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<document id="F9DE1EBDFA13592A279A10E62E19BBCA" ID-CLB-Dataset="21965" ID-DOI="10.3853/j.2201-4349.71.2019.1677" ID-GBIF-Dataset="9a9b8fd9-2aaa-4f81-b833-24bc5c400ba8" ID-ISSN="2201-4349" ID-Zenodo-Dep="3837977" ID-ZooBank="124BD25A-7712-4EC6-9A1E-48FC9C23B513" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590071897212" checkinUser="felipe" docAuthor="Smith, Graeme B., Mitchell, Andrew, Lee, Timothy R. C. &amp; Espinasa, Luis" docDate="2019" docId="3D3987FDFFA6983366B2EAC4FD7EFD22" docLanguage="en" docName="RecAustMus.71.1.1-32.pdf.imf" docOrigin="Records of the Australian Museum 71 (1)" docStyle="DocumentStyle:997BA24B490466A2F99594A367FA147F.3:RecAustMus.2019-.journal_article" docStyleId="997BA24B490466A2F99594A367FA147F" docStyleName="RecAustMus.2019-.journal_article" docStyleVersion="3" docTitle="Heterolepisma cooloola Smith &amp; Mitchell &amp; Lee &amp; Espinasa 2019, sp. nov." docType="treatment" docVersion="9" lastPageNumber="29" masterDocId="C100FF85FFB3982E653FEA5DFFB1FF86" masterDocTitle="DNA Barcoding and Integrative Taxonomy of the Heterolepisma sclerophylla species complex (Zygentoma: Lepismatidae: Heterolepismatinae) and the Description of Two New Species" masterLastPageNumber="32" masterPageNumber="1" pageNumber="22" updateTime="1698840649716" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="1F3E7802B9EC0E7D0CB31AFCD74052DC">DNA Barcoding and Integrative Taxonomy of the Heterolepisma sclerophylla species complex (Zygentoma: Lepismatidae: Heterolepismatinae) and the Description of Two New Species</mods:title>
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<taxonomicName id="72904D68FFA6983B66B2EAC4FB03FF32" authority="Smith &amp; Mitchell &amp; Lee &amp; Espinasa, 2019" authorityName="Smith &amp; Mitchell &amp; Lee &amp; Espinasa" authorityYear="2019" box="[909,1202,153,180]" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="cooloola" status="sp. nov.">
<emphasis id="87E4EAF9FFA6983B66B2EAC4FB03FF32" bold="true" box="[909,1202,153,180]" italics="true" pageId="21" pageNumber="22">Heterolepisma cooloola</emphasis>
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<figureCitation id="2DAB2A6EFFA6983B612FEA92FB26FF6C" box="[1040,1175,207,234]" captionStart-0="Figure 37" captionStart-1="Figures 3848" captionStart-2="Figures 4959" captionStart-3="Figures 6065" captionStart-4="Figures 6677" captionStart-5="Figures 7885" captionStartId-0="21.[794,863,1954,1976]" captionStartId-1="22.[166,243,1805,1827]" captionStartId-2="23.[121,198,1517,1539]" captionStartId-3="24.[166,243,1543,1565]" captionStartId-4="25.[121,198,1620,1642]" captionStartId-5="26.[166,243,1315,1337]" captionTargetBox-0="[794,1421,971,1946]" captionTargetBox-1="[174,1460,196,1772]" captionTargetBox-2="[134,1417,159,1496]" captionTargetBox-3="[177,1452,156,1521]" captionTargetBox-4="[133,1412,157,1605]" captionTargetBox-5="[177,1457,158,1295]" captionTargetId-0="figure@21.[794,1421,971,1946]" captionTargetId-1="figure@22.[166,1466,185,1777]" captionTargetId-2="figure@23.[121,1421,151,1514]" captionTargetId-3="figure@24.[166,1466,151,1532]" captionTargetId-4="figure@25.[121,1421,151,1615]" captionTargetId-5="figure@26.[166,1466,151,1313]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionTargetPageId-2="23" captionTargetPageId-3="24" captionTargetPageId-4="25" captionTargetPageId-5="26" captionText-0="Figure 37. Heterolepisma cooloola sp. nov. from leaf litter at Carlo Point." captionText-1="Figures 3848. Heterolepisma cooloola sp. nov. holotype ♀, unless indicated otherwise by specimen number(38) habitus (K.377754); (39) posterior comb of pronotum with scale; (40) head; (41) scape, pedicel and basal intervals of flagellum, from above; (42) idem, from below; (43) antenna, most distal surviving interval; (44) mandible; (45) idem, detail of molar and incisor regions; (46) maxilla, only larger setae of palp illustrated; (47) idem, lacinia and galea (QM 207012); (48) ultimate article of maxillary palp of ♂ (QM 207012). Scale bars = 0.1 mm unless otherwise indicated." captionText-2="Figures 4959. Heterolepisma cooloola sp. nov. holotype ♀ (49) labium, only large setae of palp illustrated; (50) idem, ultimate article of palp; (51) pronotum; (52) idem, right margin; (53) idem, detail of right anterior trichobothrial area; (54) idem, detail of left posterior trichobothrial area; (55) lateral margin of mesonotum; (56) idem, posterior trichobothrial area; (57) idem, anterior trichobothrial area; (58) idem, right posterior comb; (59) lateral margin of metanotum. Scale bars = 0.1 mm." captionText-3="Figures 6065. Heterolepisma cooloola sp. nov. holotype ♀ (60) presternum, prothoracic sternum and PI; (61) mesothoracic sternum and PII; (62) apex of mesothoracic sternum; (63) metathoracic sternum and PIII; (64) apex of metathoracic sternum; (65) pretarsus of PIII. Scale bars = 0.1 mm." captionText-4="Figures 6677. Heterolepisma cooloola sp. nov. holotype ♀, unless otherwise indicated by specimen number (66) urotergite IV; (67) urotergite V, right lateral comb; (68) idem, left sublateral comb; (69) idem,left submedial comb; (70) right side of urotergiteVIII (K.261189); (71) infralateral comb of urotergite IX; (72) urotergite X; (73) urotergite X of paratype (QM 207012); (74) urosternite IV; (75) posterior comb of urosternite V; (76) urosternites VII, VIII, IX and ovipositor; (77) base of stylus VII. Scale bars = 0.1 mm." captionText-5="Figures 7885. Heterolepisma cooloola sp. nov. holotype ♀ unless otherwise indicated by specimen number (78) stylet IX (QM 207012); (79) apex of anterior gonapophysis; (80) apex of posterior gonapophysis; (81) bases of terminal filaments; (82) most distal surviving divisions of cerci; (83) most distal surviving divisions of median filament; (84) coxites IX, styli and penis of male (QM 207012); (85) paramere. Scale bars = 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3837999" figureDoi-1="http://doi.org/10.5281/zenodo.3838001" figureDoi-2="http://doi.org/10.5281/zenodo.3838003" figureDoi-3="http://doi.org/10.5281/zenodo.3838007" figureDoi-4="http://doi.org/10.5281/zenodo.3838009" figureDoi-5="http://doi.org/10.5281/zenodo.3838011" httpUri-0="https://zenodo.org/record/3837999/files/figure.png" httpUri-1="https://zenodo.org/record/3838001/files/figure.png" httpUri-2="https://zenodo.org/record/3838003/files/figure.png" httpUri-3="https://zenodo.org/record/3838007/files/figure.png" httpUri-4="https://zenodo.org/record/3838009/files/figure.png" httpUri-5="https://zenodo.org/record/3838011/files/figure.png" pageId="21" pageNumber="22">Figs 3785</figureCitation>
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(
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207011 on two slides) QLD: Cooloola, Freshwater track rainforest patch,
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asl
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, Graeme Smith
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<typeStatus id="6A2B8849FFA6983B614AEB68FB6DFECC" box="[1141,1244,309,330]" pageId="21" pageNumber="22" type="paratype">Paratypes</typeStatus>
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:
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,
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specimens including
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<collectionCode id="D381AE2EFFA6983B61EEEB0DFB48FEE3" box="[1233,1273,336,357]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="21" pageNumber="22" type="Museum">QM</collectionCode>
207012 on two slides) same data as holotype
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<collectionCode id="D381AE2EFFA6983B61D4EB37FA91FEF9" box="[1259,1312,362,383]" pageId="21" pageNumber="22">AMS</collectionCode>
K.261176, K.261177 on two slides) same data as holotype
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0.74) (
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K.261178, K.261179 on two slides) same data as holotype
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<collectionCode id="D381AE2EFFA6983B6116EBE7FBD1FE49" box="[1065,1120,442,463]" pageId="21" pageNumber="22">AMS</collectionCode>
K.261186, K.261187 on two slides) same data as holotype
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;
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<specimenCount id="A396FD62FFA6983B617FEB88FBEFFE6C" box="[1088,1118,469,490]" pageId="21" pageNumber="22" type="female">1♀</specimenCount>
(
<specimenCode id="E5369E90FFA6983B6154EB88FB74FE6D" box="[1131,1221,469,491]" collectionCode="HW" pageId="21" pageNumber="22">HW 1.09</specimenCode>
) (
<collectionCode id="D381AE2EFFA6983B61E6EB88FABFFE6C" box="[1241,1294,469,490]" pageId="21" pageNumber="22">AMS</collectionCode>
K.
<quantity id="72689B0EFFA6983B6016EB88FA3CFE6D" box="[1321,1421,469,491]" metricMagnitude="3" metricUnit="m" metricValue="9.5946468" pageId="21" pageNumber="22" unit="in" value="377742.0">377742 in</quantity>
ethanol) same data as holotype
</materialsCitation>
;
<materialsCitation id="05F83CB6FFA6983B616AEBADFBD2FDA6" ID-GBIF-Occurrence="2621712314" collectingDate="2013-07-07" collectionCode="AMS" elevation="71" latitude="-25.9492" longLatPrecision="6" longitude="153.0927" pageId="21" pageNumber="22" specimenCode="HW 1.20" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="A396FD62FFA6983B616AEBADFBC2FD83" box="[1109,1139,496,517]" pageId="21" pageNumber="22" type="male">1♂</specimenCount>
(
<specimenCode id="E5369E90FFA6983B61BEEBADFB6DFD80" box="[1153,1244,496,518]" collectionCode="HW" pageId="21" pageNumber="22">HW 1.20</specimenCode>
) (
<collectionCode id="D381AE2EFFA6983B61CEEBADFA97FD83" box="[1265,1318,496,517]" pageId="21" pageNumber="22">AMS</collectionCode>
K.
<quantity id="72689B0EFFA6983B607DEBADFC9CFDA6" metricMagnitude="3" metricUnit="m" metricValue="9.5945706" pageId="21" pageNumber="22" unit="in" value="377739.0">377739 in</quantity>
ethanol) same data as holotype
</materialsCitation>
;
<materialsCitation id="05F83CB6FFA6983B6151E857FCC1FDD3" ID-GBIF-Occurrence="2621712310" collectingDate="2013-07-07" collectionCode="AMS" elevation="71" latitude="-25.9492" longLatPrecision="6" longitude="153.0927" pageId="21" pageNumber="22" specimenCount="18" specimenCount-female="6" specimenCount-juvenile="7" specimenCount-male="5" typeStatus="paratype">
<specimenCount id="A396FD62FFA6983B6151E857FB11FDA6" box="[1134,1184,522,544]" pageId="21" pageNumber="22" type="male">5♂♂</specimenCount>
,
<specimenCount id="A396FD62FFA6983B6194E857FB6DFDA6" box="[1195,1244,522,544]" pageId="21" pageNumber="22" type="female">5♀♀</specimenCount>
,
<specimenCount id="A396FD62FFA6983B61D7E857FAF9FDA6" box="[1256,1352,522,544]" pageId="21" pageNumber="22" type="juvenile">4 juvenile</specimenCount>
<specimenCount id="A396FD62FFA6983B606FE856FAC4FDA6" box="[1360,1397,523,544]" pageId="21" pageNumber="22" type="female">♀♀</specimenCount>
,
<specimenCount id="A396FD62FFA6983B60BEE857FCC0FDBD" pageId="21" pageNumber="22" type="juvenile">3 juveniles</specimenCount>
(
<collectionCode id="D381AE2EFFA6983B6640E878FC05FDBC" box="[895,948,549,570]" pageId="21" pageNumber="22">AMS</collectionCode>
K.377744 all together in ethanol) same data as holotype
</materialsCitation>
;
<materialsCitation id="05F83CB6FFA6983B6647E81DFBB6FDF6" ID-GBIF-Occurrence="2621712336" collectingDate="2013-07-07" collectionCode="AMS" elevation="71" latitude="-25.9492" longLatPrecision="6" longitude="153.0927" pageId="21" pageNumber="22" specimenCode="HW1.05" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="A396FD62FFA6983B6647E81DFC27FDD3" box="[888,918,576,597]" pageId="21" pageNumber="22" type="female">1♀</specimenCount>
(
<specimenCode id="E5369E90FFA6983B669EE862FC44FDD3" box="[929,1013,575,597]" collectionCode="HW" pageId="21" pageNumber="22">HW1.05</specimenCode>
) (
<collectionCode id="D381AE2EFFA6983B6138E81DFB8DFDD3" box="[1031,1084,576,597]" pageId="21" pageNumber="22">AMS</collectionCode>
K.261126, K.261127 on two slides) same locality as holotype
</materialsCitation>
,
<date id="C12E102BFFA6983B6130E807FBDBFDF6" box="[1039,1130,602,624]" pageId="21" pageNumber="22" value="2016-01-27">27.i.2016</date>
, Graeme Smith;
<materialsCitation id="05F83CB6FFA6983B6039E807FA3BFD0D" ID-GBIF-Occurrence="2621712342" collectingDate="2013-07-07" collectionCode="HW, AMS" elevation="71" latitude="-25.9492" longLatPrecision="6" longitude="153.0927" pageId="21" pageNumber="22" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="A396FD62FFA6983B6039E807FA92FDE9" box="[1286,1315,602,623]" pageId="21" pageNumber="22" type="male">1♂</specimenCount>
(
<collectionCode id="D381AE2EFFA6983B6011E807FAE6FDE9" box="[1326,1367,602,623]" pageId="21" pageNumber="22">HW</collectionCode>
0.98) (
<collectionCode id="D381AE2EFFA6983B661FE828FCE4FD0C" box="[800,853,629,650]" pageId="21" pageNumber="22">AMS</collectionCode>
K.261128, K.261129 on two slides) same data as previous
</materialsCitation>
;
<materialsCitation id="05F83CB6FFA6983B6625E8CDFC00FD46" ID-GBIF-Occurrence="2621712371" collectingDate="2013-07-07" collectionCode="AMS" elevation="71" latitude="-25.9492" longLatPrecision="6" longitude="153.0927" pageId="21" pageNumber="22" specimenCode="HW 1.23" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="A396FD62FFA6983B6625E8CDFC89FD23" box="[794,824,656,677]" pageId="21" pageNumber="22" type="female">1♀</specimenCount>
(
<specimenCode id="E5369E90FFA6983B6679E8CDFC13FD20" box="[838,930,656,678]" collectionCode="HW" pageId="21" pageNumber="22">HW 1.23</specimenCode>
) (
<collectionCode id="D381AE2EFFA6983B6687E8CDFC5CFD23" box="[952,1005,656,677]" pageId="21" pageNumber="22">AMS</collectionCode>
K.261130, K.261131 on two slides) same data as previous
</materialsCitation>
;
<materialsCitation id="05F83CB6FFA6983B6686E8F7FC0DFD5D" ID-GBIF-Occurrence="2621712365" collectingDate="2013-07-07" collectionCode="HW" elevation="71" latitude="-25.9492" longLatPrecision="6" longitude="153.0927" pageId="21" pageNumber="22" specimenCode="HW 1.08" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="A396FD62FFA6983B6686E8F7FC66FD39" box="[953,983,682,703]" pageId="21" pageNumber="22" type="female">1♀</specimenCount>
(
<specimenCode id="E5369E90FFA6983B66DDE8F7FB88FD46" box="[994,1081,682,704]" collectionCode="HW" pageId="21" pageNumber="22">HW 1.08</specimenCode>
) (gbs004893 in 100% ethanol) same data as previous
</materialsCitation>
;
<materialsCitation id="05F83CB6FFA6983B66F8E898FBE5FD73" ID-GBIF-Occurrence="2621712338" collectingDate="2013-07-07" collectionCode="AMS" elevation="71" latitude="-25.9492" longLatPrecision="6" longitude="153.0927" pageId="21" pageNumber="22" specimenCode="HW 1.05" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="A396FD62FFA6983B66F8E898FC54FD5C" box="[967,997,709,730]" pageId="21" pageNumber="22" type="male">1♂</specimenCount>
(
<specimenCode id="E5369E90FFA6983B66CCE898FBFEFD5D" box="[1011,1103,709,731]" collectionCode="HW" pageId="21" pageNumber="22">HW 1.05</specimenCode>
) (
<collectionCode id="D381AE2EFFA6983B615AE898FB2BFD5C" box="[1125,1178,709,730]" pageId="21" pageNumber="22">AMS</collectionCode>
K.261132, K.261133 on two slides) same data as previous
</materialsCitation>
;
<materialsCitation id="05F83CB6FFA6983B6163E8BDFB40FC96" ID-GBIF-Occurrence="2621712334" collectingDate="2013-07-07" collectionCode="HW, AMS" elevation="71" latitude="-25.9492" longLatPrecision="6" longitude="153.0927" pageId="21" pageNumber="22" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="A396FD62FFA6983B6163E8BDFBCBFD73" box="[1116,1146,736,757]" pageId="21" pageNumber="22" type="female">1♀</specimenCount>
(
<collectionCode id="D381AE2EFFA6983B61BAE8BDFB1FFD73" box="[1157,1198,736,757]" pageId="21" pageNumber="22">HW</collectionCode>
0.93) (
<collectionCode id="D381AE2EFFA6983B61D0E8BDFA95FD73" box="[1263,1316,736,757]" pageId="21" pageNumber="22">AMS</collectionCode>
K.261134, K.261135 on two slides) same data as previous
</materialsCitation>
;
<materialsCitation id="05F83CB6FFA6983B61C2E8A7FA9BFCAD" ID-GBIF-Occurrence="2621712344" collectingDate="2013-07-07" collectionCode="HW" elevation="71" latitude="-25.9492" longLatPrecision="6" longitude="153.0927" pageId="21" pageNumber="22" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="A396FD62FFA6983B61C2E8A7FAAAFC89" box="[1277,1307,762,783]" pageId="21" pageNumber="22" type="male">1♂</specimenCount>
(
<collectionCode id="D381AE2EFFA6983B6015E8A7FAE2FC89" box="[1322,1363,762,783]" pageId="21" pageNumber="22">HW</collectionCode>
0.85) (gbs004896 in 100% ethanol) same data as previous
</materialsCitation>
;
<materialsCitation id="05F83CB6FFA6983B6008E948FA85FCC3" ID-GBIF-Occurrence="2621712358" collectingDate="2013-07-07" collectionCode="HW" elevation="71" latitude="-25.9492" longLatPrecision="6" longitude="153.0927" pageId="21" pageNumber="22" specimenCode="HW 1.01" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="A396FD62FFA6983B6008E948FAE5FCAC" box="[1335,1364,789,810]" pageId="21" pageNumber="22" type="female">1♀</specimenCount>
(
<specimenCode id="E5369E90FFA6983B605BE948FCF5FCC0" collectionCode="HW" pageId="21" pageNumber="22">HW 1.01</specimenCode>
) (gbs004897 in 100% ethanol) same data as previous
</materialsCitation>
;
<materialsCitation id="05F83CB6FFA6983B6003E96DFAD6FCE6" ID-GBIF-Occurrence="2621712313" collectingDate="2013-07-07" collectionCode="HW, AMS" elevation="71" latitude="-25.9492" longLatPrecision="6" longitude="153.0927" pageId="21" pageNumber="22" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="A396FD62FFA6983B6003E96DFAE8FCC3" box="[1340,1369,816,837]" pageId="21" pageNumber="22" type="male">1♂</specimenCount>
(
<collectionCode id="D381AE2EFFA6983B605BE96DFA3CFCC3" box="[1380,1421,816,837]" pageId="21" pageNumber="22">HW</collectionCode>
0.85) (
<collectionCode id="D381AE2EFFA6983B6669E917FC3AFCD9" box="[854,907,842,863]" pageId="21" pageNumber="22">AMS</collectionCode>
K.
<quantity id="72689B0EFFA6983B669BE917FBB7FCE6" box="[932,1030,842,864]" metricMagnitude="3" metricUnit="m" metricValue="9.5949516" pageId="21" pageNumber="22" unit="in" value="377754.0">377754 in</quantity>
100% ethanol) same data as previous
</materialsCitation>
;
<materialsCitation id="05F83CB6FFA6983B604FE917FAD6FCFD" ID-GBIF-Occurrence="2621712350" collectingDate="2013-07-07" collectionCode="HW" elevation="71" latitude="-25.9492" longLatPrecision="6" longitude="153.0927" pageId="21" pageNumber="22" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="A396FD62FFA6983B604FE917FA3CFCD9" box="[1392,1421,842,863]" pageId="21" pageNumber="22" type="female">1♀</specimenCount>
(
<collectionCode id="D381AE2EFFA6983B661FE938FCF8FCFC" box="[800,841,869,890]" pageId="21" pageNumber="22">HW</collectionCode>
0.95) (gbs004899 in 100% ethanol) same data as previous
</materialsCitation>
;
<materialsCitation id="05F83CB6FFA6983B6050E938FAC8FC13" ID-GBIF-Occurrence="2621712311" collectingDate="2013-07-07" collectionCode="HW" elevation="71" latitude="-25.9492" longLatPrecision="6" longitude="153.0927" pageId="21" pageNumber="22" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="A396FD62FFA6983B6050E938FA3CFCFC" box="[1391,1421,869,890]" pageId="21" pageNumber="22" type="female">1♀</specimenCount>
(
<collectionCode id="D381AE2EFFA6983B661FE9DDFCF8FC13" box="[800,841,896,917]" pageId="21" pageNumber="22">HW</collectionCode>
0.95) (gbs004900 in 100% ethanol) same data as previous
</materialsCitation>
;
<materialsCitation id="05F83CB6FFA6983960BEE9DDFF7EF91F" ID-GBIF-Occurrence="2621712363" collectingDate="2013-07-07" collectionCode="HW" elevation="71" lastPageId="23" lastPageNumber="24" latitude="-25.9492" longLatPrecision="6" longitude="153.0927" pageId="21" pageNumber="22" specimenCount="2" specimenCount-female="1" specimenCount-subadult="1" typeStatus="paratype">
<specimenCount id="A396FD62FFA6983B60BEE9DDFCDAFC36" pageId="21" pageNumber="22" type="subadult">1 subadult</specimenCount>
<specimenCount id="A396FD62FFA6983B664FE9C6FC33FC36" box="[880,898,923,944]" pageId="21" pageNumber="22" type="female"></specimenCount>
(
<collectionCode id="D381AE2EFFA6983B66B1E9C7FC06FC29" box="[910,951,922,943]" pageId="21" pageNumber="22">HW</collectionCode>
0.90) (gbs004901 in 100% ethanol) same data as previous
</materialsCitation>
;
<materialsCitation id="05F83CB6FFA4983965E7ECDEFF5AF932" ID-GBIF-Occurrence="2621712331" collectingDate="2013-07-07" collectionCode="HW" elevation="71" latitude="-25.9492" longLatPrecision="6" longitude="153.0927" pageId="23" pageNumber="24" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="A396FD62FFA4983965E7ECDEFF46F91E" box="[216,247,1667,1688]" pageId="23" pageNumber="24" type="male">1♂</specimenCount>
(
<collectionCode id="D381AE2EFFA49839643CECDEFE9DF91E" box="[259,300,1667,1688]" pageId="23" pageNumber="24">HW</collectionCode>
0.95) (gbs004902 in 100% ethanol) same data as previous
</materialsCitation>
;
<materialsCitation id="05F83CB6FFA4983965C8ECC3FEADF948" ID-GBIF-Occurrence="2621712368" collectingDate="2013-07-07" collectionCode="HW" elevation="71" latitude="-25.9492" longLatPrecision="6" longitude="153.0927" pageId="23" pageNumber="24" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="A396FD62FFA4983965C8ECC3FEA4F935" box="[247,277,1694,1715]" pageId="23" pageNumber="24" type="male">1♂</specimenCount>
(
<collectionCode id="D381AE2EFFA49839641BECC3FEFCF935" box="[292,333,1694,1715]" pageId="23" pageNumber="24">HW</collectionCode>
0.88) (gbs004903 in 100% ethanol) same data as previous
</materialsCitation>
;
<materialsCitation id="05F83CB6FFA498396417ECE4FE27F96F" ID-GBIF-Occurrence="2621712384" collectingDate="2013-07-07" collectionCode="HW" elevation="71" latitude="-25.9492" longLatPrecision="6" longitude="153.0927" pageId="23" pageNumber="24" specimenCount="2" specimenCount-female="1" specimenCount-juvenile="1" typeStatus="paratype">
<specimenCount id="A396FD62FFA498396417ECE4FE38F948" box="[296,393,1720,1742]" pageId="23" pageNumber="24" type="juvenile">1 juvenile</specimenCount>
<specimenCount id="A396FD62FFA4983964ADECE4FE15F948" box="[402,420,1721,1742]" pageId="23" pageNumber="24" type="female"></specimenCount>
(
<collectionCode id="D381AE2EFFA49839648CECE4FE6DF948" box="[435,476,1721,1742]" pageId="23" pageNumber="24">HW</collectionCode>
0.83) (gbs004904 in 100% ethanol) same data as previous
</materialsCitation>
;
<materialsCitation id="05F83CB6FFA4983964A1EC8EFE4DF882" ID-GBIF-Occurrence="2621712360" collectingDate="2013-07-07" collectionCode="HW" elevation="71" latitude="-25.9492" longLatPrecision="6" longitude="153.0927" pageId="23" pageNumber="24" specimenCount="2" specimenCount-juvenile="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="A396FD62FFA4983964A1EC8EFE4BF96F" box="[414,506,1747,1769]" pageId="23" pageNumber="24" type="juvenile">1 juvenile</specimenCount>
<specimenCount id="A396FD62FFA4983964C1EC89FDA1F96F" box="[510,528,1748,1769]" pageId="23" pageNumber="24" type="male"></specimenCount>
(
<collectionCode id="D381AE2EFFA498396724EC8EFDF5F96E" box="[539,580,1747,1768]" pageId="23" pageNumber="24">HW</collectionCode>
0.75) (gbs004905 in 100% ethanol) same data as previous
</materialsCitation>
;
<materialsCitation id="05F83CB6FFA498396739ECB3FDDCF898" ID-GBIF-Occurrence="2621712335" collectingDate="2013-07-07" collectionCode="HW" elevation="71" latitude="-25.9492" longLatPrecision="6" longitude="153.0927" pageId="23" pageNumber="24" specimenCount="2" specimenCount-female="1" specimenCount-juvenile="1" typeStatus="paratype">
<specimenCount id="A396FD62FFA498396739ECB3FDD4F882" box="[518,613,1774,1796]" pageId="23" pageNumber="24" type="juvenile">1 juvenile</specimenCount>
<specimenCount id="A396FD62FFA498396753ECB3FDCFF885" box="[620,638,1774,1795]" pageId="23" pageNumber="24" type="female"></specimenCount>
(
<collectionCode id="D381AE2EFFA4983967B4ECB3FD05F885" box="[651,692,1774,1795]" pageId="23" pageNumber="24">HW</collectionCode>
0.75) (gbs004906 in 100% ethanol) same data as previous
</materialsCitation>
;
<materialsCitation id="05F83CB6FFA498396749ED54FD69F8BF" ID-GBIF-Occurrence="2621712375" collectingDate="2013-07-07" collectionCode="HW" elevation="71" latitude="-25.9492" longLatPrecision="6" longitude="153.0927" pageId="23" pageNumber="24" specimenCount="2" specimenCount-juvenile="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="A396FD62FFA498396749ED54FD64F898" box="[630,725,1800,1822]" pageId="23" pageNumber="24" type="juvenile">1 juvenile</specimenCount>
<specimenCount id="A396FD62FFA4983967E5ED54FD5DF898" box="[730,748,1801,1822]" pageId="23" pageNumber="24" type="male"></specimenCount>
(
<collectionCode id="D381AE2EFFA498396540ED7EFF19F8BE" box="[127,168,1827,1848]" pageId="23" pageNumber="24">HW</collectionCode>
0.70) (gbs004907 in 100% ethanol) same data as previous
</materialsCitation>
;
<materialsCitation id="05F83CB6FFA4983967DFED7EFF7EF8E8" ID-GBIF-Occurrence="2621712383" collectingDate="2013-07-07" collectionCode="HW" elevation="71" latitude="-25.9492" longLatPrecision="6" longitude="153.0927" pageId="23" pageNumber="24" specimenCount="2" specimenCount-juvenile="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="A396FD62FFA4983967DFED7EFF77F8D2" pageId="23" pageNumber="24" type="juvenile">1 juvenile</specimenCount>
<specimenCount id="A396FD62FFA4983965F3ED63FF6FF8D5" box="[204,222,1854,1875]" pageId="23" pageNumber="24" type="male"></specimenCount>
(
<collectionCode id="D381AE2EFFA4983965D5ED63FEA2F8D5" box="[234,275,1854,1875]" pageId="23" pageNumber="24">HW</collectionCode>
0.70) (gbs004908 in 100% ethanol) same data as previous
</materialsCitation>
;
<materialsCitation id="05F83CB6FFA4983965E5ED04FEF4F80F" ID-GBIF-Occurrence="2621712347" collectingDate="2013-07-07" collectionCode="HW" elevation="71" latitude="-25.9492" longLatPrecision="6" longitude="153.0927" pageId="23" pageNumber="24" specimenCount="2" specimenCount-female="1" specimenCount-juvenile="1" typeStatus="paratype">
<specimenCount id="A396FD62FFA4983965E5ED04FE8BF8E8" box="[218,314,1880,1902]" pageId="23" pageNumber="24" type="juvenile">1 juvenile</specimenCount>
<specimenCount id="A396FD62FFA49839647DED04FEE5F8E8" box="[322,340,1881,1902]" pageId="23" pageNumber="24" type="female"></specimenCount>
(
<collectionCode id="D381AE2EFFA49839645DED04FE3AF8E8" box="[354,395,1881,1902]" pageId="23" pageNumber="24">HW</collectionCode>
0.83) (gbs004909 in 100% ethanol) same data as previous
</materialsCitation>
;
<materialsCitation id="05F83CB6FFA498396472ED2EFE26F822" ID-GBIF-Occurrence="2621712386" collectingDate="2013-07-07" collectionCode="HW" elevation="71" latitude="-25.9492" longLatPrecision="6" longitude="153.0927" pageId="23" pageNumber="24" specimenCount="2" specimenCount-juvenile="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="A396FD62FFA498396472ED2EFE18F80F" box="[333,425,1907,1929]" pageId="23" pageNumber="24" type="juvenile">1 juvenile</specimenCount>
<specimenCount id="A396FD62FFA498396492ED29FE0EF80F" box="[429,447,1908,1929]" pageId="23" pageNumber="24" type="male"></specimenCount>
(
<collectionCode id="D381AE2EFFA4983964F6ED2EFE43F80E" box="[457,498,1907,1928]" pageId="23" pageNumber="24">HW</collectionCode>
0.68) (gbs004910 in 100% ethanol) same data as previous
</materialsCitation>
;
<materialsCitation id="05F83CB6FFA4983964A0EDD3FE4DF838" ID-GBIF-Occurrence="2621712352" collectingDate="2013-07-07" collectionCode="HW" elevation="71" latitude="-25.9492" longLatPrecision="6" longitude="153.0927" pageId="23" pageNumber="24" specimenCount="2" specimenCount-juvenile="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="A396FD62FFA4983964A0EDD3FE4AF822" box="[415,507,1934,1956]" pageId="23" pageNumber="24" type="juvenile">1 juvenile</specimenCount>
<specimenCount id="A396FD62FFA4983964C0EDD3FDA0F825" box="[511,529,1934,1955]" pageId="23" pageNumber="24" type="male"></specimenCount>
(
<collectionCode id="D381AE2EFFA498396724EDD3FDF5F825" box="[539,580,1934,1955]" pageId="23" pageNumber="24">HW</collectionCode>
0.75) (gbs004911 in 100% ethanol) same data as previous
</materialsCitation>
;
<materialsCitation id="05F83CB6FFA498396739EDF4FDDCF85F" ID-GBIF-Occurrence="2621712385" collectingDate="2013-07-07" collectionCode="HW" elevation="71" latitude="-25.9492" longLatPrecision="6" longitude="153.0927" pageId="23" pageNumber="24" specimenCount="2" specimenCount-female="1" specimenCount-juvenile="1" typeStatus="paratype">
<specimenCount id="A396FD62FFA498396739EDF4FDD4F838" box="[518,613,1960,1982]" pageId="23" pageNumber="24" type="juvenile">1 juvenile</specimenCount>
<specimenCount id="A396FD62FFA498396753EDF4FDCFF838" box="[620,638,1961,1982]" pageId="23" pageNumber="24" type="female"></specimenCount>
(
<collectionCode id="D381AE2EFFA4983967B4EDF4FD05F838" box="[651,692,1961,1982]" pageId="23" pageNumber="24">HW</collectionCode>
0.68) (gbs004912 in 100% ethanol) same data as previous
</materialsCitation>
;
<materialsCitation id="05F83CB6FFA498396749ED9EFA3BF91F" ID-GBIF-Occurrence="2621712324" box="[630,1418,1667,2009]" collectingDate="2013-07-07" collectionCode="HW" elevation="71" latitude="-25.9492" longLatPrecision="6" longitude="153.0927" pageId="23" pageNumber="24" specimenCount="2" specimenCount-juvenile="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="A396FD62FFA498396749ED9EFD64F85F" box="[630,725,1987,2009]" pageId="23" pageNumber="24" type="juvenile">1 juvenile</specimenCount>
<specimenCount id="A396FD62FFA4983967E5ED99FD5DF85F" box="[730,748,1988,2009]" pageId="23" pageNumber="24" type="male"></specimenCount>
(
<collectionCode id="D381AE2EFFA49839661FECDEFCF8F91E" box="[800,841,1667,1688]" pageId="23" pageNumber="24">HW</collectionCode>
0.70) (gbs004913 in 100% ethanol) same data as previous
</materialsCitation>
;
<materialsCitation id="05F83CB6FFA498396625ECC3FC3AF948" ID-GBIF-Occurrence="2621712329" collectingDate="2013-07-07" collectionCode="HW" elevation="71" latitude="-25.9492" longLatPrecision="6" longitude="153.0927" pageId="23" pageNumber="24" specimenCount="2" specimenCount-juvenile="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="A396FD62FFA498396625ECC3FCC8F932" box="[794,889,1694,1716]" pageId="23" pageNumber="24" type="juvenile">1 juvenile</specimenCount>
<specimenCount id="A396FD62FFA4983966BFECC3FC23F935" box="[896,914,1694,1715]" pageId="23" pageNumber="24" type="male"></specimenCount>
(
<collectionCode id="D381AE2EFFA4983966A0ECC3FC79F935" box="[927,968,1694,1715]" pageId="23" pageNumber="24">HW</collectionCode>
0.58) (gbs004914 in 100% ethanol) same data as previous
</materialsCitation>
;
<materialsCitation id="05F83CB6FFA4983966AAECE4FC5EF96F" ID-GBIF-Occurrence="2621712319" collectingDate="2013-07-07" collectionCode="HW" elevation="71" latitude="-25.9492" longLatPrecision="6" longitude="153.0927" pageId="23" pageNumber="24" specimenCount="1" specimenCount-juvenile="1" typeStatus="paratype">
<specimenCount id="A396FD62FFA4983966AAECE4FC47F948" box="[917,1014,1720,1742]" pageId="23" pageNumber="24" type="juvenile">1 juvenile</specimenCount>
(
<collectionCode id="D381AE2EFFA49839613CECE4FB9DF948" box="[1027,1068,1721,1742]" pageId="23" pageNumber="24">HW</collectionCode>
0.48) (gbs004915 in 100% ethanol) same data as previous
</materialsCitation>
.
</paragraph>
<caption id="E1EF6663FFA6983B6625EDFFFC3EF855" ID-DOI="http://doi.org/10.5281/zenodo.3837999" ID-Zenodo-Dep="3837999" httpUri="https://zenodo.org/record/3837999/files/figure.png" pageId="21" pageNumber="22" startId="21.[794,863,1954,1976]" targetBox="[794,1421,971,1946]" targetPageId="21">
<paragraph id="B52F36EBFFA6983B6625EDFFFC3EF855" blockId="21.[794,1422,1954,2003]" pageId="21" pageNumber="22">
<emphasis id="87E4EAF9FFA6983B6625EDFFFC31F831" bold="true" box="[794,896,1954,1976]" pageId="21" pageNumber="22">Figure 37</emphasis>
.
<taxonomicName id="72904D68FFA6983B66B0EDFFFBCCF831" authority="Smith &amp; Mitchell &amp; Lee &amp; Espinasa, 2019" authorityName="Smith &amp; Mitchell &amp; Lee &amp; Espinasa" authorityYear="2019" box="[911,1149,1954,1975]" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="cooloola" status="sp. nov.">
<emphasis id="87E4EAF9FFA6983B66B0EDFFFBCCF831" box="[911,1149,1954,1975]" italics="true" pageId="21" pageNumber="22">Heterolepisma cooloola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9CD75782FFA6983B61B9EDFEFB64F83E" box="[1158,1237,1955,1976]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
from leaf litter at Carlo Point.
</paragraph>
</caption>
<caption id="E1EF6663FFA598386599ED50FDBFF80B" ID-DOI="http://doi.org/10.5281/zenodo.3838001" ID-Zenodo-Dep="3838001" httpUri="https://zenodo.org/record/3838001/files/figure.png" pageId="22" pageNumber="23" startId="22.[166,243,1805,1827]" targetBox="[174,1460,196,1772]" targetPageId="22">
<paragraph id="B52F36EBFFA598386599ED50FDBFF80B" blockId="22.[166,1467,1804,1933]" pageId="22" pageNumber="23">
<emphasis id="87E4EAF9FFA598386599ED50FE87F8A4" bold="true" box="[166,310,1805,1827]" pageId="22" pageNumber="23">Figures 3848</emphasis>
.
<taxonomicName id="72904D68FFA59838647CED50FD9BF8A4" authority="Smith &amp; Mitchell &amp; Lee &amp; Espinasa, 2019" authorityName="Smith &amp; Mitchell &amp; Lee &amp; Espinasa" authorityYear="2019" box="[323,554,1805,1826]" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="cooloola" status="sp. nov.">
<emphasis id="87E4EAF9FFA59838647CED50FD9BF8A4" box="[323,554,1805,1826]" italics="true" pageId="22" pageNumber="23">Heterolepisma cooloola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9CD75782FFA59838670FED50FDCDF8A4" box="[560,636,1805,1826]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
holotype ♀, unless indicated otherwise by specimen number
<emphasis id="87E4EAF9FFA5983861F5ED50FB43F8A4" box="[1226,1266,1805,1826]" italics="true" pageId="22" pageNumber="23">(38)</emphasis>
habitus (K.377754);
<emphasis id="87E4EAF9FFA598386599ED7AFF7FF8BA" box="[166,206,1831,1852]" italics="true" pageId="22" pageNumber="23">(39)</emphasis>
posterior comb of pronotum with scale;
<emphasis id="87E4EAF9FFA59838676DED7AFDCBF8BA" box="[594,634,1831,1852]" italics="true" pageId="22" pageNumber="23">(40)</emphasis>
head;
<emphasis id="87E4EAF9FFA598386786ED7AFD53F8BA" box="[697,738,1831,1852]" italics="true" pageId="22" pageNumber="23">(41)</emphasis>
scape, pedicel and basal intervals of flagellum, from above;
<emphasis id="87E4EAF9FFA59838601CED7AFAFAF8BA" box="[1315,1355,1831,1852]" italics="true" pageId="22" pageNumber="23">(42)</emphasis>
idem, from below;
<emphasis id="87E4EAF9FFA5983865D3ED1FFEA5F8D1" box="[236,276,1858,1879]" italics="true" pageId="22" pageNumber="23">(43)</emphasis>
antenna, most distal surviving interval;
<emphasis id="87E4EAF9FFA5983867B1ED1FFD07F8D1" box="[654,694,1858,1879]" italics="true" pageId="22" pageNumber="23">(44)</emphasis>
mandible;
<emphasis id="87E4EAF9FFA598386620ED1FFCF6F8D1" box="[799,839,1858,1879]" italics="true" pageId="22" pageNumber="23">(45)</emphasis>
idem, detail of molar and incisor regions;
<emphasis id="87E4EAF9FFA5983861EBED1FFB4CF8D1" box="[1236,1277,1858,1879]" italics="true" pageId="22" pageNumber="23">(46)</emphasis>
maxilla, only larger setae of palp illustrated;
<emphasis id="87E4EAF9FFA5983864AFED00FE09F8F4" box="[400,440,1885,1906]" italics="true" pageId="22" pageNumber="23">(47)</emphasis>
idem, lacinia and galea (QM 207012);
<emphasis id="87E4EAF9FFA598386612ED00FCE4F8F4" box="[813,853,1885,1906]" italics="true" pageId="22" pageNumber="23">(48)</emphasis>
ultimate article of maxillary palp of ♂ (QM 207012). Scale bars = 0.1 mm unless otherwise indicated.
</paragraph>
</caption>
<caption id="E1EF6663FFA498396546EFB0FC7AF9D4" ID-DOI="http://doi.org/10.5281/zenodo.3838003" ID-Zenodo-Dep="3838003" httpUri="https://zenodo.org/record/3838003/files/figure.png" pageId="23" pageNumber="24" startId="23.[121,198,1517,1539]" targetBox="[134,1417,159,1496]" targetPageId="23">
<paragraph id="B52F36EBFFA498396546EFB0FC7AF9D4" blockId="23.[121,1421,1517,1619]" pageId="23" pageNumber="24">
<emphasis id="87E4EAF9FFA498396546EFB0FEB9F984" bold="true" box="[121,264,1517,1539]" pageId="23" pageNumber="24">Figures 4959</emphasis>
.
<taxonomicName id="72904D68FFA49839642BEFB0FE4BF984" authority="Smith &amp; Mitchell &amp; Lee &amp; Espinasa, 2019" authorityName="Smith &amp; Mitchell &amp; Lee &amp; Espinasa" authorityYear="2019" box="[276,506,1517,1538]" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="cooloola" status="sp. nov.">
<emphasis id="87E4EAF9FFA49839642BEFB0FE4BF984" box="[276,506,1517,1538]" italics="true" pageId="23" pageNumber="24">Heterolepisma cooloola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9CD75782FFA49839673EEFB0FDFDF984" box="[513,588,1517,1538]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
holotype ♀
<emphasis id="87E4EAF9FFA4983967FCEFB0FD5AF984" box="[707,747,1517,1538]" italics="true" pageId="23" pageNumber="24">(49)</emphasis>
labium, only large setae of palp illustrated;
<emphasis id="87E4EAF9FFA4983961ADEFB0FB0BF984" box="[1170,1210,1517,1538]" italics="true" pageId="23" pageNumber="24">(50)</emphasis>
idem, ultimate article of palp;
<emphasis id="87E4EAF9FFA4983965F6EC55FF43F99B" box="[201,242,1544,1565]" italics="true" pageId="23" pageNumber="24">(51)</emphasis>
pronotum;
<emphasis id="87E4EAF9FFA49839645DEC55FE38F99B" box="[354,393,1544,1565]" italics="true" pageId="23" pageNumber="24">(52)</emphasis>
idem, right margin;
<emphasis id="87E4EAF9FFA49839676FEC55FDC6F99B" box="[592,631,1544,1565]" italics="true" pageId="23" pageNumber="24">(53)</emphasis>
idem, detail of right anterior trichobothrial area;
<emphasis id="87E4EAF9FFA49839616DEC55FBC8F99B" box="[1106,1145,1544,1565]" italics="true" pageId="23" pageNumber="24">(54)</emphasis>
idem, detail of left posterior trichobothrial area; (
<emphasis id="87E4EAF9FFA49839647FEC7FFEE9F9B1" box="[320,344,1570,1591]" italics="true" pageId="23" pageNumber="24">55</emphasis>
) lateral margin of mesonotum;
<emphasis id="87E4EAF9FFA4983967B1EC7FFD07F9B1" box="[654,694,1570,1591]" italics="true" pageId="23" pageNumber="24">(56)</emphasis>
idem, posterior trichobothrial area;
<emphasis id="87E4EAF9FFA498396128EC7FFB8EF9B1" box="[1047,1087,1570,1591]" italics="true" pageId="23" pageNumber="24">(57)</emphasis>
idem, anterior trichobothrial area;
<emphasis id="87E4EAF9FFA498396546EC60FF10F9D4" box="[121,161,1597,1618]" italics="true" pageId="23" pageNumber="24">(58)</emphasis>
idem, right posterior comb;
<emphasis id="87E4EAF9FFA49839648CEC60FE6AF9D4" box="[435,475,1597,1618]" italics="true" pageId="23" pageNumber="24">(59)</emphasis>
lateral margin of metanotum. Scale bars = 0.1 mm.
</paragraph>
</caption>
<paragraph id="B52F36EBFFA498366625ED5AFA1DF85F" blockId="23.[794,1422,1799,2009]" lastBlockId="24.[839,1467,1667,2009]" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
<emphasis id="87E4EAF9FFA498396625ED5AFB86F899" bold="true" box="[794,1079,1799,1823]" pageId="23" pageNumber="24">Other material examined</emphasis>
.
<materialsCitation id="05F83CB6FFA49839617BED54FB47F8D2" ID-GBIF-Occurrence="2621712370" collectingDate="2013-07-07" collectionCode="HW, AMS" elevation="45" latitude="-25.9439" longLatPrecision="6" longitude="153.0816" pageId="23" pageNumber="24" specimenCount="1" specimenCount-male="1">
<specimenCount id="A396FD62FFA49839617BED54FBD3F898" box="[1092,1122,1801,1822]" pageId="23" pageNumber="24" type="male">1♂</specimenCount>
(
<collectionCode id="D381AE2EFFA498396151ED54FB26F898" box="[1134,1175,1801,1822]" pageId="23" pageNumber="24">HW</collectionCode>
0.95) (
<collectionCode id="D381AE2EFFA4983961E4ED54FAA1F898" box="[1243,1296,1801,1822]" pageId="23" pageNumber="24">AMS</collectionCode>
K.
<quantity id="72689B0EFFA498396015ED54FA3FF898" box="[1322,1422,1800,1822]" metricMagnitude="3" metricUnit="m" metricValue="9.5946722" pageId="23" pageNumber="24" unit="in" value="377743.0">377743 in</quantity>
ethanol) QLD: Cooloola, near start of Freshwater track,
<geoCoordinate id="D0A4502CFFA498396016ED7EFA3CF8BE" box="[1321,1421,1827,1849]" degrees="25.9439" direction="south" orientation="latitude" pageId="23" pageNumber="24" precision="5" value="-25.9439">25.9439°S</geoCoordinate>
<geoCoordinate id="D0A4502CFFA498396625ED63FC3AF8D2" box="[794,907,1854,1876]" degrees="153.0816" direction="east" orientation="longitude" pageId="23" pageNumber="24" precision="5" value="153.0816">153.0816°E</geoCoordinate>
<elevation id="3EBDD1D8FFA4983966ADED63FC56F8D2" box="[914,999,1854,1876]" metricMagnitude="1" metricUnit="m" metricValue="4.5" pageId="23" pageNumber="24" unit="m" value="45.0">
<quantity id="72689B0EFFA4983966ADED63FC75F8D5" box="[914,964,1854,1876]" metricMagnitude="1" metricUnit="m" metricValue="4.5" pageId="23" pageNumber="24" unit="m" value="45.0">45 m</quantity>
asl
</elevation>
,
<date id="C12E102BFFA4983966CDED63FBEDF8D5" box="[1010,1116,1854,1876]" pageId="23" pageNumber="24" value="2013-07-07">
<collectingDate id="D16AE9C3FFA4983966CDED63FBEDF8D5" box="[1010,1116,1854,1876]" pageId="23" pageNumber="24" value="2013-07-07">7.vii. 2013</collectingDate>
</date>
, Graeme Smith
</materialsCitation>
;
<materialsCitation id="05F83CB6FFA49839603FED63FB4AF8E8" ID-GBIF-Occurrence="2621712303" collectingDate="2013-07-07" collectionCode="HW, AMS" elevation="45" latitude="-25.9439" longLatPrecision="6" longitude="153.0816" pageId="23" pageNumber="24" specimenCount="1" specimenCount-female="1">
<specimenCount id="A396FD62FFA49839603FED63FAAFF8D5" box="[1280,1310,1854,1875]" pageId="23" pageNumber="24" type="female">1♀</specimenCount>
(
<collectionCode id="D381AE2EFFA498396013ED63FAE4F8D5" box="[1324,1365,1854,1875]" pageId="23" pageNumber="24">HW</collectionCode>
0.98) (
<collectionCode id="D381AE2EFFA49839661FED04FCE4F8E8" box="[800,853,1881,1902]" pageId="23" pageNumber="24">AMS</collectionCode>
K.
<quantity id="72689B0EFFA498396650ED05FC60F8E8" box="[879,977,1880,1902]" metricMagnitude="3" metricUnit="m" metricValue="9.5947484" pageId="23" pageNumber="24" unit="in" value="377746.0">377746 in</quantity>
ethanol) same data as previous
</materialsCitation>
;
<materialsCitation id="05F83CB6FFA49839603BED04FA3BF80F" ID-GBIF-Occurrence="2621712328" collectingDate="2013-07-07" collectionCode="AMS" elevation="45" latitude="-25.9439" longLatPrecision="6" longitude="153.0816" pageId="23" pageNumber="24" specimenCode="HW 1.01" specimenCount="1" specimenCount-male="1">
<specimenCount id="A396FD62FFA49839603BED04FA93F8E8" box="[1284,1314,1881,1902]" pageId="23" pageNumber="24" type="male">1♂</specimenCount>
(
<specimenCode id="E5369E90FFA498396012ED04FA36F8E9" box="[1325,1415,1881,1903]" collectionCode="HW" pageId="23" pageNumber="24">HW 1.01</specimenCode>
) (
<collectionCode id="D381AE2EFFA49839661FED2EFCE4F80E" box="[800,853,1907,1928]" pageId="23" pageNumber="24">AMS</collectionCode>
K.
<quantity id="72689B0EFFA49839664DED2EFC66F80F" box="[882,983,1907,1929]" metricMagnitude="3" metricUnit="m" metricValue="9.594773799999999" pageId="23" pageNumber="24" unit="in" value="377747.0">377747 in</quantity>
ethanol) same data as previous (in ethanol)
</materialsCitation>
;
<materialsCitation id="05F83CB6FFA498396625EDD3FA87F838" ID-GBIF-Occurrence="2621712343" collectingDate="2013-07-06" collectionCode="AMS" elevation="22" latitude="-25.8975" longLatPrecision="6" longitude="153.062" pageId="23" pageNumber="24" specimenCode="HW 1.01" specimenCount="1" specimenCount-male="1">
<specimenCount id="A396FD62FFA498396625EDD3FC89F825" box="[794,824,1934,1955]" pageId="23" pageNumber="24" type="male">1♂</specimenCount>
(
<specimenCode id="E5369E90FFA498396678EDD3FC15F822" box="[839,932,1934,1956]" collectionCode="HW" pageId="23" pageNumber="24">HW 1.01</specimenCode>
) (
<collectionCode id="D381AE2EFFA498396684EDD3FC41F825" box="[955,1008,1934,1955]" pageId="23" pageNumber="24">AMS</collectionCode>
K.
<quantity id="72689B0EFFA498396131EDD3FBC4F822" box="[1038,1141,1934,1956]" metricMagnitude="3" metricUnit="m" metricValue="9.5844614" pageId="23" pageNumber="24" unit="in" value="377341.0">377341 in</quantity>
ethanol) QLD: Carlo Point,
<geoCoordinate id="D0A4502CFFA498396625EDF5FCCAF838" box="[794,891,1960,1982]" degrees="25.8975" direction="south" orientation="latitude" pageId="23" pageNumber="24" precision="5" value="-25.8975">25.8975°S</geoCoordinate>
<geoCoordinate id="D0A4502CFFA498396640EDF5FC5CF838" box="[895,1005,1960,1982]" degrees="153.0620" direction="east" orientation="longitude" pageId="23" pageNumber="24" precision="5" value="153.062">153.0620°E</geoCoordinate>
<elevation id="3EBDD1D8FFA4983966CDEDF4FB8FF838" box="[1010,1086,1960,1982]" metricMagnitude="1" metricUnit="m" metricValue="2.2" pageId="23" pageNumber="24" unit="m" value="22.0">
<quantity id="72689B0EFFA4983966CDEDF4FBAEF838" box="[1010,1055,1961,1982]" metricMagnitude="1" metricUnit="m" metricValue="2.2" pageId="23" pageNumber="24" unit="m" value="22.0">22 m</quantity>
asl
</elevation>
,
<date id="C12E102BFFA498396178EDF5FB17F838" box="[1095,1190,1960,1982]" pageId="23" pageNumber="24" value="2013-07-06">
<collectingDate id="D16AE9C3FFA498396178EDF5FB17F838" box="[1095,1190,1960,1982]" pageId="23" pageNumber="24" value="2013-07-06">6.vii.2013</collectingDate>
</date>
, Graeme Smith
</materialsCitation>
;
<materialsCitation id="05F83CB6FFA498396002EDF4FA92F85F" ID-GBIF-Occurrence="2621712391" collectingDate="2013-07-06" collectionCode="AMS" elevation="22" latitude="-25.8975" longLatPrecision="6" longitude="153.062" pageId="23" pageNumber="24" specimenCode="HW 1.03" specimenCount="1" specimenCount-male="1">
<specimenCount id="A396FD62FFA498396002EDF4FAEBF838" box="[1341,1370,1961,1982]" pageId="23" pageNumber="24" type="male">1♂</specimenCount>
(
<specimenCode id="E5369E90FFA49839605AEDF4FCF5F85F" collectionCode="HW" pageId="23" pageNumber="24">HW 1.03</specimenCode>
) (
<collectionCode id="D381AE2EFFA49839666AED9EFC3BF85E" box="[853,906,1987,2008]" pageId="23" pageNumber="24">AMS</collectionCode>
K.
<quantity id="72689B0EFFA49839669DED9EFBB3F85F" box="[930,1026,1987,2009]" metricMagnitude="3" metricUnit="m" metricValue="9.5942912" pageId="23" pageNumber="24" unit="in" value="377728.0">377728 in</quantity>
ethanol) same data as previous
</materialsCitation>
;
<materialsCitation id="05F83CB6FFA498366015ED9EFEF0F932" ID-GBIF-Occurrence="2621712388" collectingDate="2013-07-06" collectionCode="AMS" elevation="22" lastPageId="24" lastPageNumber="25" latitude="-25.8975" longLatPrecision="6" longitude="153.062" pageId="23" pageNumber="24" specimenCode="HW 1.11" specimenCount="3" specimenCount-female="3">
<specimenCount id="A396FD62FFA498396015ED9EFAE8F85E" box="[1322,1369,1987,2008]" pageId="23" pageNumber="24" type="female">3♀♀</specimenCount>
(
<specimenCode id="E5369E90FFA49836605BED9EFF60F91E" collectionCode="HW" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">HW 1.11</specimenCode>
, 0.98 and 0.69) (
<collectionCode id="D381AE2EFFAB98366450ECDEFE15F91E" box="[367,420,1667,1688]" pageId="24" pageNumber="25">AMS</collectionCode>
K.377740 all together in ethanol) same data as previous
</materialsCitation>
;
<materialsCitation id="05F83CB6FFAB98366475ECC3FF53F96F" ID-GBIF-Occurrence="2621712321" collectingDate="2013-07-07" collectionCode="AMS" elevation="63" latitude="-25.996" longLatPrecision="6" longitude="153.0716" pageId="24" pageNumber="25" specimenCode="HW1.15" specimenCount="1" specimenCount-female="1">
<specimenCount id="A396FD62FFAB98366475ECC3FED6F935" box="[330,359,1694,1715]" pageId="24" pageNumber="25" type="female">1♀</specimenCount>
(
<specimenCode id="E5369E90FFAB9836644CECC3FE76F932" box="[371,455,1694,1716]" collectionCode="HW" pageId="24" pageNumber="25">HW1.15</specimenCode>
) (
<collectionCode id="D381AE2EFFAB983664E6ECC3FDBFF935" box="[473,526,1694,1715]" pageId="24" pageNumber="25">AMS</collectionCode>
K.
<quantity id="72689B0EFFAB98366717ECC3FD38F932" box="[552,649,1694,1716]" metricMagnitude="3" metricUnit="m" metricValue="9.5947992" pageId="24" pageNumber="25" unit="in" value="377748.0">377748 in</quantity>
ethanol) QLD: Cooloola,
<geoCoordinate id="D0A4502CFFAB98366433ECE5FEC1F948" box="[268,368,1720,1742]" degrees="25.9960" direction="south" orientation="latitude" pageId="24" pageNumber="25" precision="5" value="-25.996">25.9960°S</geoCoordinate>
<geoCoordinate id="D0A4502CFFAB98366446ECE5FE5BF948" box="[377,490,1720,1742]" degrees="153.0716" direction="east" orientation="longitude" pageId="24" pageNumber="25" precision="5" value="153.0716">153.0716°E</geoCoordinate>
<elevation id="3EBDD1D8FFAB983664CDECE5FDFBF948" box="[498,586,1720,1742]" metricMagnitude="1" metricUnit="m" metricValue="6.3" pageId="24" pageNumber="25" unit="m" value="63.0">
<quantity id="72689B0EFFAB983664CDECE5FD97F948" box="[498,550,1720,1742]" metricMagnitude="1" metricUnit="m" metricValue="6.3" pageId="24" pageNumber="25" unit="m" value="63.0">63 m</quantity>
asl
</elevation>
,
<date id="C12E102BFFAB98366768ECE5FD73F948" box="[599,706,1720,1742]" pageId="24" pageNumber="25" value="2013-07-07">
<collectingDate id="D16AE9C3FFAB98366768ECE5FD73F948" box="[599,706,1720,1742]" pageId="24" pageNumber="25" value="2013-07-07">7.vii. 2013</collectingDate>
</date>
, Graeme Smith
</materialsCitation>
;
<materialsCitation id="05F83CB6FFAB983665D2EC8EFEC8F882" ID-GBIF-Occurrence="2621712392" collectingDate="2013-07-07" collectionCode="AMS" elevation="63" latitude="-25.996" longLatPrecision="6" longitude="153.0716" pageId="24" pageNumber="25" specimenCode="HW 1.03" specimenCount="1" specimenCount-male="1">
<specimenCount id="A396FD62FFAB983665D2EC8EFEBAF96E" box="[237,267,1747,1768]" pageId="24" pageNumber="25" type="male">1♂</specimenCount>
(
<specimenCode id="E5369E90FFAB98366427EC8EFEC2F96F" box="[280,371,1747,1769]" collectionCode="HW" pageId="24" pageNumber="25">HW 1.03</specimenCode>
) (
<collectionCode id="D381AE2EFFAB983664B8EC8EFE0DF96E" box="[391,444,1747,1768]" pageId="24" pageNumber="25">AMS</collectionCode>
K.261188, K.261189 on two slides) same data as previous
</materialsCitation>
;
<materialsCitation id="05F83CB6FFAB983664BDECB3FE51F898" ID-GBIF-Occurrence="2621712382" collectingDate="2013-07-07" collectionCode="AMS" elevation="63" latitude="-25.996" longLatPrecision="6" longitude="153.0716" pageId="24" pageNumber="25" specimenCount="24">
<specimenCount id="A396FD62FFAB983664BDECB3FDB2F882" box="[386,515,1774,1796]" pageId="24" pageNumber="25" type="generic">24 specimens</specimenCount>
(
<collectionCode id="D381AE2EFFAB98366730ECB3FDF5F885" box="[527,580,1774,1795]" pageId="24" pageNumber="25">AMS</collectionCode>
K.377745 all together in ethanol) same data as previous
</materialsCitation>
;
<materialsCitation id="05F83CB6FFAB983664D7ED54FDAAF8D2" ID-GBIF-Occurrence="2621712389" collectingDate="2016-01-27" collectionCode="AMS" latitude="-25.8991" longLatPrecision="6" longitude="153.0615" pageId="24" pageNumber="25" specimenCode="HW 1.15" specimenCount="1" specimenCount-female="1">
<specimenCount id="A396FD62FFAB983664D7ED54FDB7F898" box="[488,518,1801,1822]" pageId="24" pageNumber="25" type="female">1♀</specimenCount>
(
<specimenCode id="E5369E90FFAB9836672DED54FDDBF898" box="[530,618,1800,1822]" collectionCode="HW" pageId="24" pageNumber="25">HW 1.15</specimenCode>
) (
<collectionCode id="D381AE2EFFAB98366743ED54FD00F898" box="[636,689,1801,1822]" pageId="24" pageNumber="25">AMS</collectionCode>
K.261180, K.261181 on two slides) QLD: Carlo Point,
<geoCoordinate id="D0A4502CFFAB9836677FED7EFD15F8BF" box="[576,676,1827,1849]" degrees="25.8991" direction="south" orientation="latitude" pageId="24" pageNumber="25" precision="5" value="-25.8991">25.8991°S</geoCoordinate>
<geoCoordinate id="D0A4502CFFAB98366797ED7EFCA8F8BF" box="[680,793,1827,1849]" degrees="153.0615" direction="east" orientation="longitude" pageId="24" pageNumber="25" precision="5" value="153.0615">153.0615°E</geoCoordinate>
near sea level,
<date id="C12E102BFFAB98366411ED63FE38F8D2" box="[302,393,1854,1876]" pageId="24" pageNumber="25" value="2016-01-27">
<collectingDate id="D16AE9C3FFAB98366411ED63FE38F8D2" box="[302,393,1854,1876]" pageId="24" pageNumber="25" value="2016-01-27">27.i.2016</collectingDate>
</date>
, Graeme Smith
</materialsCitation>
;
<materialsCitation id="05F83CB6FFAB9836671BED63FDCCF8E8" ID-GBIF-Occurrence="2621712340" collectingDate="2016-01-27" collectionCode="HW" latitude="-25.8991" longLatPrecision="6" longitude="153.0615" pageId="24" pageNumber="25" specimenCode="HW 1.10" specimenCount="1" specimenCount-female="1">
<specimenCount id="A396FD62FFAB9836671BED63FDF0F8D5" box="[548,577,1854,1875]" pageId="24" pageNumber="25" type="female">1♀</specimenCount>
(
<specimenCode id="E5369E90FFAB98366773ED63FD12F8D2" box="[588,675,1854,1876]" collectionCode="HW" pageId="24" pageNumber="25">HW 1.10</specimenCode>
) (K.261136, K.261137 on two slides) same data as previous
</materialsCitation>
;
<materialsCitation id="05F83CB6FFAB983667B6ED04FD3AF80F" ID-GBIF-Occurrence="2621712317" collectingDate="2016-01-27" collectionCode="HW" latitude="-25.8991" longLatPrecision="6" longitude="153.0615" pageId="24" pageNumber="25" specimenCount="1" specimenCount-male="1">
<specimenCount id="A396FD62FFAB983667B6ED04FD19F8E8" box="[649,680,1881,1902]" pageId="24" pageNumber="25" type="male">1♂</specimenCount>
(
<collectionCode id="D381AE2EFFAB98366789ED04FD6EF8E8" box="[694,735,1881,1902]" pageId="24" pageNumber="25">HW</collectionCode>
0.95) (gbs004869 in 100% ethanol) same data as previous
</materialsCitation>
;
<materialsCitation id="05F83CB6FFAB983667ACED2EFD3AF822" ID-GBIF-Occurrence="2621712320" collectingDate="2016-01-27" collectionCode="HW" latitude="-25.8991" longLatPrecision="6" longitude="153.0615" pageId="24" pageNumber="25" specimenCode="HW 1.10" specimenCount="1" specimenCount-female="1">
<specimenCount id="A396FD62FFAB983667ACED2EFD00F80E" box="[659,689,1907,1928]" pageId="24" pageNumber="25" type="female">1♀</specimenCount>
(
<specimenCode id="E5369E90FFAB98366784ED2EFCA2F80F" box="[699,787,1907,1929]" collectionCode="HW" pageId="24" pageNumber="25">HW 1.10</specimenCode>
) (gbs004870 in 100% ethanol) same data as previous
</materialsCitation>
;
<materialsCitation id="05F83CB6FFAB983667ACEDD3FD3AF838" ID-GBIF-Occurrence="2621712349" collectingDate="2016-01-27" collectionCode="HW" latitude="-25.8991" longLatPrecision="6" longitude="153.0615" pageId="24" pageNumber="25" specimenCode="HW 1.13" specimenCount="1" specimenCount-female="1">
<specimenCount id="A396FD62FFAB983667ACEDD3FD00F825" box="[659,689,1934,1955]" pageId="24" pageNumber="25" type="female">1♀</specimenCount>
(
<specimenCode id="E5369E90FFAB98366784EDD3FCA3F822" box="[699,786,1934,1956]" collectionCode="HW" pageId="24" pageNumber="25">HW 1.13</specimenCode>
) (gbs004871 in 100% ethanol) same data as previous
</materialsCitation>
;
<materialsCitation id="05F83CB6FFAB983667ACEDF4FD3AF85F" ID-GBIF-Occurrence="2621712327" collectingDate="2016-01-27" collectionCode="HW" latitude="-25.8991" longLatPrecision="6" longitude="153.0615" pageId="24" pageNumber="25" specimenCount="1" specimenCount-female="1">
<specimenCount id="A396FD62FFAB983667ACEDF4FD00F838" box="[659,689,1961,1982]" pageId="24" pageNumber="25" type="female">1♀</specimenCount>
(
<collectionCode id="D381AE2EFFAB98366784EDF4FD55F838" box="[699,740,1961,1982]" pageId="24" pageNumber="25">HW</collectionCode>
0.98) (gbs004872 in 100% ethanol) same data as previous
</materialsCitation>
;
<materialsCitation id="05F83CB6FFAB983667ACED9EFA06F91F" ID-GBIF-Occurrence="2621712374" box="[659,1463,1667,2009]" collectingDate="2016-01-27" collectionCode="AMS" latitude="-25.8991" longLatPrecision="6" longitude="153.0615" pageId="24" pageNumber="25" specimenCode="HW 1.05" specimenCount="1" specimenCount-male="1">
<specimenCount id="A396FD62FFAB983667ACED9EFD01F85E" box="[659,688,1987,2008]" pageId="24" pageNumber="25" type="male">1♂</specimenCount>
(
<specimenCode id="E5369E90FFAB98366784ED9EFCA3F85F" box="[699,786,1987,2009]" collectionCode="HW" pageId="24" pageNumber="25">HW 1.05</specimenCode>
) (
<collectionCode id="D381AE2EFFAB98366671ECDEFC32F91E" box="[846,899,1667,1688]" pageId="24" pageNumber="25">AMS</collectionCode>
K.261138, K.261139 on two slides) same data as previous
</materialsCitation>
;
<materialsCitation id="05F83CB6FFAB98366678ECC3FA07F932" ID-GBIF-Occurrence="2621712346" box="[839,1462,1694,1716]" collectingDate="2016-01-27" collectionCode="HW" latitude="-25.8991" longLatPrecision="6" longitude="153.0615" pageId="24" pageNumber="25" specimenCount="1" specimenCount-male="1">
<specimenCount id="A396FD62FFAB98366678ECC3FCD5F935" box="[839,868,1694,1715]" pageId="24" pageNumber="25" type="male">1♂</specimenCount>
(
<collectionCode id="D381AE2EFFAB98366650ECC3FC29F935" box="[879,920,1694,1715]" pageId="24" pageNumber="25">HW</collectionCode>
0.98) (gbs004874 in 100% ethanol) same data as previous
</materialsCitation>
;
<materialsCitation id="05F83CB6FFAB98366678ECE4FA07F948" ID-GBIF-Occurrence="2621712355" box="[839,1462,1720,1743]" collectingDate="2016-01-27" collectionCode="HW" latitude="-25.8991" longLatPrecision="6" longitude="153.0615" pageId="24" pageNumber="25" specimenCount="1" specimenCount-male="1">
<specimenCount id="A396FD62FFAB98366678ECE4FCD5F948" box="[839,868,1721,1742]" pageId="24" pageNumber="25" type="male">1♂</specimenCount>
(
<collectionCode id="D381AE2EFFAB98366650ECE4FC29F948" box="[879,920,1721,1742]" pageId="24" pageNumber="25">HW</collectionCode>
1.00) (gbs004875 in 100% ethanol) same data as previous
</materialsCitation>
;
<materialsCitation id="05F83CB6FFAB98366678EC8EFA07F96F" ID-GBIF-Occurrence="2621712353" box="[839,1462,1747,1769]" collectingDate="2016-01-27" collectionCode="HW" latitude="-25.8991" longLatPrecision="6" longitude="153.0615" pageId="24" pageNumber="25" specimenCount="1" specimenCount-female="1">
<specimenCount id="A396FD62FFAB98366678EC8EFCD4F96E" box="[839,869,1747,1768]" pageId="24" pageNumber="25" type="female">1♀</specimenCount>
(
<collectionCode id="D381AE2EFFAB98366650EC8EFC29F96E" box="[879,920,1747,1768]" pageId="24" pageNumber="25">HW</collectionCode>
0.90) (gbs004876 in 100% ethanol) same data as previous
</materialsCitation>
;
<materialsCitation id="05F83CB6FFAB98366678ECB3FC02F898" ID-GBIF-Occurrence="2621712333" collectingDate="2016-01-27" collectionCode="HW, AMS" latitude="-25.8991" longLatPrecision="6" longitude="153.0615" pageId="24" pageNumber="25" specimenCount="1" specimenCount-male="1">
<specimenCount id="A396FD62FFAB98366678ECB3FCD5F885" box="[839,868,1774,1795]" pageId="24" pageNumber="25" type="male">1♂</specimenCount>
(
<collectionCode id="D381AE2EFFAB98366650ECB3FC29F885" box="[879,920,1774,1795]" pageId="24" pageNumber="25">HW</collectionCode>
0.95) (
<collectionCode id="D381AE2EFFAB983666E7ECB3FBBCF885" box="[984,1037,1774,1795]" pageId="24" pageNumber="25">AMS</collectionCode>
K.261140, K.261141 on two slides) same data as previous
</materialsCitation>
;
<materialsCitation id="05F83CB6FFAB98366684ED54FC08F8BF" ID-GBIF-Occurrence="2621712376" collectingDate="2016-01-27" collectionCode="HW" latitude="-25.8991" longLatPrecision="6" longitude="153.0615" pageId="24" pageNumber="25" specimenCount="1" specimenCount-female="1">
<specimenCount id="A396FD62FFAB98366684ED54FC68F898" box="[955,985,1801,1822]" pageId="24" pageNumber="25" type="female">1♀</specimenCount>
(
<collectionCode id="D381AE2EFFAB983666DBED54FBBCF898" box="[996,1037,1801,1822]" pageId="24" pageNumber="25">HW</collectionCode>
0.83) (gbs004878 in 100% ethanol) same data as previous
</materialsCitation>
;
<materialsCitation id="05F83CB6FFAB983666FAED7EFBEDF8D2" ID-GBIF-Occurrence="2621712356" collectingDate="2016-01-27" collectionCode="HW, AMS" latitude="-25.8991" longLatPrecision="6" longitude="153.0615" pageId="24" pageNumber="25" specimenCount="1" specimenCount-female="1">
<specimenCount id="A396FD62FFAB983666FAED7EFC52F8BE" box="[965,995,1827,1848]" pageId="24" pageNumber="25" type="female">1♀</specimenCount>
(
<collectionCode id="D381AE2EFFAB983666CDED7EFBAAF8BE" box="[1010,1051,1827,1848]" pageId="24" pageNumber="25">HW</collectionCode>
0.93) (
<collectionCode id="D381AE2EFFAB9836615AED7EFB2BF8BE" box="[1125,1178,1827,1848]" pageId="24" pageNumber="25">AMS</collectionCode>
K.261182, K.261183 on two slides) same data as previous
</materialsCitation>
;
<materialsCitation id="05F83CB6FFAB9836615BED63FB7BF8E8" ID-GBIF-Occurrence="2621712348" collectingDate="2016-01-27" collectionCode="HW" latitude="-25.8991" longLatPrecision="6" longitude="153.0615" pageId="24" pageNumber="25" specimenCount="2" specimenCount-female="1" specimenCount-juvenile="1">
<specimenCount id="A396FD62FFAB9836615BED63FB72F8D2" box="[1124,1219,1854,1876]" pageId="24" pageNumber="25" type="juvenile">1 juvenile</specimenCount>
<specimenCount id="A396FD62FFAB983661F8ED63FB68F8D5" box="[1223,1241,1854,1875]" pageId="24" pageNumber="25" type="female"></specimenCount>
(
<collectionCode id="D381AE2EFFAB983661DAED63FABFF8D5" box="[1253,1294,1854,1875]" pageId="24" pageNumber="25">HW</collectionCode>
0.88) (gbs004880 in 100% ethanol) same data as previous
</materialsCitation>
;
<materialsCitation id="05F83CB6FFAB983661EBED04FAFCF80F" ID-GBIF-Occurrence="2621712337" collectingDate="2016-01-27" collectionCode="HW" latitude="-25.8991" longLatPrecision="6" longitude="153.0615" pageId="24" pageNumber="25" specimenCount="2" specimenCount-juvenile="1" specimenCount-male="1">
<specimenCount id="A396FD62FFAB983661EBED04FA85F8E8" box="[1236,1332,1880,1902]" pageId="24" pageNumber="25" type="juvenile">1 juvenile</specimenCount>
<specimenCount id="A396FD62FFAB98366005ED04FAFDF8E8" box="[1338,1356,1881,1902]" pageId="24" pageNumber="25" type="male"></specimenCount>
(
<collectionCode id="D381AE2EFFAB98366066ED04FA33F8E8" box="[1369,1410,1881,1902]" pageId="24" pageNumber="25">HW</collectionCode>
0.83) (gbs004881 in 100% ethanol) same data as previous
</materialsCitation>
;
<materialsCitation id="05F83CB6FFAB98366066ED2EFA25F822" ID-GBIF-Occurrence="2621712332" collectingDate="2016-01-27" collectionCode="HW" latitude="-25.8991" longLatPrecision="6" longitude="153.0615" pageId="24" pageNumber="25" specimenCount="1" specimenCount-juvenile="1">
<specimenCount id="A396FD62FFAB98366066ED2EFA0AF80F" box="[1369,1467,1907,1929]" pageId="24" pageNumber="25" type="juvenile">1 juvenile</specimenCount>
(
<collectionCode id="D381AE2EFFAB98366672EDD3FCC7F825" box="[845,886,1934,1955]" pageId="24" pageNumber="25">HW</collectionCode>
0.75) (gbs004882 in 100% ethanol) same data as previous
</materialsCitation>
;
<materialsCitation id="05F83CB6FFAB983660A2EDD3FA19F85F" ID-GBIF-Occurrence="2621712304" collectingDate="2016-01-27" collectionCode="HW, AMS" latitude="-25.8991" longLatPrecision="6" longitude="153.0616" pageId="24" pageNumber="25" specimenCount="1" specimenCount-male="1">
<specimenCount id="A396FD62FFAB983660A2EDD3FA0AF825" box="[1437,1467,1934,1955]" pageId="24" pageNumber="25" type="male">1♂</specimenCount>
(
<collectionCode id="D381AE2EFFAB98366672EDF4FCC7F838" box="[845,886,1961,1982]" pageId="24" pageNumber="25">HW</collectionCode>
0.90) (
<collectionCode id="D381AE2EFFAB98366683EDF4FC40F838" box="[956,1009,1961,1982]" pageId="24" pageNumber="25">AMS</collectionCode>
K.
<quantity id="72689B0EFFAB98366133EDF5FBDEF838" box="[1036,1135,1960,1982]" metricMagnitude="3" metricUnit="m" metricValue="9.5949262" pageId="24" pageNumber="25" unit="in" value="377753.0">377753 in</quantity>
100% ethanol) QLD: Carlo Point,
<geoCoordinate id="D0A4502CFFAB98366678ED9EFC1AF85E" box="[839,939,1987,2009]" degrees="25.8991" direction="south" orientation="latitude" pageId="24" pageNumber="25" precision="5" value="-25.8991">25.8991°S</geoCoordinate>
<geoCoordinate id="D0A4502CFFAB9836668EED9EFB92F85F" box="[945,1059,1987,2009]" degrees="153.0616" direction="east" orientation="longitude" pageId="24" pageNumber="25" precision="5" value="153.0616">153.0616°E</geoCoordinate>
near sea level,
<date id="C12E102BFFAB9836618AED9EFAA3F85F" box="[1205,1298,1987,2009]" pageId="24" pageNumber="25" value="2016-01-27">
<collectingDate id="D16AE9C3FFAB9836618AED9EFAA3F85F" box="[1205,1298,1987,2009]" pageId="24" pageNumber="25" value="2016-01-27">27.i.2016</collectingDate>
</date>
, Graeme Smith
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="E1EF6663FFAB98366599EC5AFE15F9D4" ID-DOI="http://doi.org/10.5281/zenodo.3838007" ID-Zenodo-Dep="3838007" httpUri="https://zenodo.org/record/3838007/files/figure.png" pageId="24" pageNumber="25" startId="24.[166,243,1543,1565]" targetBox="[177,1452,156,1521]" targetPageId="24">
<paragraph id="B52F36EBFFAB98366599EC5AFE15F9D4" blockId="24.[166,1466,1543,1618]" pageId="24" pageNumber="25">
<emphasis id="87E4EAF9FFAB98366599EC5AFE86F99A" bold="true" box="[166,311,1543,1565]" pageId="24" pageNumber="25">Figures 6065</emphasis>
.
<taxonomicName id="72904D68FFAB9836647CEC5AFD9AF99A" authority="Smith &amp; Mitchell &amp; Lee &amp; Espinasa, 2019" authorityName="Smith &amp; Mitchell &amp; Lee &amp; Espinasa" authorityYear="2019" box="[323,555,1543,1564]" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="cooloola" status="sp. nov.">
<emphasis id="87E4EAF9FFAB9836647CEC5AFD9AF99A" box="[323,555,1543,1564]" italics="true" pageId="24" pageNumber="25">Heterolepisma cooloola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9CD75782FFAB9836670DEC5AFDCFF99A" box="[562,638,1543,1564]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
holotype ♀
<emphasis id="87E4EAF9FFAB983667C6EC5AFC91F99A" box="[761,800,1543,1564]" italics="true" pageId="24" pageNumber="25">(60)</emphasis>
presternum, prothoracic sternum and PI;
<emphasis id="87E4EAF9FFAB98366188EC5AFB6EF99A" box="[1207,1247,1543,1564]" italics="true" pageId="24" pageNumber="25">(61)</emphasis>
mesothoracic sternum and PII;
<emphasis id="87E4EAF9FFAB983665C4EC7FFE93F9B1" box="[251,290,1570,1591]" italics="true" pageId="24" pageNumber="25">(62)</emphasis>
apex of mesothoracic sternum;
<emphasis id="87E4EAF9FFAB98366764EC7FFD32F9B1" box="[603,643,1570,1591]" italics="true" pageId="24" pageNumber="25">(63)</emphasis>
metathoracic sternum and PIII;
<emphasis id="87E4EAF9FFAB98366681EC7FFC54F9B1" box="[958,997,1570,1591]" italics="true" pageId="24" pageNumber="25">(64)</emphasis>
apex of metathoracic sternum;
<emphasis id="87E4EAF9FFAB98366025EC7FFAF3F9B1" box="[1306,1346,1570,1591]" italics="true" pageId="24" pageNumber="25">(65)</emphasis>
pretarsus of PIII. Scale bars = 0.1 mm.
</paragraph>
</caption>
<caption id="E1EF6663FFAA98376546EC09FB73F93C" ID-DOI="http://doi.org/10.5281/zenodo.3838009" ID-Zenodo-Dep="3838009" httpUri="https://zenodo.org/record/3838009/files/figure.png" pageId="25" pageNumber="26" startId="25.[121,198,1620,1642]" targetBox="[133,1412,157,1605]" targetPageId="25">
<paragraph id="B52F36EBFFAA98376546EC09FB73F93C" blockId="25.[121,1421,1620,1722]" pageId="25" pageNumber="26">
<emphasis id="87E4EAF9FFAA98376546EC09FEB8F9EF" bold="true" box="[121,265,1620,1642]" pageId="25" pageNumber="26">Figures 6677</emphasis>
.
<taxonomicName id="72904D68FFAA98376428EC09FE4FF9EF" authority="Smith &amp; Mitchell &amp; Lee &amp; Espinasa, 2019" authorityName="Smith &amp; Mitchell &amp; Lee &amp; Espinasa" authorityYear="2019" box="[279,510,1620,1641]" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="cooloola" status="sp. nov.">
<emphasis id="87E4EAF9FFAA98376428EC09FE4FF9EF" box="[279,510,1620,1641]" italics="true" pageId="25" pageNumber="26">Heterolepisma cooloola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9CD75782FFAA98376739EC08FDE3F9EC" box="[518,594,1621,1642]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
holotype ♀, unless otherwise indicated by specimen number
<emphasis id="87E4EAF9FFAA98376193EC09FB65F9EF" box="[1196,1236,1620,1641]" italics="true" pageId="25" pageNumber="26">(66)</emphasis>
urotergite IV;
<emphasis id="87E4EAF9FFAA9837605AEC09FA3CF9EF" box="[1381,1421,1620,1641]" italics="true" pageId="25" pageNumber="26">(67)</emphasis>
urotergite V, right lateral comb;
<emphasis id="87E4EAF9FFAA983764A1EC32FE74F902" box="[414,453,1647,1668]" italics="true" pageId="25" pageNumber="26">(68)</emphasis>
idem, left sublateral comb;
<emphasis id="87E4EAF9FFAA983767FAEC32FD5DF902" box="[709,748,1647,1668]" italics="true" pageId="25" pageNumber="26">(69)</emphasis>
idem, left submedial comb;
<emphasis id="87E4EAF9FFAA983766CEEC32FBA9F902" box="[1009,1048,1647,1668]" italics="true" pageId="25" pageNumber="26">(70)</emphasis>
right side of urotergite VIII (K.261189);
<emphasis id="87E4EAF9FFAA98376546ECD7FF10F919" box="[121,161,1674,1695]" italics="true" pageId="25" pageNumber="26">(71)</emphasis>
infralateral comb of urotergite IX;
<emphasis id="87E4EAF9FFAA983764CCECD7FDAAF919" box="[499,539,1674,1695]" italics="true" pageId="25" pageNumber="26">(72)</emphasis>
urotergite X;
<emphasis id="87E4EAF9FFAA9837679EECD7FD7BF919" box="[673,714,1674,1695]" italics="true" pageId="25" pageNumber="26">(73)</emphasis>
urotergite X of paratype (QM 207012);
<emphasis id="87E4EAF9FFAA98376173ECD7FBC5F919" box="[1100,1140,1674,1695]" italics="true" pageId="25" pageNumber="26">(74)</emphasis>
urosternite IV;
<emphasis id="87E4EAF9FFAA98376035ECD7FA83F919" box="[1290,1330,1674,1695]" italics="true" pageId="25" pageNumber="26">(75)</emphasis>
posterior comb of urosternite V;
<emphasis id="87E4EAF9FFAA98376468ECF9FECFF93F" box="[343,382,1700,1721]" italics="true" pageId="25" pageNumber="26">(76)</emphasis>
urosternites VII, VIII, IX and ovipositor;
<emphasis id="87E4EAF9FFAA9837662DECF9FC8BF93F" box="[786,826,1700,1721]" italics="true" pageId="25" pageNumber="26">(77)</emphasis>
base of stylus VII. Scale bars = 0.1 mm.
</paragraph>
</caption>
<subSubSection id="FD8A6560FFAA98376546ED5FFC51F82B" pageId="25" pageNumber="26" type="diagnosis">
<paragraph id="B52F36EBFFAA98376546ED5FFC51F82B" blockId="25.[121,749,1794,1994]" lastBlockId="25.[794,1422,1794,1965]" pageId="25" pageNumber="26">
<emphasis id="87E4EAF9FFAA98376546ED5FFF57F89C" bold="true" box="[121,230,1794,1818]" pageId="25" pageNumber="26">Diagnosis</emphasis>
. This species differs from other described species of
<taxonomicName id="72904D68FFAA983765A9ED7DFE83F8BE" authorityName="Escherich" authorityYear="1905" box="[150,306,1824,1848]" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="87E4EAF9FFAA983765A9ED7DFE83F8BE" box="[150,306,1824,1848]" italics="true" pageId="25" pageNumber="26">Heterolepisma</emphasis>
</taxonomicName>
that also have 2+2 combs on urotergite I and three pairs of styli in the female and only two in the male, by the presence of lanceolate scales on the femora, tibiae and clypeus, the straight anterior margin of the head devoid of macrochaetae, the single macrochaeta mediad of the anterior trichobothrium on the pronotum, the mesosternum is also slightly different with a wider glabrous apex and the combs more compact, the posterior margin of the metasternum is rounded with a comparatively wide glabrous gap with small 1+1 combs of two to three macrochaetae. The parabolic shape of urotergite X is narrower and the terminal filaments evenly pigmented.
</paragraph>
</subSubSection>
<caption id="E1EF6663FFA998346599EF7EFE66FA0F" ID-DOI="http://doi.org/10.5281/zenodo.3838011" ID-Zenodo-Dep="3838011" httpUri="https://zenodo.org/record/3838011/files/figure.png" pageId="26" pageNumber="27" startId="26.[166,243,1315,1337]" targetBox="[177,1457,158,1295]" targetPageId="26">
<paragraph id="B52F36EBFFA998346599EF7EFE66FA0F" blockId="26.[166,1467,1315,1417]" pageId="26" pageNumber="27">
<emphasis id="87E4EAF9FFA998346599EF7EFE85FABE" bold="true" box="[166,308,1315,1337]" pageId="26" pageNumber="27">Figures 7885</emphasis>
.
<taxonomicName id="72904D68FFA998346401EF7EFD92FABE" authority="Smith &amp; Mitchell &amp; Lee &amp; Espinasa, 2019" authorityName="Smith &amp; Mitchell &amp; Lee &amp; Espinasa" authorityYear="2019" box="[318,547,1315,1336]" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="cooloola" status="sp. nov.">
<emphasis id="87E4EAF9FFA998346401EF7EFD92FABE" box="[318,547,1315,1336]" italics="true" pageId="26" pageNumber="27">Heterolepisma cooloola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9CD75782FFA998346717EF79FDC0FABF" box="[552,625,1316,1337]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
holotype ♀ unless otherwise indicated by specimen number
<emphasis id="87E4EAF9FFA998346192EF7EFB64FABE" box="[1197,1237,1315,1336]" italics="true" pageId="26" pageNumber="27">(78)</emphasis>
stylet IX (QM 207012);
<emphasis id="87E4EAF9FFA998346599EF63FF7FFAD5" box="[166,206,1342,1363]" italics="true" pageId="26" pageNumber="27">(79)</emphasis>
apex of anterior gonapophysis;
<emphasis id="87E4EAF9FFA998346734EF63FD83FAD5" box="[523,562,1342,1363]" italics="true" pageId="26" pageNumber="27">(80)</emphasis>
apex of posterior gonapophysis;
<emphasis id="87E4EAF9FFA998346645EF63FC13FAD5" box="[890,930,1342,1363]" italics="true" pageId="26" pageNumber="27">(81)</emphasis>
bases of terminal filaments;
<emphasis id="87E4EAF9FFA998346181EF63FB57FAD5" box="[1214,1254,1342,1363]" italics="true" pageId="26" pageNumber="27">(82)</emphasis>
most distal surviving divisions of cerci;
<emphasis id="87E4EAF9FFA998346465EF04FE33FAE8" box="[346,386,1369,1390]" italics="true" pageId="26" pageNumber="27">(83)</emphasis>
most distal surviving divisions of median filament;
<emphasis id="87E4EAF9FFA998346642EF04FC15FAE8" box="[893,932,1369,1390]" italics="true" pageId="26" pageNumber="27">(84)</emphasis>
coxites IX, styli and penis of male (QM 207012);
<emphasis id="87E4EAF9FFA9983460ADEF04FA0BFAE8" box="[1426,1466,1369,1390]" italics="true" pageId="26" pageNumber="27">(85)</emphasis>
paramere. Scale bars = 0.1 mm.
</paragraph>
</caption>
<subSubSection id="FD8A6560FFA998326599EFEFFBB9F8F0" lastPageId="28" lastPageNumber="29" pageId="26" pageNumber="27" type="description">
<paragraph id="B52F36EBFFA998346599EFEFFE9BFA4C" blockId="26.[166,298,1458,1482]" box="[166,298,1458,1482]" pageId="26" pageNumber="27">
<emphasis id="87E4EAF9FFA998346599EFEFFE9BFA4C" bold="true" box="[166,298,1458,1482]" pageId="26" pageNumber="27">Description</emphasis>
</paragraph>
<paragraph id="B52F36EBFFA998346599EFBDFDE7F9EB" blockId="26.[166,794,1503,1645]" pageId="26" pageNumber="27">
<emphasis id="87E4EAF9FFA998346599EFBDFE96FA71" box="[166,295,1504,1527]" italics="true" pageId="26" pageNumber="27">Appearance</emphasis>
: Medium to large silverfish, scale covering in life uniform or slightly mottled grey with brown antennae, terminal filaments slightly darker than antennae with only a small portion of each annulus bearing the larger macrochaetae lighter in colour (
<figureCitation id="2DAB2A6EFFA9983464C5EC08FDFBF9EB" box="[506,586,1621,1645]" captionStart="Figure 37" captionStartId="21.[794,863,1954,1976]" captionTargetBox="[794,1421,971,1946]" captionTargetId="figure@21.[794,1421,971,1946]" captionTargetPageId="21" captionText="Figure 37. Heterolepisma cooloola sp. nov. from leaf litter at Carlo Point." figureDoi="http://doi.org/10.5281/zenodo.3837999" httpUri="https://zenodo.org/record/3837999/files/figure.png" pageId="26" pageNumber="27">Fig. 37</figureCitation>
).
</paragraph>
<paragraph id="B52F36EBFFA998346599ECDFFE72F85B" blockId="26.[166,794,1666,2013]" pageId="26" pageNumber="27">
<emphasis id="87E4EAF9FFA998346599ECDFFE92F91C" box="[166,291,1666,1690]" italics="true" pageId="26" pageNumber="27">Body length</emphasis>
: H+B up to
<quantity id="72689B0EFFA9983464A0ECDFFE5FF91C" box="[415,494,1666,1690]" metricMagnitude="-3" metricUnit="m" metricValue="9.9" pageId="26" pageNumber="27" unit="mm" value="9.9">9.9 mm</quantity>
(
<specimenCount id="A396FD62FFA9983464C2ECDFFDBFF91C" box="[509,526,1666,1690]" pageId="26" pageNumber="27" type="female"></specimenCount>
)
<quantity id="72689B0EFFA998346722ECDFFDC8F91C" box="[541,633,1666,1690]" metricMagnitude="-3" metricUnit="m" metricValue="8.25" pageId="26" pageNumber="27" unit="mm" value="8.25">8.25 mm</quantity>
(
<specimenCount id="A396FD62FFA9983467B9ECDFFD2DF91C" box="[646,668,1666,1690]" pageId="26" pageNumber="27" type="male"></specimenCount>
); maximum HW
<quantity id="72689B0EFFA9983465E6ECFDFE8DF931" box="[217,316,1696,1720]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="26" pageNumber="27" unit="mm" value="1.2">1.20 mm</quantity>
; thorax: length up to
<quantity id="72689B0EFFA99834671DECC2FD30F931" box="[546,641,1695,1719]" metricMagnitude="-3" metricUnit="m" metricValue="2.85" pageId="26" pageNumber="27" unit="mm" value="2.85">2.85 mm</quantity>
(or 0.260.32 H+B); width up to
<quantity id="72689B0EFFA998346449ECE0FE69F952" box="[374,472,1725,1749]" metricMagnitude="-3" metricUnit="m" metricValue="1.93" pageId="26" pageNumber="27" unit="mm" value="1.93">1.93 mm</quantity>
, usually slightly widest at the mesonotum; antennae damaged in all specimens, maximum preserved length of antenna
<quantity id="72689B0EFFA9983464F3ECAAFDAAF889" box="[460,539,1783,1807]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="26" pageNumber="27" unit="mm" value="5.6">5.6 mm</quantity>
(or 0.68 H+B); terminal filaments damaged in all specimens, maximum preserved length of cercus
<quantity id="72689B0EFFA99834646FED6FFE2EF8CC" box="[336,415,1842,1866]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="26" pageNumber="27" unit="mm" value="3.6">3.6 mm</quantity>
(or 0.48 H+B); maximum preserved length of median dorsal appendage 5.00 mm (or 0.60 H+B). Body neither elongate nor broad (
<figureCitation id="2DAB2A6EFFA9983464C3ED30FDF7F803" box="[508,582,1901,1925]" captionStart="Figures 3848" captionStartId="22.[166,243,1805,1827]" captionTargetBox="[174,1460,196,1772]" captionTargetId="figure@22.[166,1466,185,1777]" captionTargetPageId="22" captionText="Figures 3848. Heterolepisma cooloola sp. nov. holotype ♀, unless indicated otherwise by specimen number(38) habitus (K.377754); (39) posterior comb of pronotum with scale; (40) head; (41) scape, pedicel and basal intervals of flagellum, from above; (42) idem, from below; (43) antenna, most distal surviving interval; (44) mandible; (45) idem, detail of molar and incisor regions; (46) maxilla, only larger setae of palp illustrated; (47) idem, lacinia and galea (QM 207012); (48) ultimate article of maxillary palp of ♂ (QM 207012). Scale bars = 0.1 mm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.3838001" httpUri="https://zenodo.org/record/3838001/files/figure.png" pageId="26" pageNumber="27">Fig. 38</figureCitation>
) with thorax slightly wider than abdominal segment I, the following abdominal segments about the same width until the fourth or fifth after which it tapers posteriorly.
</paragraph>
<paragraph id="B52F36EBFFA998356678EFEEFEFDFEF9" blockId="26.[839,1467,1459,2011]" lastBlockId="27.[121,749,154,383]" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
<emphasis id="87E4EAF9FFA998346678EFEEFC6DFA4C" box="[839,988,1459,1482]" italics="true" pageId="26" pageNumber="27">Pigmentation</emphasis>
: Pigment light chestnut-brown in alcohol preserved specimens, stronger around peri-antennal and supra-ocular lines of macrochaetae and along the band of setae on the clypeus (especially laterally); pedicel and scape very lightly pigmented distally, rest of flagellum uniformly lightly pigmented becoming somewhat darker distally; all articles of maxillary palp with pigment except the most distal article, densest on article three especially distally; labium with lines of pigment around macrochaetae across the mentum, present on distal three articles of labial palp being stronger on the edges, pigment of ultimate article mostly in basal half but with a noticeable line above the more distal row of papillae. Nota with some pigment anteriorly and along margins. Legs with pigmentation along outer edge of precoxa of PI, along the length of the outer margin among the macrochaetae and only very faintly along the inner margin, distally; trochanter with light patch on margin distally; femur pigmented, darkest distally along dorsal margin and around bulge on ventral margin; tibia pigmented along edges being a little darker distally; first tarsal article pigmented distally. Urotergite X with very faint pigment along anterior lateral margins. Styli IX slightly pigmented in distal three quarters; other styli with very little or no pigment. Ovipositor with yellowish hue. Terminal filaments lightly pigmented basally becoming a lot darker distally. Some individuals show greater or lesser levels of pigmentation, with less pigmentation in juvenile specimens.
</paragraph>
<paragraph id="B52F36EBFFA898356546EBC9FEBEFE4C" blockId="27.[121,749,404,458]" pageId="27" pageNumber="28">
<emphasis id="87E4EAF9FFA898356546EBC9FEA3FE2A" box="[121,274,404,428]" italics="true" pageId="27" pageNumber="28">Macrochaetae</emphasis>
: Bifid apically, or simple, hyaline, light to darker brown.
</paragraph>
<paragraph id="B52F36EBFFA898356546EB82FEF3FD42" blockId="27.[121,749,479,708]" pageId="27" pageNumber="28">
<emphasis id="87E4EAF9FFA898356546EB82FF0CFE71" box="[121,189,479,503]" italics="true" pageId="27" pageNumber="28">Scales</emphasis>
: Unevenly rounded or ovoid, with numerous parallel dark brown ribs, that do not extend beyond the margin (
<figureCitation id="2DAB2A6EFFA8983567FEEBA1FF26FDB4" captionStart="Figures 3848" captionStartId="22.[166,243,1805,1827]" captionTargetBox="[174,1460,196,1772]" captionTargetId="figure@22.[166,1466,185,1777]" captionTargetPageId="22" captionText="Figures 3848. Heterolepisma cooloola sp. nov. holotype ♀, unless indicated otherwise by specimen number(38) habitus (K.377754); (39) posterior comb of pronotum with scale; (40) head; (41) scape, pedicel and basal intervals of flagellum, from above; (42) idem, from below; (43) antenna, most distal surviving interval; (44) mandible; (45) idem, detail of molar and incisor regions; (46) maxilla, only larger setae of palp illustrated; (47) idem, lacinia and galea (QM 207012); (48) ultimate article of maxillary palp of ♂ (QM 207012). Scale bars = 0.1 mm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.3838001" httpUri="https://zenodo.org/record/3838001/files/figure.png" pageId="27" pageNumber="28">Fig. 39</figureCitation>
); in alcohol dorsal scales and the more lateral scales of the urosternites with dark brown ribs; ventrally mostly hyaline but those towards the lateral margins with light brown ribs. Lanceolate scales present on clypeus, femora and tibia. Scales absent from flagellum of antennae, mouthparts and terminal filaments.
</paragraph>
<paragraph id="B52F36EBFFA898356546E887FB43FF49" blockId="27.[121,749,730,1986]" lastBlockId="27.[794,1421,154,207]" pageId="27" pageNumber="28">
<emphasis id="87E4EAF9FFA898356546E887FF03FD74" box="[121,178,730,754]" italics="true" pageId="27" pageNumber="28">Head</emphasis>
: Wider than long (
<figureCitation id="2DAB2A6EFFA898356442E887FE7DFD74" box="[381,460,730,754]" captionStart="Figures 3848" captionStartId="22.[166,243,1805,1827]" captionTargetBox="[174,1460,196,1772]" captionTargetId="figure@22.[166,1466,185,1777]" captionTargetPageId="22" captionText="Figures 3848. Heterolepisma cooloola sp. nov. holotype ♀, unless indicated otherwise by specimen number(38) habitus (K.377754); (39) posterior comb of pronotum with scale; (40) head; (41) scape, pedicel and basal intervals of flagellum, from above; (42) idem, from below; (43) antenna, most distal surviving interval; (44) mandible; (45) idem, detail of molar and incisor regions; (46) maxilla, only larger setae of palp illustrated; (47) idem, lacinia and galea (QM 207012); (48) ultimate article of maxillary palp of ♂ (QM 207012). Scale bars = 0.1 mm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.3838001" httpUri="https://zenodo.org/record/3838001/files/figure.png" pageId="27" pageNumber="28">Fig. 40</figureCitation>
) with marginal rows about two macrochaetae wide along the sides of the vertex, but without macrochaetae along the anterior margin, the lateral rows extending back along the margin to the eyes and extending as a single short row above the eyes, as well as a small peri-antennal group isolated from the marginal. Clypeus with numerous setae, some long and thin, others more robust but not forming combs; with a few lanceolate scales scattered among the setae. Labrum with thin setae only. Scales on top of head, those along the anterior margin overhanging the margin. Eyes dark, composed of 12 ommatidia.—Antennae long, about ⅔ H+B, scape with a subdistal rosette of setae, very conspicuous from above (
<figureCitation id="2DAB2A6EFFA898356482EE67FDBBFBD4" box="[445,522,1082,1106]" captionStart="Figures 3848" captionStartId="22.[166,243,1805,1827]" captionTargetBox="[174,1460,196,1772]" captionTargetId="figure@22.[166,1466,185,1777]" captionTargetPageId="22" captionText="Figures 3848. Heterolepisma cooloola sp. nov. holotype ♀, unless indicated otherwise by specimen number(38) habitus (K.377754); (39) posterior comb of pronotum with scale; (40) head; (41) scape, pedicel and basal intervals of flagellum, from above; (42) idem, from below; (43) antenna, most distal surviving interval; (44) mandible; (45) idem, detail of molar and incisor regions; (46) maxilla, only larger setae of palp illustrated; (47) idem, lacinia and galea (QM 207012); (48) ultimate article of maxillary palp of ♂ (QM 207012). Scale bars = 0.1 mm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.3838001" httpUri="https://zenodo.org/record/3838001/files/figure.png" pageId="27" pageNumber="28">Fig. 41</figureCitation>
), and numerous setae along the sides and over the ventral face (
<figureCitation id="2DAB2A6EFFA898356705EE0AFD3AFBE9" box="[570,651,1111,1135]" captionStart="Figures 3848" captionStartId="22.[166,243,1805,1827]" captionTargetBox="[174,1460,196,1772]" captionTargetId="figure@22.[166,1466,185,1777]" captionTargetPageId="22" captionText="Figures 3848. Heterolepisma cooloola sp. nov. holotype ♀, unless indicated otherwise by specimen number(38) habitus (K.377754); (39) posterior comb of pronotum with scale; (40) head; (41) scape, pedicel and basal intervals of flagellum, from above; (42) idem, from below; (43) antenna, most distal surviving interval; (44) mandible; (45) idem, detail of molar and incisor regions; (46) maxilla, only larger setae of palp illustrated; (47) idem, lacinia and galea (QM 207012); (48) ultimate article of maxillary palp of ♂ (QM 207012). Scale bars = 0.1 mm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.3838001" httpUri="https://zenodo.org/record/3838001/files/figure.png" pageId="27" pageNumber="28">Fig. 42</figureCitation>
); pedicel short, 0.42 times the length of the scape (range 0.330.53), with many setae mostly distally and on the ventral face; the most apical annulus of each interval in the distal end of the flagellum with a few small inconspicuous rod-like basiconic sensilla (type B of
<bibRefCitation id="D1014B1AFFA898356402EEB7FE05FA84" author="Adel" box="[317,436,1258,1282]" firstAuthor="Adel" pageId="27" pageNumber="28" pagination="191 - 217" refId="ref25550" refString="Adel, T. 1984. Sensilleninventar und sensillenmuster auf den Antennen von Thermobia domestica und Lepisma saccharina (Insecta: Zygentoma). Braunschweiger Naturkundliche Schriften 2: 191 - 217." type="journal article" year="1984">Adel, 1984</bibRefCitation>
) (
<figureCitation id="2DAB2A6EFFA8983564F6EEB7FDA5FA84" box="[457,532,1258,1282]" captionStart="Figures 3848" captionStartId="22.[166,243,1805,1827]" captionTargetBox="[174,1460,196,1772]" captionTargetId="figure@22.[166,1466,185,1777]" captionTargetPageId="22" captionText="Figures 3848. Heterolepisma cooloola sp. nov. holotype ♀, unless indicated otherwise by specimen number(38) habitus (K.377754); (39) posterior comb of pronotum with scale; (40) head; (41) scape, pedicel and basal intervals of flagellum, from above; (42) idem, from below; (43) antenna, most distal surviving interval; (44) mandible; (45) idem, detail of molar and incisor regions; (46) maxilla, only larger setae of palp illustrated; (47) idem, lacinia and galea (QM 207012); (48) ultimate article of maxillary palp of ♂ (QM 207012). Scale bars = 0.1 mm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.3838001" httpUri="https://zenodo.org/record/3838001/files/figure.png" pageId="27" pageNumber="28">Fig. 43</figureCitation>
).—Mandibles (
<figureCitation id="2DAB2A6EFFA898356782EEB7FF71FA99" captionStart="Figures 3848" captionStartId="22.[166,243,1805,1827]" captionTargetBox="[174,1460,196,1772]" captionTargetId="figure@22.[166,1466,185,1777]" captionTargetPageId="22" captionText="Figures 3848. Heterolepisma cooloola sp. nov. holotype ♀, unless indicated otherwise by specimen number(38) habitus (K.377754); (39) posterior comb of pronotum with scale; (40) head; (41) scape, pedicel and basal intervals of flagellum, from above; (42) idem, from below; (43) antenna, most distal surviving interval; (44) mandible; (45) idem, detail of molar and incisor regions; (46) maxilla, only larger setae of palp illustrated; (47) idem, lacinia and galea (QM 207012); (48) ultimate article of maxillary palp of ♂ (QM 207012). Scale bars = 0.1 mm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.3838001" httpUri="https://zenodo.org/record/3838001/files/figure.png" pageId="27" pageNumber="28">Figs 44, 45</figureCitation>
) typical for genus with well-developed molar and incisor areas; a group of about nine strong setae distally adjacent to the pectinate molar area and a bush of 50+ setae and macrochaetae externally.—Maxilla (
<figureCitation id="2DAB2A6EFFA89835670EEF02FD1CFAF1" box="[561,685,1375,1399]" captionStart="Figures 3848" captionStartId="22.[166,243,1805,1827]" captionTargetBox="[174,1460,196,1772]" captionTargetId="figure@22.[166,1466,185,1777]" captionTargetPageId="22" captionText="Figures 3848. Heterolepisma cooloola sp. nov. holotype ♀, unless indicated otherwise by specimen number(38) habitus (K.377754); (39) posterior comb of pronotum with scale; (40) head; (41) scape, pedicel and basal intervals of flagellum, from above; (42) idem, from below; (43) antenna, most distal surviving interval; (44) mandible; (45) idem, detail of molar and incisor regions; (46) maxilla, only larger setae of palp illustrated; (47) idem, lacinia and galea (QM 207012); (48) ultimate article of maxillary palp of ♂ (QM 207012). Scale bars = 0.1 mm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.3838001" httpUri="https://zenodo.org/record/3838001/files/figure.png" pageId="27" pageNumber="28">Figs 4648</figureCitation>
) with three large macrochaetae externally proximal to the palp, the lacinia with three strong teeth, one shorter than the rest, seven lamellate processes and a row of eight simple setae, the galea slightly longer than the lacinia with setulae on the outer face. Palp with rosettes of somewhat stronger setae subapically on the two basal articles, all articles with numerous fine setae, apical article of maxillary palp short, being only 0.22 times HW (range 0.200.25) and 4.5 times longer than wide (range 3.66.0) and 1.3 times longer than penultimate article (range 1.21.6), the ultimate article in both sexes with three “branched” papillae, those in the female much less robust and with fewer “arms” than those in the male.—Labium (
<figureCitation id="2DAB2A6EFFA8983565BEEC81FF61F973" box="[129,208,1756,1781]" captionStart="Figures 4959" captionStartId="23.[121,198,1517,1539]" captionTargetBox="[134,1417,159,1496]" captionTargetId="figure@23.[121,1421,151,1514]" captionTargetPageId="23" captionText="Figures 4959. Heterolepisma cooloola sp. nov. holotype ♀ (49) labium, only large setae of palp illustrated; (50) idem, ultimate article of palp; (51) pronotum; (52) idem, right margin; (53) idem, detail of right anterior trichobothrial area; (54) idem, detail of left posterior trichobothrial area; (55) lateral margin of mesonotum; (56) idem, posterior trichobothrial area; (57) idem, anterior trichobothrial area; (58) idem, right posterior comb; (59) lateral margin of metanotum. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838003" httpUri="https://zenodo.org/record/3838003/files/figure.png" pageId="27" pageNumber="28">Fig. 49</figureCitation>
) short and broad with rows of strong setae on the prementum and submentum, glossae and paraglossae quite broad with short curved setulae; labial palp short, apical article eccentric suboval, 1.05 times as long as wide (range L/W 0.91.2) with 2+3 papillae of compact type (
<figureCitation id="2DAB2A6EFFA898356740ED0FFD7AF8EC" box="[639,715,1874,1898]" captionStart="Figures 4959" captionStartId="23.[121,198,1517,1539]" captionTargetBox="[134,1417,159,1496]" captionTargetId="figure@23.[121,1421,151,1514]" captionTargetPageId="23" captionText="Figures 4959. Heterolepisma cooloola sp. nov. holotype ♀ (49) labium, only large setae of palp illustrated; (50) idem, ultimate article of palp; (51) pronotum; (52) idem, right margin; (53) idem, detail of right anterior trichobothrial area; (54) idem, detail of left posterior trichobothrial area; (55) lateral margin of mesonotum; (56) idem, posterior trichobothrial area; (57) idem, anterior trichobothrial area; (58) idem, right posterior comb; (59) lateral margin of metanotum. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838003" httpUri="https://zenodo.org/record/3838003/files/figure.png" pageId="27" pageNumber="28">Fig. 50</figureCitation>
) in a “cluster formation” where the slightly larger distal papillae curve around the two smaller proximal papillae and at least one curved club-like thin-walled basiconic sensillum (N.B. one palp of
<typeStatus id="6A2B8849FFA8983566A1EAC7FC4AFF34" box="[926,1019,154,178]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
does not show usual shape, possibly as a result of damage in the previous instar).
</paragraph>
<paragraph id="B52F36EBFFA898356625EAB9FB9DFABC" blockId="27.[794,1422,228,1983]" pageId="27" pageNumber="28">
<emphasis id="87E4EAF9FFA898356625EAB9FCD8FF7A" box="[794,873,228,252]" italics="true" pageId="27" pageNumber="28">Thorax</emphasis>
: Pronotum (
<figureCitation id="2DAB2A6EFFA8983566C6EAB9FBFDFF7A" box="[1017,1100,228,252]" captionStart="Figures 4959" captionStartId="23.[121,198,1517,1539]" captionTargetBox="[134,1417,159,1496]" captionTargetId="figure@23.[121,1421,151,1514]" captionTargetPageId="23" captionText="Figures 4959. Heterolepisma cooloola sp. nov. holotype ♀ (49) labium, only large setae of palp illustrated; (50) idem, ultimate article of palp; (51) pronotum; (52) idem, right margin; (53) idem, detail of right anterior trichobothrial area; (54) idem, detail of left posterior trichobothrial area; (55) lateral margin of mesonotum; (56) idem, posterior trichobothrial area; (57) idem, anterior trichobothrial area; (58) idem, right posterior comb; (59) lateral margin of metanotum. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838003" httpUri="https://zenodo.org/record/3838003/files/figure.png" pageId="27" pageNumber="28">Fig. 51</figureCitation>
) with narrow setal collar of short, apically bifurcated setae and cilia, quite weak in the medial part of the margin; lateral margins (
<figureCitation id="2DAB2A6EFFA8983561C0EB42FAE1FEB1" box="[1279,1360,287,311]" captionStart="Figures 4959" captionStartId="23.[121,198,1517,1539]" captionTargetBox="[134,1417,159,1496]" captionTargetId="figure@23.[121,1421,151,1514]" captionTargetPageId="23" captionText="Figures 4959. Heterolepisma cooloola sp. nov. holotype ♀ (49) labium, only large setae of palp illustrated; (50) idem, ultimate article of palp; (51) pronotum; (52) idem, right margin; (53) idem, detail of right anterior trichobothrial area; (54) idem, detail of left posterior trichobothrial area; (55) lateral margin of mesonotum; (56) idem, posterior trichobothrial area; (57) idem, anterior trichobothrial area; (58) idem, right posterior comb; (59) lateral margin of metanotum. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838003" httpUri="https://zenodo.org/record/3838003/files/figure.png" pageId="27" pageNumber="28">Fig. 52</figureCitation>
) also with numerous small to medium sized, apically bifurcate setae as well as several larger more erect submarginal macrochaetae; trichobothrial areas open and in contact with the lateral margins, the anterior one (
<figureCitation id="2DAB2A6EFFA898356194EBC9FB4AFE2A" box="[1195,1275,404,428]" captionStart="Figures 4959" captionStartId="23.[121,198,1517,1539]" captionTargetBox="[134,1417,159,1496]" captionTargetId="figure@23.[121,1421,151,1514]" captionTargetPageId="23" captionText="Figures 4959. Heterolepisma cooloola sp. nov. holotype ♀ (49) labium, only large setae of palp illustrated; (50) idem, ultimate article of palp; (51) pronotum; (52) idem, right margin; (53) idem, detail of right anterior trichobothrial area; (54) idem, detail of left posterior trichobothrial area; (55) lateral margin of mesonotum; (56) idem, posterior trichobothrial area; (57) idem, anterior trichobothrial area; (58) idem, right posterior comb; (59) lateral margin of metanotum. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838003" httpUri="https://zenodo.org/record/3838003/files/figure.png" pageId="27" pageNumber="28">Fig. 53</figureCitation>
) located near the mid-point along the margin, almost always with one large macrochaeta located mediad of the trichobothrium (this macrochaeta missing on the left side of the
<typeStatus id="6A2B8849FFA898356018EBB1FA37FD82" box="[1319,1414,492,516]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
) and with a few cilia and setulae; posterior trichobothrial area (
<figureCitation id="2DAB2A6EFFA89835661DE87AFCC3FDB9" box="[802,882,551,575]" captionStart="Figures 4959" captionStartId="23.[121,198,1517,1539]" captionTargetBox="[134,1417,159,1496]" captionTargetId="figure@23.[121,1421,151,1514]" captionTargetPageId="23" captionText="Figures 4959. Heterolepisma cooloola sp. nov. holotype ♀ (49) labium, only large setae of palp illustrated; (50) idem, ultimate article of palp; (51) pronotum; (52) idem, right margin; (53) idem, detail of right anterior trichobothrial area; (54) idem, detail of left posterior trichobothrial area; (55) lateral margin of mesonotum; (56) idem, posterior trichobothrial area; (57) idem, anterior trichobothrial area; (58) idem, right posterior comb; (59) lateral margin of metanotum. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838003" httpUri="https://zenodo.org/record/3838003/files/figure.png" pageId="27" pageNumber="28">Fig. 54</figureCitation>
) near posterior lateral corner with a submarginal macrochaeta between the trichobothrium and the margin as well as a few setulae and cilia; posterior margin slightly concave with 1+1 combs each of one macrochaeta with a smaller seta mediad and posterior to it, the insertion of this setae smaller than that of the macrochaeta on most specimens (the smaller insertion missing from the left side of the
<typeStatus id="6A2B8849FFA89835665FE8A9FC73FC8A" box="[864,962,756,780]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
), the size of the smaller insertion becomes increasingly smaller on posterior segments such that it only appears as a very small submarginal seta on urosternite VIII; these posterior notal combs are associated with two cilia and some setulae.—Mesonotum with lateral chaetotaxy similar to pronotum (
<figureCitation id="2DAB2A6EFFA89835613CE9DAFBE5FC19" box="[1027,1108,903,927]" captionStart="Figures 4959" captionStartId="23.[121,198,1517,1539]" captionTargetBox="[134,1417,159,1496]" captionTargetId="figure@23.[121,1421,151,1514]" captionTargetPageId="23" captionText="Figures 4959. Heterolepisma cooloola sp. nov. holotype ♀ (49) labium, only large setae of palp illustrated; (50) idem, ultimate article of palp; (51) pronotum; (52) idem, right margin; (53) idem, detail of right anterior trichobothrial area; (54) idem, detail of left posterior trichobothrial area; (55) lateral margin of mesonotum; (56) idem, posterior trichobothrial area; (57) idem, anterior trichobothrial area; (58) idem, right posterior comb; (59) lateral margin of metanotum. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838003" httpUri="https://zenodo.org/record/3838003/files/figure.png" pageId="27" pageNumber="28">Fig. 55</figureCitation>
), the posterior trichobothrial area (
<figureCitation id="2DAB2A6EFFA898356665E9F9FC1FFC3A" box="[858,942,932,956]" captionStart="Figures 4959" captionStartId="23.[121,198,1517,1539]" captionTargetBox="[134,1417,159,1496]" captionTargetId="figure@23.[121,1421,151,1514]" captionTargetPageId="23" captionText="Figures 4959. Heterolepisma cooloola sp. nov. holotype ♀ (49) labium, only large setae of palp illustrated; (50) idem, ultimate article of palp; (51) pronotum; (52) idem, right margin; (53) idem, detail of right anterior trichobothrial area; (54) idem, detail of left posterior trichobothrial area; (55) lateral margin of mesonotum; (56) idem, posterior trichobothrial area; (57) idem, anterior trichobothrial area; (58) idem, right posterior comb; (59) lateral margin of metanotum. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838003" httpUri="https://zenodo.org/record/3838003/files/figure.png" pageId="27" pageNumber="28">Fig. 56</figureCitation>
) in the posterolateral corners with a large macrochaeta between the trichobothrium and the margins as well as some marginal and submarginal setae, cilia and setulae; the anterior trichobothrial area (
<figureCitation id="2DAB2A6EFFA898356184E9A1FAB6FB92" box="[1211,1287,1020,1044]" captionStart="Figures 4959" captionStartId="23.[121,198,1517,1539]" captionTargetBox="[134,1417,159,1496]" captionTargetId="figure@23.[121,1421,151,1514]" captionTargetPageId="23" captionText="Figures 4959. Heterolepisma cooloola sp. nov. holotype ♀ (49) labium, only large setae of palp illustrated; (50) idem, ultimate article of palp; (51) pronotum; (52) idem, right margin; (53) idem, detail of right anterior trichobothrial area; (54) idem, detail of left posterior trichobothrial area; (55) lateral margin of mesonotum; (56) idem, posterior trichobothrial area; (57) idem, anterior trichobothrial area; (58) idem, right posterior comb; (59) lateral margin of metanotum. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838003" httpUri="https://zenodo.org/record/3838003/files/figure.png" pageId="27" pageNumber="28">Fig. 57</figureCitation>
) about ¾ the distance posteriorly along the margin, with a submarginal macrochaeta (
<subScript id="291434AEFFA89835668EEE6AFC64FBD7" attach="left" box="[945,981,1079,1105]" fontSize="6" pageId="27" pageNumber="28">m-1</subScript>
) between it and the margin, also with a few setulae and cilia; anterior to this trichobothrial area are two combs (
<subScript id="291434AEFFA89835669FEE2FFC75FB0A" attach="left" box="[928,964,1138,1164]" fontSize="6" pageId="27" pageNumber="28">m-2</subScript>
,
<subScript id="291434AEFFA8983566ECEE2FFC44FB0A" attach="left" box="[979,1013,1138,1164]" fontSize="6" pageId="27" pageNumber="28">m-3</subScript>
) each of two macrochaetae, a further two or three submarginal macrochaetae more anterior along the margin; posterior combs as for pronotum (
<figureCitation id="2DAB2A6EFFA89835602BEEF1FAD5FB42" box="[1300,1380,1196,1220]" captionStart="Figures 4959" captionStartId="23.[121,198,1517,1539]" captionTargetBox="[134,1417,159,1496]" captionTargetId="figure@23.[121,1421,151,1514]" captionTargetPageId="23" captionText="Figures 4959. Heterolepisma cooloola sp. nov. holotype ♀ (49) labium, only large setae of palp illustrated; (50) idem, ultimate article of palp; (51) pronotum; (52) idem, right margin; (53) idem, detail of right anterior trichobothrial area; (54) idem, detail of left posterior trichobothrial area; (55) lateral margin of mesonotum; (56) idem, posterior trichobothrial area; (57) idem, anterior trichobothrial area; (58) idem, right posterior comb; (59) lateral margin of metanotum. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838003" httpUri="https://zenodo.org/record/3838003/files/figure.png" pageId="27" pageNumber="28">Fig. 58</figureCitation>
).— Metanotum (
<figureCitation id="2DAB2A6EFFA898356699EE97FC46FB64" box="[934,1015,1226,1250]" captionStart="Figures 4959" captionStartId="23.[121,198,1517,1539]" captionTargetBox="[134,1417,159,1496]" captionTargetId="figure@23.[121,1421,151,1514]" captionTargetPageId="23" captionText="Figures 4959. Heterolepisma cooloola sp. nov. holotype ♀ (49) labium, only large setae of palp illustrated; (50) idem, ultimate article of palp; (51) pronotum; (52) idem, right margin; (53) idem, detail of right anterior trichobothrial area; (54) idem, detail of left posterior trichobothrial area; (55) lateral margin of mesonotum; (56) idem, posterior trichobothrial area; (57) idem, anterior trichobothrial area; (58) idem, right posterior comb; (59) lateral margin of metanotum. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838003" httpUri="https://zenodo.org/record/3838003/files/figure.png" pageId="27" pageNumber="28">Fig. 59</figureCitation>
) similar to mesonotum (the comb at position
<subScript id="291434AEFFA898356646EEB5FC2AFA87" attach="left" box="[889,923,1256,1281]" fontSize="6" pageId="27" pageNumber="28">m-2</subScript>
on the left side of the
<typeStatus id="6A2B8849FFA8983561A4EEBAFB49FB79" box="[1179,1272,1255,1279]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
composed of only one macrochaeta, suggesting that a degree of variation exists within the species).
</paragraph>
<paragraph id="B52F36EBFFA898356607EF1DFBA2F8AA" blockId="27.[794,1422,228,1983]" pageId="27" pageNumber="28">
Presternum narrow, with transverse row of strong setae and numerous cilia.—All thoracic sterna with hyaline scales. Prothoracic sternum pointed cordiform, only slightly longer than wide at its base (L/W 1.08 range 1.41.15) and reaching almost to the end of the coxa, rounded apically and with a medial furrow (
<figureCitation id="2DAB2A6EFFA89835613BEF8FFBE2FA6C" box="[1028,1107,1490,1514]" captionStart="Figures 6065" captionStartId="24.[166,243,1543,1565]" captionTargetBox="[177,1452,156,1521]" captionTargetId="figure@24.[166,1466,151,1532]" captionTargetPageId="24" captionText="Figures 6065. Heterolepisma cooloola sp. nov. holotype ♀ (60) presternum, prothoracic sternum and PI; (61) mesothoracic sternum and PII; (62) apex of mesothoracic sternum; (63) metathoracic sternum and PIII; (64) apex of metathoracic sternum; (65) pretarsus of PIII. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838007" httpUri="https://zenodo.org/record/3838007/files/figure.png" pageId="27" pageNumber="28">Fig. 60</figureCitation>
), most of lateral margins with numerous small marginal setae and cilia, with 46 larger submarginal macrochaetae forming weak combs parallel to the edges in the distal quarter.—Mesosternum (
<figureCitation id="2DAB2A6EFFA898356008EC77FA34F9C4" box="[1335,1413,1578,1602]" captionStart="Figures 6065" captionStartId="24.[166,243,1543,1565]" captionTargetBox="[177,1452,156,1521]" captionTargetId="figure@24.[166,1466,151,1532]" captionTargetPageId="24" captionText="Figures 6065. Heterolepisma cooloola sp. nov. holotype ♀ (60) presternum, prothoracic sternum and PI; (61) mesothoracic sternum and PII; (62) apex of mesothoracic sternum; (63) metathoracic sternum and PIII; (64) apex of metathoracic sternum; (65) pretarsus of PIII. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838007" httpUri="https://zenodo.org/record/3838007/files/figure.png" pageId="27" pageNumber="28">Fig. 61</figureCitation>
) slightly longer than broad (1.09 range 0.981.18) with a truncate or evenly slightly concave posterior margin, with 1+1 distal combs of two to three submarginal macrochaeta associated with some marginal setae and cilia (
<figureCitation id="2DAB2A6EFFA89835602AECC2FAD5F931" box="[1301,1380,1695,1719]" captionStart="Figures 6065" captionStartId="24.[166,243,1543,1565]" captionTargetBox="[177,1452,156,1521]" captionTargetId="figure@24.[166,1466,151,1532]" captionTargetPageId="24" captionText="Figures 6065. Heterolepisma cooloola sp. nov. holotype ♀ (60) presternum, prothoracic sternum and PI; (61) mesothoracic sternum and PII; (62) apex of mesothoracic sternum; (63) metathoracic sternum and PIII; (64) apex of metathoracic sternum; (65) pretarsus of PIII. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838007" httpUri="https://zenodo.org/record/3838007/files/figure.png" pageId="27" pageNumber="28">Fig. 62</figureCitation>
).— Metasternum (
<figureCitation id="2DAB2A6EFFA89835668CECE1FB9BF952" box="[947,1066,1724,1748]" captionStart="Figures 6065" captionStartId="24.[166,243,1543,1565]" captionTargetBox="[177,1452,156,1521]" captionTargetId="figure@24.[166,1466,151,1532]" captionTargetPageId="24" captionText="Figures 6065. Heterolepisma cooloola sp. nov. holotype ♀ (60) presternum, prothoracic sternum and PI; (61) mesothoracic sternum and PII; (62) apex of mesothoracic sternum; (63) metathoracic sternum and PIII; (64) apex of metathoracic sternum; (65) pretarsus of PIII. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838007" httpUri="https://zenodo.org/record/3838007/files/figure.png" pageId="27" pageNumber="28">Figs 63, 64</figureCitation>
) wider than long (L/W 0.75 range 0.690.84) but otherwise similar to the mesosternum; the gap between the combs 7.3 times the average width of each comb (range 5.411.1).
</paragraph>
<paragraph id="B52F36EBFFA898326607ED6FFD0DFCF4" blockId="27.[794,1422,228,1983]" lastBlockId="28.[166,795,154,882]" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
Legs fairly long (
<figureCitation id="2DAB2A6EFFA8983566C1ED6FFB1FF8CC" box="[1022,1198,1842,1866]" captionStart="Figures 6065" captionStartId="24.[166,243,1543,1565]" captionTargetBox="[177,1452,156,1521]" captionTargetId="figure@24.[166,1466,151,1532]" captionTargetPageId="24" captionText="Figures 6065. Heterolepisma cooloola sp. nov. holotype ♀ (60) presternum, prothoracic sternum and PI; (61) mesothoracic sternum and PII; (62) apex of mesothoracic sternum; (63) metathoracic sternum and PIII; (64) apex of metathoracic sternum; (65) pretarsus of PIII. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838007" httpUri="https://zenodo.org/record/3838007/files/figure.png" pageId="27" pageNumber="28">Figs 60, 61, 63</figureCitation>
), tibia L/W ratio of legs PI 2.6 (range 2.53.0), PII 3.0 (range 2.43.5, PIII 3.6 (range 3.14.1); tarsi L/W ratio PI 6.7 (range 5.47.6), PII 6.7 (range 6.08.0), PIII 8.8 (range 7.410.0). Legs increasingly longer from front to back, mean ratio PI/PIII (tibia 0.58, tarsus 0.68). PI with transverse comb of about six macrochaetae laterally on the precoxa. Coxa of all legs covered with hyaline scales and with strong macrochaetae and numerous cilia in a row about two macrochaetae wide along the external margin, a stout macrochaeta and some long fine setae on the inner margin subapically and group of about four to six stout curved macrochaetae at the apex over the articulation. Trochanter lacking scales, with fine and one stronger seta over the surface. Femur with numerous lanceolate scales on the anterior half and along the margin, rest of surface with fine setae; anterior distal end with three to six strong, quite deeply bifurcate stout macrochaetae as well as some strong setae; posterior margin with several strong macrochaetae as illustrated. Tibia with numerous setae and lanceolate scales over the ventral surface, with two stout macrochaetae on or near the anterior margin and three or four stout macrochaetae along the posterior margin (some paired with thinner macrochaetae on the dorsal side of the margin; apical spur with several setae. Tibia of PIII with a long thin, laterally projecting trichobothria-like seta inserted dorsal to the proximal stout macrochaeta on the anterior margin, which is more than twice as long as the tibia is wide. Tarsus with four articles, all with numerous setae (without lanceolate scales). Pretarsus with long curved lateral claws and a strong curved shorter medial claw (
<figureCitation id="2DAB2A6EFFAF98326760E907FD1EFCF4" box="[607,687,858,882]" captionStart="Figures 6065" captionStartId="24.[166,243,1543,1565]" captionTargetBox="[177,1452,156,1521]" captionTargetId="figure@24.[166,1466,151,1532]" captionTargetPageId="24" captionText="Figures 6065. Heterolepisma cooloola sp. nov. holotype ♀ (60) presternum, prothoracic sternum and PI; (61) mesothoracic sternum and PII; (62) apex of mesothoracic sternum; (63) metathoracic sternum and PIII; (64) apex of metathoracic sternum; (65) pretarsus of PIII. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838007" httpUri="https://zenodo.org/record/3838007/files/figure.png" pageId="28" pageNumber="29">Fig. 65</figureCitation>
).
</paragraph>
<caption id="E1EF6663FFAF98326678EAF0FB36FF64" pageId="28" pageNumber="29">
<paragraph id="B52F36EBFFAF98326678EAF0FB36FF64" blockId="28.[839,1411,173,226]" pageId="28" pageNumber="29">
<emphasis id="87E4EAF9FFAF98326678EAF0FC28FF43" bold="true" box="[839,921,173,197]" pageId="28" pageNumber="29">Table 8</emphasis>
. Number of macrochaetae per bristle comb—
<taxonomicName id="72904D68FFAF98326678EA97FBF6FF64" authority="Smith &amp; Mitchell &amp; Lee &amp; Espinasa, 2019" authorityName="Smith &amp; Mitchell &amp; Lee &amp; Espinasa" authorityYear="2019" box="[839,1095,202,226]" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="cooloola" status="sp. nov.">
<emphasis id="87E4EAF9FFAF98326678EA97FBF6FF64" box="[839,1095,202,226]" italics="true" pageId="28" pageNumber="29">Heterolepisma cooloola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9CD75782FFAF98326171EA96FB36FF64" box="[1102,1159,203,226]" pageId="28" pageNumber="29" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="B52F36EBFFAF98326671EAA7FA31FDC7" pageId="28" pageNumber="29">
<table id="C790C44BFFAF67D16671EAA7FA02FDC7" box="[846,1459,250,577]" colsContinueFrom="20.[844,1459,251,560]" gridcols="5" gridrows="10" pageId="28" pageNumber="29">
<tr id="0BA034A9FFAF67D16671EAA7FA02FE97" box="[846,1459,250,273]" gridrow="0" pageId="28" pageNumber="29" rowspan-1="1" rowspan-3="1">
<th id="48715DD5FFAF67D16671EAA7FC1BFE97" box="[846,938,250,273]" gridcol="0" gridrow="0" pageId="28" pageNumber="29">segment</th>
<th id="48715DD5FFAF67D1610FEAA7FB29FE97" box="[1072,1176,250,273]" gridcol="2" gridrow="0" pageId="28" pageNumber="29">urotergite</th>
<th id="48715DD5FFAF67D16009EAA7FA02FE97" box="[1334,1459,250,273]" gridcol="4" gridrow="0" pageId="28" pageNumber="29">urosternites</th>
</tr>
<tr id="0BA034A9FFAF67D16671EB77FA02FEC7" box="[846,1459,298,321]" gridrow="1" pageId="28" pageNumber="29" rowspan-0="1" rowspan-4="1">
<td id="48715DD5FFAF67D166F9EB77FBB6FEC7" box="[966,1031,298,321]" gridcol="1" gridrow="1" pageId="28" pageNumber="29">lateral</td>
<td id="48715DD5FFAF67D1610FEB77FB29FEC7" box="[1072,1176,298,321]" gridcol="2" gridrow="1" pageId="28" pageNumber="29">sublateral</td>
<td id="48715DD5FFAF67D16194EB77FAA9FEC7" box="[1195,1304,298,321]" gridcol="3" gridrow="1" pageId="28" pageNumber="29">submedial</td>
</tr>
<tr id="0BA034A9FFAF67D16671EB08FA02FEEA" box="[846,1459,341,364]" gridrow="2" pageId="28" pageNumber="29">
<th id="48715DD5FFAF67D16671EB08FC1BFEEA" box="[846,938,341,364]" gridcol="0" gridrow="2" pageId="28" pageNumber="29">I</th>
<td id="48715DD5FFAF67D166F9EB08FBB6FEEA" box="[966,1031,341,364]" gridcol="1" gridrow="2" pageId="28" pageNumber="29">2</td>
<td id="48715DD5FFAF67D1610FEB08FB29FEEA" box="[1072,1176,341,364]" gridcol="2" gridrow="2" pageId="28" pageNumber="29">02</td>
<td id="48715DD5FFAF67D16194EB08FAA9FEEA" box="[1195,1304,341,364]" gridcol="3" gridrow="2" pageId="28" pageNumber="29"></td>
<td id="48715DD5FFAF67D16009EB08FA02FEEA" box="[1334,1459,341,364]" gridcol="4" gridrow="2" pageId="28" pageNumber="29"></td>
</tr>
<tr id="0BA034A9FFAF67D16671EB2DFA02FE01" box="[846,1459,368,391]" gridrow="3" pageId="28" pageNumber="29">
<th id="48715DD5FFAF67D16671EB2DFC1BFE01" box="[846,938,368,391]" gridcol="0" gridrow="3" pageId="28" pageNumber="29">II</th>
<td id="48715DD5FFAF67D166F9EB2DFBB6FE01" box="[966,1031,368,391]" gridcol="1" gridrow="3" pageId="28" pageNumber="29">23</td>
<td id="48715DD5FFAF67D1610FEB2DFB29FE01" box="[1072,1176,368,391]" gridcol="2" gridrow="3" pageId="28" pageNumber="29">13</td>
<td id="48715DD5FFAF67D16194EB2DFAA9FE01" box="[1195,1304,368,391]" gridcol="3" gridrow="3" pageId="28" pageNumber="29">1</td>
<td id="48715DD5FFAF67D16009EB2DFA02FE01" box="[1334,1459,368,391]" gridcol="4" gridrow="3" pageId="28" pageNumber="29">1</td>
</tr>
<tr id="0BA034A9FFAF67D16671EBD7FA02FE27" box="[846,1459,394,417]" gridrow="4" pageId="28" pageNumber="29">
<th id="48715DD5FFAF67D16671EBD7FC1BFE27" box="[846,938,394,417]" gridcol="0" gridrow="4" pageId="28" pageNumber="29">III</th>
<td id="48715DD5FFAF67D166F9EBD7FBB6FE27" box="[966,1031,394,417]" gridcol="1" gridrow="4" pageId="28" pageNumber="29">23</td>
<td id="48715DD5FFAF67D1610FEBD7FB29FE27" box="[1072,1176,394,417]" gridcol="2" gridrow="4" pageId="28" pageNumber="29">23</td>
<td id="48715DD5FFAF67D16194EBD7FAA9FE27" box="[1195,1304,394,417]" gridcol="3" gridrow="4" pageId="28" pageNumber="29">01</td>
<td id="48715DD5FFAF67D16009EBD7FA02FE27" box="[1334,1459,394,417]" gridcol="4" gridrow="4" pageId="28" pageNumber="29">1</td>
</tr>
<tr id="0BA034A9FFAF67D16671EBF8FA02FE3A" box="[846,1459,421,444]" gridrow="5" pageId="28" pageNumber="29">
<th id="48715DD5FFAF67D16671EBF8FC1BFE3A" box="[846,938,421,444]" gridcol="0" gridrow="5" pageId="28" pageNumber="29">IV</th>
<td id="48715DD5FFAF67D166F9EBF8FBB6FE3A" box="[966,1031,421,444]" gridcol="1" gridrow="5" pageId="28" pageNumber="29">34</td>
<td id="48715DD5FFAF67D1610FEBF8FB29FE3A" box="[1072,1176,421,444]" gridcol="2" gridrow="5" pageId="28" pageNumber="29">34</td>
<td id="48715DD5FFAF67D16194EBF8FAA9FE3A" box="[1195,1304,421,444]" gridcol="3" gridrow="5" pageId="28" pageNumber="29">1</td>
<td id="48715DD5FFAF67D16009EBF8FA02FE3A" box="[1334,1459,421,444]" gridcol="4" gridrow="5" pageId="28" pageNumber="29">1</td>
</tr>
<tr id="0BA034A9FFAF67D16671EB9DFA02FE51" box="[846,1459,448,471]" gridrow="6" pageId="28" pageNumber="29">
<th id="48715DD5FFAF67D16671EB9DFC1BFE51" box="[846,938,448,471]" gridcol="0" gridrow="6" pageId="28" pageNumber="29">V</th>
<td id="48715DD5FFAF67D166F9EB9DFBB6FE51" box="[966,1031,448,471]" gridcol="1" gridrow="6" pageId="28" pageNumber="29">34</td>
<td id="48715DD5FFAF67D1610FEB9DFB29FE51" box="[1072,1176,448,471]" gridcol="2" gridrow="6" pageId="28" pageNumber="29">34</td>
<td id="48715DD5FFAF67D16194EB9DFAA9FE51" box="[1195,1304,448,471]" gridcol="3" gridrow="6" pageId="28" pageNumber="29">1</td>
<td id="48715DD5FFAF67D16009EB9DFA02FE51" box="[1334,1459,448,471]" gridcol="4" gridrow="6" pageId="28" pageNumber="29">1</td>
</tr>
<tr id="0BA034A9FFAF67D16671EB87FA02FE77" box="[846,1459,474,497]" gridrow="7" pageId="28" pageNumber="29">
<th id="48715DD5FFAF67D16671EB87FC1BFE77" box="[846,938,474,497]" gridcol="0" gridrow="7" pageId="28" pageNumber="29">VI</th>
<td id="48715DD5FFAF67D166F9EB87FBB6FE77" box="[966,1031,474,497]" gridcol="1" gridrow="7" pageId="28" pageNumber="29">34</td>
<td id="48715DD5FFAF67D1610FEB87FB29FE77" box="[1072,1176,474,497]" gridcol="2" gridrow="7" pageId="28" pageNumber="29">34</td>
<td id="48715DD5FFAF67D16194EB87FAA9FE77" box="[1195,1304,474,497]" gridcol="3" gridrow="7" pageId="28" pageNumber="29">1</td>
<td id="48715DD5FFAF67D16009EB87FA02FE77" box="[1334,1459,474,497]" gridcol="4" gridrow="7" pageId="28" pageNumber="29">1</td>
</tr>
<tr id="0BA034A9FFAF67D16671EBA8FA02FD8A" box="[846,1459,501,524]" gridrow="8" pageId="28" pageNumber="29">
<th id="48715DD5FFAF67D16671EBA8FC1BFD8A" box="[846,938,501,524]" gridcol="0" gridrow="8" pageId="28" pageNumber="29">VII</th>
<td id="48715DD5FFAF67D166F9EBA8FBB6FD8A" box="[966,1031,501,524]" gridcol="1" gridrow="8" pageId="28" pageNumber="29">3</td>
<td id="48715DD5FFAF67D1610FEBA8FB29FD8A" box="[1072,1176,501,524]" gridcol="2" gridrow="8" pageId="28" pageNumber="29">13</td>
<td id="48715DD5FFAF67D16194EBA8FAA9FD8A" box="[1195,1304,501,524]" gridcol="3" gridrow="8" pageId="28" pageNumber="29">1</td>
<td id="48715DD5FFAF67D16009EBA8FA02FD8A" box="[1334,1459,501,524]" gridcol="4" gridrow="8" pageId="28" pageNumber="29">1</td>
</tr>
<tr id="0BA034A9FFAF67D16671E84DFA02FDC7" box="[846,1459,528,577]" gridrow="9" pageId="28" pageNumber="29">
<th id="48715DD5FFAF67D16671E84DFC1BFDC7" box="[846,938,528,577]" gridcol="0" gridrow="9" pageId="28" pageNumber="29">VIII IX</th>
<td id="48715DD5FFAF67D166F9E84DFBB6FDC7" box="[966,1031,528,577]" gridcol="1" gridrow="9" pageId="28" pageNumber="29">13 2a</td>
<td id="48715DD5FFAF67D1610FE84DFB29FDC7" box="[1072,1176,528,577]" gridcol="2" gridrow="9" pageId="28" pageNumber="29">— —</td>
<td id="48715DD5FFAF67D16194E84DFAA9FDC7" box="[1195,1304,528,577]" gridcol="3" gridrow="9" pageId="28" pageNumber="29">01 —</td>
<td id="48715DD5FFAF67D16009E84DFA02FDC7" box="[1334,1459,528,577]" gridcol="4" gridrow="9" pageId="28" pageNumber="29">1 —</td>
</tr>
</table>
</paragraph>
<paragraph id="B52F36EBFFAF98326698E808FB17FDE9" blockId="28.[935,1190,597,623]" box="[935,1190,597,623]" pageId="28" pageNumber="29">
<tableNote id="48763765FFAF98326698E808FB17FDE9" box="[935,1190,597,623]" pageId="28" pageNumber="29" targetBox="[846,1459,250,577]" targetPageId="28">
<superScript id="42E59BA3FFAF98326698E808FC1FFDE5" attach="left" box="[935,942,597,611]" fontSize="6" pageId="28" pageNumber="29">a</superScript>
small infralateral setae
</tableNote>
</paragraph>
<paragraph id="B52F36EBFFAF98326599E9DAFDC9FAD1" blockId="28.[166,795,903,1983]" pageId="28" pageNumber="29">
<emphasis id="87E4EAF9FFAF98326599E9DAFEBFFC19" box="[166,270,903,927]" italics="true" pageId="28" pageNumber="29">Abdomen</emphasis>
: Urotergite I usually with 2+2 combs of 12 macrochaetae (sublateral missing on right side of
<typeStatus id="6A2B8849FFAF98326792E9F9FCBFFC3A" box="[685,782,932,956]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
), urotergites IIVII with 3+3 combs of macrochaetae (
<figureCitation id="2DAB2A6EFFAF983267F8E99FFCA2FC5C" box="[711,787,962,986]" captionStart="Figures 6677" captionStartId="25.[121,198,1620,1642]" captionTargetBox="[133,1412,157,1605]" captionTargetId="figure@25.[121,1421,151,1615]" captionTargetPageId="25" captionText="Figures 6677. Heterolepisma cooloola sp. nov. holotype ♀, unless otherwise indicated by specimen number (66) urotergite IV; (67) urotergite V, right lateral comb; (68) idem, left sublateral comb; (69) idem,left submedial comb; (70) right side of urotergiteVIII (K.261189); (71) infralateral comb of urotergite IX; (72) urotergite X; (73) urotergite X of paratype (QM 207012); (74) urosternite IV; (75) posterior comb of urosternite V; (76) urosternites VII, VIII, IX and ovipositor; (77) base of stylus VII. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838009" httpUri="https://zenodo.org/record/3838009/files/figure.png" pageId="28" pageNumber="29">Fig. 66</figureCitation>
) as in
<tableCitation id="F8120350FFAF983265DEE982FE85FC71" box="[225,308,991,1015]" captionStart="Table 8" captionStartId="28.[839,902,173,197]" captionText="Table 8. Number of macrochaetae per bristle comb— Heterolepisma cooloola n. sp." pageId="28" pageNumber="29">Table 8</tableCitation>
, noting that the macrochaeta was sometimes missing from one of the submedial combs; each comb also associated with 03 marginal setae, 05 setulae plus 14 cilia (e.g.,
<figureCitation id="2DAB2A6EFFAF983265DEEE6AFEEBFBC9" box="[225,346,1079,1103]" captionStart="Figures 6677" captionStartId="25.[121,198,1620,1642]" captionTargetBox="[133,1412,157,1605]" captionTargetId="figure@25.[121,1421,151,1615]" captionTargetPageId="25" captionText="Figures 6677. Heterolepisma cooloola sp. nov. holotype ♀, unless otherwise indicated by specimen number (66) urotergite IV; (67) urotergite V, right lateral comb; (68) idem, left sublateral comb; (69) idem,left submedial comb; (70) right side of urotergiteVIII (K.261189); (71) infralateral comb of urotergite IX; (72) urotergite X; (73) urotergite X of paratype (QM 207012); (74) urosternite IV; (75) posterior comb of urosternite V; (76) urosternites VII, VIII, IX and ovipositor; (77) base of stylus VII. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838009" httpUri="https://zenodo.org/record/3838009/files/figure.png" pageId="28" pageNumber="29">Figs 6769</figureCitation>
). Urotergite VIII (
<figureCitation id="2DAB2A6EFFAF9832672BEE6AFDD1FBC9" box="[532,608,1079,1103]" captionStart="Figures 6677" captionStartId="25.[121,198,1620,1642]" captionTargetBox="[133,1412,157,1605]" captionTargetId="figure@25.[121,1421,151,1615]" captionTargetPageId="25" captionText="Figures 6677. Heterolepisma cooloola sp. nov. holotype ♀, unless otherwise indicated by specimen number (66) urotergite IV; (67) urotergite V, right lateral comb; (68) idem, left sublateral comb; (69) idem,left submedial comb; (70) right side of urotergiteVIII (K.261189); (71) infralateral comb of urotergite IX; (72) urotergite X; (73) urotergite X of paratype (QM 207012); (74) urosternite IV; (75) posterior comb of urosternite V; (76) urosternites VII, VIII, IX and ovipositor; (77) base of stylus VII. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838009" httpUri="https://zenodo.org/record/3838009/files/figure.png" pageId="28" pageNumber="29">Fig. 70</figureCitation>
) with 2+2 combs, lacking the sublateral comb, each comb associated with 02 marginal setae, 25 setulae and 24 cilia; urotergite IX with two long thin infralateral setae on each side as well as a 12 cilia (
<figureCitation id="2DAB2A6EFFAF983265DEEEF1FE9CFB43" box="[225,301,1196,1221]" captionStart="Figures 6677" captionStartId="25.[121,198,1620,1642]" captionTargetBox="[133,1412,157,1605]" captionTargetId="figure@25.[121,1421,151,1615]" captionTargetPageId="25" captionText="Figures 6677. Heterolepisma cooloola sp. nov. holotype ♀, unless otherwise indicated by specimen number (66) urotergite IV; (67) urotergite V, right lateral comb; (68) idem, left sublateral comb; (69) idem,left submedial comb; (70) right side of urotergiteVIII (K.261189); (71) infralateral comb of urotergite IX; (72) urotergite X; (73) urotergite X of paratype (QM 207012); (74) urosternite IV; (75) posterior comb of urosternite V; (76) urosternites VII, VIII, IX and ovipositor; (77) base of stylus VII. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838009" httpUri="https://zenodo.org/record/3838009/files/figure.png" pageId="28" pageNumber="29">Fig. 71</figureCitation>
). Urotergite X fairly long and slender parabolic in both sexes (
<figureCitation id="2DAB2A6EFFAF98326479EE97FE74FB64" box="[326,453,1226,1250]" captionStart="Figures 6677" captionStartId="25.[121,198,1620,1642]" captionTargetBox="[133,1412,157,1605]" captionTargetId="figure@25.[121,1421,151,1615]" captionTargetPageId="25" captionText="Figures 6677. Heterolepisma cooloola sp. nov. holotype ♀, unless otherwise indicated by specimen number (66) urotergite IV; (67) urotergite V, right lateral comb; (68) idem, left sublateral comb; (69) idem,left submedial comb; (70) right side of urotergiteVIII (K.261189); (71) infralateral comb of urotergite IX; (72) urotergite X; (73) urotergite X of paratype (QM 207012); (74) urosternite IV; (75) posterior comb of urosternite V; (76) urosternites VII, VIII, IX and ovipositor; (77) base of stylus VII. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838009" httpUri="https://zenodo.org/record/3838009/files/figure.png" pageId="28" pageNumber="29">Figs 72, 73</figureCitation>
), L/W at base about 0.6 (range 0.540.70) with many strong setae along entire margin, often without obvious strong submarginal macrochaetae in the postero-lateral corners but sometimes with up to two submarginal insertions visible on each side.
</paragraph>
<paragraph id="B52F36EBFFAF983265FAEF01FCABF91C" blockId="28.[166,795,903,1983]" pageId="28" pageNumber="29">
<materialsCitation id="05F83CB6FFAF983265FAEF01FCA7F91C" ID-GBIF-Occurrence="2621712366" collectionCode="VII-IX" collectorName="Urosternite I &amp; Styli IX" pageId="28" pageNumber="29" specimenCount="2" specimenCount-female="2" typeStatus="holotype">
<collectorName id="1865533DFFAF983265FAEF01FEE5FAF2" box="[197,340,1372,1396]" pageId="28" pageNumber="29">Urosternite I</collectorName>
glabrous, urosternites IIVIII (
<figureCitation id="2DAB2A6EFFAF98326781EF01FCA3FAF3" box="[702,786,1372,1397]" captionStart="Figures 6677" captionStartId="25.[121,198,1620,1642]" captionTargetBox="[133,1412,157,1605]" captionTargetId="figure@25.[121,1421,151,1615]" captionTargetPageId="25" captionText="Figures 6677. Heterolepisma cooloola sp. nov. holotype ♀, unless otherwise indicated by specimen number (66) urotergite IV; (67) urotergite V, right lateral comb; (68) idem, left sublateral comb; (69) idem,left submedial comb; (70) right side of urotergiteVIII (K.261189); (71) infralateral comb of urotergite IX; (72) urotergite X; (73) urotergite X of paratype (QM 207012); (74) urosternite IV; (75) posterior comb of urosternite V; (76) urosternites VII, VIII, IX and ovipositor; (77) base of stylus VII. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838009" httpUri="https://zenodo.org/record/3838009/files/figure.png" pageId="28" pageNumber="29">Fig. 74</figureCitation>
) with 1+1 single macrochaetae (
<figureCitation id="2DAB2A6EFFAF983264C5EF27FDF8FA14" box="[506,585,1402,1426]" captionStart="Figures 6677" captionStartId="25.[121,198,1620,1642]" captionTargetBox="[133,1412,157,1605]" captionTargetId="figure@25.[121,1421,151,1615]" captionTargetPageId="25" captionText="Figures 6677. Heterolepisma cooloola sp. nov. holotype ♀, unless otherwise indicated by specimen number (66) urotergite IV; (67) urotergite V, right lateral comb; (68) idem, left sublateral comb; (69) idem,left submedial comb; (70) right side of urotergiteVIII (K.261189); (71) infralateral comb of urotergite IX; (72) urotergite X; (73) urotergite X of paratype (QM 207012); (74) urosternite IV; (75) posterior comb of urosternite V; (76) urosternites VII, VIII, IX and ovipositor; (77) base of stylus VII. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838009" httpUri="https://zenodo.org/record/3838009/files/figure.png" pageId="28" pageNumber="29">Fig. 75</figureCitation>
) (although missing from left side of urosternite II on
<typeStatus id="6A2B8849FFAF9832673CEFCAFDD4FA29" box="[515,613,1431,1455]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
), each associated with 01 marginal seta as well as a few cilia and/or setulae. Coxites of segment VII, VIII and IX in
<specimenCount id="A396FD62FFAF9832675CEF8FFDC6FA6F" box="[611,631,1490,1513]" pageId="28" pageNumber="29" type="female"></specimenCount>
(
<figureCitation id="2DAB2A6EFFAF983267B6EF8FFD68FA6C" box="[649,729,1490,1514]" captionStart="Figures 6677" captionStartId="25.[121,198,1620,1642]" captionTargetBox="[133,1412,157,1605]" captionTargetId="figure@25.[121,1421,151,1615]" captionTargetPageId="25" captionText="Figures 6677. Heterolepisma cooloola sp. nov. holotype ♀, unless otherwise indicated by specimen number (66) urotergite IV; (67) urotergite V, right lateral comb; (68) idem, left sublateral comb; (69) idem,left submedial comb; (70) right side of urotergiteVIII (K.261189); (71) infralateral comb of urotergite IX; (72) urotergite X; (73) urotergite X of paratype (QM 207012); (74) urosternite IV; (75) posterior comb of urosternite V; (76) urosternites VII, VIII, IX and ovipositor; (77) base of stylus VII. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838009" httpUri="https://zenodo.org/record/3838009/files/figure.png" pageId="28" pageNumber="29">Fig. 76</figureCitation>
) with group of several fine setae on the rounded corners on each side of the stylus insertion (
<figureCitation id="2DAB2A6EFFAF983264EDEC51FD93F9A3" box="[466,546,1548,1573]" captionStart="Figures 6677" captionStartId="25.[121,198,1620,1642]" captionTargetBox="[133,1412,157,1605]" captionTargetId="figure@25.[121,1421,151,1615]" captionTargetPageId="25" captionText="Figures 6677. Heterolepisma cooloola sp. nov. holotype ♀, unless otherwise indicated by specimen number (66) urotergite IV; (67) urotergite V, right lateral comb; (68) idem, left sublateral comb; (69) idem,left submedial comb; (70) right side of urotergiteVIII (K.261189); (71) infralateral comb of urotergite IX; (72) urotergite X; (73) urotergite X of paratype (QM 207012); (74) urosternite IV; (75) posterior comb of urosternite V; (76) urosternites VII, VIII, IX and ovipositor; (77) base of stylus VII. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838009" httpUri="https://zenodo.org/record/3838009/files/figure.png" pageId="28" pageNumber="29">Fig. 77</figureCitation>
). Styli in three pairs in the
<specimenCount id="A396FD62FFAF983265F2EC77FF50F9C7" box="[205,225,1578,1601]" pageId="28" pageNumber="29" type="female"></specimenCount>
(
<collectionCode id="D381AE2EFFAF983265D0EC77FEF5F9C4" box="[239,324,1578,1602]" pageId="28" pageNumber="29">VIIIX</collectionCode>
); all styli with several noticeably longer and stronger setae apically.
<collectorName id="1865533DFFAF9832649EEC1AFE46F9D9" box="[417,503,1607,1631]" pageId="28" pageNumber="29">Styli IX</collectorName>
three times as long as styli VII (range 2.33.7) and about two and a half times as long as stylus VIII (range 2.02.9) and much more robust (
<figureCitation id="2DAB2A6EFFAF98326780ECDFFCBCF91C" box="[703,781,1666,1690]" captionStart="Figures 7885" captionStartId="26.[166,243,1315,1337]" captionTargetBox="[177,1457,158,1295]" captionTargetId="figure@26.[166,1466,151,1313]" captionTargetPageId="26" captionText="Figures 7885. Heterolepisma cooloola sp. nov. holotype ♀ unless otherwise indicated by specimen number (78) stylet IX (QM 207012); (79) apex of anterior gonapophysis; (80) apex of posterior gonapophysis; (81) bases of terminal filaments; (82) most distal surviving divisions of cerci; (83) most distal surviving divisions of median filament; (84) coxites IX, styli and penis of male (QM 207012); (85) paramere. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838011" httpUri="https://zenodo.org/record/3838011/files/figure.png" pageId="28" pageNumber="29">Fig. 78</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="B52F36EBFFAF983265FAECC2FA09FD4B" blockId="28.[166,795,903,1983]" lastBlockId="28.[839,1467,664,952]" pageId="28" pageNumber="29">
Coxite IX of
<specimenCount id="A396FD62FFAF9832645DECFDFEC7F931" box="[354,374,1696,1719]" pageId="28" pageNumber="29" type="female"></specimenCount>
(
<figureCitation id="2DAB2A6EFFAF983264B6ECC2FE6EF931" box="[393,479,1695,1719]" captionStart="Figures 6677" captionStartId="25.[121,198,1620,1642]" captionTargetBox="[133,1412,157,1605]" captionTargetId="figure@25.[121,1421,151,1615]" captionTargetPageId="25" captionText="Figures 6677. Heterolepisma cooloola sp. nov. holotype ♀, unless otherwise indicated by specimen number (66) urotergite IV; (67) urotergite V, right lateral comb; (68) idem, left sublateral comb; (69) idem,left submedial comb; (70) right side of urotergiteVIII (K.261189); (71) infralateral comb of urotergite IX; (72) urotergite X; (73) urotergite X of paratype (QM 207012); (74) urosternite IV; (75) posterior comb of urosternite V; (76) urosternites VII, VIII, IX and ovipositor; (77) base of stylus VII. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838009" httpUri="https://zenodo.org/record/3838009/files/figure.png" pageId="28" pageNumber="29">Fig. 76</figureCitation>
), the internal process acute apically, about 4.2 times longer than the external process (range 3.36.0) and 1.8 times as long as broad at its base (range 1.62.1), reaching almost to half the length of the stylus; external and internal margins of internal process and external margin and apex of outer process with many moderately strong setae directed both up and down.— Ovipositor (
<figureCitation id="2DAB2A6EFFAF98326416ED31FEC8F802" box="[297,377,1900,1924]" captionStart="Figures 6677" captionStartId="25.[121,198,1620,1642]" captionTargetBox="[133,1412,157,1605]" captionTargetId="figure@25.[121,1421,151,1615]" captionTargetPageId="25" captionText="Figures 6677. Heterolepisma cooloola sp. nov. holotype ♀, unless otherwise indicated by specimen number (66) urotergite IV; (67) urotergite V, right lateral comb; (68) idem, left sublateral comb; (69) idem,left submedial comb; (70) right side of urotergiteVIII (K.261189); (71) infralateral comb of urotergite IX; (72) urotergite X; (73) urotergite X of paratype (QM 207012); (74) urosternite IV; (75) posterior comb of urosternite V; (76) urosternites VII, VIII, IX and ovipositor; (77) base of stylus VII. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838009" httpUri="https://zenodo.org/record/3838009/files/figure.png" pageId="28" pageNumber="29">Fig. 76</figureCitation>
) very long and thin (up to 2.30 HW), surpassing the apex of stylus IX by at least the length of the stylus (excluding terminal macrochaetae), composed of about 3440 divisions. Distal divisions of gonapophyses VIII and IX (
<figureCitation id="2DAB2A6EFFAF983266A1E8E8FBA8FD4B" box="[926,1049,693,717]" captionStart="Figures 7885" captionStartId="26.[166,243,1315,1337]" captionTargetBox="[177,1457,158,1295]" captionTargetId="figure@26.[166,1466,151,1313]" captionTargetPageId="26" captionText="Figures 7885. Heterolepisma cooloola sp. nov. holotype ♀ unless otherwise indicated by specimen number (78) stylet IX (QM 207012); (79) apex of anterior gonapophysis; (80) apex of posterior gonapophysis; (81) bases of terminal filaments; (82) most distal surviving divisions of cerci; (83) most distal surviving divisions of median filament; (84) coxites IX, styli and penis of male (QM 207012); (85) paramere. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838011" httpUri="https://zenodo.org/record/3838011/files/figure.png" pageId="28" pageNumber="29">Figs 79, 80</figureCitation>
) with only short fine setae and setulae.
</paragraph>
<paragraph id="B52F36EBFFAF9832665AE88EFBD2FC3E" blockId="28.[839,1467,664,952]" pageId="28" pageNumber="29">
Cerci (
<figureCitation id="2DAB2A6EFFAF98326694E88EFB93FD6D" box="[939,1058,723,747]" captionStart="Figures 7885" captionStartId="26.[166,243,1315,1337]" captionTargetBox="[177,1457,158,1295]" captionTargetId="figure@26.[166,1466,151,1313]" captionTargetPageId="26" captionText="Figures 7885. Heterolepisma cooloola sp. nov. holotype ♀ unless otherwise indicated by specimen number (78) stylet IX (QM 207012); (79) apex of anterior gonapophysis; (80) apex of posterior gonapophysis; (81) bases of terminal filaments; (82) most distal surviving divisions of cerci; (83) most distal surviving divisions of median filament; (84) coxites IX, styli and penis of male (QM 207012); (85) paramere. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838011" httpUri="https://zenodo.org/record/3838011/files/figure.png" pageId="28" pageNumber="29">Figs 81, 82</figureCitation>
) with basal divisions shorter than long, gradually becoming longer distally, equally wide as long by about the sixth division after which they become even longer with more annuli each with a rosette of setae and some with trichobothria with the large macrochaetae restricted to the most distal annulus of each division; the most distal surviving divisions with up to eight annuli.—Medial filament of similar arrangement (
<figureCitation id="2DAB2A6EFFAF983266E5E9FDFBE7FC3E" box="[986,1110,928,952]" captionStart="Figures 7885" captionStartId="26.[166,243,1315,1337]" captionTargetBox="[177,1457,158,1295]" captionTargetId="figure@26.[166,1466,151,1313]" captionTargetPageId="26" captionText="Figures 7885. Heterolepisma cooloola sp. nov. holotype ♀ unless otherwise indicated by specimen number (78) stylet IX (QM 207012); (79) apex of anterior gonapophysis; (80) apex of posterior gonapophysis; (81) bases of terminal filaments; (82) most distal surviving divisions of cerci; (83) most distal surviving divisions of median filament; (84) coxites IX, styli and penis of male (QM 207012); (85) paramere. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838011" httpUri="https://zenodo.org/record/3838011/files/figure.png" pageId="28" pageNumber="29">Figs 81, 83</figureCitation>
).
</paragraph>
<paragraph id="B52F36EBFFAF98326678E990FAD8FA8D" blockId="28.[839,1467,973,1291]" pageId="28" pageNumber="29">
<emphasis id="87E4EAF9FFAF98326678E990FCCFFC63" box="[839,894,973,997]" italics="true" pageId="28" pageNumber="29">Male</emphasis>
: As for female except only two pair of styli (segments VII and IX). Coxites IX (
<figureCitation id="2DAB2A6EFFAF98326166E9B6FB16FB85" box="[1113,1191,1003,1027]" captionStart="Figures 7885" captionStartId="26.[166,243,1315,1337]" captionTargetBox="[177,1457,158,1295]" captionTargetId="figure@26.[166,1466,151,1313]" captionTargetPageId="26" captionText="Figures 7885. Heterolepisma cooloola sp. nov. holotype ♀ unless otherwise indicated by specimen number (78) stylet IX (QM 207012); (79) apex of anterior gonapophysis; (80) apex of posterior gonapophysis; (81) bases of terminal filaments; (82) most distal surviving divisions of cerci; (83) most distal surviving divisions of median filament; (84) coxites IX, styli and penis of male (QM 207012); (85) paramere. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838011" httpUri="https://zenodo.org/record/3838011/files/figure.png" pageId="28" pageNumber="29">Fig. 84</figureCitation>
) with acute inner process about 1.8 times longer than wide at its base (range 1.701.93) and about four times longer than the external process which has a small preapical constriction, reaching to about half the length of the stylus. Both processes with several strong setae mostly apically emerging from both the dorsal and ventral surfaces of the processes close to or on the margin. Parameres small, slightly longer than wide, with about eight fine setae (
<figureCitation id="2DAB2A6EFFAF98326670EE88FC2EFB6B" box="[847,927,1237,1261]" captionStart="Figures 7885" captionStartId="26.[166,243,1315,1337]" captionTargetBox="[177,1457,158,1295]" captionTargetId="figure@26.[166,1466,151,1313]" captionTargetPageId="26" captionText="Figures 7885. Heterolepisma cooloola sp. nov. holotype ♀ unless otherwise indicated by specimen number (78) stylet IX (QM 207012); (79) apex of anterior gonapophysis; (80) apex of posterior gonapophysis; (81) bases of terminal filaments; (82) most distal surviving divisions of cerci; (83) most distal surviving divisions of median filament; (84) coxites IX, styli and penis of male (QM 207012); (85) paramere. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838011" httpUri="https://zenodo.org/record/3838011/files/figure.png" pageId="28" pageNumber="29">Fig. 85</figureCitation>
). Penis typical for genus with numerous glandular setae apically, each set on a protuberance (
<figureCitation id="2DAB2A6EFFAF98326033EEAEFAEDFA8D" box="[1292,1372,1267,1291]" captionStart="Figures 7885" captionStartId="26.[166,243,1315,1337]" captionTargetBox="[177,1457,158,1295]" captionTargetId="figure@26.[166,1466,151,1313]" captionTargetPageId="26" captionText="Figures 7885. Heterolepisma cooloola sp. nov. holotype ♀ unless otherwise indicated by specimen number (78) stylet IX (QM 207012); (79) apex of anterior gonapophysis; (80) apex of posterior gonapophysis; (81) bases of terminal filaments; (82) most distal surviving divisions of cerci; (83) most distal surviving divisions of median filament; (84) coxites IX, styli and penis of male (QM 207012); (85) paramere. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838011" httpUri="https://zenodo.org/record/3838011/files/figure.png" pageId="28" pageNumber="29">Fig. 84</figureCitation>
).
</paragraph>
<paragraph id="B52F36EBFFAF98326678EF7DFBD6F91E" blockId="28.[839,1467,1312,1688]" pageId="28" pageNumber="29">
<emphasis id="87E4EAF9FFAF98326678EF7DFC5DFABE" box="[839,1004,1312,1336]" italics="true" pageId="28" pageNumber="29">Subadult stages</emphasis>
: Very small specimens (HW 0.68) only have styli on coxites IX; the coxites of segment VIII are already clearly divided in a juvenile
<specimenCount id="A396FD62FFAF98326147EF06FB3DFAF4" box="[1144,1164,1371,1394]" pageId="28" pageNumber="29" type="female"></specimenCount>
of HW 0.53. The ovipositor is just beginning to appear in a
<specimenCount id="A396FD62FFAF983261AFEF25FB15FA09" box="[1168,1188,1400,1423]" pageId="28" pageNumber="29" type="female"></specimenCount>
of HW 0.83 and styli VIII are present but not styli VII; a
<specimenCount id="A396FD62FFAF983261A8EFCBFB1AFA2B" box="[1175,1195,1430,1453]" pageId="28" pageNumber="29" type="female"></specimenCount>
with HW of 0.88 had an ovipositor that just attained the end of styli IX but still lacked styli VII; by HW 0.93 all styli were present and the ovipositor was much longer than the end of stylus IX. In a
<specimenCount id="A396FD62FFAF98326000EFB3FAE2F983" box="[1343,1363,1518,1541]" pageId="28" pageNumber="29" type="male"></specimenCount>
with HW 0.96 one stylus VIII was developed but the other was only represented by a small triangular appendage. Presumably specimens of both sexes could be considered as sexually mature once HW is greater than
<quantity id="72689B0EFFAF98326199EC3EFAF0F9FC" box="[1190,1345,1635,1659]" metricMagnitude="-4" metricUnit="m" metricValue="9.45" metricValueMax="9.6" metricValueMin="9.3" pageId="28" pageNumber="29" unit="mm" value="0.945" valueMax="0.96" valueMin="0.93">0.930.96 mm</quantity>
or once all styli are clearly developed.
</paragraph>
<paragraph id="B52F36EBFFAF98326678ECF0FBB9F8F0" blockId="28.[839,1468,1709,1910]" pageId="28" pageNumber="29">
<emphasis id="87E4EAF9FFAF98326678ECF0FC11F943" bold="true" box="[839,928,1709,1733]" pageId="28" pageNumber="29">Habitat</emphasis>
.
<taxonomicName id="72904D68FFAF9832668EECF0FB08F943" authorityName="Smith &amp; Mitchell &amp; Lee &amp; Espinasa" authorityYear="2019" box="[945,1209,1709,1733]" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="cooloola">
<emphasis id="87E4EAF9FFAF9832668EECF0FB08F943" box="[945,1209,1709,1733]" italics="true" pageId="28" pageNumber="29">Heterolepisma cooloola</emphasis>
</taxonomicName>
was fairly common in the Rainbow Beach area with specimens collected in dry leaf litter accumulating in places largely protected from rainfall such as within burned out ground level tree hollows. It was also taken from the bark of ti-trees,
<taxonomicName id="72904D68FFAF98326024ED7EFA3DF8BC" box="[1307,1420,1827,1850]" class="Magnoliopsida" family="Casuarinaceae" genus="Casuarina" kingdom="Plantae" order="Fagales" pageId="28" pageNumber="29" phylum="Tracheophyta" rank="genus">
<emphasis id="87E4EAF9FFAF98326024ED7EFA3DF8BC" box="[1307,1420,1827,1850]" italics="true" pageId="28" pageNumber="29">Casuarina</emphasis>
</taxonomicName>
and
<taxonomicName id="72904D68FFAF98326678ED1DFC0FF8DE" box="[839,958,1856,1880]" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" kingdom="Plantae" order="Myrtales" pageId="28" pageNumber="29" phylum="Tracheophyta" rank="genus">
<emphasis id="87E4EAF9FFAF98326678ED1DFC0FF8DE" box="[839,958,1856,1880]" italics="true" pageId="28" pageNumber="29">Eucalyptus</emphasis>
</taxonomicName>
in heathland and on the edge of rainforest using pyrethrum sprays.
</paragraph>
</subSubSection>
<subSubSection id="FD8A6560FFAF98336678EDD6FD7EFD22" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29" type="etymology">
<paragraph id="B52F36EBFFAF98336678EDD6FD7EFD22" blockId="28.[839,1467,1931,1984]" lastBlockId="29.[121,749,154,676]" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
<emphasis id="87E4EAF9FFAF98326678EDD6FC73F825" bold="true" box="[839,962,1931,1955]" pageId="28" pageNumber="29">Etymology</emphasis>
. The species name is derived from the proper noun Cooloola referring the locality in which it was collected.
<emphasis id="87E4EAF9FFAE98336546EAC7FF4BFF34" bold="true" box="[121,250,154,178]" pageId="29" pageNumber="30">Comments</emphasis>
.
<taxonomicName id="72904D68FFAE98336433EAC7FD91FF34" authorityName="Smith &amp; Mitchell &amp; Lee &amp; Espinasa" authorityYear="2019" box="[268,544,154,178]" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="cooloola">
<emphasis id="87E4EAF9FFAE98336433EAC7FD91FF34" box="[268,544,154,178]" italics="true" pageId="29" pageNumber="30">Heterolepisma cooloola</emphasis>
</taxonomicName>
is in many ways similar to
<taxonomicName id="72904D68FFAE983365CEEAE5FE2FFF49" authorityName="Smith" authorityYear="2014" box="[241,414,183,207]" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="sclerophylla">
<emphasis id="87E4EAF9FFAE983365CEEAE5FE2FFF49" box="[241,414,183,207]" italics="true" pageId="29" pageNumber="30">H. sclerophylla</emphasis>
</taxonomicName>
(small triangular prothoracic sternum, glabrous urosternite I, urosternites IIVIII with 1+1 macrochaetae, three pairs of styli in the female and two in the male) but in other aspects it shares characters with
<taxonomicName id="72904D68FFAE98336546EB70FE90FEC2" authority="Smith" authorityName="Smith" authorityYear="2013" box="[121,289,300,324]" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="parva">
<emphasis id="87E4EAF9FFAE98336546EB70FF68FEC2" box="[121,217,301,324]" italics="true" pageId="29" pageNumber="30">H. parva</emphasis>
Smith
</taxonomicName>
from Barrow Island (notably the glabrous anterior margin to the frons, a macrochaeta mediad of the anterior trichobothrium on the pronotum and the presence of lanceolate scales).
<taxonomicName id="72904D68FFAE9833644CEBD9FDEEFE1A" authorityName="Smith" authorityYear="2013" box="[371,607,388,412]" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="parva">
<emphasis id="87E4EAF9FFAE9833644CEBD9FDEEFE1A" box="[371,607,388,412]" italics="true" pageId="29" pageNumber="30">Heterolepisma parva</emphasis>
</taxonomicName>
is however, one of the species with a medial comb on urosternite I and 1+1 combs of several macrochaetae on urosternites IIVII (VIII), a group that Mendes (pers. comm.) has suggested may be a separate group within
<taxonomicName id="72904D68FFAE9833673BEBA7FD1FFD94" authorityName="Escherich" authorityYear="1905" box="[516,686,506,530]" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="87E4EAF9FFAE9833673BEBA7FD1FFD94" box="[516,686,506,530]" italics="true" pageId="29" pageNumber="30">Heterolepisma</emphasis>
</taxonomicName>
. The combined molecular and morphological data support the view that the presence of lanceolate scales and the absence of macrochaetae from the anterior margin of the frons are more significant to phylogeny than the arrangement of styli and the shape of the thoracic sternites in
<taxonomicName id="72904D68FFAE98336713E8D1FD79FD22" authorityName="Escherich" authorityYear="1905" box="[556,712,652,676]" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="87E4EAF9FFAE98336713E8D1FD79FD22" box="[556,712,652,676]" italics="true" pageId="29" pageNumber="30">Heterolepisma</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>