336 lines
38 KiB
XML
336 lines
38 KiB
XML
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<mods:title id="376E923CA99662BD7F05BF6015599951">A revision of the genus Leptotyphlops in northeastern Africa and southwestern Arabia (Serpentes: Leptotyphlopidae)</mods:title>
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<mods:title id="8B4EA9FB0F872E55602A4B4862E84769">Zootaxa</mods:title>
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<mods:part id="E3337FF8605DD80DB919C1D6016EEF7D">
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<mods:date id="18F037C3D0B64D762C21BC66A8075FF1">2007</mods:date>
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<mods:number id="A0C513C39140BD2875F28FA1FB155F68">2007-02-15</mods:number>
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<mods:detail id="58DEDB6AAF28461F0BC75CBE00C023F0" type="volume">
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<mods:number id="1B2E2B778C7239FB5D318594F7AEB1BA">1408</mods:number>
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<mods:identifier id="B357720514F010219407C20EEDF87F36" type="DOI">10.5281/zenodo.6789060</mods:identifier>
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<treatment id="A77887C2FFF8FFDDFF0286424445B58F" ID-DOI="http://doi.org/10.5281/zenodo.6789146" ID-GBIF-Taxon="196374633" ID-Zenodo-Dep="6789146" LSID="urn:lsid:plazi:treatment:A77887C2FFF8FFDDFF0286424445B58F" httpUri="http://treatment.plazi.org/id/A77887C2FFF8FFDDFF0286424445B58F" lastPageId="39" lastPageNumber="40" pageId="37" pageNumber="38">
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<subSubSection id="67CB655FFFF8FFDFFF02864240DCB179" pageId="37" pageNumber="38" type="nomenclature">
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<paragraph id="2F6E36D4FFF8FFDFFF0286424248B11C" blockId="37.[151,778,1551,1612]" box="[151,778,1551,1577]" pageId="37" pageNumber="38">
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<heading id="742681B8FFF8FFDFFF0286424248B11C" bold="true" box="[151,778,1551,1577]" fontSize="11" level="1" pageId="37" pageNumber="38" reason="1">
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<emphasis id="1DA5EAC6FFF8FFDFFF0286424248B11C" bold="true" box="[151,778,1551,1577]" pageId="37" pageNumber="38">
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<taxonomicName id="E8D14D57FFF8FFDFFF0286424093B11C" authority="Published, 2007" authorityName="Published" authorityYear="2007" box="[151,465,1551,1577]" class="Reptilia" family="Leptotyphlopidae" genus="Leptotyphlops" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="aethiopicus" status="sp. nov.">
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<emphasis id="1DA5EAC6FFF8FFDFFF0286424093B11C" bold="true" box="[151,465,1551,1577]" italics="true" pageId="37" pageNumber="38">Leptotyphlops aethiopicus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="069657BDFFF8FFDFFE4C86424374B11C" box="[473,566,1551,1577]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
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(
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<figureCitation id="B7EA2A51FFF8FFDFFDD3864243EFB11C" box="[582,685,1551,1577]" captionStart="PLATE 10" captionStartId="35.[151,238,1819,1843]" captionTargetBox="[300,1292,358,1808]" captionTargetId="figure-177@35.[290,1299,316,1830]" captionTargetPageId="35" captionText="PLATE 10. Figure 1. Leptotyphlops monticolus. Dorsal, lateral and ventral views of the head of NMZB 15309 — Katare, Kivu, D.R.C. The line represents 1 mm to scale. Figure 2. Leptotyphlops mbanjensis sp. nov. Dorsal, lateral and ventral views of the head of the holotype, MCZ 48040 — Mbanja, Lindi District, Tanzania. The line represents 1 mm to scale. Figure 3. Leptotyphlops keniensis sp. nov. Dorsal, lateral and ventral views of the head of the holotype, NMK/O. 2903 — Nyeri, Kenya. The line represents 1 mm to scale. Figure 4. Leptotyphlops aethiopicus sp. nov. Dorsal, lateral and ventral views of the head of the holotype, BMNH 1911.12.13.17 — near Harer, Ethiopia. The line represents 1 mm to scale." figureDoi="http://doi.org/10.5281/zenodo.6789096" httpUri="https://zenodo.org/record/6789096/files/figure.png" pageId="37" pageNumber="38">Plate 10</figureCitation>
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, Fig. 4)
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="2F6E36D4FFF8FFDFFF02867F40DCB179" blockId="37.[151,778,1551,1612]" box="[151,414,1586,1612]" pageId="37" pageNumber="38">Ethiopian worm snake</paragraph>
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</subSubSection>
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<subSubSection id="67CB655FFFF8FFDFFF0286374227B004" pageId="37" pageNumber="38" type="reference_group">
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||
<paragraph id="2F6E36D4FFF8FFDFFF0286374318B184" blockId="37.[151,1436,1658,1841]" pageId="37" pageNumber="38">
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||
<treatmentCitationGroup id="0FC111FAFFF8FFDFFF0286374318B184" pageId="37" pageNumber="38">
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<emphasis id="1DA5EAC6FFF8FFDFFF028637402DB1A4" box="[151,367,1658,1681]" italics="true" pageId="37" pageNumber="38">
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<taxonomicName id="E8D14D57FFF8FFDFFF028637400EB1A4" ID-CoL="3G6GS" authorityName="Boulenger" authorityYear="1890" box="[151,332,1658,1681]" class="Squamata" family="Leptotyphlopidae" genus="Glauconia" higherTaxonomySource="CoL" kingdom="Animalia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="emini">Glauconia emini</taxonomicName>
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—
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||
</emphasis>
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<treatmentCitation id="AE7010C5FFF8FFDFFEEC8637431BB1A4" author="Sternfeld, R." box="[377,601,1658,1681]" page="239" pageId="37" pageNumber="38" year="1908">
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||
<bibRefCitation id="4B404B25FFF8FFDFFEEC8637431BB1A4" author="Sternfeld" box="[377,601,1658,1681]" firstAuthor="Sternfeld" page="239" pageId="37" pageNumber="38" pagination="236 - 247" refId="ref62641" refString="Sternfeld, R. (1908) Zur Schlangenfauna Ostafrikas. I. Schlangen aus Sud-Abessinien. II. Schlangen aus Britisch- Ostafrika. III. Schlangen aus Portugiesisch-Ostafrika. Mitteilungen aus dem Zoologischen Museum in Berlin, 4 (1), 236 - 247." type="journal article" year="1908">Sternfeld, 1908: 239</bibRefCitation>
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</treatmentCitation>
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(Lake Haramaya; Bulli Hill);
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<treatmentCitation id="AE7010C5FFF8FFDFFC3B863745E9B1A4" author="Boulenger, G. A." box="[942,1195,1658,1681]" page="643" pageId="37" pageNumber="38" year="1915">
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<bibRefCitation id="4B404B25FFF8FFDFFC3B863745E9B1A4" author="Boulenger" box="[942,1195,1658,1681]" firstAuthor="Boulenger" page="643" pageId="37" pageNumber="38" pagination="641 - 658" refId="ref57529" refString="Boulenger, G. A. (1915 c) A list of the snakes of North-East Africa, from the Tropic to the Soudan and Somaliland, including Socotra. Proceedings of the Zoological Society of London, 1915, 641 - 658." type="journal article" year="1915" yearSuffix="c">Boulenger, 1915c: 643</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="AE7010C5FFF8FFDFFB2C863744DEB1A4" author="Scortecci, G." box="[1209,1436,1658,1681]" page="128" pageId="37" pageNumber="38" year="1939">
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<bibRefCitation id="4B404B25FFF8FFDFFB2C863744DEB1A4" author="Scortecci" box="[1209,1436,1658,1681]" firstAuthor="Scortecci" page="128" pageId="37" pageNumber="38" pagination="125 - 150" refId="ref62315" refString="Scortecci, G. (1939) Reptilia. In: Missione Biologica nel Paese dei Borana. Volume secondo. Raccolte Zoologiche. Parte prima. Reale Accademia d'Italia, Roma, pp. 125 - 150." type="book chapter" year="1939">Scortecci, 1939: 128</bibRefCitation>
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</treatmentCitation>
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(Neghelli; Arero), 1943: 283 (Murle).
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||
</treatmentCitationGroup>
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||
</paragraph>
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<paragraph id="2F6E36D4FFF8FFDFFF0286F74283B1E4" blockId="37.[151,1436,1658,1841]" box="[151,961,1722,1745]" pageId="37" pageNumber="38">
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||
<treatmentCitationGroup id="0FC111FAFFF8FFDFFF0286F74283B1E4" box="[151,961,1722,1745]" pageId="37" pageNumber="38">
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||
?
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||
<emphasis id="1DA5EAC6FFF8FFDFFF3C86F74030B1E4" box="[169,370,1722,1745]" italics="true" pageId="37" pageNumber="38">
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||
<taxonomicName id="E8D14D57FFF8FFDFFF3C86F74011B1E4" ID-CoL="3G6HL" box="[169,339,1722,1745]" class="Squamata" family="Leptotyphlopidae" genus="Glauconia" higherTaxonomySource="CoL" kingdom="Animalia" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="cairi">Glauconia cairi</taxonomicName>
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||
—
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||
</emphasis>
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||
<treatmentCitation id="AE7010C5FFF8FFDFFEED86F74323B1E4" author="Boulenger, G. A." box="[376,609,1722,1745]" page="441" pageId="37" pageNumber="38" year="1906">
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||
<bibRefCitation id="4B404B25FFF8FFDFFEED86F74323B1E4" author="Boulenger" box="[376,609,1722,1745]" firstAuthor="Boulenger" page="441" pageId="37" pageNumber="38" pagination="441" refId="ref57397" refString="Boulenger, G. A. (1906) Description of a new snake of the genus Glauconia, from Somaliland. The Annals and Magazine of Natural History (London), (7) 18 (108): 441." type="journal article" year="1906">Boulenger, 1906: 441</bibRefCitation>
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||
</treatmentCitation>
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||
;
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||
<treatmentCitation id="AE7010C5FFF8FFDFFDF986F74267B1E4" author="Werner, F." box="[620,805,1722,1745]" page="42" pageId="37" pageNumber="38" year="1907">
|
||
<bibRefCitation id="4B404B25FFF8FFDFFDF986F74267B1E4" author="Werner" box="[620,805,1722,1745]" firstAuthor="Werner" page="42" pageId="37" pageNumber="38" pagination="1823 - 1926" refId="ref64136" refString="Werner, F. (1907) Ergebnisse der mit Subvention aus der Erbschaft Treitl unternommenen zoologischen Forschungsreise Dr. Franz Werner's nach dem agyptischen Sudan und Nord-Uganda. Sitzungsberichten der kaiserlichen Akademie der Wissenschaften in Wien, Mathematisch-Naturwissenschaftliche Klasse, 116 (1), 1823 - 1926, Pls. 1 - 4." type="journal article" year="1907">Werner, 1907: 42</bibRefCitation>
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||
</treatmentCitation>
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||
(part, Harrar).
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||
</treatmentCitationGroup>
|
||
</paragraph>
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||
<paragraph id="2F6E36D4FFF8FFDFFF028697430EB1C4" blockId="37.[151,1436,1658,1841]" box="[151,588,1754,1777]" pageId="37" pageNumber="38">
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||
<treatmentCitationGroup id="0FC111FAFFF8FFDFFF028697430EB1C4" box="[151,588,1754,1777]" pageId="37" pageNumber="38">
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||
<emphasis id="1DA5EAC6FFF8FFDFFF02869740CCB1C4" box="[151,398,1754,1777]" italics="true" pageId="37" pageNumber="38">
|
||
<taxonomicName id="E8D14D57FFF8FFDFFF028697402DB1C4" box="[151,367,1754,1777]" class="Reptilia" family="Leptotyphlopidae" genus="Leptotyphlops" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="emini">Leptotyphlops emini</taxonomicName>
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||
—
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||
</emphasis>
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||
<treatmentCitation id="AE7010C5FFF8FFDFFE008697430AB1C4" author="Parker, H. W." box="[405,584,1754,1777]" page="20" pageId="37" pageNumber="38" year="1949">
|
||
<bibRefCitation id="4B404B25FFF8FFDFFE008697430AB1C4" author="Parker" box="[405,584,1754,1777]" firstAuthor="Parker" page="20" pageId="37" pageNumber="38" pagination="1 - 115" refId="ref61558" refString="Parker, H. W. (1949) The snakes of Somaliland and the Sokotra Islands. Zoologische Verhandelingen (Leiden), 6, 1 - 115." type="journal article" year="1949">Parker, 1949: 20</bibRefCitation>
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||
</treatmentCitation>
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||
.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="2F6E36D4FFF8FFDFFF0286B7422EB024" blockId="37.[151,1436,1658,1841]" box="[151,876,1786,1809]" pageId="37" pageNumber="38">
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||
<emphasis id="1DA5EAC6FFF8FFDFFF0286B74092B024" box="[151,464,1786,1809]" italics="true" pageId="37" pageNumber="38">
|
||
<taxonomicName id="E8D14D57FFF8FFDFFF0286B740F3B024" authority="emini" box="[151,433,1786,1809]" class="Reptilia" family="Leptotyphlopidae" genus="Leptotyphlops" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="38" phylum="Chordata" rank="subSpecies" species="emini" subSpecies="emini">Leptotyphlops emini emini</taxonomicName>
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||
—
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||
</emphasis>
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||
<bibRefCitation id="4B404B25FFF8FFDFFE4286B743FFB024" author="Loveridge, A." box="[471,701,1786,1809]" pageId="37" pageNumber="38" pagination="1 - 19" refId="ref60934" refString="Loveridge, A. (1957 a) On a third collection of reptiles taken in Tanganyika by C. J. P. Ionides, Esq. Tanganyika Notes and Records, (43), 1 - 19." type="journal article" year="1957">Loveridge, 1957: 247</bibRefCitation>
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||
(part, footnote).
|
||
</paragraph>
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||
<paragraph id="2F6E36D4FFF8FFDFFF0287574227B004" blockId="37.[151,1436,1658,1841]" box="[151,869,1818,1841]" pageId="37" pageNumber="38">
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<treatmentCitationGroup id="0FC111FAFFF8FFDFFF0287574227B004" box="[151,869,1818,1841]" pageId="37" pageNumber="38">
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<emphasis id="1DA5EAC6FFF8FFDFFF02875740FAB004" box="[151,440,1818,1841]" italics="true" pageId="37" pageNumber="38">
|
||
<taxonomicName id="E8D14D57FFF8FFDFFF02875740DBB004" box="[151,409,1818,1841]" class="Reptilia" family="Leptotyphlopidae" genus="Leptotyphlops" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="nigricans">Leptotyphlops nigricans</taxonomicName>
|
||
—
|
||
</emphasis>
|
||
<treatmentCitation id="AE7010C5FFF8FFDFFE2B87574259B004" author="Largen, M. J. & Rasmussen, J. B." box="[446,795,1818,1841]" page="325" pageId="37" pageNumber="38" year="1993">
|
||
<bibRefCitation id="4B404B25FFF8FFDFFE2B87574259B004" author="Largen & Rasmussen" box="[446,795,1818,1841]" firstAuthor="Largen" page="325" pageId="37" pageNumber="38" pagination="313 - 434" refId="ref59920" refString="Largen, M. J. & Rasmussen, J. B. (1993) Catalogue of the snakes of Ethiopia (Reptilia Serpentes), including identification keys. Tropical Zoology (Firenze), 6, 313 - 434." type="journal article" year="1993">Largen & Rasmussen, 1993: 325</bibRefCitation>
|
||
</treatmentCitation>
|
||
(part).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="67CB655FFFF8FFDCFF02872F4425B7EF" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38" type="materials_examined">
|
||
<paragraph id="2F6E36D4FFF8FFDFFF02872F43FCB091" blockId="37.[151,1436,1890,2036]" pageId="37" pageNumber="38">
|
||
<materialsCitation id="9FB93C89FFF8FFDFFF02872F43FCB091" ID-GBIF-Occurrence="3827581326" collectionCode="BMNH" collectorName="Kristensen." country="Ethiopia" elevation="2000" latitude="9.3" location="Harrar" longLatPrecision="1300" longitude="42.133335" municipality="Harerge Region" pageId="37" pageNumber="38" specimenCode="BMNH 1911.12" specimenCount="1" stateProvince="Hareri Hizb" typeStatus="holotype">
|
||
<emphasis id="1DA5EAC6FFF8FFDFFF02872F4052B049" bold="true" box="[151,272,1890,1916]" pageId="37" pageNumber="38">
|
||
<typeStatus id="F06A8876FFF8FFDFFF02872F4049B049" box="[151,267,1890,1916]" pageId="37" pageNumber="38" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<specimenCode id="7F779EAFFFF8FFDFFE8C872F409EB049" box="[281,476,1890,1916]" collectionCode="BMNH" pageId="37" pageNumber="38">BMNH 1911.12</specimenCode>
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||
.13.17 from near
|
||
<location id="2A0E600FFFF8FFDFFD32872F43B1B049" LSID="urn:lsid:plazi:treatment:A77887C2FFF8FFDDFF0286424445B58F:2A0E600FFFF8FFDFFD32872F43B1B049" box="[679,755,1890,1916]" country="Ethiopia" latitude="9.3" longLatPrecision="1300" longitude="42.133335" municipality="Harerge Region" name="Harrar" pageId="37" pageNumber="38" stateProvince="Hareri Hizb">Harrar</location>
|
||
[=
|
||
<collectingRegion id="ED15F836FFF8FFDFFCB5872F422AB049" box="[800,872,1890,1916]" country="Ethiopia" name="Hareri Hizb" pageId="37" pageNumber="38">Harar</collectingRegion>
|
||
],
|
||
<collectingMunicipality id="CF0AACAEFFF8FFDFFCE9872F4578B049" box="[892,1082,1890,1916]" pageId="37" pageNumber="38">Harerge Region</collectingMunicipality>
|
||
,
|
||
<collectingCountry id="57C67644FFF8FFDFFBDD872F45E9B049" box="[1096,1195,1890,1916]" name="Ethiopia" pageId="37" pageNumber="38">Ethiopia</collectingCountry>
|
||
(
|
||
<geoCoordinate id="4AE55013FFF8FFDFFB28872F4467B049" box="[1213,1317,1890,1916]" degrees="09" direction="north" minutes="18" orientation="latitude" pageId="37" pageNumber="38" precision="925" value="9.3">09°18’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="4AE55013FFF8FFDFFAA6872F44D5B049" box="[1331,1431,1890,1916]" degrees="42" direction="east" minutes="08" orientation="longitude" pageId="37" pageNumber="38" precision="925" value="42.133335">42°08’E</geoCoordinate>
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||
, elevation ca.
|
||
<quantity id="E8299B31FFF8FFDFFEA787C740CCB091" box="[306,398,1930,1956]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="37" pageNumber="38" unit="m" value="2000.0">
|
||
<elevation id="A4FCD1E7FFF8FFDFFEA787C740CCB091" box="[306,398,1930,1956]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="37" pageNumber="38" unit="m" value="2000.0">2000 m</elevation>
|
||
</quantity>
|
||
), collected by
|
||
<collectorName id="82245302FFF8FFDFFDAC87C743FCB091" box="[569,702,1930,1956]" pageId="37" pageNumber="38">Kristensen.</collectorName>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="2F6E36D4FFF8FFDCFF5387FF4425B7EF" blockId="37.[151,1436,1890,2036]" lastBlockId="38.[151,1436,152,418]" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38">
|
||
<materialsCitation id="9FB93C89FFF8FFDFFF5387FF4561B0F9" ID-GBIF-Occurrence="3827581407" box="[198,1059,1970,1996]" collectionCode="BMNH" collectorName="Kristensen." country="Ethiopia" elevation="2000" latitude="9.3" location="Harrar" longLatPrecision="1300" longitude="42.133335" municipality="Harerge Region" pageId="37" pageNumber="38" specimenCount="1" stateProvince="Hareri Hizb" typeStatus="paratype">
|
||
<emphasis id="1DA5EAC6FFF8FFDFFF5387FF4009B0F9" bold="true" box="[198,331,1970,1996]" pageId="37" pageNumber="38">
|
||
<typeStatus id="F06A8876FFF8FFDFFF5387FF4004B0F9" box="[198,326,1970,1996]" pageId="37" pageNumber="38" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<collectionCode id="49C0AE11FFF8FFDFFEC787FF40E8B0F9" box="[338,426,1970,1996]" pageId="37" pageNumber="38">BMNH</collectionCode>
|
||
1911.12.13.18–19 with the same data as the holotype
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="9FB93C89FFF8FFDCFBBA87FF4063B787" ID-GBIF-Occurrence="3827581410" collectingDate="1972-04-16" collectionCode="BMNH" collectorName="M. Largen" country="Ethiopia" elevation="2000" lastPageId="38" lastPageNumber="39" latitude="9.5" location="Harerge Region" longLatPrecision="1299" longitude="41.916668" pageId="37" pageNumber="38" specimenCode="BMNH 1976.685" specimenCount="1" stateProvince="Dire Dawa" typeStatus="paratype">
|
||
<specimenCode id="7F779EAFFFF8FFDFFBBA87FF45BEB0F9" box="[1071,1276,1970,1996]" collectionCode="BMNH" pageId="37" pageNumber="38">BMNH 1976.685</specimenCode>
|
||
—
|
||
<locationDeviation id="3EDC2FF6FFF8FFDFFABD87FF407EB0C1" location="Dire Dawa" pageId="37" pageNumber="38">
|
||
<quantity id="E8299B31FFF8FFDFFABD87FF4433B0F9" box="[1320,1393,1970,1996]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="37" pageNumber="38" unit="km" value="15.0">15 km</quantity>
|
||
SE of
|
||
<collectingRegion id="ED15F836FFF8FFDFFF2D8797407EB0C1" box="[184,316,2010,2036]" country="Ethiopia" name="Dire Dawa" pageId="37" pageNumber="38">Dire Dawa</collectingRegion>
|
||
</locationDeviation>
|
||
,
|
||
<location id="2A0E600FFFF8FFDFFEDC87974345B0C1" LSID="urn:lsid:plazi:treatment:A77887C2FFF8FFDDFF0286424445B58F:2A0E600FFFF8FFDFFEDC87974345B0C1" box="[329,519,2010,2036]" country="Ethiopia" latitude="9.5" longLatPrecision="1299" longitude="41.916668" name="Harerge Region" pageId="37" pageNumber="38" stateProvince="Dire Dawa">Harerge Region</location>
|
||
,
|
||
<collectingCountry id="57C67644FFF8FFDFFD808797433AB0C1" box="[533,632,2010,2036]" name="Ethiopia" pageId="37" pageNumber="38">Ethiopia</collectingCountry>
|
||
(
|
||
<geoCoordinate id="4AE55013FFF8FFDFFD1C879743B3B0C1" box="[649,753,2010,2036]" degrees="09" direction="north" minutes="30" orientation="latitude" pageId="37" pageNumber="38" precision="925" value="9.5">09°30’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="4AE55013FFF8FFDFFD6A87974226B0C1" box="[767,868,2010,2036]" degrees="41" direction="east" minutes="55" orientation="longitude" pageId="37" pageNumber="38" precision="925" value="41.916668">41°55’E</geoCoordinate>
|
||
,
|
||
<elevation id="A4FCD1E7FFF8FFDFFCE487974501B0C1" box="[881,1091,2010,2036]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="37" pageNumber="38" unit="m" value="2000.0">
|
||
elevation
|
||
<quantity id="E8299B31FFF8FFDFFC7087974501B0C1" box="[997,1091,2010,2036]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="37" pageNumber="38" unit="m" value="2000.0">2000 m</quantity>
|
||
</elevation>
|
||
), collected by
|
||
<collectorName id="82245302FFF8FFDFFB6787974433B0C1" box="[1266,1393,2010,2036]" pageId="37" pageNumber="38">M. Largen</collectorName>
|
||
,
|
||
<date id="5B6F1014FFF8FFDCFAEA87974063B787" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38" value="1972-04-16">
|
||
<collectingDate id="4B2BE9FCFFF8FFDCFAEA87974063B787" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38" value="1972-04-16">16 April 1972</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="9FB93C89FFFBFFDCFEA180D542E8B7EF" ID-GBIF-Occurrence="3827581377" collectingDate="1976-05-17" collectionCode="BMNH" collectorName="M. Largen" country="Ethiopia" elevation="1900" latitude="4.9166665" location="Sidamo Region" longLatPrecision="1306" longitude="38.066666" pageId="38" pageNumber="39" specimenCode="BMNH 1977.2250" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="7F779EAFFFFBFFDCFEA180D54355B787" box="[308,535,152,178]" collectionCode="BMNH" pageId="38" pageNumber="39">BMNH 1977.2250</specimenCode>
|
||
—
|
||
<locationDeviation id="3EDC2FF6FFFBFFDCFDC780D54205B787" box="[594,839,152,178]" location="Javello" pageId="38" pageNumber="39">
|
||
<quantity id="E8299B31FFFBFFDCFDC780D543D6B787" box="[594,660,152,178]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="38" pageNumber="39" unit="km" value="5.0">5 km</quantity>
|
||
W of Javello
|
||
</locationDeviation>
|
||
[=Yabelo],
|
||
<location id="2A0E600FFFFBFFDCFC7480D545E2B787" LSID="urn:lsid:plazi:treatment:A77887C2FFF8FFDDFF0286424445B58F:2A0E600FFFFBFFDCFC7480D545E2B787" box="[993,1184,152,178]" country="Ethiopia" latitude="4.9166665" longLatPrecision="1306" longitude="38.066666" name="Sidamo Region" pageId="38" pageNumber="39">Sidamo Region</location>
|
||
,
|
||
<collectingCountry id="57C67644FFFBFFDCFB2180D54455B787" box="[1204,1303,152,178]" name="Ethiopia" pageId="38" pageNumber="39">Ethiopia</collectingCountry>
|
||
(
|
||
<geoCoordinate id="4AE55013FFFBFFDCFABB80D544D4B787" box="[1326,1430,152,178]" degrees="04" direction="north" minutes="55" orientation="latitude" pageId="38" pageNumber="39" precision="925" value="4.9166665">04°55’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="4AE55013FFFBFFDCFF02808D41B8B7EF" box="[151,250,192,218]" degrees="38" direction="east" minutes="04" orientation="longitude" pageId="38" pageNumber="39" precision="925" value="38.066666">38°04’E</geoCoordinate>
|
||
,
|
||
<elevation id="A4FCD1E7FFFBFFDCFE92808D4094B7EF" box="[263,470,192,218]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="38" pageNumber="39" unit="m" value="1900.0">
|
||
elevation
|
||
<quantity id="E8299B31FFFBFFDCFEEF808D4094B7EF" box="[378,470,192,218]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="38" pageNumber="39" unit="m" value="1900.0">1900 m</quantity>
|
||
</elevation>
|
||
), collected by
|
||
<collectorName id="82245302FFFBFFDCFD14808D43BDB7EF" box="[641,767,192,218]" pageId="38" pageNumber="39">M. Largen</collectorName>
|
||
,
|
||
<date id="5B6F1014FFFBFFDCFC9E808D42E8B7EF" box="[779,938,192,218]" pageId="38" pageNumber="39" value="1976-05-17">
|
||
<collectingDate id="4B2BE9FCFFFBFFDCFC9E808D42E8B7EF" box="[779,938,192,218]" pageId="38" pageNumber="39" value="1976-05-17">17 May 1976</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="9FB93C89FFFBFFDCFC23808D4421B7EF" ID-GBIF-Occurrence="3827581481" ID-GBIF-Specimen="440047136" box="[950,1379,192,218]" collectionCode="MNHN" country="Ethiopia" httpUri="https://www.gbif.org/occurrence/440047136" location="Harrar" pageId="38" pageNumber="39" specimenCode="MNHN 1906.45" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="7F779EAFFFFBFFDCFC23808D4534B7EF" box="[950,1142,192,218]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="38" pageNumber="39">MNHN 1906.45</specimenCode>
|
||
—
|
||
<location id="2A0E600FFFFBFFDCFB37808D45B2B7EF" LSID="urn:lsid:plazi:treatment:A77887C2FFF8FFDDFF0286424445B58F:2A0E600FFFFBFFDCFB37808D45B2B7EF" box="[1186,1264,192,218]" country="Ethiopia" name="Harrar" pageId="38" pageNumber="39">Harrar</location>
|
||
,
|
||
<collectingCountry id="57C67644FFFBFFDCFB69808D4421B7EF" box="[1276,1379,192,218]" name="Ethiopia" pageId="38" pageNumber="39">Ethiopia</collectingCountry>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="67CB655FFFFBFFDCFF5380AA4090B667" pageId="38" pageNumber="39" type="diagnosis">
|
||
<paragraph id="2F6E36D4FFFBFFDCFF5380AA4090B667" blockId="38.[151,1436,152,418]" pageId="38" pageNumber="39">
|
||
<emphasis id="1DA5EAC6FFFBFFDCFF5380AA4004B634" bold="true" box="[198,326,231,257]" pageId="38" pageNumber="39">Diagnosis.</emphasis>
|
||
A member of the
|
||
<taxonomicName id="E8D14D57FFFBFFDCFDB880A5420FB634" box="[557,845,232,257]" class="Reptilia" family="Leptotyphlopidae" genus="Leptotyphlops" kingdom="Animalia" order="Squamata" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="nigricans">
|
||
<emphasis id="1DA5EAC6FFFBFFDCFDB880A5420FB634" box="[557,845,232,257]" italics="true" pageId="38" pageNumber="39">Leptotyphlops nigricans</emphasis>
|
||
</taxonomicName>
|
||
species group, closest to to
|
||
<emphasis id="1DA5EAC6FFFBFFDCFB3E80A54455B634" box="[1195,1303,232,257]" italics="true" pageId="38" pageNumber="39">
|
||
<taxonomicName id="E8D14D57FFFBFFDCFB3E80A54450B634" box="[1195,1298,232,257]" class="Reptilia" family="Leptotyphlopidae" genus="Leptotyphlops" kingdom="Animalia" order="Squamata" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="emini">L. emini</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
but distinguished therefrom by its high middorsal count (239–261), slender build (total length/ diameter ratio 61–77) and white tail tip ventrally.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="67CB655FFFFBFFDCFF538112437CB697" pageId="38" pageNumber="39" type="etymology">
|
||
<paragraph id="2F6E36D4FFFBFFDCFF538112437CB697" blockId="38.[151,1436,152,418]" pageId="38" pageNumber="39">
|
||
<emphasis id="1DA5EAC6FFFBFFDCFF5381124011B64C" bold="true" box="[198,339,351,377]" pageId="38" pageNumber="39">Etymology.</emphasis>
|
||
The specific name is derived from the Latin
|
||
<taxonomicName id="E8D14D57FFFBFFDCFC12812D454CB64C" authorityName="Published" authorityYear="2007" box="[903,1038,352,377]" class="Reptilia" family="Leptotyphlopidae" genus="Leptotyphlops" kingdom="Animalia" order="Squamata" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="aethiopicus">
|
||
<emphasis id="1DA5EAC6FFFBFFDCFC12812D454CB64C" box="[903,1038,352,377]" italics="true" pageId="38" pageNumber="39">aethiopicus</emphasis>
|
||
</taxonomicName>
|
||
= of
|
||
<collectingCountry id="57C67644FFFBFFDCFBCC812D4582B64F" box="[1113,1216,352,378]" name="Ethiopia" pageId="38" pageNumber="39">Ethiopia</collectingCountry>
|
||
, as this species is endemic to the Ethiopian highlands.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="7BAE665CFFFBFFDCFF0285D541B1B2E7" ID-DOI="http://doi.org/10.5281/zenodo.6789098" ID-Zenodo-Dep="6789098" httpUri="https://zenodo.org/record/6789098/files/figure.png" pageId="38" pageNumber="39" startId="38.[151,238,1432,1456]" targetBox="[415,1166,479,1405]" targetPageId="38">
|
||
<paragraph id="2F6E36D4FFFBFFDCFF0285D541B1B2E7" blockId="38.[151,1436,1432,1490]" pageId="38" pageNumber="39">
|
||
<emphasis id="1DA5EAC6FFFBFFDCFF0285D54056B285" bold="true" box="[151,276,1432,1456]" pageId="38" pageNumber="39">PLATE 11.</emphasis>
|
||
<taxonomicName id="E8D14D57FFFBFFDCFE8985D440F1B285" baseAuthorityName="Broadley & Broadley" baseAuthorityYear="1999" box="[284,435,1433,1456]" class="Reptilia" family="Leptotyphlopidae" genus="Leptotyphlops" kingdom="Animalia" order="Squamata" pageId="38" pageNumber="39" phylum="Chordata" rank="genus">
|
||
<emphasis id="1DA5EAC6FFFBFFDCFE8985D440F1B285" box="[284,435,1433,1456]" italics="true" pageId="38" pageNumber="39">Leptotyphlops</emphasis>
|
||
</taxonomicName>
|
||
. Distribution of the
|
||
<taxonomicName id="E8D14D57FFFBFFDCFD1B85D442D6B285" box="[654,916,1433,1456]" class="Reptilia" family="Leptotyphlopidae" genus="Leptotyphlops" kingdom="Animalia" order="Squamata" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="scutifrons">
|
||
<emphasis id="1DA5EAC6FFFBFFDCFD1B85D442D6B285" box="[654,916,1433,1456]" italics="true" pageId="38" pageNumber="39">Leptotyphlops scutifrons</emphasis>
|
||
</taxonomicName>
|
||
species group in East Africa. Type localities are labelled.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="67CB655FFFFBFFDDFF53864D43ECB577" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39" type="description">
|
||
<paragraph id="2F6E36D4FFFBFFDCFF53864D42ABB176" blockId="38.[151,1436,1536,2003]" pageId="38" pageNumber="39">
|
||
<emphasis id="1DA5EAC6FFFBFFDCFF53864D4015B12F" bold="true" box="[198,343,1536,1562]" pageId="38" pageNumber="39">Description</emphasis>
|
||
(
|
||
<typeStatus id="F06A8876FFFBFFDCFEFC864C408DB12E" box="[361,463,1537,1563]" pageId="38" pageNumber="39" type="paratype">paratype</typeStatus>
|
||
variations in parentheses). Body cylindrical, with head and neck broadened and flattened, the short tail tapers slightly before a very small terminal spine.
|
||
</paragraph>
|
||
<paragraph id="2F6E36D4FFFBFFDCFF53861C42E2B0E6" blockId="38.[151,1436,1536,2003]" pageId="38" pageNumber="39">Snout rounded, rostral broad (0.52–0.57 head width, mean = 0.55) and truncated, only slightly wider than supranasals anteriorly and not extending past a line connecting the eyes anteriorly, whereas the supranasals extend to a line connecting the eyes posteriorly, rostral lacking a preoral groove ventrally. Behind rostral, upper lip bordered by infranasal (nostril midway between rostral and supranasal along nasal suture), small (moderate) anterior supralabial with width along lip equal to that of infranasal, large ocular with large eye centrally placed and tall posterior supralabial. Supraoculars pentagonal, anteriorly wedged between upper nasal and ocular, posteriorly wedged between hexagonal subequal frontal and postfrontal, which are larger than the supraocular, but smaller than the hexagonal interparietal, which is also larger than the interoccipital and the vertebral series of scales following it. Parietals oblique, subequal to the fused occipitals, in contact with the posterior supralabial. Temoral single. No mental, four infralabials.</paragraph>
|
||
<paragraph id="2F6E36D4FFFAFFDDFF5380D5403DB637" blockId="39.[151,1436,152,698]" pageId="39" pageNumber="40">Body covered with 14 rows of smooth, imbricate, subequal scales. Reduction to 10 scale rows on the tail takes place lateral to the subtriangular cloacal shield, which is entire. Total middorsals 250 (243–261), subcaudals 24 (23–29).</paragraph>
|
||
<paragraph id="2F6E36D4FFFAFFDDFF53815D4216B61F" blockId="39.[151,1436,152,698]" box="[198,852,272,298]" pageId="39" pageNumber="40">
|
||
Total length
|
||
<quantity id="E8299B31FFFAFFDDFECF815D40FCB61F" box="[346,446,272,298]" metricMagnitude="-1" metricUnit="m" metricValue="1.45" pageId="39" pageNumber="40" unit="mm" value="145.0">145 mm</quantity>
|
||
; tail 13; diameter of body
|
||
<quantity id="E8299B31FFFAFFDDFD66815D4212B61F" box="[755,848,272,298]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="39" pageNumber="40" unit="mm" value="2.2">2.2 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="2F6E36D4FFFAFFDDFF538175451AB667" blockId="39.[151,1436,152,698]" box="[198,1112,312,338]" pageId="39" pageNumber="40">Total length/diameter ratio 72 (62–77); total length/tail ratio 12.2 (10.1–12.9).</paragraph>
|
||
<paragraph id="2F6E36D4FFFAFFDDFF53812D4224B697" blockId="39.[151,1436,152,698]" pageId="39" pageNumber="40">Dorsum and venter blackish-brown to dark brown except for the first infralabials, cloacal shield, and a patch of 14 (12–14) scales under the tail tip, which are white.</paragraph>
|
||
<paragraph id="2F6E36D4FFFAFFDDFF5381E24281B6FF" blockId="39.[151,1436,152,698]" box="[198,963,431,458]" pageId="39" pageNumber="40">
|
||
<emphasis id="1DA5EAC6FFFAFFDDFF5381E24042B6FC" bold="true" box="[198,256,431,457]" pageId="39" pageNumber="40">Size.</emphasis>
|
||
Largest specimen (BMNH 1977.2250) 145 + 16 =
|
||
<quantity id="E8299B31FFFAFFDDFCCF81FD42FCB6FF" box="[858,958,432,458]" metricMagnitude="-1" metricUnit="m" metricValue="1.61" pageId="39" pageNumber="40" unit="mm" value="161.0">161 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="2F6E36D4FFFAFFDDFF53819A43ECB577" blockId="39.[151,1436,152,698]" pageId="39" pageNumber="40">
|
||
<emphasis id="1DA5EAC6FFFAFFDDFF53819A406FB6C4" bold="true" box="[198,301,471,497]" pageId="39" pageNumber="40">Habitat.</emphasis>
|
||
When the
|
||
<typeStatus id="F06A8876FFFAFFDDFE2181954359B6C7" box="[436,539,472,498]" pageId="39" pageNumber="40" type="holotype">holotype</typeStatus>
|
||
was collected there was probably some of the climax
|
||
<taxonomicName id="E8D14D57FFFAFFDDFB3E8195445DB6C4" authorityName="C.Linnaeus" authorityYear="1753" box="[1195,1311,472,497]" class="Pinopsida" family="Cupressaceae" genus="Juniperus" kingdom="Plantae" order="Pinales" pageId="39" pageNumber="40" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="1DA5EAC6FFFAFFDDFB3E8195445DB6C4" box="[1195,1311,472,497]" italics="true" pageId="39" pageNumber="40">Juniperus</emphasis>
|
||
</taxonomicName>
|
||
woodland around Harer, but this whole area is now intensively cultivated. The climax vegetation southeast of
|
||
<collectingRegion id="ED15F836FFFAFFDDFA88824D44DEB52F" box="[1309,1436,512,538]" country="Ethiopia" name="Dire Dawa" pageId="39" pageNumber="40">Dire Dawa</collectingRegion>
|
||
and near Yabelo would be
|
||
<taxonomicName id="E8D14D57FFFAFFDDFE5A8265431FB574" box="[463,605,552,577]" class="Pinopsida" family="Podocarpaceae" genus="Podocarpus" kingdom="Plantae" order="Pinales" pageId="39" pageNumber="40" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="1DA5EAC6FFFAFFDDFE5A8265431FB574" box="[463,605,552,577]" italics="true" pageId="39" pageNumber="40">Podocarpus</emphasis>
|
||
</taxonomicName>
|
||
forest.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="67CB655FFFFAFFDDFF5382024445B58F" pageId="39" pageNumber="40" type="distribution">
|
||
<paragraph id="2F6E36D4FFFAFFDDFF53820244CCB5A7" blockId="39.[151,1436,152,698]" pageId="39" pageNumber="40">
|
||
<emphasis id="1DA5EAC6FFFAFFDDFF5382024025B55C" bold="true" box="[198,359,591,617]" pageId="39" pageNumber="40">Distribution.</emphasis>
|
||
So far only recorded from the Ethiopian highlands east of the rift valley,
|
||
<quantity id="E8299B31FFFAFFDDFB2D821D4419B55F" box="[1208,1371,592,618]" metricMagnitude="3" metricUnit="m" metricValue="1.95" metricValueMax="2.0" metricValueMin="1.9" pageId="39" pageNumber="40" unit="m" value="1950.0" valueMax="2000.0" valueMin="1900.0">1900–2000 m</quantity>
|
||
, with an additional specimen (middorsals 239, subcaudals 22) from
|
||
<collectingRegion id="ED15F836FFFAFFDDFCFA82354295B5A7" box="[879,983,632,658]" country="Kenya" name="Marsabit" pageId="39" pageNumber="40">Marsabit</collectingRegion>
|
||
in northern
|
||
<collectingCountry id="57C67644FFFAFFDDFBF2823545F4B5A7" box="[1127,1206,632,658]" name="Kenya" pageId="39" pageNumber="40">Kenya</collectingCountry>
|
||
,
|
||
<quantity id="E8299B31FFFAFFDDFB5782354459B5A7" box="[1218,1307,632,658]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="39" pageNumber="40" unit="m" value="1700.0">1700 m</quantity>
|
||
(
|
||
<figureCitation id="B7EA2A51FFFAFFDDFABF8235443DB5A7" box="[1322,1407,632,658]" captionStart="PLATE 5" captionStartId="13.[151,238,1737,1761]" captionTargetBox="[268,1322,271,1704]" captionTargetId="figure-152@13.[264,1323,266,1713]" captionTargetPageId="13" captionText="PLATE 5. Leptotyphlops. Distribution of the Leptotyphlops parkeri, L. reticulatus, L. longicaudus and L. nigricans species groups in northeastern Africa and southwestern Arabia." figureDoi="http://doi.org/10.5281/zenodo.6789078" httpUri="https://zenodo.org/record/6789078/files/figure.png" pageId="39" pageNumber="40">Plate 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="2F6E36D4FFFAFFDDFF5382D24445B58F" blockId="39.[151,1436,152,698]" box="[198,1287,671,698]" pageId="39" pageNumber="40">
|
||
<emphasis id="1DA5EAC6FFFAFFDDFF5382D24086B58C" bold="true" box="[198,452,671,697]" pageId="39" pageNumber="40">Additional material.</emphasis>
|
||
<collectingCountry id="57C67644FFFAFFDDFE5982ED4371B58F" box="[460,563,672,698]" name="Kenya" pageId="39" pageNumber="40">KENYA</collectingCountry>
|
||
.
|
||
<collectingRegion id="ED15F836FFFAFFDDFDAB82ED43E4B58F" box="[574,678,672,698]" country="Kenya" name="Marsabit" pageId="39" pageNumber="40">Marsabit</collectingRegion>
|
||
CAS 122975. No locality: NHCL 1549 (Scortecci).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |