254 lines
33 KiB
XML
254 lines
33 KiB
XML
<document id="A2100FA151D4ED2DBA7B08B36B281CEB" ID-CLB-Dataset="298914" ID-DOI="10.11646/zootaxa.5474.1.3" ID-GBIF-Dataset="9a12ec35-d848-4b69-8cee-d67b7bcd5716" ID-ISSN="1175-5326" ID-Zenodo-Dep="12526909" ID-ZooBank="EE52D68D-7082-43D2-B326-026621AE36C5" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1719300134533" checkinUser="plazi" docAuthor="Moreno, Carlos & Zilberman, Bruno" docDate="2024" docId="03B187855C43FFFDABACD72BFF4E6541" docLanguage="en" docName="zootaxa.5474.1.3.pdf" docOrigin="Zootaxa 5474 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5474.1.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ceratoxenus tricornis Mann 1923" docType="treatment" docVersion="5" lastPageNumber="40" masterDocId="FF88FFFD5C40FFF8AB3BD348FF996720" masterDocTitle="Revalidation of the termitophilous genus Ceratoxenus Mann and description of two new species from the Neotropical region (Coleoptera: Staphylinidae: Aleocharinae: Corotocini)" masterLastPageNumber="49" masterPageNumber="35" pageNumber="38" updateTime="1719412577716" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
|
||
<mods:mods id="DCE0091D0A4259AEDF62B3474E9891AA" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="689828F9E1E0C96A02AD693ACBEB89C4">
|
||
<mods:title id="3623E85A92346F2C9861526E0DA802F8">Revalidation of the termitophilous genus Ceratoxenus Mann and description of two new species from the Neotropical region (Coleoptera: Staphylinidae: Aleocharinae: Corotocini)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="61CBC1D634821A7E25FC19470A284954" type="personal">
|
||
<mods:role id="1A9E3AC4E1EA0C3AE6E74E142DC10EF6">
|
||
<mods:roleTerm id="F2FB70B59D68C1A453DAF2B16F939F0E">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="B9DE95E67BD866D5B6023BABC326D09F">Moreno, Carlos</mods:namePart>
|
||
<mods:nameIdentifier id="80FD96B8BC2C22A376F317B53B3DB2BF" type="ORCID">0000-0002-6195-8648</mods:nameIdentifier>
|
||
<mods:affiliation id="3FD0996AC862D7542EC8B9349C7BA94A">Programa de Pós-Graduação em Sistemática, Taxonomia Animal e Biodiversidade, LaC-Laboratório de Coleoptera, Museu de Zoologia da Universidade de São Paulo, 04263 - 000, São Paulo, Brazil.</mods:affiliation>
|
||
<mods:nameIdentifier id="980728AAE5F044BD89F4312590BBD640" type="email">cmpiresilva@gmail.com</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="9C0DA0CCB2B2B1A700A225C32644394F" type="personal">
|
||
<mods:role id="1CA3E6A16999638501A8D286C835E440">
|
||
<mods:roleTerm id="EEA78644509F34F4A36CCA7F67C8003A">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="CE9917649FB68C79254B6926205B2524">Zilberman, Bruno</mods:namePart>
|
||
<mods:nameIdentifier id="03772BD9A6C9CE7C8C48790D3A51B190" type="ORCID">0000-0003-2613-4827</mods:nameIdentifier>
|
||
<mods:affiliation id="A6E81C7FF6FF26796E295B27133B66F7">Programa de Pós-Graduação em Sistemática, Taxonomia Animal e Biodiversidade, LaC-Laboratório de Coleoptera, Museu de Zoologia da Universidade de São Paulo, 04263 - 000, São Paulo, Brazil.</mods:affiliation>
|
||
<mods:nameIdentifier id="84F7BFB7D9CB2CDD8E3FE4781BDB8B31" type="email">brunozilberman@gmail.com</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:typeOfResource id="D9B774076E878012477E3B7CCEFC3572">text</mods:typeOfResource>
|
||
<mods:relatedItem id="CF9AF0AC5D9CE0A5DFA639833E1D3A3F" type="host">
|
||
<mods:titleInfo id="8557FD1E2E83C197C79B37D8937C5F12">
|
||
<mods:title id="8EA887E1F40B87377D3FBB78B2FB1BAC">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="573D941B30C0E0163B4E2599F5D8344B">
|
||
<mods:date id="8BA31CC38B63BD06E42A6A66D75BE548">2024</mods:date>
|
||
<mods:detail id="B00B5B7429BA6E1A9C8FD2974ACB2DC4" type="pubDate">
|
||
<mods:number id="5929D8234E3954265B90F7D45D5F0474">2024-06-25</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="0F944654235167414D6F7EFEB8FFE6B2" type="volume">
|
||
<mods:number id="C2607B8675518C2C47158EE997023F5E">5474</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="D1E6B9E97A2E53B2B04FA505B43D983E" type="issue">
|
||
<mods:number id="27E121ABACB048673B113509E21865D7">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="865F5A602EBE6054765873A9E2798304" unit="page">
|
||
<mods:start id="332E6D4B1CD3816ACB87C7DF56B21EB8">35</mods:start>
|
||
<mods:end id="69BC58A0BADE48E7BD37A154ABD1F3A8">49</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="89A11AFE69056EEB4EE9DDE818ED1EE1">
|
||
<mods:url id="CC33D5ACDD2130EF551B7B7BF663CF76">http://dx.doi.org/10.11646/zootaxa.5474.1.3</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="67ED2B6633DA26F30D940E61EF1556C7">journal article</mods:classification>
|
||
<mods:identifier id="6C8C30C40AC0C86DC7FF582ECF0FD620" type="CLB-Dataset">298914</mods:identifier>
|
||
<mods:identifier id="4F161A81FAD0AF6B971D65BDFD40E399" type="DOI">10.11646/zootaxa.5474.1.3</mods:identifier>
|
||
<mods:identifier id="11435532792C5970FB2ECEA20018058A" type="GBIF-Dataset">9a12ec35-d848-4b69-8cee-d67b7bcd5716</mods:identifier>
|
||
<mods:identifier id="2BF5D9325A13BEF036560B79C73CD0F6" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="E8AC5CD0F0449B7E84135BA4D6ABCCAF" type="Zenodo-Dep">12526909</mods:identifier>
|
||
<mods:identifier id="A23C7E050967AA09BD205E7AD9EFF839" type="ZooBank">EE52D68D-7082-43D2-B326-026621AE36C5</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03B187855C43FFFDABACD72BFF4E6541" ID-DOI="http://doi.org/10.5281/zenodo.12547281" ID-GBIF-Taxon="233791024" ID-Zenodo-Dep="12547281" LSID="urn:lsid:plazi:treatment:03B187855C43FFFDABACD72BFF4E6541" httpUri="http://treatment.plazi.org/id/03B187855C43FFFDABACD72BFF4E6541" lastPageId="5" lastPageNumber="40" pageId="3" pageNumber="38">
|
||
<subSubSection id="C30265185C43FFFBABACD72BFDA3635C" box="[151,570,1122,1149]" pageId="3" pageNumber="38" type="nomenclature">
|
||
<paragraph id="8BA736935C43FFFBABACD72BFDA3635C" blockId="3.[151,570,1122,1185]" box="[151,570,1122,1149]" pageId="3" pageNumber="38">
|
||
<heading id="D0EF81FF5C43FFFBABACD72BFDA3635C" bold="true" box="[151,570,1122,1149]" fontSize="11" level="1" pageId="3" pageNumber="38" reason="1">
|
||
<taxonomicName id="4C184D105C43FFFBABACD72BFDA3635C" ID-CoL="SQKK" authority="Mann, 1923" authorityName="Mann" authorityYear="1923" box="[151,570,1122,1149]" class="Insecta" family="Staphylinidae" genus="Ceratoxenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="38" phylum="Arthropoda" rank="species" species="tricornis">
|
||
<emphasis id="B96CEA815C43FFFBABACD72BFDA3635C" bold="true" box="[151,570,1122,1149]" pageId="3" pageNumber="38">
|
||
<emphasis id="B96CEA815C43FFFBABACD72BFE06635D" bold="true" box="[151,415,1123,1149]" italics="true" pageId="3" pageNumber="38">Ceratoxenus tricornis</emphasis>
|
||
<bibRefCitation id="EF894B625C43FFFBAA9DD72BFDA3635C" author="Mann, W. M." box="[422,570,1122,1149]" pageId="3" pageNumber="38" pagination="323 - 366" refId="ref7470" refString="Mann, W. M. (1923) New genera and species of termitophilous Coleoptera from Northern South America. Zoologica: scientific contributions of the New York Zoological Society, 3 (17), 323 - 366. https: // doi. org / 10.5962 / p. 184636" type="journal article" year="1923">Mann, 1923</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30265185C43FFFBABACD7CFFE496381" box="[151,464,1159,1185]" pageId="3" pageNumber="38" type="description">
|
||
<paragraph id="8BA736935C43FFFBABACD7CFFE496381" blockId="3.[151,570,1122,1185]" box="[151,464,1159,1185]" pageId="3" pageNumber="38">
|
||
(
|
||
<figureCitation id="13232A165C43FFFBABA4D7CFFEBC6381" box="[159,293,1159,1185]" captionStart="FIGURE 1" captionStartId="4.[151,250,1220,1244]" captionTargetBox="[153,1435,237,1197]" captionTargetId="figure-54@4.[151,1436,237,1197]" captionTargetPageId="4" captionText="FIGURE 1. Ceratoxenus tricornis Mann, 1923. Holotype. Habitus lateral (A), ventral (B), dorsal (C), and labels (D)." figureDoi="http://doi.org/10.5281/zenodo.12526911" httpUri="https://zenodo.org/record/12526911/files/figure.png" pageId="3" pageNumber="38">Figs. 1A–D</figureCitation>
|
||
,
|
||
<figureCitation id="13232A165C43FFFBAA0AD7CFFEEE6381" box="[305,375,1159,1185]" captionStart="FIGURE 2" captionStartId="4.[151,250,1989,2013]" captionTargetBox="[151,1436,1304,1965]" captionTargetId="figure-74@4.[151,1436,1304,1965]" captionTargetPageId="4" captionText="FIGURE 2. Ceratoxenus tricornis Mann, 1923. Paratype. Habitus lateral (A), dorsal (C), and labels (B)." figureDoi="http://doi.org/10.5281/zenodo.12526915" httpUri="https://zenodo.org/record/12526915/files/figure.png" pageId="3" pageNumber="38">2A–C</figureCitation>
|
||
,
|
||
<figureCitation id="13232A165C43FFFBAAB9D7CFFE5E6381" box="[386,455,1159,1185]" captionStart="FIGURE 6" captionStartId="9.[151,250,1847,1871]" captionTargetBox="[151,1436,885,1824]" captionTargetId="figure-305@9.[151,1436,885,1824]" captionTargetPageId="9" captionText="FIGURE 6. Comparative external morphology of head and thorax among Ceratoxenus species: C. tricornis Mann, 1923 (A, D), C. inflatus sp. nov. (C, E), and C. akerata sp. nov. (C, F). White arrows point to the lateral horns, yellow arrows point to the gena with a distinctive carina in C. inflatus and C. akerata, and uncarinated in C. tricornis, while cyan arrows point to the medial projection between lateral horns." figureDoi="http://doi.org/10.5281/zenodo.12526937" httpUri="https://zenodo.org/record/12526937/files/figure.png" pageId="3" pageNumber="38">6A, D</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30265185C43FFFBABACD783FEFD6201" pageId="3" pageNumber="38" type="reference_group">
|
||
<paragraph id="8BA736935C43FFFBABACD783FC9A63C3" blockId="3.[151,1436,1227,1313]" box="[151,771,1227,1251]" pageId="3" pageNumber="38">
|
||
<treatmentCitationGroup id="AB0811BD5C43FFFBABACD783FC9A63C3" box="[151,771,1227,1251]" pageId="3" pageNumber="38">
|
||
<treatmentCitation id="0AB910825C43FFFBABACD783FD8763C3" author="Mann, W. M." box="[151,542,1227,1251]" page="360" pageId="3" pageNumber="38" year="1923">
|
||
<taxonomicName id="4C184D105C43FFFBABACD783FD8763C3" ID-CoL="SQKK" authority="Mann 1923: 360" authorityName="Mann" authorityPageNumber="360" authorityYear="1923" box="[151,542,1227,1251]" class="Insecta" family="Staphylinidae" genus="Ceratoxenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="38" phylum="Arthropoda" rank="species" species="tricornis">
|
||
Ceratoxenus tricornis
|
||
<bibRefCitation id="EF894B625C43FFFBAA4CD784FD8763C3" author="Mann, W. M." box="[375,542,1227,1251]" pageId="3" pageNumber="38" pagination="323 - 366" refId="ref7470" refString="Mann, W. M. (1923) New genera and species of termitophilous Coleoptera from Northern South America. Zoologica: scientific contributions of the New York Zoological Society, 3 (17), 323 - 366. https: // doi. org / 10.5962 / p. 184636" type="journal article" year="1923">Mann 1923: 360</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
(original description);
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BA736935C43FFFBABACD7A2FEFD6201" blockId="3.[151,1436,1227,1313]" pageId="3" pageNumber="38">
|
||
<treatmentCitationGroup id="AB0811BD5C43FFFBABACD7A2FEFD6201" pageId="3" pageNumber="38">
|
||
<taxonomicName id="4C184D105C43FFFBABACD7A2FEC36221" ID-CoL="7FVVD" baseAuthorityName="Mann" baseAuthorityYear="1923" box="[151,346,1258,1282]" class="Insecta" family="Staphylinidae" genus="Xenopelta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="38" phylum="Arthropoda" rank="species" species="tricornis">
|
||
<emphasis id="B96CEA815C43FFFBABACD7A2FEC36221" box="[151,346,1258,1282]" italics="true" pageId="3" pageNumber="38">Xenopelta tricornis</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<treatmentCitation id="0AB910825C43FFFBAA5DD7A2FDB06222" author="Emerson, A. E." box="[358,553,1258,1282]" page="369" pageId="3" pageNumber="38" year="1935">
|
||
<bibRefCitation id="EF894B625C43FFFBAA5DD7A2FDB06222" author="Emerson, A. E." box="[358,553,1258,1282]" pageId="3" pageNumber="38" pagination="369 - 395" refId="ref7276" refString="Emerson, A. E. (1935) Termitophile distribution and quantitative characters as indicators of physiological speciation in British Guiana termites (Isoptera). Annals of the Entomological Society of America, 28, 369 - 395. https: // doi. org / 10.1093 / aesa / 28.3.369" type="journal article" year="1935">Emerson 1935: 369</bibRefCitation>
|
||
</treatmentCitation>
|
||
<treatmentCitation id="0AB910825C43FFFBA915D7A2FD7F6222" author="Seevers, C. H." box="[558,742,1258,1282]" page="107" pageId="3" pageNumber="38" year="1957">
|
||
<bibRefCitation id="EF894B625C43FFFBA915D7A2FD7F6222" author="Seevers, C. H." box="[558,742,1258,1282]" pageId="3" pageNumber="38" refId="ref7789" refString="Seevers, C. H. (1957) A monograph on the termitophilous Staphylinidae (Coleoptera). Chicago Natural History Museum, Chicago, Illinois, 334 pp. https: // doi. org / 10.5962 / bhl. title. 3797" type="book" year="1957">Seevers 1957: 107</bibRefCitation>
|
||
</treatmentCitation>
|
||
(redescription as
|
||
<taxonomicName id="4C184D105C43FFFBA8ADD7A3FB9E6221" ID-CoL="7FVVD" baseAuthorityName="Mann" baseAuthorityYear="1923" box="[918,1031,1258,1282]" class="Insecta" family="Staphylinidae" genus="Xenopelta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="38" phylum="Arthropoda" rank="species" species="tricornis">
|
||
<emphasis id="B96CEA815C43FFFBA8ADD7A3FB9E6221" box="[918,1031,1258,1282]" italics="true" pageId="3" pageNumber="38">X. tricornis</emphasis>
|
||
</taxonomicName>
|
||
);
|
||
<treatmentCitation id="0AB910825C43FFFBAF27D7A2FA936222" author="Jacobson, H. R. & Kistner, D. H. & Pasteels, J. M." box="[1052,1290,1258,1282]" page="64" pageId="3" pageNumber="38" year="1986">
|
||
<bibRefCitation id="EF894B625C43FFFBAF27D7A2FA936222" author="Jacobson, H. R. & Kistner, D. H. & Pasteels, J. M." box="[1052,1290,1258,1282]" pageId="3" pageNumber="38" pagination="1 - 245" refId="ref7382" refString="Jacobson, H. R., Kistner, D. H. & Pasteels, J. M. (1986) Generic revision, phylogenetic classification, and phylogeny of the termitophilous tribe Corotocini (Coleoptera: Staphylinidae). Sociobiology, 12, 1 - 245." type="journal article" year="1986">
|
||
Jacobson
|
||
<emphasis id="B96CEA815C43FFFBAF46D7A3FB316222" box="[1149,1192,1258,1282]" italics="true" pageId="3" pageNumber="38">et al</emphasis>
|
||
. 1986: 64
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
(redescription as
|
||
<taxonomicName id="4C184D105C43FFFBABD8D641FECF6200" ID-CoL="7FVVD" baseAuthorityName="Mann" baseAuthorityYear="1923" box="[227,342,1289,1312]" class="Insecta" family="Staphylinidae" genus="Xenopelta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="38" phylum="Arthropoda" rank="species" species="tricornis">
|
||
<emphasis id="B96CEA815C43FFFBABD8D641FECF6200" box="[227,342,1289,1312]" italics="true" pageId="3" pageNumber="38">X. tricornis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30265185C43FFFDABACD606FDDF6791" lastPageId="5" lastPageNumber="40" pageId="3" pageNumber="38" type="diagnosis">
|
||
<paragraph id="8BA736935C43FFFBABACD606FAD662F5" blockId="3.[151,1437,1358,2033]" pageId="3" pageNumber="38">
|
||
<emphasis id="B96CEA815C43FFFBABACD606FE936248" bold="true" box="[151,266,1358,1384]" pageId="3" pageNumber="38">Diagnosis</emphasis>
|
||
: The species is easily recognized by the presence of long bristles on vertex, head lateral projections composed of two long and broad frontal processes in front of each antennal insertion, transpassing the antennal insertions anteriorly; medial process between lateral projection weak and obtuse, gena straight, not carinate; pronotum with anterior impression longer and slightly narrower than posterior, and anterior margin straight.
|
||
</paragraph>
|
||
<paragraph id="8BA736935C43FFFBABFCD697FB4B6039" blockId="3.[151,1437,1358,2033]" pageId="3" pageNumber="38">
|
||
<emphasis id="B96CEA815C43FFFBABFCD697FE9E62D9" bold="true" box="[199,263,1503,1529]" pageId="3" pageNumber="38">Head</emphasis>
|
||
with gena straight; long bristles present on vertex; surface roughly scamose; head lateral projections at frons composed of two long and broad frontal processes in front of each antennal insertion, transpassing the antennal insertions anteriorly; medial process acutely projected, not overtaking the level of frontal elevations; frontoclypeal suture with weak and obtuse medial process, between frontal projections (
|
||
<figureCitation id="13232A165C43FFFBAF05D503FB5D6145" box="[1086,1220,1611,1637]" captionStart="FIGURE 1" captionStartId="4.[151,250,1220,1244]" captionTargetBox="[153,1435,237,1197]" captionTargetId="figure-54@4.[151,1436,237,1197]" captionTargetPageId="4" captionText="FIGURE 1. Ceratoxenus tricornis Mann, 1923. Holotype. Habitus lateral (A), ventral (B), dorsal (C), and labels (D)." figureDoi="http://doi.org/10.5281/zenodo.12526911" httpUri="https://zenodo.org/record/12526911/files/figure.png" pageId="3" pageNumber="38">Figs. 1A–B</figureCitation>
|
||
,
|
||
<figureCitation id="13232A165C43FFFBAFE9D503FA806145" box="[1234,1305,1611,1637]" captionStart="FIGURE 2" captionStartId="4.[151,250,1989,2013]" captionTargetBox="[151,1436,1304,1965]" captionTargetId="figure-74@4.[151,1436,1304,1965]" captionTargetPageId="4" captionText="FIGURE 2. Ceratoxenus tricornis Mann, 1923. Paratype. Habitus lateral (A), dorsal (C), and labels (B)." figureDoi="http://doi.org/10.5281/zenodo.12526915" httpUri="https://zenodo.org/record/12526915/files/figure.png" pageId="3" pageNumber="38">2A, C</figureCitation>
|
||
,
|
||
<figureCitation id="13232A165C43FFFBAE1DD503FAD56145" box="[1318,1356,1611,1637]" captionStart="FIGURE 6" captionStartId="9.[151,250,1847,1871]" captionTargetBox="[151,1436,885,1824]" captionTargetId="figure-305@9.[151,1436,885,1824]" captionTargetPageId="9" captionText="FIGURE 6. Comparative external morphology of head and thorax among Ceratoxenus species: C. tricornis Mann, 1923 (A, D), C. inflatus sp. nov. (C, E), and C. akerata sp. nov. (C, F). White arrows point to the lateral horns, yellow arrows point to the gena with a distinctive carina in C. inflatus and C. akerata, and uncarinated in C. tricornis, while cyan arrows point to the medial projection between lateral horns." figureDoi="http://doi.org/10.5281/zenodo.12526937" httpUri="https://zenodo.org/record/12526937/files/figure.png" pageId="3" pageNumber="38">6A</figureCitation>
|
||
). Head surface somewhat scamose. Antenna with antennomere 1 cylindrical, with almost same length of antennomeres 2–4 combined; 2–4 longer than wide, subequal in length; 5–7 shorter than previous, longer than wide, about same length; 8 slightly shorter than previous segments and longer than segments 9–10, subquadrate; 9–10 transverse, subequal. Secondary maxilar palpomere slightly rounded and inflated at apex. Postmentum occupying more than half of total head length, longer than wide, slightly longer than gula, entirely covered by bristles.
|
||
</paragraph>
|
||
<paragraph id="8BA736935C43FFFCABFCD46BFC9267F5" blockId="3.[151,1437,1358,2033]" lastBlockId="4.[151,1436,151,213]" lastPageId="4" lastPageNumber="39" pageId="3" pageNumber="38">
|
||
<emphasis id="B96CEA815C43FFFBABFCD46BFEB9601D" bold="true" box="[199,288,1827,1853]" pageId="3" pageNumber="38">Thorax</emphasis>
|
||
. Pronotal disc light brown with lateral margins darker; anterior impression longer and slightly narrower than posterior, and with anterior margin straight; bearing long bristles marginally; posterior half of pronotal disc with circular impression occupying almost entire pronotal width, with slight circular elevation medially; anterior region slightly elevated behind head, with one row of three bristles; lateral margins parallel with longitudinal row of three long bristles symmetrically distributed (
|
||
<figureCitation id="13232A165C43FFFBA9AFD4FBFD7260ED" box="[660,747,1971,1997]" captionStart="FIGURE 1" captionStartId="4.[151,250,1220,1244]" captionTargetBox="[153,1435,237,1197]" captionTargetId="figure-54@4.[151,1436,237,1197]" captionTargetPageId="4" captionText="FIGURE 1. Ceratoxenus tricornis Mann, 1923. Holotype. Habitus lateral (A), ventral (B), dorsal (C), and labels (D)." figureDoi="http://doi.org/10.5281/zenodo.12526911" httpUri="https://zenodo.org/record/12526911/files/figure.png" pageId="3" pageNumber="38">Fig. 1C</figureCitation>
|
||
,
|
||
<figureCitation id="13232A165C43FFFBA9CCD4FBFC8560ED" box="[759,796,1971,1997]" captionStart="FIGURE 2" captionStartId="4.[151,250,1989,2013]" captionTargetBox="[151,1436,1304,1965]" captionTargetId="figure-74@4.[151,1436,1304,1965]" captionTargetPageId="4" captionText="FIGURE 2. Ceratoxenus tricornis Mann, 1923. Paratype. Habitus lateral (A), dorsal (C), and labels (B)." figureDoi="http://doi.org/10.5281/zenodo.12526915" httpUri="https://zenodo.org/record/12526915/files/figure.png" pageId="3" pageNumber="38">2C</figureCitation>
|
||
). Elytra elongate, wider on posterior region; light-brown, outer margin darker. Legs with pro- and mesocoxa transverse; metacoxa triangular. Tibia slightly longer than femur, covered with long and thick bristles along entire length; outer margins of tibia slightly convex, and inner margin straight; pro-, meso-, and metatarsi increasing in length.
|
||
</paragraph>
|
||
<caption id="DF67661B5C44FFFCABACD78CFAA763FC" ID-DOI="http://doi.org/10.5281/zenodo.12526911" ID-Zenodo-Dep="12526911" box="[151,1342,1220,1244]" httpUri="https://zenodo.org/record/12526911/files/figure.png" pageId="4" pageNumber="39" startId="4.[151,250,1220,1244]" targetBox="[153,1435,237,1197]" targetPageId="4" targetType="figure">
|
||
<paragraph id="8BA736935C44FFFCABACD78CFAA763FC" blockId="4.[151,1342,1220,1244]" box="[151,1342,1220,1244]" pageId="4" pageNumber="39">
|
||
<emphasis id="B96CEA815C44FFFCABACD78CFE8A63FC" bold="true" box="[151,275,1220,1244]" pageId="4" pageNumber="39">FIGURE 1.</emphasis>
|
||
<taxonomicName id="4C184D105C44FFFCAA22D78DFDE263FC" authority="Mann, 1923" authorityName="Mann" authorityYear="1923" box="[281,635,1220,1244]" class="Insecta" family="Staphylinidae" genus="Ceratoxenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="tricornis">
|
||
<emphasis id="B96CEA815C44FFFCAA22D78DFE6E63FC" box="[281,503,1221,1244]" italics="true" pageId="4" pageNumber="39">Ceratoxenus tricornis</emphasis>
|
||
<bibRefCitation id="EF894B625C44FFFCAAC6D78DFDE263FC" author="Mann, W. M." box="[509,635,1220,1244]" pageId="4" pageNumber="39" pagination="323 - 366" refId="ref7470" refString="Mann, W. M. (1923) New genera and species of termitophilous Coleoptera from Northern South America. Zoologica: scientific contributions of the New York Zoological Society, 3 (17), 323 - 366. https: // doi. org / 10.5962 / p. 184636" type="journal article" year="1923">Mann, 1923</bibRefCitation>
|
||
</taxonomicName>
|
||
. Holotype. Habitus lateral (A), ventral (B), dorsal (C), and labels (D).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF67661B5C44FFFCABACD48DFB2560FD" ID-DOI="http://doi.org/10.5281/zenodo.12526915" ID-Zenodo-Dep="12526915" box="[151,1212,1989,2014]" httpUri="https://zenodo.org/record/12526915/files/figure.png" pageId="4" pageNumber="39" startId="4.[151,250,1989,2013]" targetBox="[151,1436,1304,1965]" targetPageId="4" targetType="figure">
|
||
<paragraph id="8BA736935C44FFFCABACD48DFB2560FD" blockId="4.[151,1212,1989,2014]" box="[151,1212,1989,2014]" pageId="4" pageNumber="39">
|
||
<emphasis id="B96CEA815C44FFFCABACD48DFE8A60FD" bold="true" box="[151,275,1989,2013]" pageId="4" pageNumber="39">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C184D105C44FFFCAA22D48EFDE260FD" authority="Mann, 1923" authorityName="Mann" authorityYear="1923" box="[281,635,1989,2013]" class="Insecta" family="Staphylinidae" genus="Ceratoxenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="tricornis">
|
||
<emphasis id="B96CEA815C44FFFCAA22D48EFE6E60FD" box="[281,503,1990,2013]" italics="true" pageId="4" pageNumber="39">Ceratoxenus tricornis</emphasis>
|
||
<bibRefCitation id="EF894B625C44FFFCAAC6D48EFDE260FD" author="Mann, W. M." box="[509,635,1989,2013]" pageId="4" pageNumber="39" pagination="323 - 366" refId="ref7470" refString="Mann, W. M. (1923) New genera and species of termitophilous Coleoptera from Northern South America. Zoologica: scientific contributions of the New York Zoological Society, 3 (17), 323 - 366. https: // doi. org / 10.5962 / p. 184636" type="journal article" year="1923">Mann, 1923</bibRefCitation>
|
||
</taxonomicName>
|
||
. Paratype. Habitus lateral (A), dorsal (C), and labels (B).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA736935C45FFFDABFCD3DEFDDF6791" blockId="5.[151,1437,150,609]" box="[199,582,150,177]" pageId="5" pageNumber="40">
|
||
<emphasis id="B96CEA815C45FFFDABFCD3DEFEDA6790" bold="true" box="[199,323,150,176]" pageId="5" pageNumber="40">Abdomen.</emphasis>
|
||
As described to genus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30265185C45FFFDABFCD3F2FE946661" pageId="5" pageNumber="40" type="materials_examined">
|
||
<paragraph id="8BA736935C45FFFDABFCD3F2FE946661" blockId="5.[151,1437,150,609]" pageId="5" pageNumber="40">
|
||
<emphasis id="B96CEA815C45FFFDABFCD3F2FE3467F4" bold="true" box="[199,429,186,212]" pageId="5" pageNumber="40">Material examined</emphasis>
|
||
.
|
||
<materialsCitation id="3B703CCE5C45FFFDAA84D3F2FE8E663D" ID-GBIF-Occurrence="4899569304" collectingDate="1920-08-22" collectorName="W. M. Mann" country="Guyana" county="Bartica District" location="Aemerson" municipality="Kartabo" pageId="5" pageNumber="40" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="B96CEA815C45FFFDAA84D3F2FDC767F4" bold="true" box="[447,606,186,212]" pageId="5" pageNumber="40">
|
||
<typeStatus id="54A388315C45FFFDAA84D3F2FDC767F4" box="[447,606,186,212]" pageId="5" pageNumber="40" type="holotype">HOLOTYPE</typeStatus>
|
||
</emphasis>
|
||
. “Kartabo P., Br. Guiana,
|
||
<location id="8EC760485C45FFFDA8AFD3F3FB9E67F5" LSID="urn:lsid:plazi:treatment:03B187855C43FFFDABACD72BFF4E6541:8EC760485C45FFFDA8AFD3F3FB9E67F5" box="[916,1031,187,213]" country="Guyana" county="Bartica District" municipality="Kartabo" name="Aemerson" pageId="5" pageNumber="40">Aemerson</location>
|
||
[col.]”; “
|
||
<collectingMunicipality id="6BC3ACE95C45FFFDAF4CD3F3FB4F67F5" box="[1143,1238,187,213]" pageId="5" pageNumber="40">Kartabo</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="62C64E1F5C45FFFDAFDED3F3FA0167F5" box="[1253,1432,187,213]" pageId="5" pageNumber="40">Bartica District</collectingCounty>
|
||
,
|
||
<collectingCountry id="F30F76035C45FFFDABACD397FEA567D9" box="[151,316,223,249]" name="Guyana" pageId="5" pageNumber="40">British Guiana</collectingCountry>
|
||
,
|
||
<date id="FFA610535C45FFFDAA7FD397FE4567D9" box="[324,476,223,249]" pageId="5" pageNumber="40" value="1920-08-22">
|
||
<collectingDate id="EFE2E9BB5C45FFFDAA7FD397FE4567D9" box="[324,476,223,249]" pageId="5" pageNumber="40" value="1920-08-22">22.VIII.1920</collectingDate>
|
||
</date>
|
||
”; “WMMann [
|
||
<collectorName id="26ED53455C45FFFDA940D397FC9567D9" box="[635,780,223,249]" pageId="5" pageNumber="40">W. M. Mann</collectorName>
|
||
], cooln, 1940”; “Type No., 64133, U.S.N.M.”; “USNMENT 01374929”
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B703CCE5C45FFFDAA13D24AFE906661" ID-GBIF-Occurrence="4899569305" collectorName="Coll. By A. Emerson" country="Guyana" location="British Guiana" pageId="5" pageNumber="40" specimenCount="1" typeStatus="paratype">
|
||
<emphasis id="B96CEA815C45FFFDAA13D24AFE25663C" bold="true" box="[296,444,258,284]" pageId="5" pageNumber="40">
|
||
<typeStatus id="54A388315C45FFFDAA13D24AFE25663C" box="[296,444,258,284]" pageId="5" pageNumber="40" type="paratype">PARATYPE</typeStatus>
|
||
</emphasis>
|
||
. “Kartabo,
|
||
<collectingCountry id="F30F76035C45FFFDA970D24BFD61663D" box="[587,760,259,285]" name="Guyana" pageId="5" pageNumber="40">British Guiana</collectingCountry>
|
||
, 15:X:1920”; “
|
||
<collectorName id="26ED53455C45FFFDA896D24BFB35663D" box="[941,1196,259,285]" pageId="5" pageNumber="40">Coll. By A. Emerson</collectorName>
|
||
#424”; “FMNHINS 4431876”
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30265185C45FFFDABFCD203FD9666F1" pageId="5" pageNumber="40" type="discussion">
|
||
<paragraph id="8BA736935C45FFFDABFCD203FD9666F1" blockId="5.[151,1437,150,609]" pageId="5" pageNumber="40">
|
||
<emphasis id="B96CEA815C45FFFDABFCD203FEAD6645" bold="true" box="[199,308,331,357]" pageId="5" pageNumber="40">Remarks</emphasis>
|
||
.
|
||
<taxonomicName id="4C184D105C45FFFDAA79D203FDAF6644" authorityName="Mann" authorityYear="1923" box="[322,566,331,357]" class="Insecta" family="Staphylinidae" genus="Ceratoxenus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="40" phylum="Arthropoda" rank="species" species="tricornis">
|
||
<emphasis id="B96CEA815C45FFFDAA79D203FDAF6644" box="[322,566,331,357]" italics="true" pageId="5" pageNumber="40">Ceratoxenus tricornis</emphasis>
|
||
</taxonomicName>
|
||
is characterized especially by the presence of the two long and narrow frontal horns anterior to and transpassing antennal insertions, presence of vertexal bristles, and a straight and uncarinated gena; pronotal impressions are also distinct, with the anterior half longer and just slightly narrower than posterior, and with anterior margin straight.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30265185C45FFFDABFCD292FDA566D5" box="[199,572,474,501]" pageId="5" pageNumber="40" type="distribution">
|
||
<paragraph id="8BA736935C45FFFDABFCD292FDA566D5" blockId="5.[151,1437,150,609]" box="[199,572,474,501]" pageId="5" pageNumber="40">
|
||
<emphasis id="B96CEA815C45FFFDABFCD292FEC066D4" bold="true" box="[199,345,474,500]" pageId="5" pageNumber="40">Distribution</emphasis>
|
||
.
|
||
<collectingCountry id="F30F76035C45FFFDAA5CD293FE2666D5" box="[359,447,475,501]" name="Guyana" pageId="5" pageNumber="40">Guyana</collectingCountry>
|
||
—Kartabo.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30265185C45FFFDABFCD2B6FF4E6541" pageId="5" pageNumber="40" type="biology_ecology">
|
||
<paragraph id="8BA736935C45FFFDABFCD2B6FF4E6541" blockId="5.[151,1437,150,609]" pageId="5" pageNumber="40">
|
||
<emphasis id="B96CEA815C45FFFDABFCD2B6FEB86538" bold="true" box="[199,289,510,536]" pageId="5" pageNumber="40">Biology</emphasis>
|
||
. The host of
|
||
<taxonomicName id="4C184D105C45FFFDAA8DD2B7FDAE6538" authorityName="Mann" authorityYear="1923" box="[438,567,511,537]" class="Insecta" family="Staphylinidae" genus="Ceratoxenus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="40" phylum="Arthropoda" rank="species" species="tricornis">
|
||
<emphasis id="B96CEA815C45FFFDAA8DD2B7FDAE6538" box="[438,567,511,537]" italics="true" pageId="5" pageNumber="40">C. tricornis</emphasis>
|
||
</taxonomicName>
|
||
recorded in
|
||
<bibRefCitation id="EF894B625C45FFFDA9FFD2B7FCCE6539" author="Mann, W. M." box="[708,855,511,537]" pageId="5" pageNumber="40" pagination="323 - 366" refId="ref7470" refString="Mann, W. M. (1923) New genera and species of termitophilous Coleoptera from Northern South America. Zoologica: scientific contributions of the New York Zoological Society, 3 (17), 323 - 366. https: // doi. org / 10.5962 / p. 184636" type="journal article" year="1923">Mann (1923)</bibRefCitation>
|
||
was
|
||
<taxonomicName id="4C184D105C45FFFDA8AAD2B7FAF46539" authority="(Holmgren, 1910)" authorityName="Mann" authorityYear="1923" baseAuthorityName="Holmgren" baseAuthorityYear="1910" box="[913,1389,511,537]" class="Insecta" family="Termitidae" genus="Nasutitermes" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="40" phylum="Arthropoda" rank="species" species="guayanae">
|
||
<emphasis id="B96CEA815C45FFFDA8AAD2B7FB006538" box="[913,1177,511,536]" italics="true" pageId="5" pageNumber="40">Nasutitermes guayanae</emphasis>
|
||
(Holmgren, 1910)
|
||
</taxonomicName>
|
||
, but data in
|
||
<bibRefCitation id="EF894B625C45FFFDABD7D16BFE3B651D" author="Emerson, A. E." box="[236,418,546,573]" pageId="5" pageNumber="40" pagination="369 - 395" refId="ref7276" refString="Emerson, A. E. (1935) Termitophile distribution and quantitative characters as indicators of physiological speciation in British Guiana termites (Isoptera). Annals of the Entomological Society of America, 28, 369 - 395. https: // doi. org / 10.1093 / aesa / 28.3.369" type="journal article" year="1935">Emerson (1935)</bibRefCitation>
|
||
leaves little doubt if the host is actually the newly described species,
|
||
<taxonomicName id="4C184D105C45FFFDAF85D16BFF4B6541" authority="Emerson, 1935" authorityName="Emerson" authorityYear="1935" class="Insecta" family="Termitidae" genus="Nasutitermes" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="40" phylum="Arthropoda" rank="species" species="similis">
|
||
<emphasis id="B96CEA815C45FFFDAF85D16BFAB0651D" box="[1214,1321,547,573]" italics="true" pageId="5" pageNumber="40">N. similis</emphasis>
|
||
<bibRefCitation id="EF894B625C45FFFDAE09D16BFF4B6541" author="Emerson, A. E." pageId="5" pageNumber="40" pagination="369 - 395" refId="ref7276" refString="Emerson, A. E. (1935) Termitophile distribution and quantitative characters as indicators of physiological speciation in British Guiana termites (Isoptera). Annals of the Entomological Society of America, 28, 369 - 395. https: // doi. org / 10.1093 / aesa / 28.3.369" type="journal article" year="1935">Emerson, 1935</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |