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<document id="3BE985E44DFE4A253D5DDB91CD4259FC" ID-DOI="10.3853/j.2201-4349.75.2023.1823" ID-ISSN="2201-4349" ID-Zenodo-Dep="7946210" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1683042549148" checkinUser="felipe" docAuthor="Mathis, Wayne N. &amp; Zatwarnicki, Tadeusz" docDate="2023" docId="AC5287BBFFF2FF93FDDCFAFD962369DE" docLanguage="en" docName="RecAustMus.75.1.1–23.pdf" docOrigin="Records of the Australian Museum (Rec. Aust. Mus.) 75 (1)" docSource="http://dx.doi.org/10.3853/j.2201-4349.75.2023.1823" docStyle="DocumentStyle:997BA24B490466A2F99594A367FA147F.3:RecAustMus.2019-.journal_article" docStyleId="997BA24B490466A2F99594A367FA147F" docStyleName="RecAustMus.2019-.journal_article" docStyleVersion="3" docTitle="Risa Becker 1907" docType="key" docVersion="5" lastPageNumber="13" masterDocId="506BFFC3FFF9FF9FFFFBFFFA937A6D2B" masterDocTitle="Phylogeny and Taxonomy of the Risa Genus-group (Diptera: Ephydridae), with Description of a New Genus from Australia" masterLastPageNumber="23" masterPageNumber="1" pageNumber="12" updateTime="1699497443028" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="F02F61F1FE2EC40A4A28F54EA4601321">Phylogeny and Taxonomy of the Risa Genus-group (Diptera: Ephydridae), with Description of a New Genus from Australia</mods:title>
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<mods:namePart id="BCB804CEF30FE6173C2A2BCE9E6B44EC">Mathis, Wayne N.</mods:namePart>
<mods:affiliation id="17964ED8248ED82953BE11D773587EBC">Department of Entomology, PO BOX 37012, MRC 169, Smithsonian Institution, Washington, DC 20013 - 7012, United States of America</mods:affiliation>
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<mods:namePart id="87F3C725D7A9CFDBE2D00F1879FB3501">Zatwarnicki, Tadeusz</mods:namePart>
<mods:affiliation id="1E3C6A38FEDAD0D13390A3135DE2E1C6">Institute of Biology, Opole University, ul. Oleska 22, 45 - 052 Opole, Poland</mods:affiliation>
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<mods:title id="1C4EB67E88576089E425D255A43A5249">Records of the Australian Museum</mods:title>
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<mods:date id="08664D9357B3996E5003EEFE0913B3C8">2023</mods:date>
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<mods:title id="0C551338BB8BE246500C481F3C1790D3">Rec. Aust. Mus.</mods:title>
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<mods:number id="2D62AE4BE68474F73B0D381C38697D6B">2023-03-15</mods:number>
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<mods:number id="687AF81294C09485E11B33A15F239968">75</mods:number>
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<mods:url id="3D616074291EE4C41E0DDC9B4F6FC7FA">http://dx.doi.org/10.3853/j.2201-4349.75.2023.1823</mods:url>
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<treatment id="AC5287BBFFF2FF93FDDCFAFD962369DE" ID-DOI="http://doi.org/10.5281/zenodo.7946327" ID-Zenodo-Dep="7946327" LSID="urn:lsid:plazi:treatment:AC5287BBFFF2FF93FDDCFAFD962369DE" httpUri="http://treatment.plazi.org/id/AC5287BBFFF2FF93FDDCFAFD962369DE" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<subSubSection id="6CE16526FFF2FF94FDDCFAFD909A680D" box="[551,992,1286,1318]" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="244436ADFFF2FF94FDDCFAFD909A680D" blockId="11.[219,1323,1286,1367]" box="[551,992,1286,1318]" pageId="11" pageNumber="12">
<heading id="7F0C81C1FFF2FF94FDDCFAFD909A680D" bold="true" box="[551,992,1286,1318]" centered="true" fontSize="12" level="2" pageId="11" pageNumber="12" reason="2">
<emphasis id="168FEABFFFF2FF94FDDCFAFD909A680D" bold="true" box="[551,992,1286,1318]" pageId="11" pageNumber="12">
Key to species of
<taxonomicName id="E3FB4D2EFFF2FF94FCD7FAFD909A680D" ID-CoL="63S35" ID-ENA="2952857" authority="Becker" authorityName="Becker" authorityYear="1907" box="[812,992,1287,1318]" class="Insecta" family="Ephydridae" genus="Risa" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="168FEABFFFF2FF94FCD7FAFD9017680D" bold="true" box="[812,877,1287,1318]" italics="true" pageId="11" pageNumber="12">Risa</emphasis>
Becker
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="6CE16526FFF2FF93FF20FAC5962369DE" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="key">
<keyStep id="9F0F2E08FFF2FF94FF20FAC596516857" pageId="11" pageNumber="12">
<paragraph id="244436ADFFF2FF94FF20FAC59651687C" blockId="11.[219,1323,1286,1367]" box="[219,1323,1343,1367]" pageId="11" pageNumber="12">
<keyLead id="9F0A9598FFF2FF94FF20FAC59651687C" box="[219,1323,1343,1367]" pageId="11" pageNumber="12">1 Palpus yellow ............................................................................................................................ 2</keyLead>
</paragraph>
<paragraph id="244436ADFFF2FF94FF20FA9F96516857" blockId="11.[219,1323,1380,1484]" box="[219,1323,1380,1404]" pageId="11" pageNumber="12">
<keyLead id="9F0A9598FFF2FF94FF20FA9F96516857" box="[219,1323,1380,1404]" pageId="11" pageNumber="12">—— Palpus black ............................................................................................................................... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="9F0F2E08FFF2FF94FF20FA6D96506B24" pageId="11" pageNumber="12">
<paragraph id="244436ADFFF2FF94FF20FA6D965068E7" blockId="11.[219,1323,1380,1484]" pageId="11" pageNumber="12">
<keyLead id="9F0A9598FFF2FF94FF20FA6D965068E7" pageId="11" pageNumber="12">
2 Forecoxa black. Ocellar triangle short, slightly protruding beyond middle of frons. Postocellar setae present ....................................
<taxonomicName id="E3FB4D2EFFF2FF94FBB2FA4F965068E7" ID-CoL="4T52C" authority="Ozerov" authorityName="Ozerov" authorityYear="1992" box="[1097,1322,1460,1484]" class="Insecta" family="Ephydridae" genus="Risa" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="flavipalpis">
<emphasis id="168FEABFFFF2FF94FBB2FA4F97AC68E7" box="[1097,1238,1460,1484]" italics="true" pageId="11" pageNumber="12">R. flavipalpis</emphasis>
Ozerov
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="244436ADFFF2FF94FF20FA2096506B24" blockId="11.[219,1323,1497,1631]" pageId="11" pageNumber="12">
<keyLead id="9F0A9598FFF2FF94FF20FA2096506B24" pageId="11" pageNumber="12">
—— Forecoxa yellow. Ocellar triangle long, reaching lunule. Postocellar setae absent ..............................................................
<taxonomicName id="E3FB4D2EFFF2FF94FC48FA0296506B24" ID-CoL="9HVWP" authority="Krivosheina &amp; Ozerov" authorityName="Krivosheina &amp; Ozerov" authorityYear="2019" box="[947,1322,1527,1551]" class="Insecta" family="Ephydridae" genus="Risa" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="flavicoxa">
<emphasis id="168FEABFFFF2FF94FC48FA02974B6B24" box="[947,1073,1527,1551]" italics="true" pageId="11" pageNumber="12">R. flavicoxa</emphasis>
Krivosheina &amp; Ozerov
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9F0F2E08FFF2FF94FF20F9D096516BD1" pageId="11" pageNumber="12">
<paragraph id="244436ADFFF2FF94FF20F9D096516B74" blockId="11.[219,1323,1497,1631]" pageId="11" pageNumber="12">
<keyLead id="9F0A9598FFF2FF94FF20F9D096516B74" pageId="11" pageNumber="12">
3 Fronto-orbital setae 3 (rarely 2 or 4), proclinate, small, hair-like, equal or subequal in size (
<figureCitation id="BCC02A28FFF2FF94FDB7F9BD91E56B74" box="[588,671,1607,1631]" captionStart="Figures 20–25" captionStartId="10.[166,243,1096,1118]" captionTargetBox="[167,1460,151,1083]" captionTargetId="figure-249@10.[167,1467,151,1084]" captionTargetPageId="10" captionText="Figures 20–25. Line drawing of frons and associated setae, dorsal view. (20) Notorisa mcalpinei sp. nov. (21) Risa longirostris Becker. (22) R. asiatica Ozerov. (23–25) Line drawing of head, lateral view. (23) R. longirostris Becker. (24) R. brevicornis Papp. (25) R. longicornuta Papp." figureDoi="http://doi.org/10.5281/zenodo.7946222" httpUri="https://zenodo.org/record/7946222/files/figure.png" pageId="11" pageNumber="12">Figs 22</figureCitation>
,
<figureCitation id="BCC02A28FFF2FF94FD51F9BD91956B74" box="[682,751,1607,1631]" captionStart="Figures 38–41" captionStartId="15.[121,198,1628,1650]" captionTargetBox="[121,1421,166,1586]" captionTargetId="figure-98@15.[121,1421,151,1618]" captionTargetPageId="15" captionText="Figures 38–41. Photographs of Risa brevirostris sp. nov. (38–39) and R. longirostris Becker (40–41). (38) Head, anterior view. (39) Same, dorso-oblique view. (40) Habitus, lateral view. (41) Head, antero-oblique view. Scale bars = 0.1 mm (38–39), = 0.2 mm (40–41)." figureDoi="http://doi.org/10.5281/zenodo.7946228" httpUri="https://zenodo.org/record/7946228/files/figure.png" pageId="11" pageNumber="12">38, 39</figureCitation>
,
<figureCitation id="BCC02A28FFF2FF94FD01F9BD90456B74" box="[762,831,1607,1631]" captionStart="Figures 46–50" captionStartId="17.[121,198,1444,1466]" captionTargetBox="[136,1421,151,1430]" captionTargetId="figure-142@17.[121,1421,151,1430]" captionTargetPageId="17" captionText="Figures 46–50. Line drawing and photographs of Risa nettae sp. nov. (46) Head, lateral view. (47) Ventral receptacle, lateral view (microscopic photograph). (48) Ventral receptacle, posterior view (microscopic photograph). (49) Head, anterior view.(50) Same, laterooblique view. Scale bars = 0.05 mm (47–48), = 0.2 mm (49–50)." figureDoi="http://doi.org/10.5281/zenodo.7946232" httpUri="https://zenodo.org/record/7946232/files/figure.png" pageId="11" pageNumber="12">49, 50</figureCitation>
,
<figureCitation id="BCC02A28FFF2FF94FCB1F9BD90F46B74" box="[842,910,1607,1631]" captionStart="Figures 57–60" captionStartId="20.[167,244,1319,1341]" captionTargetBox="[166,1466,151,1307]" captionTargetId="figure-11@20.[166,1466,151,1307]" captionTargetPageId="20" captionText="Figures 57–60. Photographs of Risa kotrbae sp. nov. (57) Head, anterior view. (58) Head and thorax, lateral view. (59) Same, dorsal view. (60) Same, dorsolatero-oblique view. Scale bars = 0.2 mm (57), = 0.5 mm (58–60)." figureDoi="http://doi.org/10.5281/zenodo.7946236" httpUri="https://zenodo.org/record/7946236/files/figure.png" pageId="11" pageNumber="12">57–60</figureCitation>
) ........................................................ 4
</keyLead>
</paragraph>
<paragraph id="244436ADFFF2FF94FF20F99796516BD1" blockId="11.[219,1323,1644,1786]" pageId="11" pageNumber="12">
<keyLead id="9F0A9598FFF2FF94FF20F99796516BD1" pageId="11" pageNumber="12">
—— Anterior fronto-orbital seta about 2–3 times longer than middle fronto-orbital seta, inclinate; posterior fronto-orbital seta usually reclinate, occasionally proclinate, usually as long as anterior fronto-orbital setae, occasionally as long as middle fronto-orbital seta (
<figureCitation id="BCC02A28FFF2FF94FDFFF91B912D6BD1" box="[516,599,1761,1786]" captionStart="Figures 20–25" captionStartId="10.[166,243,1096,1118]" captionTargetBox="[167,1460,151,1083]" captionTargetId="figure-249@10.[167,1467,151,1084]" captionTargetPageId="10" captionText="Figures 20–25. Line drawing of frons and associated setae, dorsal view. (20) Notorisa mcalpinei sp. nov. (21) Risa longirostris Becker. (22) R. asiatica Ozerov. (23–25) Line drawing of head, lateral view. (23) R. longirostris Becker. (24) R. brevicornis Papp. (25) R. longicornuta Papp." figureDoi="http://doi.org/10.5281/zenodo.7946222" httpUri="https://zenodo.org/record/7946222/files/figure.png" pageId="11" pageNumber="12">Figs 21</figureCitation>
,
<figureCitation id="BCC02A28FFF2FF94FD99F91B91DC6BD1" box="[610,678,1761,1786]" captionStart="Figures 26–33" captionStartId="13.[121,198,1368,1390]" captionTargetBox="[121,1420,151,1355]" captionTargetId="figure-9@13.[121,1421,151,1357]" captionTargetPageId="13" captionText="Figures 26–33. Photographs of Risa longirostris Becker (26–27, 29–31, 33) and Risa brevirostris sp. nov. (28, 32). (26) Head, anterooblique view. (27) Same, anterior view. (28) Proboscis, lateral view. (29) Rostrum, anterior view (microscopic photograph). (30) Same, lateral view. (31) Labellum, posterior view. (32) Labellum, anterior view. (33) Terminal female abdominal segments, anterior view. Scale bars = 0.2 mm (26–27), = 0.1 mm (28–30, 33), = 0.05 mm (31–32)." figureDoi="http://doi.org/10.5281/zenodo.7946224" httpUri="https://zenodo.org/record/7946224/files/figure.png" pageId="11" pageNumber="12">26, 27</figureCitation>
) ........................................................................................... 9
</keyLead>
</paragraph>
</keyStep>
<paragraph id="244436ADFFF2FF94FF20F8EE96516A62" blockId="11.[219,1323,1812,1932]" pageId="11" pageNumber="12">
<keyLead id="9F0A9598FFF2FF94FF20F8EE96516A62" pageId="11" pageNumber="12">
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4 Labellum about twice length of ventral margin of head in lateral view (
<figureCitation id="BCC02A28FFF2FF94FE83F8CB92B16A61" box="[376,459,1841,1866]" captionStart="Figures 20–25" captionStartId="10.[166,243,1096,1118]" captionTargetBox="[167,1460,151,1083]" captionTargetId="figure-249@10.[167,1467,151,1084]" captionTargetPageId="10" captionText="Figures 20–25. Line drawing of frons and associated setae, dorsal view. (20) Notorisa mcalpinei sp. nov. (21) Risa longirostris Becker. (22) R. asiatica Ozerov. (23–25) Line drawing of head, lateral view. (23) R. longirostris Becker. (24) R. brevicornis Papp. (25) R. longicornuta Papp." figureDoi="http://doi.org/10.5281/zenodo.7946222" httpUri="https://zenodo.org/record/7946222/files/figure.png" pageId="11" pageNumber="12">Figs 24</figureCitation>
,
<figureCitation id="BCC02A28FFF2FF94FE2DF8CB92886A62" box="[470,498,1841,1865]" captionStart="Figures 46–50" captionStartId="17.[121,198,1444,1466]" captionTargetBox="[136,1421,151,1430]" captionTargetId="figure-142@17.[121,1421,151,1430]" captionTargetPageId="17" captionText="Figures 46–50. Line drawing and photographs of Risa nettae sp. nov. (46) Head, lateral view. (47) Ventral receptacle, lateral view (microscopic photograph). (48) Ventral receptacle, posterior view (microscopic photograph). (49) Head, anterior view.(50) Same, laterooblique view. Scale bars = 0.05 mm (47–48), = 0.2 mm (49–50)." figureDoi="http://doi.org/10.5281/zenodo.7946232" httpUri="https://zenodo.org/record/7946232/files/figure.png" pageId="11" pageNumber="12">46</figureCitation>
) ...................................................................................................................... 5
</keyStep>
</keyLead>
</paragraph>
<paragraph id="244436ADFFF2FF93FF20F8AD96226DC1" blockId="11.[219,1323,1812,1932]" lastBlockId="12.[265,1368,151,234]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<keyLead id="9F0A9598FFF2FF93FF20F8AD96226DC1" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
—— Labellum about 1–1.5 times length of ventral margin of head (
<figureCitation id="BCC02A28FFF2FF94FEC6F88E92C26AA7" box="[317,440,1908,1932]" captionStart="Figures 20–25" captionStartId="10.[166,243,1096,1118]" captionTargetBox="[167,1460,151,1083]" captionTargetId="figure-249@10.[167,1467,151,1084]" captionTargetPageId="10" captionText="Figures 20–25. Line drawing of frons and associated setae, dorsal view. (20) Notorisa mcalpinei sp. nov. (21) Risa longirostris Becker. (22) R. asiatica Ozerov. (23–25) Line drawing of head, lateral view. (23) R. longirostris Becker. (24) R. brevicornis Papp. (25) R. longicornuta Papp." figureDoi="http://doi.org/10.5281/zenodo.7946222" httpUri="https://zenodo.org/record/7946222/files/figure.png" pageId="11" pageNumber="12">Figs 23, 25</figureCitation>
,
<figureCitation id="BCC02A28FFF2FF94FE38F88E91726AA7" box="[451,520,1908,1932]" captionStart="Figures 38–41" captionStartId="15.[121,198,1628,1650]" captionTargetBox="[121,1421,166,1586]" captionTargetId="figure-98@15.[121,1421,151,1618]" captionTargetPageId="15" captionText="Figures 38–41. Photographs of Risa brevirostris sp. nov. (38–39) and R. longirostris Becker (40–41). (38) Head, anterior view. (39) Same, dorso-oblique view. (40) Habitus, lateral view. (41) Head, antero-oblique view. Scale bars = 0.1 mm (38–39), = 0.2 mm (40–41)." figureDoi="http://doi.org/10.5281/zenodo.7946228" httpUri="https://zenodo.org/record/7946228/files/figure.png" pageId="11" pageNumber="12">40, 41</figureCitation>
,
<figureCitation id="BCC02A28FFF2FF94FDE8F88E91556AA7" box="[531,559,1908,1932]" captionStart="Figures 57–60" captionStartId="20.[167,244,1319,1341]" captionTargetBox="[166,1466,151,1307]" captionTargetId="figure-11@20.[166,1466,151,1307]" captionTargetPageId="20" captionText="Figures 57–60. Photographs of Risa kotrbae sp. nov. (57) Head, anterior view. (58) Head and thorax, lateral view. (59) Same, dorsal view. (60) Same, dorsolatero-oblique view. Scale bars = 0.2 mm (57), = 0.5 mm (58–60)." figureDoi="http://doi.org/10.5281/zenodo.7946236" httpUri="https://zenodo.org/record/7946236/files/figure.png" pageId="11" pageNumber="12">58</figureCitation>
) ............................................................................................................ 8
<keyStep id="9F0F2E08FFF5FF93FEF2FF6D96226DC1" pageId="12" pageNumber="13">
5 Prescutellar acrostichal setae well developed; scutellar disk bearing scattered, sparse setulae; setulae along costal margin numerous; large species (length about
<quantity id="E3039B48FFF5FF93FD05FF2890476DC1" box="[766,829,210,234]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="13" unit="mm" value="2.0">2 mm</quantity>
) ...............................................
<taxonomicName id="E3FB4D2EFFF5FF93FB6AFF2996226DC1" ID-CoL="4T52B" authority="Ozerov" authorityName="Ozerov" box="[1169,1368,210,234]" class="Insecta" family="Ephydridae" genus="Risa" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="asiatica">
<emphasis id="168FEABFFFF5FF93FB6AFF2996796DC1" box="[1169,1283,211,234]" italics="true" pageId="12" pageNumber="13">R. asiatica</emphasis>
Ozerov
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
<paragraph id="244436ADFFF5FF93FEF2FF0296226C61" blockId="12.[265,1368,247,330]" pageId="12" pageNumber="13">
<keyLead id="9F0A9598FFF5FF93FEF2FF0296226C61" pageId="12" pageNumber="13">
—— Prescutellar acrostichal setae greatly reduced or lacking (
<figureCitation id="BCC02A28FFF5FF93FC56FF0D90836C3B" box="[941,1017,247,272]" captionStart="Figures 34–37" captionStartId="14.[167,244,1531,1553]" captionTargetBox="[167,1467,151,1487]" captionTargetId="figure-11@14.[167,1467,151,1514]" captionTargetPageId="14" captionText="Figures 34–37. Line drawing and photographs of Risa. (34) Terminal female abdominal segments of R. brevirostris sp. nov., anterior view. (35) Habitus of R. brevicornis (Papp), antero-oblique view. (36) Head of R. brevirostris sp. nov., lateral view. (37) Habitus of R. salsolae (Mathis &amp; Zatwarnicki), antero-oblique view. Scale bars = 0.1 mm (34), = 0.2 mm (35, 37)." figureDoi="http://doi.org/10.5281/zenodo.7946226" httpUri="https://zenodo.org/record/7946226/files/figure.png" pageId="12" pageNumber="13">Fig. 37</figureCitation>
,
<figureCitation id="BCC02A28FFF5FF93FE98FEEF92D06C06" box="[355,426,277,301]" captionStart="Figures 57–60" captionStartId="20.[167,244,1319,1341]" captionTargetBox="[166,1466,151,1307]" captionTargetId="figure-11@20.[166,1466,151,1307]" captionTargetPageId="20" captionText="Figures 57–60. Photographs of Risa kotrbae sp. nov. (57) Head, anterior view. (58) Head and thorax, lateral view. (59) Same, dorsal view. (60) Same, dorsolatero-oblique view. Scale bars = 0.2 mm (57), = 0.5 mm (58–60)." figureDoi="http://doi.org/10.5281/zenodo.7946236" httpUri="https://zenodo.org/record/7946236/files/figure.png" pageId="12" pageNumber="13">59–60</figureCitation>
); scutellum lacking setulae; setulae along costal margin sparse; smaller species (length about
<quantity id="E3039B48FFF5FF93FD0AFEC890256C61" box="[753,863,306,330]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="12" pageNumber="13" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">1–1.5 mm</quantity>
) ...................................................................... 6
</keyLead>
</paragraph>
<keyStep id="9F0F2E08FFF5FF93FEF2FE9F96226F3C" pageId="12" pageNumber="13">
<paragraph id="244436ADFFF5FF93FEF2FE9F96236C9C" blockId="12.[265,1369,357,439]" pageId="12" pageNumber="13">
<keyLead id="9F0A9598FFF5FF93FEF2FE9F96236C9C" pageId="12" pageNumber="13">
6 Scutum distinctly microtomentose; setulae along costal margin more numerous and denser (24–34 on distalmost section). Halter knob with large dark spot ..........................................................
<taxonomicName id="E3FB4D2EFFF5FF93FBF1FE5A97A66C9C" ID-CoL="95Y6" authority="Papp" authorityName="Papp." authorityYear="2023" baseAuthorityName="Mathis &amp; Zatwarnicki" baseAuthorityYear="2023" box="[1034,1244,415,439]" class="Insecta" family="Ephydridae" genus="Achaetorisa" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="brevicornis" status="comb. nov.">
<emphasis id="168FEABFFFF5FF93FBF1FE5A97E56C9C" box="[1034,1183,415,439]" italics="true" pageId="12" pageNumber="13">R. brevicornis</emphasis>
Papp
</taxonomicName>
<taxonomicNameLabel id="0DBC57C4FFF5FF93FB18FE6596236C9C" box="[1251,1369,415,439]" pageId="12" pageNumber="13" rank="species">comb. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="244436ADFFF5FF93FEF2FE3F96226F3C" blockId="12.[265,1368,453,535]" pageId="12" pageNumber="13">
<keyLead id="9F0A9598FFF5FF93FEF2FE3F96226F3C" pageId="12" pageNumber="13">—— Scutum indistinctly microtomentose; setulae along costal margin fewer and sparser (11–16 on distalmost section). Halter knob entirely yellow or with large dark spot ............................................................................. 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="9F0F2E08FFF5FF93FEF2FDC996236FCF" pageId="12" pageNumber="13">
<paragraph id="244436ADFFF5FF93FEF2FDC996236F4C" blockId="12.[265,1369,562,616]" pageId="12" pageNumber="13">
<keyLead id="9F0A9598FFF5FF93FEF2FDC996236F4C" pageId="12" pageNumber="13">
7 Antenna and arista predominantly brownish (
<figureCitation id="BCC02A28FFF5FF93FCA9FDC890DE6F61" box="[850,932,562,586]" captionStart="Figures 34–37" captionStartId="14.[167,244,1531,1553]" captionTargetBox="[167,1467,151,1487]" captionTargetId="figure-11@14.[167,1467,151,1514]" captionTargetPageId="14" captionText="Figures 34–37. Line drawing and photographs of Risa. (34) Terminal female abdominal segments of R. brevirostris sp. nov., anterior view. (35) Habitus of R. brevicornis (Papp), antero-oblique view. (36) Head of R. brevirostris sp. nov., lateral view. (37) Habitus of R. salsolae (Mathis &amp; Zatwarnicki), antero-oblique view. Scale bars = 0.1 mm (34), = 0.2 mm (35, 37)." figureDoi="http://doi.org/10.5281/zenodo.7946226" httpUri="https://zenodo.org/record/7946226/files/figure.png" pageId="12" pageNumber="13">Fig. 37</figureCitation>
). Halter knob yellow ......................................................
<taxonomicName id="E3FB4D2EFFF5FF93FC99FDAA97A66F4C" ID-CoL="8GTCH" authority="(Mathis &amp; Zatwarnicki)" authorityName="Mathis &amp; Zatwarnicki" authorityYear="2023" baseAuthorityName="Mathis &amp; Zatwarnicki" baseAuthorityYear="2023" box="[866,1244,591,616]" class="Insecta" family="Ephydridae" genus="Achaetorisa" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="salsolae" status="comb. nov.">
<emphasis id="168FEABFFFF5FF93FC99FDAA90AC6F4C" box="[866,982,591,615]" italics="true" pageId="12" pageNumber="13">R. salsolae</emphasis>
(Mathis &amp; Zatwarnicki)
</taxonomicName>
<taxonomicNameLabel id="0DBC57C4FFF5FF93FB18FDB596236F4C" box="[1251,1369,591,615]" pageId="12" pageNumber="13" rank="species">comb. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="244436ADFFF5FF93FEF2FD8F96236FCF" blockId="12.[265,1369,629,741]" pageId="12" pageNumber="13">
<keyLead id="9F0A9598FFF5FF93FEF2FD8F96236FCF" pageId="12" pageNumber="13">
—— Antenna predominantly or entirely yellow, scape and pedicel and basal flagellomere anteroapically occasionally darker, arista yellow to brown (
<figureCitation id="BCC02A28FFF5FF93FD97FD5591946FEC" box="[620,750,687,712]" captionStart="Figures 46–50" captionStartId="17.[121,198,1444,1466]" captionTargetBox="[136,1421,151,1430]" captionTargetId="figure-142@17.[121,1421,151,1430]" captionTargetPageId="17" captionText="Figures 46–50. Line drawing and photographs of Risa nettae sp. nov. (46) Head, lateral view. (47) Ventral receptacle, lateral view (microscopic photograph). (48) Ventral receptacle, posterior view (microscopic photograph). (49) Head, anterior view.(50) Same, laterooblique view. Scale bars = 0.05 mm (47–48), = 0.2 mm (49–50)." figureDoi="http://doi.org/10.5281/zenodo.7946232" httpUri="https://zenodo.org/record/7946232/files/figure.png" pageId="12" pageNumber="13">Figs 49, 50</figureCitation>
). Halter knob with large dark spot ...........................................................................................................
<taxonomicName id="E3FB4D2EFFF5FF93FB5AFD3497856FCE" authority="Mathis &amp; Zatwarnicki, 2023" authorityName="Mathis &amp; Zatwarnicki SMNH" authorityYear="2023" box="[1185,1279,718,741]" class="Insecta" family="Ephydridae" genus="Risa" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="nettae" status="sp. nov.">
<emphasis id="168FEABFFFF5FF93FB5AFD3497856FCE" box="[1185,1279,718,741]" italics="true" pageId="12" pageNumber="13">R. nettae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0DBC57C4FFF5FF93FAFDFD3796236FCF" box="[1286,1369,717,740]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9F0F2E08FFF5FF93FEF2FCFA9623690C" pageId="12" pageNumber="13">
<paragraph id="244436ADFFF5FF93FEF2FCFA96236EA7" blockId="12.[265,1369,767,909]" pageId="12" pageNumber="13">
<keyLead id="9F0A9598FFF5FF93FEF2FCFA96236EA7" pageId="12" pageNumber="13">
8 Face nearly flat, without prominent shallow/low carina (
<figureCitation id="BCC02A28FFF5FF93FC58FD05908A6E3C" box="[931,1008,767,791]" captionStart="Figures 57–60" captionStartId="20.[167,244,1319,1341]" captionTargetBox="[166,1466,151,1307]" captionTargetId="figure-11@20.[166,1466,151,1307]" captionTargetPageId="20" captionText="Figures 57–60. Photographs of Risa kotrbae sp. nov. (57) Head, anterior view. (58) Head and thorax, lateral view. (59) Same, dorsal view. (60) Same, dorsolatero-oblique view. Scale bars = 0.2 mm (57), = 0.5 mm (58–60)." figureDoi="http://doi.org/10.5281/zenodo.7946236" httpUri="https://zenodo.org/record/7946236/files/figure.png" pageId="12" pageNumber="13">Fig. 58</figureCitation>
); antenna elongate, basal flagellomere ratio 1.75; scutum distinctly microtomentose (
<figureCitation id="BCC02A28FFF5FF93FDE1FCC091E86E79" box="[538,658,826,850]" captionStart="Figures 57–60" captionStartId="20.[167,244,1319,1341]" captionTargetBox="[166,1466,151,1307]" captionTargetId="figure-11@20.[166,1466,151,1307]" captionTargetPageId="20" captionText="Figures 57–60. Photographs of Risa kotrbae sp. nov. (57) Head, anterior view. (58) Head and thorax, lateral view. (59) Same, dorsal view. (60) Same, dorsolatero-oblique view. Scale bars = 0.2 mm (57), = 0.5 mm (58–60)." figureDoi="http://doi.org/10.5281/zenodo.7946236" httpUri="https://zenodo.org/record/7946236/files/figure.png" pageId="12" pageNumber="13">Figs 59, 60</figureCitation>
); lower portion of frons, parafacial and lateral margin of face conspicuously and finely whitish to silvery microtomentose (
<figureCitation id="BCC02A28FFF5FF93FD97FC8F919D6EA6" box="[620,743,885,909]" captionStart="Figures 57–60" captionStartId="20.[167,244,1319,1341]" captionTargetBox="[166,1466,151,1307]" captionTargetId="figure-11@20.[166,1466,151,1307]" captionTargetPageId="20" captionText="Figures 57–60. Photographs of Risa kotrbae sp. nov. (57) Head, anterior view. (58) Head and thorax, lateral view. (59) Same, dorsal view. (60) Same, dorsolatero-oblique view. Scale bars = 0.2 mm (57), = 0.5 mm (58–60)." figureDoi="http://doi.org/10.5281/zenodo.7946236" httpUri="https://zenodo.org/record/7946236/files/figure.png" pageId="12" pageNumber="13">Figs 57, 60</figureCitation>
) ............................................................
<taxonomicName id="E3FB4D2EFFF5FF93FB6BFC8C97856EA6" authority="Mathis &amp; Zatwarnicki, 2023" authorityName="Mathis &amp; Zatwarnicki SMNH" authorityYear="2023" box="[1168,1279,885,909]" class="Insecta" family="Ephydridae" genus="Risa" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="kotrbae" status="sp. nov.">
<emphasis id="168FEABFFFF5FF93FB6BFC8C97856EA6" box="[1168,1279,885,909]" italics="true" pageId="12" pageNumber="13">R. kotrbae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0DBC57C4FFF5FF93FAFDFC8F96236EA7" box="[1286,1369,885,908]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="244436ADFFF5FF93FEF2FC619623690C" blockId="12.[265,1369,922,1064]" pageId="12" pageNumber="13">
<keyLead id="9F0A9598FFF5FF93FEF2FC619623690C" pageId="12" pageNumber="13">
—— Face with distinct carina (
<figureCitation id="BCC02A28FFF5FF93FD5DFC6090426E99" box="[678,824,922,946]" captionStart="Figures 38–41" captionStartId="15.[121,198,1628,1650]" captionTargetBox="[121,1421,166,1586]" captionTargetId="figure-98@15.[121,1421,151,1618]" captionTargetPageId="15" captionText="Figures 38–41. Photographs of Risa brevirostris sp. nov. (38–39) and R. longirostris Becker (40–41). (38) Head, anterior view. (39) Same, dorso-oblique view. (40) Habitus, lateral view. (41) Head, antero-oblique view. Scale bars = 0.1 mm (38–39), = 0.2 mm (40–41)." figureDoi="http://doi.org/10.5281/zenodo.7946228" httpUri="https://zenodo.org/record/7946228/files/figure.png" pageId="12" pageNumber="13">Figs 38, 39</figureCitation>
); antenna short, basal flagellomere ratio 1; scutum practically without microtomentum; lower position of frons, parafacial and lateral margins of face almost bare of microtomentum, shiny (
<figureCitation id="BCC02A28FFF5FF93FE90FBF5929F6903" box="[363,485,1039,1064]" captionStart="Figures 38–41" captionStartId="15.[121,198,1628,1650]" captionTargetBox="[121,1421,166,1586]" captionTargetId="figure-98@15.[121,1421,151,1618]" captionTargetPageId="15" captionText="Figures 38–41. Photographs of Risa brevirostris sp. nov. (38–39) and R. longirostris Becker (40–41). (38) Head, anterior view. (39) Same, dorso-oblique view. (40) Habitus, lateral view. (41) Head, antero-oblique view. Scale bars = 0.1 mm (38–39), = 0.2 mm (40–41)." figureDoi="http://doi.org/10.5281/zenodo.7946228" httpUri="https://zenodo.org/record/7946228/files/figure.png" pageId="12" pageNumber="13">Figs 38, 39</figureCitation>
) ............................................................................................
<taxonomicName id="E3FB4D2EFFF5FF93FB9CFBEA9785690C" authority="Mathis &amp; Zatwarnicki, 2023" authorityName="Mathis &amp; Zatwarnicki SMNH" authorityYear="2023" box="[1127,1279,1039,1063]" class="Insecta" family="Ephydridae" genus="Risa" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="brevirostris" status="sp. nov.">
<emphasis id="168FEABFFFF5FF93FB9CFBEA9785690C" box="[1127,1279,1039,1063]" italics="true" pageId="12" pageNumber="13">R. brevirostris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0DBC57C4FFF5FF93FAFDFBEA9623690C" box="[1286,1369,1040,1063]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9F0F2E08FFF5FF93FEF2FBB8962369DE" pageId="12" pageNumber="13">
<paragraph id="244436ADFFF5FF93FEF2FBB8962269BF" blockId="12.[265,1368,1090,1173]" pageId="12" pageNumber="13">
<keyLead id="9F0A9598FFF5FF93FEF2FBB8962269BF" pageId="12" pageNumber="13">
9 Proboscis short, labellum about 1.25 times length of ventral margin of head (
<figureCitation id="BCC02A28FFF5FF93FDEFFBA5911E695C" box="[532,612,1119,1143]" captionStart="Figures 20–25" captionStartId="10.[166,243,1096,1118]" captionTargetBox="[167,1460,151,1083]" captionTargetId="figure-249@10.[167,1467,151,1084]" captionTargetPageId="10" captionText="Figures 20–25. Line drawing of frons and associated setae, dorsal view. (20) Notorisa mcalpinei sp. nov. (21) Risa longirostris Becker. (22) R. asiatica Ozerov. (23–25) Line drawing of head, lateral view. (23) R. longirostris Becker. (24) R. brevicornis Papp. (25) R. longicornuta Papp." figureDoi="http://doi.org/10.5281/zenodo.7946222" httpUri="https://zenodo.org/record/7946222/files/figure.png" pageId="12" pageNumber="13">Fig. 25</figureCitation>
); antenna elongate, basal flagellomere ratio 2.5–3 (
<figureCitation id="BCC02A28FFF5FF93FE1EFB87914969BE" box="[485,563,1149,1173]" captionStart="Figures 20–25" captionStartId="10.[166,243,1096,1118]" captionTargetBox="[167,1460,151,1083]" captionTargetId="figure-249@10.[167,1467,151,1084]" captionTargetPageId="10" captionText="Figures 20–25. Line drawing of frons and associated setae, dorsal view. (20) Notorisa mcalpinei sp. nov. (21) Risa longirostris Becker. (22) R. asiatica Ozerov. (23–25) Line drawing of head, lateral view. (23) R. longirostris Becker. (24) R. brevicornis Papp. (25) R. longicornuta Papp." figureDoi="http://doi.org/10.5281/zenodo.7946222" httpUri="https://zenodo.org/record/7946222/files/figure.png" pageId="12" pageNumber="13">Fig. 25</figureCitation>
) ...................................................................................
<taxonomicName id="E3FB4D2EFFF5FF93FB8EFB84962269BF" ID-CoL="4T52D" authority="Papp" authorityName="Papp" box="[1141,1368,1149,1173]" class="Insecta" family="Ephydridae" genus="Risa" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="longicornuta">
<emphasis id="168FEABFFFF5FF93FB8EFB84966669BE" box="[1141,1308,1149,1173]" italics="true" pageId="12" pageNumber="13">R. longicornuta</emphasis>
Papp
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="244436ADFFF5FF93FEF2FB59962369DE" blockId="12.[265,1369,1186,1269]" pageId="12" pageNumber="13">
<keyLead id="9F0A9598FFF5FF93FEF2FB59962369DE" pageId="12" pageNumber="13">
—— Proboscis long, labellum about 1.5 times length of ventral margin of head (
<figureCitation id="BCC02A28FFF5FF93FE3FFB45916369F3" box="[452,537,1215,1240]" captionStart="Figures 20–25" captionStartId="10.[166,243,1096,1118]" captionTargetBox="[167,1460,151,1083]" captionTargetId="figure-249@10.[167,1467,151,1084]" captionTargetPageId="10" captionText="Figures 20–25. Line drawing of frons and associated setae, dorsal view. (20) Notorisa mcalpinei sp. nov. (21) Risa longirostris Becker. (22) R. asiatica Ozerov. (23–25) Line drawing of head, lateral view. (23) R. longirostris Becker. (24) R. brevicornis Papp. (25) R. longicornuta Papp." figureDoi="http://doi.org/10.5281/zenodo.7946222" httpUri="https://zenodo.org/record/7946222/files/figure.png" pageId="12" pageNumber="13">Figs 23</figureCitation>
,
<figureCitation id="BCC02A28FFF5FF93FDDDFB3A911769F3" box="[550,621,1216,1240]" captionStart="Figures 38–41" captionStartId="15.[121,198,1628,1650]" captionTargetBox="[121,1421,166,1586]" captionTargetId="figure-98@15.[121,1421,151,1618]" captionTargetPageId="15" captionText="Figures 38–41. Photographs of Risa brevirostris sp. nov. (38–39) and R. longirostris Becker (40–41). (38) Head, anterior view. (39) Same, dorso-oblique view. (40) Habitus, lateral view. (41) Head, antero-oblique view. Scale bars = 0.1 mm (38–39), = 0.2 mm (40–41)." figureDoi="http://doi.org/10.5281/zenodo.7946228" httpUri="https://zenodo.org/record/7946228/files/figure.png" pageId="12" pageNumber="13">40, 41</figureCitation>
); antenna shorter, basal flagellomere ratio 2 (
<figureCitation id="BCC02A28FFF5FF93FE4DFB27917369DE" box="[438,521,1245,1269]" captionStart="Figures 20–25" captionStartId="10.[166,243,1096,1118]" captionTargetBox="[167,1460,151,1083]" captionTargetId="figure-249@10.[167,1467,151,1084]" captionTargetPageId="10" captionText="Figures 20–25. Line drawing of frons and associated setae, dorsal view. (20) Notorisa mcalpinei sp. nov. (21) Risa longirostris Becker. (22) R. asiatica Ozerov. (23–25) Line drawing of head, lateral view. (23) R. longirostris Becker. (24) R. brevicornis Papp. (25) R. longicornuta Papp." figureDoi="http://doi.org/10.5281/zenodo.7946222" httpUri="https://zenodo.org/record/7946222/files/figure.png" pageId="12" pageNumber="13">Figs 23</figureCitation>
,
<figureCitation id="BCC02A28FFF5FF93FDEFFB27912269DE" box="[532,600,1245,1269]" captionStart="Figures 26–33" captionStartId="13.[121,198,1368,1390]" captionTargetBox="[121,1420,151,1355]" captionTargetId="figure-9@13.[121,1421,151,1357]" captionTargetPageId="13" captionText="Figures 26–33. Photographs of Risa longirostris Becker (26–27, 29–31, 33) and Risa brevirostris sp. nov. (28, 32). (26) Head, anterooblique view. (27) Same, anterior view. (28) Proboscis, lateral view. (29) Rostrum, anterior view (microscopic photograph). (30) Same, lateral view. (31) Labellum, posterior view. (32) Labellum, anterior view. (33) Terminal female abdominal segments, anterior view. Scale bars = 0.2 mm (26–27), = 0.1 mm (28–30, 33), = 0.05 mm (31–32)." figureDoi="http://doi.org/10.5281/zenodo.7946224" httpUri="https://zenodo.org/record/7946224/files/figure.png" pageId="12" pageNumber="13">26, 27</figureCitation>
) ............................................................................
<taxonomicName id="E3FB4D2EFFF5FF93FB96FB24962369DE" ID-CoL="6X8PX" authority="Becker" authorityName="Becker" box="[1133,1369,1245,1269]" class="Insecta" family="Ephydridae" genus="Risa" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="longirostris">
<emphasis id="168FEABFFFF5FF93FB96FB24967F69DE" box="[1133,1285,1245,1269]" italics="true" pageId="12" pageNumber="13">R. longirostris</emphasis>
Becker
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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