155 lines
22 KiB
XML
155 lines
22 KiB
XML
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<mods:title id="14ABB55C838AB759F9322B77BCDFBA54">A monograph of the Australopacific Saprininae (Coleoptera, Histeridae)</mods:title>
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<mods:namePart id="3AA39AE88D00AC6FED0E4782450E8CD4">Lackner, Tomas</mods:namePart>
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<mods:namePart id="04476B069301110480A5BF0A2B9E5EC4">Leschen, Richard A. B.</mods:namePart>
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<treatment id="6650B1349FCD31C831435C95CD2FB3C3" ID-GBIF-Taxon="132648694" LSID="urn:lsid:plazi:treatment:6650B1349FCD31C831435C95CD2FB3C3" httpUri="http://treatment.plazi.org/id/6650B1349FCD31C831435C95CD2FB3C3" lastPageId="199" lastPageNumber="200" pageId="196" pageNumber="197">
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<subSubSection id="E1028619FCB8C7496942A3B83090E4A0" pageId="196" pageNumber="197" type="nomenclature">
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<paragraph id="047CD1BDDD41902E079126D9C36DEFD4" pageId="196" pageNumber="197">
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<taxonomicName id="9E4440A97FDD63088FB6810CD210F872" ID-CoL="7CH2M" authority="Mazur, 1990" authorityName="Mazur" authorityYear="1990" class="Insecta" family="Histeridae" genus="Tomogenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tomogenius motocola" order="Coleoptera" pageId="196" pageNumber="197" phylum="Arthropoda" rank="species" species="motocola">
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<pageBreakToken id="94E383E6205D1AB3420E518B294F5FB6" pageId="196" pageNumber="197" start="start">Tomogenius</pageBreakToken>
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motocola Mazur, 1990
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</taxonomicName>
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Figs 698, 699-707, 708-715, 767
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</paragraph>
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</subSubSection>
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<subSubSection id="18A1E70BD8A61D45CA366EF5A4CF30FC" pageId="196" pageNumber="197" type="reference_group">
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<paragraph id="8124906666079A5E31AAA8359336F08F" pageId="196" pageNumber="197">
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<taxonomicName id="FB5913BAE6A290235C22C0F284A27CA3" class="Insecta" family="Histeridae" genus="Tomogenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tomogenius motocola" order="Coleoptera" pageId="196" pageNumber="197" phylum="Arthropoda" rank="species" species="motocola">Tomogenius motocola</taxonomicName>
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Mazur, 1990: 774, figs. 1-4.
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</paragraph>
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</subSubSection>
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<subSubSection id="ADDAE733161E8FE19E20E7B56C01AA5E" pageId="196" pageNumber="197" type="type locality">
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<paragraph id="9918B75946A8F18D5AAB7BD2E0643544" pageId="196" pageNumber="197">Type locality.</paragraph>
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<paragraph id="D2759D064CA9B8D1CCBEA89A29552C30" pageId="196" pageNumber="197">Australia: South Australia: 18 miles ESE of Penong.</paragraph>
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<caption id="477139E0C7389781F30A2DDD25C9096E" pageId="196" pageNumber="197">
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<paragraph id="B98CEF43DC5D61B387F072AC514EE0FF" pageId="196" pageNumber="197">
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Figure 698.
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<taxonomicName id="009F8F716DEA3FD12FEABDDF5418EE24" class="Insecta" family="Histeridae" genus="Tomogenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tomogenius motocola" order="Coleoptera" pageId="196" pageNumber="197" phylum="Arthropoda" rank="species" species="motocola">Tomogenius motocola</taxonomicName>
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Mazur, 1990 habitus, dorsal view.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="35749128A8170D6B2E57ACA128D5AB34" pageId="196" pageNumber="197" type="type material">
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<paragraph id="81BC91F6932C20E9FFF43D12154AAA99" pageId="196" pageNumber="197">Type material examined.</paragraph>
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<paragraph id="BEDEBD93FF670414C18927B4154A538C" pageId="196" pageNumber="197">
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<taxonomicName id="17F082AAC52571825AD4003443CBA2D3" class="Insecta" family="Histeridae" genus="Tomogenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tomogenius motocola" order="Coleoptera" pageId="196" pageNumber="197" phylum="Arthropoda" rank="species" species="motocola">Tomogenius motocola</taxonomicName>
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Mazur, 1990: paratype, ♀, side-mounted on a triangular card: "32 mi, NW by N / of Eucla Motel / W.A. 18.Oct.1968 / Britton, Upton / Balderson" (printed); followed by: " ANIC Database No. / 25 054256" (printed); followed:
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<normalizedToken id="8E0C3ADF580DBEAAB05EAE04AF41CB78" originalValue="“PARATYPE”">"PARATYPE"</normalizedToken>
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(blue, printed label); followed by: "
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<taxonomicName id="E48EF74FA50CA4CEA41104E4F7F661F4" class="Insecta" family="Histeridae" genus="Tomogenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tomogenius" order="Coleoptera" pageId="196" pageNumber="197" phylum="Arthropoda" rank="genus">Tomogenius</taxonomicName>
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/
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<taxonomicName id="69C6924560B97A068100B58E5F95F173" class="Insecta" family="Corydiidae" genus="Histeroidea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Histeroidea motocola" order="Blattodea" pageId="196" pageNumber="197" phylum="Arthropoda" rank="species" species="motocola">motocola</taxonomicName>
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/ det. S. Mazur" (printed-written); followed by:
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<normalizedToken id="7880FDC4DD04DE60F75E9D737A47AF6B" originalValue="“Paratypus”">"Paratypus"</normalizedToken>
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(beige printed label); followed label: "
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<taxonomicName id="CE34DEDDFB51BA584F1A51CA3458B1D6" class="Insecta" family="Histeridae" genus="Tomogenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tomogenius" order="Coleoptera" pageId="196" pageNumber="197" phylum="Arthropoda" rank="genus">Tomogenius</taxonomicName>
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/
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<taxonomicName id="C05FD2EEE075F5F92B9DA0485D1A7DCD" class="Insecta" family="Corydiidae" genus="Histeroidea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Histeroidea motocola" order="Blattodea" pageId="196" pageNumber="197" phylum="Arthropoda" rank="species" species="motocola">motocola</taxonomicName>
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" (written); followed by:
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<normalizedToken id="4DC54DE5C631B8B72588650AB800F910" originalValue="“10-141”">"10-141"</normalizedToken>
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(yellow, pencil-written label; added by the senior author) (ANIC).
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</paragraph>
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</subSubSection>
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<subSubSection id="70F065A9200F91F246E4E744358AE1D8" pageId="196" pageNumber="197" type="materials_examined">
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<paragraph id="75D27D10A63AE0908731C1A55B43D39A" pageId="196" pageNumber="197">Additional material examined.</paragraph>
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<paragraph id="3C0BDCCE0BD77670980AEE0199F082DB" pageId="196" pageNumber="197">
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AUSTRALIA. Western Australia: 2 ♂♂, Nullarbor Pl., Dingo Cave, 28.x.1968, J. Lowry (ANIC; 1 ♂ in coll. TLAN); 1 ♀ & 1 ♂, Madura Cave, 27.x.1968, J. Lowry (ANIC); 2 specs., King
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<normalizedToken id="C798B08ADC1AD9A5238CE73AF8F4CC64" originalValue="George’s">George's</normalizedToken>
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Sound, without further details (AMS). South Australia: 1 ♀, Koonalda Cave, 3.iv.1970, J. Lowry (ANIC); 12 specs., Fisher E.W. Line, coll. Toughlow & Wright, further details missing (AMS; 3 specs. in coll. TLAN); 1 spec., Fisher, Troughlow & Wright coll.; K.K.Spence collection, further details absent (AMS); 1 spec., Murray River, A.H. Elston coll., without further details (AMS).
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</paragraph>
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</subSubSection>
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<subSubSection id="F3F062589F166C18AA5248AFC8399887" pageId="196" pageNumber="197" type="biology_ecology">
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<paragraph id="ABB79ADCBDD44E1A098A9F33FBE835AD" pageId="196" pageNumber="197">Biology.</paragraph>
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<paragraph id="516BF9C72B89D1FC64D8C11E460586EE" pageId="196" pageNumber="197">This species is found in caves.</paragraph>
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</subSubSection>
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<subSubSection id="533EEE4507E0D6619EE5CC76002947AB" pageId="196" pageNumber="197" type="distribution">
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<paragraph id="3F44A0C9D6F30780A5B39E9B3BE3ADF7" pageId="196" pageNumber="197">Distribution.</paragraph>
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<paragraph id="038F619C4C26C0D2134192BC2548784D" pageId="196" pageNumber="197">South and Western Australia (Fig. 767).</paragraph>
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</subSubSection>
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<subSubSection id="AFB11167C8AC050E9827F6C169AD160B" lastPageId="199" lastPageNumber="200" pageId="196" pageNumber="197" type="description">
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<paragraph id="D8A00CCDBEA9A567710ADFC4F37623B2" pageId="196" pageNumber="197">Re-description.</paragraph>
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<paragraph id="000063FA6A01F10767F3470C8C1CB3E6" pageId="196" pageNumber="197">Body length: PEL: 3.25-3.40 mm; APW: 1.50-1.60 mm; PPW: 2.50-2.60 mm; EL: 2.20-2.25 mm; EW: 2.75-2.85 mm.</paragraph>
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<paragraph id="9FD067498FC185DA2297988835441E38" pageId="196" pageNumber="197">Body (Fig. 698) ovoid, moderately convex from above, underside slightly flattened, pronotum narrower than elytra, cuticle chestnut brown, without metallic luster; legs, mouthparts and antennae appendages similarly colored.</paragraph>
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<paragraph id="BB7AD1DB35361B93BE6F1BD14A094CAC" pageId="196" pageNumber="197">Antennal scape (Fig. 699) not particularly thickened, with two short setae; club rather large, oval, dorsally with two horizontal slit-like pits, ventrally with two vertical slit-like pits, entirely covered in dense short sensilla, intermingled with sparse longer erect sensilla; sensory structures of antennal club apart from sensory slit-like pits present dorsally and ventrally, not examined.</paragraph>
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<caption id="174A1B3E7FA161F8FBCBE0B1B964C10B" pageId="196" pageNumber="197">
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<paragraph id="7744B9152D8D4B7678B1182AEAB3CAF5" pageId="196" pageNumber="197">
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Figures 699-707. 699
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<taxonomicName id="866459C35AD65CF2BCB503061D6EA7B8" class="Insecta" family="Histeridae" genus="Tomogenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tomogenius motocola" order="Coleoptera" pageId="196" pageNumber="197" phylum="Arthropoda" rank="species" species="motocola">Tomogenius motocola</taxonomicName>
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Mazur, 1990 head, dorsal view 700 mentum, ventral view 701 propygidium + pygidium 702 prosternum 703 mesoventrite 704 metaventrite 705 lateral disc of metaventrite + metepisternum 706 protibia, dorsal view 707 ditto, ventral view.
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</paragraph>
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</caption>
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<paragraph id="4BF1006ECE8D2B4A63515C8ED7A7F33E" pageId="196" pageNumber="197">Mandibles (Fig. 699) impunctate, with rounded outer margin, acutely pointed, sub-apical tooth on inner margin of left mandible very small, almost inconspicuous; labrum even, sparsely punctate, approximately twice as wide as long, labral pits each with two moderately long labral setae; terminal labial palpomere elongated, its width about one-fourth its length; mentum (Fig. 700) sub-trapezoid, anterior angles slightly produced, anterior margin with a shallow median notch, surface around it two long setae from each side, lateral margins with a single one row of much shorter sparse ramose setae, disc with few scattered setae; rest of the mouthparts not examined.</paragraph>
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<paragraph id="18095DA28D4DC41CF1A09EA7059B431E" pageId="196" pageNumber="197">Clypeus (Fig. 699) even, rounded laterally, on anterior half with sparse fine punctures, separated by several times their diameter, punctures becoming larger and coarser on posterior half; frontal and supraorbital striae absent, occipital stria present, fine; frontal disc (Fig. 699) with fine round to ellipsoid punctures separated by several times their diameter; eyes convex, well visible from above.</paragraph>
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<paragraph id="D95D8364A0ACDB2BD5BCF2353AAB5969" lastPageId="197" lastPageNumber="198" pageId="196" pageNumber="197">
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Pronotal sides (Fig. 698) feebly convergent anteriorly, apical angles obtuse, marginal pronotal stria complete, thin, slightly carinate, somewhat weakened behind head; disc entirely punctate, medially punctures separated by several times their diameter, fine, becoming coarser and denser laterally, especially near apical angles where they
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<pageBreakToken id="B0952B89DA056583B8576B286E32D2F5" pageId="197" pageNumber="198" start="start">are</pageBreakToken>
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separated by less than their own diameter, large and ovoid; pronotal hypomeron glabrous; scutellum very small.
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</paragraph>
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<paragraph id="B6ED93C681DC60EC2B4601899963DE93" lastPageId="198" lastPageNumber="199" pageId="197" pageNumber="198">
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Elytral epipleura with scattered punctures of various sizes; marginal epipleural stria double, both striae weakly impressed but complete; marginal elytral stria well impressed, continuous along elytral apex as very weakened apical elytral stria; humeral elytral stria inconspicuous (absent?); inner subhumeral stria present as a short median fragment; elytral disc with four dorsal elytral striae 1-4, all striae in punctures, first originating at elytral base, other three striae slightly distanced from elytral base and their basal ends curved; first and second reaching about two-thirds of elytral length apically, third and fourth striae slightly shorter than first and second, surpassing elytral length apically; between fourth dorsal elytral and sutural striae a characteristic hooked appendix present, basally connected to basal fragment of sutural elytral stria; sutural elytral stria apart from this basal fragment evanescent on basal third, on apical two-thirds (roughly) present, fine, becoming weaker apically. Entire elytral disc punctate, on basal half (roughly) punctures very fine and sparse, separated several times their diameter, on apical half (roughly) punctures larger and denser, separated approximately
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<pageBreakToken id="CC8CC47D5504D1D96075F023541DAF05" pageId="198" pageNumber="199" start="start">by</pageBreakToken>
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their diameter, becoming denser apically; punctures near extreme elytral apex with minuscule striolae among them.
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</paragraph>
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<paragraph id="F84F618B455D67A6BAB259B685D897B9" pageId="198" pageNumber="199">Propygidium (Fig. 701) transverse, about four times as broad as long, completely exposed, with very dense punctures separated by less than their diameter; pygidium (Fig. 701) with sparser round punctures, separated by about their diameter, becoming sparser and finer towards apex.</paragraph>
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<paragraph id="79E58E1F2F3EE010796DDDAF7BC50D4A" pageId="198" pageNumber="199">Anterior margin of median portion of prosternum (Fig. 702) rounded; marginal prosternal stria present only laterally; prosternal process slightly convex on apical third between carinal prosternal striae, broad, dorso-laterally with sparse fine punctures; carinal prosternal striae (Fig. 702) straight, parallel, terminating near large and deep apical foveae separated by apex of prosternal process; lateral prosternal striae carinate, slightly convergent anteriorly, attaining apices of carinal prosternal striae.</paragraph>
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<paragraph id="EF5DAC8BF47982F9004B055DF50202BD" pageId="198" pageNumber="199">Discal marginal mesoventral stria (Fig. 703) well impressed, somewhat carinate, straight medially; disc flattened, with round punctures separated by about their diameter, punctures becoming finer medially, in female paratype examined punctures of mesoventrite with very short setae; meso-metaventral suture distinct, straight meso-metaventral sutural stria absent.</paragraph>
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<paragraph id="2F100327AE104F6A7BACB2B2F6025C8E" pageId="198" pageNumber="199">Intercoxal disc of metaventrite (Fig. 704) in examined female paratype medially with shallow depression; disc of metaventrite punctate, medially punctures finer and sparser, in male with minuscule setae becoming larger and coarser along lateral and basal margins. Lateral metaventral stria well impressed, carinate, slightly bisinuate, almost straight, not reaching metacoxa; lateral disc of metaventrite (Fig. 705) flattened, with round shallow large punctures; metepisternum + fused metepimeron (Fig. 705) evenly with much coarser and denser punctation; lateral metepisternal stria present, deeply impressed and almost complete.</paragraph>
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<paragraph id="B6F78F2BE580B2DFE307F983740A7275" pageId="198" pageNumber="199">Intercoxal disc of first abdominal ventrite flattened, completely striate laterally; surface of disc with scattered oblong punctation, punctures becoming sparser and finer medially.</paragraph>
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<paragraph id="D2598E9706BEE3CD9A380C4E29DFD8F8" pageId="198" pageNumber="199">Protibia (Fig. 706) flattened and somewhat dilated, outer margin with three low apical teeth topped by short denticle, followed by five widely spaced short denticles; setae of outer row short, moderately dense; setae of median row similarly dense and regular, slightly shorter than those of outer row; protarsal groove shallow; anterior protibial stria complete, costate; two thin, short tarsal denticles present apically; protibial spur short, straight, growing out from apical protibial margin; apical margin of protibia posteriorly with three tiny apical denticles; outer part of posterior surface of protibia (Fig. 707) finely imbricate, with a row of short setae; median part of posterior surface with an additional row of minuscule setae; posterior protibial stria complete, with scattered minuscule setae turning into a row of tightly-spaced short setae near apical margin; inner margin with double row of dense short setae.</paragraph>
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<paragraph id="D6EB8A177BCD387C32A62584268CEBBE" lastPageId="199" lastPageNumber="200" pageId="198" pageNumber="199">
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Mesotibia slender, outer margin with a single row of dense thin denticles growing in size apically; setae of outer row sparse, regular, short, growing somewhat longer apically; setae of median row irregular, much shorter than those of outer row; posterior mesotibial stria complete; anterior surface of mesotibia with dense row of well
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<pageBreakToken id="1A48E7EE22FB13110D569655E759D1A3" pageId="199" pageNumber="200" start="start">sclerotized</pageBreakToken>
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short denticles, with another similar row of much shorter and finer setae situated below it; anterior mesotibial stria complete, terminating in several tiny inner anterior denticles; mesotibial spur stout, short; apical margin with two tiny denticles; mesotarsus shorter than mesotibia; claws of apical tarsomere about half its length; metatibia basically similar to mesotibia, but denticles of outer margin much sparser than those of mesotibia; claws of apical tarsomere in both cases slightly bent, shorter than half its length.
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</paragraph>
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<paragraph id="80D550EB10C73BF1DEDB64050DD9F392" pageId="199" pageNumber="200">
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Male genitalia. Eighth sternite (Figs 708-709) fused on its entire length; vela laterally with two brushes of regular short setae creating a
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<normalizedToken id="1DB363D6B27F49451713252C8EB07222" originalValue="‘suction-cup’">'suction-cup'</normalizedToken>
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like appearance best observable from lateral view; eighth tergite apically straight, laterally with numerous pseudo-pores; eighth tergite and eighth sternite not fused laterally (Fig. 710). Ninth tergite (Figs 711-712) longitudinally divided medially; tenth tergite conspicuously small, basally slightly inwardly arcuate, apically rounded; spiculum gastrale (Fig. 711) gradually dilated near its apical end, basal end slightly dilated, spoon-like, outwardly arcuate. Aedeagus (Figs 713-714) slender, parallel-sided, parameres not fused on their apical third, apex of aedeagus acute (Fig. 715); basal piece of aedeagus short, ratio of its length: length of parameres 1: 3; aedeagus curved from lateral view (Fig. 714).
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</paragraph>
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<caption id="C7057058CE64C356562D1CBD03251F28" pageId="199" pageNumber="200">
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<paragraph id="E602D7EF85E4C3306767CCE81938CE12" pageId="199" pageNumber="200">
|
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Figures 708-715. 708
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<taxonomicName id="34B910DECD079A50FB623E0FFB6E4B37" class="Insecta" family="Histeridae" genus="Tomogenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tomogenius motocola" order="Coleoptera" pageId="199" pageNumber="200" phylum="Arthropoda" rank="species" species="motocola">Tomogenius motocola</taxonomicName>
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Mazur, 1990 male terminalia: 8th sternite + 8th tergite, ventral view 709 ditto, dorsal view 710 ditto, lateral view 711 male terminalia: 9th + 10th tergites, dorsal view; spiculum gastrale, ventral view 712 male terminalia: 9th + 10th tergites; spiculum gastrale, lateral view 713 male terminalia: aedeagus, dorsal view 714 ditto, lateral view 715 male terminalia: apex of aedeagus, frontal view.
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</paragraph>
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||
</caption>
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||
</subSubSection>
|
||
</treatment>
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</document> |