177 lines
24 KiB
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177 lines
24 KiB
XML
<document ID-CLB-Dataset="177622" ID-DOI="http://dx.doi.org/10.3897/fr.25.83781" ID-GBIF-Dataset="879a081e-5c23-41a3-b500-84ae793fb677" ID-Pensoft-Pub="2193-0074-1-99" ID-Pensoft-UUID="2F5D6AE2EEB55A17ACF1623B06B4EA8D" ID-ZooBank="7A16698D4F1848D29D9651A6E0CC15AC" ModsDocID="2193-0074-25-1-99" checkinTime="1667366289781" checkinUser="pensoft" docAuthor="Vasilyan, Davit, Cernansky, Andrej, Szyndlar, Zbigniew & Moers, Thomas" docDate="2022" docId="355978966049518B8F26EF3AEE1387DA" docLanguage="en" docName="FossilRecord 25(1): 99-145" docOrigin="Fossil Record 25 (1)" docPubDate="2022-05-10" docSource="http://dx.doi.org/10.3897/fr.25.83781" docTitle="Chalcides Chalcides sp." docType="treatment" docVersion="5" id="2F5D6AE2EEB55A17ACF1623B06B4EA8D" lastPageNumber="99" masterDocId="2F5D6AE2EEB55A17ACF1623B06B4EA8D" masterDocTitle="Amphibian and reptilian fauna from the early Miocene of Echzell, Germany" masterLastPageNumber="145" masterPageNumber="99" pageNumber="99" updateTime="1732674429517" updateUser="ExternalLinkService">
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<mods:titleInfo id="CBB0082256483EF2DFFB19BC046006B2">
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<mods:title id="A29F4D3DB013B931B16AE361598596D4">Amphibian and reptilian fauna from the early Miocene of Echzell, Germany</mods:title>
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<mods:name id="57F2DFE86C9AE48F696FAE07B822330D" type="personal">
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<mods:namePart id="3651E154453E94E9FF7010570C873162">Vasilyan, Davit</mods:namePart>
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<mods:nameIdentifier id="C10C9B36C01AF5BE05849123E4E75172" type="ORCID">https://orcid.org/0000-0001-8712-0678</mods:nameIdentifier>
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<mods:affiliation id="AFF7A223342A08BE96523F183EA11809">JURASSICA Museum, Route de Fontenais 21. 2900 Porrentruy, Switzerland & Department of Geosciences, University of Fribourg, Chemin du musee 6, 1700, Fribourg, Switzerland</mods:affiliation>
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<mods:nameIdentifier id="A78615B813166F785577D9355D862B9D" type="email">davit.vasilyan@jurassica.ch</mods:nameIdentifier>
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<mods:namePart id="AD85097E9EB8947FC985F1C095E17AE5">Cernansky, Andrej</mods:namePart>
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<mods:nameIdentifier id="DC04FC1E10459AFAA9769318E4D16FB2" type="ORCID">https://orcid.org/0000-0001-8920-2503</mods:nameIdentifier>
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<mods:affiliation id="8031687C6C4BB2E4D287A30FADC92A99">Department of Ecology, Laboratory of Evolutionary Biology, Faculty of Natural Sciences, Comenius University in Bratislava, Mlynska dolina, Ilkovicova 6, 842 15 Bratislava, Slovakia</mods:affiliation>
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</mods:name>
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<mods:name id="A22C12EE5B09D6FC92F0E666271EA4F1" type="personal">
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<mods:roleTerm id="9B0072342492D385584BA4F8486967C6">Author</mods:roleTerm>
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<mods:namePart id="2E5D8DFC05667CB912B3673998C5BFF3">Szyndlar, Zbigniew</mods:namePart>
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<mods:affiliation id="B15B72A4495A7D6E669A058827D4AF2B">Institute of Systematics and Evolution of Animals, Polish Academy of Sciences, Slawkowska 17, 31 - 016 Krakow, Poland</mods:affiliation>
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<mods:namePart id="C7CC28914FA972ACDB17F4188A84CAB3">Moers, Thomas</mods:namePart>
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<mods:nameIdentifier id="188C2EC27F83DDCE5B5EDC011D1A27E2" type="ORCID">https://orcid.org/0000-0003-2268-5824</mods:nameIdentifier>
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<mods:affiliation id="812F14F9FE3F6E5CEA8C5E6259680426">Department of Palaeobiology, Swedish Museum of Natural History, P. O. Box 50007, 104 05 Stockholm, Sweden & Bolin Centre for Climate Research, Stockholm University, Stockholm, Sweden</mods:affiliation>
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<mods:typeOfResource id="512F2F38A420262A32955F8EFFD37DDE">text</mods:typeOfResource>
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<mods:title id="2A67AE405205475DDB5CB6E664A96C14">Fossil Record</mods:title>
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<mods:part id="5F0584AC2EFC491AA1901BA48E1BA97B">
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<mods:date id="C3B4A4B5C86FC01C2140C2E4281F2A69">2022</mods:date>
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<mods:number id="FCC44442293AAE18260F2518837EB31D">2022-05-10</mods:number>
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<mods:number id="335CB37155F778B2FE397F786D1318A7">25</mods:number>
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<mods:number id="15F392B539555F76749DF5AA57FAACD0">1</mods:number>
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<mods:start id="16191DD09836E5E36D529ACFE3E7DD96">99</mods:start>
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<mods:classification id="641821691B3E18047AF92E6C63E3C08A">journal article</mods:classification>
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<mods:identifier id="A3BE42691A30327FF75E0A833170D438" type="DOI">http://dx.doi.org/10.3897/fr.25.83781</mods:identifier>
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<treatment id="355978966049518B8F26EF3AEE1387DA" ID-GBIF-Taxon="203870949" LSID="urn:lsid:plazi:treatment:355978966049518B8F26EF3AEE1387DA" httpUri="http://treatment.plazi.org/id/355978966049518B8F26EF3AEE1387DA" lastPageNumber="99" pageId="0" pageNumber="99" scope_family="Scincidae" scope_order="Squamata" scope_subClass="Lissamphibia" scope_subOrder="Iguania">
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<subSubSection id="21429052C82B3EA12E5696591057F7C0" pageId="0" pageNumber="99" type="nomenclature">
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<paragraph id="AEEE69592535BD3262CF05D5E0DE4EC0" pageId="0" pageNumber="99">
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cf.
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<taxonomicName id="5A9322DD2B8CCE6F3F439DD382142DDB" LSID="35597896-6049-518B-8F26-EF3AEE1387DA" authority="Chalcides sp." authorityName="sp." class="Reptilia" family="Scincidae" genus="Chalcides" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Chalcides Chalcides" order="Squamata" pageId="0" pageNumber="99" rank="species" species="Chalcides">Chalcides Chalcides sp.</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="0723E46821F58BA0D17562DAB4FB087A" pageId="0" pageNumber="99" type="description">
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<paragraph id="87F3BC3EEDC19DB74DD65E109DACB361" pageId="0" pageNumber="99">
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<figureCitation id="EF83E6DCF68C0160A7FE39D57876730B" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Lacertidae indet. (A, B) and cf. Chalcides sp. (C-H) from the Echzell locality. Right dentary HLMD-Ez 1992 in (A) lateral and (B) medial views with a detail of dentition. Right maxilla HLMD-Ez 1990 in (C) lateral and (D) medial views with a detail of dentition. Right dentary HLMD-Ez 1994 in (E) lateral and (F) medial views with a detail of dentition. Right dentary HLMD-Ez 1993 in (G) lateral, and (H) medial views." figureDoi="10.3897/fr.25.83781.figure12" httpUri="https://binary.pensoft.net/fig/682232" pageId="0" pageNumber="99">Fig. 12C-H</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="2BC453E0485538EE07E567173CCDCA70" pageId="0" pageNumber="99" type="materials_examined">
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<paragraph id="C366E471C3C01398FFAB71932AEDBB45" pageId="0" pageNumber="99">Material.</paragraph>
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<paragraph id="87BB5A95E37776EFDDD49279151225CF" pageId="0" pageNumber="99">Right maxilla HLMD-Ez 1990, six right dentaries HLMD-Ez 1993-1998, one left dentary HLMD-Ez 1999.</paragraph>
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</subSubSection>
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<subSubSection id="9FDF24EBE9B9F77966AA5A49249096AF" pageId="0" pageNumber="99" type="description">
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<paragraph id="F1F437E3D0C63D07A7C868847D93585A" pageId="0" pageNumber="99">Description.</paragraph>
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<paragraph id="D1A0985995AC7D540087823AD0494BC3" pageId="0" pageNumber="99">
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Maxilla: One fragment of a right maxilla is preserved. The specimen HLMD-Ez 1990 (Fig.
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<figureCitation id="8C281E5B8B2786F5EACED30A7D4767C1" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Lacertidae indet. (A, B) and cf. Chalcides sp. (C-H) from the Echzell locality. Right dentary HLMD-Ez 1992 in (A) lateral and (B) medial views with a detail of dentition. Right maxilla HLMD-Ez 1990 in (C) lateral and (D) medial views with a detail of dentition. Right dentary HLMD-Ez 1994 in (E) lateral and (F) medial views with a detail of dentition. Right dentary HLMD-Ez 1993 in (G) lateral, and (H) medial views." figureDoi="10.3897/fr.25.83781.figure12" httpUri="https://binary.pensoft.net/fig/682232" pageId="0" pageNumber="99">12C, D</figureCitation>
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) represents the posterior region of the maxilla. The lateral surface of this specimen is completely smooth. This maxillary fragment bears eight tooth positions with five teeth still attached). The nasal process is partly preserved. Note, however, that the dorsoventral height of the posteriorly located posteroventral process of the maxilla is still significant. Thus, it forms the wall along the entire length of the process here rather than narrowing posteriorly into a tip. Moreover, the shallow notch is developed posteriorly between the dental crest supporting teeth and the dorsally located wall. Although the posterior portion of this dorsal wall is broken off, it can be estimated that, when completely preserved, it exceeds the dental part posteriorly. The dorsal margin of the posteroventral process is slightly damaged, but this portion has a thicker appearance than the ventrally located region possessing a longitudinal depression. This depression partly forms a jugal facet. The ventrally located supradental shelf is thin and expands laterally. It is dorsally convex, but only a small portion is completely preserved. No alveolar superior foramen is preserved here. This highlights a possible anterior position of this foramen.
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</paragraph>
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<paragraph id="62C32A6FB4675BD14BF44B5529235813" pageId="0" pageNumber="99">
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Dentary: The description is based on several fragments (Fig.
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<figureCitation id="43E1629E81ED4791999EDDB359853612" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Lacertidae indet. (A, B) and cf. Chalcides sp. (C-H) from the Echzell locality. Right dentary HLMD-Ez 1992 in (A) lateral and (B) medial views with a detail of dentition. Right maxilla HLMD-Ez 1990 in (C) lateral and (D) medial views with a detail of dentition. Right dentary HLMD-Ez 1994 in (E) lateral and (F) medial views with a detail of dentition. Right dentary HLMD-Ez 1993 in (G) lateral, and (H) medial views." figureDoi="10.3897/fr.25.83781.figure12" httpUri="https://binary.pensoft.net/fig/682232" pageId="0" pageNumber="99">12E-H</figureCitation>
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), most of which represent more-or-less anterior sections. The most complete specimen bears 20 tooth positions; however, its posterior region is broken off. The real tooth number is undoubtedly slightly higher. The dentary is slender, anteroposteriorly elongated. The bone gradually narrows anteriorly. In dorsal view, its anterior portion has a small medial curvature. The otherwise smooth lateral surface is pierced by several labial foramina located in the mid-portion of the bone. In medial view, the
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<normalizedToken id="04A9E0D16FF2C20B5C84238EFBEEF1DD" originalValue="Meckel`s">Meckel's</normalizedToken>
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groove is narrow, but entirely open. The subdental shelf is medially protruded and robust, especially in the anterior region. It narrows posteriorly because of the presence of the splenial articulation facet. The symphysis is small, rectangular and somewhat narrow.
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</paragraph>
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<paragraph id="1C941001ACD5B3D3EAD4D4F3FC34BE7E" pageId="0" pageNumber="99">Dentition: The tooth implantation is pleurodont. The teeth are conical and high. They are closely spaced with small interdental gaps. The tooth crowns are mediolaterally compressed. Thus, the necks have a slightly lingually enlarged appearance. The tooth crowns have blunt apices. In medial view, they have a labial and lingual cusp. The lingual side, bordered by the culmen lateralis posterior and anterior, has striation formed by apicobasal ridges. They are more-or-less parallel to each other and their number usually varies from around five to eight. The labial aspect of the teeth appears smooth. Resorption pits pierce the tooth bases of some teeth.</paragraph>
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</subSubSection>
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<subSubSection id="EB4D60B85735B90E21C09B3186E1B8E9" pageId="0" pageNumber="99" type="remarks">
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<paragraph id="D5A1A56E0A6E9067B6BF11C8B4AA7EAE" pageId="0" pageNumber="99">Remarks.</paragraph>
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<paragraph id="62CC9CE1A69F97E651BE092FC4A8B080" pageId="0" pageNumber="99">
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All scincid elements here are assigned to one species based on the significant similarity in the dentition. Moreover, all elements are from the same locality and comparable in size. Besides tooth morphology (see Kosma 2004), the Echzell skink material can be allocated to the clade
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<taxonomicName id="A5110303D8818FD64B157F0A82F732F1" class="Reptilia" family="Scincidae" kingdom="Animalia" lsidName="" order="Squamata" pageId="0" pageNumber="99" phylum="Chordata" rank="family">Scincidae</taxonomicName>
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(sensu
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<bibRefCitation id="D439AC50AF6822BD6D8931CAF0281164" DOI="https://doi.org/10.11646/zootaxa.3765.4.2" author="Hedges, SB" journalOrPublisher="Zootaxa" pageId="0" pageNumber="99" pagination="317 - 338" refId="B59" refString="Hedges, SB, 2014. The high-level classification of skinks (Reptilia, Squamata, Scincomorpha). Zootaxa 3765: 317 - 338, DOI: https://doi.org/10.11646/zootaxa.3765.4.2" title="The high-level classification of skinks (Reptilia, Squamata, Scincomorpha)." url="https://doi.org/10.11646/zootaxa.3765.4.2" volume="3765" year="2014">Hedges 2014</bibRefCitation>
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;
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<taxonomicName id="015D7603AD1C8E09A482BAA77BC0E502" authorityName="Gray" authorityYear="1825" lsidName="" pageId="0" pageNumber="99" rank="subFamily" subFamily="Scincinae">Scincinae</taxonomicName>
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sensu
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<bibRefCitation id="C5186FAF4128537DB30DE576C9B30883" author="Estes, R" editor="Estes, R" journalOrPublisher="Stanford University Press, Stanford, California" pageId="0" pageNumber="99" pagination="119 - 281" refId="B38" refString="Estes, R, Queiroz, K, Gauthier, J, 1988. Phylogenetic relationships within Squamata. In: Estes, R, Pregill, G, Eds., Phylogenetic relationships of the lizard families. Stanford University Press, Stanford, California: 119 - 281" title="Phylogenetic relationships within Squamata." volumeTitle="Phylogenetic relationships of the lizard families." year="1988">Estes et al. 1988</bibRefCitation>
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) based on the narrow but fully open Meckelian groove in the dentary. This is present in members of the
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<taxonomicName id="576949490ECA1E51585B46E7B764169E" class="Reptilia" family="Scincidae" kingdom="Animalia" lsidName="" order="Squamata" pageId="0" pageNumber="99" phylum="Chordata" rank="family">Scincidae</taxonomicName>
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clade and in, e.g.,
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<taxonomicName id="6A0C9640DA0CA5D18241E4E0F2D1A2F8" authorityName="Gray" authorityYear="1845" class="Reptilia" family="Scincidae" genus="Ateuchosaurus" kingdom="Animalia" lsidName="Ateuchosaurus" order="Squamata" pageId="0" pageNumber="99" phylum="Chordata" rank="genus">
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<emphasis id="D322A3BB8E7C350B4C2ADDBA0A7BB1C1" italics="true" pageId="0" pageNumber="99">Ateuchosaurus</emphasis>
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</taxonomicName>
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, whereas it is closed in members of
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<taxonomicName id="9A9397D800FF627CF170F8B28C993152" authorityName="Gray" authorityYear="1845" class="Reptilia" family="Acontiidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Squamata" pageId="0" pageNumber="99" phylum="Chordata" rank="family">Acontiidae</taxonomicName>
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,
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<taxonomicName id="0E7766ABD41227AA13887F0B0C22615F" family="Sphenomorphidae" lsidName="" pageId="0" pageNumber="99" rank="family">Sphenomorphidae</taxonomicName>
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,
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<taxonomicName id="C63BB4CC2686C2B07100D0F48B24D375" family="Eugongylidae" lsidName="" pageId="0" pageNumber="99" rank="family">Eugongylidae</taxonomicName>
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,
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<taxonomicName id="F6326FE340557208BD8A045EB9518B3D" family="Lygosomidae" lsidName="" pageId="0" pageNumber="99" rank="family">Lygosomidae</taxonomicName>
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,
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<taxonomicName id="D6B6C7788FE294F32763861EA93DA00D" family="Egerniidae" lsidName="" pageId="0" pageNumber="99" rank="family">Egerniidae</taxonomicName>
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and
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<taxonomicName id="382FA5BE4EF9D4BB62AF598EE2ECEC0A" authorityName="Mittleman" authorityYear="1952" class="Reptilia" family="Mabuyidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Squamata" pageId="0" pageNumber="99" phylum="Chordata" rank="family">Mabuyidae</taxonomicName>
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(see, e.g.,
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<bibRefCitation id="53CB1FC3BC40548079D1D2ED6CF9172B" author="Greer, AE" journalOrPublisher="Bulletin of the Museum of Comparative Zoology" pageId="0" pageNumber="99" pagination="151 - 183" refId="B55" refString="Greer, AE, 1970. A subfamilial classification of scincid lizards. Bulletin of the Museum of Comparative Zoology 139 (3): 151 - 183" title="A subfamilial classification of scincid lizards." volume="139" year="1970">Greer 1970</bibRefCitation>
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,
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<bibRefCitation id="339C110ADE30D525667D264C2201937B" DOI="https://doi.org/10.1071/AJZS031" author="Greer, AE" journalOrPublisher="Australian Journal of Zoology Supplementary Series" pageId="0" pageNumber="99" pagination="1 - 67" refId="B56" refString="Greer, AE, 1974. The genetic relationships of the Scincid lizard genus Leiolopisma and its relatives. Australian Journal of Zoology Supplementary Series 22 (31): 1 - 67, DOI: https://doi.org/10.1071/AJZS031" title="The genetic relationships of the Scincid lizard genus Leiolopisma and its relatives." url="https://doi.org/10.1071/AJZS031" volume="22" year="1974">1974</bibRefCitation>
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;
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<bibRefCitation id="920DA220CA0AB49CABCFE16DBD6D9B06" author="Evans, SE" editor="Gans, C" journalOrPublisher="Society for the Study of Amphibians and Reptiles, Ithaca" pageId="0" pageNumber="99" pagination="1 - 347" refId="B39" refString="Evans, SE, 2008. The skull of lizards and tuatara. In: Gans, C, Ed., The skull of Lepidosauria. Society for the Study of Amphibians and Reptiles, Ithaca: 1 - 347" title="The skull of lizards and tuatara." volumeTitle="The skull of Lepidosauria." year="2008">Evans 2008</bibRefCitation>
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;
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<bibRefCitation id="C6745CD713796AD6F73CB57F4CFE6CE8" DOI="https://doi.org/10.1670/08-126R.1" author="Hutchinson, MN" journalOrPublisher="Journal of Herpetology" pageId="0" pageNumber="99" pagination="139 - 147" refId="B65" refString="Hutchinson, MN, Scanlon, JD, 2009. New and Unusual Plio-Pleistocene Lizard (Reptilia: Scincidae) from Wellington Caves, New South Wales, Australia. Journal of Herpetology 43: 139 - 147, DOI: https://doi.org/10.1670/08-126R.1" title="New and Unusual Plio-Pleistocene Lizard (Reptilia: Scincidae) from Wellington Caves, New South Wales, Australia." url="https://doi.org/10.1670/08-126R.1" volume="43" year="2009">Hutchinson and Scanlon 2009</bibRefCitation>
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;
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<bibRefCitation id="4014DF156387F69CE1B03DFAC3349C78" DOI="https://doi.org/10.1002/ar.24289" author="Cernansky, A" journalOrPublisher="The Anatomical Record" pageId="0" pageNumber="99" pagination="1901 - 1934" refId="B29" refString="Cernansky, A, Syromyatnikova, EV, Kovalenko, ES, Podurets, KM, Kaloyan, AA, 2020b. The Key to Understanding the European Miocene Chalcides (Squamata, Scincidae) Comes from Asia: The Lizards of the East Siberian Tagay Locality (Baikal Lake) in Russia. The Anatomical Record 303: 1901 - 1934, DOI: https://doi.org/10.1002/ar.24289" title="The Key to Understanding the European Miocene Chalcides (Squamata, Scincidae) Comes from Asia: The Lizards of the East Siberian Tagay Locality (Baikal Lake) in Russia." url="https://doi.org/10.1002/ar.24289" volume="303" year="2020 b">
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<normalizedToken id="026DCDAC7C6B25265FFC59D49904A794" originalValue="Čerňanský">Cernansky</normalizedToken>
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et al. 2020b
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</bibRefCitation>
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;
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<bibRefCitation id="A7025EB68432F0AD0646489B21CB4AA1" DOI="https://doi.org/10.1002/spp2.1279" author="Cernansky, A" journalOrPublisher="Papers in Palaeontology" pageId="0" pageNumber="99" pagination="195 - 215" refId="B24" refString="Cernansky, A, Syromyatnikova, EV, 2021. The first pre-Quaternary fossil record of the clade Mabuyidae with a comment on the enclosure of the Meckelian canal in skinks. Papers in Palaeontology 7: 195 - 215, DOI: https://doi.org/10.1002/spp2.1279" title="The first pre-Quaternary fossil record of the clade Mabuyidae with a comment on the enclosure of the Meckelian canal in skinks." url="https://doi.org/10.1002/spp2.1279" volume="7" year="2021">
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<normalizedToken id="A78A41770908C74E901EE7B02FD41610" originalValue="Čerňanský">Cernansky</normalizedToken>
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and Syromyatnikova 2021
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</bibRefCitation>
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).
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</paragraph>
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<paragraph id="6CAAC320A9628ECA83E713B17936B819" pageId="0" pageNumber="99">
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The specimens resemble members of the genus
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<taxonomicName id="6617F501C6BB304CB8BFF70CB4171E8E" class="Reptilia" family="Scincidae" genus="Chalcides" kingdom="Animalia" lsidName="Chalcides" order="Squamata" pageId="0" pageNumber="99" rank="genus">
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<emphasis id="9590539D580FE2F37A800A8A0A1FC65D" italics="true" pageId="0" pageNumber="99">Chalcides</emphasis>
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</taxonomicName>
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. The Miocene species of this taxon is represented by
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<taxonomicName id="75A31B1651108697CAAB31BE3AE1F491" class="Reptilia" family="Scincidae" genus="Chalcides" kingdom="Animalia" lsidName="Chalcides augei" order="Squamata" pageId="0" pageNumber="99" rank="species" species="augei">
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<emphasis id="91B5A6373F8920DD54C840C21B979DA6" italics="true" pageId="0" pageNumber="99">Chalcides augei</emphasis>
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</taxonomicName>
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(see
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<bibRefCitation id="123E80697F97362684C7027F00CC076B" DOI="https://doi.org/10.1002/ar.24289" author="Cernansky, A" journalOrPublisher="The Anatomical Record" pageId="0" pageNumber="99" pagination="1901 - 1934" refId="B29" refString="Cernansky, A, Syromyatnikova, EV, Kovalenko, ES, Podurets, KM, Kaloyan, AA, 2020b. The Key to Understanding the European Miocene Chalcides (Squamata, Scincidae) Comes from Asia: The Lizards of the East Siberian Tagay Locality (Baikal Lake) in Russia. The Anatomical Record 303: 1901 - 1934, DOI: https://doi.org/10.1002/ar.24289" title="The Key to Understanding the European Miocene Chalcides (Squamata, Scincidae) Comes from Asia: The Lizards of the East Siberian Tagay Locality (Baikal Lake) in Russia." url="https://doi.org/10.1002/ar.24289" volume="303" year="2020 b">
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<normalizedToken id="D8ABEDB82E37D15C1862C4DD84E94E65" originalValue="Čerňanský">Cernansky</normalizedToken>
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et al. 2020b
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</bibRefCitation>
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) and the Echzell material shows similarities to this taxon. Note that usually five striations are reported in
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<taxonomicName id="5648A0C9C3DAE294CD630AE3A8FEE950" class="Reptilia" kingdom="Animalia" lsidName="Ch. augei" order="Squamata" pageId="0" pageNumber="99" phylum="Chordata" rank="species" species="augei">
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<emphasis id="960C8EC3C5C2425B7A26FB2D728CD5CF" italics="true" pageId="0" pageNumber="99">Ch. augei</emphasis>
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</taxonomicName>
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, it can range from five to seven, and the small variation in the number of striae (from five to seven ot eight) can be seen in specimens from Austria as well (
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<bibRefCitation id="D5AADDAE037D081B536BAFAA4680B9D4" DOI="https://doi.org/10.1007/s12542-016-0329-1" author="Cernansky, A" journalOrPublisher="Palaeontologische Zeitschrift" pageId="0" pageNumber="99" pagination="723 - 746" refId="B20" refString="Cernansky, A, 2016. Another piece of the puzzle: The first report on the Early Miocene lizard fauna from Austria (Ottnangian, MN 4; Oberdorf locality). Palaeontologische Zeitschrift 90: 723 - 746, DOI: https://doi.org/10.1007/s12542-016-0329-1" title="Another piece of the puzzle: The first report on the Early Miocene lizard fauna from Austria (Ottnangian, MN 4; Oberdorf locality)." url="https://doi.org/10.1007/s12542-016-0329-1" volume="90" year="2016">
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<normalizedToken id="1AB91B0BB39C262AB8832F19DD98C057" originalValue="Čerňanský">Cernansky</normalizedToken>
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2016
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</bibRefCitation>
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). However, the limitation of the Echzell material does not allow alpha taxonomy and caution is needed here.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |