226 lines
16 KiB
XML
226 lines
16 KiB
XML
<document ID-DOI="10.26879/891" ID-Zenodo-Dep="3990295" approvalRequired="202" approvalRequired_for_document="2" approvalRequired_for_illustrations="2" approvalRequired_for_textStreams="175" approvalRequired_for_treatments="23" checkinTime="1597772865363" checkinUser="torsten" docAuthor="Dale E. Greenwalt, Daniel J. Bickel, Peter H. Kerr, Gregory R. Curler, Brian V. Brown, Herman de Jong, Scott J. Fitzgerald, Torsten Dikow, Michal Tkoč, Christian Kehlmaier & Dalton De Souza Amorim" docDate="2019" docId="571F246BFFB8FFA51067FC70ADDFC5C4" docLanguage="en" docName="greenwalt_etal_2019.imf" docOrigin="Paleontologia Electronica 22 (2), No. 50" docStyle="DocumentStyle{}" docTitle="Sciara lacoei Cockerell 1915" docType="treatment" docVersion="4" lastPageId="23" lastPageNumber="24" masterDocId="AB265C13FFAEFFB21318FF0FAF31C65B" masterDocTitle="Diptera of the middle Eocene Kishenehn Formation. I. Documenting of diversity at the family level" masterLastPageNumber="56" masterPageNumber="1" pageId="22" pageNumber="23" updateTime="1636465287172" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Diptera of the middle Eocene Kishenehn Formation. I. Documenting of diversity at the family level</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Dale E. Greenwalt</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Daniel J. Bickel</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Peter H. Kerr</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Gregory R. Curler</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Brian V. Brown</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Herman de Jong</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Scott J. Fitzgerald</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Torsten Dikow</mods:namePart>
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<mods:nameIdentifier type="LSID">F8869067-4618-4CCE-960C-E8A107F162FB</mods:nameIdentifier>
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<mods:nameIdentifier type="ORCID">0000-0003-4816-2909</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Michal Tkoč</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Christian Kehlmaier</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Dalton De Souza Amorim</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Paleontologia Electronica</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>22</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>2</mods:number>
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</mods:detail>
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<mods:detail type="numero">
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<mods:number>50</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>56</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.26879/891</mods:identifier>
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<mods:identifier type="Zenodo-Dep">3990295</mods:identifier>
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<mods:identifier type="ZooBank">A6C79E56-3CCC-484E-B6AF-EAEEE1695FF6</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:571F246BFFB8FFA51067FC70ADDFC5C4" httpUri="http://treatment.plazi.org/id/571F246BFFB8FFA51067FC70ADDFC5C4" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
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<subSubSection pageId="22" pageNumber="23" type="nomenclature">
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<paragraph blockId="22.[840,1441,326,1188]" pageId="22" pageNumber="23">
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<taxonomicName authority="Cockerell, 1915" authorityName="Cockerell" authorityYear="1915" box="[895,1244,895,921]" class="Insecta" family="Sciaridae" genus="Sciara" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="lacoei">
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<emphasis box="[895,1048,895,920]" italics="true" pageId="22" pageNumber="23">Sciara lacoei</emphasis>
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<bibRefCitation author="Cockerell, T. D. A." box="[1055,1244,895,921]" journalOrPublisher="Proceedings of the United States National Museum" pageId="22" pageNumber="23" pagination="469 - 499" part="49" refId="ref35367" refString="Cockerell, T. D. A. 1915. British fossil insects. Proceedings of the United States National Museum, 49: 469 - 499." title="British fossil insects" type="journal article" year="1915">Cockerell, 1915</bibRefCitation>
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</taxonomicName>
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, reassigned Figure 19.2-3
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</paragraph>
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</subSubSection>
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<materialsCitation collectionCode="USNM" collectorName="T.D.A. Cockerell" country="United States of America" location="National Museum of Natural History" municipality="Eocene" pageId="22" pageNumber="23" specimenCode="USNM 61436" specimenCount="1" stateProvince="Washington" typeStatus="Holotype">
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<subSubSection pageId="22" pageNumber="23" type="materials_examined">
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<paragraph blockId="22.[840,1441,326,1188]" pageId="22" pageNumber="23">
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<emphasis bold="true" box="[840,1081,970,995]" pageId="22" pageNumber="23">Material examined.</emphasis>
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<typeStatus box="[1092,1202,970,995]" pageId="22" pageNumber="23">Holotype</typeStatus>
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,
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<specimenCode box="[1215,1382,970,996]" collectionCode="USNM" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="22" pageNumber="23" type="Museum">
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<collectionCode box="[1215,1293,970,996]" pageId="22" pageNumber="23">USNM</collectionCode>
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61436
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</specimenCode>
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, and specimen 7583, both housed in the
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<location LSID="urn:lsid:plazi:treatment:571F246BFFB8FFA51067FC70ADDFC5C4:DA69C3A6FFB8FFA41626FCE5AB99C278" country="United States of America" name="National Museum of Natural History" pageId="22" pageNumber="23" stateProvince="Washington">National Museum of Natural History</location>
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,
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<collectingRegion box="[1213,1357,1034,1059]" country="United States of America" name="Washington" pageId="22" pageNumber="23">Washington</collectingRegion>
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,
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<collectorName box="[1379,1436,1034,1060]" name="T.D.A. Cockerell" pageId="22" pageNumber="23">D.C.</collectorName>
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, USA.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="22" pageNumber="23" type="distribution">
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<paragraph blockId="22.[840,1441,326,1188]" pageId="22" pageNumber="23">
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<emphasis bold="true" box="[840,1029,1098,1123]" pageId="22" pageNumber="23">
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<typeStatus box="[840,900,1098,1123]" pageId="22" pageNumber="23">Type</typeStatus>
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horizon.
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</emphasis>
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<location LSID="urn:lsid:plazi:treatment:571F246BFFB8FFA51067FC70ADDFC5C4:DA69C3A6FFB8FFA41707FB45ABCFC238" box="[1055,1278,1098,1123]" country="United Kingdom" municipality="Eocene" name="Bembridge Marls" pageId="22" pageNumber="23">Bembridge Marls</location>
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,
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<collectingCountry box="[1306,1347,1098,1123]" name="United Kingdom" pageId="22" pageNumber="23" value="United Kingdom">UK</collectingCountry>
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, latest
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<collectingMunicipality box="[840,930,1130,1155]" pageId="22" pageNumber="23">Eocene</collectingMunicipality>
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</paragraph>
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<paragraph blockId="22.[840,1441,326,1188]" box="[840,1375,1162,1188]" pageId="22" pageNumber="23">
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<emphasis bold="true" box="[840,1005,1162,1187]" pageId="22" pageNumber="23">
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<typeStatus box="[840,900,1162,1187]" pageId="22" pageNumber="23">Type</typeStatus>
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locality.
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</emphasis>
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<location LSID="urn:lsid:plazi:treatment:571F246BFFB8FFA51067FC70ADDFC5C4:DA69C3A6FFB8FFA410ECFB85ABBCC2F8" box="[1012,1165,1162,1188]" country="United Kingdom" name="Gurnard Bay" pageId="22" pageNumber="23" stateProvince="Isle of Wight">Gurnard Bay</location>
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,
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<collectingRegion box="[1176,1328,1162,1187]" country="United Kingdom" name="Isle of Wight" pageId="22" pageNumber="23">Isle of Wight</collectingRegion>
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,
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<collectingCountry box="[1338,1375,1162,1187]" name="United Kingdom" pageId="22" pageNumber="23" value="United Kingdom">UK</collectingCountry>
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</paragraph>
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</subSubSection>
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</materialsCitation>
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<subSubSection lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23" type="discussion">
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<paragraph blockId="22.[840,953,1210,1235]" box="[840,953,1210,1235]" pageId="22" pageNumber="23">
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<heading bold="true" box="[840,953,1210,1235]" fontSize="10" level="3" pageId="22" pageNumber="23" reason="2">
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<emphasis bold="true" box="[840,953,1210,1235]" pageId="22" pageNumber="23">Remarks</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="22.[840,1441,1258,1906]" pageId="22" pageNumber="23">This fossil is of the dorsal aspect of the insect (Figure 19.2). The specimen is an impression fossil with the head, with well-preserved and clearly separated eyes, having formed a deep hemispherical depression in fine-grained mud; the face of the head contacted the bottom of the depression. Poorly preserved remnants of the basal antennomeres lie on the surface of the fossil above the hemispherical depression. The posterior portion of the head is not preserved and it is therefore impossible to determine whether or not an eye bridge is present.</paragraph>
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<paragraph blockId="22.[840,1441,1258,1906]" lastBlockId="23.[192,793,169,1205]" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
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C, R
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<subScript attach="left" box="[959,971,1653,1673]" fontSize="8" pageId="22" pageNumber="23">1</subScript>
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and R
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<subScript attach="left" box="[1066,1102,1653,1673]" fontSize="8" pageId="22" pageNumber="23">4+5</subScript>
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are clearly present; Sc is absent (there is a gouge in the fossil at that point). R
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<subScript attach="left" box="[859,871,1722,1742]" fontSize="8" pageId="22" pageNumber="23">1</subScript>
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and R
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<subScript attach="left" box="[952,988,1722,1742]" fontSize="8" pageId="22" pageNumber="23">4+5</subScript>
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extend to 0.48 and 0.89 of the wing’s total length. The cubital veins are well preserved, but the medial veins are not. The fossil contains several creases that confound identification of these veins. In Cockerell’s figure, there is a short apical portion of M
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<subScript attach="left" box="[1063,1075,1886,1906]" fontSize="8" pageId="22" pageNumber="23">2</subScript>
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preserved, and this vein does indeed appear to be present in the fossil. M
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<subScript attach="left" box="[773,785,179,199]" fontSize="8" pageId="23" pageNumber="24">1</subScript>
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, labeled with a “?” in figure 18.3, if not a crease, takes an uncharacteristically straight path to the margin; the base of the medial fork appears to be too basal for
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<taxonomicName authorityName="Billberg" authorityYear="1820" box="[361,477,301,327]" class="Insecta" family="Sciaridae" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="family">Sciaridae</taxonomicName>
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. As discussed above, M
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<subScript attach="left" box="[780,792,312,332]" fontSize="8" pageId="23" pageNumber="24">4</subScript>
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originating very basally in the wing in this species is seen in some
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<taxonomicName box="[389,568,370,396]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="family">Cecidomyiidae</taxonomicName>
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,
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<taxonomicName authorityName="Jaschhof & Didham" authorityYear="2002" box="[580,792,370,395]" class="Insecta" family="Rangomaramidae" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="family">Rangomaramidae</taxonomicName>
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and
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<taxonomicName authorityName="Newman" authorityYear="1834" box="[250,432,402,427]" class="Insecta" family="Mycetophilidae" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="family">Mycetophilidae</taxonomicName>
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. The very long R
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<subScript attach="left" box="[653,665,413,433]" fontSize="8" pageId="23" pageNumber="24">5</subScript>
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also suggests that
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<taxonomicName authorityName="Cockerell" authorityYear="1915" box="[318,422,439,464]" class="Insecta" family="Sciaridae" genus="Sciara" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="lacoei">
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<emphasis box="[318,422,439,464]" italics="true" pageId="23" pageNumber="24">S. lacoei</emphasis>
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</taxonomicName>
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is not a sciarid. We, therefore, assign this specimen to Sciaroidea incertae sedis.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="23" pageNumber="24" type="materials_examined">
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<paragraph blockId="23.[192,793,169,1205]" pageId="23" pageNumber="24">
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A second specimen (#7583, also labelled
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<collectionCode box="[192,270,535,561]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="23" pageNumber="24" type="Museum">USNM</collectionCode>
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61436) is not a type but is labelled
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<taxonomicName authorityName="Cockerell" authorityYear="1915" class="Insecta" family="Sciaridae" genus="Sciara" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="lacoei">
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<emphasis italics="true" pageId="23" pageNumber="24">Sciara lacoei</emphasis>
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</taxonomicName>
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. This specimen (Figure 19.3) is
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<quantity box="[639,727,567,592]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="23" pageNumber="24" unit="mm" value="1.7">1.7 mm</quantity>
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long, although the end of the abdomen is not visible; it contains no structures of value other than the wing. The preserved portion of the wing (the apical portion is missing) is approximately
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<quantity box="[583,673,695,720]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="23" pageNumber="24" unit="mm" value="1.4">1.4 mm</quantity>
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in length, the same length as that of the
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<typeStatus box="[560,665,727,752]" pageId="23" pageNumber="24">holotype</typeStatus>
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. R
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<subScript attach="left" box="[696,708,738,758]" fontSize="8" pageId="23" pageNumber="24">1</subScript>
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is preserved, but the end of R
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<subScript attach="left" box="[488,524,775,795]" fontSize="8" pageId="23" pageNumber="24">4+5</subScript>
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is not. Apical portions of M
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<subScript attach="left" box="[247,259,811,831]" fontSize="8" pageId="23" pageNumber="24">1</subScript>
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and M
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<subScript attach="left" box="[348,360,811,831]" fontSize="8" pageId="23" pageNumber="24">2</subScript>
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are preserved, but the base of the fork is not. This specimen appears to be a sciarid, although it is impossible to assign this specimen to
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<taxonomicName authorityName="Meigen" authorityYear="1803" box="[192,267,902,927]" class="Insecta" family="Sciaridae" genus="Sciara" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
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<emphasis box="[192,267,902,927]" italics="true" pageId="23" pageNumber="24">Sciara</emphasis>
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</taxonomicName>
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; we assign it to
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<taxonomicName authorityName="Billberg" authorityYear="1820" box="[459,570,902,928]" class="Insecta" family="Sciaridae" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="family">Sciaridae</taxonomicName>
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incertae sedis.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |