476 lines
41 KiB
XML
476 lines
41 KiB
XML
<document ID-DOI="10.26879/891" ID-Zenodo-Dep="3990295" approvalRequired="202" approvalRequired_for_document="2" approvalRequired_for_illustrations="2" approvalRequired_for_textStreams="175" approvalRequired_for_treatments="23" checkinTime="1597772865363" checkinUser="torsten" docAuthor="Dale E. Greenwalt, Daniel J. Bickel, Peter H. Kerr, Gregory R. Curler, Brian V. Brown, Herman de Jong, Scott J. Fitzgerald, Torsten Dikow, Michal Tkoč, Christian Kehlmaier & Dalton De Souza Amorim" docDate="2019" docId="571F246BFFBDFFA71095FD21AB33C2F5" docLanguage="en" docName="greenwalt_etal_2019.imf" docOrigin="Paleontologia Electronica 22 (2), No. 50" docStyle="DocumentStyle{}" docTitle="Eosciarites hermes Dale E. Greenwalt & Daniel J. Bickel & Peter H. Kerr & Gregory R. Curler & Brian V. Brown & Herman de Jong & Scott J. Fitzgerald & Torsten Dikow & Michal Tkoč & Christian Kehlmaier & Dalton De Souza Amorim 2019, sp. nov." docType="treatment" docVersion="5" lastPageId="21" lastPageNumber="22" masterDocId="AB265C13FFAEFFB21318FF0FAF31C65B" masterDocTitle="Diptera of the middle Eocene Kishenehn Formation. I. Documenting of diversity at the family level" masterLastPageNumber="56" masterPageNumber="1" pageId="19" pageNumber="20" updateTime="1636465287172" updateUser="ExternalLinkService">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>Diptera of the middle Eocene Kishenehn Formation. I. Documenting of diversity at the family level</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Dale E. Greenwalt</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Daniel J. Bickel</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Peter H. Kerr</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Gregory R. Curler</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Brian V. Brown</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Herman de Jong</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Scott J. Fitzgerald</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Torsten Dikow</mods:namePart>
|
||
<mods:nameIdentifier type="LSID">F8869067-4618-4CCE-960C-E8A107F162FB</mods:nameIdentifier>
|
||
<mods:nameIdentifier type="ORCID">0000-0003-4816-2909</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Michal Tkoč</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Christian Kehlmaier</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Dalton De Souza Amorim</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>Paleontologia Electronica</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2019</mods:date>
|
||
<mods:detail type="volume">
|
||
<mods:number>22</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="issue">
|
||
<mods:number>2</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="numero">
|
||
<mods:number>50</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>1</mods:start>
|
||
<mods:end>56</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">10.26879/891</mods:identifier>
|
||
<mods:identifier type="Zenodo-Dep">3990295</mods:identifier>
|
||
<mods:identifier type="ZooBank">A6C79E56-3CCC-484E-B6AF-EAEEE1695FF6</mods:identifier>
|
||
</mods:mods>
|
||
<treatment LSID="urn:lsid:plazi:treatment:571F246BFFBDFFA71095FD21AB33C2F5" httpUri="http://treatment.plazi.org/id/571F246BFFBDFFA71095FD21AB33C2F5" lastPageId="21" lastPageNumber="22" pageId="19" pageNumber="20">
|
||
<subSubSection box="[909,1371,558,584]" pageId="19" pageNumber="20" type="nomenclature">
|
||
<paragraph blockId="19.[840,1441,168,1436]" box="[909,1371,558,584]" pageId="19" pageNumber="20">
|
||
<taxonomicName authority="Greenwalt" authorityName="Dale E. Greenwalt & Daniel J. Bickel & Peter H. Kerr & Gregory R. Curler & Brian V. Brown & Herman de Jong & Scott J. Fitzgerald & Torsten Dikow & Michal Tkoč & Christian Kehlmaier & Dalton De Souza Amorim" authorityYear="2019" box="[909,1269,558,584]" class="Insecta" family="Sciaridae" genus="Eosciarites" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="hermes" status="sp. nov.">
|
||
<emphasis box="[909,1136,558,583]" italics="true" pageId="19" pageNumber="20">Eosciarites hermes</emphasis>
|
||
Greenwalt
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel box="[1279,1371,558,583]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="19" pageNumber="20" type="multiple">
|
||
<paragraph blockId="19.[840,1441,168,1436]" pageId="19" pageNumber="20">
|
||
<figureCitation box="[1055,1184,590,615]" captionStart="FIGURE 16" captionStartId="20.[197,288,1383,1406]" captionTargetBox="[196,793,165,1361]" captionTargetId="figure@20.[196,793,165,1362]" captionTargetPageId="20" captionText="FIGURE 16. Eosciarites hermes sp. nov., USNM 624633. 1, Habitus; 2, Right wing, arrow points to the halter; 3, Line drawing of the right wing. Scale bars equal 1.0 mm (1) and 0.5 mm (2)." figureDoi="http://doi.org/10.5281/zenodo.3990329" httpUri="https://zenodo.org/record/3990329/files/figure.png" pageId="19" pageNumber="20">Figures 16</figureCitation>
|
||
,
|
||
<figureCitation box="[1196,1226,590,615]" captionStart="FIGURE 17" captionStartId="20.[843,933,753,776]" captionTargetBox="[843,1440,168,735]" captionTargetId="figure@20.[843,1440,167,735]" captionTargetPageId="20" captionText="FIGURE 17. Eosciarites hermes sp. nov., USNM 624633. 1, Head, arrow points to the labellum; 2, Tibial spur of foreleg (left arrow) and tibial spur of midleg (right arrow). Arrowheads denote a small dark patch of setae; 3, Terminalia, female. Arrow and arrow head denote the two-segmented cercus and sternite 10, respectively; Scale bars equal 0.3 mm (1) and 0.5 mm (2, 3)." figureDoi="http://doi.org/10.5281/zenodo.3990327" httpUri="https://zenodo.org/record/3990327/files/figure.png" pageId="19" pageNumber="20">17</figureCitation>
|
||
<emphasis bold="true" box="[840,1408,633,653]" pageId="19" pageNumber="20">
|
||
zoobank.org/
|
||
<uuid box="[978,1408,633,653]" pageId="19" pageNumber="20">C0541528-F65E-4593-9B09-9C1CB318C6C2</uuid>
|
||
</emphasis>
|
||
<emphasis bold="true" box="[840,982,670,695]" pageId="19" pageNumber="20">Etymology.</emphasis>
|
||
The generic epithet is a combination of the greek word Eos (early, dawn), the genus name
|
||
<taxonomicName authorityName="Meigen" authorityYear="1803" box="[914,989,734,759]" class="Insecta" family="Sciaridae" genus="Sciara" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[914,989,734,759]" italics="true" pageId="19" pageNumber="20">Sciara</emphasis>
|
||
</taxonomicName>
|
||
and the suffix “-ites” (Latin for ”having the nature of”).
|
||
<taxonomicName authorityName="Greenwalt" authorityYear="2019" box="[1021,1151,766,791]" class="Insecta" family="Scatopsidae" genus="Eosciarites" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1021,1151,766,791]" italics="true" pageId="19" pageNumber="20">Eosciarites</emphasis>
|
||
</taxonomicName>
|
||
is a collective parataxon as defined by
|
||
<bibRefCitation author="Rasnitsyn, A. P." box="[1007,1203,798,823]" journalOrPublisher="Paleontologicheskii Zhurnal" pageId="19" pageNumber="20" pagination="11 - 21" part="3" refId="ref42630" refString="Rasnitsyn, A. P. 1986. Parataxon and paranomenclature. Paleontologicheskii Zhurnal, 3: 11 - 21. (Paleontological Journal, 20 [3]: 25 - 33)" title="Parataxon and paranomenclature" type="journal article" year="1986">Rasnitsyn (1986</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Rasnitsyn, A. P." box="[1214,1279,798,823]" journalOrPublisher="Contributions to Zoology" pageId="19" pageNumber="20" pagination="3 - 41" part="66" refId="ref42664" refString="Rasnitsyn, A. P. 1996. Conceptual issues in phylogeny, taxonomy, and nomenclature. Contributions to Zoology, 66: 3 - 41." title="Conceptual issues in phylogeny, taxonomy, and nomenclature" type="journal article" year="1996">1996</bibRefCitation>
|
||
). The specific epithet is the Greek word Hermes (mythical messenger of the gods).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<materialsCitation collectionCode="USNM" country="United States of America" county="Smithsonian Institution" location="Department of Paleobiology" municipality="National Museum of Natural History" pageId="19" pageNumber="20" specimenCode="USNM 624633" specimenCount="1" stateProvince="Kishenehn Formation" typeStatus="Holotype">
|
||
<subSubSection pageId="19" pageNumber="20" type="materials_examined">
|
||
<paragraph blockId="19.[840,1441,168,1436]" pageId="19" pageNumber="20">
|
||
<emphasis bold="true" box="[840,961,894,919]" pageId="19" pageNumber="20">
|
||
<typeStatus box="[840,957,894,919]" pageId="19" pageNumber="20">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<specimenCode box="[968,1147,894,920]" collectionCode="USNM" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="19" pageNumber="20" type="Museum">
|
||
<collectionCode box="[968,1046,894,920]" pageId="19" pageNumber="20">USNM</collectionCode>
|
||
624633
|
||
</specimenCode>
|
||
, deposited in the
|
||
<location LSID="urn:lsid:plazi:treatment:571F246BFFBDFFA71095FD21AB33C2F5:DA69C3A6FFBDFFA11650FC71AB77C5EC" country="United States of America" county="Smithsonian Institution" municipality="National Museum of Natural History" name="Department of Paleobiology" pageId="19" pageNumber="20" stateProvince="Washington">Department of Paleobiology</location>
|
||
,
|
||
<collectingMunicipality pageId="19" pageNumber="20">National Museum of Natural History</collectingMunicipality>
|
||
(NMNH),
|
||
<collectingCounty box="[1044,1318,958,984]" pageId="19" pageNumber="20">Smithsonian Institution</collectingCounty>
|
||
,
|
||
<collectingRegion country="United States of America" name="Washington" pageId="19" pageNumber="20">Washington</collectingRegion>
|
||
,
|
||
<collectingRegion box="[892,1128,990,1016]" country="United States of America" name="District of Columbia" pageId="19" pageNumber="20">District of Columbia</collectingRegion>
|
||
,
|
||
<collectingCountry box="[1138,1197,990,1016]" name="United States of America" pageId="19" pageNumber="20" value="United States of America">USA</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="19" pageNumber="20" type="distribution">
|
||
<paragraph blockId="19.[840,1441,168,1436]" pageId="19" pageNumber="20">
|
||
<emphasis bold="true" box="[840,1016,1022,1047]" pageId="19" pageNumber="20">Type horizon.</emphasis>
|
||
<location LSID="urn:lsid:plazi:treatment:571F246BFFBDFFA71095FD21AB33C2F5:DA69C3A6FFBDFFA1171CFCF1AA61C243" box="[1028,1360,1022,1048]" name="Middle Eocene Coal Creek" pageId="19" pageNumber="20" stateProvince="Kishenehn Formation">Middle Eocene Coal Creek</location>
|
||
Member,
|
||
<collectingRegion box="[892,1149,1054,1079]" pageId="19" pageNumber="20">Kishenehn Formation</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="19.[840,1441,168,1436]" pageId="19" pageNumber="20">
|
||
<emphasis bold="true" box="[840,1008,1086,1111]" pageId="19" pageNumber="20">Type locality.</emphasis>
|
||
Dakin site, Middle Fork of the Flathead River (Pinnacle,
|
||
<collectingRegion box="[1102,1210,1118,1143]" country="United States of America" name="Montana" pageId="19" pageNumber="20">Montana</collectingRegion>
|
||
,
|
||
<collectingCountry box="[1220,1281,1118,1144]" name="United States of America" pageId="19" pageNumber="20" value="United States of America">USA</collectingCountry>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</materialsCitation>
|
||
<subSubSection pageId="19" pageNumber="20" type="diagnosis">
|
||
<paragraph blockId="19.[840,1441,168,1436]" pageId="19" pageNumber="20">
|
||
<emphasis bold="true" box="[840,1124,1150,1175]" pageId="19" pageNumber="20">Differential Diagnosis.</emphasis>
|
||
The absence of macrotrichia on veins M and Cu and flagellomeres with cylindrical nodes or necks differentiates this specimen from
|
||
<taxonomicName authorityName="Meigen" authorityYear="1803" box="[971,1046,1246,1271]" class="Insecta" family="Sciaridae" genus="Sciara" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[971,1046,1246,1271]" italics="true" pageId="19" pageNumber="20">Sciara</emphasis>
|
||
</taxonomicName>
|
||
. A three-segmented palpus, R
|
||
<subScript attach="left" box="[1428,1440,1256,1276]" fontSize="8" pageId="19" pageNumber="20">1</subScript>
|
||
joining C prior to medial fork and significantly longer than half the length of R, M+CuA significantly greater than bm-m, middle and hind tibia with a single apical tibial spur are all diagnostic of this specimen.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20" type="description">
|
||
<paragraph blockId="19.[840,987,1459,1484]" box="[840,987,1459,1484]" pageId="19" pageNumber="20">
|
||
<heading bold="true" box="[840,987,1459,1484]" fontSize="10" level="3" pageId="19" pageNumber="20" reason="2">
|
||
<emphasis bold="true" box="[840,987,1459,1484]" pageId="19" pageNumber="20">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="19.[840,1441,1507,1884]" pageId="19" pageNumber="20">
|
||
<emphasis bold="true" box="[840,1114,1507,1532]" pageId="19" pageNumber="20">Female (Figure 16.1).</emphasis>
|
||
Total length
|
||
<quantity box="[1279,1376,1507,1532]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="19" pageNumber="20" unit="mm" value="2.2">2.2 mm</quantity>
|
||
, light brown in colour.
|
||
</paragraph>
|
||
<paragraph blockId="19.[840,1441,1507,1884]" pageId="19" pageNumber="20">
|
||
<emphasis bold="true" box="[840,912,1571,1596]" pageId="19" pageNumber="20">Head.</emphasis>
|
||
Black, spherical,
|
||
<quantity box="[1145,1255,1571,1596]" metricMagnitude="-4" metricUnit="m" metricValue="2.7" pageId="19" pageNumber="20" unit="mm" value="0.27">0.27 mm</quantity>
|
||
diameter; eye about
|
||
<quantity box="[914,1018,1603,1628]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" pageId="19" pageNumber="20" unit="mm" value="0.12">0.12 mm</quantity>
|
||
in diameter. Maxillary palpus threesegmented, about
|
||
<quantity box="[1063,1168,1635,1660]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="19" pageNumber="20" unit="mm" value="0.19">0.19 mm</quantity>
|
||
in length, terminal two segments slightly longer than wide. Antennal base situated in a distinct depression, flagellomeres brown, quadrate basally. Apical flagellomeres not preserved (Figure 17.1).
|
||
</paragraph>
|
||
<paragraph blockId="19.[840,1441,1507,1884]" lastBlockId="20.[192,793,1552,1865]" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
|
||
<emphasis bold="true" box="[840,936,1795,1820]" pageId="19" pageNumber="20">Thorax.</emphasis>
|
||
Black to dark red, legs light brown. Scutum covered with short scattered setae. Mediotergite with several bristles on its posterior margin, katepisternum not attentuated. Haltere light brown, knob about
|
||
<quantity box="[331,435,1584,1609]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="20" pageNumber="21" unit="mm" value="0.09">0.09 mm</quantity>
|
||
wide and
|
||
<quantity box="[557,652,1584,1609]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="20" pageNumber="21" unit="mm" value="0.16">0.16mm</quantity>
|
||
long.
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3990327" ID-Zenodo-Dep="3990327" httpUri="https://zenodo.org/record/3990327/files/figure.png" pageId="20" pageNumber="21" startId="20.[843,933,753,776]" targetBox="[843,1440,168,735]" targetPageId="20">
|
||
<paragraph blockId="20.[843,1439,753,953]" pageId="20" pageNumber="21">
|
||
<emphasis bold="true" box="[843,986,753,777]" pageId="20" pageNumber="21">FIGURE 17.</emphasis>
|
||
<taxonomicName authority="Dale E. Greenwalt & Daniel J. Bickel & Peter H. Kerr & Gregory R. Curler & Brian V. Brown & Herman de Jong & Scott J. Fitzgerald & Torsten Dikow & Michal Tkoč & Christian Kehlmaier & Dalton De Souza Amorim, 2019" authorityName="Dale E. Greenwalt & Daniel J. Bickel & Peter H. Kerr & Gregory R. Curler & Brian V. Brown & Herman de Jong & Scott J. Fitzgerald & Torsten Dikow & Michal Tkoč & Christian Kehlmaier & Dalton De Souza Amorim" authorityYear="2019" box="[1007,1224,754,777]" class="Insecta" family="Scatopsidae" genus="Eosciarites" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="hermes" status="sp. nov.">
|
||
<emphasis box="[1007,1224,754,777]" italics="true" pageId="20" pageNumber="21">Eosciarites hermes</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1244,1345,754,777]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, USNM 624633.
|
||
<emphasis bold="true" box="[937,950,783,806]" pageId="20" pageNumber="21">1</emphasis>
|
||
, Head, arrow points to the labellum;
|
||
<emphasis bold="true" box="[1353,1366,783,806]" pageId="20" pageNumber="21">2</emphasis>
|
||
, Tibial spur of foreleg (left arrow) and tibial spur of midleg (right arrow). Arrowheads denote a small dark patch of setae;
|
||
<emphasis bold="true" box="[843,856,871,894]" pageId="20" pageNumber="21">3</emphasis>
|
||
, Terminalia, female. Arrow and arrow head denote the two-segmented cercus and sternite 10, respectively; Scale bars equal 0.3 mm (
|
||
<emphasis bold="true" box="[1122,1135,930,953]" pageId="20" pageNumber="21">1</emphasis>
|
||
) and 0.5 mm (
|
||
<emphasis bold="true" box="[1291,1304,930,953]" pageId="20" pageNumber="21">2</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[1318,1331,930,953]" pageId="20" pageNumber="21">3</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3990329" ID-Zenodo-Dep="3990329" httpUri="https://zenodo.org/record/3990329/files/figure.png" pageId="20" pageNumber="21" startId="20.[197,288,1383,1406]" targetBox="[196,793,165,1361]" targetPageId="20">
|
||
<paragraph blockId="20.[197,788,1383,1494]" pageId="20" pageNumber="21">
|
||
<emphasis bold="true" box="[197,340,1383,1406]" pageId="20" pageNumber="21">FIGURE 16.</emphasis>
|
||
<taxonomicName authority="Dale E. Greenwalt & Daniel J. Bickel & Peter H. Kerr & Gregory R. Curler & Brian V. Brown & Herman de Jong & Scott J. Fitzgerald & Torsten Dikow & Michal Tkoč & Christian Kehlmaier & Dalton De Souza Amorim, 2019" authorityName="Dale E. Greenwalt & Daniel J. Bickel & Peter H. Kerr & Gregory R. Curler & Brian V. Brown & Herman de Jong & Scott J. Fitzgerald & Torsten Dikow & Michal Tkoč & Christian Kehlmaier & Dalton De Souza Amorim" authorityYear="2019" box="[360,577,1383,1406]" class="Insecta" family="Scatopsidae" genus="Eosciarites" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="hermes" status="sp. nov.">
|
||
<emphasis box="[360,577,1383,1406]" italics="true" pageId="20" pageNumber="21">Eosciarites hermes</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[596,695,1383,1406]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, USNM 624633.
|
||
<emphasis bold="true" box="[294,307,1413,1436]" pageId="20" pageNumber="21">1</emphasis>
|
||
, Habitus;
|
||
<emphasis bold="true" box="[421,434,1413,1436]" pageId="20" pageNumber="21">2</emphasis>
|
||
, Right wing, arrow points to the halter;
|
||
<emphasis bold="true" box="[276,289,1442,1465]" pageId="20" pageNumber="21">3</emphasis>
|
||
, Line drawing of the right wing. Scale bars equal 1.0 mm (
|
||
<emphasis bold="true" box="[357,370,1471,1494]" pageId="20" pageNumber="21">1</emphasis>
|
||
) and 0.5 mm (
|
||
<emphasis bold="true" box="[526,539,1471,1494]" pageId="20" pageNumber="21">2</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="20.[192,793,1552,1865]" pageId="20" pageNumber="21">
|
||
<emphasis bold="true" box="[203,276,1616,1641]" pageId="20" pageNumber="21">Legs.</emphasis>
|
||
Setose, light brown basally, tibia and tarsi darker. Forecoxa with long setae along ventral margin, hind tibia with line of spines posteriorly; middle and hind tibia with one and possibly two apical spurs respectively, spurs longer than tibia diameter; foretibia with a spur longer than foretibial diameter on either side of a dark triangular area of setae (Figure 17.2).
|
||
</paragraph>
|
||
<paragraph blockId="20.[840,1441,1027,1668]" pageId="20" pageNumber="21">
|
||
<emphasis bold="true" box="[840,927,1027,1052]" pageId="20" pageNumber="21">Wings.</emphasis>
|
||
Length:
|
||
<quantity box="[1036,1146,1027,1052]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" pageId="20" pageNumber="21" unit="mm" value="1.45">1.45 mm</quantity>
|
||
, width:
|
||
<quantity box="[1239,1345,1027,1052]" metricMagnitude="-4" metricUnit="m" metricValue="5.2" pageId="20" pageNumber="21" unit="mm" value="0.52">0.52 mm</quantity>
|
||
(Figure 16.2-3). Membrane with microtrichia, macrotrichia restricted to C, R
|
||
<subScript attach="left" box="[1048,1060,1101,1121]" fontSize="8" pageId="20" pageNumber="21">1</subScript>
|
||
and R
|
||
<subScript attach="left" box="[1144,1180,1101,1121]" fontSize="8" pageId="20" pageNumber="21">4+5</subScript>
|
||
. C extending beyond apex of R
|
||
<subScript attach="left" box="[956,993,1138,1158]" fontSize="8" pageId="20" pageNumber="21">4+5</subScript>
|
||
, about half way to M
|
||
<subScript attach="left" box="[1244,1256,1138,1158]" fontSize="8" pageId="20" pageNumber="21">1</subScript>
|
||
. Sc short, free, R
|
||
<subScript attach="left" box="[859,871,1175,1195]" fontSize="8" pageId="20" pageNumber="21">1</subScript>
|
||
significantly longer than half of R, R
|
||
<subScript attach="left" box="[1339,1351,1175,1195]" fontSize="8" pageId="20" pageNumber="21">1</subScript>
|
||
with a short, brief posterior turn basally, ending well-short of medial fork. R
|
||
<subScript attach="left" box="[1032,1068,1244,1264]" fontSize="8" pageId="20" pageNumber="21">4+5</subScript>
|
||
reaching C well before wing tip. Rs oblique to R
|
||
<subScript attach="left" box="[1022,1058,1281,1301]" fontSize="8" pageId="20" pageNumber="21">4+5</subScript>
|
||
. Base of M and stem of M
|
||
<subScript attach="left" box="[1369,1381,1281,1301]" fontSize="8" pageId="20" pageNumber="21">4</subScript>
|
||
both shorter than r-m cross vein, ratio of r-m to M-petiole 1:2.6. M
|
||
<subScript attach="left" box="[984,1019,1350,1370]" fontSize="8" pageId="20" pageNumber="21">1+2</subScript>
|
||
inconspicuous, slightly longer than medial fork, M
|
||
<subScript attach="left" box="[1016,1028,1387,1407]" fontSize="8" pageId="20" pageNumber="21">1</subScript>
|
||
and M
|
||
<subScript attach="left" box="[1118,1130,1387,1407]" fontSize="8" pageId="20" pageNumber="21">2</subScript>
|
||
slightly divergent; cubital fork very long, with origin basal to origin of M
|
||
<subScript attach="left" box="[1396,1432,1424,1444]" fontSize="8" pageId="20" pageNumber="21">1+2</subScript>
|
||
. CuP inconspicuous or not preserved.
|
||
</paragraph>
|
||
<paragraph blockId="20.[840,1441,1027,1668]" pageId="20" pageNumber="21">
|
||
<emphasis bold="true" box="[840,1139,1482,1507]" pageId="20" pageNumber="21">Abdomen and genitalia.</emphasis>
|
||
Female, length
|
||
<quantity box="[1329,1437,1482,1507]" metricMagnitude="-3" metricUnit="m" metricValue="1.58" pageId="20" pageNumber="21" unit="mm" value="1.58">1.58 mm</quantity>
|
||
, brown; basal tergites wider than long. Tergites 3–5 with short setae at posterior margin; cercus twosegmented (Figure 17.3).
|
||
</paragraph>
|
||
<paragraph blockId="20.[840,1441,1027,1668]" box="[840,1140,1610,1635]" pageId="20" pageNumber="21">
|
||
<emphasis bold="true" box="[840,952,1610,1635]" pageId="20" pageNumber="21">
|
||
<typeStatus box="[840,948,1610,1635]" pageId="20" pageNumber="21">Allotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Male unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="discussion">
|
||
<paragraph blockId="20.[840,1441,1027,1668]" box="[840,1435,1642,1668]" pageId="20" pageNumber="21">
|
||
<emphasis bold="true" box="[840,1077,1642,1668]" pageId="20" pageNumber="21">Syncompressions.</emphasis>
|
||
Thysanoptera (1),
|
||
<taxonomicName authorityName="Linnaeus" authorityYear="1758" box="[1302,1388,1642,1667]" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="order">Diptera</taxonomicName>
|
||
(1).
|
||
</paragraph>
|
||
<paragraph blockId="20.[840,953,1690,1715]" box="[840,953,1690,1715]" pageId="20" pageNumber="21">
|
||
<heading bold="true" box="[840,953,1690,1715]" fontSize="10" level="3" pageId="20" pageNumber="21" reason="2">
|
||
<emphasis bold="true" box="[840,953,1690,1715]" pageId="20" pageNumber="21">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="20.[840,1441,1738,1891]" lastBlockId="21.[192,796,168,1899]" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
|
||
The family has 92 genera and 2,455 species, with a very complex taxonomy.
|
||
<taxonomicName authority="Meigen, 1803" authorityName="Meigen" authorityYear="1803" class="Insecta" family="Sciaridae" genus="Sciara" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1253,1329,1770,1795]" italics="true" pageId="20" pageNumber="21">Sciara</emphasis>
|
||
<bibRefCitation author="Meigen, J. W." journalOrPublisher="Magazin fur Insektenkunde" pageId="20" pageNumber="21" pagination="259 - 281" part="2" refId="ref39950" refString="Meigen, J. W. 1803. Versuch einer neuen Gattungs-Eintheilung der europaischen zweiflugligen Insekten. Magazin fur Insektenkunde, 2: 259 - 281." title="Versuch einer neuen Gattungs-Eintheilung der europaischen zweiflugligen Insekten" type="journal article" year="1803">Meigen, 1803</bibRefCitation>
|
||
</taxonomicName>
|
||
, with approximately 700 extant species, is one of the most species-rich genera in the class
|
||
<taxonomicName authorityName="Linnaeus" authorityYear="1758" box="[840,926,1866,1891]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="21" phylum="Arthropoda" rank="class">Insecta</taxonomicName>
|
||
— although many of these species may be misplaced.
|
||
<taxonomicName authorityName="Meigen" authorityYear="1803" box="[327,403,168,194]" class="Insecta" family="Sciaridae" genus="Sciara" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="24" phylum="Arthropoda" rank="genus">Sciara</taxonomicName>
|
||
is the largest genus in the family
|
||
<taxonomicName authorityName="Billberg" authorityYear="1820" box="[192,303,200,226]" class="Insecta" family="Sciaridae" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="family">Sciaridae</taxonomicName>
|
||
and constitutes nearly 30% of the family’s extant species.
|
||
<taxonomicName authorityName="Meigen" authorityYear="1803" box="[427,503,232,257]" class="Insecta" family="Sciaridae" genus="Sciara" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[427,503,232,257]" italics="true" pageId="21" pageNumber="22">Sciara</emphasis>
|
||
</taxonomicName>
|
||
has obviously served as a default assignment for poorly preserved fossils; 89 of all 168 fossil
|
||
<taxonomicName authorityName="Billberg" authorityYear="1820" box="[432,543,296,322]" class="Insecta" family="Sciaridae" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="family">Sciaridae</taxonomicName>
|
||
are assigned to the genus (
|
||
<bibRefCitation accessDate="invalid" author="PBDB" box="[279,434,329,354]" pageId="21" pageNumber="22" publicationUrl="https://paleobiodb.org//" refId="ref41742" refString="PBDB, 2018. Paleobiology Database. https: // paleobiodb. org / / (accessed April 27, 2018 and other dates)." title="Paleobiology Database" type="url" year="2018">PBDB, 2018</bibRefCitation>
|
||
). Species epithets such as
|
||
<emphasis italics="true" pageId="21" pageNumber="22">rottensis</emphasis>
|
||
,
|
||
<emphasis box="[284,414,360,385]" italics="true" pageId="21" pageNumber="22">defectuosa</emphasis>
|
||
,
|
||
<emphasis box="[437,676,361,386]" italics="true" pageId="21" pageNumber="22">deperdita, diabolica</emphasis>
|
||
,
|
||
<emphasis box="[699,785,360,385]" italics="true" pageId="21" pageNumber="22">difficilis</emphasis>
|
||
,
|
||
<emphasis box="[192,288,393,418]" italics="true" pageId="21" pageNumber="22">ignorata</emphasis>
|
||
, etc. may reflect authors’ frustrations in the identification of their fossil specimens (
|
||
<bibRefCitation author="Heyden, C. H. G., von" journalOrPublisher="Palaeontographica" pageId="21" pageNumber="22" pagination="237 - 266" part="17" refId="ref38076" refString="Heyden, C. H. G., von. 1870. Fossile Dipteren aus der Braunkohle von Rott im Siebengebirge. Palaeontographica, 17: 237 - 266." title="Fossile Dipteren aus der Braunkohle von Rott im Siebengebirge" type="journal article" year="1870">Heyden, 1870</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Scudder, S. H." box="[269,447,456,482]" journalOrPublisher="Geological Survey of Canada, Montreal" pageId="21" pageNumber="22" refId="ref43484" refString="Scudder, S. H. 1878. Additions to the Insect Fauna of the Tertiary Beds at Quesnel (British Colombia). Report of Progress of the Geological Survey of Canada 1876 - 1877. Geological Survey of Canada, Montreal." type="journal article" volumeTitle="Additions to the Insect Fauna of the Tertiary Beds at Quesnel (British Colombia). Report of Progress of the Geological Survey of Canada 1876 - 1877" year="1878">Scudder, 1878</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Meunier, F." box="[462,655,457,482]" journalOrPublisher="Annales de la Societe Scientifique de Bruxelles" pageId="21" pageNumber="22" pagination="12 - 92" part="28" refId="ref40296" refString="Meunier, F. 1904 b. Monographie des Cecidomyidae, des Sciaridae, des Mycetophilidae et des Chironomidae de l'ambre de la Baltique. Annales de la Societe Scientifique de Bruxelles, 28: 12 - 92." title="Monographie des Cecidomyidae, des Sciaridae, des Mycetophilidae et des Chironomidae de l'ambre de la Baltique" type="journal article" year="1904">Meunier, 1904b</bibRefCitation>
|
||
). The presence or absence of macrotrichia on veins M and/or Cu is an essential morphological character in the identification of
|
||
<taxonomicName authorityName="Meigen" authorityYear="1803" box="[388,464,552,577]" class="Insecta" family="Sciaridae" genus="Sciara" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[388,464,552,577]" italics="true" pageId="21" pageNumber="22">Sciara</emphasis>
|
||
</taxonomicName>
|
||
. A common question with fossil specimens is whether the macrothrichia were ever present or whether they were just not preserved. The near universal presence of these structures on the costal and radial veins however, should serve as an internal control. Neither macrotrichia nor their sockets are preserved in veins C, R
|
||
<subScript attach="left" box="[211,223,787,807]" fontSize="8" pageId="21" pageNumber="22">1</subScript>
|
||
and R
|
||
<subScript attach="left" box="[319,355,787,807]" fontSize="8" pageId="21" pageNumber="22">
|
||
4+
|
||
<quantity box="[343,392,777,807]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="21" pageNumber="22" unit="in" value="5.0">5 in</quantity>
|
||
</subScript>
|
||
the three species re-examined below.
|
||
</paragraph>
|
||
<paragraph blockId="21.[192,796,168,1899]" pageId="21" pageNumber="22">
|
||
While this new specimen does not appear to belong to the genus
|
||
<taxonomicName authorityName="Meigen" authorityYear="1803" box="[439,519,877,903]" class="Insecta" family="Sciaridae" genus="Sciara" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="24" phylum="Arthropoda" rank="genus">Sciara</taxonomicName>
|
||
, due to the absence of macrotrichia on veins M, the fact that it is a female and the inability to determine morphological characters such as the absence or presence of a neck on the flagellomeres (Figure 17.1), make it difficult to assign the specimen to an extant genus. For an impression fossil, preservation of this specimen is exceptional, but although the claws do not appear to have teeth and the labial palps do not appear to have setae, their absence can not be definitively established.
|
||
<emphasis box="[348,494,1197,1222]" italics="true" pageId="21" pageNumber="22">Dolichociara</emphasis>
|
||
and
|
||
<taxonomicName authorityName="Rudzinski" authorityYear="1997" box="[566,735,1197,1222]" class="Insecta" family="Sciaridae" genus="Angustosciara" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[566,735,1197,1222]" italics="true" pageId="21" pageNumber="22">Angustosciara</emphasis>
|
||
</taxonomicName>
|
||
can be eliminated due to the absence of macrotrichia on the M veins;
|
||
<emphasis box="[379,525,1261,1286]" italics="true" pageId="21" pageNumber="22">Sciarotrichia</emphasis>
|
||
can be eliminated due to the length of the terminal segments of its labial palps;
|
||
<taxonomicName box="[273,384,1325,1350]" class="Insecta" family="Sciaridae" genus="Edidapus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[273,384,1325,1350]" italics="true" pageId="21" pageNumber="22">Edidapus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Johannsen" authorityYear="1912" box="[449,529,1325,1350]" class="Insecta" family="Sciaridae" genus="Pnyxia" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[449,529,1325,1350]" italics="true" pageId="21" pageNumber="22">Pnyxia</emphasis>
|
||
</taxonomicName>
|
||
can be eliminated as they lack a patch of anteroapical setae on their foretibia and the presence of an attenuated katepisternum;
|
||
<taxonomicName box="[386,540,1421,1446]" class="Insecta" family="Sciaridae" genus="Scatopsciara" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[386,540,1421,1446]" italics="true" pageId="21" pageNumber="22">Scatopsciara</emphasis>
|
||
</taxonomicName>
|
||
can be eliminated due to R
|
||
<subScript attach="left" box="[296,308,1464,1484]" fontSize="8" pageId="21" pageNumber="22">1</subScript>
|
||
less than half the length of R,
|
||
<taxonomicName authorityName="Enderlein" authorityYear="1911" box="[680,792,1454,1479]" class="Insecta" family="Sciaridae" genus="Euricrium" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[680,792,1454,1479]" italics="true" pageId="21" pageNumber="22">Euricrium</emphasis>
|
||
</taxonomicName>
|
||
can be eliminated as it has an M fork bell-shaped, wider basally than distally;
|
||
<taxonomicName authorityName="Winnertz" authorityYear="1867" box="[525,629,1522,1547]" class="Insecta" family="Sciaridae" genus="Bradysia" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[525,629,1522,1547]" italics="true" pageId="21" pageNumber="22">Bradysia</emphasis>
|
||
</taxonomicName>
|
||
is eliminated as its foretibia have two apical spurs;
|
||
<taxonomicName authorityName="Lengersdorf" authorityYear="1931" class="Insecta" family="Sciaridae" genus="Rhynchomegalosphys" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="21" pageNumber="22">Rhynchomegalosphys</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Enderlein" authorityYear="1911" box="[452,639,1586,1612]" class="Insecta" family="Sciaridae" genus="Scythropochroa" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[452,639,1586,1612]" italics="true" pageId="21" pageNumber="22">Scythropochroa</emphasis>
|
||
</taxonomicName>
|
||
can be eliminated as M+CuA is much longer than bm-m;
|
||
<taxonomicName authorityName="Enderlein" authorityYear="1911" box="[192,349,1650,1675]" class="Insecta" family="Sciaridae" genus="Ceratiosciara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[192,349,1650,1675]" italics="true" pageId="21" pageNumber="22">Ceratiosciara</emphasis>
|
||
</taxonomicName>
|
||
is eliminated as it has strongly shortened convex flagellomeres;
|
||
<taxonomicName authorityName="Coquillett" authorityYear="1896" box="[550,677,1682,1707]" class="Insecta" family="Sciaridae" genus="Eugnoriste" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[550,677,1682,1707]" italics="true" pageId="21" pageNumber="22">Eugnoriste</emphasis>
|
||
</taxonomicName>
|
||
is eliminated due to its greatly elongated mouthparts;
|
||
<taxonomicName box="[192,285,1746,1772]" class="Insecta" family="Sciaridae" genus="Cratyna" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[192,285,1746,1772]" italics="true" pageId="21" pageNumber="22">Cratyna</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Mohrig & Roschmann" authorityYear="2005" box="[344,492,1746,1771]" class="Insecta" family="Sciaridae" genus="Archicratyna" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[344,492,1746,1771]" italics="true" pageId="21" pageNumber="22">Archicratyna</emphasis>
|
||
</taxonomicName>
|
||
are eliminated due to the presence of two midtibial spurs;
|
||
<taxonomicName authorityName="Schmitz" authorityYear="1918" box="[595,736,1778,1803]" class="Insecta" family="Sciaridae" genus="Hyperlasion" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[595,736,1778,1803]" italics="true" pageId="21" pageNumber="22">Hyperlasion</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Frey" authorityYear="1942" box="[192,347,1810,1835]" class="Insecta" family="Sciaridae" genus="Cosmosciara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[192,347,1810,1835]" italics="true" pageId="21" pageNumber="22">Cosmosciara</emphasis>
|
||
</taxonomicName>
|
||
are eliminated due to the presence of an attenuated katepisternum (Menzel and Smith, 2017). This does not mean that the specimen does
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3990331" ID-Zenodo-Dep="3990331" httpUri="https://zenodo.org/record/3990331/files/figure.png" pageId="21" pageNumber="22" startId="21.[845,935,881,904]" targetBox="[844,1439,165,859]" targetPageId="21">
|
||
<paragraph blockId="21.[845,1439,881,1024]" pageId="21" pageNumber="22">
|
||
<emphasis bold="true" box="[845,986,881,904]" pageId="21" pageNumber="22">FIGURE 18.</emphasis>
|
||
<taxonomicName authority="Cockerell 1916" authorityName="Cockerell" authorityYear="1916" box="[1004,1435,881,904]" class="Insecta" family="Sciaridae" genus="Sciara" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="florissantensis">
|
||
<emphasis box="[1004,1242,881,904]" italics="true" pageId="21" pageNumber="22">Sciara florissantensis</emphasis>
|
||
Cockerell 1916
|
||
</taxonomicName>
|
||
, USNM 61995.
|
||
<emphasis bold="true" box="[1013,1026,910,933]" pageId="21" pageNumber="22">1</emphasis>
|
||
, Habitus;
|
||
<emphasis bold="true" box="[1145,1158,910,933]" pageId="21" pageNumber="22">2</emphasis>
|
||
, Head and antennae;
|
||
<emphasis bold="true" box="[1418,1431,910,933]" pageId="21" pageNumber="22">3</emphasis>
|
||
, Right forewing. Long arrow denotes 1
|
||
<superScript attach="left" box="[1250,1265,940,958]" fontSize="7" pageId="21" pageNumber="22">st</superScript>
|
||
abscissa of Rs; the arrowhead points to remnants of the r-m crossvein. Scale bar equals 1.0 mm (
|
||
<emphasis bold="true" box="[1125,1138,1001,1024]" pageId="21" pageNumber="22">1</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[1152,1165,1001,1024]" pageId="21" pageNumber="22">3</emphasis>
|
||
), 0.25 mm (
|
||
<emphasis bold="true" box="[1294,1307,1001,1024]" pageId="21" pageNumber="22">2</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="21.[840,1441,1077,1476]" pageId="21" pageNumber="22">
|
||
not belong to an extant genus. However, until additional specimens are collected that may assist in such an assignment, the generic epithet
|
||
<taxonomicName authorityName="Greenwalt" authorityYear="2019" class="Insecta" family="Scatopsidae" genus="Eosciarites" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="42" phylum="Arthropoda" rank="genus">Eosciarites</taxonomicName>
|
||
is provided.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |