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<document id="11EAEDE1AD75B227877CA14EDB28CC3E" ID-DOI="10.11646/zootaxa.4755.2.3" ID-GBIF-Dataset="60cf9eb6-409b-40db-a6e6-a0bfadeb06fa" ID-ISSN="1175-5334" ID-Zenodo-Dep="3731581" ID-ZooBank="urn:lsid:zoobank.org:pub:8D3A61EC-17A4-4824-9D67-CAF1DECEDB36" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1585338343226" checkinUser="carolina" docAuthor="Jałoszyński, Paweł" docDate="2020" docId="038587BCC07A4C783C8EFF30D904FC24" docLanguage="en" docName="zootaxa.4755.2.3.pdf.imf" docOrigin="Zootaxa 4755 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Venezolanoconnus asserculatus Jałoszyński 2020, comb. n." docType="treatment" docVersion="7" lastPageNumber="268" masterDocId="FFBCFFC4C06A4C693C19FFA9DC31FFA7" masterDocTitle="Corrections to the classification of the Neotropical ant-like stone beetles misplaced in Euconnus Thomson (Coleoptera, Staphylinidae, Scydmaeninae). Part 1." masterLastPageNumber="270" masterPageNumber="251" pageNumber="267" updateTime="1698732931681" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="4339FBC29FB1A43530EE324DFC90E9AA">Corrections to the classification of the Neotropical ant-like stone beetles misplaced in Euconnus Thomson (Coleoptera, Staphylinidae, Scydmaeninae). Part 1.</mods:title>
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<paragraph id="8B9336AAC07A4C793C8EFF30DF5CFF13" blockId="16.[151,877,153,180]" box="[151,877,153,180]" pageId="16" pageNumber="267">
<heading id="D0DB81C6C07A4C793C8EFF30DF5CFF13" bold="true" box="[151,877,153,180]" fontSize="10" level="2" pageId="16" pageNumber="267" reason="2">
<emphasis id="B958EAB8C07A4C793C8EFF30DF5CFF13" bold="true" box="[151,877,153,180]" pageId="16" pageNumber="267">
<taxonomicName id="4C2C4D29C07A4C793C8EFF30DEC7FF14" ID-CoL="7FLN5" authority="(L. W. Schaufuss)" authorityName="Jałoszyński" authorityYear="2020" baseAuthorityName="L. W. Schaufuss" baseAuthorityYear="2020" box="[151,758,153,180]" class="Insecta" family="Staphylinidae" genus="Venezolanoconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="267" phylum="Arthropoda" rank="species" species="asserculatus" status="comb. nov.">
<emphasis id="B958EAB8C07A4C793C8EFF30DE26FF13" bold="true" box="[151,535,153,180]" italics="true" pageId="16" pageNumber="267">Venezolanoconnus asserculatus</emphasis>
(L.W. Schaufuss)
</taxonomicName>
,
<taxonomicNameLabel id="A26B57C3C07A4C793F1BFF33DF5CFF13" box="[770,877,154,180]" pageId="16" pageNumber="267" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C3366521C07A4C793C8EFF77DFF7FE94" pageId="16" pageNumber="267" type="reference_group">
<paragraph id="8B9336AAC07A4C793C8EFF77DE9AFF51" blockId="16.[151,966,221,343]" box="[151,683,221,246]" pageId="16" pageNumber="267">
<treatmentCitationGroup id="AB3C1184C07A4C793C8EFF77DE9AFF51" box="[151,683,221,246]" pageId="16" pageNumber="267">
<treatmentCitation id="0A8D10BBC07A4C793C8EFF77DE97FF51" author="Schaufuss, L. W." box="[151,678,221,246]" page="68" pageId="16" pageNumber="267" year="1866">
<taxonomicName id="4C2C4D29C07A4C793C8EFF77DE97FF51" ID-CoL="4W4BV" authority="L. W. Schaufuss, 1866: 68" authorityName="L. W. Schaufuss" authorityPageNumber="68" authorityYear="1866" box="[151,678,221,246]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="267" phylum="Arthropoda" rank="species" species="asserculatus">
<emphasis id="B958EAB8C07A4C793C8EFF77DDA8FF51" box="[151,409,222,246]" italics="true" pageId="16" pageNumber="267">Scydmaenus asserculatus</emphasis>
L.W.
<bibRefCitation id="EFBD4B5BC07A4C793DCEFF74DE97FF51" author="Schaufuss, L. W." box="[471,678,221,246]" pageId="16" pageNumber="267" pagination="1 - 103" refId="ref11922" refString="Schaufuss, L. W. (1866) Monographie der Scydmaeniden Central- und Sudamerika's. Novorum Actorum Academiae Caesareae Leopoldino-Carolinae Germanicae Naturae Curiosorum, 33 (6), 1 - 103, pls. 1 - 4. [1867]" type="journal article" year="1866">Schaufuss, 1866: 68</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9336AAC07A4C793C8EFF54DF0DFEB2" blockId="16.[151,966,221,343]" box="[151,828,252,277]" pageId="16" pageNumber="267">
<treatmentCitationGroup id="AB3C1184C07A4C793C8EFF54DF0DFEB2" box="[151,828,252,277]" pageId="16" pageNumber="267">
<taxonomicName id="4C2C4D29C07A4C793C8EFF54DEB5FEB2" authority="(L. W. Schaufuss)" baseAuthorityName="L. W. Schaufuss" box="[151,644,252,277]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="267" phylum="Arthropoda" rank="species" species="asserculatus">
<emphasis id="B958EAB8C07A4C793C8EFF54DCCAFEB3" box="[151,251,253,276]" italics="true" pageId="16" pageNumber="267">Euconnus</emphasis>
(s. str.)
<emphasis id="B958EAB8C07A4C793D54FF55DDFAFEB3" box="[333,459,252,276]" italics="true" pageId="16" pageNumber="267">asserculatus</emphasis>
(L.W. Schaufuss)
</taxonomicName>
;
<treatmentCitation id="0A8D10BBC07A4C793E96FF55DF06FEB2" author="Csiki, E." box="[655,823,252,277]" page="45" pageId="16" pageNumber="267" year="1919">
<bibRefCitation id="EFBD4B5BC07A4C793E96FF55DF23FEB3" author="Csiki, E." box="[655,786,252,277]" pageId="16" pageNumber="267" pagination="1 - 106" refId="ref11279" refString="Csiki, E. (1919) Scydmaenidae. In: Schenkling, S. &amp; Junk, W. (Eds.), Coleopterorum Catalogus. Pars 70. W Junk, Berlin, pp. 1 - 106." type="book chapter" year="1919">Csiki (1919)</bibRefCitation>
: 45
</treatmentCitation>
.
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</paragraph>
<paragraph id="8B9336AAC07A4C793C8EFEB5DFF7FE94" blockId="16.[151,966,221,343]" box="[151,966,282,307]" pageId="16" pageNumber="267">
<treatmentCitationGroup id="AB3C1184C07A4C793C8EFEB5DFF7FE94" box="[151,966,282,307]" pageId="16" pageNumber="267">
<taxonomicName id="4C2C4D29C07A4C793C8EFEB5DCCAFE94" authorityName="in Reitter &amp; Croissandeau" authorityYear="1890" box="[151,251,284,307]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="267" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAB8C07A4C793C8EFEB5DCCAFE94" box="[151,251,284,307]" italics="true" pageId="16" pageNumber="267">Euconnus</emphasis>
</taxonomicName>
(
<emphasis id="B958EAB8C07A4C793D13FEB5DDA7FE94" box="[266,406,283,307]" italics="true" pageId="16" pageNumber="267">incertae sedis</emphasis>
)
<taxonomicName id="4C2C4D29C07A4C793DBCFEB2DEEDFE94" authority="(L. W. Schaufuss)" baseAuthorityName="L. W. Schaufuss" box="[421,732,282,307]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="267" phylum="Arthropoda" rank="species" species="asserculatus">
<emphasis id="B958EAB8C07A4C793DBCFEB2DE12FE94" box="[421,547,283,307]" italics="true" pageId="16" pageNumber="267">asserculatus</emphasis>
(L.W. Schaufuss)
</taxonomicName>
;
<treatmentCitation id="0A8D10BBC07A4C793EFFFEB2DFBFFE94" author="Franz, H." box="[742,910,282,307]" page="697" pageId="16" pageNumber="267" year="1967">
<bibRefCitation id="EFBD4B5BC07A4C793EFFFEB2DFBFFE94" author="Franz, H." box="[742,910,282,307]" pageId="16" pageNumber="267" pagination="611 - 724" refId="ref11321" refString="Franz, H. (1967) Zur Kenntnis der Scydmaenidenfauna von Lateinamerika. I Teil: Scydmaeniden aus Chile und dem sudlichen Argentinien. II Teil: Scydmaeniden aus dem subtropischen und tropischen Amerika. In: Delamare Deboutteville, C. &amp; Rappaport, E. (Eds.), Biologie de l'Amerique Australe. Vol. III. Etudes sur la faune du sol, documents biogeographiques. Centre National de la Recherche Scientifique, Paris, pp. 611 - 724." type="book chapter" year="1967">Franz 1967: 697</bibRefCitation>
</treatmentCitation>
, 715.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3366521C07A4C793C8EFE94DD16FEF0" box="[151,295,317,343]" pageId="16" pageNumber="267" type="description">
<paragraph id="8B9336AAC07A4C793C8EFE94DD16FEF0" blockId="16.[151,966,221,343]" box="[151,295,317,343]" pageId="16" pageNumber="267">
(
<figureCitation id="13172A2FC07A4C793C86FE94DD11FEF0" box="[159,288,317,343]" captionStart="FIGURES 4043" captionStartId="16.[151,264,1587,1612]" captionTargetBox="[206,1380,950,1555]" captionTargetId="figure@16.[201,1386,941,1561]" captionTargetPageId="16" captionText="FIGURES 4043. Venezolanoconnus asserculatus (L.W. Schaufuss) comb. n., syntype male. Dorsal habitus (40); original set of labels (41); aedeagus in ventral (42) and lateral (43) views." figureDoi="http://doi.org/10.5281/zenodo.3731605" httpUri="https://zenodo.org/record/3731605/files/figure.png" pageId="16" pageNumber="267">Figs 4043</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3366521C07A4C793C8EFE2FDD94FD88" pageId="16" pageNumber="267" type="materials_examined">
<paragraph id="8B9336AAC07A4C793C8EFE2FDD94FD88" blockId="16.[151,1437,389,920]" pageId="16" pageNumber="267">
<emphasis id="B958EAB8C07A4C793C8EFE2FDD78FE07" bold="true" box="[151,329,389,416]" pageId="16" pageNumber="267">Type material.</emphasis>
<materialsCitation id="3B443CF7C07A4C793D4DFE2CDD94FD88" ID-GBIF-Occurrence="2592198291" collectionCode="SDEI" collectorName="C. Schaufuss &amp; L. W. Schaufuss" country="Colombia" determinerName="P. Jaloszynski" location="Granada" municipality="Schf" pageId="16" pageNumber="267" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="syntype">
<emphasis id="B958EAB8C07A4C793D4DFE2CDE41FE07" bold="true" box="[340,624,389,416]" pageId="16" pageNumber="267">
<typeStatus id="54978808C07A4C793D4DFE2CDD88FE07" box="[340,441,389,416]" pageId="16" pageNumber="267" type="syntype">Syntype</typeStatus>
:
<collectingCountry id="F33B763AC07A4C793DD1FE2CDE5AFE07" box="[456,619,389,416]" name="Colombia" pageId="16" pageNumber="267">COLOMBIA</collectingCountry>
:
</emphasis>
<specimenCount id="9D2AFD23C07A4C793E62FE2FDEA3FE07" box="[635,658,390,416]" pageId="16" pageNumber="267" type="male"></specimenCount>
, five historical labels (
<figureCitation id="13172A2FC07A4C793FB9FE2CDFC9FE38" box="[928,1016,389,415]" captionStart="FIGURES 4043" captionStartId="16.[151,264,1587,1612]" captionTargetBox="[206,1380,950,1555]" captionTargetId="figure@16.[201,1386,941,1561]" captionTargetPageId="16" captionText="FIGURES 4043. Venezolanoconnus asserculatus (L.W. Schaufuss) comb. n., syntype male. Dorsal habitus (40); original set of labels (41); aedeagus in ventral (42) and lateral (43) views." figureDoi="http://doi.org/10.5281/zenodo.3731605" httpUri="https://zenodo.org/record/3731605/files/figure.png" pageId="16" pageNumber="267">Fig. 41</figureCitation>
): “asserculatus /
<collectingMunicipality id="6BF7ACD0C07A4C7938D8FE2CD8C5FE38" box="[1217,1268,389,415]" pageId="16" pageNumber="267">Schf</collectingMunicipality>
/ N
<location id="8EF36071C07A4C793928FE2CD9A2FE38" LSID="urn:lsid:plazi:treatment:038587BCC07A4C783C8EFF30D904FC24:8EF36071C07A4C793928FE2CD9A2FE38" box="[1329,1427,389,415]" country="Colombia" municipality="Schf" name="Granada" pageId="16" pageNumber="267">Granada</location>
” [originally white, now darkened], “asserculatus /
<location id="8EF36071C07A4C793ED4FE00DFEDFE64" LSID="urn:lsid:plazi:treatment:038587BCC07A4C783C8EFF30D904FC24:8EF36071C07A4C793ED4FE00DFEDFE64" box="[717,988,425,452]" country="Colombia" municipality="Schf" name="Schauf. / Nov. Granada" pageId="16" pageNumber="267">Schauf. / Nov. Granada</location>
” [originally white, now darkened], “
<collectorName id="26D9537CC07A4C79399BFE00DD38FE40" pageId="16" pageNumber="267">C. Schaufuss</collectorName>
1930” [white, printed], “
<typeStatus id="54978808C07A4C793E3FFE64DEA2FE40" box="[550,659,461,487]" pageId="16" pageNumber="267" type="syntype">Syntypus</typeStatus>
” [red, printed], “
<specimenCount id="9D2AFD23C07A4C793F41FE64DF5BFE40" box="[856,874,461,487]" pageId="16" pageNumber="267" type="female"></specimenCount>
” [white, handwritten]; during the present study a new label was added: “
<taxonomicName id="4C2C4D29C07A4C793D83FE5BDE8BFDAB" authorityName="Franz. Each" authorityYear="1987" box="[410,698,498,524]" class="Insecta" family="Staphylinidae" genus="Venezolanoconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="267" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAB8C07A4C793D83FE5BDE8BFDAB" box="[410,698,498,524]" italics="true" pageId="16" pageNumber="267">VENEZOLANOCONNUS</emphasis>
</taxonomicName>
/
<taxonomicName id="4C2C4D29C07A4C793ED6FE58D858FDAB" authority="(L. W. Schaufuss, 1866)" baseAuthorityName="L. W. Schaufuss" baseAuthorityYear="1866" box="[719,1129,497,524]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="267" phylum="Arthropoda" rank="species" species="asserculatus">
<emphasis id="B958EAB8C07A4C793ED6FE58DF6BFDAC" box="[719,858,497,523]" italics="true" pageId="16" pageNumber="267">asserculatus</emphasis>
(
<collectorName id="26D9537CC07A4C793F73FE58D82DFDAC" box="[874,1052,497,523]" pageId="16" pageNumber="267">
L.W.
<bibRefCitation id="EFBD4B5BC07A4C793FB1FE58D82DFDAC" author="Schaufuss, L. W." box="[936,1052,497,523]" pageId="16" pageNumber="267" pagination="1 - 103" refId="ref11922" refString="Schaufuss, L. W. (1866) Monographie der Scydmaeniden Central- und Sudamerika's. Novorum Actorum Academiae Caesareae Leopoldino-Carolinae Germanicae Naturae Curiosorum, 33 (6), 1 - 103, pls. 1 - 4. [1867]" type="journal article" year="1866">Schaufuss</bibRefCitation>
</collectorName>
, 1866)
</taxonomicName>
/ det.
<determinerName id="7E4FDA6EC07A4C7938B6FE5BD97CFDAC" box="[1199,1357,497,524]" pageId="16" pageNumber="267">P. Jałoszyński</determinerName>
, 2020” [white, printed] (
<collectionCode id="ED3DAE6FC07A4C793D4CFDBCDDA8FD88" box="[341,409,533,559]" pageId="16" pageNumber="267">SDEI</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3366521C07A4C793CDEFD90DF6CFDD0" pageId="16" pageNumber="267" type="diagnosis">
<paragraph id="8B9336AAC07A4C793CDEFD90DF6CFDD0" blockId="16.[151,1437,389,920]" pageId="16" pageNumber="267">
<emphasis id="B958EAB8C07A4C793CDEFD90DD73FDF3" bold="true" box="[199,322,569,596]" pageId="16" pageNumber="267">Diagnosis.</emphasis>
Apical region of median lobe in lateral view projecting distad and curved ventrad; median lobe with long and sparse setae, distributed along its laterodistal margins.
</paragraph>
</subSubSection>
<subSubSection id="C3366521C07A4C783CDEFD28DDAFFF5C" lastPageId="17" lastPageNumber="268" pageId="16" pageNumber="267" type="description">
<paragraph id="8B9336AAC07A4C793CDEFD28DD33FD67" blockId="16.[151,1437,389,920]" pageId="16" pageNumber="267">
<emphasis id="B958EAB8C07A4C793CDEFD28DD44FD3B" bold="true" box="[199,373,641,668]" pageId="16" pageNumber="267">Redescription.</emphasis>
Body of male (
<figureCitation id="13172A2FC07A4C793E38FD28DE43FD3C" box="[545,626,641,667]" captionStart="FIGURES 4043" captionStartId="16.[151,264,1587,1612]" captionTargetBox="[206,1380,950,1555]" captionTargetId="figure@16.[201,1386,941,1561]" captionTargetPageId="16" captionText="FIGURES 4043. Venezolanoconnus asserculatus (L.W. Schaufuss) comb. n., syntype male. Dorsal habitus (40); original set of labels (41); aedeagus in ventral (42) and lateral (43) views." figureDoi="http://doi.org/10.5281/zenodo.3731605" httpUri="https://zenodo.org/record/3731605/files/figure.png" pageId="16" pageNumber="267">Fig. 40</figureCitation>
) stout and strongly convex, brown, setae slightly lighter than cuticle; BL
<quantity id="4CD49B4FC07A4C793C8EFD0CDCCFFD67" box="[151,254,677,704]" metricMagnitude="-3" metricUnit="m" metricValue="1.58" pageId="16" pageNumber="267" unit="mm" value="1.58">1.58 mm</quantity>
.
</paragraph>
<paragraph id="8B9336AAC07A4C793CDEFD60DECCFC30" blockId="16.[151,1437,389,920]" pageId="16" pageNumber="267">
Head (
<figureCitation id="13172A2FC07A4C793D0AFD60DD54FD44" box="[275,357,713,739]" captionStart="FIGURES 4043" captionStartId="16.[151,264,1587,1612]" captionTargetBox="[206,1380,950,1555]" captionTargetId="figure@16.[201,1386,941,1561]" captionTargetPageId="16" captionText="FIGURES 4043. Venezolanoconnus asserculatus (L.W. Schaufuss) comb. n., syntype male. Dorsal habitus (40); original set of labels (41); aedeagus in ventral (42) and lateral (43) views." figureDoi="http://doi.org/10.5281/zenodo.3731605" httpUri="https://zenodo.org/record/3731605/files/figure.png" pageId="16" pageNumber="267">Fig. 40</figureCitation>
) broadest at large, strongly convex eyes, HL
<quantity id="4CD49B4FC07A4C793F72FD60DFE3FD43" box="[875,978,713,740]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="16" pageNumber="267" unit="mm" value="0.25">0.25 mm</quantity>
, HW
<quantity id="4CD49B4FC07A4C79380FFD60D84FFD43" box="[1046,1150,713,740]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="16" pageNumber="267" unit="mm" value="0.3">0.30 mm</quantity>
; frons and vertex confluent, weakly convex, together about as long as broad; supraantennal tubercles weakly elevated; tempora distinctly shorter than eye in dorsal view, strongly convergent posteromesad, rounded. Vertex and frons with fine, inconspicuous punctures; setae short, sparse, suberect. Antennae (
<figureCitation id="13172A2FC07A4C793F18FC9CDF63FCE8" box="[769,850,821,847]" captionStart="FIGURES 4043" captionStartId="16.[151,264,1587,1612]" captionTargetBox="[206,1380,950,1555]" captionTargetId="figure@16.[201,1386,941,1561]" captionTargetPageId="16" captionText="FIGURES 4043. Venezolanoconnus asserculatus (L.W. Schaufuss) comb. n., syntype male. Dorsal habitus (40); original set of labels (41); aedeagus in ventral (42) and lateral (43) views." figureDoi="http://doi.org/10.5281/zenodo.3731605" httpUri="https://zenodo.org/record/3731605/files/figure.png" pageId="16" pageNumber="267">Fig. 40</figureCitation>
) slender, AnL
<quantity id="4CD49B4FC07A4C793FE1FC9CD851FCF7" box="[1016,1120,821,848]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="16" pageNumber="267" unit="mm" value="0.68">0.68 mm</quantity>
; antennomeres I and II each elongate (II weakly so), IIIV each about as long as broad; VIX each distinctly transverse; XI indistinctly shorter than IX and X combined, about 1.3 × as long as broad.
</paragraph>
<caption id="DF536622C07A4C793C8EF99ADF38F9D7" ID-DOI="http://doi.org/10.5281/zenodo.3731605" ID-Zenodo-Dep="3731605" httpUri="https://zenodo.org/record/3731605/files/figure.png" pageId="16" pageNumber="267" startId="16.[151,264,1587,1612]" targetBox="[206,1380,950,1555]" targetPageId="16">
<paragraph id="8B9336AAC07A4C793C8EF99ADF38F9D7" blockId="16.[151,1436,1587,1648]" pageId="16" pageNumber="267">
<emphasis id="B958EAB8C07A4C793C8EF99ADD64F9EB" bold="true" box="[151,341,1587,1612]" pageId="16" pageNumber="267">FIGURES 4043.</emphasis>
<taxonomicName id="4C2C4D29C07A4C793D42F99ADF62F9EB" authority="(L. W. Schaufuss)" authorityName="Jałoszyński" authorityYear="2020" baseAuthorityName="L. W. Schaufuss" baseAuthorityYear="2020" box="[347,851,1587,1612]" class="Insecta" family="Staphylinidae" genus="Venezolanoconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="267" phylum="Arthropoda" rank="species" species="asserculatus" status="comb. nov.">
<emphasis id="B958EAB8C07A4C793D42F99ADEAAF9EC" box="[347,667,1587,1611]" italics="true" pageId="16" pageNumber="267">Venezolanoconnus asserculatus</emphasis>
(L.W. Schaufuss)
</taxonomicName>
<emphasis id="B958EAB8C07A4C793F40F99DDF87F9EB" bold="true" box="[857,950,1588,1612]" pageId="16" pageNumber="267">
<taxonomicNameLabel id="A26B57C3C07A4C793F40F99DDF87F9EB" box="[857,950,1588,1612]" pageId="16" pageNumber="267" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
, syntype male. Dorsal habitus (40); original set of labels (41); aedeagus in ventral (42) and lateral (43) views.
</paragraph>
</caption>
<paragraph id="8B9336AAC07A4C793CDEF937DF1FF883" blockId="16.[151,1437,1694,2044]" pageId="16" pageNumber="267">
Pronotum (
<figureCitation id="13172A2FC07A4C793D5FF937DDA8F91F" box="[326,409,1694,1720]" captionStart="FIGURES 4043" captionStartId="16.[151,264,1587,1612]" captionTargetBox="[206,1380,950,1555]" captionTargetId="figure@16.[201,1386,941,1561]" captionTargetPageId="16" captionText="FIGURES 4043. Venezolanoconnus asserculatus (L.W. Schaufuss) comb. n., syntype male. Dorsal habitus (40); original set of labels (41); aedeagus in ventral (42) and lateral (43) views." figureDoi="http://doi.org/10.5281/zenodo.3731605" httpUri="https://zenodo.org/record/3731605/files/figure.png" pageId="16" pageNumber="267">Fig. 40</figureCitation>
) broadest near posterior 1/4; PL
<quantity id="4CD49B4FC07A4C793F09F937DF49F91F" box="[784,888,1694,1720]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="16" pageNumber="267" unit="mm" value="0.4">0.40 mm</quantity>
, PW
<quantity id="4CD49B4FC07A4C793FAFF937D82FF91F" box="[950,1054,1694,1720]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="16" pageNumber="267" unit="mm" value="0.43">0.43 mm</quantity>
. All margins rounded; inner antebasal pits present but indistinct, small, shallow and diffuse, connected by shallow but distinct transverse groove. Punctures on pronotal disc fine, inconspicuous; setae on dorsal region short, sparse, suberect, on sides replaced by long erect bristles (most broken off in the studied
<typeStatus id="54978808C07A4C793EDFF8A3DF13F883" box="[710,802,1802,1828]" pageId="16" pageNumber="267" type="syntype">syntype</typeStatus>
).
</paragraph>
<paragraph id="8B9336AAC07A4C793CDEF887DEFCF837" blockId="16.[151,1437,1694,2044]" pageId="16" pageNumber="267">
Elytra (
<figureCitation id="13172A2FC07A4C793D03F887DD5AF8EF" box="[282,363,1838,1864]" captionStart="FIGURES 4043" captionStartId="16.[151,264,1587,1612]" captionTargetBox="[206,1380,950,1555]" captionTargetId="figure@16.[201,1386,941,1561]" captionTargetPageId="16" captionText="FIGURES 4043. Venezolanoconnus asserculatus (L.W. Schaufuss) comb. n., syntype male. Dorsal habitus (40); original set of labels (41); aedeagus in ventral (42) and lateral (43) views." figureDoi="http://doi.org/10.5281/zenodo.3731605" httpUri="https://zenodo.org/record/3731605/files/figure.png" pageId="16" pageNumber="267">Fig. 40</figureCitation>
) together oval, broadest distinctly behind middle; EL
<quantity id="4CD49B4FC07A4C793FDAF887D818F8EF" box="[963,1065,1838,1864]" metricMagnitude="-4" metricUnit="m" metricValue="9.3" pageId="16" pageNumber="267" unit="mm" value="0.93">0.93 mm</quantity>
, EW
<quantity id="4CD49B4FC07A4C79387CF887D8FBF8EF" box="[1125,1226,1838,1864]" metricMagnitude="-4" metricUnit="m" metricValue="7.8" pageId="16" pageNumber="267" unit="mm" value="0.78">0.78 mm</quantity>
, EI 1.19; basal impressions short and shallow, humeral calli distinct, elongate, elytral apices separately rounded. Punctures and setae similar to those on median region of pronotal disc.
</paragraph>
<paragraph id="8B9336AAC07A4C793CDEF833DDE2F813" blockId="16.[151,1437,1694,2044]" box="[199,467,1946,1972]" pageId="16" pageNumber="267">Hind wings not studied.</paragraph>
<paragraph id="8B9336AAC07A4C793CDEF817DEEBF87F" blockId="16.[151,1437,1694,2044]" box="[199,730,1982,2008]" pageId="16" pageNumber="267">Legs moderately long and slender; unmodified.</paragraph>
<paragraph id="8B9336AAC07A4C783CDEF84BDD68FF70" blockId="16.[151,1437,1694,2044]" lastBlockId="17.[151,1437,153,899]" lastPageId="17" lastPageNumber="268" pageId="16" pageNumber="267">
Aedeagus (
<figureCitation id="13172A2FC07A4C793D5AF84BDDF2F85B" box="[323,451,2018,2044]" captionStart="FIGURES 4043" captionStartId="16.[151,264,1587,1612]" captionTargetBox="[206,1380,950,1555]" captionTargetId="figure@16.[201,1386,941,1561]" captionTargetPageId="16" captionText="FIGURES 4043. Venezolanoconnus asserculatus (L.W. Schaufuss) comb. n., syntype male. Dorsal habitus (40); original set of labels (41); aedeagus in ventral (42) and lateral (43) views." figureDoi="http://doi.org/10.5281/zenodo.3731605" httpUri="https://zenodo.org/record/3731605/files/figure.png" pageId="16" pageNumber="267">Figs 4243</figureCitation>
) stout; AeL
<quantity id="4CD49B4FC07A4C793E50F84BDE9EF85B" box="[585,687,2018,2044]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="16" pageNumber="267" unit="mm" value="0.2">0.20 mm</quantity>
. Median lobe in ventral view broadest in subapical region, in lateral view with dorsal and ventral walls sinuate; endophallus with long C-shaped flagellum; each paramere with several long apical setae.
</paragraph>
<paragraph id="8B9336AAC07B4C783CDEFF48DDAFFF5C" blockId="17.[151,1437,153,899]" box="[199,414,225,251]" pageId="17" pageNumber="268">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C3366521C07B4C783CDEFEADD9B1FEB8" box="[199,1408,260,288]" pageId="17" pageNumber="268" type="distribution">
<paragraph id="8B9336AAC07B4C783CDEFEADD9B1FEB8" blockId="17.[151,1437,153,899]" box="[199,1408,260,288]" pageId="17" pageNumber="268">
<emphasis id="B958EAB8C07B4C783CDEFEADDD50FEB8" bold="true" box="[199,353,260,287]" pageId="17" pageNumber="268">Distribution.</emphasis>
Western
<collectingCountry id="F33B763AC07B4C783DD0FEACDE06FEB8" box="[457,567,261,287]" name="Colombia" pageId="17" pageNumber="268">Colombia</collectingCountry>
(“ad flumen Rio Madalena”); “
<collectingCountry id="F33B763AC07B4C783F80FEACDFD3FEB8" box="[921,994,261,287]" name="Brazil" pageId="17" pageNumber="268">Brazil</collectingCountry>
” in
<bibRefCitation id="EFBD4B5BC07B4C783817FEACD8DCFEB8" author="Asenjo, A. &amp; Irmler, U. &amp; Klimaszewski, J. &amp; Herman, L. K. &amp; Chandler, D. S." box="[1038,1261,261,287]" pageId="17" pageNumber="268" pagination="1 - 419" refId="ref11218" refString="Asenjo, A., Irmler, U., Klimaszewski, J., Herman, L. K. &amp; Chandler, D. S. (2013) A complete checklist with new records and geographical distribution of the rove beetles (Coleoptera, Staphylinidae) of Brazil. Insecta Mundi, 0277, 1 - 419." type="book chapter" year="2013">Asenjo et al. (2013)</bibRefCitation>
was in error.
</paragraph>
</subSubSection>
<subSubSection id="C3366521C07B4C783CDEFE80D904FC24" pageId="17" pageNumber="268" type="discussion">
<paragraph id="8B9336AAC07B4C783CDEFE80DEF3FE74" blockId="17.[151,1437,153,899]" pageId="17" pageNumber="268">
<emphasis id="B958EAB8C07B4C783CDEFE80DD0AFEE4" bold="true" box="[199,315,297,323]" pageId="17" pageNumber="268">Remarks.</emphasis>
Although
<bibRefCitation id="EFBD4B5BC07B4C783DA8FE80DE73FEE4" author="Franz, H." box="[433,578,297,323]" pageId="17" pageNumber="268" pagination="59 - 84" refId="ref11481" refString="Franz, H. (1988) Nachtrag zur monographie der Scydmaeniden Venezuelas (Coleoptera, Scydmaenidae). Elytron, 1, 59 - 84. [1987]" type="journal article" year="1988">Franz (1988)</bibRefCitation>
described
<taxonomicName id="4C2C4D29C07B4C783EA3FE80DFB8FEE4" authorityName="Franz. Each" authorityYear="1987" box="[698,905,297,323]" class="Insecta" family="Staphylinidae" genus="Venezolanoconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="268" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAB8C07B4C783EA3FE80DFB8FEE4" box="[698,905,297,323]" italics="true" pageId="17" pageNumber="268">Venezolanoconnus</emphasis>
</taxonomicName>
, he apparently did not recognize the previously (
<bibRefCitation id="EFBD4B5BC07B4C783C86FEE4DD10FECF" author="Franz, H." box="[159,289,333,360]" pageId="17" pageNumber="268" pagination="611 - 724" refId="ref11321" refString="Franz, H. (1967) Zur Kenntnis der Scydmaenidenfauna von Lateinamerika. I Teil: Scydmaeniden aus Chile und dem sudlichen Argentinien. II Teil: Scydmaeniden aus dem subtropischen und tropischen Amerika. In: Delamare Deboutteville, C. &amp; Rappaport, E. (Eds.), Biologie de l'Amerique Australe. Vol. III. Etudes sur la faune du sol, documents biogeographiques. Centre National de la Recherche Scientifique, Paris, pp. 611 - 724." type="book chapter" year="1967">Franz 1967</bibRefCitation>
) redescribed
<taxonomicName id="4C2C4D29C07B4C783DA3FEE4DEE6FEC0" authorityName="L. W. Schaufuss" authorityYear="1866" box="[442,727,333,359]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="268" phylum="Arthropoda" rank="species" species="asserculatus">
<emphasis id="B958EAB8C07B4C783DA3FEE4DEE6FEC0" box="[442,727,333,359]" italics="true" pageId="17" pageNumber="268">Scydmaenus asserculatus</emphasis>
</taxonomicName>
as a member of this genus. Moreover,
<bibRefCitation id="EFBD4B5BC07B4C783888FEE4D912FECF" author="Franz, H." box="[1169,1315,333,360]" pageId="17" pageNumber="268" pagination="611 - 724" refId="ref11321" refString="Franz, H. (1967) Zur Kenntnis der Scydmaenidenfauna von Lateinamerika. I Teil: Scydmaeniden aus Chile und dem sudlichen Argentinien. II Teil: Scydmaeniden aus dem subtropischen und tropischen Amerika. In: Delamare Deboutteville, C. &amp; Rappaport, E. (Eds.), Biologie de l'Amerique Australe. Vol. III. Etudes sur la faune du sol, documents biogeographiques. Centre National de la Recherche Scientifique, Paris, pp. 611 - 724." type="book chapter" year="1967">Franz (1967)</bibRefCitation>
misidentified the
<typeStatus id="54978808C07B4C783CEDFED8DD7AFE2C" box="[244,331,369,395]" pageId="17" pageNumber="268" type="syntype">syntype</typeStatus>
housed at SDEI as a female, whereas it is a male. Species of
<taxonomicName id="4C2C4D29C07B4C78381AFED8D8E3FE2C" authorityName="Franz. Each" authorityYear="1987" box="[1027,1234,369,395]" class="Insecta" family="Staphylinidae" genus="Venezolanoconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="268" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAB8C07B4C78381AFED8D8E3FE2C" box="[1027,1234,369,395]" italics="true" pageId="17" pageNumber="268">Venezolanoconnus</emphasis>
</taxonomicName>
have small, sacklike and lightly sclerotized aedeagi, and it is easy to overlook and even lose the male copulatory organ during a careless dissection; this can explain Franzs error.
</paragraph>
<paragraph id="8B9336AAC07B4C783CDEFE74D902FDC4" blockId="17.[151,1437,153,899]" pageId="17" pageNumber="268">
The species originally described as
<taxonomicName id="4C2C4D29C07B4C783E47FE74DF4DFE50" authorityName="L. W. Schaufuss" authorityYear="1866" box="[606,892,477,503]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="268" phylum="Arthropoda" rank="species" species="asserculatus">
<emphasis id="B958EAB8C07B4C783E47FE74DF4DFE50" box="[606,892,477,503]" italics="true" pageId="17" pageNumber="268">Scydmaenus asserculatus</emphasis>
</taxonomicName>
can be easily recognized as a member of
<emphasis id="B958EAB8C07B4C783947FE77DD00FDBC" italics="true" pageId="17" pageNumber="268">Vene- zolanoconnus</emphasis>
by its general body form. The unique structure of the aedeagus confirms this placement. Strongly modified parameres, almost surrounding the median lobe and at least in their basal half as broad as the median lobe, co-occurring with an asymmetrical, broad and curved flagellum are found only in this Neotropical genus.
</paragraph>
<paragraph id="8B9336AAC07B4C783CDEFDC4D904FC24" blockId="17.[151,1437,153,899]" pageId="17" pageNumber="268">
<taxonomicName id="4C2C4D29C07B4C783CDEFDC4DE19FD20" authorityName="Jałoszyński" authorityYear="2020" baseAuthorityName="L. W. Schaufuss" baseAuthorityYear="2020" box="[199,552,621,647]" class="Insecta" family="Staphylinidae" genus="Venezolanoconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="268" phylum="Arthropoda" rank="species" species="asserculatus">
<emphasis id="B958EAB8C07B4C783CDEFDC4DE19FD20" box="[199,552,621,647]" italics="true" pageId="17" pageNumber="268">Venezolanoconnus asserculatus</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4C2C4D29C07B4C783EECFDC7DFC5FD20" authority="Franz, 1988" authorityName="Franz" authorityYear="1988" box="[757,1012,621,647]" class="Insecta" family="Staphylinidae" genus="Venezolanoconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="268" phylum="Arthropoda" rank="species" species="andinus">
<emphasis id="B958EAB8C07B4C783EECFDC7DF58FD20" box="[757,873,621,647]" italics="true" pageId="17" pageNumber="268">V. andinus</emphasis>
<bibRefCitation id="EFBD4B5BC07B4C783F76FDC4DFC5FD20" author="Franz, H." box="[879,1012,621,647]" pageId="17" pageNumber="268" pagination="59 - 84" refId="ref11481" refString="Franz, H. (1988) Nachtrag zur monographie der Scydmaeniden Venezuelas (Coleoptera, Scydmaenidae). Elytron, 1, 59 - 84. [1987]" type="journal article" year="1988">Franz, 1988</bibRefCitation>
</taxonomicName>
of western
<collectingCountry id="F33B763AC07B4C78386FFDC4D8D8FD20" box="[1142,1257,621,647]" name="Venezuela" pageId="17" pageNumber="268">Venezuela</collectingCountry>
(redescribed by
<bibRefCitation id="EFBD4B5BC07B4C783C8EFD38DD6CFD0C" author="Jaloszynski, P." box="[151,349,657,683]" pageId="17" pageNumber="268" pagination="55 - 82" refId="ref11596" refString="Jaloszynski, P. (2012) Taxonomy of ' Euconnus complex'. Part I. Morphology of Euconnus s. str. and revision of Euconnomorphus Franz and Venezolanoconnus Franz (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 3555 (1), 55 - 82. https: // doi. org / 10.11646 / zootaxa. 3555.1.3" type="journal article" year="2012">Jałoszyński 2012</bibRefCitation>
); especially their aedeagi have a similar shape and structure. However, these species clearly dif- fer in pronotal structures: in
<taxonomicName id="4C2C4D29C07B4C783DC0FD1FDE7FFD68" authorityName="Franz" authorityYear="1988" box="[473,590,693,719]" class="Insecta" family="Staphylinidae" genus="Venezolanoconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="268" phylum="Arthropoda" rank="species" species="andinus">
<emphasis id="B958EAB8C07B4C783DC0FD1FDE7FFD68" box="[473,590,693,719]" italics="true" pageId="17" pageNumber="268">V. andinus</emphasis>
</taxonomicName>
the pronotum lacks antebasal pits and groove, whereas in
<taxonomicName id="4C2C4D29C07B4C7838F8FD1FD9B9FD68" authorityName="Jałoszyński" authorityYear="2020" baseAuthorityName="L. W. Schaufuss" baseAuthorityYear="2020" box="[1249,1416,693,719]" class="Insecta" family="Staphylinidae" genus="Venezolanoconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="268" phylum="Arthropoda" rank="species" species="asserculatus">
<emphasis id="B958EAB8C07B4C7838F8FD1FD9B9FD68" box="[1249,1416,693,719]" italics="true" pageId="17" pageNumber="268">V. asserculatus</emphasis>
</taxonomicName>
a shallow but well visible transverse groove is present, connecting a pair of indistinct pits. Moreover, in the aedeagus of
<taxonomicName id="4C2C4D29C07B4C783CACFD57DD18FCB0" authorityName="Franz" authorityYear="1988" box="[181,297,765,791]" class="Insecta" family="Staphylinidae" genus="Venezolanoconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="268" phylum="Arthropoda" rank="species" species="andinus">
<emphasis id="B958EAB8C07B4C783CACFD57DD18FCB0" box="[181,297,765,791]" italics="true" pageId="17" pageNumber="268">V. andinus</emphasis>
</taxonomicName>
, there are numerous (~ 15) short setae distributed from middle to distal margin of the median lobe, not visible or poorly visible in the ventral view. In
<taxonomicName id="4C2C4D29C07B4C783EB6FC8BDF67FC9C" authorityName="Jałoszyński" authorityYear="2020" baseAuthorityName="L. W. Schaufuss" baseAuthorityYear="2020" box="[687,854,801,827]" class="Insecta" family="Staphylinidae" genus="Venezolanoconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="268" phylum="Arthropoda" rank="species" species="asserculatus">
<emphasis id="B958EAB8C07B4C783EB6FC8BDF67FC9C" box="[687,854,801,827]" italics="true" pageId="17" pageNumber="268">V. asserculatus</emphasis>
</taxonomicName>
, the aedeagus bears only a few conspicuously long setae distributed along the distal margin of the median lobe, and most of them are well-visible in the ventral view. The apical region of the median lobe in the lateral view has also a clearly different shape in these species.
</paragraph>
</subSubSection>
</treatment>
</document>