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<mods:title id="7DAC9B2E3ABF25862024C4146EDB8FE0">Ilyocypris (Crustacea: Ostracoda) species in North East Asian rice fields description of one new species, and redescriptions of Ilyocypris dentifera Sars 1903 and Ilyocypris japonica Okubo, 1990</mods:title>
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<treatment id="A90187A1BA46FF9A128BD0804FF6F9FF" ID-DOI="http://doi.org/10.5281/zenodo.5610190" ID-GBIF-Taxon="159043965" ID-Zenodo-Dep="5610190" LSID="urn:lsid:plazi:treatment:A90187A1BA46FF9A128BD0804FF6F9FF" httpUri="http://treatment.plazi.org/id/A90187A1BA46FF9A128BD0804FF6F9FF" lastPageId="14" pageId="3">
<subSubSection id="69B2653CBA46FF97128BD0804845FA89" box="[151,549,1254,1281]" pageId="3" type="nomenclature">
<paragraph id="211736B7BA46FF97128BD0804845FA89" blockId="3.[151,552,1254,1317]" box="[151,549,1254,1281]" pageId="3">
<heading id="7A5F81DBBA46FF97128BD0804845FA89" bold="true" box="[151,549,1254,1281]" fontSize="11" level="1" pageId="3" reason="1">
<taxonomicName id="E6A84D34BA46FF97128BD0804845FA89" authority="Okubo, 1990" authorityName="Okubo" authorityYear="1990" box="[151,549,1254,1281]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="3" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="13DCEAA5BA46FF97128BD0804845FA89" bold="true" box="[151,549,1254,1281]" pageId="3">
<emphasis id="13DCEAA5BA46FF97128BD0804B1DFA89" bold="true" box="[151,381,1254,1281]" italics="true" pageId="3">Ilyocypris japonica</emphasis>
<bibRefCitation id="45394B46BA46FF971399D0814845FA89" author="Okubo, I." box="[389,549,1255,1281]" pageId="3" pagination="39 - 50" refId="ref22357" refString="Okubo, I. (1990) Sixteen species of freshwater Ostracoda from Japan. Bulletin of the Biogeographical Society of Japan, 45, 39 - 50." type="journal article" year="1990">Okubo, 1990</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="69B2653CBA46FF97128BD16D4E01FA4B" pageId="3" type="description">
<paragraph id="211736B7BA46FF97128BD16D4848FAAD" blockId="3.[151,552,1254,1317]" box="[151,552,1290,1317]" pageId="3">
(
<figureCitation id="B9932A32BA46FF971283D16D4B60FAAD" box="[159,256,1291,1317]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="5.[151,250,1676,1701]" captionStartId-1="6.[151,250,1743,1768]" captionStartId-2="7.[151,250,1839,1864]" captionStartId-3="8.[151,248,1767,1792]" captionTargetBox-0="[294,1293,179,1649]" captionTargetBox-1="[271,1316,181,1717]" captionTargetBox-2="[239,1349,181,1814]" captionTargetBox-3="[262,1321,181,1741]" captionTargetId-0="figure@5.[294,1293,179,1649]" captionTargetId-1="figure@6.[271,1316,181,1718]" captionTargetId-2="figure@7.[239,1349,181,1814]" captionTargetId-3="figure@8.[261,1326,181,1741]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionText-0="FIGURE 1. Ilyocypris japonica Okubo, 1990. Japanese specimens. A, female left valve, external view (LBM 1430009143, sample 6; see Table 1). B, female right valve, external view (LBM 1430009143, sample 6). C, female left valve, internal view (LBM 1430009143, sample 6). D, female right valve, internal view (LBM 1430009143, sample 6). E, female left valve, internal view, detail of postero-ventral margin (LBM 1430009143, sample 6). F, female right valve, internal view, detail of antero-ventral margin (LBM 1430009143, sample 6). G, female left valve, internal view, muscle scars (LBM 1430009143, sample 6). H, male right valve, external view (LBM 1430009141, sample 6). I, male carapace, dorsal view, anterior to right (LBM 1430009160, sample 25). J, female carapace, dorsal view, anterior to right (LBM 1430009142, sample 6). K, female carapace, tuberculated form, right lateral view (LBM 1430009159, sample 23). L, female carapace, tuberculated form, dorsal view, anterior to right (LBM 1430009159, sample 23). M, male left valve, ventral view (LBM 1430009141, sample 6). Scale bar = 500 µm for AD, HM, 174 µm for EG." captionText-1="FIGURE 2. Ilyocypris japonica Okubo, 1990. Korean specimens. A, female left valve, external view (LBM 1430009147, sample 15, see Table 1). B, female right valve, external view (LBM 1430009147, sample 15). C, female left valve, internal view (LBM 1430009147, sample 15). D, female right valve, internal view (LBM 1430009147, sample 15). E, female left valve, internal view, detail of postero-ventral margin (LBM 1430009147, sample 15). F, female right valve, internal view, detail of anteroventral margin (LBM 1430009147, sample 15). G, female left valve, internal view, muscle scars (LBM 1430009147, sample 15). H, male right valve, external view (LBM 1430009146, sample 15). I, male left valve, external view (LBM 1430009146, sample 15). J, female carapace, tuberculated form, right lateral view (LBM 1430009149, sample 16). K, female carapace, tuberculated form, oblique view of right side (LBM 1430009149, sample 16). Scale bar = 500 µm for AD, HK, 174 µm for EG." captionText-2="FIGURE 3. Ilyocypris japonica Okubo, 1990. A, antennule, male (LBM 1430009154). B, antenna, male (LBM 1430009155). C, mandibular palp, male (LBM 1430009158). D, mandibular coxa, male (LBM 1430009155). E, maxillula palp and endites (setae on endites not drawn), male (LBM 1430009154). F, fifth limb, female (LBM 1430009157). G, fifth limb palp, male (LBM 1430009154). H, sixth limb, male (LBM 1430009154), arrow and dashed line indicate length of first endopodal segment. I, caudal ramus, male (LBM 1430009155). All specimens from sample 22; see Table 1." captionText-3="FIGURE 4. AI, Ilyocypris japonica Okubo, 1990. A, detail of female antenna (LBM 1430009156, sample 22; see Table 1). B, detail of male antenna (LBM 1430009155, sample 22). C, seventh limb, male (LBM 1430009154, sample 22). D, outline of female reproductive organ, arrow indicates anterior (LBM 1430009157, sample 22). E, hemipenis (LBM 1430009154, sample 22). F, copulatory process of hemipenis (LBM 1430009154, sample 22). G, hemipenis (LBM 1430009148, sample 15). H, copulatory process of hemipenis (LBM 1430009148, sample 15). I, outlines of the hemipenes of six specimens, some reflected on the vertical axis to facilitate comparisons (left to right LBM 1430009153, sample 22; LBM 1430009152, sample 21; LBM 1430009146, sample 15; LBM 1430009145, sample 10; LBM 1430009140, sample 5; LBM 1430009139, sample 3). J &amp; K, Ilyocypris mongolica. J, hemipenis (LBM 1430009161, sample 17). K, copulatory process of hemipenis (LBM 1430009161, sample 17)." httpUri-0="https://zenodo.org/record/3363535/files/figure.png" httpUri-1="https://zenodo.org/record/3363537/files/figure.png" httpUri-2="https://zenodo.org/record/3363539/files/figure.png" httpUri-3="https://zenodo.org/record/3363541/files/figure.png" pageId="3">Figs 14</figureCitation>
AI, 5A &amp; B, 6A, 7 &amp; 17)
</paragraph>
<paragraph id="211736B7BA46FF97128BD1294902FAEF" blockId="3.[151,1246,1358,2027]" box="[151,866,1358,1383]" pageId="3">
1989
<taxonomicName id="E6A84D34BA46FF9712CCD1294B55FAEF" authorityName="Brady &amp; Norman" authorityYear="1889" box="[208,309,1359,1383]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="3" phylum="Arthropoda" rank="genus">
<emphasis id="13DCEAA5BA46FF9712CCD1294B55FAEF" box="[208,309,1359,1383]" italics="true" pageId="3">Ilyocypris</emphasis>
</taxonomicName>
sp. A.—Okubo &amp; Ida: 105, pl. 4gl, fide
<bibRefCitation id="45394B46BA46FF9710C1D128493EFAEF" author="Okubo, I." box="[733,862,1358,1383]" pageId="3" pagination="39 - 50" refId="ref22357" refString="Okubo, I. (1990) Sixteen species of freshwater Ostracoda from Japan. Bulletin of the Biogeographical Society of Japan, 45, 39 - 50." type="journal article" year="1990">Okubo 1990</bibRefCitation>
.
</paragraph>
<paragraph id="211736B7BA46FF97128BD10848B3FA0E" blockId="3.[151,1246,1358,2027]" box="[151,723,1389,1414]" pageId="3">
<bibRefCitation id="45394B46BA46FF97128BD1084AAAFA0E" author="Okubo, I." box="[151,202,1390,1414]" pageId="3" pagination="39 - 50" refId="ref22357" refString="Okubo, I. (1990) Sixteen species of freshwater Ostracoda from Japan. Bulletin of the Biogeographical Society of Japan, 45, 39 - 50." type="journal article" year="1990">1990</bibRefCitation>
<taxonomicName id="E6A84D34BA46FF9712CCD1084BF5FA0D" authority="Smith &amp; Zhai &amp; Chang, 2019" authorityName="Smith &amp; Zhai &amp; Chang" authorityYear="2019" box="[208,405,1390,1414]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="3" phylum="Arthropoda" rank="species" species="japonica" status="sp. nov.">
<emphasis id="13DCEAA5BA46FF9712CCD1084BF5FA0D" box="[208,405,1390,1414]" italics="true" pageId="3">Ilyocypris japonica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="08EF57DEBA46FF971387D1084B89FA0E" box="[411,489,1389,1414]" pageId="3" rank="species">sp. nov.</taxonomicNameLabel>
—Okubo: 40, fig. 1fi.
</paragraph>
<paragraph id="211736B7BA46FF97128BD1EA4E23FA2C" blockId="3.[151,1246,1358,2027]" box="[151,1091,1420,1445]" pageId="3">
1992
<taxonomicName id="E6A84D34BA46FF9712CCD1EA4BA3FA2C" authority="Smith &amp; Zhai &amp; Chang, 2019" authorityName="Smith &amp; Zhai &amp; Chang" authorityYear="2019" box="[208,451,1420,1444]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="3" phylum="Arthropoda" rank="species" species="haterumensis" status="sp. nov.">
<emphasis id="13DCEAA5BA46FF9712CCD1EA4BA3FA2C" box="[208,451,1420,1444]" italics="true" pageId="3">Ilyocypris haterumensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="08EF57DEBA46FF9713D5D1EB4876FA2D" box="[457,534,1420,1445]" pageId="3" rank="species">sp. nov.</taxonomicNameLabel>
—Okubo &amp; Terauchi: 101, fig. 1af, fide
<bibRefCitation id="45394B46BA46FF9711A2D1EA4E5FFA2C" author="Okubo I." box="[958,1087,1420,1445]" pageId="3" refId="ref22386" refString="Okubo I. (2004) Nihon tansui san kaimijinko rui ni tsuite. Kabushikigaisha Sanmon Insatsusho, Okayama, 72 pp. [In Japanese, privately published]" type="book" year="2004">Okubo 2004</bibRefCitation>
.
</paragraph>
<paragraph id="211736B7BA46FF97128BD1CD4E01FA4B" blockId="3.[151,1246,1358,2027]" box="[151,1121,1450,1475]" pageId="3">
? 2002
<taxonomicName id="E6A84D34BA46FF9712FED1CD4877FA4B" authority="Sars, 1903" authorityName="G. O. Sars-Chen" authorityYear="1903" box="[226,535,1450,1475]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="3" phylum="Arthropoda" rank="species" species="dentifera">
<emphasis id="13DCEAA5BA46FF9712FED1CD4BC8FA4B" box="[226,424,1451,1475]" italics="true" pageId="3">Ilyocypris dentifera</emphasis>
<bibRefCitation id="45394B46BA46FF9713B3D1CD4877FA4B" author="Sars, G. O." box="[431,535,1450,1475]" pageId="3" pagination="3 - 44" refId="ref22910" refString="Sars, G. O. (1903) Fresh-water Entomostraca from China and Sumatra. Archiv for Mathematik og Naturvidenskab, 25, 3 - 44." type="journal article" year="1903">Sars, 1903</bibRefCitation>
</taxonomicName>
—Nakao &amp; Tsukagoshi: 71, figs 2ad, tables 25, herein.
</paragraph>
</subSubSection>
<subSubSection id="69B2653CBA46FF97128BD1AC48F5F9B6" pageId="3" type="reference_group">
<paragraph id="211736B7BA46FF97128BD1AC49CAFA6A" blockId="3.[151,1246,1358,2027]" box="[151,938,1481,1506]" pageId="3">
2003
<taxonomicName id="E6A84D34BA46FF9712CCD1AC487FFA6A" authority="Okubo, 1990" authorityName="Okubo" authorityYear="1990" box="[208,543,1481,1506]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="3" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="13DCEAA5BA46FF9712CCD1AC4BF5FA69" box="[208,405,1482,1506]" italics="true" pageId="3">Ilyocypris japonica</emphasis>
<bibRefCitation id="45394B46BA46FF971387D1AF487FFA6A" author="Okubo, I." box="[411,543,1481,1506]" pageId="3" pagination="39 - 50" refId="ref22357" refString="Okubo, I. (1990) Sixteen species of freshwater Ostracoda from Japan. Bulletin of the Biogeographical Society of Japan, 45, 39 - 50." type="journal article" year="1990">Okubo, 1990</bibRefCitation>
</taxonomicName>
<emphasis id="13DCEAA5BA46FF971003D1AC4856FA69" box="[543,566,1482,1505]" italics="true" pageId="3"></emphasis>
<collectingRegion id="E36CF855BA46FF97102AD1AF48C8FA6A" box="[566,680,1481,1506]" country="Japan" name="Yamaguchi" pageId="3">Yamaguchi</collectingRegion>
&amp; Endo:
<figureCitation id="B9932A32BA46FF971117D1AF4935FA6A" box="[779,853,1481,1506]" captionStart="FIGURE 4" captionStartId="8.[151,248,1767,1792]" captionTargetBox="[262,1321,181,1741]" captionTargetId="figure@8.[261,1326,181,1741]" captionTargetPageId="8" captionText="FIGURE 4. AI, Ilyocypris japonica Okubo, 1990. A, detail of female antenna (LBM 1430009156, sample 22; see Table 1). B, detail of male antenna (LBM 1430009155, sample 22). C, seventh limb, male (LBM 1430009154, sample 22). D, outline of female reproductive organ, arrow indicates anterior (LBM 1430009157, sample 22). E, hemipenis (LBM 1430009154, sample 22). F, copulatory process of hemipenis (LBM 1430009154, sample 22). G, hemipenis (LBM 1430009148, sample 15). H, copulatory process of hemipenis (LBM 1430009148, sample 15). I, outlines of the hemipenes of six specimens, some reflected on the vertical axis to facilitate comparisons (left to right LBM 1430009153, sample 22; LBM 1430009152, sample 21; LBM 1430009146, sample 15; LBM 1430009145, sample 10; LBM 1430009140, sample 5; LBM 1430009139, sample 3). J &amp; K, Ilyocypris mongolica. J, hemipenis (LBM 1430009161, sample 17). K, copulatory process of hemipenis (LBM 1430009161, sample 17)." httpUri="https://zenodo.org/record/3363541/files/figure.png" pageId="3">
Fig.
<quantity id="E6509B52BA46FF971126D1AC4935FA6A" box="[826,853,1482,1506]" metricMagnitude="-3" metricUnit="kg" metricValue="4.0" pageId="3" unit="g" value="4.0">4g</quantity>
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, table 1.
</paragraph>
<paragraph id="211736B7BA46FF97128BD18E492AF988" blockId="3.[151,1246,1358,2027]" box="[151,842,1512,1537]" pageId="3">
2004
<taxonomicName id="E6A84D34BA46FF9712CCD18E487FF988" authority="Okubo, 1990" authorityName="Okubo" authorityYear="1990" box="[208,543,1512,1537]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="3" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="13DCEAA5BA46FF9712CCD18E4BF5F988" box="[208,405,1512,1536]" italics="true" pageId="3">Ilyocypris japonica</emphasis>
<bibRefCitation id="45394B46BA46FF971387D18E487FF988" author="Okubo, I." box="[411,543,1512,1537]" pageId="3" pagination="39 - 50" refId="ref22357" refString="Okubo, I. (1990) Sixteen species of freshwater Ostracoda from Japan. Bulletin of the Biogeographical Society of Japan, 45, 39 - 50." type="journal article" year="1990">Okubo, 1990</bibRefCitation>
</taxonomicName>
<emphasis id="13DCEAA5BA46FF971003D18F4856F988" box="[543,566,1513,1536]" italics="true" pageId="3"></emphasis>
Okubo: 1718, figs 4waa.
</paragraph>
<paragraph id="211736B7BA46FF97128BD26149A6F997" blockId="3.[151,1246,1358,2027]" box="[151,966,1542,1567]" pageId="3">
2007
<taxonomicName id="E6A84D34BA46FF9712CCD261487DF997" authority="Okubo, 1990" authorityName="Okubo" authorityYear="1990" box="[208,541,1542,1567]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="3" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="13DCEAA5BA46FF9712CCD2614BF5F997" box="[208,405,1543,1567]" italics="true" pageId="3">Ilyocypris japonica</emphasis>
<bibRefCitation id="45394B46BA46FF971387D260487DF997" author="Okubo, I." box="[411,541,1542,1567]" pageId="3" pagination="39 - 50" refId="ref22357" refString="Okubo, I. (1990) Sixteen species of freshwater Ostracoda from Japan. Bulletin of the Biogeographical Society of Japan, 45, 39 - 50." type="journal article" year="1990">Okubo, 1990</bibRefCitation>
</taxonomicName>
—Yamada: 46, figs
<date id="55161077BA46FF9710F6D260489EF997" box="[746,766,1542,1567]" pageId="3">2i</date>
&amp; 3n, tables 1 &amp; 2.
</paragraph>
<paragraph id="211736B7BA46FF97128BD24048F5F9B6" blockId="3.[151,1246,1358,2027]" box="[151,661,1573,1598]" pageId="3">
2008
<taxonomicName id="E6A84D34BA46FF9712CCD240482AF9B6" authority="- Tinn &amp; Oakley" authorityName="- Tinn &amp; Oakley" box="[208,586,1573,1598]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="3" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="13DCEAA5BA46FF9712CCD2404BF5F9B5" box="[208,405,1574,1598]" italics="true" pageId="3">Ilyocypris japonica</emphasis>
—Tinn &amp; Oakley
</taxonomicName>
:
<figureCitation id="B9932A32BA46FF971048D24348F0F9B6" box="[596,656,1573,1598]" captionStart="FIGURE 1" captionStartId="5.[151,250,1676,1701]" captionTargetBox="[294,1293,179,1649]" captionTargetId="figure@5.[294,1293,179,1649]" captionTargetPageId="5" captionText="FIGURE 1. Ilyocypris japonica Okubo, 1990. Japanese specimens. A, female left valve, external view (LBM 1430009143, sample 6; see Table 1). B, female right valve, external view (LBM 1430009143, sample 6). C, female left valve, internal view (LBM 1430009143, sample 6). D, female right valve, internal view (LBM 1430009143, sample 6). E, female left valve, internal view, detail of postero-ventral margin (LBM 1430009143, sample 6). F, female right valve, internal view, detail of antero-ventral margin (LBM 1430009143, sample 6). G, female left valve, internal view, muscle scars (LBM 1430009143, sample 6). H, male right valve, external view (LBM 1430009141, sample 6). I, male carapace, dorsal view, anterior to right (LBM 1430009160, sample 25). J, female carapace, dorsal view, anterior to right (LBM 1430009142, sample 6). K, female carapace, tuberculated form, right lateral view (LBM 1430009159, sample 23). L, female carapace, tuberculated form, dorsal view, anterior to right (LBM 1430009159, sample 23). M, male left valve, ventral view (LBM 1430009141, sample 6). Scale bar = 500 µm for AD, HM, 174 µm for EG." httpUri="https://zenodo.org/record/3363535/files/figure.png" pageId="3">Fig. 1</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="69B2653CBA46FF97128BD2224EBEF9D5" box="[151,1246,1604,1629]" pageId="3" type="description">
<paragraph id="211736B7BA46FF97128BD2224EBEF9D5" blockId="3.[151,1246,1358,2027]" box="[151,1246,1604,1629]" pageId="3">
? 2008
<taxonomicName id="E6A84D34BA46FF9712FED2224877F9D5" authority="Sars, 1903" authorityName="G. O. Sars-Chen" authorityYear="1903" box="[226,535,1604,1629]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="3" phylum="Arthropoda" rank="species" species="dentifera">
<emphasis id="13DCEAA5BA46FF9712FED2224BC8F9D4" box="[226,424,1604,1628]" italics="true" pageId="3">Ilyocypris dentifera</emphasis>
<bibRefCitation id="45394B46BA46FF9713B3D2224877F9D5" author="Sars, G. O." box="[431,535,1604,1629]" pageId="3" pagination="3 - 44" refId="ref22910" refString="Sars, G. O. (1903) Fresh-water Entomostraca from China and Sumatra. Archiv for Mathematik og Naturvidenskab, 25, 3 - 44." type="journal article" year="1903">Sars, 1903</bibRefCitation>
</taxonomicName>
—Nakao &amp; Tsukagoshi: 251, 263, plate 1ad, tables 1, 1214, herein.
</paragraph>
</subSubSection>
<subSubSection id="69B2653CBA46FF90128BD20548E3FF58" lastPageId="4" pageId="3" type="reference_group">
<paragraph id="211736B7BA46FF97128BD205488FF9F2" blockId="3.[151,1246,1358,2027]" box="[151,751,1634,1659]" pageId="3">
2009
<taxonomicName id="E6A84D34BA46FF9712CCD205487FF9F3" authority="Okubo, 1990" authorityName="Okubo" authorityYear="1990" box="[208,543,1634,1659]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="3" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="13DCEAA5BA46FF9712CCD2054BF5F9F3" box="[208,405,1635,1659]" italics="true" pageId="3">Ilyocypris japonica</emphasis>
<bibRefCitation id="45394B46BA46FF971387D204487FF9F3" author="Okubo, I." box="[411,543,1634,1659]" pageId="3" pagination="39 - 50" refId="ref22357" refString="Okubo, I. (1990) Sixteen species of freshwater Ostracoda from Japan. Bulletin of the Biogeographical Society of Japan, 45, 39 - 50." type="journal article" year="1990">Okubo, 1990</bibRefCitation>
</taxonomicName>
<emphasis id="13DCEAA5BA46FF971003D2024856F9F3" box="[543,566,1636,1659]" italics="true" pageId="3"></emphasis>
Hayashi
<emphasis id="13DCEAA5BA46FF971093D20248DCF9F3" box="[655,700,1635,1659]" italics="true" pageId="3">et al</emphasis>
.: 77.
</paragraph>
<paragraph id="211736B7BA46FF97128BD2E448AAF912" blockId="3.[151,1246,1358,2027]" box="[151,714,1665,1690]" pageId="3">
2010
<taxonomicName id="E6A84D34BA46FF9712CCD2E44810F912" authority="- Kaji &amp; Tsukagoshi" authorityName="- Kaji &amp; Tsukagoshi" box="[208,624,1665,1690]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="3" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="13DCEAA5BA46FF9712CCD2E44BF5F911" box="[208,405,1666,1690]" italics="true" pageId="3">Ilyocypris japonica</emphasis>
—Kaji &amp; Tsukagoshi
</taxonomicName>
:
<tableCitation id="6C2A030CBA46FF971065D2E748A6F912" box="[633,710,1665,1690]" captionStart="TABLE 1" captionStartId="1.[151,244,1160,1187]" captionTargetBox="[163,1412,1310,1957]" captionText="TABLE 1. Adult Ilyocypris specimens examined for this study. Initial discriminations of species were based on lateral views of carapaces, and confirmed by dissections of male specimens. Figures in brackets indicate the number of specimens with tubercles on the carapace. Note that tubercles vary from weakly to well-developed and were no- ticeable on whole specimens or valves in dorsal view using a stereo optical microscope." httpUri="http://table.plazi.org/id/75D7663FBA44FF95128BD0EE4E0FFA87" pageId="3" tableUuid="75D7663FBA44FF95128BD0EE4E0FFA87">Table 1</tableCitation>
.
</paragraph>
<paragraph id="211736B7BA46FF97128BD2C649AEF930" blockId="3.[151,1246,1358,2027]" box="[151,974,1696,1721]" pageId="3">
2011
<taxonomicName id="E6A84D34BA46FF9712D3D2C6482BF930" authority="Okubo, 1992" authorityName="Okubo" authorityYear="1992" box="[207,587,1696,1721]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="3" phylum="Arthropoda" rank="species" species="haterumensis">
<emphasis id="13DCEAA5BA46FF9712D3D2C64BA2F930" box="[207,450,1696,1720]" italics="true" pageId="3">Ilyocypris haterumensis</emphasis>
<bibRefCitation id="45394B46BA46FF9713D4D2C6482BF930" author="Okubo, I. &amp; Terauchi, M." box="[456,587,1696,1721]" pageId="3" pagination="101 - 108" refId="ref22454" refString="Okubo, I. &amp; Terauchi, M. (1992) Freshwater Ostracoda from the Southwestern Islands of Japan. Proceedings of the Japanese Society of Systematic Zoology, 46, 101 - 108." type="journal article" year="1992">Okubo, 1992</bibRefCitation>
</taxonomicName>
—Martens &amp; Savatenalinton: Table 2.
</paragraph>
<paragraph id="211736B7BA46FF97128BD2D949C0F95F" blockId="3.[151,1246,1358,2027]" box="[151,928,1726,1751]" pageId="3">
2011
<taxonomicName id="E6A84D34BA46FF9712D3D2D9487CF95F" authority="Okubo, 1990" authorityName="Okubo" authorityYear="1990" box="[207,540,1726,1751]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="3" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="13DCEAA5BA46FF9712D3D2D94BF4F95F" box="[207,404,1727,1751]" italics="true" pageId="3">Ilyocypris japonica</emphasis>
<bibRefCitation id="45394B46BA46FF971386D2D8487CF95F" author="Okubo, I." box="[410,540,1726,1751]" pageId="3" pagination="39 - 50" refId="ref22357" refString="Okubo, I. (1990) Sixteen species of freshwater Ostracoda from Japan. Bulletin of the Biogeographical Society of Japan, 45, 39 - 50." type="journal article" year="1990">Okubo, 1990</bibRefCitation>
</taxonomicName>
—Martens &amp; Savatenalinton: Table 2.
</paragraph>
<paragraph id="211736B7BA46FF97128BD2B84881F97E" blockId="3.[151,1246,1358,2027]" box="[151,737,1757,1782]" pageId="3">
<date id="55161077BA46FF97128BD2B84A80F97D" box="[151,224,1758,1782]" pageId="3" value="2012-01">
2012
<taxonomicName id="E6A84D34BA46FF9712CCD2B84A80F97D" authority="Okubo 1992" authorityName="Okubo" authorityYear="1992" box="[208,224,1758,1781]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="3" phylum="Arthropoda" rank="species" species="haterumensis">
<emphasis id="13DCEAA5BA46FF9712CCD2B84A80F97D" box="[208,224,1758,1781]" italics="true" pageId="3">I.</emphasis>
</taxonomicName>
</date>
haterumensis
<bibRefCitation id="45394B46BA46FF97136FD2BB4B90F97E" author="Okubo, I. &amp; Terauchi, M." box="[371,496,1757,1782]" pageId="3" pagination="101 - 108" refId="ref22454" refString="Okubo, I. &amp; Terauchi, M. (1992) Freshwater Ostracoda from the Southwestern Islands of Japan. Proceedings of the Japanese Society of Systematic Zoology, 46, 101 - 108." type="journal article" year="1992">Okubo 1992</bibRefCitation>
—Karanovic: 204, 207.
</paragraph>
<paragraph id="211736B7BA46FF97128BD29A4974F89C" blockId="3.[151,1246,1358,2027]" box="[151,788,1788,1813]" pageId="3">
2012
<taxonomicName id="E6A84D34BA46FF9712CCD29A4878F89D" authority="Okubo 1990" authorityName="Okubo" authorityYear="1990" box="[208,536,1788,1813]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="3" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="13DCEAA5BA46FF9712CCD29A4BF5F89C" box="[208,405,1788,1812]" italics="true" pageId="3">Ilyocypris japonica</emphasis>
<bibRefCitation id="45394B46BA46FF971387D29A4878F89D" author="Okubo, I." box="[411,536,1788,1813]" pageId="3" pagination="39 - 50" refId="ref22357" refString="Okubo, I. (1990) Sixteen species of freshwater Ostracoda from Japan. Bulletin of the Biogeographical Society of Japan, 45, 39 - 50." type="journal article" year="1990">Okubo 1990</bibRefCitation>
</taxonomicName>
a—Karanovic: 205, 208.
</paragraph>
<paragraph id="211736B7BA46FF97128BD37D490FF8BB" blockId="3.[151,1246,1358,2027]" box="[151,879,1818,1843]" pageId="3">
2012
<taxonomicName id="E6A84D34BA46FF9712CCD37D487FF8BB" authority="Okubo, 1990" authorityName="Okubo" authorityYear="1990" box="[208,543,1818,1843]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="3" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="13DCEAA5BA46FF9712CCD37D4BF5F8BB" box="[208,405,1819,1843]" italics="true" pageId="3">Ilyocypris japonica</emphasis>
<bibRefCitation id="45394B46BA46FF971387D37C487FF8BB" author="Okubo, I." box="[411,543,1818,1843]" pageId="3" pagination="39 - 50" refId="ref22357" refString="Okubo, I. (1990) Sixteen species of freshwater Ostracoda from Japan. Bulletin of the Biogeographical Society of Japan, 45, 39 - 50." type="journal article" year="1990">Okubo, 1990</bibRefCitation>
</taxonomicName>
<emphasis id="13DCEAA5BA46FF971003D37A4856F8BB" box="[543,566,1820,1843]" italics="true" pageId="3"></emphasis>
Sidorov &amp; Semenchenko: 233.
</paragraph>
<paragraph id="211736B7BA46FF97128BD35C488FF8DA" blockId="3.[151,1246,1358,2027]" box="[151,751,1849,1874]" pageId="3">
2012
<taxonomicName id="E6A84D34BA46FF9712CCD35C487FF8DA" authority="Okubo, 1990" authorityName="Okubo" authorityYear="1990" box="[208,543,1849,1874]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="3" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="13DCEAA5BA46FF9712CCD35C4BF5F8D9" box="[208,405,1850,1874]" italics="true" pageId="3">Ilyocypris japonica</emphasis>
<bibRefCitation id="45394B46BA46FF971387D35F487FF8DA" author="Okubo, I." box="[411,543,1849,1874]" pageId="3" pagination="39 - 50" refId="ref22357" refString="Okubo, I. (1990) Sixteen species of freshwater Ostracoda from Japan. Bulletin of the Biogeographical Society of Japan, 45, 39 - 50." type="journal article" year="1990">Okubo, 1990</bibRefCitation>
</taxonomicName>
<emphasis id="13DCEAA5BA46FF971003D35C4856F8D9" box="[543,566,1850,1873]" italics="true" pageId="3"></emphasis>
Hayashi
<emphasis id="13DCEAA5BA46FF971093D35C48DCF8DA" box="[655,700,1850,1874]" italics="true" pageId="3">et al</emphasis>
.: 61.
</paragraph>
<paragraph id="211736B7BA46FF97128BD33E495AF8F9" blockId="3.[151,1246,1358,2027]" box="[151,826,1880,1905]" pageId="3">
<date id="55161077BA46FF97128BD33E4A80F8F8" box="[151,224,1880,1904]" pageId="3" value="2013-01">
2013
<taxonomicName id="E6A84D34BA46FF9712CCD33F4A80F8F8" authority="Okubo, 1992" authorityName="Okubo" authorityYear="1992" box="[208,224,1881,1904]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="3" phylum="Arthropoda" rank="species" species="haterumensis">
<emphasis id="13DCEAA5BA46FF9712CCD33F4A80F8F8" box="[208,224,1881,1904]" italics="true" pageId="3">I.</emphasis>
</taxonomicName>
</date>
haterumensis
<bibRefCitation id="45394B46BA46FF97136FD33E4B95F8F9" author="Okubo, I. &amp; Terauchi, M." box="[371,501,1880,1905]" pageId="3" pagination="101 - 108" refId="ref22454" refString="Okubo, I. &amp; Terauchi, M. (1992) Freshwater Ostracoda from the Southwestern Islands of Japan. Proceedings of the Japanese Society of Systematic Zoology, 46, 101 - 108." type="journal article" year="1992">Okubo, 1992</bibRefCitation>
—Karanovic &amp; Lee: 41, 58, 64.
</paragraph>
<paragraph id="211736B7BA46FF97128BD3114915F807" blockId="3.[151,1246,1358,2027]" box="[151,885,1910,1935]" pageId="3">
<date id="55161077BA46FF97128BD3114A80F807" box="[151,224,1911,1935]" pageId="3" value="2013-01">
2013
<taxonomicName id="E6A84D34BA46FF9712CCD31E4A80F807" authority="Okubo, 1990" authorityName="Okubo" authorityYear="1990" box="[208,224,1912,1935]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="3" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="13DCEAA5BA46FF9712CCD31E4A80F807" box="[208,224,1912,1935]" italics="true" pageId="3">I.</emphasis>
</taxonomicName>
</date>
japonica
<bibRefCitation id="45394B46BA46FF971358D3104BA7F807" author="Okubo, I." box="[324,455,1910,1935]" pageId="3" pagination="39 - 50" refId="ref22357" refString="Okubo, I. (1990) Sixteen species of freshwater Ostracoda from Japan. Bulletin of the Biogeographical Society of Japan, 45, 39 - 50." type="journal article" year="1990">Okubo, 1990</bibRefCitation>
—Karanovic &amp; Lee: 41, 58, 64, appendix.
</paragraph>
<paragraph id="211736B7BA46FF97128BD3F0485DF826" blockId="3.[151,1246,1358,2027]" box="[151,573,1941,1966]" pageId="3">
<date id="55161077BA46FF97128BD3F04A80F825" box="[151,224,1942,1966]" pageId="3" value="2013-01">
2013
<emphasis id="13DCEAA5BA46FF9712CCD3F04A80F825" box="[208,224,1942,1965]" italics="true" pageId="3">I.</emphasis>
</date>
haterumensis—Zhai &amp; Xiao: 468.
</paragraph>
<paragraph id="211736B7BA46FF97128BD3D249ECF845" blockId="3.[151,1246,1358,2027]" box="[151,908,1972,1997]" pageId="3">
2013
<taxonomicName id="E6A84D34BA46FF9712CCD3D2487DF844" authority="Okubo, 1990" authorityName="Okubo" authorityYear="1990" box="[208,541,1972,1997]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="3" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="13DCEAA5BA46FF9712CCD3D24BF5F844" box="[208,405,1972,1996]" italics="true" pageId="3">Ilyocypris japonica</emphasis>
<bibRefCitation id="45394B46BA46FF971387D3D2487DF844" author="Okubo, I." box="[411,541,1972,1997]" pageId="3" pagination="39 - 50" refId="ref22357" refString="Okubo, I. (1990) Sixteen species of freshwater Ostracoda from Japan. Bulletin of the Biogeographical Society of Japan, 45, 39 - 50." type="journal article" year="1990">Okubo, 1990</bibRefCitation>
</taxonomicName>
—Martens
<emphasis id="13DCEAA5BA46FF97108ED3D348DEF844" box="[658,702,1972,1996]" italics="true" pageId="3">et al</emphasis>
.: No page numbers.
</paragraph>
<paragraph id="211736B7BA46FF97128BD3B549DAF863" blockId="3.[151,1246,1358,2027]" box="[151,954,2002,2027]" pageId="3">
2013
<taxonomicName id="E6A84D34BA46FF9712CCD3B5482CF863" authority="Okubo, 1992" authorityName="Okubo" authorityYear="1992" box="[208,588,2002,2027]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="3" phylum="Arthropoda" rank="species" species="haterumensis">
<emphasis id="13DCEAA5BA46FF9712CCD3B54BA3F863" box="[208,451,2003,2027]" italics="true" pageId="3">Ilyocypris haterumensis</emphasis>
<bibRefCitation id="45394B46BA46FF9713D5D3B4482CF863" author="Okubo, I. &amp; Terauchi, M." box="[457,588,2002,2027]" pageId="3" pagination="101 - 108" refId="ref22454" refString="Okubo, I. &amp; Terauchi, M. (1992) Freshwater Ostracoda from the Southwestern Islands of Japan. Proceedings of the Japanese Society of Systematic Zoology, 46, 101 - 108." type="journal article" year="1992">Okubo, 1992</bibRefCitation>
</taxonomicName>
—Martens
<emphasis id="13DCEAA5BA46FF9710DCD3B2488DF863" box="[704,749,2003,2027]" italics="true" pageId="3">et al</emphasis>
.: No page numbers.
</paragraph>
<paragraph id="211736B7BA41FF90128BD4FE4897FF39" blockId="4.[151,1109,152,361]" box="[151,759,152,177]" pageId="4">
<date id="55161077BA41FF90128BD4FE4A80FF38" box="[151,224,152,177]" pageId="4" value="2015-01">
2015
<taxonomicName id="E6A84D34BA41FF9012CCD4FF4A80FF38" authority="Okubo, 1992" authorityName="Okubo" authorityYear="1992" box="[208,224,153,176]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="4" phylum="Arthropoda" rank="species" species="haterumensis">
<emphasis id="13DCEAA5BA41FF9012CCD4FF4A80FF38" box="[208,224,153,176]" italics="true" pageId="4">I.</emphasis>
</taxonomicName>
</date>
haterumensis
<bibRefCitation id="45394B46BA41FF90136FD4FE4B95FF39" author="Okubo, I. &amp; Terauchi, M." box="[371,501,152,177]" pageId="4" pagination="101 - 108" refId="ref22454" refString="Okubo, I. &amp; Terauchi, M. (1992) Freshwater Ostracoda from the Southwestern Islands of Japan. Proceedings of the Japanese Society of Systematic Zoology, 46, 101 - 108." type="journal article" year="1992">Okubo, 1992</bibRefCitation>
—DAmbrosio
<emphasis id="13DCEAA5BA41FF90108BD4FF48A3FF39" box="[663,707,153,177]" italics="true" pageId="4">et al</emphasis>
.: 65.
</paragraph>
<paragraph id="211736B7BA41FF90128BD4D148E3FF58" blockId="4.[151,1109,152,361]" box="[151,643,183,208]" pageId="4">
2016b
<emphasis id="13DCEAA5BA41FF9012C1D4D14BD7FF47" box="[221,439,183,207]" italics="true" pageId="4">
<taxonomicName id="E6A84D34BA41FF9012C1D4D14B22FF47" authorityName="Brady &amp; Norman" authorityYear="1889" box="[221,322,183,207]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="4" phylum="Arthropoda" rank="genus">Ilyocypris</taxonomicName>
japonica—
</emphasis>
Smith
<emphasis id="13DCEAA5BA41FF9013E7D4DE484EFF47" box="[507,558,183,207]" italics="true" pageId="4">et al.</emphasis>
:
<figureCitation id="B9932A32BA41FF901027D4D148E0FF58" box="[571,640,183,208]" captionStart="FIGURE 2" captionStartId="6.[151,250,1743,1768]" captionTargetBox="[271,1316,181,1717]" captionTargetId="figure@6.[271,1316,181,1718]" captionTargetPageId="6" captionText="FIGURE 2. Ilyocypris japonica Okubo, 1990. Korean specimens. A, female left valve, external view (LBM 1430009147, sample 15, see Table 1). B, female right valve, external view (LBM 1430009147, sample 15). C, female left valve, internal view (LBM 1430009147, sample 15). D, female right valve, internal view (LBM 1430009147, sample 15). E, female left valve, internal view, detail of postero-ventral margin (LBM 1430009147, sample 15). F, female right valve, internal view, detail of anteroventral margin (LBM 1430009147, sample 15). G, female left valve, internal view, muscle scars (LBM 1430009147, sample 15). H, male right valve, external view (LBM 1430009146, sample 15). I, male left valve, external view (LBM 1430009146, sample 15). J, female carapace, tuberculated form, right lateral view (LBM 1430009149, sample 16). K, female carapace, tuberculated form, oblique view of right side (LBM 1430009149, sample 16). Scale bar = 500 µm for AD, HK, 174 µm for EG." httpUri="https://zenodo.org/record/3363537/files/figure.png" pageId="4">Fig. 2i</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="69B2653CBA41FF90128BD4B34837FEA4" pageId="4" type="description">
<paragraph id="211736B7BA41FF90128BD4B34E35FF66" blockId="4.[151,1109,152,361]" box="[151,1109,213,238]" pageId="4">
2016a
<taxonomicName id="E6A84D34BA41FF9012C7D4B04849FF66" authority="Okubo, 1990" authorityName="Okubo" authorityYear="1990" box="[219,553,213,238]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="4" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="13DCEAA5BA41FF9012C7D4B04BC0FF66" box="[219,416,214,238]" italics="true" pageId="4">Ilyocypris japonica</emphasis>
<bibRefCitation id="45394B46BA41FF9013BAD4B34849FF66" author="Okubo, I." box="[422,553,213,238]" pageId="4" pagination="39 - 50" refId="ref22357" refString="Okubo, I. (1990) Sixteen species of freshwater Ostracoda from Japan. Bulletin of the Biogeographical Society of Japan, 45, 39 - 50." type="journal article" year="1990">Okubo, 1990</bibRefCitation>
</taxonomicName>
—Smith
<emphasis id="13DCEAA5BA41FF90109BD4B148D3FF66" box="[647,691,214,238]" italics="true" pageId="4">et al</emphasis>
.: 484488, 492495, fig. 6, tables 1 &amp; 2.
</paragraph>
<paragraph id="211736B7BA41FF90128BD493484CFE85" blockId="4.[151,1109,152,361]" box="[151,556,244,269]" pageId="4">
<date id="55161077BA41FF90128BD4934A80FE84" box="[151,224,245,269]" pageId="4" value="2017-01">
2017
<taxonomicName id="E6A84D34BA41FF9012CCD4934A80FE84" authority="- Ghaouaci et al." authorityName="Okubo" authorityYear="1990" box="[208,224,245,268]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="4" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="13DCEAA5BA41FF9012CCD4934A80FE84" box="[208,224,245,268]" italics="true" pageId="4">I.</emphasis>
</taxonomicName>
</date>
japonica—Ghaouaci
<emphasis id="13DCEAA5BA41FF9013A3D4934B8BFE85" box="[447,491,245,269]" italics="true" pageId="4">et al</emphasis>
.: 145.
</paragraph>
<paragraph id="211736B7BA41FF90128BD5754837FEA4" blockId="4.[151,1109,152,361]" box="[151,599,275,300]" pageId="4">
<date id="55161077BA41FF90128BD5754A80FEA3" box="[151,224,275,299]" pageId="4" value="2017-01">
2017
<emphasis id="13DCEAA5BA41FF9012CCD5724A80FEA3" box="[208,224,276,299]" italics="true" pageId="4">I.</emphasis>
</date>
haterumensis—Ghaouaci
<emphasis id="13DCEAA5BA41FF9013F6D5724877FEA3" box="[490,535,275,299]" italics="true" pageId="4">et al</emphasis>
.: 145.
</paragraph>
</subSubSection>
<subSubSection id="69B2653CBA41FF90128BD5544889FEE1" pageId="4" type="reference_group">
<paragraph id="211736B7BA41FF90128BD5544945FEC2" blockId="4.[151,1109,152,361]" box="[151,805,305,330]" pageId="4">
2018
<taxonomicName id="E6A84D34BA41FF9012CCD554487DFEC2" authority="Okubo, 1990" authorityName="Okubo" authorityYear="1990" box="[208,541,305,330]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="4" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="13DCEAA5BA41FF9012CCD5544BF5FEC2" box="[208,405,306,330]" italics="true" pageId="4">Ilyocypris japonica</emphasis>
<bibRefCitation id="45394B46BA41FF901387D557487DFEC2" author="Okubo, I." box="[411,541,305,330]" pageId="4" pagination="39 - 50" refId="ref22357" refString="Okubo, I. (1990) Sixteen species of freshwater Ostracoda from Japan. Bulletin of the Biogeographical Society of Japan, 45, 39 - 50." type="journal article" year="1990">Okubo, 1990</bibRefCitation>
</taxonomicName>
—Smith
<emphasis id="13DCEAA5BA41FF901067D55548C8FEC2" box="[635,680,306,330]" italics="true" pageId="4">et al</emphasis>
.: Appendix.
</paragraph>
<paragraph id="211736B7BA41FF90128BD5374889FEE1" blockId="4.[151,1109,152,361]" box="[151,745,336,361]" pageId="4">
2019
<taxonomicName id="E6A84D34BA41FF9012CCD537487DFEE1" authority="Okubo, 1990" authorityName="Okubo" authorityYear="1990" box="[208,541,336,361]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="4" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="13DCEAA5BA41FF9012CCD5374BF5FEE0" box="[208,405,337,361]" italics="true" pageId="4">Ilyocypris japonica</emphasis>
<bibRefCitation id="45394B46BA41FF901387D536487DFEE1" author="Okubo, I." box="[411,541,336,361]" pageId="4" pagination="39 - 50" refId="ref22357" refString="Okubo, I. (1990) Sixteen species of freshwater Ostracoda from Japan. Bulletin of the Biogeographical Society of Japan, 45, 39 - 50." type="journal article" year="1990">Okubo, 1990</bibRefCitation>
</taxonomicName>
—Meisch
<emphasis id="13DCEAA5BA41FF901095D53748D6FEE1" box="[649,694,337,361]" italics="true" pageId="4">et al</emphasis>
.: 71.
</paragraph>
</subSubSection>
<subSubSection id="69B2653CBA41FF9A128BD5F74F0CFDB7" lastPageId="14" pageId="4" type="materials_examined">
<paragraph id="211736B7BA41FF90128BD5F74980FD44" blockId="4.[151,1437,401,2045]" pageId="4">
<emphasis id="13DCEAA5BA41FF90128BD5F74B72FE24" bold="true" box="[151,274,401,428]" pageId="4">Diagnosis.</emphasis>
Carapace anterior margin slightly more inflated than posterior margin, dorsal margin sloping down posteriorly. Surface covered in many well-developed small pits, and a small number of tiny spines near posterior and anterior margins (sometimes missing). Up to six tubercles sometimes present, rounded to angular. Anterior inner calcified lamellae of both valves with lists. Swimming setae of antenna very long, extending significantly beyond end of claws. Sixth limb with five segments (second endopodal segment divided). Seventh limb with one seta on basis, two setae on third segment (= second endopodal segment), and small, hook-like setules distally on same segment. Anterior end of Zenker organ large and inflated, much larger than posterior end. Inner lobe of hemipenis long and thin, with inflated, rounded tip, and noticeably longer than outer lobe, middle and outer lobes overlapping, copulatory process slender, with curved outer edge and sharply pointed tip.
</paragraph>
<paragraph id="211736B7BA41FF9012DBD6B04882FCB0" blockId="4.[151,1437,401,2045]" pageId="4">
<emphasis id="13DCEAA5BA41FF9012DBD6B04B06FD78" bold="true" box="[199,358,725,752]" pageId="4">
<typeStatus id="FE138815BA41FF9012DBD6B04B62FD78" box="[199,258,726,752]" pageId="4">Type</typeStatus>
locality.
</emphasis>
A
<typeStatus id="FE138815BA41FF90139BD6B04BD7FD78" box="[391,439,726,752]" pageId="4">type</typeStatus>
locality was not explicitly designated by
<bibRefCitation id="45394B46BA41FF901197D6B04E4CFD78" author="Okubo, I." box="[907,1068,726,752]" pageId="4" pagination="39 - 50" refId="ref22357" refString="Okubo, I. (1990) Sixteen species of freshwater Ostracoda from Japan. Bulletin of the Biogeographical Society of Japan, 45, 39 - 50." type="journal article" year="1990">Okubo (1990)</bibRefCitation>
. The
<typeStatus id="FE138815BA41FF901676D6B04EFAFD78" box="[1130,1178,726,752]" pageId="4">type</typeStatus>
material was collected from “paddy fields near Tsukuda Station of the
<collectingCountry id="59BF7627BA41FF9010A2D69C489EFC9C" box="[702,766,762,788]" name="Japan" pageId="4">Japan</collectingCountry>
Railway, Gumma Prefecture” (
<bibRefCitation id="45394B46BA41FF90167BD69C4E9BFC9C" author="Okubo, I." box="[1127,1275,762,788]" pageId="4" pagination="39 - 50" refId="ref22357" refString="Okubo, I. (1990) Sixteen species of freshwater Ostracoda from Japan. Bulletin of the Biogeographical Society of Japan, 45, 39 - 50." type="journal article" year="1990">Okubo 1990</bibRefCitation>
), approximate coordinates (herein given)
<geoCoordinate id="449C5070BA41FF9013D8D77B4826FCB0" box="[452,582,797,824]" degrees="36.5610" direction="north" orientation="latitude" pageId="4" precision="5" value="36.561">36.5610º N</geoCoordinate>
,
<geoCoordinate id="449C5070BA41FF90104DD77848BEFCB0" box="[593,734,798,824]" degrees="139.0427" direction="east" orientation="longitude" pageId="4" precision="5" value="139.0427">139.0427º E</geoCoordinate>
.
</paragraph>
<paragraph id="211736B7BA41FF9012DBD7244B67FC2C" blockId="4.[151,1437,401,2045]" pageId="4">
<emphasis id="13DCEAA5BA41FF9012DBD7244B10FCD4" bold="true" box="[199,368,833,860]" pageId="4">Type material</emphasis>
(as designated by
<bibRefCitation id="45394B46BA41FF901054D72448BAFCD4" author="Okubo, I." box="[584,730,834,860]" pageId="4" pagination="39 - 50" refId="ref22357" refString="Okubo, I. (1990) Sixteen species of freshwater Ostracoda from Japan. Bulletin of the Biogeographical Society of Japan, 45, 39 - 50." type="journal article" year="1990">Okubo 1990</bibRefCitation>
).
<typeStatus id="FE138815BA41FF9010F3D724493CFCD4" box="[751,860,834,860]" pageId="4" type="holotype">Holotype</typeStatus>
: male (FO-652).
<typeStatus id="FE138815BA41FF901603D7244EE5FCD4" box="[1055,1157,834,860]" pageId="4" type="allotype">Allotype</typeStatus>
: female (FO-654).
<typeStatus id="FE138815BA41FF90177DD7244AB6FC08" pageId="4" type="paratype">Paratypes</typeStatus>
: male (FO-651), male (FO-655). Numbers were provisional. Location of type material not stated and currently unknown.
</paragraph>
<paragraph id="211736B7BA41FF9012DBD7CB4820FC40" blockId="4.[151,1437,401,2045]" box="[199,576,941,968]" pageId="4">
<emphasis id="13DCEAA5BA41FF9012DBD7CB4BD1FC40" bold="true" box="[199,433,941,968]" pageId="4">Material examined.</emphasis>
See
<tableCitation id="6C2A030CBA41FF9013FAD7C8485AFC40" box="[486,570,942,968]" captionStart="TABLE 1" captionStartId="1.[151,244,1160,1187]" captionTargetBox="[163,1412,1310,1957]" captionText="TABLE 1. Adult Ilyocypris specimens examined for this study. Initial discriminations of species were based on lateral views of carapaces, and confirmed by dissections of male specimens. Figures in brackets indicate the number of specimens with tubercles on the carapace. Note that tubercles vary from weakly to well-developed and were no- ticeable on whole specimens or valves in dorsal view using a stereo optical microscope." httpUri="http://table.plazi.org/id/75D7663FBA44FF95128BD0EE4E0FFA87" pageId="4" tableUuid="75D7663FBA44FF95128BD0EE4E0FFA87">Table 1</tableCitation>
.
</paragraph>
<paragraph id="211736B7BA41FF9012DBD7B749EBFA48" blockId="4.[151,1437,401,2045]" pageId="4">
<emphasis id="13DCEAA5BA41FF9012DBD7B74B39FC64" bold="true" box="[199,345,977,1004]" pageId="4">Description.</emphasis>
Male carapace length
<quantity id="E6509B52BA41FF901041D7B74890FC64" box="[605,752,977,1004]" metricMagnitude="-1" metricUnit="m" metricValue="7.409999999999999" metricValueMax="8.23" metricValueMin="6.59" pageId="4" unit="mm" value="741.0" valueMax="823.0" valueMin="659.0">659823 µm</quantity>
, height
<quantity id="E6509B52BA41FF901150D7B449BFFC64" box="[844,991,978,1004]" metricMagnitude="-1" metricUnit="m" metricValue="3.9899999999999998" metricValueMax="4.3100000000000005" metricValueMin="3.6700000000000004" pageId="4" unit="mm" value="399.0" valueMax="431.0" valueMin="367.0">367431 µm</quantity>
, height/length 0.500.56. Female carapace length
<quantity id="E6509B52BA41FF90133BD7904BDCFB98" box="[295,444,1014,1040]" metricMagnitude="-1" metricUnit="m" metricValue="8.245" metricValueMax="9.27" metricValueMin="7.22" pageId="4" unit="mm" value="824.5" valueMax="927.0" valueMin="722.0">722927 µm</quantity>
, height
<quantity id="E6509B52BA41FF901001D79348D2FB98" box="[541,690,1013,1040]" metricMagnitude="-1" metricUnit="m" metricValue="4.525" metricValueMax="5.0" metricValueMin="4.05" pageId="4" unit="mm" value="452.5" valueMax="500.0" valueMin="405.0">405500 µm</quantity>
, height/length 0.540.56. Anterior margin approximately evenly curved, more inflated than posterior margin (
<figureCitation id="B9932A32BA41FF9010BBD07C4892FBBC" box="[679,754,1050,1076]" captionStart="FIGURE 1" captionStartId="5.[151,250,1676,1701]" captionTargetBox="[294,1293,179,1649]" captionTargetId="figure@5.[294,1293,179,1649]" captionTargetPageId="5" captionText="FIGURE 1. Ilyocypris japonica Okubo, 1990. Japanese specimens. A, female left valve, external view (LBM 1430009143, sample 6; see Table 1). B, female right valve, external view (LBM 1430009143, sample 6). C, female left valve, internal view (LBM 1430009143, sample 6). D, female right valve, internal view (LBM 1430009143, sample 6). E, female left valve, internal view, detail of postero-ventral margin (LBM 1430009143, sample 6). F, female right valve, internal view, detail of antero-ventral margin (LBM 1430009143, sample 6). G, female left valve, internal view, muscle scars (LBM 1430009143, sample 6). H, male right valve, external view (LBM 1430009141, sample 6). I, male carapace, dorsal view, anterior to right (LBM 1430009160, sample 25). J, female carapace, dorsal view, anterior to right (LBM 1430009142, sample 6). K, female carapace, tuberculated form, right lateral view (LBM 1430009159, sample 23). L, female carapace, tuberculated form, dorsal view, anterior to right (LBM 1430009159, sample 23). M, male left valve, ventral view (LBM 1430009141, sample 6). Scale bar = 500 µm for AD, HM, 174 µm for EG." httpUri="https://zenodo.org/record/3363535/files/figure.png" pageId="4">Figs 1</figureCitation>
&amp;
<figureCitation id="B9932A32BA41FF901102D07C494FFBBC" box="[798,815,1050,1076]" captionStart="FIGURE 2" captionStartId="6.[151,250,1743,1768]" captionTargetBox="[271,1316,181,1717]" captionTargetId="figure@6.[271,1316,181,1718]" captionTargetPageId="6" captionText="FIGURE 2. Ilyocypris japonica Okubo, 1990. Korean specimens. A, female left valve, external view (LBM 1430009147, sample 15, see Table 1). B, female right valve, external view (LBM 1430009147, sample 15). C, female left valve, internal view (LBM 1430009147, sample 15). D, female right valve, internal view (LBM 1430009147, sample 15). E, female left valve, internal view, detail of postero-ventral margin (LBM 1430009147, sample 15). F, female right valve, internal view, detail of anteroventral margin (LBM 1430009147, sample 15). G, female left valve, internal view, muscle scars (LBM 1430009147, sample 15). H, male right valve, external view (LBM 1430009146, sample 15). I, male left valve, external view (LBM 1430009146, sample 15). J, female carapace, tuberculated form, right lateral view (LBM 1430009149, sample 16). K, female carapace, tuberculated form, oblique view of right side (LBM 1430009149, sample 16). Scale bar = 500 µm for AD, HK, 174 µm for EG." httpUri="https://zenodo.org/record/3363537/files/figure.png" pageId="4">2</figureCitation>
). Dorsal margin noticeably sloping down posteriorly, ventral margin distinctly concave. Maximum height at anterior end of hinge. Well-defined bifurcated sulcus near dorsal margin above and slightly anterior of adductor muscle scars. Both valves without outer lists (
<figureCitation id="B9932A32BA41FF9016EFD0044F31FBF4" box="[1267,1361,1122,1148]" captionStart="FIGURE 1" captionStartId="5.[151,250,1676,1701]" captionTargetBox="[294,1293,179,1649]" captionTargetId="figure@5.[294,1293,179,1649]" captionTargetPageId="5" captionText="FIGURE 1. Ilyocypris japonica Okubo, 1990. Japanese specimens. A, female left valve, external view (LBM 1430009143, sample 6; see Table 1). B, female right valve, external view (LBM 1430009143, sample 6). C, female left valve, internal view (LBM 1430009143, sample 6). D, female right valve, internal view (LBM 1430009143, sample 6). E, female left valve, internal view, detail of postero-ventral margin (LBM 1430009143, sample 6). F, female right valve, internal view, detail of antero-ventral margin (LBM 1430009143, sample 6). G, female left valve, internal view, muscle scars (LBM 1430009143, sample 6). H, male right valve, external view (LBM 1430009141, sample 6). I, male carapace, dorsal view, anterior to right (LBM 1430009160, sample 25). J, female carapace, dorsal view, anterior to right (LBM 1430009142, sample 6). K, female carapace, tuberculated form, right lateral view (LBM 1430009159, sample 23). L, female carapace, tuberculated form, dorsal view, anterior to right (LBM 1430009159, sample 23). M, male left valve, ventral view (LBM 1430009141, sample 6). Scale bar = 500 µm for AD, HM, 174 µm for EG." httpUri="https://zenodo.org/record/3363535/files/figure.png" pageId="4">Fig. 1M</figureCitation>
). With or without tubercles; when present, one rounded to angular tubercle each side of sulcus (posterior one called lateral tubercle by some authors), one in mid-position of sulcus, one towards postero-ventral region, and two tubercles in central ventral region (
<figureCitation id="B9932A32BA41FF90138ED0A84878FB60" box="[402,536,1230,1256]" captionStart="FIGURE 2" captionStartId="6.[151,250,1743,1768]" captionTargetBox="[271,1316,181,1717]" captionTargetId="figure@6.[271,1316,181,1718]" captionTargetPageId="6" captionText="FIGURE 2. Ilyocypris japonica Okubo, 1990. Korean specimens. A, female left valve, external view (LBM 1430009147, sample 15, see Table 1). B, female right valve, external view (LBM 1430009147, sample 15). C, female left valve, internal view (LBM 1430009147, sample 15). D, female right valve, internal view (LBM 1430009147, sample 15). E, female left valve, internal view, detail of postero-ventral margin (LBM 1430009147, sample 15). F, female right valve, internal view, detail of anteroventral margin (LBM 1430009147, sample 15). G, female left valve, internal view, muscle scars (LBM 1430009147, sample 15). H, male right valve, external view (LBM 1430009146, sample 15). I, male left valve, external view (LBM 1430009146, sample 15). J, female carapace, tuberculated form, right lateral view (LBM 1430009149, sample 16). K, female carapace, tuberculated form, oblique view of right side (LBM 1430009149, sample 16). Scale bar = 500 µm for AD, HK, 174 µm for EG." httpUri="https://zenodo.org/record/3363537/files/figure.png" pageId="4">Fig. 2J &amp; K</figureCitation>
). Six adductor muscle scars, location visible externally as pit below posterior part of sulcus. Surface of carapace covered with well defined pits, and small spines near anterior and posterior margins. Anterior inner calcified lamellae of both valves with lists. Inner side of lists uneven and with striae. Postero-ventral region of calcified inner lamella of left valve with shallow depression, internally roughened, but not with well-defined marginal ripplets. Dorsal view more rounded posteriorly than anteriorly, maximum width posterior of midlength. Males generally smaller and slightly more elongate than females. Dorsal view of male similar to female, but with more rounded posterior margin and slightly more compressed.
</paragraph>
<paragraph id="211736B7BA41FF9012DBD1AC4E94F91F" blockId="4.[151,1437,401,2045]" pageId="4">
Antennule (
<figureCitation id="B9932A32BA41FF901350D1AC4BC5FA6C" box="[332,421,1482,1508]" captionStart="FIGURE 3" captionStartId="7.[151,250,1839,1864]" captionTargetBox="[239,1349,181,1814]" captionTargetId="figure@7.[239,1349,181,1814]" captionTargetPageId="7" captionText="FIGURE 3. Ilyocypris japonica Okubo, 1990. A, antennule, male (LBM 1430009154). B, antenna, male (LBM 1430009155). C, mandibular palp, male (LBM 1430009158). D, mandibular coxa, male (LBM 1430009155). E, maxillula palp and endites (setae on endites not drawn), male (LBM 1430009154). F, fifth limb, female (LBM 1430009157). G, fifth limb palp, male (LBM 1430009154). H, sixth limb, male (LBM 1430009154), arrow and dashed line indicate length of first endopodal segment. I, caudal ramus, male (LBM 1430009155). All specimens from sample 22; see Table 1." httpUri="https://zenodo.org/record/3363539/files/figure.png" pageId="4">Fig. 3A</figureCitation>
) with seven articulated segments. First segment with two long sub-apical setae on ventral margin and one medium-length seta on dorsal margin. Second segment with relatively long apical-dorsal seta. Third segment with one short apical-dorsal seta and one short apical-ventral seta. Fourth segment with two long apicaldorsal setae, and two shorter apical-ventral setae. Fifth segment with two long apical-dorsal setae, and two shorter, stout, apical-ventral setae. Sixth segment with four long setae and relatively long alpha seta. Terminal segment apically with two short setae, one of which quite stout, one long seta and relatively short aesthetasc ya.
</paragraph>
<paragraph id="211736B7BA41FF9012DBD2C64E99F8AD" blockId="4.[151,1437,401,2045]" pageId="4">
Male antennae with fourth segment undivided (
<figureCitation id="B9932A32BA41FF9010CAD2C6494EF932" box="[726,814,1696,1722]" captionStart="FIGURE 3" captionStartId="7.[151,250,1839,1864]" captionTargetBox="[239,1349,181,1814]" captionTargetId="figure@7.[239,1349,181,1814]" captionTargetPageId="7" captionText="FIGURE 3. Ilyocypris japonica Okubo, 1990. A, antennule, male (LBM 1430009154). B, antenna, male (LBM 1430009155). C, mandibular palp, male (LBM 1430009158). D, mandibular coxa, male (LBM 1430009155). E, maxillula palp and endites (setae on endites not drawn), male (LBM 1430009154). F, fifth limb, female (LBM 1430009157). G, fifth limb palp, male (LBM 1430009154). H, sixth limb, male (LBM 1430009154), arrow and dashed line indicate length of first endopodal segment. I, caudal ramus, male (LBM 1430009155). All specimens from sample 22; see Table 1." httpUri="https://zenodo.org/record/3363539/files/figure.png" pageId="4">Figs 3B</figureCitation>
&amp;
<figureCitation id="B9932A32BA41FF90114DD2C64915F932" box="[849,885,1696,1722]" captionStart="FIGURE 4" captionStartId="8.[151,248,1767,1792]" captionTargetBox="[262,1321,181,1741]" captionTargetId="figure@8.[261,1326,181,1741]" captionTargetPageId="8" captionText="FIGURE 4. AI, Ilyocypris japonica Okubo, 1990. A, detail of female antenna (LBM 1430009156, sample 22; see Table 1). B, detail of male antenna (LBM 1430009155, sample 22). C, seventh limb, male (LBM 1430009154, sample 22). D, outline of female reproductive organ, arrow indicates anterior (LBM 1430009157, sample 22). E, hemipenis (LBM 1430009154, sample 22). F, copulatory process of hemipenis (LBM 1430009154, sample 22). G, hemipenis (LBM 1430009148, sample 15). H, copulatory process of hemipenis (LBM 1430009148, sample 15). I, outlines of the hemipenes of six specimens, some reflected on the vertical axis to facilitate comparisons (left to right LBM 1430009153, sample 22; LBM 1430009152, sample 21; LBM 1430009146, sample 15; LBM 1430009145, sample 10; LBM 1430009140, sample 5; LBM 1430009139, sample 3). J &amp; K, Ilyocypris mongolica. J, hemipenis (LBM 1430009161, sample 17). K, copulatory process of hemipenis (LBM 1430009161, sample 17)." httpUri="https://zenodo.org/record/3363541/files/figure.png" pageId="4">4B</figureCitation>
). Swimming setae very long, extending considerably beyond end of claws; shorter accompanying seta also long, reaching to mid-point of claws. G3 reduced to short seta, but extending slightly beyond end of terminal segment. Claw z1 small and slender, not reaching to end of claw G2. Female antenna claw G2 long, longer than G1 (
<figureCitation id="B9932A32BA41FF9010C7D36D4956F8AD" box="[731,822,1803,1829]" captionStart="FIGURE 4" captionStartId="8.[151,248,1767,1792]" captionTargetBox="[262,1321,181,1741]" captionTargetId="figure@8.[261,1326,181,1741]" captionTargetPageId="8" captionText="FIGURE 4. AI, Ilyocypris japonica Okubo, 1990. A, detail of female antenna (LBM 1430009156, sample 22; see Table 1). B, detail of male antenna (LBM 1430009155, sample 22). C, seventh limb, male (LBM 1430009154, sample 22). D, outline of female reproductive organ, arrow indicates anterior (LBM 1430009157, sample 22). E, hemipenis (LBM 1430009154, sample 22). F, copulatory process of hemipenis (LBM 1430009154, sample 22). G, hemipenis (LBM 1430009148, sample 15). H, copulatory process of hemipenis (LBM 1430009148, sample 15). I, outlines of the hemipenes of six specimens, some reflected on the vertical axis to facilitate comparisons (left to right LBM 1430009153, sample 22; LBM 1430009152, sample 21; LBM 1430009146, sample 15; LBM 1430009145, sample 10; LBM 1430009140, sample 5; LBM 1430009139, sample 3). J &amp; K, Ilyocypris mongolica. J, hemipenis (LBM 1430009161, sample 17). K, copulatory process of hemipenis (LBM 1430009161, sample 17)." httpUri="https://zenodo.org/record/3363541/files/figure.png" pageId="4">Fig. 4A</figureCitation>
). Claw Gm ca. 80% length of claw GM.
</paragraph>
<paragraph id="211736B7BA41FF9012DBD349485CF874" blockId="4.[151,1437,401,2045]" pageId="4">
Mandibular coxa small, with well-developed teeth on endite (
<figureCitation id="B9932A32BA41FF901162D34949B9F8C1" box="[894,985,1839,1865]" captionStart="FIGURE 3" captionStartId="7.[151,250,1839,1864]" captionTargetBox="[239,1349,181,1814]" captionTargetId="figure@7.[239,1349,181,1814]" captionTargetPageId="7" captionText="FIGURE 3. Ilyocypris japonica Okubo, 1990. A, antennule, male (LBM 1430009154). B, antenna, male (LBM 1430009155). C, mandibular palp, male (LBM 1430009158). D, mandibular coxa, male (LBM 1430009155). E, maxillula palp and endites (setae on endites not drawn), male (LBM 1430009154). F, fifth limb, female (LBM 1430009157). G, fifth limb palp, male (LBM 1430009154). H, sixth limb, male (LBM 1430009154), arrow and dashed line indicate length of first endopodal segment. I, caudal ramus, male (LBM 1430009155). All specimens from sample 22; see Table 1." httpUri="https://zenodo.org/record/3363539/files/figure.png" pageId="4">Fig. 3D</figureCitation>
). Mandibular palp four-segmented (
<figureCitation id="B9932A32BA41FF901773D3494ADBF8E5" captionStart="FIGURE 3" captionStartId="7.[151,250,1839,1864]" captionTargetBox="[239,1349,181,1814]" captionTargetId="figure@7.[239,1349,181,1814]" captionTargetPageId="7" captionText="FIGURE 3. Ilyocypris japonica Okubo, 1990. A, antennule, male (LBM 1430009154). B, antenna, male (LBM 1430009155). C, mandibular palp, male (LBM 1430009158). D, mandibular coxa, male (LBM 1430009155). E, maxillula palp and endites (setae on endites not drawn), male (LBM 1430009154). F, fifth limb, female (LBM 1430009157). G, fifth limb palp, male (LBM 1430009154). H, sixth limb, male (LBM 1430009154), arrow and dashed line indicate length of first endopodal segment. I, caudal ramus, male (LBM 1430009155). All specimens from sample 22; see Table 1." httpUri="https://zenodo.org/record/3363539/files/figure.png" pageId="4">Fig. 3C</figureCitation>
). S1 seta on first segment longer than S2, alpha seta small and slender. Second segment with two setae on outer edge, and 3+1+beta setae on inner edge. Beta seta short, stubby and hirsute. Third segment with four setae on outer apical corner, three setae on inner apical corner, and one apical seta between other two groups. Terminal segment with three stout apical setae, two thinner and shorter apical setae and one seta protruding from area proximal of segment mid-length, towards outer edge.
</paragraph>
<caption id="75D7663FBA40FF91128BD2EA4BE8F861" httpUri="https://zenodo.org/record/3363535/files/figure.png" pageId="5" startId="5.[151,250,1676,1701]" targetBox="[294,1293,179,1649]" targetPageId="5">
<paragraph id="211736B7BA40FF91128BD2EA4BE8F861" blockId="5.[151,1437,1676,2025]" pageId="5">
<emphasis id="13DCEAA5BA40FF91128BD2EA4B75F92D" bold="true" box="[151,277,1676,1701]" pageId="5">FIGURE 1.</emphasis>
<taxonomicName id="E6A84D34BA40FF911302D2EB4817F92D" authority="Okubo, 1990" authorityName="Okubo" authorityYear="1990" box="[286,631,1676,1701]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="5" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="13DCEAA5BA40FF911302D2EB4B85F92C" box="[286,485,1677,1701]" italics="true" pageId="5">Ilyocypris japonica</emphasis>
<bibRefCitation id="45394B46BA40FF9113F1D2EA4817F92D" author="Okubo, I." box="[493,631,1676,1701]" pageId="5" pagination="39 - 50" refId="ref22357" refString="Okubo, I. (1990) Sixteen species of freshwater Ostracoda from Japan. Bulletin of the Biogeographical Society of Japan, 45, 39 - 50." type="journal article" year="1990">Okubo, 1990</bibRefCitation>
</taxonomicName>
. Japanese specimens. A, female left valve, external view (LBM 1430009143, sample 6; see Table 1). B, female right valve, external view (LBM 1430009143, sample 6). C, female left valve, internal view (LBM 1430009143, sample 6). D, female right valve, internal view (LBM 1430009143, sample 6). E, female left valve, internal view, detail of postero-ventral margin (LBM 1430009143, sample 6). F, female right valve, internal view, detail of antero-ventral margin (LBM 1430009143, sample 6). G, female left valve, internal view, muscle scars (LBM 1430009143, sample 6). H, male right valve, external view (LBM 1430009141, sample 6). I, male carapace, dorsal view, anterior to right (LBM 1430009160, sample 25). J, female carapace, dorsal view, anterior to right (LBM 1430009142, sample 6). K, female carapace, tuberculated form, right lateral view (LBM 1430009159, sample 23). L, female carapace, tuberculated form, dorsal view, anterior to right (LBM 1430009159, sample 23). M, male left valve, ventral view (LBM 1430009141, sample 6). Scale bar = 500 µm for AD, HM, 174 µm for EG.
</paragraph>
</caption>
<caption id="75D7663FBA43FF92128BD2A94FF7F86C" httpUri="https://zenodo.org/record/3363537/files/figure.png" pageId="6" startId="6.[151,250,1743,1768]" targetBox="[271,1316,181,1717]" targetPageId="6">
<paragraph id="211736B7BA43FF92128BD2A94FF7F86C" blockId="6.[151,1436,1743,2020]" pageId="6">
<emphasis id="13DCEAA5BA43FF92128BD2A94B73F960" bold="true" box="[151,275,1743,1768]" pageId="6">FIGURE 2.</emphasis>
<taxonomicName id="E6A84D34BA43FF921305D2B6480AF960" authority="Okubo, 1990" authorityName="Okubo" authorityYear="1990" box="[281,618,1743,1768]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="6" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="13DCEAA5BA43FF921305D2B64BBDF960" box="[281,477,1744,1768]" italics="true" pageId="6">Ilyocypris japonica</emphasis>
<bibRefCitation id="45394B46BA43FF9213FED2A9480AF960" author="Okubo, I." box="[482,618,1743,1768]" pageId="6" pagination="39 - 50" refId="ref22357" refString="Okubo, I. (1990) Sixteen species of freshwater Ostracoda from Japan. Bulletin of the Biogeographical Society of Japan, 45, 39 - 50." type="journal article" year="1990">Okubo, 1990</bibRefCitation>
</taxonomicName>
. Korean specimens. A, female left valve, external view (LBM 1430009147, sample 15, see Table 1). B, female right valve, external view (LBM 1430009147, sample 15). C, female left valve, internal view (LBM 1430009147, sample 15). D, female right valve, internal view (LBM 1430009147, sample 15). E, female left valve, internal view, detail of postero-ventral margin (LBM 1430009147, sample 15). F, female right valve, internal view, detail of anteroventral margin (LBM 1430009147, sample 15). G, female left valve, internal view, muscle scars (LBM 1430009147, sample 15). H, male right valve, external view (LBM 1430009146, sample 15). I, male left valve, external view (LBM 1430009146, sample 15). J, female carapace, tuberculated form, right lateral view (LBM 1430009149, sample 16). K, female carapace, tuberculated form, oblique view of right side (LBM 1430009149, sample 16). Scale bar = 500 µm for AD, HK, 174 µm for EG.
</paragraph>
</caption>
<caption id="75D7663FBA42FF93128BD3494994F850" httpUri="https://zenodo.org/record/3363539/files/figure.png" pageId="7" startId="7.[151,250,1839,1864]" targetBox="[239,1349,181,1814]" targetPageId="7">
<paragraph id="211736B7BA42FF93128BD3494994F850" blockId="7.[151,1437,1839,2008]" pageId="7">
<emphasis id="13DCEAA5BA42FF93128BD3494B73F8C0" bold="true" box="[151,275,1839,1864]" pageId="7">FIGURE 3.</emphasis>
<taxonomicName id="E6A84D34BA42FF931306D356480EF8C0" authority="Okubo, 1990" authorityName="Okubo" authorityYear="1990" box="[282,622,1839,1864]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="7" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="13DCEAA5BA42FF931306D3564BBFF8C0" box="[282,479,1840,1864]" italics="true" pageId="7">Ilyocypris japonica</emphasis>
<bibRefCitation id="45394B46BA42FF9313FAD349480EF8C0" author="Okubo, I." box="[486,622,1839,1864]" pageId="7" pagination="39 - 50" refId="ref22357" refString="Okubo, I. (1990) Sixteen species of freshwater Ostracoda from Japan. Bulletin of the Biogeographical Society of Japan, 45, 39 - 50." type="journal article" year="1990">Okubo, 1990</bibRefCitation>
</taxonomicName>
. A, antennule, male (LBM 1430009154). B, antenna, male (LBM 1430009155). C, mandibular palp, male (LBM 1430009158). D, mandibular coxa, male (LBM 1430009155). E, maxillula palp and endites (setae on endites not drawn), male (LBM 1430009154). F, fifth limb, female (LBM 1430009157). G, fifth limb palp, male (LBM 1430009154). H, sixth limb, male (LBM 1430009154), arrow and dashed line indicate length of first endopodal segment. I, caudal ramus, male (LBM 1430009155). All specimens from sample 22; see Table 1.
</paragraph>
</caption>
<caption id="75D7663FBA4DFF9C128BD2814E8FF873" httpUri="https://zenodo.org/record/3363541/files/figure.png" pageId="8" startId="8.[151,248,1767,1792]" targetBox="[262,1321,181,1741]" targetPageId="8">
<paragraph id="211736B7BA4DFF9C128BD2814E8FF873" blockId="8.[151,1437,1767,2044]" pageId="8">
<emphasis id="13DCEAA5BA4DFF9C128BD2814B72F888" bold="true" box="[151,274,1767,1792]" pageId="8">FIGURE 4.</emphasis>
AI,
<taxonomicName id="E6A84D34BA4DFF9C1351D28148FBF977" authority="Okubo, 1990" authorityName="Okubo" authorityYear="1990" box="[333,667,1767,1792]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="8" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="13DCEAA5BA4DFF9C1351D281486EF977" box="[333,526,1767,1791]" italics="true" pageId="8">Ilyocypris japonica</emphasis>
<bibRefCitation id="45394B46BA4DFF9C1009D28148FBF977" author="Okubo, I." box="[533,667,1767,1792]" pageId="8" pagination="39 - 50" refId="ref22357" refString="Okubo, I. (1990) Sixteen species of freshwater Ostracoda from Japan. Bulletin of the Biogeographical Society of Japan, 45, 39 - 50." type="journal article" year="1990">Okubo, 1990</bibRefCitation>
</taxonomicName>
. A, detail of female antenna (LBM 1430009156, sample 22; see Table 1). B, detail of male antenna (LBM 1430009155, sample 22). C, seventh limb, male (LBM 1430009154, sample 22). D, outline of female reproductive organ, arrow indicates anterior (LBM 1430009157, sample 22). E, hemipenis (LBM 1430009154, sample 22). F, copulatory process of hemipenis (LBM 1430009154, sample 22). G, hemipenis (LBM 1430009148, sample 15). H, copulatory process of hemipenis (LBM 1430009148, sample 15). I, outlines of the hemipenes of six specimens, some reflected on the vertical axis to facilitate comparisons (left to right LBM 1430009153, sample 22; LBM 1430009152, sample 21; LBM 1430009146, sample 15; LBM 1430009145, sample 10; LBM 1430009140, sample 5; LBM 1430009139, sample 3). J &amp; K,
<taxonomicName id="E6A84D34BA4DFF9C16ACD3D94FE1F85F" authorityName="Martens" authorityYear="1991" box="[1200,1409,1983,2007]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="8" phylum="Arthropoda" rank="species" species="mongolica">
<emphasis id="13DCEAA5BA4DFF9C16ACD3D94FE1F85F" box="[1200,1409,1983,2007]" italics="true" pageId="8">Ilyocypris mongolica</emphasis>
</taxonomicName>
. J, hemipenis (LBM 1430009161, sample 17). K, copulatory process of hemipenis (LBM 1430009161, sample 17).
</paragraph>
</caption>
<caption id="75D7663FBA4CFF9D128BD1A54B8BF9E4" httpUri="https://zenodo.org/record/3363543/files/figure.png" pageId="9" startId="9.[151,250,1475,1500]" targetBox="[151,1436,181,1446]" targetPageId="9">
<paragraph id="211736B7BA4CFF9D128BD1A54B8BF9E4" blockId="9.[151,1436,1475,1644]" pageId="9">
<emphasis id="13DCEAA5BA4CFF9D128BD1A54B73FA54" bold="true" box="[151,275,1475,1500]" pageId="9">FIGURE 5.</emphasis>
Light microscopy photographs of
<taxonomicName id="E6A84D34BA4CFF9D106AD1A548BBFA53" authorityName="Brady &amp; Norman" authorityYear="1889" box="[630,731,1475,1499]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="9" phylum="Arthropoda" rank="genus">
<emphasis id="13DCEAA5BA4CFF9D106AD1A548BBFA53" box="[630,731,1475,1499]" italics="true" pageId="9">Ilyocypris</emphasis>
</taxonomicName>
hemipenes. A,
<taxonomicName id="E6A84D34BA4CFF9D1165D1A54EAAFA54" authority="Okubo, 1990" authorityName="Okubo" authorityYear="1990" box="[889,1226,1475,1500]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="9" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="13DCEAA5BA4CFF9D1165D1A54E5FFA53" box="[889,1087,1475,1499]" italics="true" pageId="9">Ilyocypris japonica</emphasis>
<bibRefCitation id="45394B46BA4CFF9D1659D1A54EAAFA54" author="Okubo, I." box="[1093,1226,1475,1500]" pageId="9" pagination="39 - 50" refId="ref22357" refString="Okubo, I. (1990) Sixteen species of freshwater Ostracoda from Japan. Bulletin of the Biogeographical Society of Japan, 45, 39 - 50." type="journal article" year="1990">Okubo, 1990</bibRefCitation>
</taxonomicName>
(LBM 1430009150, sample 18; see Table 1). B,
<taxonomicName id="E6A84D34BA4CFF9D13B0D181489AF988" authority="Okubo, 1990" authorityName="Okubo" authorityYear="1990" box="[428,762,1511,1536]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="9" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="13DCEAA5BA4CFF9D13B0D1814810FA77" box="[428,624,1511,1535]" italics="true" pageId="9">Ilyocypris japonica</emphasis>
<bibRefCitation id="45394B46BA4CFF9D106AD181489AF988" author="Okubo, I." box="[630,762,1511,1536]" pageId="9" pagination="39 - 50" refId="ref22357" refString="Okubo, I. (1990) Sixteen species of freshwater Ostracoda from Japan. Bulletin of the Biogeographical Society of Japan, 45, 39 - 50." type="journal article" year="1990">Okubo, 1990</bibRefCitation>
</taxonomicName>
(LBM 1430009151, sample 18). C,
<taxonomicName id="E6A84D34BA4CFF9D167BD1814AAAF9AC" authority="Martens, 1991" authorityName="Martens" authorityYear="1991" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="9" phylum="Arthropoda" rank="species" species="mongolica">
<emphasis id="13DCEAA5BA4CFF9D167BD1814F5DFA77" box="[1127,1341,1511,1535]" italics="true" pageId="9">Ilyocypris mongolica</emphasis>
<bibRefCitation id="45394B46BA4CFF9D175FD18E4AAAF9AC" author="Martens, K." pageId="9" pagination="53 - 60" refId="ref21598" refString="Martens, K. (1991) On a small collection of non-marine ostracods from Mongolia, with the description of a new species (Crustacea, Ostracoda). Miscellanea Zoologica Hungarica, 6, 53 - 60." type="journal article" year="1991">Martens, 1991</bibRefCitation>
</taxonomicName>
(LBM 1430009161, sample 17). D,
<taxonomicName id="E6A84D34BA4CFF9D1020D26D4915F9AC" authority="Sars, 1903" authorityName="G. O. Sars-Chen" authorityYear="1903" box="[572,885,1547,1572]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="9" phylum="Arthropoda" rank="species" species="dentifera">
<emphasis id="13DCEAA5BA4CFF9D1020D26D4963F9AB" box="[572,771,1547,1571]" italics="true" pageId="9">Ilyocypris dentifera</emphasis>
<bibRefCitation id="45394B46BA4CFF9D1116D26A4915F9AC" author="Sars, G. O." box="[778,885,1548,1572]" pageId="9" pagination="3 - 44" refId="ref22910" refString="Sars, G. O. (1903) Fresh-water Entomostraca from China and Sumatra. Archiv for Mathematik og Naturvidenskab, 25, 3 - 44." type="journal article" year="1903">Sars, 1903</bibRefCitation>
</taxonomicName>
(LBM 1430009119, sample 6). E,
<taxonomicName id="E6A84D34BA4CFF9D16CBD26D4B62F9C0" authority="Sars, 1903" authorityName="Sars" authorityYear="1903" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="9" phylum="Arthropoda" rank="species" species="angulata">
<emphasis id="13DCEAA5BA4CFF9D16CBD26D4FFCF9AB" box="[1239,1436,1547,1571]" italics="true" pageId="9">Ilyocypris angulata</emphasis>
<bibRefCitation id="45394B46BA4CFF9D128BD2564B62F9C0" author="Sars, G. O." box="[151,258,1584,1608]" pageId="9" pagination="3 - 44" refId="ref22910" refString="Sars, G. O. (1903) Fresh-water Entomostraca from China and Sumatra. Archiv for Mathematik og Naturvidenskab, 25, 3 - 44." type="journal article" year="1903">Sars, 1903</bibRefCitation>
</taxonomicName>
(LBM 1430009112, sample 20). F,
<taxonomicName id="E6A84D34BA4CFF9D1074D2494967F9CF" authority="Smith &amp; Zhai &amp; Chang, 2019" authorityName="Smith &amp; Zhai &amp; Chang" authorityYear="2019" box="[616,775,1583,1607]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="9" phylum="Arthropoda" rank="species" species="incus" status="sp. nov.">
<emphasis id="13DCEAA5BA4CFF9D1074D2494967F9CF" box="[616,775,1583,1607]" italics="true" pageId="9">Ilyocypris incus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="08EF57DEBA4CFF9D1111D256493CF9CF" box="[781,860,1584,1607]" pageId="9" rank="species">sp. nov.</taxonomicNameLabel>
(DB40054, sample 14). E has been reflected on the vertical axis to facilitate comparisons.
</paragraph>
</caption>
<paragraph id="211736B7BA4CFF9D12DBD2FC4E22F974" blockId="9.[151,1437,1690,2040]" pageId="9">
Maxillula palp first segment with three sub-equal, medium-length setae on outer apical corner, one shorter seta positioned on apical edge offset from apical corner, and one sub-apical seta in mid-width position (
<figureCitation id="B9932A32BA4CFF9D16FCD2D84F57F950" box="[1248,1335,1726,1752]" captionStart="FIGURE 3" captionStartId="7.[151,250,1839,1864]" captionTargetBox="[239,1349,181,1814]" captionTargetId="figure@7.[239,1349,181,1814]" captionTargetPageId="7" captionText="FIGURE 3. Ilyocypris japonica Okubo, 1990. A, antennule, male (LBM 1430009154). B, antenna, male (LBM 1430009155). C, mandibular palp, male (LBM 1430009158). D, mandibular coxa, male (LBM 1430009155). E, maxillula palp and endites (setae on endites not drawn), male (LBM 1430009154). F, fifth limb, female (LBM 1430009157). G, fifth limb palp, male (LBM 1430009154). H, sixth limb, male (LBM 1430009154), arrow and dashed line indicate length of first endopodal segment. I, caudal ramus, male (LBM 1430009155). All specimens from sample 22; see Table 1." httpUri="https://zenodo.org/record/3363539/files/figure.png" pageId="9">Fig. 3E</figureCitation>
). Second segment wider than long, apically with three claw-like setae and three shorter setae.
</paragraph>
<paragraph id="211736B7BA4CFF9D12DBD3604E26F8E0" blockId="9.[151,1437,1690,2040]" pageId="9">
Fifth limb with two a setae on dorsal margin, and approximately 17 setae along dorsal and anterior edge of basis (
<figureCitation id="B9932A32BA4CFF9D12BCD34C4A94F8CC" box="[160,244,1834,1860]" captionStart="FIGURE 3" captionStartId="7.[151,250,1839,1864]" captionTargetBox="[239,1349,181,1814]" captionTargetId="figure@7.[239,1349,181,1814]" captionTargetPageId="7" captionText="FIGURE 3. Ilyocypris japonica Okubo, 1990. A, antennule, male (LBM 1430009154). B, antenna, male (LBM 1430009155). C, mandibular palp, male (LBM 1430009158). D, mandibular coxa, male (LBM 1430009155). E, maxillula palp and endites (setae on endites not drawn), male (LBM 1430009154). F, fifth limb, female (LBM 1430009157). G, fifth limb palp, male (LBM 1430009154). H, sixth limb, male (LBM 1430009154), arrow and dashed line indicate length of first endopodal segment. I, caudal ramus, male (LBM 1430009155). All specimens from sample 22; see Table 1." httpUri="https://zenodo.org/record/3363539/files/figure.png" pageId="9">Fig. 3F</figureCitation>
). Branchial plate with six rays. Palp (endopodite) of female two-segmented, terminating with three setae of differing lengths. Palps of male symmetrical, with slender elongate hooks (
<figureCitation id="B9932A32BA4CFF9D11C3D3284E59F8E0" box="[991,1081,1870,1896]" captionStart="FIGURE 3" captionStartId="7.[151,250,1839,1864]" captionTargetBox="[239,1349,181,1814]" captionTargetId="figure@7.[239,1349,181,1814]" captionTargetPageId="7" captionText="FIGURE 3. Ilyocypris japonica Okubo, 1990. A, antennule, male (LBM 1430009154). B, antenna, male (LBM 1430009155). C, mandibular palp, male (LBM 1430009158). D, mandibular coxa, male (LBM 1430009155). E, maxillula palp and endites (setae on endites not drawn), male (LBM 1430009154). F, fifth limb, female (LBM 1430009157). G, fifth limb palp, male (LBM 1430009154). H, sixth limb, male (LBM 1430009154), arrow and dashed line indicate length of first endopodal segment. I, caudal ramus, male (LBM 1430009155). All specimens from sample 22; see Table 1." httpUri="https://zenodo.org/record/3363539/files/figure.png" pageId="9">Fig. 3G</figureCitation>
).
</paragraph>
<paragraph id="211736B7BA4CFF9D12DBD3144F22F870" blockId="9.[151,1437,1690,2040]" pageId="9">
Sixth limb five-segmented (
<figureCitation id="B9932A32BA4CFF9D13E2D3144838F804" box="[510,600,1906,1932]" captionStart="FIGURE 3" captionStartId="7.[151,250,1839,1864]" captionTargetBox="[239,1349,181,1814]" captionTargetId="figure@7.[239,1349,181,1814]" captionTargetPageId="7" captionText="FIGURE 3. Ilyocypris japonica Okubo, 1990. A, antennule, male (LBM 1430009154). B, antenna, male (LBM 1430009155). C, mandibular palp, male (LBM 1430009158). D, mandibular coxa, male (LBM 1430009155). E, maxillula palp and endites (setae on endites not drawn), male (LBM 1430009154). F, fifth limb, female (LBM 1430009157). G, fifth limb palp, male (LBM 1430009154). H, sixth limb, male (LBM 1430009154), arrow and dashed line indicate length of first endopodal segment. I, caudal ramus, male (LBM 1430009155). All specimens from sample 22; see Table 1." httpUri="https://zenodo.org/record/3363539/files/figure.png" pageId="9">Fig. 3H</figureCitation>
). First segment (= protopodite) with short d1 seta. Second, third and fourth segments (= first, second and third endopodal segments respectively) with short e, f and g setae respectively. Terminal segment (= fourth endopodal segment) with relatively long h1 seta, short h3 and well-developed h2 claw, 1.9x length of anterior sclerotized margin of first endopodal segment (indicated with black triangle on
<figureCitation id="B9932A32BA4CFF9D16C0D3B84F55F870" box="[1244,1333,2014,2040]" captionStart="FIGURE 3" captionStartId="7.[151,250,1839,1864]" captionTargetBox="[239,1349,181,1814]" captionTargetId="figure@7.[239,1349,181,1814]" captionTargetPageId="7" captionText="FIGURE 3. Ilyocypris japonica Okubo, 1990. A, antennule, male (LBM 1430009154). B, antenna, male (LBM 1430009155). C, mandibular palp, male (LBM 1430009158). D, mandibular coxa, male (LBM 1430009155). E, maxillula palp and endites (setae on endites not drawn), male (LBM 1430009154). F, fifth limb, female (LBM 1430009157). G, fifth limb palp, male (LBM 1430009154). H, sixth limb, male (LBM 1430009154), arrow and dashed line indicate length of first endopodal segment. I, caudal ramus, male (LBM 1430009155). All specimens from sample 22; see Table 1." httpUri="https://zenodo.org/record/3363539/files/figure.png" pageId="9">Fig. 3H</figureCitation>
).
</paragraph>
<paragraph id="211736B7BA4FFF9E12DBD4FF4F3FFF73" blockId="10.[151,1436,153,323]" pageId="10">
Seventh limb with four segments (
<figureCitation id="B9932A32BA4FFF9E1052D4FF48C8FF3B" box="[590,680,153,179]" captionStart="FIGURE 4" captionStartId="8.[151,248,1767,1792]" captionTargetBox="[262,1321,181,1741]" captionTargetId="figure@8.[261,1326,181,1741]" captionTargetPageId="8" captionText="FIGURE 4. AI, Ilyocypris japonica Okubo, 1990. A, detail of female antenna (LBM 1430009156, sample 22; see Table 1). B, detail of male antenna (LBM 1430009155, sample 22). C, seventh limb, male (LBM 1430009154, sample 22). D, outline of female reproductive organ, arrow indicates anterior (LBM 1430009157, sample 22). E, hemipenis (LBM 1430009154, sample 22). F, copulatory process of hemipenis (LBM 1430009154, sample 22). G, hemipenis (LBM 1430009148, sample 15). H, copulatory process of hemipenis (LBM 1430009148, sample 15). I, outlines of the hemipenes of six specimens, some reflected on the vertical axis to facilitate comparisons (left to right LBM 1430009153, sample 22; LBM 1430009152, sample 21; LBM 1430009146, sample 15; LBM 1430009145, sample 10; LBM 1430009140, sample 5; LBM 1430009139, sample 3). J &amp; K, Ilyocypris mongolica. J, hemipenis (LBM 1430009161, sample 17). K, copulatory process of hemipenis (LBM 1430009161, sample 17)." httpUri="https://zenodo.org/record/3363541/files/figure.png" pageId="10">Fig. 4C</figureCitation>
). One seta (d1) on basal segment. Second segment with one apical seta (e). Third segment with two setae on outer edge (f and g) (junior synonym
<taxonomicName id="E6A84D34BA4FFF9E1600D4D84EA9FF5F" authorityName="Okubo" authorityYear="1992" box="[1052,1225,189,215]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="10" phylum="Arthropoda" rank="species" species="haterumensis">
<emphasis id="13DCEAA5BA4FFF9E1600D4D84EA9FF5F" box="[1052,1225,189,215]" italics="true" pageId="10">I. haterumensis</emphasis>
</taxonomicName>
figured with one), and group of hook-like setules apically. Fourth segment with two long and one medium-length setae apically.
</paragraph>
<paragraph id="211736B7BA4FFF9E12DBD5634AB0FECB" blockId="10.[151,1436,153,323]" pageId="10">
Caudal ramus typical of genus, ramus curved (
<figureCitation id="B9932A32BA4FFF9E10F8D5634955FE97" box="[740,821,261,287]" captionStart="FIGURE 3" captionStartId="7.[151,250,1839,1864]" captionTargetBox="[239,1349,181,1814]" captionTargetId="figure@7.[239,1349,181,1814]" captionTargetPageId="7" captionText="FIGURE 3. Ilyocypris japonica Okubo, 1990. A, antennule, male (LBM 1430009154). B, antenna, male (LBM 1430009155). C, mandibular palp, male (LBM 1430009158). D, mandibular coxa, male (LBM 1430009155). E, maxillula palp and endites (setae on endites not drawn), male (LBM 1430009154). F, fifth limb, female (LBM 1430009157). G, fifth limb palp, male (LBM 1430009154). H, sixth limb, male (LBM 1430009154), arrow and dashed line indicate length of first endopodal segment. I, caudal ramus, male (LBM 1430009155). All specimens from sample 22; see Table 1." httpUri="https://zenodo.org/record/3363539/files/figure.png" pageId="10">
Fig.
<date id="55161077BA4FFF9E1107D5634955FE97" box="[795,821,261,287]" pageId="10">3I</date>
</figureCitation>
). Claw Gp approximately same length as Ga, seta sp long.
</paragraph>
<caption id="75D7663FBA4FFF9E128BD2A7495FF8AA" httpUri="https://zenodo.org/record/3363545/files/figure.png" pageId="10" startId="10.[151,250,1729,1754]" targetBox="[151,1436,345,1704]" targetPageId="10">
<paragraph id="211736B7BA4FFF9E128BD2A7495FF8AA" blockId="10.[151,1436,1729,1826]" pageId="10">
<emphasis id="13DCEAA5BA4FFF9E128BD2A74B74F952" bold="true" box="[151,276,1729,1754]" pageId="10">FIGURE 6.</emphasis>
Light microscopy photographs of
<taxonomicName id="E6A84D34BA4FFF9E1064D2A4494CF952" authority="Zenker" authorityName="Zenker" box="[632,812,1729,1754]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="10" phylum="Arthropoda" rank="genus">
<emphasis id="13DCEAA5BA4FFF9E1064D2A448BDF952" box="[632,733,1730,1754]" italics="true" pageId="10">Ilyocypris</emphasis>
Zenker
</taxonomicName>
organs. Black triangles mark the anterior ends. A,
<taxonomicName id="E6A84D34BA4FFF9E172BD2A44B1BF976" authority="Okubo, 1990" authorityName="Okubo" authorityYear="1990" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="10" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="13DCEAA5BA4FFF9E172BD2A44A90F976" italics="true" pageId="10">Ilyocypris japonica</emphasis>
<bibRefCitation id="45394B46BA4FFF9E12EAD2834B1BF976" author="Okubo, I." box="[246,379,1765,1790]" pageId="10" pagination="39 - 50" refId="ref22357" refString="Okubo, I. (1990) Sixteen species of freshwater Ostracoda from Japan. Bulletin of the Biogeographical Society of Japan, 45, 39 - 50." type="journal article" year="1990">Okubo, 1990</bibRefCitation>
</taxonomicName>
(LBM 1430009144, sample 9; see Table 1). B,
<taxonomicName id="E6A84D34BA4FFF9E1141D2804EF5F976" authority="Sars, 1903" authorityName="G. O. Sars-Chen" authorityYear="1903" box="[861,1173,1766,1790]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="10" phylum="Arthropoda" rank="species" species="dentifera">
<emphasis id="13DCEAA5BA4FFF9E1141D2804E43F976" box="[861,1059,1766,1790]" italics="true" pageId="10">Ilyocypris dentifera</emphasis>
<bibRefCitation id="45394B46BA4FFF9E1636D2804EF5F976" author="Sars, G. O." box="[1066,1173,1766,1790]" pageId="10" pagination="3 - 44" refId="ref22910" refString="Sars, G. O. (1903) Fresh-water Entomostraca from China and Sumatra. Archiv for Mathematik og Naturvidenskab, 25, 3 - 44." type="journal article" year="1903">Sars, 1903</bibRefCitation>
</taxonomicName>
(LBM 1430009119, sam- ple 6). C,
<taxonomicName id="E6A84D34BA4FFF9E12E0D36C4BFCF8AA" authority="Smith &amp; Zhai &amp; Chang, 2019" authorityName="Smith &amp; Zhai &amp; Chang" authorityYear="2019" box="[252,412,1802,1826]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="10" phylum="Arthropoda" rank="species" species="incus" status="sp. nov.">
<emphasis id="13DCEAA5BA4FFF9E12E0D36C4BFCF8AA" box="[252,412,1802,1826]" italics="true" pageId="10">Ilyocypris incus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="08EF57DEBA4FFF9E13BFD36C4B92F8A9" box="[419,498,1802,1825]" pageId="10" rank="species">sp. nov.</taxonomicNameLabel>
(LBM 1430009131, sample 14).
</paragraph>
</caption>
<paragraph id="211736B7BA4FFF9E12DBD33649D9F806" blockId="10.[151,1437,1872,2043]" pageId="10">
Female reproductive organ with one flattened lobe, sub-angular towards anterior and posterior corners (
<figureCitation id="B9932A32BA4FFF9E1773D3364ADEF806" captionStart="FIGURE 4" captionStartId="8.[151,248,1767,1792]" captionTargetBox="[262,1321,181,1741]" captionTargetId="figure@8.[261,1326,181,1741]" captionTargetPageId="8" captionText="FIGURE 4. AI, Ilyocypris japonica Okubo, 1990. A, detail of female antenna (LBM 1430009156, sample 22; see Table 1). B, detail of male antenna (LBM 1430009155, sample 22). C, seventh limb, male (LBM 1430009154, sample 22). D, outline of female reproductive organ, arrow indicates anterior (LBM 1430009157, sample 22). E, hemipenis (LBM 1430009154, sample 22). F, copulatory process of hemipenis (LBM 1430009154, sample 22). G, hemipenis (LBM 1430009148, sample 15). H, copulatory process of hemipenis (LBM 1430009148, sample 15). I, outlines of the hemipenes of six specimens, some reflected on the vertical axis to facilitate comparisons (left to right LBM 1430009153, sample 22; LBM 1430009152, sample 21; LBM 1430009146, sample 15; LBM 1430009145, sample 10; LBM 1430009140, sample 5; LBM 1430009139, sample 3). J &amp; K, Ilyocypris mongolica. J, hemipenis (LBM 1430009161, sample 17). K, copulatory process of hemipenis (LBM 1430009161, sample 17)." httpUri="https://zenodo.org/record/3363541/files/figure.png" pageId="10">Fig. 4D</figureCitation>
). Organ delicate and easily distorted or damaged during dissections.
</paragraph>
<paragraph id="211736B7BA4FFF9F12DBD3FE4966FF5F" blockId="10.[151,1437,1872,2043]" lastBlockId="11.[151,1437,153,324]" lastPageId="11" pageId="10">
Hemipenes with outer lobe rounded distally, overlapping middle lobe (
<figureCitation id="B9932A32BA4FFF9E11C0D3FE4E42F83A" box="[988,1058,1944,1970]" captionStart="FIGURE 4" captionStartId="8.[151,248,1767,1792]" captionTargetBox="[262,1321,181,1741]" captionTargetId="figure@8.[261,1326,181,1741]" captionTargetPageId="8" captionText="FIGURE 4. AI, Ilyocypris japonica Okubo, 1990. A, detail of female antenna (LBM 1430009156, sample 22; see Table 1). B, detail of male antenna (LBM 1430009155, sample 22). C, seventh limb, male (LBM 1430009154, sample 22). D, outline of female reproductive organ, arrow indicates anterior (LBM 1430009157, sample 22). E, hemipenis (LBM 1430009154, sample 22). F, copulatory process of hemipenis (LBM 1430009154, sample 22). G, hemipenis (LBM 1430009148, sample 15). H, copulatory process of hemipenis (LBM 1430009148, sample 15). I, outlines of the hemipenes of six specimens, some reflected on the vertical axis to facilitate comparisons (left to right LBM 1430009153, sample 22; LBM 1430009152, sample 21; LBM 1430009146, sample 15; LBM 1430009145, sample 10; LBM 1430009140, sample 5; LBM 1430009139, sample 3). J &amp; K, Ilyocypris mongolica. J, hemipenis (LBM 1430009161, sample 17). K, copulatory process of hemipenis (LBM 1430009161, sample 17)." httpUri="https://zenodo.org/record/3363541/files/figure.png" pageId="10">Figs 4</figureCitation>
EI, black triangle indicates overlap, 5A &amp; B). Middle lobe sub-quadrate to triangular, with thick base, widening distally, distal inner corner sharply angled, outer edge angled to rounded. Inner lobe long and thin, protruding beyond outer lobe, terminating with bul- bous end; distally variable, ranging from narrow and hook-like to slightly thicker and bulbous. Copulatory process slender, with curved outer edge and sharply pointed tip.
</paragraph>
<paragraph id="211736B7BA4EFF9F12DBD4874A8CFECC" blockId="11.[151,1437,153,324]" pageId="11">
Zenker organ with proximal end significantly inflated compared with posterior end, and with 1517 internal rosettes (
<figureCitation id="B9932A32BA4EFF9F12E1D5634B38FEA8" box="[253,344,261,288]" captionStart="FIGURE 6" captionStartId="10.[151,250,1729,1754]" captionTargetBox="[151,1436,345,1704]" captionTargetId="figure@10.[151,1436,345,1704]" captionTargetPageId="10" captionText="FIGURE 6. Light microscopy photographs of Ilyocypris Zenker organs. Black triangles mark the anterior ends. A, Ilyocypris japonica Okubo, 1990 (LBM 1430009144, sample 9; see Table 1). B, Ilyocypris dentifera Sars, 1903 (LBM 1430009119, sam- ple 6). C, Ilyocypris incus sp. nov. (LBM 1430009131, sample 14)." httpUri="https://zenodo.org/record/3363545/files/figure.png" pageId="11">Fig. 6A</figureCitation>
). Spermatozoa lengths ranging from
<quantity id="E6509B52BA4EFF9F10E0D56349A0FE97" box="[764,960,261,287]" metricMagnitude="0" metricUnit="m" metricValue="1.1195" metricValueMax="1.232" metricValueMin="1.007" pageId="11" unit="mm" value="1119.5" valueMax="1232.0" valueMin="1007.0">
<date id="55161077BA4EFF9F10E0D56349F2FE97" box="[764,914,261,287]" pageId="11" value="1007" valueMax="1232">1007 to 1232</date>
µm
</quantity>
with an average of
<quantity id="E6509B52BA4EFF9F16BED5634F67FE97" box="[1186,1287,261,287]" metricMagnitude="0" metricUnit="m" metricValue="1.139" pageId="11" unit="mm" value="1139.0">1139 µm</quantity>
(
<bibRefCitation id="45394B46BA4EFF9F170BD5634A81FECC" author="Smith, R. J. &amp; Matzke-Karasz, R. &amp; Kamiya, T." pageId="11" pagination="483 - 497" refId="ref23242" refString="Smith, R. J., Matzke-Karasz, R. &amp; Kamiya, T. (2016 a) Sperm length variations in five species of cypridoidean non-marine ostracods (Crustacea). Cell and Tissue Research, 366, 483 - 497. https: // doi. org / 10.1007 / s 00441 - 016 - 2459 - x" type="journal article" year="2016">
Smith
<emphasis id="13DCEAA5BA4EFF9F177FD5604FF5FE97" box="[1379,1429,261,287]" italics="true" pageId="11">et al</emphasis>
. 2016a
</bibRefCitation>
).
</paragraph>
<caption id="75D7663FBA4EFF9F128BD1444907FA2F" httpUri="https://zenodo.org/record/3363547/files/figure.png" pageId="11" startId="11.[151,250,1314,1339]" targetBox="[151,1434,345,1289]" targetPageId="11">
<paragraph id="211736B7BA4EFF9F128BD1444907FA2F" blockId="11.[151,1437,1314,1448]" pageId="11">
<emphasis id="13DCEAA5BA4EFF9F128BD1444B74FAB3" bold="true" box="[151,276,1314,1339]" pageId="11">FIGURE 7.</emphasis>
Two male specimens of
<taxonomicName id="E6A84D34BA4EFF9F100FD14548B8FAB3" authorityName="Okubo" authorityYear="1990" box="[531,728,1315,1339]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="11" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="13DCEAA5BA4EFF9F100FD14548B8FAB3" box="[531,728,1315,1339]" italics="true" pageId="11">Ilyocypris japonica</emphasis>
</taxonomicName>
from the same sample (Lake Biwa Museum rice field, 14 June 2013, sample 18; see Table 1), showing the two size types, large and small. A (LBM 1430009151) &amp; B (LBM 1430009150), external view of right valves. C (LBM 1430009151) &amp; D (LBM 1430009150), details of surface of valves, locations marked by dotted rectangles on A &amp; B. Scale bar = 500 µm for A &amp; B, 94 µm for C &amp; D.
</paragraph>
</caption>
<paragraph id="211736B7BA4EFF9F12DBD1B34B74F8BB" blockId="11.[151,1437,1493,1988]" pageId="11">
<emphasis id="13DCEAA5BA4EFF9F12DBD1B34BBBFA78" bold="true" box="[199,475,1493,1520]" pageId="11">Intraspecific variation.</emphasis>
13% of specimens studied have tubercles present on each valve (see description for more details) (
<tableCitation id="6C2A030CBA4EFF9F1320D19F4BF1F99C" box="[316,401,1529,1556]" captionStart="TABLE 1" captionStartId="1.[151,244,1160,1187]" captionTargetBox="[163,1412,1310,1957]" captionText="TABLE 1. Adult Ilyocypris specimens examined for this study. Initial discriminations of species were based on lateral views of carapaces, and confirmed by dissections of male specimens. Figures in brackets indicate the number of specimens with tubercles on the carapace. Note that tubercles vary from weakly to well-developed and were no- ticeable on whole specimens or valves in dorsal view using a stereo optical microscope." httpUri="http://table.plazi.org/id/75D7663FBA44FF95128BD0EE4E0FFA87" pageId="11" tableUuid="75D7663FBA44FF95128BD0EE4E0FFA87">Table 1</tableCitation>
), ranging from slightly to well developed, rounded to angular (
<figureCitation id="B9932A32BA4EFF9F167BD19F4ED0F99C" box="[1127,1200,1529,1556]" captionStart="FIGURE 1" captionStartId="5.[151,250,1676,1701]" captionTargetBox="[294,1293,179,1649]" captionTargetId="figure@5.[294,1293,179,1649]" captionTargetPageId="5" captionText="FIGURE 1. Ilyocypris japonica Okubo, 1990. Japanese specimens. A, female left valve, external view (LBM 1430009143, sample 6; see Table 1). B, female right valve, external view (LBM 1430009143, sample 6). C, female left valve, internal view (LBM 1430009143, sample 6). D, female right valve, internal view (LBM 1430009143, sample 6). E, female left valve, internal view, detail of postero-ventral margin (LBM 1430009143, sample 6). F, female right valve, internal view, detail of antero-ventral margin (LBM 1430009143, sample 6). G, female left valve, internal view, muscle scars (LBM 1430009143, sample 6). H, male right valve, external view (LBM 1430009141, sample 6). I, male carapace, dorsal view, anterior to right (LBM 1430009160, sample 25). J, female carapace, dorsal view, anterior to right (LBM 1430009142, sample 6). K, female carapace, tuberculated form, right lateral view (LBM 1430009159, sample 23). L, female carapace, tuberculated form, dorsal view, anterior to right (LBM 1430009159, sample 23). M, male left valve, ventral view (LBM 1430009141, sample 6). Scale bar = 500 µm for AD, HM, 174 µm for EG." httpUri="https://zenodo.org/record/3363535/files/figure.png" pageId="11">Figs 1</figureCitation>
&amp;
<figureCitation id="B9932A32BA4EFF9F16CBD19C4E88F99C" box="[1239,1256,1530,1556]" captionStart="FIGURE 2" captionStartId="6.[151,250,1743,1768]" captionTargetBox="[271,1316,181,1717]" captionTargetId="figure@6.[271,1316,181,1718]" captionTargetPageId="6" captionText="FIGURE 2. Ilyocypris japonica Okubo, 1990. Korean specimens. A, female left valve, external view (LBM 1430009147, sample 15, see Table 1). B, female right valve, external view (LBM 1430009147, sample 15). C, female left valve, internal view (LBM 1430009147, sample 15). D, female right valve, internal view (LBM 1430009147, sample 15). E, female left valve, internal view, detail of postero-ventral margin (LBM 1430009147, sample 15). F, female right valve, internal view, detail of anteroventral margin (LBM 1430009147, sample 15). G, female left valve, internal view, muscle scars (LBM 1430009147, sample 15). H, male right valve, external view (LBM 1430009146, sample 15). I, male left valve, external view (LBM 1430009146, sample 15). J, female carapace, tuberculated form, right lateral view (LBM 1430009149, sample 16). K, female carapace, tuberculated form, oblique view of right side (LBM 1430009149, sample 16). Scale bar = 500 µm for AD, HK, 174 µm for EG." httpUri="https://zenodo.org/record/3363537/files/figure.png" pageId="11">2</figureCitation>
). The inner and middle lobes of the hemipenis are somewhat variable in shape; some specimens have a middle lobe that is slightly wider distally than others, and the outer corner ranges from quite angular to rounded (
<figureCitation id="B9932A32BA4EFF9F166CD2274F7BF9D4" box="[1136,1307,1601,1628]" captionStart="FIGURE 4" captionStartId="8.[151,248,1767,1792]" captionTargetBox="[262,1321,181,1741]" captionTargetId="figure@8.[261,1326,181,1741]" captionTargetPageId="8" captionText="FIGURE 4. AI, Ilyocypris japonica Okubo, 1990. A, detail of female antenna (LBM 1430009156, sample 22; see Table 1). B, detail of male antenna (LBM 1430009155, sample 22). C, seventh limb, male (LBM 1430009154, sample 22). D, outline of female reproductive organ, arrow indicates anterior (LBM 1430009157, sample 22). E, hemipenis (LBM 1430009154, sample 22). F, copulatory process of hemipenis (LBM 1430009154, sample 22). G, hemipenis (LBM 1430009148, sample 15). H, copulatory process of hemipenis (LBM 1430009148, sample 15). I, outlines of the hemipenes of six specimens, some reflected on the vertical axis to facilitate comparisons (left to right LBM 1430009153, sample 22; LBM 1430009152, sample 21; LBM 1430009146, sample 15; LBM 1430009145, sample 10; LBM 1430009140, sample 5; LBM 1430009139, sample 3). J &amp; K, Ilyocypris mongolica. J, hemipenis (LBM 1430009161, sample 17). K, copulatory process of hemipenis (LBM 1430009161, sample 17)." httpUri="https://zenodo.org/record/3363541/files/figure.png" pageId="11">Fig. 4E, G &amp; I</figureCitation>
). The male copulatory process is slightly narrower distally in some specimens compared with others. Sizes of individuals were variable, even within one sample, with some adult specimens noticeably larger (approximately 20% longer and 14% higher) than other adults (
<figureCitation id="B9932A32BA4EFF9F13A6D2CB4B9FF940" box="[442,511,1709,1736]" captionStart="FIGURE 7" captionStartId="11.[151,250,1314,1339]" captionTargetBox="[151,1434,345,1289]" captionTargetId="figure@11.[151,1435,345,1289]" captionTargetPageId="11" captionText="FIGURE 7. Two male specimens of Ilyocypris japonica from the same sample (Lake Biwa Museum rice field, 14 June 2013, sample 18; see Table 1), showing the two size types, large and small. A (LBM 1430009151) &amp; B (LBM 1430009150), external view of right valves. C (LBM 1430009151) &amp; D (LBM 1430009150), details of surface of valves, locations marked by dotted rectangles on A &amp; B. Scale bar = 500 µm for A &amp; B, 94 µm for C &amp; D." httpUri="https://zenodo.org/record/3363547/files/figure.png" pageId="11">Fig. 7</figureCitation>
). The hemipenis morphology of the two size
<typeStatus id="FE138815BA4EFF9F11E1D2C84E58F940" box="[1021,1080,1710,1736]" pageId="11">types</typeStatus>
are very similar (
<figureCitation id="B9932A32BA4EFF9F171CD2CB4FEEF940" box="[1280,1422,1709,1736]" captionStart="FIGURE 5" captionStartId="9.[151,250,1475,1500]" captionTargetBox="[151,1436,181,1446]" captionTargetId="figure@9.[151,1436,181,1449]" captionTargetPageId="9" captionText="FIGURE 5. Light microscopy photographs of Ilyocypris hemipenes. A, Ilyocypris japonica Okubo, 1990 (LBM 1430009150, sample 18; see Table 1). B, Ilyocypris japonica Okubo, 1990 (LBM 1430009151, sample 18). C, Ilyocypris mongolica Martens, 1991 (LBM 1430009161, sample 17). D, Ilyocypris dentifera Sars, 1903 (LBM 1430009119, sample 6). E, Ilyocypris angulata Sars, 1903 (LBM 1430009112, sample 20). F, Ilyocypris incus sp. nov. (DB40054, sample 14). E has been reflected on the vertical axis to facilitate comparisons." httpUri="https://zenodo.org/record/3363543/files/figure.png" pageId="11">Fig. 5A &amp; B</figureCitation>
), so we conclude that they are the same species (see Discussion). The roughened surface of the postero-ventral region of the calcified inner lamella of the left valve varies in extent, extending higher up the posterior margin in some specimens.
</paragraph>
<paragraph id="211736B7BA4EFF9F12DBD3584F2DF84B" blockId="11.[151,1437,1493,1988]" pageId="11">
<emphasis id="13DCEAA5BA4EFF9F12DBD3584B54F8D0" bold="true" box="[199,308,1854,1880]" pageId="11">Remarks</emphasis>
. The locations of the
<typeStatus id="FE138815BA4EFF9F1033D358483FF8D0" box="[559,607,1854,1880]" pageId="11">type</typeStatus>
specimens of
<taxonomicName id="E6A84D34BA4EFF9F1119D3584980F8DF" authorityName="Okubo" authorityYear="1990" box="[773,992,1854,1880]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="11" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="13DCEAA5BA4EFF9F1119D3584980F8DF" box="[773,992,1854,1880]" italics="true" pageId="11">Ilyocypris japonica</emphasis>
</taxonomicName>
and its junior synonym
<taxonomicName id="E6A84D34BA4EFF9F16EBD3584BC3F8F4" authority="Okubo, 1992" authorityName="Okubo" authorityYear="1992" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="11" phylum="Arthropoda" rank="species" species="haterumensis">
<emphasis id="13DCEAA5BA4EFF9F16EBD3584B69F8F3" italics="true" pageId="11">Ilyocypris haterumensis</emphasis>
<bibRefCitation id="45394B46BA4EFF9F130CD3074BC3F8F4" author="Okubo, I. &amp; Terauchi, M." box="[272,419,1889,1916]" pageId="11" pagination="101 - 108" refId="ref22454" refString="Okubo, I. &amp; Terauchi, M. (1992) Freshwater Ostracoda from the Southwestern Islands of Japan. Proceedings of the Japanese Society of Systematic Zoology, 46, 101 - 108." type="journal article" year="1992">Okubo, 1992</bibRefCitation>
</taxonomicName>
(in
<bibRefCitation id="45394B46BA4EFF9F13CCD3074882F8F4" author="Okubo, I. &amp; Terauchi, M." box="[464,738,1889,1916]" pageId="11" pagination="101 - 108" refId="ref22454" refString="Okubo, I. &amp; Terauchi, M. (1992) Freshwater Ostracoda from the Southwestern Islands of Japan. Proceedings of the Japanese Society of Systematic Zoology, 46, 101 - 108." type="journal article" year="1992">Okubo &amp; Terauchi 1992</bibRefCitation>
) were not mentioned in the original publications and enquires were unsuccessful in determining their whereabouts. The material must therefore be considered lost, and only the figures in
<bibRefCitation id="45394B46BA4EFF9F1315D3CF4BC0F84C" author="Okubo, I." box="[265,416,1961,1988]" pageId="11" pagination="39 - 50" refId="ref22357" refString="Okubo, I. (1990) Sixteen species of freshwater Ostracoda from Japan. Bulletin of the Biogeographical Society of Japan, 45, 39 - 50." type="journal article" year="1990">Okubo (1990</bibRefCitation>
;
<bibRefCitation id="45394B46BA4EFF9F13B0D3CC4B86F84C" author="Okubo I." box="[428,486,1962,1988]" pageId="11" refId="ref22386" refString="Okubo I. (2004) Nihon tansui san kaimijinko rui ni tsuite. Kabushikigaisha Sanmon Insatsusho, Okayama, 72 pp. [In Japanese, privately published]" type="book" year="2004">2004</bibRefCitation>
) and
<bibRefCitation id="45394B46BA4EFF9F1038D3CF4926F84C" author="Okubo, I. &amp; Terauchi, M." box="[548,838,1961,1988]" pageId="11" pagination="101 - 108" refId="ref22454" refString="Okubo, I. &amp; Terauchi, M. (1992) Freshwater Ostracoda from the Southwestern Islands of Japan. Proceedings of the Japanese Society of Systematic Zoology, 46, 101 - 108." type="journal article" year="1992">Okubo &amp; Terauchi (1992)</bibRefCitation>
are available for identification of this species.
</paragraph>
<caption id="75D7663FBA49FF98128BD2394E99F8A4" httpUri="https://zenodo.org/record/3363549/files/figure.png" pageId="12" startId="12.[151,250,1631,1656]" targetBox="[151,1435,181,1605]" targetPageId="12">
<paragraph id="211736B7BA49FF98128BD2394E99F8A4" blockId="12.[151,1436,1631,1836]" pageId="12">
<emphasis id="13DCEAA5BA49FF98128BD2394B72F9F0" bold="true" box="[151,274,1631,1656]" pageId="12">FIGURE 8.</emphasis>
<taxonomicName id="E6A84D34BA49FF98130BD20648E7F9F0" authority="Martens, 1991" authorityName="Martens" authorityYear="1991" box="[279,647,1632,1656]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="12" phylum="Arthropoda" rank="species" species="mongolica">
<emphasis id="13DCEAA5BA49FF98130BD2064B8DF9F0" box="[279,493,1632,1656]" italics="true" pageId="12">Ilyocypris mongolica</emphasis>
<bibRefCitation id="45394B46BA49FF9813EED20648E7F9F0" author="Martens, K." box="[498,647,1632,1656]" pageId="12" pagination="53 - 60" refId="ref21598" refString="Martens, K. (1991) On a small collection of non-marine ostracods from Mongolia, with the description of a new species (Crustacea, Ostracoda). Miscellanea Zoologica Hungarica, 6, 53 - 60." type="journal article" year="1991">Martens, 1991</bibRefCitation>
</taxonomicName>
. A, female left valve, external view (LBM 1430009163). B, female right valve, external view (LBM 1430009163). C, female left valve, internal view (LBM 1430009163). D, female right valve, internal view (LBM 1430009163). E, female left valve, internal view, detail of postero-ventral margin (LBM 1430009163). F, female right valve, internal view, detail of antero-ventral margin (LBM 1430009163). G, female left valve, internal view, muscle scars (LBM 1430009163). H, male right valve, external view (LBM 1430009161). I, female carapace, dorsal view, anterior to right (LBM 1430009162). Scale bar = 500 µm for AD, HI, 174 µm for EG. All specimens from sample 17; see Table 1.
</paragraph>
</caption>
<caption id="75D7663FBA48FF99128BD2524A82F875" httpUri="https://zenodo.org/record/3363551/files/figure.png" pageId="13" startId="13.[151,250,1588,1613]" targetBox="[325,1263,199,1562]" targetPageId="13">
<paragraph id="211736B7BA48FF99128BD2524A82F875" blockId="13.[151,1437,1588,2045]" pageId="13">
<emphasis id="13DCEAA5BA48FF99128BD2524B73F9C5" bold="true" box="[151,275,1588,1613]" pageId="13">FIGURE 9.</emphasis>
Distribution maps of
<taxonomicName id="E6A84D34BA48FF9913EED25348D9F9C5" authorityName="G. O. Sars-Chen" authorityYear="1903" box="[498,697,1589,1613]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="13" phylum="Arthropoda" rank="species" species="dentifera">
<emphasis id="13DCEAA5BA48FF9913EED25348D9F9C5" box="[498,697,1589,1613]" italics="true" pageId="13">Ilyocypris dentifera</emphasis>
</taxonomicName>
,
<taxonomicName id="E6A84D34BA48FF9910DAD2534956F9C5" authorityName="Sars" authorityYear="1903" box="[710,822,1589,1613]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="13" phylum="Arthropoda" rank="species" species="angulata">
<emphasis id="13DCEAA5BA48FF9910DAD2534956F9C5" box="[710,822,1589,1613]" italics="true" pageId="13">I. angulata</emphasis>
</taxonomicName>
,
<taxonomicName id="E6A84D34BA48FF99115FD25349D1F9C4" authorityName="Okubo" authorityYear="1990" box="[835,945,1589,1612]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="13" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="13DCEAA5BA48FF99115FD25349D1F9C4" box="[835,945,1589,1612]" italics="true" pageId="13">I. japonica</emphasis>
</taxonomicName>
,
<taxonomicName id="E6A84D34BA48FF9911A1D2534E5EF9C5" authorityName="Martens" authorityYear="1991" box="[957,1086,1589,1613]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="13" phylum="Arthropoda" rank="species" species="mongolica">
<emphasis id="13DCEAA5BA48FF9911A1D2534E5EF9C5" box="[957,1086,1589,1613]" italics="true" pageId="13">I. mongolica</emphasis>
</taxonomicName>
, and
<taxonomicName id="E6A84D34BA48FF99166AD2534F76F9C4" authority="Smith &amp; Zhai &amp; Chang, 2019" authorityName="Smith &amp; Zhai &amp; Chang" authorityYear="2019" box="[1142,1302,1589,1613]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="13" phylum="Arthropoda" rank="species" species="incus" status="sp. nov.">
<emphasis id="13DCEAA5BA48FF99166AD2534F76F9C4" box="[1142,1302,1589,1613]" italics="true" pageId="13">Ilyocypris incus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="08EF57DEBA48FF991701D2534F0CF9C4" box="[1309,1388,1589,1612]" pageId="13" rank="species">sp. nov.</taxonomicNameLabel>
One additional fossil record of
<taxonomicName id="E6A84D34BA48FF9913BFD23F4873F9F9" authorityName="G. O. Sars-Chen" authorityYear="1903" box="[419,531,1625,1649]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="13" phylum="Arthropoda" rank="species" species="dentifera">
<emphasis id="13DCEAA5BA48FF9913BFD23F4873F9F9" box="[419,531,1625,1649]" italics="true" pageId="13">I. dentifera</emphasis>
</taxonomicName>
exists from Manitoba, Canada (
<bibRefCitation id="45394B46BA48FF991144D23E4E40F9F9" author="Klassen, R. W. &amp; Delorme, L. D. &amp; Mott, R. J." box="[856,1056,1624,1649]" pageId="13" pagination="433 - 447" refId="ref21165" refString="Klassen, R. W., Delorme, L. D. &amp; Mott, R. J. (1967) Geology and paleontology of Pleistocene deposits in southwestern Manitoba. Canadian Journal of Earth Sciences, 4, 433 - 447. https: // doi. org / 10.1139 / e 67 - 022" type="journal article" year="1967">
Klassen
<emphasis id="13DCEAA5BA48FF9911ADD23F4984F9F9" box="[945,996,1625,1649]" italics="true" pageId="13">et al.</emphasis>
1967
</bibRefCitation>
), but is not shown on the map. As the records of
<taxonomicName id="E6A84D34BA48FF99131ED21B4BA8F91D" authorityName="G. O. Sars-Chen" authorityYear="1903" box="[258,456,1661,1685]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="13" phylum="Arthropoda" rank="species" species="dentifera">
<emphasis id="13DCEAA5BA48FF99131ED21B4BA8F91D" box="[258,456,1661,1685]" italics="true" pageId="13">Ilyocypris dentifera</emphasis>
</taxonomicName>
of Kim &amp; Min (1991) probably at least in part refer to different species, their records are not plotted on the map. Records of
<taxonomicName id="E6A84D34BA48FF9913AED2C74843F931" authorityName="G. O. Sars-Chen" authorityYear="1903" box="[434,547,1697,1721]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="13" phylum="Arthropoda" rank="species" species="dentifera">
<emphasis id="13DCEAA5BA48FF9913AED2C74843F931" box="[434,547,1697,1721]" italics="true" pageId="13">I. dentifera</emphasis>
</taxonomicName>
by
<bibRefCitation id="45394B46BA48FF991056D2C748B8F931" author="Huang, M. X." box="[586,728,1696,1721]" pageId="13" pagination="8 - 10" refId="ref20359" refString="Huang, M. X. (1986) A general survey of ecological distribution of Ostracoda (Crustacea) in Sichuan Province. Sichuan Journal of Zoology, 5, 8 - 10. [In Chinese]" type="journal article" year="1986">Huang (1986)</bibRefCitation>
and
<bibRefCitation id="45394B46BA48FF991117D2C749B3F931" author="Huang, M. X. &amp; Du, J. &amp; Long, Z. H." box="[779,979,1696,1721]" pageId="13" pagination="128 - 149" refId="ref20398" refString="Huang, M. X., Du, J. &amp; Long, Z. H. (2006) The crustacean of Sichuan. Sichuan Science and Technology Publishers, Chengdu, pp. 128 - 149. [In Chinese]" type="book chapter" year="2006">
Huang
<emphasis id="13DCEAA5BA48FF991149D2C749E2F931" box="[853,898,1697,1721]" italics="true" pageId="13">et al</emphasis>
. (2006)
</bibRefCitation>
are also not plotted as they are likely to also be misidentifications. Other reports with no figures cannot be confirmed but are plotted (see text for more details).? indicates uncertain records. Abbreviations and colours of points refer to Köppen climatic zones, following
<bibRefCitation id="45394B46BA48FF99166ED28E4F44F889" author="Peel, M. C. &amp; Finlayson, B. L. &amp; McMahon, T. A." box="[1138,1316,1768,1793]" pageId="13" pagination="1633 - 1644" refId="ref22573" refString="Peel, M. C., Finlayson, B. L. &amp; McMahon, T. A. (2007) Updated world map of the Koppen-Geiger climate classification. Hydrology and Earth System Sciences Discussions, European Geosciences Union, 11, 1633 - 1644. https: // doi. org / 10.5194 / hess- 11 - 1633 - 2007" type="journal article" year="2007">
Peel
<emphasis id="13DCEAA5BA48FF9916B8D28F4EB1F889" box="[1188,1233,1769,1793]" italics="true" pageId="13">et al</emphasis>
. (2007)
</bibRefCitation>
. Data from: 1,
<bibRefCitation id="45394B46BA48FF9912ADD36A4B1AF8AD" author="Kaiser, K. &amp; Kempf, E. K. &amp; Leroi-Gourhan, A. &amp; Schutt, H." box="[177,378,1804,1829]" pageId="13" pagination="263 - 353" refId="ref20442" refString="Kaiser, K., Kempf, E. K., Leroi-Gourhan, A. &amp; Schutt, H. (1973) Quartarstratigraphische Untersuchungen aus dem Damaskus- Becken und seiner Umgebung. Zeitschrift fur Geomorphologie, N. F., 17, 263 - 353." type="journal article" year="1973">
Kaiser
<emphasis id="13DCEAA5BA48FF9912E6D36B4B4EF8AD" box="[250,302,1805,1829]" italics="true" pageId="13">et al.</emphasis>
(1973)
</bibRefCitation>
. 2,
<bibRefCitation id="45394B46BA48FF991382D36A480BF8AD" author="Shinde, Y. S. &amp; Victor, R. &amp; Pai, K." box="[414,619,1804,1829]" pageId="13" pagination="5667 - 5670" refId="ref22938" refString="Shinde, Y. S., Victor, R. &amp; Pai, K. (2014) Freshwater ostracods (Crustacea: Ostracoda) of the plateaus of the northern Western Ghats, India. Journal of Threatened Taxa, 6, 5667 - 5670. https: // doi. org / 10.11609 / JoTT. o 3610.5667 - 70" type="journal article" year="2014">
Shinde
<emphasis id="13DCEAA5BA48FF9913F7D36B4878F8AD" box="[491,536,1805,1829]" italics="true" pageId="13">et al</emphasis>
. (2014)
</bibRefCitation>
. 3,
<bibRefCitation id="45394B46BA48FF991093D36A4910F8AD" author="Kulkarni, M. R. &amp; Padhye, S. &amp; Vanjare, A. I. &amp; Jakhalekar, S. S. &amp; Shinde, Y. S. &amp; Paripatyadar, S. V. &amp; Sheth, S. D. &amp; Kulkarni, S. &amp; Phuge, S. K. &amp; Bhakare, K. &amp; Klukarni, A. S. &amp; Pai, K. &amp; Ghate, H. V." box="[655,880,1804,1829]" pageId="13" pagination="7196 - 7210" refId="ref21249" refString="Kulkarni, M. R., Padhye, S., Vanjare, A. I., Jakhalekar, S. S., Shinde, Y. S., Paripatyadar, S. V., Sheth, S. D., Kulkarni, S., Phuge, S. K., Bhakare, K., Klukarni, A. S., Pai, K. &amp; Ghate, H. V. (2015) Documenting the fauna of a small temporary pond from Pune, Maharashtra, India. Journal of Threatened Taxa, 7, 7196 - 7210. https: // doi. org / 10.11609 / JoTT. o 4190.7196 - 210" type="journal article" year="2015">
Kulkarni
<emphasis id="13DCEAA5BA48FF9910ECD36B497DF8AD" box="[752,797,1805,1829]" italics="true" pageId="13">et al</emphasis>
. (2015)
</bibRefCitation>
. 4,
<bibRefCitation id="45394B46BA48FF991189D36A4E05F8AD" author="Zhai, D. &amp; Zhao, W." box="[917,1125,1804,1829]" pageId="13" pagination="985 - 1026" refId="ref24541" refString="Zhai, D. &amp; Zhao, W. (2014) On some Recent non-marine ostracods from northern China, with descriptions of one new species. Crustaceana, 87, 985 - 1026. https: // doi. org / 10.1163 / 15685403 - 00003339" type="journal article" year="2014">Zhai &amp; Zhao (2014)</bibRefCitation>
. 5,
<bibRefCitation id="45394B46BA48FF991695D36A4F64F8AD" author="Chen, S. Z." box="[1161,1284,1804,1829]" pageId="13" pagination="48 - 63" refId="ref19473" refString="Chen, S. Z. (1982) Subclass Ostracoda. In: Dong, Y. M., Dai, A. Y., Jiang, X. Z., Chen, S. Z., Chen, Y. S. &amp; Cai, R. X. (Eds.), Atlas of the Animals in China: Crustaceans. Volume 1, Version 2. Science Press, Beijing, pp. 48 - 63. [In Chinese]" type="book chapter" year="1982">Chen (1982</bibRefCitation>
;
<bibRefCitation id="45394B46BA48FF991713D36B4F26F8AD" author="Chen, S. Z." box="[1295,1350,1805,1829]" pageId="13" pagination="253 - 256" refId="ref19563" refString="Chen, S. Z. (1990) Description of male Ilyocypris angulata (Crustacea, Ostracoda). Acta Micropalaeontologica Sinica, 7, 253 - 256." type="journal article" year="1990">1990</bibRefCitation>
). 6,
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,
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.
</paragraph>
</caption>
<paragraph id="211736B7BA4BFF9A12DBD4FF4F0CFDB7" blockId="14.[151,1437,153,1655]" pageId="14">
In the original description of
<taxonomicName id="E6A84D34BA4BFF9A1012D4FC48D8FF3B" authorityName="Okubo" authorityYear="1992" box="[526,696,153,179]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="14" phylum="Arthropoda" rank="species" species="haterumensis">
<emphasis id="13DCEAA5BA4BFF9A1012D4FC48D8FF3B" box="[526,696,153,179]" italics="true" pageId="14">I. haterumensis</emphasis>
</taxonomicName>
it was noted that it was similar to
<taxonomicName id="E6A84D34BA4BFF9A1621D4FC4ED5FF3B" authorityName="Okubo" authorityYear="1990" box="[1085,1205,154,179]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="14" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="13DCEAA5BA4BFF9A1621D4FC4ED5FF3B" box="[1085,1205,154,179]" italics="true" pageId="14">I. japonica</emphasis>
</taxonomicName>
, but the middle lobe of the hemipenis was slightly different, and this was used to justify the erection of a new species (
<bibRefCitation id="45394B46BA4BFF9A16D1D4DB4AB2FF73" author="Okubo, I. &amp; Terauchi, M." pageId="14" pagination="101 - 108" refId="ref22454" refString="Okubo, I. &amp; Terauchi, M. (1992) Freshwater Ostracoda from the Southwestern Islands of Japan. Proceedings of the Japanese Society of Systematic Zoology, 46, 101 - 108." type="journal article" year="1992">Okubo &amp; Terauchi 1992</bibRefCitation>
). However, it was later synonymised with
<taxonomicName id="E6A84D34BA4BFF9A10B8D484497BFF73" authorityName="Okubo" authorityYear="1990" box="[676,795,226,251]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="14" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="13DCEAA5BA4BFF9A10B8D484497BFF73" box="[676,795,226,251]" italics="true" pageId="14">I. japonica</emphasis>
</taxonomicName>
by
<bibRefCitation id="45394B46BA4BFF9A1158D4874981FF73" author="Okubo I." box="[836,993,225,251]" pageId="14" refId="ref22386" refString="Okubo I. (2004) Nihon tansui san kaimijinko rui ni tsuite. Kabushikigaisha Sanmon Insatsusho, Okayama, 72 pp. [In Japanese, privately published]" type="book" year="2004">Okubo (2004)</bibRefCitation>
, but without an explanation. The middle lobe of the hemipenis of
<taxonomicName id="E6A84D34BA4BFF9A13B6D5604841FE97" authorityName="Okubo" authorityYear="1990" box="[426,545,262,287]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="14" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="13DCEAA5BA4BFF9A13B6D5604841FE97" box="[426,545,262,287]" italics="true" pageId="14">I. japonica</emphasis>
</taxonomicName>
is somewhat variable in shape (
<figureCitation id="B9932A32BA4BFF9A119DD5634E48FE97" box="[897,1064,261,287]" captionStart="FIGURE 4" captionStartId="8.[151,248,1767,1792]" captionTargetBox="[262,1321,181,1741]" captionTargetId="figure@8.[261,1326,181,1741]" captionTargetPageId="8" captionText="FIGURE 4. AI, Ilyocypris japonica Okubo, 1990. A, detail of female antenna (LBM 1430009156, sample 22; see Table 1). B, detail of male antenna (LBM 1430009155, sample 22). C, seventh limb, male (LBM 1430009154, sample 22). D, outline of female reproductive organ, arrow indicates anterior (LBM 1430009157, sample 22). E, hemipenis (LBM 1430009154, sample 22). F, copulatory process of hemipenis (LBM 1430009154, sample 22). G, hemipenis (LBM 1430009148, sample 15). H, copulatory process of hemipenis (LBM 1430009148, sample 15). I, outlines of the hemipenes of six specimens, some reflected on the vertical axis to facilitate comparisons (left to right LBM 1430009153, sample 22; LBM 1430009152, sample 21; LBM 1430009146, sample 15; LBM 1430009145, sample 10; LBM 1430009140, sample 5; LBM 1430009139, sample 3). J &amp; K, Ilyocypris mongolica. J, hemipenis (LBM 1430009161, sample 17). K, copulatory process of hemipenis (LBM 1430009161, sample 17)." httpUri="https://zenodo.org/record/3363541/files/figure.png" pageId="14">Figs 4E, G &amp; I</figureCitation>
) and the figured middle lobe of
<taxonomicName id="E6A84D34BA4BFF9A1790D5604B49FECB" authorityName="Okubo" authorityYear="1992" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="14" phylum="Arthropoda" rank="species" species="haterumensis">
<emphasis id="13DCEAA5BA4BFF9A1790D5604B49FECB" italics="true" pageId="14">I. haterumensis</emphasis>
</taxonomicName>
does fit into this range. In the figures of
<taxonomicName id="E6A84D34BA4BFF9A10F4D54C49F1FECB" authorityName="Okubo" authorityYear="1992" box="[744,913,297,323]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="14" phylum="Arthropoda" rank="species" species="haterumensis">
<emphasis id="13DCEAA5BA4BFF9A10F4D54C49F1FECB" box="[744,913,297,323]" italics="true" pageId="14">I. haterumensis</emphasis>
</taxonomicName>
provided by
<bibRefCitation id="45394B46BA4BFF9A163ED54F4F5FFECB" author="Okubo, I. &amp; Terauchi, M." box="[1058,1343,297,323]" pageId="14" pagination="101 - 108" refId="ref22454" refString="Okubo, I. &amp; Terauchi, M. (1992) Freshwater Ostracoda from the Southwestern Islands of Japan. Proceedings of the Japanese Society of Systematic Zoology, 46, 101 - 108." type="journal article" year="1992">Okubo &amp; Terauchi (1992)</bibRefCitation>
the seventh limb has one seta on the third segment, and the sp seta on the caudal ramus is short, while
<taxonomicName id="E6A84D34BA4BFF9A168ED5284F67FEEF" authorityName="Okubo" authorityYear="1990" box="[1170,1287,334,359]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="14" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="13DCEAA5BA4BFF9A168ED5284F67FEEF" box="[1170,1287,334,359]" italics="true" pageId="14">I. japonica</emphasis>
</taxonomicName>
has two setae on the same seventh limb segment, and the sp seta on the caudal ramus is noticeably longer. These discrepancies cannot be checked because of the lack of
<typeStatus id="FE138815BA4BFF9A1012D5F3485DFE27" box="[526,573,405,431]" pageId="14">type</typeStatus>
material. We therefore follow
<bibRefCitation id="45394B46BA4BFF9A1191D5F34E4AFE27" author="Okubo I." box="[909,1066,405,431]" pageId="14" refId="ref22386" refString="Okubo I. (2004) Nihon tansui san kaimijinko rui ni tsuite. Kabushikigaisha Sanmon Insatsusho, Okayama, 72 pp. [In Japanese, privately published]" type="book" year="2004">Okubo (2004)</bibRefCitation>
, who observed specimens of both species and considered that the two names are synonyms. We assume that the discrepancies are due to errors, damage or abnormalities with the specimen of
<taxonomicName id="E6A84D34BA4BFF9A1022D5B84887FE7F" authorityName="Okubo" authorityYear="1992" box="[574,743,477,503]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="14" phylum="Arthropoda" rank="species" species="haterumensis">
<emphasis id="13DCEAA5BA4BFF9A1022D5B84887FE7F" box="[574,743,477,503]" italics="true" pageId="14">I. haterumensis</emphasis>
</taxonomicName>
figured by
<bibRefCitation id="45394B46BA4BFF9A117AD5BB4EE7FE7F" author="Okubo, I. &amp; Terauchi, M." box="[870,1159,477,503]" pageId="14" pagination="101 - 108" refId="ref22454" refString="Okubo, I. &amp; Terauchi, M. (1992) Freshwater Ostracoda from the Southwestern Islands of Japan. Proceedings of the Japanese Society of Systematic Zoology, 46, 101 - 108." type="journal article" year="1992">Okubo &amp; Terauchi (1992)</bibRefCitation>
. This is supported by our material, some of which has a very similar hemipenis morphology to that of
<taxonomicName id="E6A84D34BA4BFF9A11CAD6644E1EFD93" authorityName="Okubo" authorityYear="1992" box="[982,1150,513,539]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="14" phylum="Arthropoda" rank="species" species="haterumensis">
<emphasis id="13DCEAA5BA4BFF9A11CAD6644E1EFD93" box="[982,1150,513,539]" italics="true" pageId="14">I. haterumensis</emphasis>
</taxonomicName>
(i.e. a very similar middle lobe morphology), but two setae on the third segment of the seventh limb and a long sp seta on the caudal ramus.
</paragraph>
</subSubSection>
<subSubSection id="69B2653CBA4BFF9A12DBD62F4F13FBBF" pageId="14" type="reference_group">
<paragraph id="211736B7BA4BFF9A12DBD62F4E7CFCD7" blockId="14.[151,1437,153,1655]" pageId="14">
<taxonomicName id="E6A84D34BA4BFF9A12DBD62F4BFFFDEB" authorityName="Okubo" authorityYear="1990" box="[199,415,585,611]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="14" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="13DCEAA5BA4BFF9A12DBD62F4BFFFDEB" box="[199,415,585,611]" italics="true" pageId="14">Ilyocypris japonica</emphasis>
</taxonomicName>
is morphologically very similar to
<taxonomicName id="E6A84D34BA4BFF9A113ED62F4ED8FDEB" authority="Martens, 1991" authorityName="Martens" authorityYear="1991" box="[802,1208,585,611]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="14" phylum="Arthropoda" rank="species" species="mongolica">
<emphasis id="13DCEAA5BA4BFF9A113ED62F4E6EFDEB" box="[802,1038,585,611]" italics="true" pageId="14">Ilyocypris mongolica</emphasis>
<bibRefCitation id="45394B46BA4BFF9A1608D62F4ED8FDEB" author="Martens, K." box="[1044,1208,585,611]" pageId="14" pagination="53 - 60" refId="ref21598" refString="Martens, K. (1991) On a small collection of non-marine ostracods from Mongolia, with the description of a new species (Crustacea, Ostracoda). Miscellanea Zoologica Hungarica, 6, 53 - 60." type="journal article" year="1991">Martens, 1991</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="E6A84D34BA4BFF9A16DDD62F4AD1FD0F" authorityName="Martens" authorityYear="1991" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="14" phylum="Arthropoda" rank="species" species="mongolica">
<emphasis id="13DCEAA5BA4BFF9A16DDD62F4AD1FD0F" italics="true" pageId="14">Ilyocypris mongolica</emphasis>
</taxonomicName>
differs from
<taxonomicName id="E6A84D34BA4BFF9A135ED6084BD9FD0F" authorityName="Okubo" authorityYear="1990" box="[322,441,622,647]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="14" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="13DCEAA5BA4BFF9A135ED6084BD9FD0F" box="[322,441,622,647]" italics="true" pageId="14">I. japonica</emphasis>
</taxonomicName>
in the following characters: it is slightly more rectangular in lateral view, with a shallower sloping dorsal margin; the pitting on the carapace is coarser; the inner lists on the calcified inner lamella are closer to the inner edge; and the copulatory process, although similar, is differently shaped (
<figureCitation id="B9932A32BA4BFF9A162ED6D34EDEFD47" box="[1074,1214,693,719]" captionStart="FIGURE 4" captionStartId="8.[151,248,1767,1792]" captionTargetBox="[262,1321,181,1741]" captionTargetId="figure@8.[261,1326,181,1741]" captionTargetPageId="8" captionText="FIGURE 4. AI, Ilyocypris japonica Okubo, 1990. A, detail of female antenna (LBM 1430009156, sample 22; see Table 1). B, detail of male antenna (LBM 1430009155, sample 22). C, seventh limb, male (LBM 1430009154, sample 22). D, outline of female reproductive organ, arrow indicates anterior (LBM 1430009157, sample 22). E, hemipenis (LBM 1430009154, sample 22). F, copulatory process of hemipenis (LBM 1430009154, sample 22). G, hemipenis (LBM 1430009148, sample 15). H, copulatory process of hemipenis (LBM 1430009148, sample 15). I, outlines of the hemipenes of six specimens, some reflected on the vertical axis to facilitate comparisons (left to right LBM 1430009153, sample 22; LBM 1430009152, sample 21; LBM 1430009146, sample 15; LBM 1430009145, sample 10; LBM 1430009140, sample 5; LBM 1430009139, sample 3). J &amp; K, Ilyocypris mongolica. J, hemipenis (LBM 1430009161, sample 17). K, copulatory process of hemipenis (LBM 1430009161, sample 17)." httpUri="https://zenodo.org/record/3363541/files/figure.png" pageId="14">Figs 4J &amp; K</figureCitation>
,
<figureCitation id="B9932A32BA4BFF9A16D5D6D24E8CFD47" box="[1225,1260,692,719]" captionStart="FIGURE 5" captionStartId="9.[151,250,1475,1500]" captionTargetBox="[151,1436,181,1446]" captionTargetId="figure@9.[151,1436,181,1449]" captionTargetPageId="9" captionText="FIGURE 5. Light microscopy photographs of Ilyocypris hemipenes. A, Ilyocypris japonica Okubo, 1990 (LBM 1430009150, sample 18; see Table 1). B, Ilyocypris japonica Okubo, 1990 (LBM 1430009151, sample 18). C, Ilyocypris mongolica Martens, 1991 (LBM 1430009161, sample 17). D, Ilyocypris dentifera Sars, 1903 (LBM 1430009119, sample 6). E, Ilyocypris angulata Sars, 1903 (LBM 1430009112, sample 20). F, Ilyocypris incus sp. nov. (DB40054, sample 14). E has been reflected on the vertical axis to facilitate comparisons." httpUri="https://zenodo.org/record/3363543/files/figure.png" pageId="14">5C</figureCitation>
,
<figureCitation id="B9932A32BA4BFF9A16EBD6D34F67FD47" box="[1271,1287,693,719]" captionStart="FIGURE 8" captionStartId="12.[151,250,1631,1656]" captionTargetBox="[151,1435,181,1605]" captionTargetId="figure@12.[151,1435,181,1606]" captionTargetPageId="12" captionText="FIGURE 8. Ilyocypris mongolica Martens, 1991.A, female left valve, external view (LBM 1430009163). B, female right valve, external view (LBM 1430009163). C, female left valve, internal view (LBM 1430009163). D, female right valve, internal view (LBM 1430009163). E, female left valve, internal view, detail of postero-ventral margin (LBM 1430009163). F, female right valve, internal view, detail of antero-ventral margin (LBM 1430009163). G, female left valve, internal view, muscle scars (LBM 1430009163). H, male right valve, external view (LBM 1430009161). I, female carapace, dorsal view, anterior to right (LBM 1430009162). Scale bar = 500 µm for AD, HI, 174 µm for EG. All specimens from sample 17; see Table 1." httpUri="https://zenodo.org/record/3363549/files/figure.png" pageId="14">8</figureCitation>
). The middle lobe of the hemipenis is also generally larger distally than that of
<taxonomicName id="E6A84D34BA4BFF9A1163D6BC4999FD7B" authorityName="Okubo" authorityYear="1990" box="[895,1017,730,755]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="14" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="13DCEAA5BA4BFF9A1163D6BC4999FD7B" box="[895,1017,730,755]" italics="true" pageId="14">I. japonica</emphasis>
</taxonomicName>
, but the variation of this character in
<taxonomicName id="E6A84D34BA4BFF9A128BD6984B71FC9F" authorityName="Okubo" authorityYear="1990" box="[151,273,766,791]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="14" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="13DCEAA5BA4BFF9A128BD6984B71FC9F" box="[151,273,766,791]" italics="true" pageId="14">I. japonica</emphasis>
</taxonomicName>
is quite large, so that the largest example is similar to that of
<taxonomicName id="E6A84D34BA4BFF9A11D9D6984E33FC9F" authorityName="Martens" authorityYear="1991" box="[965,1107,765,791]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="14" phylum="Arthropoda" rank="species" species="mongolica">
<emphasis id="13DCEAA5BA4BFF9A11D9D6984E33FC9F" box="[965,1107,765,791]" italics="true" pageId="14">I. mongolica</emphasis>
</taxonomicName>
. These morphological differences are small, but due to the different geographical (and climatic) distributions and habitats of the two species (see Distribution section below), we provisionally treat them as two separate species.
</paragraph>
<paragraph id="211736B7BA4BFF9A12DBD70F4F13FBBF" blockId="14.[151,1437,153,1655]" pageId="14">
Contrary to our results, a cladistic study by
<bibRefCitation id="45394B46BA4BFF9A10A5D70F49AFFC0B" author="Karanovic, I. &amp; Lee, W." box="[697,975,873,899]" pageId="14" pagination="39 - 71" refId="ref20665" refString="Karanovic, I. &amp; Lee, W. (2013) On the ostracod genus Ilyocypris, with description of one new species from Korea and the first report of males of I. bradyi (Crustacea: Ostracoda: Podocopida). Proceedings of the Biological Society of Washington, 126, 39 - 71. https: // doi. org / 10.2988 / 0006 - 324 X- 126.1.39" type="journal article" year="2013">Karanovic &amp; Lee (2013)</bibRefCitation>
of 21
<taxonomicName id="E6A84D34BA4BFF9A1600D70F4EEBFC0B" authorityName="Zenker" box="[1052,1163,873,899]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="14" phylum="Arthropoda" rank="genus">
<emphasis id="13DCEAA5BA4BFF9A1600D70F4EEBFC0B" box="[1052,1163,873,899]" italics="true" pageId="14">Ilyocypris</emphasis>
</taxonomicName>
species indicated that
<taxonomicName id="E6A84D34BA4BFF9A1790D70C4A98FC2F" authorityName="Okubo" authorityYear="1990" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="14" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="13DCEAA5BA4BFF9A1790D70C4A98FC2F" italics="true" pageId="14">I. japonica</emphasis>
</taxonomicName>
and
<taxonomicName id="E6A84D34BA4BFF9A1332D7E84BDAFC2F" authorityName="Martens" authorityYear="1991" box="[302,442,909,935]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="14" phylum="Arthropoda" rank="species" species="mongolica">
<emphasis id="13DCEAA5BA4BFF9A1332D7E84BDAFC2F" box="[302,442,909,935]" italics="true" pageId="14">I. mongolica</emphasis>
</taxonomicName>
are only distantly related (see cladistic diagram
<figureCitation id="B9932A32BA4BFF9A11CFD7EB4E42FC2F" box="[979,1058,908,935]" captionStart="FIGURE 15" captionStartId="25.[151,250,1839,1864]" captionTargetBox="[238,1350,181,1814]" captionTargetId="figure@25.[238,1350,181,1814]" captionTargetPageId="25" captionText="FIGURE 15. Ilyocypris incus sp. nov. A, antennule, male (DB40054). B, antenna, male (LBM 1430009132). C, mandibular palp, female (LBM 1430009136). D, mandibular coxa, female (LBM 1430009134). E, maxillula palp and endites (setae on endites not drawn), female (LBM 1430009136). F, fifth limb, female (DB40055). G, sixth limb, male (LBM 1430009132), arrow and dashed line indicate length of first endopodal segment. H, fifth limb palp, male (DB40054). All specimens from sample 14; see Table 1." httpUri="https://zenodo.org/record/3363563/files/figure.png" pageId="14">Fig. 15</figureCitation>
of
<bibRefCitation id="45394B46BA4BFF9A165BD7EB4F2AFC2F" author="Karanovic, I. &amp; Lee, W." box="[1095,1354,909,935]" pageId="14" pagination="39 - 71" refId="ref20665" refString="Karanovic, I. &amp; Lee, W. (2013) On the ostracod genus Ilyocypris, with description of one new species from Korea and the first report of males of I. bradyi (Crustacea: Ostracoda: Podocopida). Proceedings of the Biological Society of Washington, 126, 39 - 71. https: // doi. org / 10.2988 / 0006 - 324 X- 126.1.39" type="journal article" year="2013">Karanovic &amp; Lee 2013</bibRefCitation>
). However, characters for some species were incorrectly coded in the study of
<bibRefCitation id="45394B46BA4BFF9A11ACD7D74EA0FC43" author="Karanovic, I. &amp; Lee, W." box="[944,1216,945,971]" pageId="14" pagination="39 - 71" refId="ref20665" refString="Karanovic, I. &amp; Lee, W. (2013) On the ostracod genus Ilyocypris, with description of one new species from Korea and the first report of males of I. bradyi (Crustacea: Ostracoda: Podocopida). Proceedings of the Biological Society of Washington, 126, 39 - 71. https: // doi. org / 10.2988 / 0006 - 324 X- 126.1.39" type="journal article" year="2013">Karanovic &amp; Lee (2013)</bibRefCitation>
(e.g. for
<taxonomicName id="E6A84D34BA4BFF9A1739D7D44FFCFC43" authorityName="Okubo" authorityYear="1990" box="[1317,1436,946,971]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="14" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="13DCEAA5BA4BFF9A1739D7D44FFCFC43" box="[1317,1436,946,971]" italics="true" pageId="14">I. japonica</emphasis>
</taxonomicName>
characters 3, 11, 12, and 14 were incorrectly coded), and this undoubtedly accounts for the discrepancy between their cladistic results and our detailed morphological analysis. A corrected cladistic analysis of
<taxonomicName id="E6A84D34BA4BFF9A1692D79F4E9DFB9B" authorityName="Zenker" box="[1166,1277,1017,1043]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="14" phylum="Arthropoda" rank="genus">
<emphasis id="13DCEAA5BA4BFF9A1692D79F4E9DFB9B" box="[1166,1277,1017,1043]" italics="true" pageId="14">Ilyocypris</emphasis>
</taxonomicName>
species is not given herein, as an informative analysis is hindered by the lack of comprehensive descriptions for many species.
</paragraph>
</subSubSection>
<subSubSection id="69B2653CBA4BFF9A12DBD0274FE7FBF7" pageId="14" type="description">
<paragraph id="211736B7BA4BFF9A12DBD0274FE7FBF7" blockId="14.[151,1437,153,1655]" pageId="14">
The record of
<taxonomicName id="E6A84D34BA4BFF9A1379D0244B81FBD3" authorityName="G. O. Sars-Chen" authorityYear="1903" box="[357,481,1089,1115]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="14" phylum="Arthropoda" rank="species" species="dentifera">
<emphasis id="13DCEAA5BA4BFF9A1379D0244B81FBD3" box="[357,481,1089,1115]" italics="true" pageId="14">I. dentifera</emphasis>
</taxonomicName>
reported from the brackish part of the Obitsu River estuary in
<collectingCountry id="59BF7627BA4BFF9A168BD0274EB7FBD3" box="[1175,1239,1089,1115]" name="Japan" pageId="14">Japan</collectingCountry>
by
<bibRefCitation id="45394B46BA4BFF9A171CD0274B5AFBF7" author="Nakao Y. &amp; Tsukagoshi, A." pageId="14" pagination="67 - 115" refId="ref22230" refString="Nakao Y. &amp; Tsukagoshi, A. (2002) Brackish-water Ostracoda (Crustacea) from Obitsu River estuary, Central Japan. Species Diversity, 7, 67 - 115. https: // doi. org / 10.12782 / specdiv. 7.67" type="journal article" year="2002">Nakao &amp; Tsukagoshi (2002</bibRefCitation>
;
<bibRefCitation id="45394B46BA4BFF9A1359D0034B1FFBF7" author="Nakao, Y. &amp; Tsukagoshi, A." box="[325,383,1125,1151]" pageId="14" pagination="249 - 275" refId="ref22275" refString="Nakao, Y. &amp; Tsukagoshi, A. (2008) Seasonal and decadal changes in distribution and abundance of Ostracoda (Crustacea) in the brackish-water environments. Proceedings of the Institute of Natural Sciences, Nihon University, 43, 249 - 275." type="journal article" year="2008">2008</bibRefCitation>
) is possibly a misidentification of
<taxonomicName id="E6A84D34BA4BFF9A111ED000491CFBF7" authorityName="Okubo" authorityYear="1990" box="[770,892,1126,1151]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="14" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="13DCEAA5BA4BFF9A111ED000491CFBF7" box="[770,892,1126,1151]" italics="true" pageId="14">I. japonica</emphasis>
</taxonomicName>
. See under
<taxonomicName id="E6A84D34BA4BFF9A161CD0004E1CFBF7" authorityName="G. O. Sars-Chen" authorityYear="1903" box="[1024,1148,1125,1151]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="14" phylum="Arthropoda" rank="species" species="dentifera">
<emphasis id="13DCEAA5BA4BFF9A161CD0004E1CFBF7" box="[1024,1148,1125,1151]" italics="true" pageId="14">I. dentifera</emphasis>
</taxonomicName>
below for more details.
</paragraph>
</subSubSection>
<subSubSection id="69B2653CBA4BFF9A12DBD0EE4FF6F9FF" pageId="14" type="materials_examined">
<paragraph id="211736B7BA4BFF9A12DBD0EE4896FA17" blockId="14.[151,1437,153,1655]" pageId="14">
<emphasis id="13DCEAA5BA4BFF9A12DBD0EE4B01FB2B" bold="true" box="[199,353,1160,1187]" pageId="14">Distribution.</emphasis>
In
<collectingCountry id="59BF7627BA4BFF9A1399D0EF4BA5FB2B" box="[389,453,1161,1187]" name="Japan" pageId="14">Japan</collectingCountry>
<taxonomicName id="E6A84D34BA4BFF9A13D7D0EC4824FB2B" authorityName="Okubo" authorityYear="1990" box="[459,580,1162,1187]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="14" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="13DCEAA5BA4BFF9A13D7D0EC4824FB2B" box="[459,580,1162,1187]" italics="true" pageId="14">I. japonica</emphasis>
</taxonomicName>
is known from
<collectingRegion id="E36CF855BA4BFF9A10E8D0EF4929FB2B" box="[756,841,1161,1187]" country="Japan" name="Gunma" pageId="14">Gunma</collectingRegion>
,
<collectingRegion id="E36CF855BA4BFF9A1148D0EF49D9FB2B" box="[852,953,1161,1187]" country="Japan" name="Okinawa" pageId="14">Okinawa</collectingRegion>
and
<collectingRegion id="E36CF855BA4BFF9A11F3D0EF4E4FFB2B" box="[1007,1071,1161,1187]" country="Japan" name="Shiga" pageId="14">Shiga</collectingRegion>
Prefectures (
<bibRefCitation id="45394B46BA4BFF9A16DDD0EF4FF7FB2B" author="Okubo, I. &amp; Ida, K." box="[1217,1431,1161,1187]" pageId="14" pagination="105 - 107" refId="ref22416" refString="Okubo, I. &amp; Ida, K. (1989) Freshwater Ostracoda from Gunma Prefecture, Japan. Bulletin of the Biogeographical Society of Japan, 44, 105 - 107. [In Japanese]" type="journal article" year="1989">Okubo &amp; Ida 1989</bibRefCitation>
;
<bibRefCitation id="45394B46BA4BFF9A128BD0CB4B47FB4F" author="Okubo, I." box="[151,295,1197,1223]" pageId="14" pagination="39 - 50" refId="ref22357" refString="Okubo, I. (1990) Sixteen species of freshwater Ostracoda from Japan. Bulletin of the Biogeographical Society of Japan, 45, 39 - 50." type="journal article" year="1990">Okubo 1990</bibRefCitation>
;
<bibRefCitation id="45394B46BA4BFF9A1328D0CB482CFB4F" author="Okubo, I. &amp; Terauchi, M." box="[308,588,1197,1223]" pageId="14" pagination="101 - 108" refId="ref22454" refString="Okubo, I. &amp; Terauchi, M. (1992) Freshwater Ostracoda from the Southwestern Islands of Japan. Proceedings of the Japanese Society of Systematic Zoology, 46, 101 - 108." type="journal article" year="1992">Okubo &amp; Terauchi 1992</bibRefCitation>
; herein) (
<figureCitation id="B9932A32BA4BFF9A10A5D0CB489FFB4F" box="[697,767,1197,1223]" captionStart="FIGURE 9" captionStartId="13.[151,250,1588,1613]" captionTargetBox="[325,1263,199,1562]" captionTargetId="figure@13.[325,1263,199,1562]" captionTargetPageId="13" captionText="FIGURE 9. Distribution maps of Ilyocypris dentifera, I. angulata, I. japonica, I. mongolica, and Ilyocypris incus sp. nov. One additional fossil record of I. dentifera exists from Manitoba, Canada (Klassen et al. 1967), but is not shown on the map. As the records of Ilyocypris dentifera of Kim &amp; Min (1991) probably at least in part refer to different species, their records are not plotted on the map. Records of I. dentifera by Huang (1986) and Huang et al. (2006) are also not plotted as they are likely to also be misidentifications. Other reports with no figures cannot be confirmed but are plotted (see text for more details).? indicates uncertain records. Abbreviations and colours of points refer to Köppen climatic zones, following Peel et al. (2007). Data from: 1, Kaiser et al. (1973). 2, Shinde et al. (2014). 3, Kulkarni et al. (2015). 4, Zhai &amp; Zhao (2014). 5, Chen (1982; 1990). 6, Sars (1903), Chen (1982). 7, Yu et al. (2005). 8, Ma &amp; Yu (2018). 9, Yu (2014). 10, Yu et al. (2009). 11, Victor &amp; Fernando (1980). 12, Zenina &amp; Schornikov (2008). 13, Sidorov &amp; Semenchenko (2012). 14, Lee et al. (2000). 15, This study. 16, Tabuki &amp; Tsuhako (2015). 17, Okubo (1990). 18, Broodbakker (1988). 19, Okubo (2004), this study. 20, Tanaka et al. (2015). 21, Sánchez-Bayo &amp; Goka (2006a; 2006b). 22, Van der Meeren et al. (2010). 23, Martens (1991). 24, Zhai &amp; Zhao (2014). 25, This study. 26, Okubo &amp; Terauchi (1992). 27, This study. 28, Okubo (1990). 29, Nakao &amp; Tsukagoshi (2002; 2008). 30, This study. 31, Okubo (1974)." httpUri="https://zenodo.org/record/3363551/files/figure.png" pageId="14">Fig. 9</figureCitation>
). The records of
<taxonomicName id="E6A84D34BA4BFF9A11D8D0C84E5FFB4F" authorityName="Okubo" authorityYear="1990" box="[964,1087,1198,1223]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="14" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="13DCEAA5BA4BFF9A11D8D0C84E5FFB4F" box="[964,1087,1198,1223]" italics="true" pageId="14">I. japonica</emphasis>
</taxonomicName>
reported herein from
<collectingRegion id="E36CF855BA4BFF9A1726D0CB4B40FB63" country="South Korea" name="Gyeongnam" pageId="14">Gyeongsangnam-do</collectingRegion>
and
<collectingRegion id="E36CF855BA4BFF9A1344D0B74853FB63" box="[344,563,1233,1259]" country="South Korea" name="Gyeongbuk" pageId="14">Gyeongsangbuk-do</collectingRegion>
in the south east of
<collectingCountry id="59BF7627BA4BFF9A1104D0B749C5FB63" box="[792,933,1233,1259]" name="South Korea" pageId="14">South Korea</collectingCountry>
are the first for that country (see
<tableCitation id="6C2A030CBA4BFF9A173DD0B74F14FB63" box="[1313,1396,1233,1259]" captionStart="TABLE 1" captionStartId="1.[151,244,1160,1187]" captionTargetBox="[163,1412,1310,1957]" captionText="TABLE 1. Adult Ilyocypris specimens examined for this study. Initial discriminations of species were based on lateral views of carapaces, and confirmed by dissections of male specimens. Figures in brackets indicate the number of specimens with tubercles on the carapace. Note that tubercles vary from weakly to well-developed and were no- ticeable on whole specimens or valves in dorsal view using a stereo optical microscope." httpUri="http://table.plazi.org/id/75D7663FBA44FF95128BD0EE4E0FFA87" pageId="14" tableUuid="75D7663FBA44FF95128BD0EE4E0FFA87">Table 1</tableCitation>
for more details). Its distribution corresponds to temperate to cold Köppen climatic zones with hot summers, and no dry season (zones Dfa and Cfa;
<bibRefCitation id="45394B46BA4BFF9A101CD17F48D5FABB" author="Peel, M. C. &amp; Finlayson, B. L. &amp; McMahon, T. A." box="[512,693,1305,1331]" pageId="14" pagination="1633 - 1644" refId="ref22573" refString="Peel, M. C., Finlayson, B. L. &amp; McMahon, T. A. (2007) Updated world map of the Koppen-Geiger climate classification. Hydrology and Earth System Sciences Discussions, European Geosciences Union, 11, 1633 - 1644. https: // doi. org / 10.5194 / hess- 11 - 1633 - 2007" type="journal article" year="2007">
Peel
<emphasis id="13DCEAA5BA4BFF9A1024D17C4809FABB" box="[568,617,1305,1331]" italics="true" pageId="14">et al</emphasis>
. 2007
</bibRefCitation>
). The morphologically similar species
<taxonomicName id="E6A84D34BA4BFF9A1674D17C4E96FABB" authorityName="Martens" authorityYear="1991" box="[1128,1270,1305,1331]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="14" phylum="Arthropoda" rank="species" species="mongolica">
<emphasis id="13DCEAA5BA4BFF9A1674D17C4E96FABB" box="[1128,1270,1305,1331]" italics="true" pageId="14">I. mongolica</emphasis>
</taxonomicName>
, in contrast, is known from
<collectingCountry id="59BF7627BA4BFF9A133BD15B4BF4FADF" box="[295,404,1341,1367]" name="Mongolia" pageId="14">Mongolia</collectingCountry>
and
<collectingRegion id="E36CF855BA4BFF9A13D5D15B481BFADF" box="[457,635,1341,1367]" country="China" name="Nei Mongol" pageId="14">Inner Mongolia</collectingRegion>
,
<collectingCountry id="59BF7627BA4BFF9A109AD15B48A9FADF" box="[646,713,1341,1367]" name="China" pageId="14">China</collectingCountry>
(
<bibRefCitation id="45394B46BA4BFF9A10CBD15B4916FADF" author="Martens, K." box="[727,886,1341,1367]" pageId="14" pagination="53 - 60" refId="ref21598" refString="Martens, K. (1991) On a small collection of non-marine ostracods from Mongolia, with the description of a new species (Crustacea, Ostracoda). Miscellanea Zoologica Hungarica, 6, 53 - 60." type="journal article" year="1991">Martens 1991</bibRefCitation>
;
<bibRefCitation id="45394B46BA4BFF9A119DD15B4ED3FADF" author="Van der Meeren, T. &amp; Almendinger, J. E. &amp; Ito, E. &amp; Martens, K." box="[897,1203,1341,1367]" pageId="14" pagination="253 - 273" refId="ref23665" refString="Van der Meeren, T., Almendinger, J. E., Ito, E. &amp; Martens, K. (2010) The ecology of ostracodes (Ostracoda, Crustacea) in western Mongolia. Hydrobiologia, 641, 253 - 273. https: // doi. org / 10.1007 / s 10750 - 010 - 0089 - y" type="journal article" year="2010">
Van der Meeren
<emphasis id="13DCEAA5BA4BFF9A1625D1584E0AFADF" box="[1081,1130,1341,1367]" italics="true" pageId="14">et al</emphasis>
. 2010
</bibRefCitation>
;
<bibRefCitation id="45394B46BA4BFF9A16A2D15B4FF0FADF" author="Zhai, D. &amp; Zhao, W." box="[1214,1424,1341,1367]" pageId="14" pagination="985 - 1026" refId="ref24541" refString="Zhai, D. &amp; Zhao, W. (2014) On some Recent non-marine ostracods from northern China, with descriptions of one new species. Crustaceana, 87, 985 - 1026. https: // doi. org / 10.1163 / 15685403 - 00003339" type="journal article" year="2014">Zhai &amp; Zhao 2014</bibRefCitation>
), corresponding to Köppen climatic zones of arid, cold steppe (BSk), arid, cold desert (BWk) and cold, dry winters, with warm summers (Dwb) (
<bibRefCitation id="45394B46BA4BFF9A13C6D1E348ECFA17" author="Peel, M. C. &amp; Finlayson, B. L. &amp; McMahon, T. A." box="[474,652,1413,1439]" pageId="14" pagination="1633 - 1644" refId="ref22573" refString="Peel, M. C., Finlayson, B. L. &amp; McMahon, T. A. (2007) Updated world map of the Koppen-Geiger climate classification. Hydrology and Earth System Sciences Discussions, European Geosciences Union, 11, 1633 - 1644. https: // doi. org / 10.5194 / hess- 11 - 1633 - 2007" type="journal article" year="2007">
Peel
<emphasis id="13DCEAA5BA4BFF9A100FD1E04825FA17" box="[531,581,1413,1439]" italics="true" pageId="14">et al</emphasis>
. 2007
</bibRefCitation>
) (
<figureCitation id="B9932A32BA4BFF9A10BFD1E34887FA17" box="[675,743,1413,1439]" captionStart="FIGURE 9" captionStartId="13.[151,250,1588,1613]" captionTargetBox="[325,1263,199,1562]" captionTargetId="figure@13.[325,1263,199,1562]" captionTargetPageId="13" captionText="FIGURE 9. Distribution maps of Ilyocypris dentifera, I. angulata, I. japonica, I. mongolica, and Ilyocypris incus sp. nov. One additional fossil record of I. dentifera exists from Manitoba, Canada (Klassen et al. 1967), but is not shown on the map. As the records of Ilyocypris dentifera of Kim &amp; Min (1991) probably at least in part refer to different species, their records are not plotted on the map. Records of I. dentifera by Huang (1986) and Huang et al. (2006) are also not plotted as they are likely to also be misidentifications. Other reports with no figures cannot be confirmed but are plotted (see text for more details).? indicates uncertain records. Abbreviations and colours of points refer to Köppen climatic zones, following Peel et al. (2007). Data from: 1, Kaiser et al. (1973). 2, Shinde et al. (2014). 3, Kulkarni et al. (2015). 4, Zhai &amp; Zhao (2014). 5, Chen (1982; 1990). 6, Sars (1903), Chen (1982). 7, Yu et al. (2005). 8, Ma &amp; Yu (2018). 9, Yu (2014). 10, Yu et al. (2009). 11, Victor &amp; Fernando (1980). 12, Zenina &amp; Schornikov (2008). 13, Sidorov &amp; Semenchenko (2012). 14, Lee et al. (2000). 15, This study. 16, Tabuki &amp; Tsuhako (2015). 17, Okubo (1990). 18, Broodbakker (1988). 19, Okubo (2004), this study. 20, Tanaka et al. (2015). 21, Sánchez-Bayo &amp; Goka (2006a; 2006b). 22, Van der Meeren et al. (2010). 23, Martens (1991). 24, Zhai &amp; Zhao (2014). 25, This study. 26, Okubo &amp; Terauchi (1992). 27, This study. 28, Okubo (1990). 29, Nakao &amp; Tsukagoshi (2002; 2008). 30, This study. 31, Okubo (1974)." httpUri="https://zenodo.org/record/3363551/files/figure.png" pageId="14">Fig. 9</figureCitation>
).
</paragraph>
<paragraph id="211736B7BA4BFF9A12DBD1CF4FF6F9FF" blockId="14.[151,1437,153,1655]" pageId="14">
<emphasis id="13DCEAA5BA4BFF9A12DBD1CF4B4AFA4B" bold="true" box="[199,298,1449,1475]" pageId="14">Ecology.</emphasis>
This species can be abundant in rice fields in
<collectingCountry id="59BF7627BA4BFF9A1127D1CF491CFA4B" box="[827,892,1449,1475]" name="Japan" pageId="14">Japan</collectingCountry>
, found in the top few millimetres of mud in the oxygenated zone. In
<collectingCountry id="59BF7627BA4BFF9A1363D1AB4BDDFA6F" box="[383,445,1485,1511]" name="Japan" pageId="14">Japan</collectingCountry>
it has only been found in rice fields, but in
<collectingCountry id="59BF7627BA4BFF9A11B4D1AB498BFA6F" box="[936,1003,1485,1511]" name="South Korea" pageId="14">Korea</collectingCountry>
it was also collected from a lotus field (
<tableCitation id="6C2A030CBA4BFF9A1282D1974A91F983" box="[158,241,1521,1547]" captionStart="TABLE 1" captionStartId="1.[151,244,1160,1187]" captionTargetBox="[163,1412,1310,1957]" captionText="TABLE 1. Adult Ilyocypris specimens examined for this study. Initial discriminations of species were based on lateral views of carapaces, and confirmed by dissections of male specimens. Figures in brackets indicate the number of specimens with tubercles on the carapace. Note that tubercles vary from weakly to well-developed and were no- ticeable on whole specimens or valves in dorsal view using a stereo optical microscope." httpUri="http://table.plazi.org/id/75D7663FBA44FF95128BD0EE4E0FFA87" pageId="14" tableUuid="75D7663FBA44FF95128BD0EE4E0FFA87">Table 1</tableCitation>
). This species has so far only been found from mid May through to late July (
<bibRefCitation id="45394B46BA4BFF9A1654D1974EB6F983" author="Okubo, I." box="[1096,1238,1521,1547]" pageId="14" pagination="39 - 50" refId="ref22357" refString="Okubo, I. (1990) Sixteen species of freshwater Ostracoda from Japan. Bulletin of the Biogeographical Society of Japan, 45, 39 - 50." type="journal article" year="1990">Okubo 1990</bibRefCitation>
; herein), although its appearance and disappearance in rice fields is probably strongly influenced by the rice-growing cycle rather than the seasons per se.
<taxonomicName id="E6A84D34BA4BFF9A1374D25F4830F9DB" authorityName="Martens" authorityYear="1991" box="[360,592,1593,1619]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="14" phylum="Arthropoda" rank="species" species="mongolica">
<emphasis id="13DCEAA5BA4BFF9A1374D25F4830F9DB" box="[360,592,1593,1619]" italics="true" pageId="14">Ilyocypris mongolica</emphasis>
</taxonomicName>
, on the other hand, has not been reported from rice fields, but from a puddle, lakes, and a saline ditch on the flood plain of a river (
<bibRefCitation id="45394B46BA4BFF9A10C0D23B4918F9FF" author="Martens, K." box="[732,888,1629,1655]" pageId="14" pagination="53 - 60" refId="ref21598" refString="Martens, K. (1991) On a small collection of non-marine ostracods from Mongolia, with the description of a new species (Crustacea, Ostracoda). Miscellanea Zoologica Hungarica, 6, 53 - 60." type="journal article" year="1991">Martens 1991</bibRefCitation>
;
<bibRefCitation id="45394B46BA4BFF9A119ED23B4ECFF9FF" author="Van der Meeren, T. &amp; Almendinger, J. E. &amp; Ito, E. &amp; Martens, K." box="[898,1199,1629,1655]" pageId="14" pagination="253 - 273" refId="ref23665" refString="Van der Meeren, T., Almendinger, J. E., Ito, E. &amp; Martens, K. (2010) The ecology of ostracodes (Ostracoda, Crustacea) in western Mongolia. Hydrobiologia, 641, 253 - 273. https: // doi. org / 10.1007 / s 10750 - 010 - 0089 - y" type="journal article" year="2010">
Van der Meeren
<emphasis id="13DCEAA5BA4BFF9A1624D2384E07F9FF" box="[1080,1127,1629,1655]" italics="true" pageId="14">et al</emphasis>
. 2010
</bibRefCitation>
;
<bibRefCitation id="45394B46BA4BFF9A16A6D23B4FEAF9FF" author="Zhai, D. &amp; Zhao, W." box="[1210,1418,1629,1655]" pageId="14" pagination="985 - 1026" refId="ref24541" refString="Zhai, D. &amp; Zhao, W. (2014) On some Recent non-marine ostracods from northern China, with descriptions of one new species. Crustaceana, 87, 985 - 1026. https: // doi. org / 10.1163 / 15685403 - 00003339" type="journal article" year="2014">Zhai &amp; Zhao 2014</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>