13" pageNumber="94">
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<heading id="D0EF81EDFFE49D044BA505ADDB5E20FC" bold="true" box="[151,541,151,178]" fontSize="11" level="1" pageId="13" pageNumber="94" reason="1">
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<taxonomicName id="4C184D02FFE49D044BA505ADDB5E20FC" ID-CoL="78RVF" authority="Shelley, 2003" authorityName="Shelley" authorityYear="2003" box="[151,541,151,178]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="13" pageNumber="94" phylum="Arthropoda" rank="species" species="leonardi">
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<emphasis id="B96CEA93FFE49D044BA505ADDB5E20FC" bold="true" box="[151,541,151,178]" pageId="13" pageNumber="94">
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<emphasis id="B96CEA93FFE49D044BA505ADD83720FF" bold="true" box="[151,372,151,178]" italics="true" pageId="13" pageNumber="94">Retrorsia leonardi</emphasis>
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<bibRefCitation id="EF894B70FFE49D044A4905ADDB5E20FC" author="Shelley, R. M." box="[379,541,151,177]" pageId="13" pageNumber="94" pagination="1 - 12" refId="ref20733" refString="Shelley, R. M. (2003) A new polydesmid genus and two new species from Oregon and Washington, U. S. A., with a review of Bidentogon Buckett & Gardner, 1968, and a summary of the family in western North America (Polydesmida: Polydesmidae). Zootaxa, 296, 1 - 12. https: // doi. org / 10.11646 / zootaxa. 296.1.1" type="journal article" year="2003">Shelley, 2003</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</paragraph>
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<subSubSection id="C302650AFFE49D044BA50581D8C72098" box="[151,388,187,213]" pageId="13" pageNumber="94" type="description">
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<paragraph id="8BA73681FFE49D044BA50581D8C72098" blockId="13.[151,541,151,213]" box="[151,388,187,213]" pageId="13" pageNumber="94">
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<figureCitation id="13232A04FFE49D044BA50581D8542098" box="[151,279,187,213]" captionStart-0="FIGURES 21–26" captionStart-1="FIGURES 27, 28" captionStartId-0="13.[151,264,1925,1949]" captionStartId-1="14.[151,264,1981,2005]" captionTargetBox-0="[151,1434,414,1899]" captionTargetBox-1="[153,1436,1066,1957]" captionTargetId-0="figure-50@13.[151,1436,412,1902]" captionTargetId-1="figure-465@14.[151,1436,1064,1957]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="FIGURES 21–26. Retrorsia leonardi Shelley. 21. Head and anterior rings, dorsal view. 22. Pygidium, lateral view. 23. Midbody ring, dorsal view. 24. Retracted female genitalia, ventral view. 25. Tarsus of left leg 7, posterior view. 26. Sphaerotrichomes on tarsus of leg 7. SEM stub WS32-15" captionText-1="FIGURES 27, 28. Retrorsia leonardi Shelley gonopods. 27. Gonopods in situ, posterior view. 28. Gonopod tips. SEM stub WS32-15. Abbreviations: ac, acropodite; rp, retrorse process; sm, solenomere; tz, terminal zone." figureDoi-0="http://doi.org/10.5281/zenodo.4804935" figureDoi-1="http://doi.org/10.5281/zenodo.4804941" httpUri-0="https://zenodo.org/record/4804935/files/figure.png" httpUri-1="https://zenodo.org/record/4804941/files/figure.png" pageId="13" pageNumber="94">Figs 21–28</figureCitation>
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,
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<figureCitation id="13232A04FFE49D044A100581D80D2098" box="[290,334,187,213]" captionStart="FIGURES 121–126" captionStartId="40.[151,264,1408,1432]" captionTargetBox="[160,1431,185,1372]" captionTargetId="figure-24@40.[151,1436,181,1384]" captionTargetPageId="40" captionText="FIGURES 121–126. Drawings of gonopods of polydesmid species. 121. Left gonopod of Retrorisia leonardi, mesal view. 122. Left gonopod of Retrorsia benedictae, mesal view. 123. Left gonopod of Retrorsia richarti,n. sp., mesal view. 124. Right gonopod of Retrorsia gracilis, n. sp., lateral view. 125, Gonopod tip of Retrorsia gracilis, mesal view. 126. Right gonopod of Retrorsia simplicissima, n. sp., lateral view." figureDoi="http://doi.org/10.5281/zenodo.4805071" httpUri="https://zenodo.org/record/4805071/files/figure.png" pageId="13" pageNumber="94">121</figureCitation>
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,
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<figureCitation id="13232A04FFE49D044A680581D8C72098" box="[346,388,187,213]" captionStart="FIGURE 138" captionStartId="43.[151,250,1002,1026]" captionTargetBox="[179,1408,181,979]" captionTargetId="figure-17@43.[179,1408,181,981]" captionTargetPageId="43" captionText="FIGURE 138. Retrorsia leonardi Shelley, living specimen in habitat. Photograph by William Leonard." figureDoi="http://doi.org/10.5281/zenodo.4805087" httpUri="https://zenodo.org/record/4805087/files/figure.png" pageId="13" pageNumber="94">138</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C302650AFFE49D044BA5043AD8BF215A" box="[151,508,255,280]" pageId="13" pageNumber="94" type="reference_group">
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<paragraph id="8BA73681FFE49D044BA5043AD8BF215A" blockId="13.[151,508,255,280]" box="[151,508,255,280]" pageId="13" pageNumber="94">
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<treatmentCitationGroup id="AB0811AFFFE49D044BA5043AD8BF215A" box="[151,508,255,280]" pageId="13" pageNumber="94">
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<treatmentCitation id="0AB91090FFE49D044BA5043AD8BB215A" ID-DOI="http://doi.org/10.5281/zenodo.5095716" ID-Zenodo-Dep="5095716" author="Shelley, R. M." box="[151,504,255,280]" httpUri="http://treatment.plazi.org/id/03A3096E3A11FFFAF462FC164D33C18B" page="5" pageId="13" pageNumber="94" year="2003">
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<taxonomicName id="4C184D02FFE49D044BA5043AD8BB215A" ID-CoL="78RVF" authority="Shelley, 2003: 5" authorityName="Shelley" authorityPageNumber="5" authorityYear="2003" box="[151,504,255,280]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="13" pageNumber="94" phylum="Arthropoda" rank="species" species="leonardi">
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<emphasis id="B96CEA93FFE49D044BA5043AD8122155" box="[151,337,256,280]" italics="true" pageId="13" pageNumber="94">Retrorsia leonardi</emphasis>
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<bibRefCitation id="EF894B70FFE49D044A6A043AD8BB215A" author="Shelley, R. M." box="[344,504,255,280]" pageId="13" pageNumber="94" pagination="1 - 12" refId="ref20733" refString="Shelley, R. M. (2003) A new polydesmid genus and two new species from Oregon and Washington, U. S. A., with a review of Bidentogon Buckett & Gardner, 1968, and a summary of the family in western North America (Polydesmida: Polydesmidae). Zootaxa, 296, 1 - 12. https: // doi. org / 10.11646 / zootaxa. 296.1.1" type="journal article" year="2003">Shelley, 2003:5</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</paragraph>
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<subSubSection id="C302650AFFE49D044BA5047CD89821C9" pageId="13" pageNumber="94" type="materials_examined">
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<paragraph id="8BA73681FFE49D044BA5047CD89821C9" blockId="13.[151,1436,325,388]" pageId="13" pageNumber="94">
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<materialsCitation id="3B703CDCFFE49D044BA5047CD89421C9" ID-GBIF-Occurrence="3128519314" county="Multnomah County" location="National Scenic Area" municipality="Ainsworth State Park" pageId="13" pageNumber="94" specimenCount="1" stateProvince="Oregon" typeStatus="holotype">
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<emphasis id="B96CEA93FFE49D044BA5047CD8752112" bold="true" box="[151,310,325,352]" pageId="13" pageNumber="94">
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<typeStatus id="54A38823FFE49D044BA5047CD991212D" box="[151,210,326,352]" pageId="13" pageNumber="94">Type</typeStatus>
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locality.
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</emphasis>
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<collectingMunicipality id="6BC3ACFBFFE49D044A0E047CDB6C212D" box="[316,559,326,352]" pageId="13" pageNumber="94">Ainsworth State Park</collectingMunicipality>
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,
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<taxonomicName id="4C184D02FFE49D044908047CDA79212D" authority="River Gorge" authorityName="River Gorge" box="[570,826,326,352]" family="Percopsidae" genus="Columbia" higherTaxonomySource="CoL" kingdom="Animalia" order="Percopsiformes" pageId="13" pageNumber="94" phylum="Chordata" rank="genus">Columbia River Gorge</taxonomicName>
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<location id="8EC7605AFFE49D044873047CDD71212D" LSID="urn:lsid:plazi:treatment:03B18797FFE49D074BA505ADDDCD245D:8EC7605AFFE49D044873047CDD71212D" box="[833,1074,326,352]" county="Multnomah County" municipality="Ainsworth State Park" name="National Scenic Area" pageId="13" pageNumber="94" stateProvince="Oregon">National Scenic Area</location>
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,
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<quantity id="4CE09B64FFE49D044F0F047CDD3D212D" box="[1085,1150,326,352]" metricMagnitude="4" metricUnit="m" metricValue="4.828032" pageId="13" pageNumber="94" unit="mi" value="30.0">30 mi</quantity>
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(
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<quantity id="4CE09B64FFE49D044FBF047CDD94212C" box="[1165,1239,326,353]" metricMagnitude="4" metricUnit="m" metricValue="4.8" pageId="13" pageNumber="94" unit="km" value="48.0">48 km</quantity>
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) east of
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<location id="8EC7605AFFE49D044E04047CDCDB212D" LSID="urn:lsid:plazi:treatment:03B18797FFE49D074BA505ADDDCD245D:8EC7605AFFE49D044E04047CDCDB212D" box="[1334,1432,326,352]" county="Multnomah County" municipality="Ainsworth State Park" name="Portland" pageId="13" pageNumber="94" stateProvince="Oregon">Portland</location>
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,
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<collectingCounty id="62C64E0DFFE49D044BA50450D83621C9" box="[151,373,362,388]" pageId="13" pageNumber="94">Multnomah County</collectingCounty>
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,
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<collectingRegion id="49DCF863FFE49D044A4D0450D89421C9" box="[383,471,362,388]" country="United States of America" name="Oregon" pageId="13" pageNumber="94">Oregon</collectingRegion>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<caption id="DF676609FFE49D044BA502BFD8BD27A8" ID-DOI="http://doi.org/10.5281/zenodo.4804935" ID-Zenodo-Dep="4804935" httpUri="https://zenodo.org/record/4804935/files/figure.png" pageId="13" pageNumber="94" startId="13.[151,264,1925,1949]" targetBox="[151,1434,414,1899]" targetPageId="13">
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<paragraph id="8BA73681FFE49D044BA502BFD8BD27A8" blockId="13.[151,1436,1925,2022]" pageId="13" pageNumber="94">
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<emphasis id="B96CEA93FFE49D044BA502BFD80F27D0" bold="true" box="[151,332,1925,1949]" pageId="13" pageNumber="94">FIGURES 21–26</emphasis>
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.
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||
<taxonomicName id="4C184D02FFE49D044A6A02BCDB2B27D0" authority="Shelley." authorityName="Shelley." box="[344,616,1925,1949]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="13" pageNumber="94" phylum="Arthropoda" rank="species" species="leonardi">
|
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<emphasis id="B96CEA93FFE49D044A6A02BCDB5227D0" box="[344,529,1925,1949]" italics="true" pageId="13" pageNumber="94">Retrorsia leonardi</emphasis>
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Shelley.
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</taxonomicName>
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21. Head and anterior rings, dorsal view.
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||
<emphasis id="B96CEA93FFE49D044F3502BFDD6327D0" bold="true" box="[1031,1056,1925,1949]" pageId="13" pageNumber="94">22</emphasis>
|
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. Pygidium, lateral view.
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<emphasis id="B96CEA93FFE49D044E2802BFDC7027D0" bold="true" box="[1306,1331,1925,1949]" pageId="13" pageNumber="94">23</emphasis>
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. Midbody ring, dorsal view.
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<emphasis id="B96CEA93FFE49D044A7F0293D825278C" bold="true" box="[333,358,1961,1985]" pageId="13" pageNumber="94">24</emphasis>
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||
. Retracted female genitalia, ventral view.
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||
<emphasis id="B96CEA93FFE49D0448210293DA6F278C" bold="true" box="[787,812,1961,1985]" pageId="13" pageNumber="94">25</emphasis>
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||
. Tarsus of left leg 7, posterior view.
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<emphasis id="B96CEA93FFE49D044F920293DD83278C" bold="true" box="[1184,1216,1961,1985]" pageId="13" pageNumber="94">26.</emphasis>
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Sphaerotrichomes on tarsus of leg 7. SEM stub WS32-15
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</paragraph>
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||
</caption>
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||
<subSubSection id="C302650AFFE79D074BF505ACD8E62254" pageId="14" pageNumber="95" type="diagnosis">
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<paragraph id="8BA73681FFE79D074BF505ACDC5B20B5" blockId="14.[151,1437,150,1041]" pageId="14" pageNumber="95">
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||
<emphasis id="B96CEA93FFE79D074BF505ACD80120FD" bold="true" box="[199,322,150,176]" pageId="14" pageNumber="95">Diagnosis.</emphasis>
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The retrorse process (
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<emphasis id="B96CEA93FFE79D07490A05ACDB1820FD" bold="true" box="[568,603,150,176]" pageId="14" pageNumber="95">rp,</emphasis>
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||
<figureCitation id="13232A04FFE79D07495305ACDBF220FD" box="[609,689,150,176]" captionStart="FIGURES 27, 28" captionStartId="14.[151,264,1981,2005]" captionTargetBox="[153,1436,1066,1957]" captionTargetId="figure-465@14.[151,1436,1064,1957]" captionTargetPageId="14" captionText="FIGURES 27, 28. Retrorsia leonardi Shelley gonopods. 27. Gonopods in situ, posterior view. 28. Gonopod tips. SEM stub WS32-15. Abbreviations: ac, acropodite; rp, retrorse process; sm, solenomere; tz, terminal zone." figureDoi="http://doi.org/10.5281/zenodo.4804941" httpUri="https://zenodo.org/record/4804941/files/figure.png" pageId="14" pageNumber="95">Fig. 28</figureCitation>
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) terminating the terminal zone is strongly curved and the subterminal process (
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<emphasis id="B96CEA93FFE79D074A160580D8032099" bold="true" box="[292,320,186,212]" pageId="14" pageNumber="95">st,</emphasis>
|
||
<figureCitation id="13232A04FFE79D074A740580D8D52099" box="[326,406,186,212]" captionStart="FIGURES 27, 28" captionStartId="14.[151,264,1981,2005]" captionTargetBox="[153,1436,1066,1957]" captionTargetId="figure-465@14.[151,1436,1064,1957]" captionTargetPageId="14" captionText="FIGURES 27, 28. Retrorsia leonardi Shelley gonopods. 27. Gonopods in situ, posterior view. 28. Gonopod tips. SEM stub WS32-15. Abbreviations: ac, acropodite; rp, retrorse process; sm, solenomere; tz, terminal zone." figureDoi="http://doi.org/10.5281/zenodo.4804941" httpUri="https://zenodo.org/record/4804941/files/figure.png" pageId="14" pageNumber="95">Fig. 28</figureCitation>
|
||
) is broadly spatulate. The notch above the pulvillus seems almost entirely suppressed in this species. The anterior marginal row of setae on the collum may consist of as many as 20 setae (
|
||
<figureCitation id="13232A04FFE79D074F8505E4DC4820B5" box="[1207,1291,222,248]" captionStart="FIGURES 21–26" captionStartId="13.[151,264,1925,1949]" captionTargetBox="[151,1434,414,1899]" captionTargetId="figure-50@13.[151,1436,412,1902]" captionTargetPageId="13" captionText="FIGURES 21–26. Retrorsia leonardi Shelley. 21. Head and anterior rings, dorsal view. 22. Pygidium, lateral view. 23. Midbody ring, dorsal view. 24. Retracted female genitalia, ventral view. 25. Tarsus of left leg 7, posterior view. 26. Sphaerotrichomes on tarsus of leg 7. SEM stub WS32-15" figureDoi="http://doi.org/10.5281/zenodo.4804935" httpUri="https://zenodo.org/record/4804935/files/figure.png" pageId="14" pageNumber="95">Fig. 21</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA73681FFE79D074BF50438DB3621E1" blockId="14.[151,1437,150,1041]" pageId="14" pageNumber="95">
|
||
<emphasis id="B96CEA93FFE79D074BF50438D8522151" bold="true" box="[199,273,258,284]" pageId="14" pageNumber="95">Notes.</emphasis>
|
||
Males have sphaerotrichomes on the tarsi (
|
||
<figureCitation id="13232A04FFE79D0748360438DA142150" box="[772,855,258,285]" captionStart="FIGURES 21–26" captionStartId="13.[151,264,1925,1949]" captionTargetBox="[151,1434,414,1899]" captionTargetId="figure-50@13.[151,1436,412,1902]" captionTargetPageId="13" captionText="FIGURES 21–26. Retrorsia leonardi Shelley. 21. Head and anterior rings, dorsal view. 22. Pygidium, lateral view. 23. Midbody ring, dorsal view. 24. Retracted female genitalia, ventral view. 25. Tarsus of left leg 7, posterior view. 26. Sphaerotrichomes on tarsus of leg 7. SEM stub WS32-15" figureDoi="http://doi.org/10.5281/zenodo.4804935" httpUri="https://zenodo.org/record/4804935/files/figure.png" pageId="14" pageNumber="95">Fig. 25</figureCitation>
|
||
) of the anterior legs, the bases of which are proximodistally flattened and have fingerprint-like wrinkles (
|
||
<figureCitation id="13232A04FFE79D07483B041CDA1E210C" box="[777,861,294,321]" captionStart="FIGURES 21–26" captionStartId="13.[151,264,1925,1949]" captionTargetBox="[151,1434,414,1899]" captionTargetId="figure-50@13.[151,1436,412,1902]" captionTargetPageId="13" captionText="FIGURES 21–26. Retrorsia leonardi Shelley. 21. Head and anterior rings, dorsal view. 22. Pygidium, lateral view. 23. Midbody ring, dorsal view. 24. Retracted female genitalia, ventral view. 25. Tarsus of left leg 7, posterior view. 26. Sphaerotrichomes on tarsus of leg 7. SEM stub WS32-15" figureDoi="http://doi.org/10.5281/zenodo.4804935" httpUri="https://zenodo.org/record/4804935/files/figure.png" pageId="14" pageNumber="95">Fig. 26</figureCitation>
|
||
). The cuticular fimbriae around the seminal pore of the pulvillus extend distally along the posterior surface of the gonopod in a diminishing line (
|
||
<figureCitation id="13232A04FFE79D074FA50470DDAE2129" box="[1175,1261,330,356]" captionStart="FIGURES 27, 28" captionStartId="14.[151,264,1981,2005]" captionTargetBox="[153,1436,1066,1957]" captionTargetId="figure-465@14.[151,1436,1064,1957]" captionTargetPageId="14" captionText="FIGURES 27, 28. Retrorsia leonardi Shelley gonopods. 27. Gonopods in situ, posterior view. 28. Gonopod tips. SEM stub WS32-15. Abbreviations: ac, acropodite; rp, retrorse process; sm, solenomere; tz, terminal zone." figureDoi="http://doi.org/10.5281/zenodo.4804941" httpUri="https://zenodo.org/record/4804941/files/figure.png" pageId="14" pageNumber="95">Figs 27</figureCitation>
|
||
,
|
||
<figureCitation id="13232A04FFE79D074FC50470DC662129" box="[1271,1317,330,356]" captionStart="FIGURES 117–120" captionStartId="39.[151,264,1199,1223]" captionTargetBox="[151,1436,181,1176]" captionTargetId="figure-17@39.[151,1436,181,1176]" captionTargetPageId="39" captionText="FIGURES 117–120. Species of Benedictesmus, n. gen. 117. Gonopod tips of Benedictesmus yaquina, n. sp., posterior view. SEM stub WS33-14. Figs 118–120. Benedictesmus timber, n. sp. 118. Head and anterior rings, dorsal view. 119. Midbody ring, dorsal view. 120. Pygidium, lateral view. SEM stub WS33-12/ Abbreviations: p, pulvillus; pp, pulvillar process; tz, terminal zone." figureDoi="http://doi.org/10.5281/zenodo.4805069" httpUri="https://zenodo.org/record/4805069/files/figure.png" pageId="14" pageNumber="95">120</figureCitation>
|
||
). Posterior metazonites may have up to six rows of setae, but the rows tend to become less distinct on rings 16–18. The epiproct is short, not decurved or swollen (
|
||
<figureCitation id="13232A04FFE79D07492604A8DB2B21E1" box="[532,616,402,428]" captionStart="FIGURES 21–26" captionStartId="13.[151,264,1925,1949]" captionTargetBox="[151,1434,414,1899]" captionTargetId="figure-50@13.[151,1436,412,1902]" captionTargetPageId="13" captionText="FIGURES 21–26. Retrorsia leonardi Shelley. 21. Head and anterior rings, dorsal view. 22. Pygidium, lateral view. 23. Midbody ring, dorsal view. 24. Retracted female genitalia, ventral view. 25. Tarsus of left leg 7, posterior view. 26. Sphaerotrichomes on tarsus of leg 7. SEM stub WS32-15" figureDoi="http://doi.org/10.5281/zenodo.4804935" httpUri="https://zenodo.org/record/4804935/files/figure.png" pageId="14" pageNumber="95">Fig. 22</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA73681FFE79D074BF5048CD8E62254" blockId="14.[151,1437,150,1041]" pageId="14" pageNumber="95">
|
||
The female genitalia may be unusual and deserve further study. In their retracted position (
|
||
<figureCitation id="13232A04FFE79D074FF2048CDC52219D" box="[1216,1297,438,464]" captionStart="FIGURES 21–26" captionStartId="13.[151,264,1925,1949]" captionTargetBox="[151,1434,414,1899]" captionTargetId="figure-50@13.[151,1436,412,1902]" captionTargetPageId="13" captionText="FIGURES 21–26. Retrorsia leonardi Shelley. 21. Head and anterior rings, dorsal view. 22. Pygidium, lateral view. 23. Midbody ring, dorsal view. 24. Retracted female genitalia, ventral view. 25. Tarsus of left leg 7, posterior view. 26. Sphaerotrichomes on tarsus of leg 7. SEM stub WS32-15" figureDoi="http://doi.org/10.5281/zenodo.4804935" httpUri="https://zenodo.org/record/4804935/files/figure.png" pageId="14" pageNumber="95">Fig. 24</figureCitation>
|
||
) paired projecting structures can be seen which have the same cuticular sculpture as the sternite. It is not clear if these are the cyphopod valves or not.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C302650AFFE79D074BF50718DDCD245D" pageId="14" pageNumber="95" type="distribution">
|
||
<paragraph id="8BA73681FFE79D074BF50718DCDA2271" blockId="14.[151,1437,150,1041]" box="[199,1433,546,573]" pageId="14" pageNumber="95">
|
||
<emphasis id="B96CEA93FFE79D074BF50718D8192271" bold="true" box="[199,346,546,572]" pageId="14" pageNumber="95">Distribution.</emphasis>
|
||
The new records for this species expand the range into
|
||
<collectingRegion id="49DCF863FFE79D0748980718DD6A2271" box="[938,1065,546,572]" country="United States of America" name="Washington" pageId="14" pageNumber="95">Washington</collectingRegion>
|
||
well north of the
|
||
<taxonomicName id="4C184D02FFE79D074FD40718DCDA2271" authority="River." authorityName="River." box="[1254,1433,546,572]" family="Percopsidae" genus="Columbia" higherTaxonomySource="CoL" kingdom="Animalia" order="Percopsiformes" pageId="14" pageNumber="95" phylum="Chordata" rank="genus">Columbia River.</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="8BA73681FFE79D074BF5077CDDCD245D" blockId="14.[151,1437,150,1041]" pageId="14" pageNumber="95">
|
||
<emphasis id="B96CEA93FFE79D074BF5077CD827222D" bold="true" box="[199,356,582,608]" pageId="14" pageNumber="95">New records.</emphasis>
|
||
<collectingRegion id="49DCF863FFE79D074A59077CD8A7222D" box="[363,484,582,608]" country="United States of America" name="Oregon" pageId="14" pageNumber="95">OREGON</collectingRegion>
|
||
:
|
||
<emphasis id="B96CEA93FFE79D074ADC077CDBD7222C" box="[494,660,582,609]" italics="true" pageId="14" pageNumber="95">
|
||
<taxonomicName id="4C184D02FFE79D074ADC077CDBCE222C" authority="Co." authorityName="Co." box="[494,653,582,609]" family="Percopsidae" genus="Columbia" higherTaxonomySource="CoL" kingdom="Animalia" order="Percopsiformes" pageId="14" pageNumber="95" phylum="Chordata" rank="genus">Columbia Co.</taxonomicName>
|
||
:
|
||
</emphasis>
|
||
Keystone Creek, Clatskanie Valley,
|
||
<quantity id="4CE09B64FFE79D074F1F077CDD2C222D" box="[1069,1135,582,608]" metricMagnitude="1" metricUnit="m" metricValue="4.02336" pageId="14" pageNumber="95" unit="ft" value="132.0">132 ft</quantity>
|
||
asl, 40.0822°, -123.1569°,
|
||
<date id="FFA61041FFE79D074BA50750D81B22C9" box="[151,344,618,644]" pageId="14" pageNumber="95" value="2002-02-08">8 February 2002</date>
|
||
, W. Leonard, C. Richart, mm ff.
|
||
<emphasis id="B96CEA93FFE79D0749EB0750DA2422C8" box="[729,871,618,645]" italics="true" pageId="14" pageNumber="95">Yamhill Co.:</emphasis>
|
||
near Amity, 45.4594°, -122.4600°,
|
||
<date id="FFA61041FFE79D074E360750D99122E4" pageId="14" pageNumber="95" value="1915-12-30">30 December 1915</date>
|
||
, P. Nosler, mm ff.
|
||
<collectingRegion id="49DCF863FFE79D074A9507B4DB2522E5" box="[423,614,654,680]" country="United States of America" name="Washington" pageId="14" pageNumber="95">WASHINGTON</collectingRegion>
|
||
:
|
||
<emphasis id="B96CEA93FFE79D07494307B4DA4322E4" box="[625,768,654,681]" italics="true" pageId="14" pageNumber="95">Cowlitz Co.:</emphasis>
|
||
<quantity id="4CE09B64FFE79D07483A07B4DA1D22E5" box="[776,862,654,680]" metricMagnitude="4" metricUnit="m" metricValue="1.83465216" pageId="14" pageNumber="95" unit="mi" value="11.4">11.4 mi</quantity>
|
||
east of I-
|
||
<quantity id="4CE09B64FFE79D0748FC07B4DD6F22E5" box="[974,1068,654,681]" metricMagnitude="3" metricUnit="m" metricValue="1.648968" pageId="14" pageNumber="95" unit="ft" value="5410.0">5, 410 ft</quantity>
|
||
asl,
|
||
<date id="FFA61041FFE79D074F5107B5DC4622E5" box="[1123,1285,654,681]" pageId="14" pageNumber="95" value="2004-03-07">7 March 2004</date>
|
||
, W. Leonard, mm ff; Pin Creek, Kool Road
|
||
<quantity id="4CE09B64FFE79D074AC00788DB7F2281" box="[498,572,690,716]" metricMagnitude="2" metricUnit="m" metricValue="6.437376" pageId="14" pageNumber="95" unit="mi" value="0.4">0.4 mi</quantity>
|
||
west of Fishpond Road,
|
||
<quantity id="4CE09B64FFE79D0748690788DADC2281" box="[859,927,690,716]" metricMagnitude="1" metricUnit="m" metricValue="5.4864" pageId="14" pageNumber="95" unit="ft" value="180.0">180 ft</quantity>
|
||
asl, 47.0742°, -122.8470°,
|
||
<date id="FFA61041FFE79D074FEB0788DCDB2281" box="[1241,1432,690,717]" pageId="14" pageNumber="95" value="2004-01-29">29 January 2004</date>
|
||
, W. Leonard, C. Richart, mm ff; SR503,
|
||
<quantity id="4CE09B64FFE79D07495207ECDBE922BD" box="[608,682,726,752]" metricMagnitude="3" metricUnit="m" metricValue="6.2764416" pageId="14" pageNumber="95" unit="mi" value="3.9">3.9 mi</quantity>
|
||
east of I-5, hillside north of Lewis River, 45.9412°, -122.6874°,
|
||
<date id="FFA61041FFE79D074EBC07EDD8672359" pageId="14" pageNumber="95" value="2004-03-07">7 March 2004</date>
|
||
, W. Leonard, mm ff; SR504,
|
||
<quantity id="4CE09B64FFE79D07494A07C0DB812359" box="[632,706,762,788]" metricMagnitude="3" metricUnit="m" metricValue="6.2764416" pageId="14" pageNumber="95" unit="mi" value="3.9">3.9 mi</quantity>
|
||
east of Toutle,
|
||
<quantity id="4CE09B64FFE79D07484407C0DAFA2359" box="[886,953,762,789]" metricMagnitude="2" metricUnit="m" metricValue="1.524" pageId="14" pageNumber="95" unit="ft" value="500.0">500 ft</quantity>
|
||
asl, 46.3483°, -122.7067°.
|
||
<date id="FFA61041FFE79D074FC707C0DCDB2358" box="[1269,1432,762,789]" pageId="14" pageNumber="95" value="2005-03-01">1 March 2005</date>
|
||
, W. Leonard, C. Richart.
|
||
<emphasis id="B96CEA93FFE79D074A9D0625DB6A2374" box="[431,553,799,825]" italics="true" pageId="14" pageNumber="95">Lewis Co.:</emphasis>
|
||
<quantity id="4CE09B64FFE79D0749000624DB3F2375" box="[562,636,798,825]" metricMagnitude="4" metricUnit="m" metricValue="1.3679424" pageId="14" pageNumber="95" unit="mi" value="8.5">8.5 mi</quantity>
|
||
south of Rendell on FS25, Gifford Pinchot National Forest, 46.4415°, -121.9982°,
|
||
<date id="FFA61041FFE79D074A100678D8BA2311" box="[290,505,834,860]" pageId="14" pageNumber="95" value="2003-12-21">21 December 2003</date>
|
||
, W. Leonard, m ff; same locality,
|
||
<date id="FFA61041FFE79D07485C0678DD742311" box="[878,1079,834,860]" pageId="14" pageNumber="95" value="2003-12-06">6 December 2003</date>
|
||
, W. Leonard, C. Richart, mm ff; Iron Creek Campground,
|
||
<quantity id="4CE09B64FFE79D074A85065CD8BC23CD" box="[439,511,870,896]" metricMagnitude="4" metricUnit="m" metricValue="1.54497024" pageId="14" pageNumber="95" unit="mi" value="9.6">9.6 mi</quantity>
|
||
south of Rendell on FS25, Gifford Pinchot National Forest, 45.4297°, -121.9869°,
|
||
<date id="FFA61041FFE79D074BA506B0D82D23E9" box="[151,366,906,932]" pageId="14" pageNumber="95" value="2003-12-21">21 December 2003</date>
|
||
, W. Leonard, m; Shafer Park,
|
||
<quantity id="4CE09B64FFE79D07498806B0DBBF23E9" box="[698,764,906,932]" metricMagnitude="1" metricUnit="m" metricValue="7.43712" pageId="14" pageNumber="95" unit="ft" value="244.0">244 ft</quantity>
|
||
asl, 46.7530°, -122.9380°,
|
||
<date id="FFA61041FFE79D074F1F06B0DDB623E9" box="[1069,1269,906,932]" pageId="14" pageNumber="95" value="2004-02-29">29 February 2004</date>
|
||
, C. Richart, m; SR508 at Bremer,
|
||
<quantity id="4CE09B64FFE79D074A540694D8E42385" box="[358,423,942,969]" metricMagnitude="2" metricUnit="m" metricValue="2.286" pageId="14" pageNumber="95" unit="ft" value="750.0">750 ft</quantity>
|
||
asl, 45.5917°, -122.4319°,
|
||
<date id="FFA61041FFE79D0749E80694DAE72385" box="[730,932,942,968]" pageId="14" pageNumber="95" value="2006-12-06">6 December 2006</date>
|
||
, W. Leonard, C. Richart, mm ff.
|
||
<emphasis id="B96CEA93FFE79D074E230694D98B23A0" italics="true" pageId="14" pageNumber="95">Wahakiaium Co.:</emphasis>
|
||
County Line Par, north of SR14, 46.1743°, -121.2183°,
|
||
<date id="FFA61041FFE79D07487006E8DD4F23A1" box="[834,1036,978,1004]" pageId="14" pageNumber="95" value="2004-02-22">22 February 2004</date>
|
||
, W. Leonard, mm ff; Rock Creek at Elochoman River, 46.2763°, -123.2796°,
|
||
<date id="FFA61041FFE79D07495B06CCDA66245D" box="[617,805,1014,1041]" pageId="14" pageNumber="95" value="2004-01-10">10 January 2004</date>
|
||
, W. Leonard, C. Richart, mm ff.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |