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<mods:title id="D0C035C96B215E82517DFCF6CAC77102">Scorpions of Sri Lanka (Scorpiones Buthidae, Chaerilidae, Scorpionidae) with description of four new species of the genera Charmus Karsch, 1879 and Reddyanus Vachon, 1972, stat n</mods:title>
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<taxonomicName id="FFD84D4AA83BFFBCFEFAFA419CCAFA13" ID-CoL="6MZ76" authority="Pocock, 1897" authorityName="Pocock" authorityYear="1897" box="[314,680,1430,1454]" class="Arachnida" family="Buthidae" genus="Isometrus" kingdom="Animalia" order="Scorpiones" pageId="44" pageNumber="43" phylum="Arthropoda" rank="species" species="thwaitesi">
<emphasis id="0AACEADBA83BFFBCFEFAFA419C72FA13" bold="true" box="[314,528,1430,1454]" italics="true" pageId="44" pageNumber="43">Isometrus thwaitesi</emphasis>
<bibRefCitation id="5C494B38A83BFFBCFDD7FA409CCAFA13" author="Pocock" box="[535,680,1431,1454]" firstAuthor="Pocock" journalOrPublisher="Journal of the Bombay Natural History Society" pageId="44" pageNumber="43" pagination="102 - 117" part="11" refId="ref43758" refString="POCOCK, R. I. 1897. Descriptions of some new species of scorpions from India. Journal of the Bombay Natural History Society, 11: 102 - 117." title="Descriptions of some new species of scorpions from India" type="journal article" year="1897">Pocock, 1897</bibRefCitation>
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<paragraph id="386736C9A83BFFBCFF0EFA629CECFA56" blockId="44.[206,788,1430,1515]" pageId="44" pageNumber="43">
(
<figureCitation id="A0E32A4CA83BFFBCFF17FA629F4FFA71" box="[215,301,1461,1484]" captionStart="Figures 1215" captionStartId="6.[192,278,1885,1909]" captionTargetBox="[191,1440,167,1856]" captionTargetId="figure-11@6.[192,1439,168,1855]" captionTargetPageId="6" captionText="Figures 1215: Maps showing the known geographical distribution of the Sri Lankan species." figureDoi="http://doi.org/10.5281/zenodo.7124476" httpUri="https://zenodo.org/record/7124476/files/figure.png" pageId="44" pageNumber="43">Figs. 13</figureCitation>
,
<figureCitation id="A0E32A4CA83BFFBCFEFAFA629FF5FA71" box="[314,407,1461,1484]" captionStart-0="Figures 138141" captionStart-1="Figures 142145" captionStart-2="Figures 146149" captionStart-3="Figures 150151" captionStartId-0="36.[192,278,1582,1606]" captionStartId-1="37.[192,278,1844,1868]" captionStartId-2="38.[192,278,1446,1470]" captionStartId-3="39.[192,278,1829,1853]" captionTargetBox-0="[210,1401,172,1544]" captionTargetBox-1="[213,1417,167,1815]" captionTargetBox-2="[194,1436,179,1409]" captionTargetBox-3="[191,1440,197,1801]" captionTargetId-0="figure-72@36.[192,1439,168,1553]" captionTargetId-1="figure-15@37.[214,1416,168,1814]" captionTargetId-2="figure-11@38.[192,1439,168,1417]" captionTargetId-3="figure-6@39.[192,1439,197,1800]" captionTargetPageId-0="36" captionTargetPageId-1="37" captionTargetPageId-2="38" captionTargetPageId-3="39" captionText-0="Figures 138141: Isometrus thwaitesi. Figures 138139. Male from locality 15CH in dorsal (138) and ventral (139) views. Figures 140141. Female from locality 15CO in dorsal (140) and ventral (141) views." captionText-1="Figures 142145: Isometrus thwaitesi. Figures 142, 144. Male from locality 15CH, chelicerae, carapace and tergites IIII (142) and sternopectinal region and sternite III (144). Figures 143, 145. Female from locality 15CO, chelicerae, carapace and tergites IIII (143) and sternopectinal region and sternite III (145)." captionText-2="Figures 146149: Left hemispermatophore of Isometrus thwaitesi from locality 15CH. External (146), dorsal (147), internal (148) and ventral (149) views. Note: distal part of flagellum and basal part of trunk (foot) were damaged during dissection. Scale bar: 500 µm. Abbreviations: bl, basal lobe; f, flagellum; ml, median lobe; mlc, median lobe carina; pr, pars recta of flagellum; prf, pars reflecta of flagellum; t, trunk." captionText-3="Figures 150151: Isometrus thwaitesi from locality 15CF, male (150) and female (151)." figureDoi-0="http://doi.org/10.5281/zenodo.7124565" figureDoi-1="http://doi.org/10.5281/zenodo.7124569" figureDoi-2="http://doi.org/10.5281/zenodo.7124574" figureDoi-3="http://doi.org/10.5281/zenodo.7124576" httpUri-0="https://zenodo.org/record/7124565/files/figure.png" httpUri-1="https://zenodo.org/record/7124569/files/figure.png" httpUri-2="https://zenodo.org/record/7124574/files/figure.png" httpUri-3="https://zenodo.org/record/7124576/files/figure.png" pageId="44" pageNumber="43">138151</figureCitation>
,
<figureCitation id="A0E32A4CA83BFFBCFE65FA629FAFFA71" box="[421,461,1461,1484]" captionStart="Figures 193200" captionStartId="48.[192,278,1070,1094]" captionTargetBox="[213,1446,171,1037]" captionTargetId="figure-11@48.[205,1452,168,1041]" captionTargetPageId="48" captionText="Figures 193200: Distal segments of leg IV, retroventral view of Sri Lankan Buthidae and Chaerilidae genera. Buthoscorpio sarasinorum, male from locality 15CF (193), Charmus laneus, male from locality 15CO (194), Hottentotta tamulus, male from locality 15CK (195), Isometrus maculatus, female from locality 15CI (196), Isometrus thwaitesi, female from locality 15CO (197), Lychas srilankensis, male from locality 15CJ (198), Reddyanus basilicus, male from locality 15CS (199), and Chaerilus ceylonensis, male from locality 15CD (200)." figureDoi="http://doi.org/10.5281/zenodo.7124608" httpUri="https://zenodo.org/record/7124608/files/figure.png" pageId="44" pageNumber="43">197</figureCitation>
,
<figureCitation id="A0E32A4CA83BFFBCFE1AFA629C55FA71" box="[474,567,1461,1484]" captionStart="Figures 209224" captionStartId="50.[192,278,1736,1760]" captionTargetBox="[208,1434,174,1703]" captionTargetId="figure-11@50.[205,1452,168,1707]" captionTargetPageId="50" captionText="Figures 209224: Metasomal segments IIIII, lateral view of Sri Lankan Reddyanus and Isometrus species. Figures 209210. Reddyanus basilicus, male (209) and female (210) from locality 15CR. Figures 211212. R. besucheti, male holotype (211) and female from locality 15CG (212). Figures 213214. R. ceylonensis sp. n., male holotype (213) and female paratype from locality 15CI (214). Figures 215216. R. jayarathnei sp. n., male (215) and female (216) paratypes. Figures 217218. R. loebli, male (217) and female (218) from locality 15CG. Figures 219220. R. ranawanai sp. n., male holotype (219) and female paratype (220). Figures 221222. Isometrus maculatus, male from locality 15CP (221) and female from locality 15CI (222). Figures 223 224. I. thwaitesi, male from locality 15CH (223) and female from locality 15CO (224)." figureDoi="http://doi.org/10.5281/zenodo.7124622" httpUri="https://zenodo.org/record/7124622/files/figure.png" pageId="44" pageNumber="43">223224</figureCitation>
,
<figureCitation id="A0E32A4CA83BFFBCFD85FA629F19FA56" captionStart-0="Figures 225240" captionStart-1="Figures 241259" captionStartId-0="51.[192,278,1493,1517]" captionStartId-1="52.[192,278,1733,1757]" captionTargetBox-0="[193,1432,180,1451]" captionTargetBox-1="[238,1387,153,1699]" captionTargetId-0="figure-6@51.[192,1439,179,1464]" captionTargetId-1="figure-11@52.[234,1392,152,1704]" captionTargetPageId-0="51" captionTargetPageId-1="52" captionText-0="Figures 225240: Pedipalp, dorsal view of Sri Lankan Reddyanus and Isometrus species. Figures 225226. Reddyanus basilicus, male (225) and female (226) from locality 15CR. Figures 227228. R. besucheti, male from locality 15CD (227) and female from locality 15CG (228). Figures 229230. R. ceylonensis sp. n., male holotype (229) and female paratype from locality 15CI (230). Figures 231232. R. jayarathnei sp. n., male (231) and female (232) paratypes. Figures 233234. R. loebli, male (233) and female (234) from locality 15CG. Figures 235236. R. ranawanai sp. n., male holotype (235) and female paratype (236). Figures 237238. Isometrus maculatus, male from locality 15CP (237) and female from locality 15CI (238). Figures 239240. I. thwaitesi, male from locality 15CH (239) and female from locality 15CO (240)." captionText-1="Figures 241259: Pedipalp chela dorsal (241), movable finger (242250), and fixed finger (251259) of Sri Lankan Isometrus, Lychas, and Reddyanus species. Figures 241242, 252. Isometrus maculatus, female from locality 15CO (241) and male from locality 15CP (242, 252). Figures 243244, 253. I. thwaitesi, male from locality 15CH (243) and female from locality 15CO (244, 253). Figures 245, 254. Reddyanus basilicus, male from locality 15CS. Figures 246, 255. R. besucheti, male from locality 15CD. Figures 247, 256. R. ceylonensis sp. n., male holotype. Figures 248, 257. R. jayarathnei sp. n., male paratype. Figures 249, 258. R. loebli, male from locality 15CG. Figures 250, 259. R. ranawanai sp. n., male holotype. Figure 251. Lychas srilankensis, male from locality 15CO." figureDoi-0="http://doi.org/10.5281/zenodo.7124624" figureDoi-1="http://doi.org/10.5281/zenodo.7124626" httpUri-0="https://zenodo.org/record/7124624/files/figure.png" httpUri-1="https://zenodo.org/record/7124626/files/figure.png" pageId="44" pageNumber="43">239241, 243244, 253</figureCitation>
,
<figureCitation id="A0E32A4CA83BFFBCFE48FA039F8BFA56" box="[392,489,1492,1515]" captionStart="Figures 403429" captionStartId="85.[192,278,744,768]" captionTargetId="figure-11@84.[172,1464,185,1878]" captionTargetPageId="84" captionText="Figures 403429: Telson, lateral view of Sri Lankan Buthidae and Chaerilidae species. Figures 403404. Isometrus maculatus, male from locality 15CP (403) and female from locality 15CI (404). Figures 405406. I. thwaitesi, male from locality 15CH (405) and female from locality 15CO (406). Figures 407408. Lychas srilankensis, male (407) and female (408) from locality 15CJ. Figures 409410. Reddyanus basilicus, male (409) and female (410) from locality 15CR. Figures 411412. R. besucheti, male holotype (411) and female from locality 15CG (412). Figures 413414. R. ceylonensis sp. n., male holotype (413) and female paratype (414) from locality 15CI. Figures 415416. R. jayarathnei sp. n., male (415) and female (416) paratypes. Figures 417418. R. loebli, male (417) and female (418) from locality 15CG. Figures 419420. R. ranawanai sp. n., male holotype (419) and female paratype (420). Figures 421422: Buthoscorpio sarasinorum, male from locality 15CF (421) and female from locality 15CG (422). Figures 423424. Charmus laneus, male (423) and female (424) from locality 15CO. Figures 425426. Charmus saradieli sp. n. male paratype (425) (MHNG) and female holotype (426). Figurs 427. Hottentotta tamulus, male from locality 15CK (427). Figures 428429. Chaerilus ceylonensis, male (428) and female (429) from locality 15CD." figureDoi="http://doi.org/10.5281/zenodo.7124696" httpUri="https://zenodo.org/record/7124696/files/figure.png" pageId="44" pageNumber="43">405406</figureCitation>
, 549, 558560)
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<treatmentCitation id="B97910D8A83BFFBCFF00F9CF9C07F992" author="POCOCK, R. I." box="[192,613,1560,1583]" page="114" pageId="44" pageNumber="43" year="1897">
<taxonomicName id="FFD84D4AA83BFFBCFF00F9CF9C07F992" authority="Pocock, 1897: 114" authorityName="Pocock" authorityPageNumber="114" authorityYear="1897" box="[192,613,1560,1583]" class="Arachnida" family="Buthidae" genus="Isometrus" kingdom="Animalia" order="Scorpiones" pageId="44" pageNumber="43" phylum="Arthropoda" rank="species" species="thwaitesii">
<emphasis id="0AACEADBA83BFFBCFF00F9CF9FFAF992" box="[192,408,1560,1583]" italics="true" pageId="44" pageNumber="43">Isometrus thwaitesii</emphasis>
<bibRefCitation id="5C494B38A83BFFBCFE5EF9CF9C07F992" author="Pocock" box="[414,613,1560,1583]" firstAuthor="Pocock" journalOrPublisher="Journal of the Bombay Natural History Society" page="114" pageId="44" pageNumber="43" pagination="102 - 117" part="11" refId="ref43758" refString="POCOCK, R. I. 1897. Descriptions of some new species of scorpions from India. Journal of the Bombay Natural History Society, 11: 102 - 117." title="Descriptions of some new species of scorpions from India" type="journal article" year="1897">Pocock, 1897: 114</bibRefCitation>
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.
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<treatmentCitation id="B97910D8A83BFFBCFF00F9E09D7FF9F3" author="VACHON, M." box="[192,797,1591,1614]" page="90" pageId="44" pageNumber="43" year="1982">
<taxonomicName id="FFD84D4AA83BFFBCFF00F9E09D7FF9F3" ID-CoL="6MZ76" authority=": Vachon, 1982: 88 - 90" authorityName=": Vachon" authorityPageNumber="88 - 90" authorityYear="1982" box="[192,797,1591,1614]" class="Arachnida" family="Buthidae" genus="Isometrus" kingdom="Animalia" order="Scorpiones" pageId="44" pageNumber="43" phylum="Arthropoda" rank="species" species="thwaitesi" subGenus="Isometrus">
<emphasis id="0AACEADBA83BFFBCFF00F9E09F4BF9F3" box="[192,297,1591,1614]" italics="true" pageId="44" pageNumber="43">Isometrus</emphasis>
(
<emphasis id="0AACEADBA83BFFBCFEFDF9E09FC4F9F3" box="[317,422,1591,1614]" italics="true" pageId="44" pageNumber="43">Isometrus</emphasis>
)
<emphasis id="0AACEADBA83BFFBCFE7AF9E09C78F9F3" box="[442,538,1591,1614]" italics="true" pageId="44" pageNumber="43">thwaitesi</emphasis>
:
<bibRefCitation id="5C494B38A83BFFBCFDECF9E09D7FF9F3" author="Vachon" box="[556,797,1591,1614]" firstAuthor="Vachon" journalOrPublisher="Revue suisse de Zoologie" page="88 - 90" pageId="44" pageNumber="43" pagination="77 - 114" part="89" refId="ref45190" refString="VACHON, M. 1982. Les scorpions de Sri Lanka (Recherches sur les scorpions appartenant ou deposes au Museum d`Histoire naturelle de Geneve III.). Revue suisse de Zoologie, 89: 77 - 114." title="Les scorpions de Sri Lanka (Recherches sur les scorpions appartenant ou deposes au Museum d`Histoire naturelle de Geneve III.)" type="journal article" year="1982">Vachon, 1982: 8890</bibRefCitation>
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</treatmentCitation>
, figs. 3639;
<treatmentCitation id="B97910D8A83BFFBCFE48F9819CC5F9D0" author="FET, V. &amp; W. D. SISSOM &amp; G. LOWE &amp; M. E. BRAUNWALDER" box="[392,679,1622,1645]" page="150" pageId="44" pageNumber="43" year="2000">
<bibRefCitation id="5C494B38A83BFFBCFE48F9819CC5F9D0" author="Fet &amp; Lowe" bookContentInfo="689 pp." box="[392,679,1622,1645]" firstAuthor="Fet" journalOrPublisher="New York: The New York Entomological Society" page="150" pageId="44" pageNumber="43" refId="ref40873" refString="FET, V., W. D. SISSOM, G. LOWE &amp; M. E. BRAUNWALDER. 2000. Catalog of the Scorpions of the World (1758 - 1998). New York: The New York Entomological Society, 689 pp." title="Catalog of the Scorpions of the World (1758 - 1998)" type="book" year="2000">Fet &amp; Lowe, 2000: 150</bibRefCitation>
</treatmentCitation>
(complete reference list until 1998);
<treatmentCitation id="B97910D8A83BFFBCFDC6F9A19CDDF930" author="Kovarik" box="[518,703,1654,1677]" httpUri="http://treatment.plazi.org/id/4703879F451A2706FCA1AB13201585C9" page="4" pageId="44" pageNumber="43" year="2003">
<bibRefCitation id="5C494B38A83BFFBCFDC6F9A19CDDF930" author="Kovarik" box="[518,703,1654,1677]" firstAuthor="Kovarik" page="4" pageId="44" pageNumber="43" pagination="1 - 19" refId="ref41718" refString="KOVARIK, F. 2003. A review of the genus Isometrus Ehrenberg, 1828 (Scorpiones: Buthidae) with descriptions of four new species from Asia and Australia. Euscorpius, 10: 1 - 19." type="journal article" year="2003">Kovařík, 2003: 4</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="B97910D8A83BFFBCFD0AF9A19F97F911" author="KOVARIK, F &amp; A. A. OJANGUREN AFFILASTRO" page="184" pageId="44" pageNumber="43" year="2013">
<bibRefCitation id="5C494B38A83BFFBCFD0AF9A19F97F911" author="Kovarik &amp; Ojanguren" bookContentInfo="400 pp." firstAuthor="Kovarik" journalOrPublisher="Clairon Production, Prague" page="184" pageId="44" pageNumber="43" refId="ref42080" refString="KOVARIK, F &amp; A. A. OJANGUREN AFFILASTRO. 2013. Illustrated catalog of scorpions Part II. Bothriuridae; Chaerilidae; Buthidae I., genera Compsobuthus, Hottentotta, Isometrus, Lychas, and Sassanidotus. Clairon Production, Prague, 400 pp." title="Illustrated catalog of scorpions Part II. Bothriuridae; Chaerilidae; Buthidae I., genera Compsobuthus, Hottentotta, Isometrus, Lychas, and Sassanidotus" type="book" year="2013">Kovařík &amp; Ojanguren, 2013: 184</bibRefCitation>
</treatmentCitation>
, 347, figs. 12601263.
</treatmentCitationGroup>
</paragraph>
<paragraph id="386736C9A83BFFBCFF00F9639F8BF8B7" blockId="44.[192,802,1560,1802]" pageId="44" pageNumber="43">
<treatmentCitationGroup id="18C811E7A83BFFBCFF00F9639F8BF8B7" pageId="44" pageNumber="43">
=
<treatmentCitation id="B97910D8A83BFFBCFF19F9639FDBF957" author="LOURENCO, W. R. &amp; D. HUBER" page="266" pageId="44" pageNumber="43" year="2002">
<taxonomicName id="FFD84D4AA83BFFBCFF19F9639FDBF957" authority="Lourenco &amp; Huber, 2002: 266" authorityName="Lourenco &amp; Huber" authorityPageNumber="266" authorityYear="2002" class="Arachnida" family="Buthidae" genus="Isometrus" kingdom="Animalia" order="Scorpiones" pageId="44" pageNumber="43" phylum="Arthropoda" rank="subSpecies" species="thwaitesi" subGenus="Isometrus" subSpecies="pallidus">
<emphasis id="0AACEADBA83BFFBCFF19F9639F20F976" box="[217,322,1716,1739]" italics="true" pageId="44" pageNumber="43">Isometrus</emphasis>
(
<emphasis id="0AACEADBA83BFFBCFE96F9639FDDF976" box="[342,447,1716,1739]" italics="true" pageId="44" pageNumber="43">Isometrus</emphasis>
)
<emphasis id="0AACEADBA83BFFBCFE12F9639CF0F976" box="[466,658,1716,1739]" italics="true" pageId="44" pageNumber="43">thwaitesi pallidus</emphasis>
<bibRefCitation id="5C494B38A83BFFBCFD5CF9639FDBF957" author="Lourenco &amp; Huber" firstAuthor="Lourenco" journalOrPublisher="Revue suisse de Zoologie" page="266" pageId="44" pageNumber="43" pagination="265 - 275" part="109" refId="ref43004" refString="LOURENCO, W. R. &amp; D. HUBER. 2002. New addition to the scorpion fauna (Arachnida: Scorpiones) of Sri Lanka. Revue suisse de Zoologie, 109 (2): 265 - 275." title="New addition to the scorpion fauna (Arachnida: Scorpiones) of Sri Lanka" type="journal article" year="2002">Lourenço &amp; Huber, 2002: 266</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs. 17 (syn. by
<treatmentCitation id="B97910D8A83BFFBCFD68F9049FBBF8B7" author="KOVARIK, F &amp; A. A. OJANGUREN AFFILASTRO" page="184" pageId="44" pageNumber="43" year="2013">
<bibRefCitation id="5C494B38A83BFFBCFD68F9049FBBF8B7" author="Kovarik &amp; Ojanguren" bookContentInfo="400 pp." firstAuthor="Kovarik" journalOrPublisher="Clairon Production, Prague" page="184" pageId="44" pageNumber="43" refId="ref42080" refString="KOVARIK, F &amp; A. A. OJANGUREN AFFILASTRO. 2013. Illustrated catalog of scorpions Part II. Bothriuridae; Chaerilidae; Buthidae I., genera Compsobuthus, Hottentotta, Isometrus, Lychas, and Sassanidotus. Clairon Production, Prague, 400 pp." title="Illustrated catalog of scorpions Part II. Bothriuridae; Chaerilidae; Buthidae I., genera Compsobuthus, Hottentotta, Isometrus, Lychas, and Sassanidotus" type="book" year="2013">Kovařík &amp; Ojanguren, 2013: 184</bibRefCitation>
</treatmentCitation>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="70C26542A83BFFBDFF00F8E09F74F8DB" lastPageId="45" lastPageNumber="44" pageId="44" pageNumber="43" type="materials_examined">
<paragraph id="386736C9A83BFFBCFF00F8E09F0AF8D1" blockId="44.[192,802,1847,1900]" pageId="44" pageNumber="43">
<typeStatus id="E763886BA83BFFBCFF00F8E09E9AF8F0" box="[192,248,1847,1870]" pageId="44" pageNumber="43">TYPE</typeStatus>
LOCALITY AND
<typeStatus id="E763886BA83BFFBCFE65F8ED9FBBF8F0" box="[421,473,1850,1869]" pageId="44" pageNumber="43">TYPE</typeStatus>
REPOSITORY.
<materialsCitation id="88B03C94A83BFFBCFDB1F8E09F0AF8D1" ID-GBIF-Occurrence="3913992381" collectionCode="BMNH" country="Sri Lanka" location="Sri Lanka" pageId="44" pageNumber="43" specimenCount="1" typeStatus="holotype">
Ceylon, now
<collectingCountry id="40CF7659A83BFFBCFCC3F8E09F64F8D1" name="Sri Lanka" pageId="44" pageNumber="43">Sri Lanka</collectingCountry>
;
<collectionCode id="5EC9AE0CA83BFFBCFED1F8829F06F8D1" box="[273,356,1877,1900]" pageId="44" pageNumber="43">BMNH</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="386736C9A83BFFBDFCFEFB039F74F8DB" blockId="44.[830,1442,1236,1903]" lastBlockId="45.[192,802,1810,1894]" lastPageId="45" lastPageNumber="44" pageId="44" pageNumber="43">
MATERIAL EXAMINED.
<materialsCitation id="88B03C94A83BFFBCFBFDFB039DF2FA95" ID-GBIF-Occurrence="3913992327" collectingDate="1995-11-30" collectionCode="FKCP" collectorName="S. Becvar &amp; Kostal" location="Sri Lanka" municipality="Ratnapura District" pageId="44" pageNumber="43" specimenCount="1" specimenCount-female="1">
<location id="3D076012A83BFFBCFBFDFB039ACBFB56" LSID="urn:lsid:plazi:treatment:B07187DFA83BFFA0FEFAFA419FB4F91F:3D076012A83BFFBCFBFDFB039ACBFB56" box="[1085,1193,1236,1259]" municipality="Ratnapura District" name="Sri Lanka" pageId="44" pageNumber="43">Sri Lanka</location>
,
<collectingMunicipality id="D803ACB3A83BFFBCFB7AFB039BE7FB56" box="[1210,1413,1236,1259]" pageId="44" pageNumber="43">Ratnapura District</collectingMunicipality>
,
<locationDeviation id="29D52FEBA83BFFBCFA53FB039DAAFAB7" location="Hayes" pageId="44" pageNumber="43">
<quantity id="FF209B2CA83BFFBCFA53FB039D02FAB7" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="44" pageNumber="43" unit="km" value="2.0">2 km</quantity>
S Hayes
</locationDeviation>
,
<date id="4C661009A83BFFBCFC16FB249A2DFAB7" box="[982,1103,1267,1290]" pageId="44" pageNumber="43" value="1995-11-30">
<collectingDate id="5C22E9E1A83BFFBCFC16FB249A2DFAB7" box="[982,1103,1267,1290]" pageId="44" pageNumber="43" value="1995-11-30">30.XI.1995</collectingDate>
</date>
,
<specimenCount id="2EDEFD40A83BFFBCFBA0FB249AE0FAB7" box="[1120,1154,1267,1290]" pageId="44" pageNumber="43" type="female">1♀</specimenCount>
,
<collectionCode id="5EC9AE0CA83BFFBCFB52FB249ABAFAB7" box="[1170,1240,1267,1290]" pageId="44" pageNumber="43">FKCP</collectionCode>
, leg.
<collectorName id="952D531FA83BFFBCFAD7FB249BE0FAB7" box="[1303,1410,1267,1290]" pageId="44" pageNumber="43">S. Bečvář</collectorName>
&amp;
<collectorName id="952D531FA83BFFBCFCFEFAC69DE9FA95" box="[830,907,1297,1320]" pageId="44" pageNumber="43">Košťál</collectorName>
;
</materialsCitation>
<materialsCitation id="88B03C94A83BFFBCFC5BFAC69D8FFA1E" ID-GBIF-Occurrence="3913992372" collectionCode="FKCP" collectorName="Kovarik" country="Switzerland" county="Locality" elevation="101" latitude="8.144611" location="North Central Province" longLatPrecision="15" longitude="80.85111" municipality="Kaudulla National Park" pageId="44" pageNumber="43" specimenCount="1" specimenCount-female="1" specimenCount-male="1">
<location id="3D076012A83BFFBCFC5BFAC69AC0FA95" LSID="urn:lsid:plazi:treatment:B07187DFA83BFFA0FEFAFA419FB4F91F:3D076012A83BFFBCFC5BFAC69AC0FA95" box="[923,1186,1297,1320]" county="Locality" latitude="8.144611" longLatPrecision="15" longitude="80.85111" municipality="Kaudulla National Park" name="North Central Province" pageId="44" pageNumber="43">North Central Province</location>
,
<location id="3D076012A83BFFBCFB71FAC69BFEFA95" LSID="urn:lsid:plazi:treatment:B07187DFA83BFFA0FEFAFA419FB4F91F:3D076012A83BFFBCFB71FAC69BFEFA95" box="[1201,1436,1297,1320]" county="Locality" latitude="8.144611" longLatPrecision="15" longitude="80.85111" municipality="Kaudulla National Park" name="Polonnaruwa District" pageId="44" pageNumber="43">Polonnaruwa District</location>
, near
<collectingMunicipality id="D803ACB3A83BFFBCFCBFFAE79AF0FAFA" box="[895,1170,1328,1351]" pageId="44" pageNumber="43">Kaudulla National Park</collectingMunicipality>
,
<geoCoordinate id="5DEC500EA83BFFBCFB6AFAE79B59FAFA" box="[1194,1339,1328,1351]" degrees="08" direction="north" minutes="08" orientation="latitude" pageId="44" pageNumber="43" precision="1" seconds="40. 6" value="8.144611">08°08'40.6&quot;N</geoCoordinate>
<geoCoordinate id="5DEC500EA83BFFBCFA8EFAE79D15FADB" degrees="080" direction="east" minutes="51" orientation="longitude" pageId="44" pageNumber="43" precision="15" seconds="04" value="80.85111">080°51' 04&quot;E</geoCoordinate>
,
<elevation id="B3F5D1FAA83BFFBCFC48FA989A73FADB" box="[904,1041,1359,1382]" metricMagnitude="2" metricUnit="m" metricValue="1.01" pageId="44" pageNumber="43" unit="m" value="101.0">
<quantity id="FF209B2CA83BFFBCFC48FA989DB1FADB" box="[904,979,1359,1382]" metricMagnitude="2" metricUnit="m" metricValue="1.01" pageId="44" pageNumber="43" unit="m" value="101.0">101 m</quantity>
a.s.l.
</elevation>
(
<collectingCounty id="D1064E45A83BFFBCFBE5FA989AE2FADB" box="[1061,1152,1359,1382]" pageId="44" pageNumber="43">Locality</collectingCounty>
<emphasis id="0AACEADBA83BFFBCFB4DFA999AAEFADB" bold="true" box="[1165,1228,1358,1382]" pageId="44" pageNumber="43">15CF</emphasis>
,
<figureCitation id="A0E32A4CA83BFFBCFB20FA989B20FADB" box="[1248,1346,1359,1382]" captionStart="Figures 579581" captionStartId="121.[192,278,1661,1685]" captionTargetBox="[213,1417,167,1633]" captionTargetId="figure-89@121.[214,1416,168,1632]" captionTargetPageId="121" captionText="Figures 579581: Figures 579580. Locality 15CE, Sri Lanka, North Central Province, Polonnaruwa District, Giritale, 08°01'26.0&quot;N 080°54'37.2&quot;E, 233 m a.s.l. During night collecting on 22.23.IV.2015 in the hotel garden (UV detection) we recorded Heterometrus swammerdami (inside the termite constructions, Fig. 580) and Heterometrus gravimanus inside burrows in open terrain. The mothers had 30 to 60 cm long oblique burrows usually once curved from which several narrow burrows created by their juveniles separated under surface. Every burrow of juveniles has a proper exit 15 to 90 cm from the main exit of the motherś burrow. On the locality we recorded night temperature 32.8 ºC 26.5 ºC (minimum temperature) and night humidity varied between 51% and 79%. Figure 581. Locality 15CF, Sri Lanka, North Central Province, Polonnaruwa District, near Kaudulla National Park, 08°08'40.6&quot;N 080°51'04&quot;E, 101 m a.s.l. During a day 23.IV.2015 we recorded Buthoscorpio sarasinorum (under stones or woods), Isometrus thwaitesi (under dry bark of standing trees), Reddyanus besucheti (under stones), and Chaerilus ceylonensis (under stones)." figureDoi="http://doi.org/10.5281/zenodo.7124777" httpUri="https://zenodo.org/record/7124777/files/figure.png" pageId="44" pageNumber="43">Fig. 581</figureCitation>
),
<date id="4C661009A83BFFBCFA9CFA989BFFFADB" box="[1372,1437,1359,1382]" pageId="44" pageNumber="43" value="2015-04-23">
23.
<collectionCode id="5EC9AE0CA83BFFBCFABDFA989BFFFADB" box="[1405,1437,1359,1382]" pageId="44" pageNumber="43">IV</collectionCode>
</date>
. 2015,
<specimenCount id="2EDEFD40A83BFFBCFC48FABA9DCBFA39" box="[904,937,1389,1412]" pageId="44" pageNumber="43" type="male">1♂</specimenCount>
(
<figureCitation id="A0E32A4CA83BFFBCFC7DFABA9A43FA39" box="[957,1057,1389,1412]" captionStart="Figures 150151" captionStartId="39.[192,278,1829,1853]" captionTargetBox="[191,1440,197,1801]" captionTargetId="figure-6@39.[192,1439,197,1800]" captionTargetPageId="39" captionText="Figures 150151: Isometrus thwaitesi from locality 15CF, male (150) and female (151)." figureDoi="http://doi.org/10.5281/zenodo.7124576" httpUri="https://zenodo.org/record/7124576/files/figure.png" pageId="44" pageNumber="43">Fig. 150</figureCitation>
,
<figureCitation id="A0E32A4CA83BFFBCFBF2FABA9A3EFA39" box="[1074,1116,1389,1412]" captionStart="Figures 547554" captionStartId="114.[192,278,810,834]" captionTargetBox="[191,1440,191,782]" captionTargetId="figure-538@114.[192,1440,192,781]" captionTargetPageId="114" captionText="Figures 547554: The mitotic metaphases of buthid males from Sri Lanka. Buthoscorpio sarasinorum from locality 15CF (2n=14) (547), Charmus laneus from locality 15CO (2n=9) (548), Isometrus thwaitesi from locality 15CF (2n=8) (549), Lychas srilankensis from locality 15CN (2n=16) (550), Reddyanus basilicus from locality 15CS (2n=16) (551), Reddyanus basilicus from locality 15CR (2n=15) (552), Reddyanus ceylonensis from locality 15CI (2n=16) (553), Reddyanus loebli from locality 15CH (2n=17) (554). Arrowheads indicate extra large odd chromosomes. Scale bar = 5 µm for 547554." figureDoi="http://doi.org/10.5281/zenodo.7124765" httpUri="https://zenodo.org/record/7124765/files/figure.png" pageId="44" pageNumber="43">549</figureCitation>
)
<specimenCount id="2EDEFD40A83BFFBCFBAFFABA9AF3FA39" box="[1135,1169,1389,1412]" pageId="44" pageNumber="43" type="female">1♀</specimenCount>
(
<figureCitation id="A0E32A4CA83BFFBCFB65FABA9B65FA39" box="[1189,1287,1389,1412]" captionStart="Figures 150151" captionStartId="39.[192,278,1829,1853]" captionTargetBox="[191,1440,197,1801]" captionTargetId="figure-6@39.[192,1439,197,1800]" captionTargetPageId="39" captionText="Figures 150151: Isometrus thwaitesi from locality 15CF, male (150) and female (151)." figureDoi="http://doi.org/10.5281/zenodo.7124576" httpUri="https://zenodo.org/record/7124576/files/figure.png" pageId="44" pageNumber="43">Fig. 151</figureCitation>
),
<collectionCode id="5EC9AE0CA83BFFBCFAE2FABA9B07FA39" box="[1314,1381,1389,1412]" pageId="44" pageNumber="43">FKCP</collectionCode>
, leg.
<collectorName id="952D531FA83BFFBCFCFEFA5B9DF4FA1E" box="[830,918,1420,1443]" pageId="44" pageNumber="43">Kovařík</collectorName>
et al.;
</materialsCitation>
<materialsCitation id="88B03C94A83BFFBCFC3EFA5B9B50F9A0" ID-GBIF-Occurrence="3913992360" collectionCode="FKCP" collectorName="Kovarik" country="Switzerland" county="Locality" elevation="90" latitude="8.807305" location="Northern Province" longLatPrecision="15" longitude="80.17389" municipality="Madhu Road" pageId="44" pageNumber="43" specimenCount="1" specimenCount-male="1">
<location id="3D076012A83BFFBCFC3EFA5B9AACFA1E" LSID="urn:lsid:plazi:treatment:B07187DFA83BFFA0FEFAFA419FB4F91F:3D076012A83BFFBCFC3EFA5B9AACFA1E" box="[1022,1230,1420,1443]" country="Switzerland" county="Locality" latitude="8.807305" longLatPrecision="15" longitude="80.17389" municipality="Madhu Road" name="Northern Province" pageId="44" pageNumber="43">Northern Province</location>
,
<location id="3D076012A83BFFBCFB25FA5B9BFEFA1E" LSID="urn:lsid:plazi:treatment:B07187DFA83BFFA0FEFAFA419FB4F91F:3D076012A83BFFBCFB25FA5B9BFEFA1E" box="[1253,1436,1420,1443]" country="Switzerland" county="Locality" latitude="8.807305" longLatPrecision="15" longitude="80.17389" municipality="Madhu Road" name="Mannar District" pageId="44" pageNumber="43">Mannar District</location>
,
<collectingMunicipality id="D803ACB3A83BFFBCFCFEFA7D9DB2FA7C" box="[830,976,1450,1473]" pageId="44" pageNumber="43">Madhu Road</collectingMunicipality>
,
<geoCoordinate id="5DEC500EA83BFFBCFC25FA7D9A15FA7C" box="[997,1143,1450,1473]" degrees="08" direction="north" minutes="48" orientation="latitude" pageId="44" pageNumber="43" precision="1" seconds="26.3" value="8.807305">08°48'26.3&quot;N</geoCoordinate>
<geoCoordinate id="5DEC500EA83BFFBCFB45FA7D9B72FA7C" box="[1157,1296,1450,1473]" degrees="080" direction="east" minutes="10" orientation="longitude" pageId="44" pageNumber="43" precision="15" seconds="26" value="80.17389">080°10'26&quot;E</geoCoordinate>
,
<elevation id="B3F5D1FAA83BFFBCFAE2FA7D9BC2FA7C" box="[1314,1440,1450,1473]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="44" pageNumber="43" unit="m" value="90.0">
<quantity id="FF209B2CA83BFFBCFAE2FA7D9B02FA7C" box="[1314,1376,1450,1473]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="44" pageNumber="43" unit="m" value="90.0">90 m</quantity>
a.s.l.
</elevation>
(
<collectingCounty id="D1064E45A83BFFBCFC86FA1E9DC3FA5D" box="[838,929,1481,1504]" pageId="44" pageNumber="43">Locality</collectingCounty>
<emphasis id="0AACEADBA83BFFBCFC6BFA1F9D8CFA5D" bold="true" box="[939,1006,1480,1504]" pageId="44" pageNumber="43">
15
<collectingCountry id="40CF7659A83BFFBCFC06FA1F9D8CFA5D" box="[966,1006,1480,1504]" name="Switzerland" pageId="44" pageNumber="43">CH</collectingCountry>
</emphasis>
,
<figureCitation id="A0E32A4CA83BFFBCFBC0FA1E9A3CFA5D" box="[1024,1118,1481,1504]" captionStart="Figures 582584" captionStartId="122.[192,278,1697,1721]" captionTargetBox="[191,1440,167,1669]" captionTargetId="figure-235@122.[192,1439,168,1668]" captionTargetPageId="122" captionText="Figures 582584: Figures 582583. Locality 15CG, Sri Lanka, Central Province, Matale District, Habarana, Wananiwahana Resort, 07°59'25.8&quot;N 080°43'24.6&quot;E, 280 m a.s.l. During night collecting on 23.24.IV.2015 in Wananiwahana resort garden (UV detection) we recorded Buthoscorpio sarasinorum (running on the land among leaves), Reddyanus besucheti (on the land among leaves) and Reddyanus loebli (hidden under scales of barks of standing tree trunks, see the blue arrows on the Fig. 583). On the locality we recorded night temperature 28.4 ºC 24 ºC (minimum temperature) and night humidity varied between 55% and 87%. Figure 584. Locality 15CH, Sri Lanka, Northern Province, Mannar District, Madhu Road, 08°48'26.3&quot;N 080°10'26&quot;E, 90 m a.s.l. During night collecting on 24.25.IV.2015 (UV detection) we recorded Isometrus maculatus and Isometrus thwaitesii sympatrically (running on branches and trunks of trees, also siting on leaves one to four metres high), Reddyanus ceylonensis sp. n. (on the land among leaves), Reddyanus loebli (hidden under scales of barks of standing tree trunks)." figureDoi="http://doi.org/10.5281/zenodo.7124779" httpUri="https://zenodo.org/record/7124779/files/figure.png" pageId="44" pageNumber="43">Fig. 584</figureCitation>
),
<date id="4C661009A83BFFBCFBB9FA1E9A92FA5D" box="[1145,1264,1481,1504]" pageId="44" pageNumber="43" value="2015-04-24" valueMax="2015-04-25" valueMin="2015-04-24">
24. 25.
<collectionCode id="5EC9AE0CA83BFFBCFB14FA1E9A92FA5D" box="[1236,1264,1481,1504]" pageId="44" pageNumber="43">IV</collectionCode>
</date>
.2015,
<specimenCount id="2EDEFD40A83BFFBCFAFEFA1E9B02FA5D" box="[1342,1376,1481,1504]" pageId="44" pageNumber="43" type="male">1♂</specimenCount>
(
<figureCitation id="A0E32A4CA83BFFBCFAB2FA1E9DB4FA42" captionStart="Figures 138141" captionStartId="36.[192,278,1582,1606]" captionTargetBox="[210,1401,172,1544]" captionTargetId="figure-72@36.[192,1439,168,1553]" captionTargetPageId="36" captionText="Figures 138141: Isometrus thwaitesi. Figures 138139. Male from locality 15CH in dorsal (138) and ventral (139) views. Figures 140141. Female from locality 15CO in dorsal (140) and ventral (141) views." figureDoi="http://doi.org/10.5281/zenodo.7124565" httpUri="https://zenodo.org/record/7124565/files/figure.png" pageId="44" pageNumber="43">Figures 138139</figureCitation>
,
<figureCitation id="A0E32A4CA83BFFBCFC24FA3F9A2AFA42" box="[996,1096,1512,1535]" captionStart="Figures 142145" captionStartId="37.[192,278,1844,1868]" captionTargetBox="[213,1417,167,1815]" captionTargetId="figure-15@37.[214,1416,168,1814]" captionTargetPageId="37" captionText="Figures 142145: Isometrus thwaitesi. Figures 142, 144. Male from locality 15CH, chelicerae, carapace and tergites IIII (142) and sternopectinal region and sternite III (144). Figures 143, 145. Female from locality 15CO, chelicerae, carapace and tergites IIII (143) and sternopectinal region and sternite III (145)." figureDoi="http://doi.org/10.5281/zenodo.7124569" httpUri="https://zenodo.org/record/7124569/files/figure.png" pageId="44" pageNumber="43">142, 144</figureCitation>
,
<figureCitation id="A0E32A4CA83BFFBCFB97FA3F9AD5FA42" box="[1111,1207,1512,1535]" captionStart="Figures 146149" captionStartId="38.[192,278,1446,1470]" captionTargetBox="[194,1436,179,1409]" captionTargetId="figure-11@38.[192,1439,168,1417]" captionTargetPageId="38" captionText="Figures 146149: Left hemispermatophore of Isometrus thwaitesi from locality 15CH. External (146), dorsal (147), internal (148) and ventral (149) views. Note: distal part of flagellum and basal part of trunk (foot) were damaged during dissection. Scale bar: 500 µm. Abbreviations: bl, basal lobe; f, flagellum; ml, median lobe; mlc, median lobe carina; pr, pars recta of flagellum; prf, pars reflecta of flagellum; t, trunk." figureDoi="http://doi.org/10.5281/zenodo.7124574" httpUri="https://zenodo.org/record/7124574/files/figure.png" pageId="44" pageNumber="43">146149</figureCitation>
,
<figureCitation id="A0E32A4CA83BFFBCFB05FA3F9A8FFA42" box="[1221,1261,1512,1535]" captionStart="Figures 209224" captionStartId="50.[192,278,1736,1760]" captionTargetBox="[208,1434,174,1703]" captionTargetId="figure-11@50.[205,1452,168,1707]" captionTargetPageId="50" captionText="Figures 209224: Metasomal segments IIIII, lateral view of Sri Lankan Reddyanus and Isometrus species. Figures 209210. Reddyanus basilicus, male (209) and female (210) from locality 15CR. Figures 211212. R. besucheti, male holotype (211) and female from locality 15CG (212). Figures 213214. R. ceylonensis sp. n., male holotype (213) and female paratype from locality 15CI (214). Figures 215216. R. jayarathnei sp. n., male (215) and female (216) paratypes. Figures 217218. R. loebli, male (217) and female (218) from locality 15CG. Figures 219220. R. ranawanai sp. n., male holotype (219) and female paratype (220). Figures 221222. Isometrus maculatus, male from locality 15CP (221) and female from locality 15CI (222). Figures 223 224. I. thwaitesi, male from locality 15CH (223) and female from locality 15CO (224)." figureDoi="http://doi.org/10.5281/zenodo.7124622" httpUri="https://zenodo.org/record/7124622/files/figure.png" pageId="44" pageNumber="43">223</figureCitation>
,
<figureCitation id="A0E32A4CA83BFFBCFB3FFA3F9B48FA42" box="[1279,1322,1512,1535]" captionStart="Figures 225240" captionStartId="51.[192,278,1493,1517]" captionTargetBox="[193,1432,180,1451]" captionTargetId="figure-6@51.[192,1439,179,1464]" captionTargetPageId="51" captionText="Figures 225240: Pedipalp, dorsal view of Sri Lankan Reddyanus and Isometrus species. Figures 225226. Reddyanus basilicus, male (225) and female (226) from locality 15CR. Figures 227228. R. besucheti, male from locality 15CD (227) and female from locality 15CG (228). Figures 229230. R. ceylonensis sp. n., male holotype (229) and female paratype from locality 15CI (230). Figures 231232. R. jayarathnei sp. n., male (231) and female (232) paratypes. Figures 233234. R. loebli, male (233) and female (234) from locality 15CG. Figures 235236. R. ranawanai sp. n., male holotype (235) and female paratype (236). Figures 237238. Isometrus maculatus, male from locality 15CP (237) and female from locality 15CI (238). Figures 239240. I. thwaitesi, male from locality 15CH (239) and female from locality 15CO (240)." figureDoi="http://doi.org/10.5281/zenodo.7124624" httpUri="https://zenodo.org/record/7124624/files/figure.png" pageId="44" pageNumber="43">239</figureCitation>
,
<figureCitation id="A0E32A4CA83BFFBCFAF8FA3F9B02FA42" box="[1336,1376,1512,1535]" captionStart="Figures 241259" captionStartId="52.[192,278,1733,1757]" captionTargetBox="[238,1387,153,1699]" captionTargetId="figure-11@52.[234,1392,152,1704]" captionTargetPageId="52" captionText="Figures 241259: Pedipalp chela dorsal (241), movable finger (242250), and fixed finger (251259) of Sri Lankan Isometrus, Lychas, and Reddyanus species. Figures 241242, 252. Isometrus maculatus, female from locality 15CO (241) and male from locality 15CP (242, 252). Figures 243244, 253. I. thwaitesi, male from locality 15CH (243) and female from locality 15CO (244, 253). Figures 245, 254. Reddyanus basilicus, male from locality 15CS. Figures 246, 255. R. besucheti, male from locality 15CD. Figures 247, 256. R. ceylonensis sp. n., male holotype. Figures 248, 257. R. jayarathnei sp. n., male paratype. Figures 249, 258. R. loebli, male from locality 15CG. Figures 250, 259. R. ranawanai sp. n., male holotype. Figure 251. Lychas srilankensis, male from locality 15CO." figureDoi="http://doi.org/10.5281/zenodo.7124626" httpUri="https://zenodo.org/record/7124626/files/figure.png" pageId="44" pageNumber="43">243</figureCitation>
,
<figureCitation id="A0E32A4CA83BFFBCFAB1FA3F9BFBFA42" box="[1393,1433,1512,1535]" captionStart="Figures 403429" captionStartId="85.[192,278,744,768]" captionTargetId="figure-11@84.[172,1464,185,1878]" captionTargetPageId="84" captionText="Figures 403429: Telson, lateral view of Sri Lankan Buthidae and Chaerilidae species. Figures 403404. Isometrus maculatus, male from locality 15CP (403) and female from locality 15CI (404). Figures 405406. I. thwaitesi, male from locality 15CH (405) and female from locality 15CO (406). Figures 407408. Lychas srilankensis, male (407) and female (408) from locality 15CJ. Figures 409410. Reddyanus basilicus, male (409) and female (410) from locality 15CR. Figures 411412. R. besucheti, male holotype (411) and female from locality 15CG (412). Figures 413414. R. ceylonensis sp. n., male holotype (413) and female paratype (414) from locality 15CI. Figures 415416. R. jayarathnei sp. n., male (415) and female (416) paratypes. Figures 417418. R. loebli, male (417) and female (418) from locality 15CG. Figures 419420. R. ranawanai sp. n., male holotype (419) and female paratype (420). Figures 421422: Buthoscorpio sarasinorum, male from locality 15CF (421) and female from locality 15CG (422). Figures 423424. Charmus laneus, male (423) and female (424) from locality 15CO. Figures 425426. Charmus saradieli sp. n. male paratype (425) (MHNG) and female holotype (426). Figurs 427. Hottentotta tamulus, male from locality 15CK (427). Figures 428429. Chaerilus ceylonensis, male (428) and female (429) from locality 15CD." figureDoi="http://doi.org/10.5281/zenodo.7124696" httpUri="https://zenodo.org/record/7124696/files/figure.png" pageId="44" pageNumber="43">405</figureCitation>
, 558560)
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et al.;
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,
<location id="3D076012A83BFFBCFC6BF9F29A3AF981" LSID="urn:lsid:plazi:treatment:B07187DFA83BFFA0FEFAFA419FB4F91F:3D076012A83BFFBCFC6BF9F29A3AF981" box="[939,1112,1573,1596]" country="Switzerland" county="Locality" latitude="8.558972" longLatPrecision="15" longitude="79.9475" municipality="Wilpattu National Park" name="Mannar District" pageId="44" pageNumber="43">Mannar District</location>
,
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env, border of
<collectingMunicipality id="D803ACB3A83BFFBCFC9DF9939A37F9E6" box="[861,1109,1604,1627]" pageId="44" pageNumber="43">Wilpattu National Park</collectingMunicipality>
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<quantity id="FF209B2CA83BFFBCFA53F9939D36F9C4" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="44" pageNumber="43" unit="m" value="7.0">7 m</quantity>
a.s.l.
</elevation>
(
<collectingCounty id="D1064E45A83BFFBCFC5CF9B59D95F9C4" box="[924,1015,1634,1657]" pageId="44" pageNumber="43">Locality</collectingCounty>
<emphasis id="0AACEADBA83BFFBCFBC1F9B59A5AF9C7" bold="true" box="[1025,1080,1634,1658]" pageId="44" pageNumber="43">15CI</emphasis>
,
<figureCitation id="A0E32A4CA83BFFBCFB86F9B59AC0F9C4" box="[1094,1186,1634,1657]" captionStart="Figures 585587" captionStartId="123.[192,278,1541,1565]" captionTargetBox="[233,1393,189,1513]" captionTargetId="figure-293@123.[234,1392,190,1512]" captionTargetPageId="123" captionText="Figures 585587: Figure 585. Locality 15CI, Sri Lanka, Northern Province, Mannar District, Marichchukkaddi env, border of Wilpattu National Park, 08°33'32.3&quot;N 079°56'51&quot;E, 7 m a.s.l. During night collecting on 25.26.IV.2015 (UV detection) we recorded Isometrus maculatus and Isometrus thwaitesii sympatrically (running on branches and trunks of trees, also sitting on leaves one to four metres high), Reddyanus ceylonensis sp. n. (on the land among leaves; the type locality), Reddyanus loebli (hidden under scales of bark of standing tree trunks). Near to the locality we recorded temperature between 37.6 ºC (maximum daytime temperature and 26.2 ºC (minimum nighttime temperature) shortly before sunrise. Humidity varied between 67% and 81%. Figures 586587. Locality 15CJ, Sri Lanka, Northern Province, Jaffna District, 09°42'51.6&quot;N 080°04'44.8&quot;E, 19 m a.s.l. During night collecting on 26.27.IV.2015 (UV detection) we recorded Lychas srilankensis (on the land usually under branches, leaves or in their vicinity), Heterometrus swammerdami (inside the termite constructions or inside heaps of garden &quot;waste&quot;) and Heterometrus gravimanus inside burrows in open terrain (Fig. 587). The females had 30 to 60 cm long oblique burrows once or twice curved. A female shortly after maturity ecdysis was in the burrow together with a male; another female was in the burrow together with three juveniles which had their own narrow burrows separated under surface from the motherś burrow; a juvenile after the third ecdysis had a proper burrow 30 cm long oblique and once curved. There also lives Isometrus maculatus. On the locality we recorded night temperature 31.6 ºC 27.2 ºC (minimum temperature) and night humidity varied between 60% and 87%." figureDoi="http://doi.org/10.5281/zenodo.7124781" httpUri="https://zenodo.org/record/7124781/files/figure.png" pageId="44" pageNumber="43">Fig. 585</figureCitation>
),
<date id="4C661009A83BFFBCFB79F9B59B4FF9C4" box="[1209,1325,1634,1657]" pageId="44" pageNumber="43" value="2015-04-25" valueMax="2015-04-26" valueMin="2015-04-25">
25. 26.
<collectionCode id="5EC9AE0CA83BFFBCFACEF9B59B4FF9C4" box="[1294,1325,1634,1657]" pageId="44" pageNumber="43">IV</collectionCode>
</date>
.2015,
<specimenCount id="2EDEFD40A83BFFBCFAB8F9B59BF8F9C4" box="[1400,1434,1634,1657]" pageId="44" pageNumber="43" type="female">1♀</specimenCount>
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<collectionCode id="5EC9AE0CA83BFFBCFCFEF9569DE6F925" box="[830,900,1665,1688]" pageId="44" pageNumber="43">FKCP</collectionCode>
, leg.
<collectorName id="952D531FA83BFFBCFC10F9569A4AF925" box="[976,1064,1665,1688]" pageId="44" pageNumber="43">Kovařík</collectorName>
et al.;
</materialsCitation>
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<location id="3D076012A83BFFBCFB4DF9569BFFF925" LSID="urn:lsid:plazi:treatment:B07187DFA83BFFA0FEFAFA419FB4F91F:3D076012A83BFFBCFB4DF9569BFFF925" box="[1165,1437,1665,1688]" country="Switzerland" county="Locality" latitude="8.347722" longLatPrecision="1" longitude="80.5077" municipality="Mihintale" name="North Central Province" pageId="44" pageNumber="43">North Central Province</location>
,
<location id="3D076012A83BFFBCFCFEF9779A4DF90A" LSID="urn:lsid:plazi:treatment:B07187DFA83BFFA0FEFAFA419FB4F91F:3D076012A83BFFBCFCFEF9779A4DF90A" box="[830,1071,1696,1719]" country="Switzerland" county="Locality" latitude="8.347722" longLatPrecision="1" longitude="80.5077" municipality="Mihintale" name="Anuradhapura District" pageId="44" pageNumber="43">Anuradhapura District</location>
,
<collectingMunicipality id="D803ACB3A83BFFBCFBFFF9779ACAF90A" box="[1087,1192,1696,1719]" pageId="44" pageNumber="43">Mihintale</collectingMunicipality>
,
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<geoCoordinate id="5DEC500EA83BFFBCFA8EF9779DEAF968" degrees="080" direction="east" minutes="30" orientation="longitude" pageId="44" pageNumber="43" precision="1" seconds="27.7" value="80.5077">080°30' 27.7&quot;E</geoCoordinate>
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a.s.l.
</elevation>
(
<collectingCounty id="D1064E45A83BFFBCFBF4F9699AEDF968" box="[1076,1167,1726,1749]" pageId="44" pageNumber="43">Locality</collectingCounty>
<emphasis id="0AACEADBA83BFFBCFB5AF9699AB8F96B" bold="true" box="[1178,1242,1726,1750]" pageId="44" pageNumber="43">15CL</emphasis>
,
<figureCitation id="A0E32A4CA83BFFBCFB2CF9699B28F968" box="[1260,1354,1726,1749]" captionStart="Figures 588589" captionStartId="124.[192,278,1745,1769]" captionTargetBox="[191,1440,191,1716]" captionTargetId="figure-185@124.[192,1439,192,1715]" captionTargetPageId="124" captionText="Figures 588589: Figure 588. Locality 15CK, Sri Lanka, Northern Province, Jaffna District, 09°49'15.4&quot;N 080°08'41.6&quot;E, 19 m a.s.l. During day 27.IV.2015 we recorded Hottentotta tamulus under stones or woods and also in vicinity of houses. Figure 589. Locality 15CL, Sri Lanka, North Central Province, Anuradhapura District, Mihintale, 08°20'51.8&quot;N 080°30'27.7&quot;E, 156 m a.s.l. During night collecting on 27.28.IV.2015 (UV detection) we recorded Isometrus thwaitesii (running on branches and trunks of trees, also sitting on leaves), Reddyanus besucheti (on the ground among leaves), Reddyanus loebli (hidden under scales of bark of standing tree trunks) and Heterometrus swammerdami (inside burrows). On the locality we recorded night temperature 27.4 ºC 24.5 ºC (minimum temperature) and night humidity varied between 64% and 87%." figureDoi="http://doi.org/10.5281/zenodo.7124783" httpUri="https://zenodo.org/record/7124783/files/figure.png" pageId="44" pageNumber="43">Fig. 589</figureCitation>
),
<date id="4C661009A83BFFBCFAA6F9699D1DF949" pageId="44" pageNumber="43" value="2015-04-27" valueMax="2015-04-28" valueMin="2015-04-27">
27. 28.
<collectionCode id="5EC9AE0CA83BFFBCFC9FF90A9D1DF949" box="[863,895,1757,1780]" pageId="44" pageNumber="43">IV</collectionCode>
</date>
.2015,
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,
<collectionCode id="5EC9AE0CA83BFFBCFC39F90A9A5DF949" box="[1017,1087,1757,1780]" pageId="44" pageNumber="43">FKCP</collectionCode>
, leg.
<collectorName id="952D531FA83BFFBCFBBDF90A9AB6F949" box="[1149,1236,1757,1780]" pageId="44" pageNumber="43">Kovařík</collectorName>
et al.;
</materialsCitation>
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<location id="3D076012A83BFFBCFAE5F90A9DB3F8AE" LSID="urn:lsid:plazi:treatment:B07187DFA83BFFA0FEFAFA419FB4F91F:3D076012A83BFFBCFAE5F90A9DB3F8AE" country="Switzerland" county="Locality" latitude="8.2875" longLatPrecision="15" longitude="79.844086" municipality="Eluwankulam" name="North Central Province" pageId="44" pageNumber="43">North Central Province</location>
,
<location id="3D076012A83BFFBCFC22F92B9AC3F8AE" LSID="urn:lsid:plazi:treatment:B07187DFA83BFFA0FEFAFA419FB4F91F:3D076012A83BFFBCFC22F92B9AC3F8AE" box="[994,1185,1788,1811]" country="Switzerland" county="Locality" latitude="8.2875" longLatPrecision="15" longitude="79.844086" municipality="Eluwankulam" name="Puttalam District" pageId="44" pageNumber="43">Puttalam District</location>
,
<collectingMunicipality id="D803ACB3A83BFFBCFB73F92B9B25F8AE" box="[1203,1351,1788,1811]" pageId="44" pageNumber="43">Eluwankulam</collectingMunicipality>
,
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<geoCoordinate id="5DEC500EA83BFFBCFC43F8CD9A43F88C" box="[899,1057,1818,1841]" degrees="079" direction="east" minutes="50" orientation="longitude" pageId="44" pageNumber="43" precision="1" seconds="38.7" value="79.844086">079°50'38.7&quot;E</geoCoordinate>
,
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<quantity id="FF209B2CA83BFFBCFBF1F8CD9A0EF88C" box="[1073,1132,1818,1841]" metricMagnitude="1" metricUnit="m" metricValue="3.8" pageId="44" pageNumber="43" unit="m" value="38.0">38 m</quantity>
a.s.l.
</elevation>
(
<collectingCounty id="D1064E45A83BFFBCFB7BF8CD9B74F88C" box="[1211,1302,1818,1841]" pageId="44" pageNumber="43">Locality</collectingCounty>
<emphasis id="0AACEADBA83BFFBCFAE1F8CD9B06F88F" bold="true" box="[1313,1380,1818,1842]" pageId="44" pageNumber="43">15CO</emphasis>
,
<figureCitation id="A0E32A4CA83BFFBCFAB6F8CD9D0AF8ED" captionStart="Figures 592594" captionStartId="126.[192,278,1617,1641]" captionTargetBox="[213,1417,167,1588]" captionTargetId="figure-251@126.[214,1416,168,1587]" captionTargetPageId="126" captionText="Figures 592594: Figure 592. Locality 15CO, Sri Lanka, North Central Province, Puttalam District, Eluwankulam, 08°17'15&quot;N 079°50'38.7&quot;E, 38 m a.s.l. During night collecting from 10.30 p.m. on 28.IV.2015 to 4.00 a.m. on 29.IV.2015 (UV detection) we recorded Charmus laneus (running on the ground among leaves), Isometrus thwaitesii (running on branches of bushes, also sitting on leaves), Lychas srilankensis (on the ground under branches, leaves or in their vicinity), Reddyanus ceylonensis sp. n. (on the ground), Chaerilus ceylonensis (inside oblique usually not curved burrows 15 to 30 cm long) and Heterometrus swammerdami (inside long burrows). On the locality we recorded between 10.00 p.m. and 4.00 a.m. temperature 25.9 ºC and humidity between 58% and 70%. On the locality we recorded 6 scorpion species and genera of all three Sri Lankan families. Figures 593594. Locality 15CP, Sri Lanka, Southern Province, Matara District, Kekanadura village, 05°58'28.2&quot;N 080°36'20.5&quot;E, 40 m a.s.l. During night collecting on 30.IV.2015 (UV detection) we recorded Isometrus maculatus (inside a house, the species occupy the roof of the house, Fig. 593) and Heterometrus serratus (inside almost straight burrows in a ground wall). On the locality we recorded night temperature 28.4 ºC 25.2 ºC (minimum temperature) and night humidity varied between 63% and 89%." figureDoi="http://doi.org/10.5281/zenodo.7124791" httpUri="https://zenodo.org/record/7124791/files/figure.png" pageId="44" pageNumber="43">Fig. 592</figureCitation>
),
<date id="4C661009A83BFFBCFC43F8EE9DA2F8ED" box="[899,960,1849,1872]" pageId="44" pageNumber="43" value="2015-04-28">
28.
<collectionCode id="5EC9AE0CA83BFFBCFC61F8EE9DA2F8ED" box="[929,960,1849,1872]" pageId="44" pageNumber="43">IV</collectionCode>
</date>
.2015,
<specimenCount id="2EDEFD40A83BFFBCFBD0F8EE9A53F8ED" box="[1040,1073,1849,1872]" pageId="44" pageNumber="43" type="male">1♂</specimenCount>
<specimenCount id="2EDEFD40A83BFFBCFBF1F8EE9A31F8ED" box="[1073,1107,1849,1872]" pageId="44" pageNumber="43" type="female">1♀</specimenCount>
(
<figureCitation id="A0E32A4CA83BFFBCFBA8F8EE9B40F8ED" box="[1128,1314,1849,1872]" captionStart="Figures 138141" captionStartId="36.[192,278,1582,1606]" captionTargetBox="[210,1401,172,1544]" captionTargetId="figure-72@36.[192,1439,168,1553]" captionTargetPageId="36" captionText="Figures 138141: Isometrus thwaitesi. Figures 138139. Male from locality 15CH in dorsal (138) and ventral (139) views. Figures 140141. Female from locality 15CO in dorsal (140) and ventral (141) views." figureDoi="http://doi.org/10.5281/zenodo.7124565" httpUri="https://zenodo.org/record/7124565/files/figure.png" pageId="44" pageNumber="43">Figures 140141</figureCitation>
,
<figureCitation id="A0E32A4CA83BFFBCFAF6F8EE9BFBF8ED" box="[1334,1433,1849,1872]" captionStart="Figures 142145" captionStartId="37.[192,278,1844,1868]" captionTargetBox="[213,1417,167,1815]" captionTargetId="figure-15@37.[214,1416,168,1814]" captionTargetPageId="37" captionText="Figures 142145: Isometrus thwaitesi. Figures 142, 144. Male from locality 15CH, chelicerae, carapace and tergites IIII (142) and sternopectinal region and sternite III (144). Figures 143, 145. Female from locality 15CO, chelicerae, carapace and tergites IIII (143) and sternopectinal region and sternite III (145)." figureDoi="http://doi.org/10.5281/zenodo.7124569" httpUri="https://zenodo.org/record/7124569/files/figure.png" pageId="44" pageNumber="43">143, 145</figureCitation>
,
<figureCitation id="A0E32A4CA83BFFBCFCFEF88F9D04F8D2" box="[830,870,1880,1903]" captionStart="Figures 193200" captionStartId="48.[192,278,1070,1094]" captionTargetBox="[213,1446,171,1037]" captionTargetId="figure-11@48.[205,1452,168,1041]" captionTargetPageId="48" captionText="Figures 193200: Distal segments of leg IV, retroventral view of Sri Lankan Buthidae and Chaerilidae genera. Buthoscorpio sarasinorum, male from locality 15CF (193), Charmus laneus, male from locality 15CO (194), Hottentotta tamulus, male from locality 15CK (195), Isometrus maculatus, female from locality 15CI (196), Isometrus thwaitesi, female from locality 15CO (197), Lychas srilankensis, male from locality 15CJ (198), Reddyanus basilicus, male from locality 15CS (199), and Chaerilus ceylonensis, male from locality 15CD (200)." figureDoi="http://doi.org/10.5281/zenodo.7124608" httpUri="https://zenodo.org/record/7124608/files/figure.png" pageId="44" pageNumber="43">197</figureCitation>
,
<figureCitation id="A0E32A4CA83BFFBCFCBAF88F9DC0F8D2" box="[890,930,1880,1903]" captionStart="Figures 209224" captionStartId="50.[192,278,1736,1760]" captionTargetBox="[208,1434,174,1703]" captionTargetId="figure-11@50.[205,1452,168,1707]" captionTargetPageId="50" captionText="Figures 209224: Metasomal segments IIIII, lateral view of Sri Lankan Reddyanus and Isometrus species. Figures 209210. Reddyanus basilicus, male (209) and female (210) from locality 15CR. Figures 211212. R. besucheti, male holotype (211) and female from locality 15CG (212). Figures 213214. R. ceylonensis sp. n., male holotype (213) and female paratype from locality 15CI (214). Figures 215216. R. jayarathnei sp. n., male (215) and female (216) paratypes. Figures 217218. R. loebli, male (217) and female (218) from locality 15CG. Figures 219220. R. ranawanai sp. n., male holotype (219) and female paratype (220). Figures 221222. Isometrus maculatus, male from locality 15CP (221) and female from locality 15CI (222). Figures 223 224. I. thwaitesi, male from locality 15CH (223) and female from locality 15CO (224)." figureDoi="http://doi.org/10.5281/zenodo.7124622" httpUri="https://zenodo.org/record/7124622/files/figure.png" pageId="44" pageNumber="43">224</figureCitation>
,
<figureCitation id="A0E32A4CA83BFFBCFC76F88F9AEEF8D2" box="[950,1164,1880,1903]" captionStart-0="Figures 225240" captionStart-1="Figures 241259" captionStartId-0="51.[192,278,1493,1517]" captionStartId-1="52.[192,278,1733,1757]" captionTargetBox-0="[193,1432,180,1451]" captionTargetBox-1="[238,1387,153,1699]" captionTargetId-0="figure-6@51.[192,1439,179,1464]" captionTargetId-1="figure-11@52.[234,1392,152,1704]" captionTargetPageId-0="51" captionTargetPageId-1="52" captionText-0="Figures 225240: Pedipalp, dorsal view of Sri Lankan Reddyanus and Isometrus species. Figures 225226. Reddyanus basilicus, male (225) and female (226) from locality 15CR. Figures 227228. R. besucheti, male from locality 15CD (227) and female from locality 15CG (228). Figures 229230. R. ceylonensis sp. n., male holotype (229) and female paratype from locality 15CI (230). Figures 231232. R. jayarathnei sp. n., male (231) and female (232) paratypes. Figures 233234. R. loebli, male (233) and female (234) from locality 15CG. Figures 235236. R. ranawanai sp. n., male holotype (235) and female paratype (236). Figures 237238. Isometrus maculatus, male from locality 15CP (237) and female from locality 15CI (238). Figures 239240. I. thwaitesi, male from locality 15CH (239) and female from locality 15CO (240)." captionText-1="Figures 241259: Pedipalp chela dorsal (241), movable finger (242250), and fixed finger (251259) of Sri Lankan Isometrus, Lychas, and Reddyanus species. Figures 241242, 252. Isometrus maculatus, female from locality 15CO (241) and male from locality 15CP (242, 252). Figures 243244, 253. I. thwaitesi, male from locality 15CH (243) and female from locality 15CO (244, 253). Figures 245, 254. Reddyanus basilicus, male from locality 15CS. Figures 246, 255. R. besucheti, male from locality 15CD. Figures 247, 256. R. ceylonensis sp. n., male holotype. Figures 248, 257. R. jayarathnei sp. n., male paratype. Figures 249, 258. R. loebli, male from locality 15CG. Figures 250, 259. R. ranawanai sp. n., male holotype. Figure 251. Lychas srilankensis, male from locality 15CO." figureDoi-0="http://doi.org/10.5281/zenodo.7124624" figureDoi-1="http://doi.org/10.5281/zenodo.7124626" httpUri-0="https://zenodo.org/record/7124624/files/figure.png" httpUri-1="https://zenodo.org/record/7124626/files/figure.png" pageId="44" pageNumber="43">240241, 244, 253</figureCitation>
,
<figureCitation id="A0E32A4CA83BFFBCFB59F88F9AA1F8D2" box="[1177,1219,1880,1903]" captionStart="Figures 403429" captionStartId="85.[192,278,744,768]" captionTargetId="figure-11@84.[172,1464,185,1878]" captionTargetPageId="84" captionText="Figures 403429: Telson, lateral view of Sri Lankan Buthidae and Chaerilidae species. Figures 403404. Isometrus maculatus, male from locality 15CP (403) and female from locality 15CI (404). Figures 405406. I. thwaitesi, male from locality 15CH (405) and female from locality 15CO (406). Figures 407408. Lychas srilankensis, male (407) and female (408) from locality 15CJ. Figures 409410. Reddyanus basilicus, male (409) and female (410) from locality 15CR. Figures 411412. R. besucheti, male holotype (411) and female from locality 15CG (412). Figures 413414. R. ceylonensis sp. n., male holotype (413) and female paratype (414) from locality 15CI. Figures 415416. R. jayarathnei sp. n., male (415) and female (416) paratypes. Figures 417418. R. loebli, male (417) and female (418) from locality 15CG. Figures 419420. R. ranawanai sp. n., male holotype (419) and female paratype (420). Figures 421422: Buthoscorpio sarasinorum, male from locality 15CF (421) and female from locality 15CG (422). Figures 423424. Charmus laneus, male (423) and female (424) from locality 15CO. Figures 425426. Charmus saradieli sp. n. male paratype (425) (MHNG) and female holotype (426). Figurs 427. Hottentotta tamulus, male from locality 15CK (427). Figures 428429. Chaerilus ceylonensis, male (428) and female (429) from locality 15CD." figureDoi="http://doi.org/10.5281/zenodo.7124696" httpUri="https://zenodo.org/record/7124696/files/figure.png" pageId="44" pageNumber="43">406</figureCitation>
)
<specimenCount id="2EDEFD40A83BFFBCFB17F88F9B6FF8D2" box="[1239,1293,1880,1903]" pageId="44" pageNumber="43" type="generic">1juv.</specimenCount>
,
<collectionCode id="5EC9AE0CA83BFFBCFAE1F88F9B05F8D2" box="[1313,1383,1880,1903]" pageId="44" pageNumber="43">FKCP</collectionCode>
, leg.
<collectorName id="952D531FA83AFFBDFF00F8C59F7AF894" box="[192,280,1810,1833]" pageId="45" pageNumber="44">Kovařík</collectorName>
et al.;
</materialsCitation>
<materialsCitation id="88B03C94A83AFFBDFE50F8C59F74F8DB" ID-GBIF-Occurrence="3913992359" collectionCode="UPSL" collectorName="Kovarik" country="Switzerland" county="Locality" elevation="156" latitude="8.347722" location="North Central Province" longLatPrecision="1" longitude="80.5077" municipality="Mihintale" pageId="45" pageNumber="44" specimenCount="1" specimenCount-female="1" specimenCount-male="1">
<location id="3D076012A83AFFBDFE50F8C59CCEF894" LSID="urn:lsid:plazi:treatment:B07187DFA83BFFA0FEFAFA419FB4F91F:3D076012A83AFFBDFE50F8C59CCEF894" box="[400,684,1810,1833]" country="Switzerland" latitude="7.973778" longLatPrecision="1" longitude="80.749916" municipality="Sigiriya" name="North Central Province" pageId="45" pageNumber="44">North Central Province</location>
,
<collectingMunicipality id="D803ACB3A83AFFBDFD06F8C59D78F894" box="[710,794,1810,1833]" pageId="45" pageNumber="44">Sigiriya</collectingMunicipality>
,
<geoCoordinate id="5DEC500EA83AFFBDFF00F8E69F26F8F5" box="[192,324,1841,1864]" degrees="7" direction="north" minutes="58" orientation="latitude" pageId="45" pageNumber="44" precision="1" seconds="25. 6" value="7.973778">7°58'25.6&quot;N</geoCoordinate>
<geoCoordinate id="5DEC500EA83AFFBDFE97F8E69F87F8F5" box="[343,485,1841,1864]" degrees="80" direction="east" minutes="44" orientation="longitude" pageId="45" pageNumber="44" precision="1" seconds="59.7" value="80.749916">80°44'59.7&quot;E</geoCoordinate>
,
<specimenCount id="2EDEFD40A83AFFBDFE3FF8E69C42F8F5" box="[511,544,1841,1864]" pageId="45" pageNumber="44" type="female">1♀</specimenCount>
,
<collectionCode id="5EC9AE0CA83AFFBDFDF9F8E69C18F8F5" box="[569,634,1841,1864]" pageId="45" pageNumber="44">UPSL</collectionCode>
, leg.
<collectorName id="952D531FA83AFFBDFD0DF8E69F74F8DB" pageId="45" pageNumber="44">S. Jayarathne.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="6CA76641A83AFFBDFF00F9AC9BE6F960" ID-DOI="http://doi.org/10.5281/zenodo.7124594" ID-Zenodo-Dep="7124594" httpUri="https://zenodo.org/record/7124594/files/figure.png" pageId="45" pageNumber="44" startId="45.[192,278,1659,1683]" targetBox="[196,1422,175,1626]" targetPageId="45">
<paragraph id="386736C9A83AFFBDFF00F9AC9BE6F960" blockId="45.[192,1440,1659,1757]" pageId="45" pageNumber="44">
<emphasis id="0AACEADBA83AFFBDFF00F9AC9FEBF92E" bold="true" box="[192,393,1659,1683]" pageId="45" pageNumber="44">Figures 183186:</emphasis>
Right hemispermatophore of
<taxonomicName id="FFD84D4AA83AFFBDFD01F9A99DE6F92E" authorityName="Lourenco" authorityYear="1997" box="[705,900,1662,1683]" class="Arachnida" family="Buthidae" genus="Lychas" kingdom="Animalia" order="Scorpiones" pageId="45" pageNumber="44" phylum="Arthropoda" rank="species" species="srilankensis">
<emphasis id="0AACEADBA83AFFBDFD01F9A99DE6F92E" box="[705,900,1662,1683]" italics="true" pageId="45" pageNumber="44">Lychas srilankensis</emphasis>
</taxonomicName>
from locality 15CN (presented as mirror image for comparison to other species). External (183), dorsal (184), internal (185) and ventral (186) views. Extracted after 3rd ecdysis. Hemispermatophores examined from a second male after 4th ecdysis were similar. Scale bar: 500 µm. Abbreviations:
<emphasis id="0AACEADBA83AFFBDFA90F9789B03F979" box="[1360,1377,1711,1732]" italics="true" pageId="45" pageNumber="44">bl</emphasis>
, basal lobe;
<emphasis id="0AACEADBA83AFFBDFF36F91F9E9FF960" box="[246,253,1736,1757]" italics="true" pageId="45" pageNumber="44">f</emphasis>
, flagellum;
<emphasis id="0AACEADBA83AFFBDFEB1F91F9FEBF960" box="[369,393,1736,1757]" italics="true" pageId="45" pageNumber="44">ml</emphasis>
, median lobe;
<emphasis id="0AACEADBA83AFFBDFDD8F91F9C58F960" box="[536,570,1736,1757]" italics="true" pageId="45" pageNumber="44">mlc</emphasis>
, median lobe carina;
<emphasis id="0AACEADBA83AFFBDFCCAF91F9D7DF960" box="[778,799,1736,1757]" italics="true" pageId="45" pageNumber="44">pr</emphasis>
, pars recta of flagellum;
<emphasis id="0AACEADBA83AFFBDFBD1F91F9A51F960" box="[1041,1075,1736,1757]" italics="true" pageId="45" pageNumber="44">prf,</emphasis>
pars reflecta of flagellum;
<emphasis id="0AACEADBA83AFFBDFAF7F91F9B5CF960" box="[1335,1342,1736,1757]" italics="true" pageId="45" pageNumber="44">t</emphasis>
, trunk.
</paragraph>
</caption>
<subSubSection id="70C26542A83AFFA0FCFEF8C59CD8F9D8" lastPageId="48" lastPageNumber="47" pageId="45" pageNumber="44" type="diagnosis">
<paragraph id="386736C9A83AFFA0FCFEF8C59C37FAD2" blockId="45.[830,1440,1810,1894]" lastBlockId="48.[192,802,1246,1391]" lastPageId="48" lastPageNumber="47" pageId="45" pageNumber="44">
DIAGNOSIS. Total length
<quantity id="FF209B2CA83AFFBDFB90F8C59AA7F894" box="[1104,1221,1810,1833]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="45" pageNumber="44" unit="mm" value="40.0" valueMax="50.0" valueMin="30.0">3050 mm</quantity>
. Manus of pedipalp very thin, width in male equal to that that of patella and femur. Pedipalps and legs yellow, with spots. Fingers and manus of pedipalps of the same color, spotted. Mesosomal segments light-colored. First (basal) middle lamella of both sexes pecten rounded. Posterior margin of sternite V strongly convex medially. Subaculear tooth spinoid. Pectinal teeth number 1316.
</paragraph>
<caption id="6CA76641A839FFBEFF00F8E39E9BF8C0" ID-DOI="http://doi.org/10.5281/zenodo.7124596" ID-Zenodo-Dep="7124596" httpUri="https://zenodo.org/record/7124596/files/figure.png" pageId="46" pageNumber="45" startId="46.[192,278,1844,1868]" targetBox="[191,1440,167,1815]" targetPageId="46">
<paragraph id="386736C9A839FFBEFF00F8E39E9BF8C0" blockId="46.[192,1440,1844,1917]" pageId="46" pageNumber="45">
<emphasis id="0AACEADBA839FFBEFF00F8E39FE7F8F1" bold="true" box="[192,389,1844,1868]" pageId="46" pageNumber="45">Figures 187190:</emphasis>
<taxonomicName id="FFD84D4AA839FFBEFE4EF8E19C2CF8F6" authorityName="Lourenco" authorityYear="1997" box="[398,590,1846,1867]" class="Arachnida" family="Buthidae" genus="Lychas" kingdom="Animalia" order="Scorpiones" pageId="46" pageNumber="45" phylum="Arthropoda" rank="species" species="srilankensis">
<emphasis id="0AACEADBA839FFBEFE4EF8E19C2CF8F6" box="[398,590,1846,1867]" italics="true" pageId="46" pageNumber="45">Lychas srilankensis</emphasis>
</taxonomicName>
.
<emphasis id="0AACEADBA839FFBEFD9CF8E29CB4F8F7" bold="true" box="[604,726,1845,1867]" pageId="46" pageNumber="45">Figures 187</emphasis>
. Male after 5th ecdysis at locality 15CJ.
<emphasis id="0AACEADBA839FFBEFBB5F8E29A8DF8F7" bold="true" box="[1141,1263,1845,1867]" pageId="46" pageNumber="45">Figures 188</emphasis>
. Female after 5th ecdysis at locality 15CO.
<emphasis id="0AACEADBA839FFBEFE7AF8999C50F8DE" bold="true" box="[442,562,1870,1892]" pageId="46" pageNumber="45">Figures 189</emphasis>
. Male after 3rd ecdysis at locality 15CN.
<emphasis id="0AACEADBA839FFBEFC0AF8999A20F8DE" bold="true" box="[970,1090,1870,1892]" pageId="46" pageNumber="45">Figures 190</emphasis>
. Female after 5th ecdysis at locality 15CJ.
</paragraph>
</caption>
<caption id="6CA76641A838FFBFFF00F8E69FC4F8DC" ID-DOI="http://doi.org/10.5281/zenodo.7124604" ID-Zenodo-Dep="7124604" httpUri="https://zenodo.org/record/7124604/files/figure.png" pageId="47" pageNumber="46" startId="47.[192,278,1841,1865]" targetBox="[213,1417,167,1812]" targetPageId="47">
<paragraph id="386736C9A838FFBFFF00F8E69FC4F8DC" blockId="47.[192,1441,1841,1889]" pageId="47" pageNumber="46">
<emphasis id="0AACEADBA838FFBFFF00F8E69FE1F8F4" bold="true" box="[192,387,1841,1865]" pageId="47" pageNumber="46">Figures 191192:</emphasis>
<taxonomicName id="FFD84D4AA838FFBFFE49F8E49C25F8F5" authorityName="Lourenco" authorityYear="1997" box="[393,583,1843,1864]" class="Arachnida" family="Buthidae" genus="Lychas" kingdom="Animalia" order="Scorpiones" pageId="47" pageNumber="46" phylum="Arthropoda" rank="species" species="srilankensis">
<emphasis id="0AACEADBA838FFBFFE49F8E49C25F8F5" box="[393,583,1843,1864]" italics="true" pageId="47" pageNumber="46">Lychas srilankensis</emphasis>
</taxonomicName>
, female from locality 1 5CN with newborns before first ecdysis (191), and with juveniles after first ecdysis (192).
</paragraph>
</caption>
<caption id="6CA76641A827FFA0FF00FBF99C0EFB14" ID-DOI="http://doi.org/10.5281/zenodo.7124608" ID-Zenodo-Dep="7124608" httpUri="https://zenodo.org/record/7124608/files/figure.png" pageId="48" pageNumber="47" startId="48.[192,278,1070,1094]" targetBox="[213,1446,171,1037]" targetPageId="48">
<paragraph id="386736C9A827FFA0FF00FBF99C0EFB14" blockId="48.[192,1442,1070,1193]" pageId="48" pageNumber="47">
<emphasis id="0AACEADBA827FFA0FF00FBF99FE6FBFB" bold="true" box="[192,388,1070,1094]" pageId="48" pageNumber="47">Figures 193200:</emphasis>
Distal segments of leg IV, retroventral view of Sri Lankan
<taxonomicName id="FFD84D4AA827FFA0FC14FBE79A49FBF8" authorityName="C. L. Koch" authorityYear="1837" box="[980,1067,1072,1093]" class="Arachnida" family="Buthidae" kingdom="Animalia" order="Scorpiones" pageId="48" pageNumber="47" phylum="Arthropoda" rank="family">Buthidae</taxonomicName>
and
<taxonomicName id="FFD84D4AA827FFA0FB9DFBE79AA8FBF8" authorityName="Pocock" authorityYear="1893" box="[1117,1226,1072,1093]" class="Arachnida" family="Chaerilidae" kingdom="Animalia" order="Scorpiones" pageId="48" pageNumber="47" phylum="Arthropoda" rank="family">Chaerilidae</taxonomicName>
genera.
<taxonomicName id="FFD84D4AA827FFA0FAE0FBE79F5BFBE3" authorityName=": Fet" authorityYear="1997" class="Arachnida" family="Buthidae" genus="Buthoscorpio" kingdom="Animalia" order="Scorpiones" pageId="48" pageNumber="47" phylum="Arthropoda" rank="species" species="sarasinorum">
<emphasis id="0AACEADBA827FFA0FAE0FBE79F5BFBE3" italics="true" pageId="48" pageNumber="47">Buthoscorpio sarasinorum</emphasis>
</taxonomicName>
, male from locality 15CF (193),
<taxonomicName id="FFD84D4AA827FFA0FD44FB9E9D42FBE3" authorityName="Karsch" authorityYear="1879" box="[644,800,1097,1118]" class="Arachnida" family="Buthidae" genus="Charmus" kingdom="Animalia" order="Scorpiones" pageId="48" pageNumber="47" phylum="Arthropoda" rank="species" species="laneus">
<emphasis id="0AACEADBA827FFA0FD44FB9E9D42FBE3" box="[644,800,1097,1118]" italics="true" pageId="48" pageNumber="47">Charmus laneus</emphasis>
</taxonomicName>
, male from locality 15CO (194),
<taxonomicName id="FFD84D4AA827FFA0FBAEFB9E9B4FFBE3" authorityName=": Kovarik &amp; Ojanguren Affilastro" authorityYear="2013" box="[1134,1325,1097,1118]" class="Arachnida" family="Buthidae" genus="Hottentotta" kingdom="Animalia" order="Scorpiones" pageId="48" pageNumber="47" phylum="Arthropoda" rank="species" species="tamulus">
<emphasis id="0AACEADBA827FFA0FBAEFB9E9B4FFBE3" box="[1134,1325,1097,1118]" italics="true" pageId="48" pageNumber="47">Hottentotta tamulus</emphasis>
</taxonomicName>
, male from locality 15CK (195),
<taxonomicName id="FFD84D4AA827FFA0FE58FBB59C01FBCA" authorityName=": Vachon" authorityYear="1972" box="[408,611,1122,1143]" class="Arachnida" family="Buthidae" genus="Isometrus" kingdom="Animalia" order="Scorpiones" pageId="48" pageNumber="47" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="0AACEADBA827FFA0FE58FBB59C01FBCA" box="[408,611,1122,1143]" italics="true" pageId="48" pageNumber="47">Isometrus maculatus</emphasis>
</taxonomicName>
, female from locality 15CI (196),
<taxonomicName id="FFD84D4AA827FFA0FC04FBB59AE0FBCA" authorityName=": Vachon" authorityYear="1982" box="[964,1154,1122,1143]" class="Arachnida" family="Buthidae" genus="Isometrus" kingdom="Animalia" order="Scorpiones" pageId="48" pageNumber="47" phylum="Arthropoda" rank="species" species="thwaitesi">
<emphasis id="0AACEADBA827FFA0FC04FBB59AE0FBCA" box="[964,1154,1122,1143]" italics="true" pageId="48" pageNumber="47">Isometrus thwaitesi</emphasis>
</taxonomicName>
, female from locality 15CO (197),
<taxonomicName id="FFD84D4AA827FFA0FEC2FBAC9FA3FB2D" authorityName="Lourenco" authorityYear="1997" box="[258,449,1147,1168]" class="Arachnida" family="Buthidae" genus="Lychas" kingdom="Animalia" order="Scorpiones" pageId="48" pageNumber="47" phylum="Arthropoda" rank="species" species="srilankensis">
<emphasis id="0AACEADBA827FFA0FEC2FBAC9FA3FB2D" box="[258,449,1147,1168]" italics="true" pageId="48" pageNumber="47">Lychas srilankensis</emphasis>
</taxonomicName>
, male from locality 15CJ (198),
<taxonomicName id="FFD84D4AA827FFA0FCC8FBAC9DAFFB2D" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" baseAuthorityName="Karsch" baseAuthorityYear="1879" box="[776,973,1147,1168]" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="48" pageNumber="47" phylum="Arthropoda" rank="species" species="basilicus">
<emphasis id="0AACEADBA827FFA0FCC8FBAC9DAFFB2D" box="[776,973,1147,1168]" italics="true" pageId="48" pageNumber="47">Reddyanus basilicus</emphasis>
</taxonomicName>
, male from locality 15CS (199), and
<taxonomicName id="FFD84D4AA827FFA0FA83FBAC9F4DFB14" authorityName="Pocock" authorityYear="1894" class="Arachnida" family="Chaerilidae" genus="Chaerilus" kingdom="Animalia" order="Scorpiones" pageId="48" pageNumber="47" phylum="Arthropoda" rank="species" species="ceylonensis">
<emphasis id="0AACEADBA827FFA0FA83FBAC9F4DFB14" italics="true" pageId="48" pageNumber="47">Chaerilus ceylonensis</emphasis>
</taxonomicName>
, male from locality 15CD (200).
</paragraph>
</caption>
<paragraph id="386736C9A827FFA0FF00FA419CD8F9D8" blockId="48.[192,802,1430,1637]" pageId="48" pageNumber="47">
HEMISPERMATOPHORE (
<figureCitation id="A0E32A4CA827FFA0FE08FA419C0FFA10" box="[456,621,1430,1453]" captionStart="Figures 146149" captionStartId="38.[192,278,1446,1470]" captionTargetBox="[194,1436,179,1409]" captionTargetId="figure-11@38.[192,1439,168,1417]" captionTargetPageId="38" captionText="Figures 146149: Left hemispermatophore of Isometrus thwaitesi from locality 15CH. External (146), dorsal (147), internal (148) and ventral (149) views. Note: distal part of flagellum and basal part of trunk (foot) were damaged during dissection. Scale bar: 500 µm. Abbreviations: bl, basal lobe; f, flagellum; ml, median lobe; mlc, median lobe carina; pr, pars recta of flagellum; prf, pars reflecta of flagellum; t, trunk." figureDoi="http://doi.org/10.5281/zenodo.7124574" httpUri="https://zenodo.org/record/7124574/files/figure.png" pageId="48" pageNumber="47">Figs. 146149</figureCitation>
). Trunk broad, short, only slightly longer than capsule region. Capsule region broad. Flagellum laminiform, short, basally broadened and fused with median lobe. Median lobe rather narrow, distally truncate, with strong, curved dorsal carina. Basal lobe well developed, a broad hook-like process arising dorsally at base of median lobe.
</paragraph>
</subSubSection>
<subSubSection id="70C26542A827FFA0FF00F95C9FB4F91F" box="[192,470,1675,1698]" pageId="48" pageNumber="47" type="distribution">
<paragraph id="386736C9A827FFA0FF00F95C9FB4F91F" blockId="48.[192,470,1675,1698]" box="[192,470,1675,1698]" pageId="48" pageNumber="47">
DISTRIBUTION.
<collectingCountry id="40CF7659A827FFA0FEA7F95C9FADF91F" box="[359,463,1675,1698]" name="Sri Lanka" pageId="48" pageNumber="47">Sri Lanka</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>