620 lines
66 KiB
XML
620 lines
66 KiB
XML
<document id="A48D2F254606039812ACCEBCEA77A973" ID-DOI="10.5281/zenodo.198798" ID-GBIF-Dataset="7e6ef230-5a66-4dd0-b41e-411da4adee7b" ID-ISSN="1175-5326" ID-Zenodo-Dep="198798" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460314459685" checkinUser="plazi" docAuthor="Stuart, Bryan L., Phimmachak, Somphouthone, Sivongxay, Niane & Robichaud, William G." docDate="2010" docId="4A38FA2DFFB5FFCDE5F4B8BB321FF86B" docLanguage="en" docName="zt02650p032.pdf" docOrigin="Zootaxa 2650" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tylototriton notialis Stuart, Phimmachak, Sivongxay & Robichaud, 2010, sp. nov." docType="treatment" docVersion="7" lastPageNumber="27" masterDocId="B6018255FFB1FFC5E563BD4C3104FF99" masterDocTitle="A new species in the Tylototriton asperrimus group (Caudata: Salamandridae) from central Laos" masterLastPageNumber="32" masterPageNumber="19" pageNumber="23" updateTime="1698254641828" updateUser="plazi">
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<mods:mods id="EA955BA3DA458FA2F31356452379126B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="7CAD83313D1F7D6937934FB1E9573EFC">
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<mods:title id="AEE32DAA1D6EDE3F1C34CFDBB69A0D58">A new species in the Tylototriton asperrimus group (Caudata: Salamandridae) from central Laos</mods:title>
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</mods:titleInfo>
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<mods:name id="3B6EF7134BCD51679C5A151F0F0CBF4D" type="personal">
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<mods:role id="7798DFC0D2F2813F6D858DDEA228AB4D">
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<mods:roleTerm id="08C1C3C5E1C3CFBCF040CC43246CA4D2">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="87D84CC3C5666776470D1618303DDE54">Stuart, Bryan L.</mods:namePart>
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</mods:name>
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<mods:name id="558E7A76AE3282DF6137E0156DD04E99" type="personal">
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<mods:role id="12F95695ADA58D83CCCF4061E8EB5952">
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<mods:roleTerm id="4B66A8D236D29D7A90406831CA83B95B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="DD8888076DECA571214D80C3A393710E">Phimmachak, Somphouthone</mods:namePart>
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</mods:name>
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<mods:name id="9B5275F3BED5A9DBCC52DC171594D376" type="personal">
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<mods:role id="691F4B0785E63647841F4624EC818F5F">
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<mods:roleTerm id="DDF79818114C49D91FD96779AD630388">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="08174C19E4AFF475819A967533AB0979">Sivongxay, Niane</mods:namePart>
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</mods:name>
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<mods:name id="820C5C14F406E80FFDD0288E36E8D2B7" type="personal">
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<mods:role id="7604E40C7DC85429A090515F79AD6A66">
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<mods:roleTerm id="8625B78D47DC7B859B60E15FF19DEB90">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D20DA0FA460096833B22C27F6FF98297">Robichaud, William G.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="7AD29EFB1B78BEB84326E524312FDE95">text</mods:typeOfResource>
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<mods:titleInfo id="8A10186835887B042EF94A9DD1D75D6E">
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<mods:title id="75B2E55D6ABABBF35694336276E6FF39">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="8115C3F9DB6CC97AB553AC72BC6ADDB2">
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<mods:date id="1806CE09EEEF2573EAB5572BB10CC7F7">2010</mods:date>
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<mods:detail id="1962D336D9556A14D74A0098CF1BD468" type="volume">
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<mods:number id="4F619AF4932A8519501FDA3B255223C1">2650</mods:number>
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<mods:start id="25B587BCF8CCF62D67824A131A27C1F7">19</mods:start>
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<mods:end id="4B6694ED12F2D414E73F5592CE1B4F92">32</mods:end>
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<mods:classification id="C4EAC3C8CEFE7CE8DA680B37D32658E6">journal article</mods:classification>
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<mods:identifier id="823F75C58A8EFE48363753805675BF43" type="DOI">10.5281/zenodo.198798</mods:identifier>
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<mods:identifier id="49FA921EB34B8BEB0D979404A3B6F562" type="GBIF-Dataset">7e6ef230-5a66-4dd0-b41e-411da4adee7b</mods:identifier>
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<mods:identifier id="32271CFA7C661760F2BB9DD8A17DAF49" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="E1E85085A2270C91A6823F051CD59776" type="Zenodo-Dep">198798</mods:identifier>
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<treatment id="4A38FA2DFFB5FFCDE5F4B8BB321FF86B" ID-DOI="http://doi.org/10.5281/zenodo.6196181" ID-GBIF-Taxon="119401160" ID-Zenodo-Dep="6196181" LSID="urn:lsid:plazi:treatment:4A38FA2DFFB5FFCDE5F4B8BB321FF86B" httpUri="http://treatment.plazi.org/id/4A38FA2DFFB5FFCDE5F4B8BB321FF86B" lastPageId="8" lastPageNumber="27" pageId="4" pageNumber="23">
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<subSubSection id="8A8B18B0FFB5FFC1E5F4B8BB30EFF988" box="[151,491,1527,1553]" pageId="4" pageNumber="23" type="nomenclature">
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<paragraph id="C22E4B3BFFB5FFC1E5F4B8BB30EFF988" blockId="4.[151,491,1527,1553]" box="[151,491,1527,1553]" pageId="4" pageNumber="23">
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<heading id="9966FC57FFB5FFC1E5F4B8BB30EFF988" bold="true" box="[151,491,1527,1553]" fontSize="11" level="1" pageId="4" pageNumber="23" reason="1">
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||
<emphasis id="F0E59729FFB5FFC1E5F4B8BB30EFF988" bold="true" box="[151,491,1527,1553]" pageId="4" pageNumber="23">
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||
<taxonomicName id="059130B8FFB5FFC1E5F4B8BB3082F988" ID-CoL="7DBM4" box="[151,390,1527,1553]" class="Amphibia" family="Salamandridae" genus="Tylototriton" kingdom="Animalia" order="Caudata" pageId="4" pageNumber="23" phylum="Chordata" rank="species" species="notialis" status="sp. nov.">
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<emphasis id="F0E59729FFB5FFC1E5F4B8BB3082F988" bold="true" box="[151,390,1527,1553]" italics="true" pageId="4" pageNumber="23">Tylototriton notialis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="EBD62A52FFB5FFC1E4EEB8BB30EFF988" box="[397,491,1527,1553]" pageId="4" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="8A8B18B0FFB5FFC1E5F4BB7333D9F86D" pageId="4" pageNumber="23" type="materials_examined">
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||
<paragraph id="C22E4B3BFFB5FFC1E5F4BB7333FDF950" blockId="4.[151,1438,1599,2036]" pageId="4" pageNumber="23">
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||
<emphasis id="F0E59729FFB5FFC1E5F4BB73301AF9C0" bold="true" box="[151,286,1599,1625]" pageId="4" pageNumber="23">
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||
<typeStatus id="1D2AF599FFB5FFC1E5F4BB73301CF9C0" box="[151,280,1599,1625]" pageId="4" pageNumber="23" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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<collectionCode id="A480D3FEFFB5FFC1E444BB733085F9C0" LSID="urn:lsid:biocol.org:col:34795" box="[295,385,1599,1625]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="4" pageNumber="23">FMNH</collectionCode>
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<date id="B62F6DFBFFB5FFC1E4E9BB7330ECF9C0" box="[394,488,1599,1625]" pageId="4" pageNumber="23" value="21-11-27">271121</date>
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(field tag BLS 10775), adult male (
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<figureCitation id="5AAA57BEFFB5FFC1E6DFBB733508F9C0" box="[956,1036,1599,1625]" captionStart="FIGURE 3" captionStartId="6.[151,255,1780,1804]" captionTargetBox="[152,1436,287,1739]" captionTargetId="figure@6.[151,1436,269,1760]" captionTargetPageId="6" captionText="FIGURE 3. Tylototriton notialis sp. nov. in life. From top to bottom: dorsal view of holotype (FMNH 271121), dorsal view of paratype (FMNH 271120), ventral view of holotype (FMNH 271121), ventral view of paratype (FMNH 271120)." httpUri="https://zenodo.org/record/198801/files/figure.png" pageId="4" pageNumber="23">Fig. 3</figureCitation>
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),
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<collectingCountry id="BA860BABFFB5FFC1E144BB73356DF9C0" box="[1063,1129,1599,1625]" name="Laos" pageId="4" pageNumber="23">Laos</collectingCountry>
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, Khammouan Province, Boualapha District, Nakai-Nam Theun National Protected Area, Nam On River catchment, Phou Ak escarpment,
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<geoCoordinate id="A7A52DFCFFB5FFC1E456BBC630DBF93D" box="[309,479,1674,1700]" direction="north" orientation="latitude" pageId="4" pageNumber="23" precision="1" value="17.644333">17°38'39.6"N</geoCoordinate>
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<geoCoordinate id="A7A52DFCFFB5FFC1E48BBBC6339AF93D" box="[488,670,1674,1700]" direction="east" orientation="longitude" pageId="4" pageNumber="23" precision="1" value="105.73675">105°44'12.3"E</geoCoordinate>
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(
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<figureCitation id="5AAA57BEFFB5FFC1E7D3BBC633FAF93D" box="[688,766,1674,1700]" captionStart="FIGURE 4" captionStartId="7.[151,255,1834,1858]" captionTargetBox="[375,1212,1236,1822]" captionTargetId="figure@7.[375,1212,1236,1824]" captionTargetPageId="7" captionText="FIGURE 4. Map illustrating the type localities of Tylototriton notialis sp. nov. (star) and T. vietnamensis (circle)." httpUri="https://zenodo.org/record/198802/files/figure.png" pageId="4" pageNumber="23">Fig. 4</figureCitation>
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),
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<quantity id="0569E6DEFFB5FFC1E674BBC63262F93D" box="[791,870,1674,1700]" metricMagnitude="2" metricUnit="m" metricValue="9.8" pageId="4" pageNumber="23" unit="m" value="980.0">980 m</quantity>
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elev., coll.
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<date id="B62F6DFBFFB5FFC1E699BBC635A5F93D" box="[1018,1185,1674,1700]" pageId="4" pageNumber="23" value="2007-05-27">27 May 2007</date>
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by Bryan L. Stuart, Somphouthone Phimmachak, and Niane Sivongxay.
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</paragraph>
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<paragraph id="C22E4B3BFFB5FFC1E5A5BB983343F8A0" blockId="4.[151,1438,1599,2036]" pageId="4" pageNumber="23">
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||
<emphasis id="F0E59729FFB5FFC1E5A5BB983049F977" bold="true" box="[198,333,1748,1774]" pageId="4" pageNumber="23">
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<typeStatus id="1D2AF599FFB5FFC1E5A5BB98304CF977" box="[198,328,1748,1774]" pageId="4" pageNumber="23" type="paratype">Paratypes</typeStatus>
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:
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</emphasis>
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<collectionCode id="A480D3FEFFB5FFC1E436BB9830AEF977" LSID="urn:lsid:biocol.org:col:34795" box="[341,426,1748,1774]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="4" pageNumber="23">FMNH</collectionCode>
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||
<date id="B62F6DFBFFB5FFC1E4D2BB98330FF977" box="[433,523,1748,1774]" pageId="4" pageNumber="23" value="20-11-27">271120</date>
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, adult male (
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||
<figureCitation id="5AAA57BEFFB5FFC1E7C0BB9833EEF977" box="[675,746,1748,1774]" captionStart="FIGURE 3" captionStartId="6.[151,255,1780,1804]" captionTargetBox="[152,1436,287,1739]" captionTargetId="figure@6.[151,1436,269,1760]" captionTargetPageId="6" captionText="FIGURE 3. Tylototriton notialis sp. nov. in life. From top to bottom: dorsal view of holotype (FMNH 271121), dorsal view of paratype (FMNH 271120), ventral view of holotype (FMNH 271121), ventral view of paratype (FMNH 271120)." httpUri="https://zenodo.org/record/198801/files/figure.png" pageId="4" pageNumber="23">Fig. 3</figureCitation>
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||
), same data as
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<typeStatus id="1D2AF599FFB5FFC1E6FEBB983500F977" box="[925,1028,1748,1774]" pageId="4" pageNumber="23" type="holotype">holotype</typeStatus>
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||
except coll.
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<date id="B62F6DFBFFB5FFC1E1F8BB98343FF977" box="[1179,1339,1748,1774]" pageId="4" pageNumber="23" value="2007-05-26">26 May 2007</date>
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||
.
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||
<collectionCode id="A480D3FEFFB5FFC1E024BB983498F977" LSID="urn:lsid:biocol.org:col:34795" box="[1351,1436,1748,1774]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="4" pageNumber="23">FMNH</collectionCode>
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<date id="B62F6DFBFFB5FFC1E5F4BBB631F6F88D" box="[151,242,1786,1812]" pageId="4" pageNumber="23" value="22-11-27">271122</date>
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, adult female, same data as
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<typeStatus id="1D2AF599FFB5FFC1E75CBBB633A2F88D" box="[575,678,1786,1812]" pageId="4" pageNumber="23" type="holotype">holotype</typeStatus>
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||
except
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<geoCoordinate id="A7A52DFCFFB5FFC1E667BBB632A0F88D" box="[772,932,1786,1812]" direction="north" orientation="latitude" pageId="4" pageNumber="23" precision="1" value="17.650944">17°39'03.4"N</geoCoordinate>
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||
<geoCoordinate id="A7A52DFCFFB5FFC1E6CEBBB63558F88D" box="[941,1116,1786,1812]" direction="east" orientation="longitude" pageId="4" pageNumber="23" precision="1" value="105.74033">105°44'25.2"E</geoCoordinate>
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||
, ca.
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<quantity id="0569E6DEFFB5FFC1E1F0BBB635F0F88D" box="[1171,1268,1786,1812]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="23" unit="m" value="1000.0">1,000 m</quantity>
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elev., coll.
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<date id="B62F6DFBFFB5FFC1E01DBBB6300BF8A0" pageId="4" pageNumber="23" value="2006-05-22">22 May 2006</date>
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by William G. Robichaud.
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</paragraph>
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<paragraph id="C22E4B3BFFB5FFC1E5A6BA083275F81D" blockId="4.[151,1438,1599,2036]" pageId="4" pageNumber="23">
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||
<emphasis id="F0E59729FFB5FFC1E5A6BA0830B4F8C7" bold="true" box="[197,432,1860,1886]" pageId="4" pageNumber="23">Referred material:</emphasis>
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||
<collectionCode id="A480D3FEFFB5FFC1E4D4BA083308F8C7" LSID="urn:lsid:biocol.org:col:34795" box="[439,524,1860,1886]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="4" pageNumber="23">FMNH</collectionCode>
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<date id="B62F6DFBFFB5FFC1E770BA08336EF8C7" box="[531,618,1860,1886]" pageId="4" pageNumber="23" value="25-11-27">271125</date>
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||
(one larva;
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||
<figureCitation id="5AAA57BEFFB5FFC1E79BBA08323AF8C7" box="[760,830,1860,1886]" captionStart="FIGURE 5" captionStartId="8.[151,255,1159,1183]" captionTargetBox="[195,1391,493,1138]" captionTargetId="figure@8.[195,1392,493,1138]" captionTargetPageId="8" captionText="FIGURE 5. Dorsal and lateral view of larva (FMNH 271129) of Tylototriton notialis in preservative (total length 20.9 mm)." httpUri="https://zenodo.org/record/198803/files/figure.png" pageId="4" pageNumber="23">Fig. 5</figureCitation>
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||
), same data as
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<typeStatus id="1D2AF599FFB5FFC1E692BA083552F8C7" box="[1009,1110,1860,1886]" pageId="4" pageNumber="23" type="paratype">paratype</typeStatus>
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||
male.
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||
<collectionCode id="A480D3FEFFB5FFC1E1C6BA0835FEF8C7" LSID="urn:lsid:biocol.org:col:34795" box="[1189,1274,1860,1886]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="4" pageNumber="23">FMNH</collectionCode>
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||
<date id="B62F6DFBFFB5FFC1E062BA08345CF8C7" box="[1281,1368,1860,1886]" pageId="4" pageNumber="23" value="29-11-27">271129</date>
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||
(nine larvae), same data as
|
||
<typeStatus id="1D2AF599FFB5FFC1E4F1BA2630F3F81D" box="[402,503,1898,1924]" pageId="4" pageNumber="23" type="paratype">paratype</typeStatus>
|
||
male except coll.
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||
<date id="B62F6DFBFFB5FFC1E7AEBA263268F81D" box="[717,876,1898,1924]" pageId="4" pageNumber="23" value="2007-05-29">29 May 2007</date>
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||
.
|
||
</paragraph>
|
||
<paragraph id="C22E4B3BFFB5FFC1E5A5BAC333D9F86D" blockId="4.[151,1438,1599,2036]" pageId="4" pageNumber="23">
|
||
<emphasis id="F0E59729FFB5FFC1E5A5BAC3304CF830" bold="true" box="[198,328,1935,1961]" pageId="4" pageNumber="23">Subgenus:</emphasis>
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||
The new species is assigned to the subgenus
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<emphasis id="F0E59729FFB5FFC1E6FEBADC3503F830" box="[925,1031,1936,1961]" italics="true" pageId="4" pageNumber="23">Yaotriton</emphasis>
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||
<bibRefCitation id="A60036CAFFB5FFC1E17BBAC33454F830" author="Dubois" box="[1048,1360,1935,1961]" pageId="4" pageNumber="23" refString="Dubois, A. & Raffaelli, J. (2009) A new ergotaxonomy of the family Salamandridae Goldfuss, 1820 (Amphibia, Urodela). Alytes, 26, 1 - 85." type="journal article" year="2009">Dubois & Raffaëlli 2009</bibRefCitation>
|
||
(=
|
||
<taxonomicName id="059130B8FFB5FFC1E0E5BADC301FF857" class="Amphibia" family="Salamandridae" genus="Tylototriton" kingdom="Animalia" order="Caudata" pageId="4" pageNumber="23" phylum="Chordata" rank="species" species="asperrimus">
|
||
<emphasis id="F0E59729FFB5FFC1E0E5BADC301FF857" italics="true" pageId="4" pageNumber="23">T. asperrimus</emphasis>
|
||
</taxonomicName>
|
||
species group of
|
||
<bibRefCitation id="A60036CAFFB5FFC1E484BAF83394F857" author="Fei" box="[487,656,1972,1998]" pageId="4" pageNumber="23" refString="Fei, L., Ye, C., Jiang, J. & Xie, F. (2005) An Illustrated Key to Chinese Amphibians. Sichuan Publishing House of Science and Technology, Chongqing, Sichuan, 340 pp." type="book" year="2005">
|
||
Fei
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||
<emphasis id="F0E59729FFB5FFC1E777BAF9334AF857" box="[532,590,1973,1998]" italics="true" pageId="4" pageNumber="23">et al.</emphasis>
|
||
2005
|
||
</bibRefCitation>
|
||
) based on its phylogenetic position (
|
||
<figureCitation id="5AAA57BEFFB5FFC1E15FBAF8357BF857" box="[1084,1151,1972,1998]" captionStart="FIGURE 1" captionStartId="4.[151,260,1356,1380]" captionTargetBox="[151,1436,485,1341]" captionTargetId="figure@4.[151,1436,483,1344]" captionTargetPageId="4" captionText="FIGURE 1. Fifty percent majority-rule consensus phylogram resulting from mixed-model Bayesian analysis of mitochondrial DNA from salamandrids. Numbers at nodes are Bayesian posterior probabilities. Numbers in parentheses refer to ID in Table 1." httpUri="https://zenodo.org/record/198799/files/figure.png" pageId="4" pageNumber="23">Fig. 1</figureCitation>
|
||
) and mostly dark dorsal and ventral coloration (
|
||
<bibRefCitation id="A60036CAFFB5FFC1E4C4BA9633D4F86D" author="Dubois" box="[423,720,2010,2036]" pageId="4" pageNumber="23" refString="Dubois, A. & Raffaelli, J. (2009) A new ergotaxonomy of the family Salamandridae Goldfuss, 1820 (Amphibia, Urodela). Alytes, 26, 1 - 85." type="journal article" year="2009">Dubois & Raffaëlli 2009</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8A8B18B0FFB4FFC0E5A5BDDB356EFF4E" pageId="5" pageNumber="24" type="etymology">
|
||
<paragraph id="C22E4B3BFFB4FFC0E5A5BDDB356EFF4E" blockId="5.[151,1435,151,215]" pageId="5" pageNumber="24">
|
||
<emphasis id="F0E59729FFB4FFC0E5A5BDDB3053FF28" bold="true" box="[198,343,151,177]" pageId="5" pageNumber="24">Etymology:</emphasis>
|
||
The specific epithet taken from
|
||
<taxonomicName id="059130B8FFB4FFC0E78DBDD43242FF28" box="[750,838,152,177]" class="Amphibia" family="Salamandridae" genus="Tylototriton" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="24" phylum="Chordata" rank="species" species="notialis">
|
||
<emphasis id="F0E59729FFB4FFC0E78DBDD43242FF28" box="[750,838,152,177]" italics="true" pageId="5" pageNumber="24">notialis</emphasis>
|
||
</taxonomicName>
|
||
L. for southern, in reference to the new species having the southernmost known locality of any member in the
|
||
<taxonomicName id="059130B8FFB4FFC0E615BDF13513FF4F" box="[886,1047,189,214]" class="Amphibia" family="Salamandridae" genus="Tylototriton" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="24" phylum="Chordata" rank="species" species="asperrimus">
|
||
<emphasis id="F0E59729FFB4FFC0E615BDF13513FF4F" box="[886,1047,189,214]" italics="true" pageId="5" pageNumber="24">T. asperrimus</emphasis>
|
||
</taxonomicName>
|
||
group.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="96EE1BB3FFB4FFC0E5F4B9533306FBEE" httpUri="https://zenodo.org/record/198800/files/figure.png" pageId="5" pageNumber="24" targetBox="[151,1435,257,1031]" targetPageId="5">
|
||
<paragraph id="C22E4B3BFFB4FFC0E5F4B9533306FBEE" blockId="5.[151,1436,1055,1143]" pageId="5" pageNumber="24">
|
||
<emphasis id="F0E59729FFB4FFC0E5F4B953301EFBAE" bold="true" box="[151,282,1055,1079]" pageId="5" pageNumber="24">FIGURE 2.</emphasis>
|
||
Fifty percent majority-rule consensus phylogram resulting from mixed-model Bayesian analysis of nuclear DNA (part of the POMC gene) from salamandrids. Numbers at nodes are Bayesian posterior probabilities. Numbers in parentheses refer to ID in Table 1.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="8A8B18B0FFB4FFC0E5A6B9E93351FAB6" pageId="5" pageNumber="24" type="diagnosis">
|
||
<paragraph id="C22E4B3BFFB4FFC0E5A6B9E93351FAB6" blockId="5.[151,1437,1189,2036]" pageId="5" pageNumber="24">
|
||
<emphasis id="F0E59729FFB4FFC0E5A6B9E93043FB26" bold="true" box="[197,327,1189,1215]" pageId="5" pageNumber="24">Diagnosis:</emphasis>
|
||
<taxonomicName id="059130B8FFB4FFC0E42CB9E9333CFB27" box="[335,568,1189,1214]" class="Amphibia" family="Salamandridae" genus="Tylototriton" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="24" phylum="Chordata" rank="species" species="notialis">
|
||
<emphasis id="F0E59729FFB4FFC0E42CB9E9333CFB27" box="[335,568,1189,1214]" italics="true" pageId="5" pageNumber="24">Tylototriton notialis</emphasis>
|
||
</taxonomicName>
|
||
is a
|
||
<taxonomicName id="059130B8FFB4FFC0E713B9E93285FB26" authority="Yaotriton" authorityName="Yaotriton" box="[624,897,1189,1215]" class="Amphibia" family="Salamandridae" genus="Tylototriton" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="24" phylum="Chordata" rank="genus">
|
||
<emphasis id="F0E59729FFB4FFC0E713B9E933FEFB27" box="[624,762,1189,1214]" italics="true" pageId="5" pageNumber="24">Tylototriton</emphasis>
|
||
(
|
||
<emphasis id="F0E59729FFB4FFC0E668B9E93273FB27" box="[779,887,1189,1214]" italics="true" pageId="5" pageNumber="24">Yaotriton</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
having the combination of distinct, knob-like rib nodules (dorsolateral row of glandular warts); glandular warts on most of the remaining dorsal and ventral surfaces; very dark brown to black coloration on dorsum and venter; and bright orange coloration on rib nodules and posterior end of parotoid.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8A8B18B0FFB4FFC2E5A5B876342AF833" lastPageId="7" lastPageNumber="26" pageId="5" pageNumber="24" type="description">
|
||
<paragraph id="C22E4B3BFFB4FFC0E5A5B8763351F8C6" blockId="5.[151,1437,1189,2036]" pageId="5" pageNumber="24">
|
||
<emphasis id="F0E59729FFB4FFC0E5A5B87630F3FACD" bold="true" box="[198,503,1338,1364]" pageId="5" pageNumber="24">
|
||
Description of
|
||
<typeStatus id="1D2AF599FFB4FFC0E4E1B87630F6FACD" box="[386,498,1338,1364]" pageId="5" pageNumber="24" type="holotype">holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
Habitus moderately stout. Head broader than body, longer than wide, slightly sloping in profile. Snout short, truncate in dorsal view, rounded in profile, exceeding beyond lower jaw. Nostrils close to snout tip, right nostril slightly visible from above. Vomeropalatine teeth in two rows, anteriorly in contact and beginning posterior to the anterior margin of the choanae, converging into two parallel rows briefly, then diverging from one another. Glandular ridge on midline of crown from above anterior edge of eye to middle of head. Glandular ridge on outer margin of crown from above eye to base of parotoid. Parotoids enlarged, projecting backward. Glandular patch of skin on nape. Distinct vertebral tubercular ridge from posterior end of crown to base of tail, separated from ridge on midline of crown. Dorsolateral row of approximately 14 large glandular warts (rib nodules) on each side from level of axilla to base of tail, distinctly knob-like anteriorly, becoming smaller posteriorly, merging at level of groin. Smaller, glandular warts on most of remaining dorsal and ventral surfaces, warts on crown, nape, and back with clusters of glands and sometimes conical, those on throat granular and widely spaced, those on belly arranged in striations perpendicular to body axis, with a smooth, glandular, ovoid patch of skin on chest. Weak gular fold present. Four fingers, five toes, all without webbing. Tail laterally compressed, narrow dorsal fin, smooth ventral ridge, tip acuminate in profile.
|
||
</paragraph>
|
||
<paragraph id="C22E4B3BFFB4FFC0E5A5BA263499F86D" blockId="5.[151,1437,1189,2036]" pageId="5" pageNumber="24">
|
||
<emphasis id="F0E59729FFB4FFC0E5A5BA263309F81D" bold="true" box="[198,525,1898,1924]" pageId="5" pageNumber="24">
|
||
Color of
|
||
<typeStatus id="1D2AF599FFB4FFC0E423BA2630A8F81D" box="[320,428,1898,1924]" pageId="5" pageNumber="24" type="holotype">holotype</typeStatus>
|
||
in life:
|
||
</emphasis>
|
||
Body very dark brown. Margin of upper and lower lip, posterior end of parotoid, dorsolateral glands (rib nodules), dorsal margin of tail, outer margin of upper surface of hand and finger tips, lower surface of hand except center of palm, dorsal surface of toe tips, ventral surface of toes and extending from fifth toe to base of foot, cloacal region continuing to ventral ridge of tail bright orange.
|
||
</paragraph>
|
||
<paragraph id="C22E4B3BFFB7FFC3E5F4BDD435D8FF4E" blockId="6.[151,1436,152,215]" pageId="6" pageNumber="25">Anterior surface of forelimb from axilla to base of first finger with narrow, broken, yellow stripe. Anterior and ventral surfaces of hindlimbs with small, scattered yellow spots. Iris dark brown, pupil black.</paragraph>
|
||
<caption id="96EE1BB3FFB7FFC3E5F4BBB83499F8B5" httpUri="https://zenodo.org/record/198801/files/figure.png" pageId="6" pageNumber="25" targetBox="[152,1436,287,1739]" targetPageId="6">
|
||
<paragraph id="C22E4B3BFFB7FFC3E5F4BBB83499F8B5" blockId="6.[151,1437,1780,1836]" pageId="6" pageNumber="25">
|
||
<emphasis id="F0E59729FFB7FFC3E5F4BBB8301EF895" bold="true" box="[151,282,1780,1804]" pageId="6" pageNumber="25">FIGURE 3.</emphasis>
|
||
<taxonomicName id="059130B8FFB7FFC3E441BBB930FCF895" box="[290,504,1781,1804]" class="Amphibia" family="Salamandridae" genus="Tylototriton" kingdom="Animalia" order="Caudata" pageId="6" pageNumber="25" phylum="Chordata" rank="species" species="notialis" status="sp. nov.">
|
||
<emphasis id="F0E59729FFB7FFC3E441BBB930FCF895" box="[290,504,1781,1804]" italics="true" pageId="6" pageNumber="25">Tylototriton notialis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EBD62A52FFB7FFC3E763BBB83353F895" box="[512,599,1780,1804]" pageId="6" pageNumber="25" rank="species">
|
||
<emphasis id="F0E59729FFB7FFC3E763BBB83353F895" bold="true" box="[512,599,1780,1804]" pageId="6" pageNumber="25">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
in life. From top to bottom: dorsal view of holotype (FMNH 271121), dorsal view of paratype (FMNH 271120), ventral view of holotype (FMNH 271121), ventral view of paratype (FMNH 271120).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="C22E4B3BFFB7FFC3E5A5BA1630C7F803" blockId="6.[151,1436,1882,2020]" pageId="6" pageNumber="25">
|
||
<emphasis id="F0E59729FFB7FFC3E5A5BA163377F8ED" bold="true" box="[198,627,1882,1908]" pageId="6" pageNumber="25">
|
||
Color of
|
||
<typeStatus id="1D2AF599FFB7FFC3E45ABA1630A0F8ED" box="[313,420,1882,1908]" pageId="6" pageNumber="25" type="holotype">holotype</typeStatus>
|
||
in preservative:
|
||
</emphasis>
|
||
Body faded to a lighter shade of brown. Orange markings faded to yellow or creamy-yellow.
|
||
</paragraph>
|
||
<paragraph id="C22E4B3BFFB7FFC3E5A5BAE933E3F87D" blockId="6.[151,1436,1882,2020]" pageId="6" pageNumber="25">
|
||
<emphasis id="F0E59729FFB7FFC3E5A5BAE9331FF826" bold="true" box="[198,539,1957,1983]" pageId="6" pageNumber="25">
|
||
Measurements of
|
||
<typeStatus id="1D2AF599FFB7FFC3E4C5BAE93312F826" box="[422,534,1957,1983]" pageId="6" pageNumber="25" type="holotype">holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
SVLA 60.2; SVLP 69.1; AXGR 30.5; TTL 130.4; TAL 63.1; TAD 6.5; HL 19.6; HW 15.9; EN 4.0; IN 4.9; AL 23.7; PL 23.0.
|
||
</paragraph>
|
||
<paragraph id="C22E4B3BFFB6FFC2E5A5BDDB30D1FEBB" blockId="7.[151,1436,151,290]" pageId="7" pageNumber="26">
|
||
<emphasis id="F0E59729FFB6FFC2E5A5BDDB3047FF28" bold="true" box="[198,323,151,177]" pageId="7" pageNumber="26">Variation:</emphasis>
|
||
The body coloration of the
|
||
<typeStatus id="1D2AF599FFB6FFC2E7F0BDD433FCFF2B" box="[659,760,152,178]" pageId="7" pageNumber="26" type="paratype">paratype</typeStatus>
|
||
male (collected on land) was black rather than very dark brown as in the
|
||
<typeStatus id="1D2AF599FFB6FFC2E43ABDF130C6FF4E" box="[345,450,189,215]" pageId="7" pageNumber="26" type="holotype">holotype</typeStatus>
|
||
(collected in water) and
|
||
<typeStatus id="1D2AF599FFB6FFC2E78CBDF13252FF4E" box="[751,854,189,215]" pageId="7" pageNumber="26" type="paratype">paratype</typeStatus>
|
||
female (collected on land).
|
||
<bibRefCitation id="A60036CAFFB6FFC2E1CABDF13498FF4E" box="[1193,1436,189,215]" pageId="7" pageNumber="26" refString="Bohme, W., Schottler, T., Nguyen, T. Q. & Kohler, J. (2005) A new species of salamander, genus Tylototriton (Urodela: Salamandridae), from northern Vietnam. Salamandra, 41, 215 - 220." type="journal article">
|
||
Böhme
|
||
<emphasis id="F0E59729FFB6FFC2E064BDF13441FF4F" box="[1287,1349,189,214]" italics="true" pageId="7" pageNumber="26">et al.</emphasis>
|
||
(2005)
|
||
</bibRefCitation>
|
||
reported that
|
||
<taxonomicName id="059130B8FFB6FFC2E451BDAE30E1FF62" box="[306,485,226,251]" class="Amphibia" family="Salamandridae" genus="Tylototriton" kingdom="Animalia" order="Caudata" pageId="7" pageNumber="26" phylum="Chordata" rank="species" species="vietnamensis">
|
||
<emphasis id="F0E59729FFB6FFC2E451BDAE30E1FF62" box="[306,485,226,251]" italics="true" pageId="7" pageNumber="26">T. vietnamensis</emphasis>
|
||
</taxonomicName>
|
||
taken on land were darker in coloration than those living in water. Measurements are summarized in
|
||
<tableCitation id="8F137E80FFB6FFC2E415BC4430CBFEBB" box="[374,463,264,290]" captionStart="TABLE 3" captionStartId="7.[151,239,332,356]" captionTargetBox="[151,1435,415,1184]" captionTargetPageId="7" captionText="TABLE 3. Measurements (mm) of Tylototriton notialis sp. nov., T. vietnamensis, and T. cf. vietnamensis. Abbreviations defined in the text." httpUri="http://table.plazi.org/id/96EE1BB3FFB6FFC2E5F4BC00305BFE1D" pageId="7" pageNumber="26" tableUuid="96EE1BB3FFB6FFC2E5F4BC00305BFE1D">Table 3</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="96EE1BB3FFB6FFC2E5F4BC00305BFE1D" ID-Table-UUID="96EE1BB3FFB6FFC2E5F4BC00305BFE1D" httpUri="http://table.plazi.org/id/96EE1BB3FFB6FFC2E5F4BC00305BFE1D" pageId="7" pageNumber="26" targetBox="[151,1435,415,1184]" targetIsTable="true" targetPageId="7">
|
||
<paragraph id="C22E4B3BFFB6FFC2E5F4BC00305BFE1D" blockId="7.[151,1435,332,388]" pageId="7" pageNumber="26">
|
||
<emphasis id="F0E59729FFB6FFC2E5F4BC00300DFEFD" bold="true" box="[151,265,332,356]" pageId="7" pageNumber="26">TABLE 3.</emphasis>
|
||
Measurements (mm) of
|
||
<taxonomicName id="059130B8FFB6FFC2E770BC0133E3FEFD" box="[531,743,333,356]" class="Amphibia" family="Salamandridae" genus="Tylototriton" kingdom="Animalia" order="Caudata" pageId="7" pageNumber="26" phylum="Chordata" rank="species" species="notialis" status="sp. nov.">
|
||
<emphasis id="F0E59729FFB6FFC2E770BC0133E3FEFD" box="[531,743,333,356]" italics="true" pageId="7" pageNumber="26">Tylototriton notialis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EBD62A52FFB6FFC2E78DBC003240FEFD" box="[750,836,332,356]" pageId="7" pageNumber="26" rank="species">
|
||
<emphasis id="F0E59729FFB6FFC2E78DBC003240FEFD" bold="true" box="[750,836,332,356]" pageId="7" pageNumber="26">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="059130B8FFB6FFC2E632BC0132F2FEFD" box="[849,1014,333,356]" class="Amphibia" family="Salamandridae" genus="Tylototriton" kingdom="Animalia" order="Caudata" pageId="7" pageNumber="26" phylum="Chordata" rank="species" species="vietnamensis">
|
||
<emphasis id="F0E59729FFB6FFC2E632BC0132F2FEFD" box="[849,1014,333,356]" italics="true" pageId="7" pageNumber="26">T. vietnamensis</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="059130B8FFB6FFC2E152BC0135F3FEFD" box="[1073,1271,333,356]" class="Amphibia" family="Salamandridae" genus="Tylototriton" kingdom="Animalia" order="Caudata" pageId="7" pageNumber="26" phylum="Chordata" rank="species" species="vietnamensis">
|
||
<emphasis id="F0E59729FFB6FFC2E152BC013547FEFD" box="[1073,1091,333,356]" italics="true" pageId="7" pageNumber="26">T.</emphasis>
|
||
cf.
|
||
<emphasis id="F0E59729FFB6FFC2E10EBC0135F3FEFD" box="[1133,1271,333,356]" italics="true" pageId="7" pageNumber="26">vietnamensis</emphasis>
|
||
</taxonomicName>
|
||
. Abbreviations defined in the text.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="C22E4B3BFFB6FFC2E5FCBCD33316FB39" pageId="7" pageNumber="26">
|
||
<table id="B091B99BFFB6003AE5F4BCD3349FFB39" box="[151,1435,415,1184]" gridcols="4" gridrows="13" pageId="7" pageNumber="26">
|
||
<tr id="7CA14979FFB6003AE5F4BCD3349FFDDB" box="[151,1435,415,578]" gridrow="0" pageId="7" pageNumber="26">
|
||
<th id="3F702005FFB6003AE5F4BCD3335FFDDB" box="[151,603,415,578]" gridcol="0" gridrow="0" pageId="7" pageNumber="26">
|
||
Measurement
|
||
<emphasis id="F0E59729FFB6FFC2E45DBCD330F6FE2D" bold="true" box="[318,498,415,436]" pageId="7" pageNumber="26">
|
||
<taxonomicName id="059130B8FFB6FFC2E45DBCD3309BFE2D" box="[318,415,415,436]" class="Amphibia" family="Salamandridae" genus="Tylototriton" kingdom="Animalia" order="Caudata" pageId="7" pageNumber="26" phylum="Chordata" rank="species" species="notialis" status="sp. nov.">
|
||
<emphasis id="F0E59729FFB6FFC2E45DBCD3309BFE2D" bold="true" box="[318,415,415,436]" italics="true" pageId="7" pageNumber="26">T. notialis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EBD62A52FFB6FFC2E4C6BCD330F6FE2D" box="[421,498,415,436]" pageId="7" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Khammouan, Laos Males (
|
||
<emphasis id="F0E59729FFB6FFC2E4E5BCAB3096FE65" box="[390,402,487,508]" italics="true" pageId="7" pageNumber="26">n</emphasis>
|
||
=2) Female (
|
||
<emphasis id="F0E59729FFB6FFC2E74DBCAB333EFE65" box="[558,570,487,508]" italics="true" pageId="7" pageNumber="26">n</emphasis>
|
||
=1) Range SVLA 56.5–60.2 73.4
|
||
</th>
|
||
<th id="3F702005FFB6003AE7E3BCD3326BFDDB" box="[640,879,415,578]" gridcol="1" gridrow="0" pageId="7" pageNumber="26">
|
||
<taxonomicName id="059130B8FFB6FFC2E7E3BCEE3217FE2E" box="[640,787,418,439]" class="Amphibia" family="Salamandridae" genus="Tylototriton" kingdom="Animalia" order="Caudata" pageId="7" pageNumber="26" phylum="Chordata" rank="species" species="vietnamensis">
|
||
<emphasis id="F0E59729FFB6FFC2E7E3BCEE3217FE2E" box="[640,787,418,439]" italics="true" pageId="7" pageNumber="26">T. vietnamensis</emphasis>
|
||
</taxonomicName>
|
||
types a Bac Giang, Vietnam Males (
|
||
<emphasis id="F0E59729FFB6FFC2E7ABBCAB33D0FE65" box="[712,724,487,508]" italics="true" pageId="7" pageNumber="26">n</emphasis>
|
||
=3) Range; Mean ± SD 48.3–53.6; 50.7 ± 2.7
|
||
</th>
|
||
<th id="3F702005FFB6003AE6E9BCD33585FDDB" box="[906,1153,415,578]" gridcol="2" gridrow="0" pageId="7" pageNumber="26">
|
||
<taxonomicName id="059130B8FFB6FFC2E6E9BCEC3519FE2C" box="[906,1053,416,437]" class="Amphibia" family="Salamandridae" genus="Tylototriton" kingdom="Animalia" order="Caudata" pageId="7" pageNumber="26" phylum="Chordata" rank="species" species="vietnamensis">
|
||
<emphasis id="F0E59729FFB6FFC2E6E9BCEC3519FE2C" box="[906,1053,416,437]" italics="true" pageId="7" pageNumber="26">T. vietnamensis</emphasis>
|
||
</taxonomicName>
|
||
topotypes Bac Giang, Vietnam Males (
|
||
<emphasis id="F0E59729FFB6FFC2E6B0BCAB32DBFE65" box="[979,991,487,508]" italics="true" pageId="7" pageNumber="26">n</emphasis>
|
||
=2) Range 61.3–62.8
|
||
</th>
|
||
<th id="3F702005FFB6003AE1F6BCD3349FFDDB" box="[1173,1435,415,578]" gridcol="3" gridrow="0" pageId="7" pageNumber="26">
|
||
<taxonomicName id="059130B8FFB6FFC2E1F6BCEC344CFE2C" box="[1173,1352,415,437]" class="Amphibia" family="Salamandridae" genus="Tylototriton" kingdom="Animalia" order="Caudata" pageId="7" pageNumber="26" phylum="Chordata" rank="species" species="vietnamensis">
|
||
<emphasis id="F0E59729FFB6FFC2E1F6BCEC35A3FE2C" box="[1173,1191,416,437]" italics="true" pageId="7" pageNumber="26">T.</emphasis>
|
||
cf.
|
||
<emphasis id="F0E59729FFB6FFC2E1AFBCEC344CFE2C" box="[1228,1352,416,437]" italics="true" pageId="7" pageNumber="26">vietnamensis</emphasis>
|
||
</taxonomicName>
|
||
Cao Bang, Vietnam Males (
|
||
<emphasis id="F0E59729FFB6FFC2E1BDBCAB35EEFE65" box="[1246,1258,487,508]" italics="true" pageId="7" pageNumber="26">n</emphasis>
|
||
=30) Range; Mean ± SD 56.1–68.4; 61.7 ± 3.2
|
||
</th>
|
||
</tr>
|
||
<tr id="7CA14979FFB6003AE5F4BF17349FFDE9" box="[151,1435,603,624]" gridrow="1" pageId="7" pageNumber="26">
|
||
<th id="3F702005FFB6003AE5F4BF17335FFDE9" box="[151,603,603,624]" gridcol="0" gridrow="1" pageId="7" pageNumber="26">SVLP 64.5–69.1 78.0</th>
|
||
<td id="3F702005FFB6003AE7E3BF17326BFDE9" box="[640,879,603,624]" gridcol="1" gridrow="1" pageId="7" pageNumber="26">-</td>
|
||
<td id="3F702005FFB6003AE6E9BF173585FDE9" box="[906,1153,603,624]" gridcol="2" gridrow="1" pageId="7" pageNumber="26">66.0–69.8</td>
|
||
<td id="3F702005FFB6003AE1F6BF17349FFDE9" box="[1173,1435,603,624]" gridcol="3" gridrow="1" pageId="7" pageNumber="26">61.5–74.3; 67.7 ± 3.5</td>
|
||
</tr>
|
||
<tr id="7CA14979FFB6003AE5F4BFC4349FFD04" box="[151,1435,648,669]" gridrow="2" pageId="7" pageNumber="26">
|
||
<th id="3F702005FFB6003AE5F4BFC4335FFD04" box="[151,603,648,669]" gridcol="0" gridrow="2" pageId="7" pageNumber="26">AXGR 27.1–30.5 39.3</th>
|
||
<td id="3F702005FFB6003AE7E3BFC4326BFD04" box="[640,879,648,669]" gridcol="1" gridrow="2" pageId="7" pageNumber="26">-</td>
|
||
<td id="3F702005FFB6003AE6E9BFC43585FD04" box="[906,1153,648,669]" gridcol="2" gridrow="2" pageId="7" pageNumber="26">30.6–33.7</td>
|
||
<td id="3F702005FFB6003AE1F6BFC4349FFD04" box="[1173,1435,648,669]" gridcol="3" gridrow="2" pageId="7" pageNumber="26">28.4–39.4; 34.0 ± 2.4</td>
|
||
</tr>
|
||
<tr id="7CA14979FFB6003AE5F4BFFB349FFD57" box="[151,1435,695,718]" gridrow="3" pageId="7" pageNumber="26">
|
||
<th id="3F702005FFB6003AE5F4BFFB335FFD57" box="[151,603,695,718]" gridcol="0" gridrow="3" pageId="7" pageNumber="26">TTL 109.1–130.4b 141.8</th>
|
||
<td id="3F702005FFB6003AE7E3BFFB326BFD57" box="[640,879,695,718]" gridcol="1" gridrow="3" pageId="7" pageNumber="26">113.9–121.8; 118.4 ± 4.1</td>
|
||
<td id="3F702005FFB6003AE6E9BFFB3585FD57" box="[906,1153,695,718]" gridcol="2" gridrow="3" pageId="7" pageNumber="26">125.3–125.4</td>
|
||
<td id="3F702005FFB6003AE1F6BFFB349FFD57" box="[1173,1435,695,718]" gridcol="3" gridrow="3" pageId="7" pageNumber="26">109.6–142.3; 126.1 ± 7.5</td>
|
||
</tr>
|
||
<tr id="7CA14979FFB6003AE5F4BFA8349FFD65" box="[151,1435,740,764]" gridrow="4" pageId="7" pageNumber="26">
|
||
<th id="3F702005FFB6003AE5F4BFA8335FFD65" box="[151,603,740,764]" gridcol="0" gridrow="4" pageId="7" pageNumber="26">TAL 45.3–63.1 2 65.2</th>
|
||
<td id="3F702005FFB6003AE7E3BFA8326BFD65" box="[640,879,740,764]" gridcol="1" gridrow="4" pageId="7" pageNumber="26">62.4–63.9; 63.1 ± 0.8</td>
|
||
<td id="3F702005FFB6003AE6E9BFA83585FD65" box="[906,1153,740,764]" gridcol="2" gridrow="4" pageId="7" pageNumber="26">55.6–59.4</td>
|
||
<td id="3F702005FFB6003AE1F6BFA8349FFD65" box="[1173,1435,740,764]" gridcol="3" gridrow="4" pageId="7" pageNumber="26">44.7–68.5; 58.4 ± 5.4</td>
|
||
</tr>
|
||
<tr id="7CA14979FFB6003AE5F4BE5F349FFCB1" box="[151,1435,787,808]" gridrow="5" pageId="7" pageNumber="26">
|
||
<th id="3F702005FFB6003AE5F4BE5F335FFCB1" box="[151,603,787,808]" gridcol="0" gridrow="5" pageId="7" pageNumber="26">TAD 6.5–7.1 7.4</th>
|
||
<td id="3F702005FFB6003AE7E3BE5F326BFCB1" box="[640,879,787,808]" gridcol="1" gridrow="5" pageId="7" pageNumber="26">6.6–8.0; 7.4 ± 0.7</td>
|
||
<td id="3F702005FFB6003AE6E9BE5F3585FCB1" box="[906,1153,787,808]" gridcol="2" gridrow="5" pageId="7" pageNumber="26">8.0–8.1</td>
|
||
<td id="3F702005FFB6003AE1F6BE5F349FFCB1" box="[1173,1435,787,808]" gridcol="3" gridrow="5" pageId="7" pageNumber="26">6.4–9.2; 7.6 ± 0.7</td>
|
||
</tr>
|
||
<tr id="7CA14979FFB6003AE5F4BE0C349FFCCC" box="[151,1435,832,853]" gridrow="6" pageId="7" pageNumber="26">
|
||
<th id="3F702005FFB6003AE5F4BE0C335FFCCC" box="[151,603,832,853]" gridcol="0" gridrow="6" pageId="7" pageNumber="26">HL 19.4–19.6 21.5</th>
|
||
<td id="3F702005FFB6003AE7E3BE0C326BFCCC" box="[640,879,832,853]" gridcol="1" gridrow="6" pageId="7" pageNumber="26">15.3–18.6; 17.2 ± 1.7</td>
|
||
<td id="3F702005FFB6003AE6E9BE0C3585FCCC" box="[906,1153,832,853]" gridcol="2" gridrow="6" pageId="7" pageNumber="26">18.3–18.4</td>
|
||
<td id="3F702005FFB6003AE1F6BE0C349FFCCC" box="[1173,1435,832,853]" gridcol="3" gridrow="6" pageId="7" pageNumber="26">17.5–21.2; 19.3 ± 0.9</td>
|
||
</tr>
|
||
<tr id="7CA14979FFB6003AE5F4BE23349FFC1D" box="[151,1435,879,900]" gridrow="7" pageId="7" pageNumber="26">
|
||
<th id="3F702005FFB6003AE5F4BE23335FFC1D" box="[151,603,879,900]" gridcol="0" gridrow="7" pageId="7" pageNumber="26">HW 15.8–15.9 19.4</th>
|
||
<td id="3F702005FFB6003AE7E3BE23326BFC1D" box="[640,879,879,900]" gridcol="1" gridrow="7" pageId="7" pageNumber="26">15.7–17.0; 16.5 ± 0.7</td>
|
||
<td id="3F702005FFB6003AE6E9BE233585FC1D" box="[906,1153,879,900]" gridcol="2" gridrow="7" pageId="7" pageNumber="26">16.8</td>
|
||
<td id="3F702005FFB6003AE1F6BE23349FFC1D" box="[1173,1435,879,900]" gridcol="3" gridrow="7" pageId="7" pageNumber="26">15.7–18.9; 17.2 ± 0.9</td>
|
||
</tr>
|
||
<tr id="7CA14979FFB6003AE5F4BED1349FFC2B" box="[151,1435,925,946]" gridrow="8" pageId="7" pageNumber="26">
|
||
<th id="3F702005FFB6003AE5F4BED1335FFC2B" box="[151,603,925,946]" gridcol="0" gridrow="8" pageId="7" pageNumber="26">EN 3.5–4.0 4.7</th>
|
||
<td id="3F702005FFB6003AE7E3BED1326BFC2B" box="[640,879,925,946]" gridcol="1" gridrow="8" pageId="7" pageNumber="26">4.0–4.4; 4.1 ± 0.2</td>
|
||
<td id="3F702005FFB6003AE6E9BED13585FC2B" box="[906,1153,925,946]" gridcol="2" gridrow="8" pageId="7" pageNumber="26">3.8–4.4</td>
|
||
<td id="3F702005FFB6003AE1F6BED1349FFC2B" box="[1173,1435,925,946]" gridcol="3" gridrow="8" pageId="7" pageNumber="26">3.2–4.3; 3.7 ± 0.3</td>
|
||
</tr>
|
||
<tr id="7CA14979FFB6003AE5F4BE87349FFC79" box="[151,1435,971,992]" gridrow="9" pageId="7" pageNumber="26">
|
||
<th id="3F702005FFB6003AE5F4BE87335FFC79" box="[151,603,971,992]" gridcol="0" gridrow="9" pageId="7" pageNumber="26">IN 4.9–5.0 5.8</th>
|
||
<td id="3F702005FFB6003AE7E3BE87326BFC79" box="[640,879,971,992]" gridcol="1" gridrow="9" pageId="7" pageNumber="26">5.4–6.1; 5.7 ± 0.4</td>
|
||
<td id="3F702005FFB6003AE6E9BE873585FC79" box="[906,1153,971,992]" gridcol="2" gridrow="9" pageId="7" pageNumber="26">5.6–5.7</td>
|
||
<td id="3F702005FFB6003AE1F6BE87349FFC79" box="[1173,1435,971,992]" gridcol="3" gridrow="9" pageId="7" pageNumber="26">4.3–6.2; 5.4 ± 0.4</td>
|
||
</tr>
|
||
<tr id="7CA14979FFB6003AE5F4BEB4349FFB94" box="[151,1435,1016,1037]" gridrow="10" pageId="7" pageNumber="26">
|
||
<th id="3F702005FFB6003AE5F4BEB4335FFB94" box="[151,603,1016,1037]" gridcol="0" gridrow="10" pageId="7" pageNumber="26">AL 22.9–23.7 26.6</th>
|
||
<td id="3F702005FFB6003AE7E3BEB4326BFB94" box="[640,879,1016,1037]" gridcol="1" gridrow="10" pageId="7" pageNumber="26">16.0–19.8; 18.2 ± 2.0</td>
|
||
<td id="3F702005FFB6003AE6E9BEB43585FB94" box="[906,1153,1016,1037]" gridcol="2" gridrow="10" pageId="7" pageNumber="26">20.9–22.4</td>
|
||
<td id="3F702005FFB6003AE1F6BEB4349FFB94" box="[1173,1435,1016,1037]" gridcol="3" gridrow="10" pageId="7" pageNumber="26">18.5–25.9; 22.2 ± 1.6</td>
|
||
</tr>
|
||
<tr id="7CA14979FFB6003AE5F4B96B349FFBA5" box="[151,1435,1063,1084]" gridrow="11" pageId="7" pageNumber="26">
|
||
<th id="3F702005FFB6003AE5F4B96B335FFBA5" box="[151,603,1063,1084]" gridcol="0" gridrow="11" pageId="7" pageNumber="26">PL 22.7–23.0 28.0</th>
|
||
<td id="3F702005FFB6003AE7E3B96B326BFBA5" box="[640,879,1063,1084]" gridcol="1" gridrow="11" pageId="7" pageNumber="26">20.1–21.3; 20.7 ± 0.6</td>
|
||
<td id="3F702005FFB6003AE6E9B96B3585FBA5" box="[906,1153,1063,1084]" gridcol="2" gridrow="11" pageId="7" pageNumber="26">22.2–23.5</td>
|
||
<td id="3F702005FFB6003AE1F6B96B349FFBA5" box="[1173,1435,1063,1084]" gridcol="3" gridrow="11" pageId="7" pageNumber="26">18.6–24.1; 21.5 ± 1.6</td>
|
||
</tr>
|
||
<tr id="7CA14979FFB6003AE5F4B929349FFB39" box="[151,1435,1125,1184]" gridrow="12" pageId="7" pageNumber="26" rowspan-1="1" rowspan-2="1" rowspan-3="1">
|
||
<th id="3F702005FFB6003AE5F4B929335FFB39" box="[151,603,1125,1184]" gridcol="0" gridrow="12" pageId="7" pageNumber="26">
|
||
Data from Böhme
|
||
<emphasis id="F0E59729FFB6FFC2E4EBB92930BCFBE3" box="[392,440,1125,1146]" italics="true" pageId="7" pageNumber="26">et al.</emphasis>
|
||
(2005) Tail tip missing in paratype male
|
||
</th>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption id="96EE1BB3FFB6FFC2E5F4BA663457F8D8" box="[151,1363,1834,1858]" httpUri="https://zenodo.org/record/198802/files/figure.png" pageId="7" pageNumber="26" targetBox="[375,1212,1236,1822]" targetPageId="7">
|
||
<paragraph id="C22E4B3BFFB6FFC2E5F4BA663457F8D8" blockId="7.[151,1363,1834,1858]" box="[151,1363,1834,1858]" pageId="7" pageNumber="26">
|
||
<emphasis id="F0E59729FFB6FFC2E5F4BA66301DF8DB" bold="true" box="[151,281,1834,1858]" pageId="7" pageNumber="26">FIGURE 4.</emphasis>
|
||
Map illustrating the type localities of
|
||
<taxonomicName id="059130B8FFB6FFC2E7D2BA663281F8D8" box="[689,901,1834,1857]" class="Amphibia" family="Salamandridae" genus="Tylototriton" kingdom="Animalia" order="Caudata" pageId="7" pageNumber="26" phylum="Chordata" rank="species" species="notialis" status="sp. nov.">
|
||
<emphasis id="F0E59729FFB6FFC2E7D2BA663281F8D8" box="[689,901,1834,1857]" italics="true" pageId="7" pageNumber="26">Tylototriton notialis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EBD62A52FFB6FFC2E6EFBA6632E5F8DB" box="[908,993,1834,1858]" pageId="7" pageNumber="26" rank="species">
|
||
<emphasis id="F0E59729FFB6FFC2E6EFBA6632E5F8DB" bold="true" box="[908,993,1834,1858]" pageId="7" pageNumber="26">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(star) and
|
||
<taxonomicName id="059130B8FFB6FFC2E137BA6635FCF8D8" box="[1108,1272,1834,1857]" class="Amphibia" family="Salamandridae" genus="Tylototriton" kingdom="Animalia" order="Caudata" pageId="7" pageNumber="26" phylum="Chordata" rank="species" species="vietnamensis">
|
||
<emphasis id="F0E59729FFB6FFC2E137BA6635FCF8D8" box="[1108,1272,1834,1857]" italics="true" pageId="7" pageNumber="26">T. vietnamensis</emphasis>
|
||
</taxonomicName>
|
||
(circle).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="C22E4B3BFFB6FFC2E5A5BA26342AF833" blockId="7.[151,1437,1898,2036]" pageId="7" pageNumber="26">
|
||
<emphasis id="F0E59729FFB6FFC2E5A5BA26302DF81D" bold="true" box="[198,297,1898,1924]" pageId="7" pageNumber="26">Larvae:</emphasis>
|
||
Two size classes of larvae were obtained in syntopy with the
|
||
<typeStatus id="1D2AF599FFB6FFC2E14FBA263593F81D" box="[1068,1175,1898,1924]" pageId="7" pageNumber="26" type="holotype">holotype</typeStatus>
|
||
: FMNH
|
||
<date id="B62F6DFBFFB6FFC2E066BA263458F81D" box="[1285,1372,1898,1924]" pageId="7" pageNumber="26" value="25-11-27">271125</date>
|
||
(one larva) with TTL 33.1, and FMNH
|
||
<date id="B62F6DFBFFB6FFC2E749BADC3385F833" box="[554,641,1936,1962]" pageId="7" pageNumber="26" value="29-11-27">271129</date>
|
||
(nine larvae) with TTL 17.8–20.9 (mean 19.4 ± S.D. 0.9).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8A8B18B0FFB6FFCDE5A5BAF9321FF86B" lastPageId="8" lastPageNumber="27" pageId="7" pageNumber="26" type="distribution">
|
||
<paragraph id="C22E4B3BFFB6FFCDE5A5BAF9336DFE2E" blockId="7.[151,1437,1898,2036]" lastBlockId="8.[151,1436,152,439]" lastPageId="8" lastPageNumber="27" pageId="7" pageNumber="26">
|
||
<emphasis id="F0E59729FFB6FFC2E5A5BAF93376F856" bold="true" box="[198,626,1973,1999]" pageId="7" pageNumber="26">Distribution and natural history:</emphasis>
|
||
The new species is currently known only from the
|
||
<typeStatus id="1D2AF599FFB6FFC2E063BAF93437F856" box="[1280,1331,1973,1999]" pageId="7" pageNumber="26">type</typeStatus>
|
||
locality, although it likely also occurs in adjacent
|
||
<collectingCountry id="BA860BABFFB6FFC2E7C1BA96320FF86D" box="[674,779,2010,2036]" name="Vietnam" pageId="7" pageNumber="26">Vietnam</collectingCountry>
|
||
given its proximity to the border. All were taken in evergreen mixed with deciduous and pine forest (
|
||
<figureCitation id="5AAA57BEFFB9FFCDE78EBDD4323CFF2B" box="[749,824,152,178]" captionStart="FIGURE 6" captionStartId="9.[151,255,1160,1184]" captionTargetBox="[151,1436,185,1148]" captionTargetId="figure@9.[151,1436,185,1149]" captionTargetPageId="9" captionText="FIGURE 6. Habitat of Tylototriton notialis at the type locality." httpUri="https://zenodo.org/record/198804/files/figure.png" pageId="8" pageNumber="27">Fig. 6</figureCitation>
|
||
). The
|
||
<typeStatus id="1D2AF599FFB9FFCDE6E5BDD432EBFF2B" box="[902,1007,152,178]" pageId="8" pageNumber="27" type="holotype">holotype</typeStatus>
|
||
and larvae were collected at
|
||
<date id="B62F6DFBFFB9FFCDE031BDD431D6FF4E" pageId="8" pageNumber="27" value="1940" valueMax="2042">1940– 2042</date>
|
||
h on the stream bottom under
|
||
<quantity id="0569E6DEFFB9FFCDE757BDF133ACFF4E" box="[564,680,189,215]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="8" pageNumber="27" unit="cm" value="20.0" valueMax="30.0" valueMin="10.0">10–30 cm</quantity>
|
||
of water (
|
||
<collectingCountry id="BA860BABFFB9FFCDE67EBDF13246FF4E" box="[797,834,189,215]" name="Philippines" pageId="8" pageNumber="27">pH</collectingCountry>
|
||
5.0, temperature 19.5ºC) in a
|
||
<quantity id="0569E6DEFFB9FFCDE1C3BDF135C9FF4E" box="[1184,1229,189,215]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="27" unit="m" value="3.0">3 m</quantity>
|
||
wide stream with slow current and sand and bedrock substrate, including leaf litter-filled potholes, just upstream from a
|
||
<quantity id="0569E6DEFFB9FFCDE00DBDAE3498FF65" box="[1390,1436,226,252]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="27" unit="m" value="3.0">3 m</quantity>
|
||
high chute over bedrock. The
|
||
<typeStatus id="1D2AF599FFB9FFCDE773BC44337FFEBB" box="[528,635,264,290]" pageId="8" pageNumber="27" type="paratype">paratype</typeStatus>
|
||
male was found at 1030 h,
|
||
<quantity id="0569E6DEFFB9FFCDE6BABC443522FEBB" box="[985,1062,264,290]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="27" unit="cm" value="30.0">30 cm</quantity>
|
||
above the ground on a
|
||
<quantity id="0569E6DEFFB9FFCDE033BC443498FEBB" box="[1360,1436,264,290]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="27" unit="cm" value="20.0">20 cm</quantity>
|
||
diameter log covered by another log, approximately
|
||
<quantity id="0569E6DEFFB9FFCDE79EBC61323DFEDE" box="[765,825,301,327]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="27" unit="m" value="20.0">20 m</quantity>
|
||
from a steep escarpment. The
|
||
<typeStatus id="1D2AF599FFB9FFCDE1C2BC6135D7FEDE" box="[1185,1235,301,327]" pageId="8" pageNumber="27">type</typeStatus>
|
||
locality lies near the former Ho Chi Minh Trail, and a large, unexploded bomb dropped by American forces during the war was embedded in the ground approximately
|
||
<quantity id="0569E6DEFFB9FFCDE70CBC343398FE0B" box="[623,668,376,402]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="27" unit="m" value="3.0">3 m</quantity>
|
||
from the
|
||
<typeStatus id="1D2AF599FFB9FFCDE672BC343272FE0B" box="[785,886,376,402]" pageId="8" pageNumber="27" type="paratype">paratype</typeStatus>
|
||
male (
|
||
<figureCitation id="5AAA57BEFFB9FFCDE6ABBC343515FE0B" box="[968,1041,376,402]" captionStart="FIGURE 7" captionStartId="10.[151,258,1874,1898]" captionTargetBox="[171,1414,194,1852]" captionTargetId="figure@10.[171,1415,194,1853]" captionTargetPageId="10" captionText="FIGURE 7. Microhabitat (log in foreground) of the paratype (FMNH 271120) of Tylototriton notialis. The arrow indicates an unexploded bomb dropped by American forces during the war." httpUri="https://zenodo.org/record/198805/files/figure.png" pageId="8" pageNumber="27">Fig. 7</figureCitation>
|
||
). The
|
||
<typeStatus id="1D2AF599FFB9FFCDE13EBC3435C6FE0B" box="[1117,1218,376,402]" pageId="8" pageNumber="27" type="paratype">paratype</typeStatus>
|
||
female was found during the day on the damp forest floor.
|
||
</paragraph>
|
||
<caption id="96EE1BB3FFB9FFCDE5F4B9CB31D4FB27" httpUri="https://zenodo.org/record/198803/files/figure.png" pageId="8" pageNumber="27" targetBox="[195,1391,493,1138]" targetPageId="8">
|
||
<paragraph id="C22E4B3BFFB9FFCDE5F4B9CB31D4FB27" blockId="8.[151,1436,1159,1214]" pageId="8" pageNumber="27">
|
||
<emphasis id="F0E59729FFB9FFCDE5F4B9CB301FFB06" bold="true" box="[151,283,1159,1183]" pageId="8" pageNumber="27">FIGURE 5.</emphasis>
|
||
Dorsal and lateral view of larva (FMNH 271129) of
|
||
<taxonomicName id="059130B8FFB9FFCDE63FB9C43530FB06" box="[860,1076,1160,1183]" class="Amphibia" family="Salamandridae" genus="Tylototriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="27" phylum="Chordata" rank="species" species="notialis">
|
||
<emphasis id="F0E59729FFB9FFCDE63FB9C43530FB06" box="[860,1076,1160,1183]" italics="true" pageId="8" pageNumber="27">Tylototriton notialis</emphasis>
|
||
</taxonomicName>
|
||
in preservative (total length 20.9 mm).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="C22E4B3BFFB9FFCDE5A6B9A133ECFACB" blockId="8.[151,1437,1261,2034]" pageId="8" pageNumber="27">
|
||
The testes of the
|
||
<typeStatus id="1D2AF599FFB9FFCDE4EFB9A130F6FA9E" box="[396,498,1261,1287]" pageId="8" pageNumber="27" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="1D2AF599FFB9FFCDE748B9A13394FA9E" box="[555,656,1261,1287]" pageId="8" pageNumber="27" type="paratype">paratype</typeStatus>
|
||
males are mature but not enlarged, suggesting these males were in their first reproductive season. The
|
||
<typeStatus id="1D2AF599FFB9FFCDE726B85E33A8FAB5" box="[581,684,1298,1324]" pageId="8" pageNumber="27" type="paratype">paratype</typeStatus>
|
||
female has enlarged oviducts, indicating that she had recently laid eggs. Eggs of the new species were not found.
|
||
</paragraph>
|
||
<paragraph id="C22E4B3BFFB9FFCDE5A6B81035FEF9E5" blockId="8.[151,1437,1261,2034]" pageId="8" pageNumber="27">
|
||
<emphasis id="F0E59729FFB9FFCDE5A6B8103072FAEF" bold="true" box="[197,374,1372,1398]" pageId="8" pageNumber="27">Conservation:</emphasis>
|
||
Overharvesting for traditional medicine and the international pet trade has been identified as a major threat to Asian salamandrids (
|
||
<bibRefCitation id="A60036CAFFB9FFCDE7E0B8CE3274FA05" author="Rowley" box="[643,880,1410,1436]" pageId="8" pageNumber="27" refString="Rowley, J., Brown, R., Bain, R., Kusrini, M., Inger, R., Stuart, B., Wogan, G., Neang, T., Chan-ard, T., Cao, T. T., Diesmos, A., Iskandar, D. T., Lau, M., Leong, M. T., Makchai, S., Nguyen, T. Q. & Phimmachak, S. (2010) Impending conservation crisis for Southeast Asian amphibians. Biology Letters, 6, 336 - 338." type="journal article" year="2010">
|
||
Rowley
|
||
<emphasis id="F0E59729FFB9FFCDE789B8CE3223FA02" box="[746,807,1410,1435]" italics="true" pageId="8" pageNumber="27">et al.</emphasis>
|
||
2010
|
||
</bibRefCitation>
|
||
). The formal description of
|
||
<taxonomicName id="059130B8FFB9FFCDE1A2B8CE30CBFA5B" authority="Stuart & Papenfuss 2002" authorityName="Stuart & Papenfuss" authorityYear="2002" class="Amphibia" family="Salamandridae" genus="Laotriton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="27" phylum="Chordata" rank="species" species="laoensis">
|
||
<emphasis id="F0E59729FFB9FFCDE1A2B8CE3498FA02" box="[1217,1436,1410,1435]" italics="true" pageId="8" pageNumber="27">Laotriton laoensis</emphasis>
|
||
(
|
||
<bibRefCitation id="A60036CAFFB9FFCDE5FCB8E430C3FA5B" author="Stuart" box="[159,455,1448,1474]" pageId="8" pageNumber="27" refString="Stuart, B. L. & Papenfuss, T. J. (2002) A new salamander of the genus Paramesotriton (Caudata: Salamandridae) from Laos. Journal of Herpetology, 36, 145 - 148." type="journal article" year="2002">Stuart & Papenfuss 2002</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, the first salamandrid known from
|
||
<collectingCountry id="BA860BABFFB9FFCDE60BB8E432A0FA5B" box="[872,932,1448,1474]" name="Laos" pageId="8" pageNumber="27">Laos</collectingCountry>
|
||
, inadvertently led to its exploitation for the international pet trade (
|
||
<bibRefCitation id="A60036CAFFB9FFCDE4D4B8813390FA7E" author="Stuart" box="[439,660,1485,1511]" pageId="8" pageNumber="27" refString="Stuart, B. L., Rhodin, A. G. J., Grismer, L. L. & Hansel, T. (2006) Scientific description can imperil species. Science, 312, 1137." type="journal article" year="2006">
|
||
Stuart
|
||
<emphasis id="F0E59729FFB9FFCDE768B881334DFA7F" box="[523,585,1485,1510]" italics="true" pageId="8" pageNumber="27">et al.</emphasis>
|
||
2006
|
||
</bibRefCitation>
|
||
). That species is now protected from commercial collecting by national legislation in
|
||
<collectingCountry id="BA860BABFFB9FFCDE4C6B8BE30E7F995" box="[421,483,1522,1548]" name="Laos" pageId="8" pageNumber="27">Laos</collectingCountry>
|
||
. It is hoped that the muted coloration and superficial similarity of
|
||
<taxonomicName id="059130B8FFB9FFCDE067B8BE3479F992" box="[1284,1405,1522,1547]" class="Amphibia" family="Salamandridae" genus="Tylototriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="27" phylum="Chordata" rank="species" species="notialis">
|
||
<emphasis id="F0E59729FFB9FFCDE067B8BE3479F992" box="[1284,1405,1522,1547]" italics="true" pageId="8" pageNumber="27">T. notialis</emphasis>
|
||
</taxonomicName>
|
||
to other members of the
|
||
<taxonomicName id="059130B8FFB9FFCDE4FCBB543346F9A8" box="[415,578,1560,1585]" class="Amphibia" family="Salamandridae" genus="Tylototriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="27" phylum="Chordata" rank="species" species="asperrimus">
|
||
<emphasis id="F0E59729FFB9FFCDE4FCBB543346F9A8" box="[415,578,1560,1585]" italics="true" pageId="8" pageNumber="27">T. asperrimus</emphasis>
|
||
</taxonomicName>
|
||
group will minimize its demand in the international pet trade. The new species’ occurrence in a remote area within Nakai-Nam Theun National Protected Area, which is one of Laos’ largest and best-funded national protected areas (
|
||
<bibRefCitation id="A60036CAFFB9FFCDE7B7BB2E3271F9E5" author="WMPA" box="[724,885,1634,1660]" pageId="8" pageNumber="27" refString="Watershed Management and Protection Authority (WMPA) (2005) Social and environmental management framework and 1 st operational plan. Nam Theun 2 WMPA, Vientiane, Laos." type="book chapter" year="2005">WMPA 2005</bibRefCitation>
|
||
), should also afford it protection.
|
||
</paragraph>
|
||
<paragraph id="C22E4B3BFFB9FFCDE5A6BBCB321FF86B" blockId="8.[151,1437,1261,2034]" pageId="8" pageNumber="27">
|
||
<emphasis id="F0E59729FFB9FFCDE5A6BBCB307CF938" bold="true" box="[197,376,1671,1697]" pageId="8" pageNumber="27">Comparisons:</emphasis>
|
||
<taxonomicName id="059130B8FFB9FFCDE4E3BBC43374F938" box="[384,624,1672,1697]" class="Amphibia" family="Salamandridae" genus="Tylototriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="27" phylum="Chordata" rank="species" species="notialis">
|
||
<emphasis id="F0E59729FFB9FFCDE4E3BBC43374F938" box="[384,624,1672,1697]" italics="true" pageId="8" pageNumber="27">Tylototriton notialis</emphasis>
|
||
</taxonomicName>
|
||
differs from all other species in the
|
||
<taxonomicName id="059130B8FFB9FFCDE14ABBC435CAF938" box="[1065,1230,1672,1697]" class="Amphibia" family="Salamandridae" genus="Tylototriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="27" phylum="Chordata" rank="species" species="asperrimus">
|
||
<emphasis id="F0E59729FFB9FFCDE14ABBC435CAF938" box="[1065,1230,1672,1697]" italics="true" pageId="8" pageNumber="27">T. asperrimus</emphasis>
|
||
</taxonomicName>
|
||
group by having bright orange coloration on the posterior end of the parotoids (parotoid coloration dark like dorsum in all other species).
|
||
<taxonomicName id="059130B8FFB9FFCDE46CBB9E3315F972" box="[271,529,1746,1771]" class="Amphibia" family="Salamandridae" genus="Tylototriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="27" phylum="Chordata" rank="species" species="notialis">
|
||
<emphasis id="F0E59729FFB9FFCDE46CBB9E3315F972" box="[271,529,1746,1771]" italics="true" pageId="8" pageNumber="27">Tylototriton notialis</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="059130B8FFB9FFCDE7DEBB9E3268F972" box="[701,876,1746,1771]" class="Amphibia" family="Salamandridae" genus="Tylototriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="27" phylum="Chordata" rank="species" species="asperrimus">
|
||
<emphasis id="F0E59729FFB9FFCDE7DEBB9E3268F972" box="[701,876,1746,1771]" italics="true" pageId="8" pageNumber="27">T. asperrimus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="059130B8FFB9FFCDE61DBB9E354CF972" box="[894,1096,1746,1771]" class="Amphibia" family="Salamandridae" genus="Tylototriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="27" phylum="Chordata" rank="species" species="wenxianensis">
|
||
<emphasis id="F0E59729FFB9FFCDE61DBB9E354CF972" box="[894,1096,1746,1771]" italics="true" pageId="8" pageNumber="27">T. wenxianensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="059130B8FFB9FFCDE139BB9E3417F972" box="[1114,1299,1746,1771]" class="Amphibia" family="Salamandridae" genus="Tylototriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="27" phylum="Chordata" rank="species" species="hainanensis">
|
||
<emphasis id="F0E59729FFB9FFCDE139BB9E3417F972" box="[1114,1299,1746,1771]" italics="true" pageId="8" pageNumber="27">T. hainanensis</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="059130B8FFB9FFCDE038BB9E3033F888" class="Amphibia" family="Salamandridae" genus="Tylototriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="27" phylum="Chordata" rank="species" species="vietnamensis">
|
||
<emphasis id="F0E59729FFB9FFCDE038BB9E3475F972" box="[1371,1393,1746,1771]" italics="true" pageId="8" pageNumber="27">T.</emphasis>
|
||
cf.
|
||
<emphasis id="F0E59729FFB9FFCDE5F4BBB43033F888" box="[151,311,1784,1809]" italics="true" pageId="8" pageNumber="27">vietnamensis</emphasis>
|
||
</taxonomicName>
|
||
from Cao Bang Province,
|
||
<collectingCountry id="BA860BABFFB9FFCDE7E4BBB433F7F88B" box="[647,755,1784,1810]" name="Vietnam" pageId="8" pageNumber="27">Vietnam</collectingCountry>
|
||
, by having bright orange coloration on the rib nodules (uniformly dark dorsum in
|
||
<taxonomicName id="059130B8FFB9FFCDE494BA5133A3F8AF" box="[503,679,1821,1846]" class="Amphibia" family="Salamandridae" genus="Tylototriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="27" phylum="Chordata" rank="species" species="asperrimus">
|
||
<emphasis id="F0E59729FFB9FFCDE494BA5133A3F8AF" box="[503,679,1821,1846]" italics="true" pageId="8" pageNumber="27">T. asperrimus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="059130B8FFB9FFCDE7DABA513280F8AF" box="[697,900,1821,1846]" class="Amphibia" family="Salamandridae" genus="Tylototriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="27" phylum="Chordata" rank="species" species="wenxianensis">
|
||
<emphasis id="F0E59729FFB9FFCDE7DABA513280F8AF" box="[697,900,1821,1846]" italics="true" pageId="8" pageNumber="27">T. wenxianensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="059130B8FFB9FFCDE6F5BA51354AF8AF" box="[918,1102,1821,1846]" class="Amphibia" family="Salamandridae" genus="Tylototriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="27" phylum="Chordata" rank="species" species="hainanensis">
|
||
<emphasis id="F0E59729FFB9FFCDE6F5BA51354AF8AF" box="[918,1102,1821,1846]" italics="true" pageId="8" pageNumber="27">T. hainanensis</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="059130B8FFB9FFCDE1FBBA51348DF8AF" box="[1176,1417,1821,1847]" class="Amphibia" family="Salamandridae" genus="Tylototriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="27" phylum="Chordata" rank="species" species="vietnamensis">
|
||
<emphasis id="F0E59729FFB9FFCDE1FBBA5135AAF8AF" box="[1176,1198,1821,1846]" italics="true" pageId="8" pageNumber="27">T.</emphasis>
|
||
cf.
|
||
<emphasis id="F0E59729FFB9FFCDE187BA51348DF8AF" box="[1252,1417,1821,1846]" italics="true" pageId="8" pageNumber="27">vietnamensis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="059130B8FFB9FFCDE5F4BA0E3085F8C2" box="[151,385,1858,1883]" class="Amphibia" family="Salamandridae" genus="Tylototriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="27" phylum="Chordata" rank="species" species="notialis">
|
||
<emphasis id="F0E59729FFB9FFCDE5F4BA0E3085F8C2" box="[151,385,1858,1883]" italics="true" pageId="8" pageNumber="27">Tylototriton notialis</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="059130B8FFB9FFCDE77EBA0E33D6F8C2" box="[541,722,1858,1883]" class="Amphibia" family="Salamandridae" genus="Tylototriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="27" phylum="Chordata" rank="species" species="vietnamensis">
|
||
<emphasis id="F0E59729FFB9FFCDE77EBA0E33D6F8C2" box="[541,722,1858,1883]" italics="true" pageId="8" pageNumber="27">T. vietnamensis</emphasis>
|
||
</taxonomicName>
|
||
by having glandular warts on most of the dorsal and ventral surfaces and very dark brown to black body coloration (
|
||
<taxonomicName id="059130B8FFB9FFCDE64DBA2432E7F818" box="[814,995,1896,1921]" class="Amphibia" family="Salamandridae" genus="Tylototriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="27" phylum="Chordata" rank="species" species="vietnamensis">
|
||
<emphasis id="F0E59729FFB9FFCDE64DBA2432E7F818" box="[814,995,1896,1921]" italics="true" pageId="8" pageNumber="27">T. vietnamensis</emphasis>
|
||
</taxonomicName>
|
||
with mostly smooth skin and gray to tan body coloration).
|
||
<taxonomicName id="059130B8FFB9FFCDE4C9BAC133ACF83F" box="[426,680,1933,1958]" class="Amphibia" family="Salamandridae" genus="Tylototriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="27" phylum="Chordata" rank="species" species="notialis">
|
||
<emphasis id="F0E59729FFB9FFCDE4C9BAC133ACF83F" box="[426,680,1933,1958]" italics="true" pageId="8" pageNumber="27">Tylototriton notialis</emphasis>
|
||
</taxonomicName>
|
||
further differs from
|
||
<taxonomicName id="059130B8FFB9FFCDE6D1BAC13563F83F" box="[946,1127,1933,1958]" class="Amphibia" family="Salamandridae" genus="Tylototriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="27" phylum="Chordata" rank="species" species="hainanensis">
|
||
<emphasis id="F0E59729FFB9FFCDE6D1BAC13563F83F" box="[946,1127,1933,1958]" italics="true" pageId="8" pageNumber="27">T. hainanensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="059130B8FFB9FFCDE118BAC1343AF83F" box="[1147,1342,1933,1958]" class="Amphibia" family="Salamandridae" genus="Tylototriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="27" phylum="Chordata" rank="species" species="vietnamensis">
|
||
<emphasis id="F0E59729FFB9FFCDE118BAC1343AF83F" box="[1147,1342,1933,1958]" italics="true" pageId="8" pageNumber="27">T. vietnamensis</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="059130B8FFB9FFCDE0E6BAC1303AF852" class="Amphibia" family="Salamandridae" genus="Tylototriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="27" phylum="Chordata" rank="species" species="wenxianensis">
|
||
<emphasis id="F0E59729FFB9FFCDE0E6BAC1303AF852" italics="true" pageId="8" pageNumber="27">T. wenxianensis</emphasis>
|
||
</taxonomicName>
|
||
by having knob-like rib nodules (slightly flattened rib nodules in
|
||
<taxonomicName id="059130B8FFB9FFCDE1E6BAFE3443F852" box="[1157,1351,1970,1995]" class="Amphibia" family="Salamandridae" genus="Tylototriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="27" phylum="Chordata" rank="species" species="vietnamensis">
|
||
<emphasis id="F0E59729FFB9FFCDE1E6BAFE3443F852" box="[1157,1351,1970,1995]" italics="true" pageId="8" pageNumber="27">T. vietnamensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="059130B8FFB9FFCDE0E6BAFE3027F868" class="Amphibia" family="Salamandridae" genus="Tylototriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="27" phylum="Chordata" rank="species" species="hainanensis">
|
||
<emphasis id="F0E59729FFB9FFCDE0E6BAFE3027F868" italics="true" pageId="8" pageNumber="27">T. hainanensis</emphasis>
|
||
</taxonomicName>
|
||
and indistinct nodules in
|
||
<taxonomicName id="059130B8FFB9FFCDE732BA94320DF868" box="[593,777,2008,2033]" class="Amphibia" family="Salamandridae" genus="Tylototriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="27" phylum="Chordata" rank="species" species="wenxianensis">
|
||
<emphasis id="F0E59729FFB9FFCDE732BA94320DF868" box="[593,777,2008,2033]" italics="true" pageId="8" pageNumber="27">T. wenxianensis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |