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<document id="8BBA4FEEA9676F5DBB3D6D766C53D272" ID-DOI="10.26107/RBZ-2019-0043" ID-ISSN="2345-7600" ID-Zenodo-Dep="5343617" ID-ZooBank="0F0AB24A-A37E-4DE0-B367-E2F5476FBB0E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630197339872" checkinUser="diego" docAuthor="Dayrat, Benoît, Goulding, Tricia C., Bourke, Adam J., Khalil, Munawar &amp; Tan, Shau Hwai" docDate="2019" docId="664587E01448FFC855378890FA5A5A7A" docLanguage="en" docName="RBZ-2019-0043.pdf" docOrigin="Raffles Bulletin of Zoology 67" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Melayonchis Dayrat &amp; Goulding 2017" docType="treatment" docVersion="3" lastPageNumber="569" masterDocId="9A7CFF981443FFC455418E70FF985E1E" masterDocTitle="New species and new records of Melayonchis slugs (Gastropoda: Euthyneura: Pulmonata: Onchidiidae)" masterLastPageNumber="585" masterPageNumber="557" pageNumber="568" updateTime="1699183113432" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="923E4E9FDC5C60000F11370AFE9F79E7">New species and new records of Melayonchis slugs (Gastropoda: Euthyneura: Pulmonata: Onchidiidae)</mods:title>
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<mods:namePart id="B39E869F6E8EE3F4589022AEC48FA09A">Dayrat, Benoît</mods:namePart>
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<mods:namePart id="2C449477E1E077A0ECFDB5B49D71F71D">Goulding, Tricia C.</mods:namePart>
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<mods:namePart id="0349FA55FF40ACB73DC6F4BFB6DBCFB9">Bourke, Adam J.</mods:namePart>
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<mods:namePart id="A0F16A8BA0DC9F37E3EF1EE3CD52646F">Khalil, Munawar</mods:namePart>
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<mods:namePart id="1528657A5E3131AE942AF8B84EEE5558">Tan, Shau Hwai</mods:namePart>
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<mods:title id="22F7B88C0510927927F4086C1B0C000A">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="B739FB7148D9FFCA2CBD6B4EB2F62DE1">2019</mods:date>
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<mods:number id="91035E205FAB8A0B3BAA42D55EF32FF6">2019-11-21</mods:number>
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<mods:number id="545D0F32447B9CCC773A67774CEFB10B">67</mods:number>
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<subSubSection id="A6F6657D1448FFCF55378890FD7358E7" box="[118,747,1760,1785]" pageId="11" pageNumber="568" type="nomenclature">
<paragraph id="EE5336F61448FFCF55378890FD7358E7" blockId="11.[118,747,1760,1785]" box="[118,747,1760,1785]" pageId="11" pageNumber="568">
<heading id="B51B819A1448FFCF55378890FD7358E7" bold="true" box="[118,747,1760,1785]" centered="true" fontSize="10" level="2" pageId="11" pageNumber="568" reason="2">
<emphasis id="DC98EAE41448FFCF55378890FD7358E7" bold="true" box="[118,747,1760,1785]" pageId="11" pageNumber="568">
<taxonomicName id="29EC4D751448FFCF55378890FE7658E7" ID-CoL="7P8NQ" authority="Dayrat &amp; Goulding" authorityName="Dayrat &amp; Goulding" authorityYear="2017" box="[118,494,1760,1785]" class="Gastropoda" family="Onchidiidae" genus="Melayonchis" kingdom="Animalia" order="Systellommatophora" pageId="11" pageNumber="568" phylum="Mollusca" rank="genus">
<emphasis id="DC98EAE41448FFCF55378890FE9B58E6" bold="true" box="[118,259,1760,1784]" italics="true" pageId="11" pageNumber="568">Melayonchis</emphasis>
Dayrat &amp; Goulding
</taxonomicName>
<emphasis id="DC98EAE41448FFCF54B78891FD9458E7" bold="true" box="[502,524,1761,1785]" italics="true" pageId="11" pageNumber="568">in</emphasis>
Dayrat et al., 2017
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A6F6657D1448FFCF55308951FE2F5947" pageId="11" pageNumber="568" type="type_taxon">
<paragraph id="EE5336F61448FFCF55308951FE2F5947" blockId="11.[113,752,1825,1881]" pageId="11" pageNumber="568">
<emphasis id="DC98EAE41448FFCF55308951FE915927" bold="true" box="[113,265,1825,1849]" pageId="11" pageNumber="568">
<typeStatus id="315788541448FFCF55308951FF335927" box="[113,171,1825,1849]" pageId="11" pageNumber="568">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="29EC4D751448FFCF54508951FDA65927" ID-CoL="3ZDG3" authority="Dayrat" authorityName="Dayrat" authorityYear="2017" box="[273,574,1825,1849]" class="Gastropoda" family="Onchidiidae" genus="Melayonchis" kingdom="Animalia" order="Systellommatophora" pageId="11" pageNumber="568" phylum="Mollusca" rank="species" species="eloisae">
<emphasis id="DC98EAE41448FFCF54508951FE755927" box="[273,493,1825,1849]" italics="true" pageId="11" pageNumber="568">Melayonchis eloisae</emphasis>
Dayrat
</taxonomicName>
<emphasis id="DC98EAE41448FFCF57078952FDC35927" box="[582,603,1826,1849]" italics="true" pageId="11" pageNumber="568">in</emphasis>
Dayrat et al., 2017, by original designation.
</paragraph>
</subSubSection>
<subSubSection id="A6F6657D1448FFCF553089F1FAD358A7" pageId="11" pageNumber="568" type="etymology">
<paragraph id="EE5336F61448FFCF553089F1FE4359C7" blockId="11.[113,752,1921,2009]" pageId="11" pageNumber="568">
<emphasis id="DC98EAE41448FFCF553089F1FF6C5987" bold="true" box="[113,244,1921,1945]" pageId="11" pageNumber="568">Etymology.</emphasis>
Combination of Melayu, the Malay word for Malays, and
<emphasis id="DC98EAE41448FFCF55B989D1FEDC59A7" box="[248,324,1953,1977]" italics="true" pageId="11" pageNumber="568">Onchis</emphasis>
, one of the old names used for onchidiid slugs (Dayrat et al., 2017: 1860).
</paragraph>
<paragraph id="EE5336F61448FFCF566C8811FAD358A7" blockId="11.[813,1451,1633,1722]" pageId="11" pageNumber="568">
<emphasis id="DC98EAE41448FFCF566C8811FC115867" bold="true" box="[813,905,1633,1657]" pageId="11" pageNumber="568">Gender.</emphasis>
Masculine, gender of
<emphasis id="DC98EAE41448FFCF513A8811FB595867" box="[1147,1217,1633,1657]" italics="true" pageId="11" pageNumber="568">onchis</emphasis>
(
<bibRefCitation id="8A7D4B071448FFCF518E8811FACD5867" author="ICZN" box="[1231,1365,1633,1657]" pageId="11" pageNumber="568" refId="ref18917" refString="ICZN (1999) International Code of Zoological Nomenclature. Fourth Edition. The International Trust for Zoological Nomenclature, London, UK, 306 pp." type="book" year="1999">ICZN, 1999</bibRefCitation>
; Article 30.1.1), a word derived from the masculine Greek word ὁ ὂγκος (
<emphasis id="DC98EAE41448FFCF563C88D1FC5158A7" box="[893,969,1697,1721]" italics="true" pageId="11" pageNumber="568">onchos</emphasis>
), which means ʻmassʼ or ʻtumour.ʼ
</paragraph>
</subSubSection>
<subSubSection id="A6F6657D1448FFC8566C8891FDAF5BBA" lastPageId="12" lastPageNumber="569" pageId="11" pageNumber="568" type="diagnosis">
<paragraph id="EE5336F61448FFC8566C8891FDBD5DFA" blockId="11.[813,1452,1761,2073]" lastBlockId="12.[136,775,940,996]" lastPageId="12" lastPageNumber="569" pageId="11" pageNumber="568">
<emphasis id="DC98EAE41448FFCF566C8891FC3958E7" bold="true" box="[813,929,1761,1785]" pageId="11" pageNumber="568">Diagnosis.</emphasis>
Body not flattened. No dorsal gills. Dorsal eyes present on notum. Retractable, central papilla present or absent. Foot wide. Eye tentacles short and narrow. Eyes at tip of ocular tentacles. Elongated, transversal protuberance on each oral lobe. Male opening below and on left of right ocular tentacle (in dorsal view). Pneumostome median. Female opening close to anus. Intestinal loops of
<typeStatus id="315788541448FFCF507089D2FAF159A4" box="[1329,1385,1954,1978]" pageId="11" pageNumber="568">types</typeStatus>
II and III. Rectal gland present. Lateral radular teeth with a strong protuberance on their inner lateral margin. Accessory penial gland and hollow spine present or absent. When present, accessory penial gland with no muscular sac. Penis short (&lt;
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), soft papilla with no hooks.
</paragraph>
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<paragraph id="EE5336F6144FFFC855C98D55FCFD5D6B" blockId="12.[136,1474,805,885]" pageId="12" pageNumber="569">
Fig. 7. Habitats,
<taxonomicName id="29EC4D75144FFFC854728D55FE625D24" authorityName="Dayrat &amp; Goulding" authorityYear="2019" box="[307,506,805,826]" class="Gastropoda" family="Onchidiidae" genus="Melayonchis" kingdom="Animalia" order="Systellommatophora" pageId="12" pageNumber="569" phylum="Mollusca" rank="species" species="eberlyi">
<emphasis id="DC98EAE4144FFFC854728D55FE625D24" box="[307,506,805,826]" italics="true" pageId="12" pageNumber="569">Melayonchis eberlyi</emphasis>
</taxonomicName>
, unique site (type locality) where
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<emphasis id="DC98EAE4144FFFC856188D55FBB95D24" box="[857,1057,805,826]" italics="true" pageId="12" pageNumber="569">Melayonchis eberlyi</emphasis>
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was found and which corresponds to the stations 27 (in 2011), 262 (in 2016), and 263 (in 2016) (A, B). Old forest with tall
<taxonomicName id="29EC4D75144FFFC856F78D32FBBE5D49" box="[950,1062,834,855]" class="Magnoliopsida" family="Rhizophoraceae" genus="Rhizophora" kingdom="Plantae" order="Malpighiales" pageId="12" pageNumber="569" phylum="Tracheophyta" rank="genus">
<emphasis id="DC98EAE4144FFFC856F78D32FBBE5D49" box="[950,1062,834,855]" italics="true" pageId="12" pageNumber="569">Rhizophora</emphasis>
</taxonomicName>
trees, high in the tidal zone (ferns), in the educational mangrove preserve near Kuala Sepatang, Peninsular Malaysia.
</paragraph>
</caption>
<paragraph id="EE5336F6144FFFC855C98A7CFDAF5BBA" blockId="12.[136,775,1036,1444]" pageId="12" pageNumber="569">
<emphasis id="DC98EAE4144FFFC855C98A7CFEF75A3A" bold="true" box="[136,367,1036,1060]" pageId="12" pageNumber="569">Distinctive features.</emphasis>
Externally,
<taxonomicName id="29EC4D75144FFFC854B98A7CFD185A3A" authorityName="Dayrat &amp; Goulding" authorityYear="2017" box="[504,640,1036,1060]" class="Gastropoda" family="Onchidiidae" genus="Melayonchis" kingdom="Animalia" order="Systellommatophora" pageId="12" pageNumber="569" phylum="Mollusca" rank="genus">
<emphasis id="DC98EAE4144FFFC854B98A7CFD185A3A" box="[504,640,1036,1060]" italics="true" pageId="12" pageNumber="569">Melayonchis</emphasis>
</taxonomicName>
slugs could be easily confused with
<taxonomicName id="29EC4D75144FFFC854D18A5CFC9F5A5A" authority="Dayrat &amp; Goulding" authorityName="Dayrat &amp; Goulding" authorityYear="2018" box="[400,775,1068,1093]" class="Gastropoda" family="Onchidiidae" genus="Marmaronchis" kingdom="Animalia" order="Systellommatophora" pageId="12" pageNumber="569" phylum="Mollusca" rank="genus">
<emphasis id="DC98EAE4144FFFC854D18A5CFDB75A5A" box="[400,559,1068,1092]" italics="true" pageId="12" pageNumber="569">Marmaronchis</emphasis>
Dayrat &amp; Goulding
</taxonomicName>
<emphasis id="DC98EAE4144FFFC855C88A3DFF065A7A" box="[137,158,1101,1124]" italics="true" pageId="12" pageNumber="569">in</emphasis>
<bibRefCitation id="8A7D4B07144FFFC855E48A3DFEE95A7A" author="Dayrat B &amp; Goulding TC &amp; Khalil M &amp; Lozouet P &amp; Tan SH" box="[165,369,1100,1125]" pageId="12" pageNumber="569" pagination="814 - 837" refId="ref18385" refString="Dayrat B, Goulding TC, Khalil M, Lozouet P &amp; Tan SH (2018) Systematic revision one clade at a time: A new genus of onchidiid slugs from the Indo-West Pacific (Gastropoda: Euthyneura: Onchidiidae). Raffles Bulletin of Zoology, 66: 814 - 837." type="journal article" year="2018">Dayrat et al., 2018</bibRefCitation>
, or
<taxonomicName id="29EC4D75144FFFC854DC8A3CFC9F5A7B" authority="Goulding &amp; Dayrat" authorityName="Goulding &amp; Dayrat" authorityYear="2018" box="[413,775,1100,1125]" class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" kingdom="Animalia" order="Systellommatophora" pageId="12" pageNumber="569" phylum="Mollusca" rank="genus">
<emphasis id="DC98EAE4144FFFC854DC8A3CFDB65A7A" box="[413,558,1100,1124]" italics="true" pageId="12" pageNumber="569">Wallaconchis</emphasis>
Goulding &amp; Dayrat
</taxonomicName>
<emphasis id="DC98EAE4144FFFC855C88A1DFF065A9A" box="[137,158,1133,1156]" italics="true" pageId="12" pageNumber="569">in</emphasis>
<bibRefCitation id="8A7D4B07144FFFC855E68A1CFE3C5A9A" author="Goulding TC &amp; Khalil M &amp; Tan SH &amp; Dayrat B" box="[167,420,1132,1156]" pageId="12" pageNumber="569" pagination="1 - 111" refId="ref18681" refString="Goulding TC, Khalil M, Tan SH &amp; Dayrat B (2018 b) Integrative taxonomy of a new and highly-diverse genus of onchidiid slugs from the Coral Triangle (Gastropoda: Pulmonata: Onchidiidae). ZooKeys, 763: 1 - 111." type="journal article" year="2018">Goulding et al., 2018b</bibRefCitation>
, even though the colour pattern of some
<taxonomicName id="29EC4D75144FFFC855B48AFCFE725ABA" box="[245,490,1164,1188]" class="Gastropoda" family="Onchidiidae" genus="Melayonchis" kingdom="Animalia" order="Systellommatophora" pageId="12" pageNumber="569" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="DC98EAE4144FFFC855B48AFCFE125ABA" box="[245,394,1164,1188]" italics="true" pageId="12" pageNumber="569">Melayonchis</emphasis>
species
</taxonomicName>
is very distinctive and unmistakable. Internally, however, the short (&lt;
<quantity id="29149B13144FFFC857CE8ADCFD4A5ADA" box="[655,722,1196,1220]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="569" unit="mm" value="1.0">1 mm</quantity>
), the soft papilla with no hooks is unique to
<taxonomicName id="29EC4D75144FFFC8577B8ABCFD5A5AFA" authorityName="Dayrat &amp; Goulding" authorityYear="2017" box="[570,706,1228,1252]" class="Gastropoda" family="Onchidiidae" genus="Melayonchis" kingdom="Animalia" order="Systellommatophora" pageId="12" pageNumber="569" phylum="Mollusca" rank="genus">
<emphasis id="DC98EAE4144FFFC8577B8ABCFD5A5AFA" box="[570,706,1228,1252]" italics="true" pageId="12" pageNumber="569">Melayonchis</emphasis>
</taxonomicName>
slugs. In addition, the lateral radular teeth of
<taxonomicName id="29EC4D75144FFFC8570F8A9CFD435B1A" authorityName="Dayrat &amp; Goulding" authorityYear="2017" box="[590,731,1260,1284]" class="Gastropoda" family="Onchidiidae" genus="Melayonchis" kingdom="Animalia" order="Systellommatophora" pageId="12" pageNumber="569" phylum="Mollusca" rank="genus">
<emphasis id="DC98EAE4144FFFC8570F8A9CFD435B1A" box="[590,731,1260,1284]" italics="true" pageId="12" pageNumber="569">Melayonchis</emphasis>
</taxonomicName>
are characterised by a strong protuberance on their inner lateral margin. A similar protuberance is also present in radular teeth of
<taxonomicName id="29EC4D75144FFFC8559F8B3CFEC45B7A" authorityName="Baker" authorityYear="1938" box="[222,348,1356,1380]" class="Gastropoda" family="Onchidiidae" genus="Platevindex" kingdom="Animalia" order="Systellommatophora" pageId="12" pageNumber="569" phylum="Mollusca" rank="genus">
<emphasis id="DC98EAE4144FFFC8559F8B3CFEC45B7A" box="[222,348,1356,1380]" italics="true" pageId="12" pageNumber="569">Platevindex</emphasis>
</taxonomicName>
and
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<emphasis id="DC98EAE4144FFFC854CE8B3CFDB65B7A" box="[399,558,1356,1380]" italics="true" pageId="12" pageNumber="569">Marmaronchis</emphasis>
</taxonomicName>
slugs. However, the protuberance of
<taxonomicName id="29EC4D75144FFFC854798B1CFE585B9A" authorityName="Dayrat &amp; Goulding" authorityYear="2017" box="[312,448,1388,1412]" class="Gastropoda" family="Onchidiidae" genus="Melayonchis" kingdom="Animalia" order="Systellommatophora" pageId="12" pageNumber="569" phylum="Mollusca" rank="genus">
<emphasis id="DC98EAE4144FFFC854798B1CFE585B9A" box="[312,448,1388,1412]" italics="true" pageId="12" pageNumber="569">Melayonchis</emphasis>
</taxonomicName>
radular teeth is much stronger than in
<taxonomicName id="29EC4D75144FFFC8559F8BFCFEC45BBA" authorityName="Baker" authorityYear="1938" box="[222,348,1420,1444]" class="Gastropoda" family="Onchidiidae" genus="Platevindex" kingdom="Animalia" order="Systellommatophora" pageId="12" pageNumber="569" phylum="Mollusca" rank="genus">
<emphasis id="DC98EAE4144FFFC8559F8BFCFEC45BBA" box="[222,348,1420,1444]" italics="true" pageId="12" pageNumber="569">Platevindex</emphasis>
</taxonomicName>
and
<taxonomicName id="29EC4D75144FFFC854D58BFCFDA95BBA" authorityName="Dayrat &amp; Goulding" authorityYear="2018" box="[404,561,1420,1444]" class="Gastropoda" family="Onchidiidae" genus="Marmaronchis" kingdom="Animalia" order="Systellommatophora" pageId="12" pageNumber="569" phylum="Mollusca" rank="genus">
<emphasis id="DC98EAE4144FFFC854D58BFCFDA95BBA" box="[404,561,1420,1444]" italics="true" pageId="12" pageNumber="569">Marmaronchis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="A6F6657D144FFFC855C98BBCFD9158DA" pageId="12" pageNumber="569" type="distribution">
<paragraph id="EE5336F6144FFFC855C98BBCFD9158DA" blockId="12.[135,775,1484,1732]" pageId="12" pageNumber="569">
<emphasis id="DC98EAE4144FFFC855C98BBCFE105BFA" bold="true" box="[136,392,1484,1508]" pageId="12" pageNumber="569">
Distribution (
<figureCitation id="76D72A73144FFFC8546E8BBCFEE15BFA" box="[303,377,1484,1508]" captionStart="Fig" captionStartId="10.[135,170,1978,2000]" captionTargetBox="[162,1448,189,1964]" captionTargetId="figure-8@10.[134,1471,172,1981]" captionTargetPageId="10" captionText="Fig. 6. Geographic distribution of the six known Melayonchis species. The many blue dots correspond to regions where we collected gastropods since 2010 (with several sites per dot), clearly indicating where we have not found Melayonchis (e.g., Philippines, Sulawesi, Queensland). So far, Melayonchis was only found in the Andaman Islands, on both sides of the Strait of Malacca (western Peninsular Malaysia and eastern Sumatra), Singapore, southern Vietnam, Brunei Darussalam, and Northern territory. The colours used for each species are the same as those used in Figs. 3–5." figureDoi="http://doi.org/10.5281/zenodo.5343623" httpUri="https://zenodo.org/record/5343623/files/figure.png" pageId="12" pageNumber="569">Fig. 6</figureCitation>
).
</emphasis>
<taxonomicName id="29EC4D75144FFFC854D28BBCFDBA5BFA" authorityName="Dayrat &amp; Goulding" authorityYear="2017" box="[403,546,1484,1508]" class="Gastropoda" family="Onchidiidae" genus="Melayonchis" kingdom="Animalia" order="Systellommatophora" pageId="12" pageNumber="569" phylum="Mollusca" rank="genus">
<emphasis id="DC98EAE4144FFFC854D28BBCFDBA5BFA" box="[403,546,1484,1508]" italics="true" pageId="12" pageNumber="569">Melayonchis</emphasis>
</taxonomicName>
is known from the Andaman Islands, both sides of the Strait of
<collectingCountry id="96FB7666144FFFC857E68B9CFC9E581A" box="[679,774,1516,1540]" name="Malaysia" pageId="12" pageNumber="569">Malacca</collectingCountry>
(western Peninsular
<collectingCountry id="96FB7666144FFFC85433887CFE4F583A" box="[370,471,1548,1572]" name="Malaysia" pageId="12" pageNumber="569">Malaysia</collectingCountry>
and eastern Sumatra), the South
<collectingCountry id="96FB7666144FFFC85591885CFE88585A" box="[208,272,1580,1604]" name="China" pageId="12" pageNumber="569">China</collectingCountry>
Sea (southern
<collectingCountry id="96FB7666144FFFC854FB885CFD81585A" box="[442,537,1580,1604]" name="Vietnam" pageId="12" pageNumber="569">Vietnam</collectingCountry>
,
<collectingCountry id="96FB7666144FFFC85766885CFD63585A" box="[551,763,1580,1604]" name="Brunei" pageId="12" pageNumber="569">Brunei Darussalam</collectingCountry>
), and the
<collectingRegion id="2C28F814144FFFC8559D883CFE3E587A" box="[220,422,1612,1636]" country="Australia" name="Northern Territory" pageId="12" pageNumber="569">Northern Territory</collectingRegion>
,
<collectingCountry id="96FB7666144FFFC854F1883CFD8D587A" box="[432,533,1612,1636]" name="Australia" pageId="12" pageNumber="569">Australia</collectingCountry>
. The Strait of
<collectingCountry id="96FB7666144FFFC857EC883CFC9F587A" box="[685,775,1612,1636]" name="Malaysia" pageId="12" pageNumber="569">Malacca</collectingCountry>
is the core of the geographic distribution of
<taxonomicName id="29EC4D75144FFFC85711881CFD40589A" authorityName="Dayrat &amp; Goulding" authorityYear="2017" box="[592,728,1644,1668]" class="Gastropoda" family="Onchidiidae" genus="Melayonchis" kingdom="Animalia" order="Systellommatophora" pageId="12" pageNumber="569" phylum="Mollusca" rank="genus">
<emphasis id="DC98EAE4144FFFC85711881CFD40589A" box="[592,728,1644,1668]" italics="true" pageId="12" pageNumber="569">Melayonchis</emphasis>
</taxonomicName>
: the Matang mangrove, near Kuala Sepatang, western Peninsular
<collectingCountry id="96FB7666144FFFC855C688DCFF7358DA" box="[135,235,1708,1732]" name="Malaysia" pageId="12" pageNumber="569">Malaysia</collectingCountry>
, hosts all but one species.
</paragraph>
</subSubSection>
<subSubSection id="A6F6657D144FFFC855C9889CFA5A5A7A" pageId="12" pageNumber="569" type="discussion">
<paragraph id="EE5336F6144FFFC855C9889CFA5A5A7A" blockId="12.[135,775,1772,2116]" lastBlockId="12.[835,1475,940,1124]" pageId="12" pageNumber="569">
<emphasis id="DC98EAE4144FFFC855C9889CFF64591A" bold="true" box="[136,252,1772,1796]" pageId="12" pageNumber="569">Remarks.</emphasis>
Differences among
<taxonomicName id="29EC4D75144FFFC854AC889CFD41591A" box="[493,729,1772,1796]" class="Gastropoda" family="Onchidiidae" genus="Melayonchis" kingdom="Animalia" order="Systellommatophora" pageId="12" pageNumber="569" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="DC98EAE4144FFFC854AC889CFDE4591A" box="[493,636,1772,1796]" italics="true" pageId="12" pageNumber="569">Melayonchis</emphasis>
species
</taxonomicName>
are summarised in
<tableCitation id="A36E034D144FFFC85466897CFEEF593A" box="[295,375,1804,1828]" captionStart="Table 3" captionStartId="11.[113,168,189,211]" captionText="Table 3. Morphological differences among Melayonchis species. Observations between parentheses are only occasional. All traits are subject to individual variation. Information regarding M. aileenae, M. annae, M. eloisae, and M. siongkiati is from Dayrat et al. (2017). The orientation of the transitional loop (TL) is provided along with the type of intestinal loops. For the radular formulae, the range of number of rows (e.g., 70 to 80 rows in M. eberlyi) and the range of number of lateral teeth per half row (e.g., 140 to 180 in M. eberlyi) are provided." pageId="12" pageNumber="569">Table 3</tableCitation>
. Both the diagnosis and the distinctive features above are slightly updated from our previous description (Dayrat et al., 2017: 1860) to incorporate new findings: the retractable central papilla is absent in some species (described as present in the 2017 diagnosis) and, when present, is only slightly larger than the other dorsal papillae; there are no intestinal loops “intermediary between
<typeStatus id="31578854144FFFC855C9899DFF58561B" box="[136,192,2029,2053]" pageId="12" pageNumber="569">types</typeStatus>
II and III” and intestinal loops are clearly of
<typeStatus id="31578854144FFFC857FF899DFD74561B" box="[702,748,2029,2053]" pageId="12" pageNumber="569">type</typeStatus>
II or of
<typeStatus id="31578854144FFFC8558B867DFF61563B" box="[202,249,2061,2085]" pageId="12" pageNumber="569">type</typeStatus>
III (
<figureCitation id="76D72A73144FFFC85470867CFEEF563A" box="[305,375,2060,2084]" captionStart="Fig" captionStartId="5.[113,148,1724,1746]" captionTargetBox="[118,1455,271,1708]" captionTargetId="figure-13@5.[105,1470,265,1732]" captionTargetPageId="5" captionText="Fig. 2. Onchidiid intestinal type III. Small black arrows indicate the direction of the intestinal transport, which starts in the blue loop. A blue loop turns clockwise. A yellow loop turns counterclockwise. A green loop is transitional between a blue loop and a yellow loop. The orientation of the transitional (green) loop is indicated with a red arrow (e.g., at 3 o’clock in C, at 6 o’clock in I, and at 12 o’clock in J). For details about the Melayonchis and Onchidium individuals illustrated here, see Dayrat et al. (2016, 2017). A, Plate’s (1893: pl. 8, fig. 31a) original figure of a type III, Onchidium nigrum (junior synonym of Onchidium stuxbergi), redrawn (scale unclear), colours indicate Plate’s interpretation; B, same as A, colours indicate a new interpretation based on two parts (in red) drawn by mistake by Plate (see the text); C, O. stuxbergi (from Dayrat et al., 2016: fig. 11B), scale bar = 5 mm; D, O. stuxbergi, (PNM 041200), scale bar = 4 mm; E, O. stuxbergi, [5605], scale bar = 5 mm; F, Melayonchis eloisae, [1011], scale bar = 2 mm; G, M. eloisae, [922], scale bar = 2 mm; H, M. eloisae, [5607], scale bar = 1 mm; I, M. eloisae, [951], scale bar = 2 mm; J, M. aileenae, [5631], scale bar = 2 mm; K, M. aileenae (from Dayrat et al., 2017: fig. 23a), scale bar = 3 mm; L, M. aileenae, [970], scale bar = 2 mm; M, M. siongkiati, [1052], scale bar = 2 mm; N, M. siongkiati, [5608], scale bar = 3 mm; O, M. siongkiati (from Dayrat et al., 2017: fig. 12a), scale bar = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5343621" httpUri="https://zenodo.org/record/5343621/files/figure.png" pageId="12" pageNumber="569">Fig. 2</figureCitation>
); finally, in 2017, the protuberance on the inner lateral margin of the lateral radular teeth was mentioned as a distinctive feature unique to
<taxonomicName id="29EC4D75144FFFC850728DDCFA235DDA" authorityName="Dayrat &amp; Goulding" authorityYear="2017" box="[1331,1467,940,964]" class="Gastropoda" family="Onchidiidae" genus="Melayonchis" kingdom="Animalia" order="Systellommatophora" pageId="12" pageNumber="569" phylum="Mollusca" rank="genus">
<emphasis id="DC98EAE4144FFFC850728DDCFA235DDA" box="[1331,1467,940,964]" italics="true" pageId="12" pageNumber="569">Melayonchis</emphasis>
</taxonomicName>
, but, after dissecting many other onchidiid slugs since then (
<bibRefCitation id="8A7D4B07144FFFC8560B8D9CFBBA5A1A" author="Dayrat B &amp; Goulding TC &amp; Khalil M &amp; Lozouet P &amp; Tan SH" box="[842,1058,1004,1028]" pageId="12" pageNumber="569" pagination="814 - 837" refId="ref18385" refString="Dayrat B, Goulding TC, Khalil M, Lozouet P &amp; Tan SH (2018) Systematic revision one clade at a time: A new genus of onchidiid slugs from the Indo-West Pacific (Gastropoda: Euthyneura: Onchidiidae). Raffles Bulletin of Zoology, 66: 814 - 837." type="journal article" year="2018">Dayrat et al., 2018</bibRefCitation>
,
<bibRefCitation id="8A7D4B07144FFFC851718D9CFBED5A1A" author="Dayrat B &amp; Goulding TC &amp; Khalil M &amp; Apte D &amp; Bourke AJ &amp; Comendador J &amp; Tan SH" box="[1072,1141,1004,1028]" pageId="12" pageNumber="569" pagination="1 - 77" refId="ref18439" refString="Dayrat B, Goulding TC, Khalil M, Apte D, Bourke AJ, Comendador J &amp; Tan SH (2019 a) A new genus and three new species of mangrove slugs from the Indo-West Pacific (Gastropoda: Euthyneura: Onchidiidae). European Journal of Taxonomy, 500: 1 - 77." type="journal article" year="2019">2019a</bibRefCitation>
, b;
<bibRefCitation id="8A7D4B07144FFFC851E38D9CFA385A1A" author="Goulding TC &amp; Khalil M &amp; Tan SH &amp; Dayrat B" box="[1186,1440,1004,1028]" pageId="12" pageNumber="569" pagination="337 - 349" refId="ref18634" refString="Goulding TC, Khalil M, Tan SH &amp; Dayrat B (2018 a) A new genus and a new species of onchidiid slugs from eastern Indonesia (Gastropoda: Euthyneura: Onchidiidae). Raffles Bulletin of Zoology, 66: 337 - 349." type="journal article" year="2018">Goulding et al., 2018a</bibRefCitation>
, b, c), we know now that a smaller but similar protuberance is found in
<taxonomicName id="29EC4D75144FFFC856E98A5CFBBE5A5A" authorityName="Baker" authorityYear="1938" box="[936,1062,1068,1092]" class="Gastropoda" family="Onchidiidae" genus="Platevindex" kingdom="Animalia" order="Systellommatophora" pageId="12" pageNumber="569" phylum="Mollusca" rank="genus">
<emphasis id="DC98EAE4144FFFC856E98A5CFBBE5A5A" box="[936,1062,1068,1092]" italics="true" pageId="12" pageNumber="569">Platevindex</emphasis>
</taxonomicName>
and
<taxonomicName id="29EC4D75144FFFC8511C8A5CFB625A5A" authorityName="Dayrat &amp; Goulding" authorityYear="2018" box="[1117,1274,1068,1092]" class="Gastropoda" family="Onchidiidae" genus="Marmaronchis" kingdom="Animalia" order="Systellommatophora" pageId="12" pageNumber="569" phylum="Mollusca" rank="genus">
<emphasis id="DC98EAE4144FFFC8511C8A5CFB625A5A" box="[1117,1274,1068,1092]" italics="true" pageId="12" pageNumber="569">Marmaronchis</emphasis>
</taxonomicName>
, but that the short (&lt;
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), soft penial papilla is truly unique to
<taxonomicName id="29EC4D75144FFFC850728A3CFA235A7A" authorityName="Dayrat &amp; Goulding" authorityYear="2017" box="[1331,1467,1100,1124]" class="Gastropoda" family="Onchidiidae" genus="Melayonchis" kingdom="Animalia" order="Systellommatophora" pageId="12" pageNumber="569" phylum="Mollusca" rank="genus">
<emphasis id="DC98EAE4144FFFC850728A3CFA235A7A" box="[1331,1467,1100,1124]" italics="true" pageId="12" pageNumber="569">Melayonchis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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