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<document id="C2AB7BA88065F3471D0818AE1DAE5C5C" ID-CLB-Dataset="6634" ID-DOI="10.5852/ejt.2021.749.1355" ID-GBIF-Dataset="83109e54-596a-4be0-ac2c-c4e1702cfb8a" ID-ISSN="2118-9773" ID-Zenodo-Dep="4770293" ID-ZooBank="25FD5744-861D-45E4-B6AD-350716AA29BF" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1621349695117" checkinUser="felipe" docAuthor="Gao, Chuan-bu &amp; Coca-Abia, María Milagro" docDate="2021" docId="03B56743A6780664FE07FD33FE7DE2A5" docLanguage="en" docName="ejt-749_1-94.pdf" docOrigin="European Journal of Taxonomy 749" docSource="http://dx.doi.org/10.5852/ejt.2021.749.1355" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Miridiba ciliatipennis Gao &amp; Coca-Abia 2021, comb. nov." docType="treatment" docVersion="9" lastPageNumber="79" masterDocId="FF8C1F3BA635062AFFFBFFA9FFE2E431" masterDocTitle="Revision of the genus Miridiba Reitter, 1902 (Coleoptera, Scarabaeidae, Melolonthinae): genital morphotypes and new taxonomic data" masterLastPageNumber="94" masterPageNumber="1" pageNumber="78" updateTime="1698921317801" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="4884AFA0AB2D4CFB2FBB4E7FBADC4B23">Revision of the genus Miridiba Reitter, 1902 (Coleoptera, Scarabaeidae, Melolonthinae): genital morphotypes and new taxonomic data</mods:title>
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<mods:namePart id="F662B3FA7DF7C2D72DB79B967BAE4778">Gao, Chuan-bu</mods:namePart>
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<mods:affiliation id="6E6ACC7F147B35F401B68E4290A1D0EA">Guangdong Key Laboratory of Animal Conservation and Resource Utilization, Guangdong Public Laboratory of Wild Animal Conservation and Utilization, Institute of Zoology, Guangdong Academy of Sciences, 105 Xingang Road W., Guangzhou 510260, China. Unidad de Protección Vegetal. Centro de Investigación y Tecnología Agroalimentaria (CITA) e Instituto Agroalimentario de Aragón (IA 2) (CITA-Universidad de Zaragoza), Avenida Montañana, 930. 50059 Zaragoza, Spain. &amp; Buzh.</mods:affiliation>
<mods:nameIdentifier id="B005630A80EBFAA38C881DE2B2DD221C" type="email">Buzh.Gao@msn.com</mods:nameIdentifier>
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<mods:namePart id="1CD494C896C8696D3BEF173643508323">Coca-Abia, María Milagro</mods:namePart>
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<mods:date id="AD25F1CEC3B5414AB4971D719E140CEF">2021</mods:date>
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<subSubSection id="C30685DEA6780667FE07FD33FBDAE684" box="[508,1080,666,693]" pageId="77" pageNumber="78" type="nomenclature">
<paragraph id="8BA3D655A6780667FE07FD33FBDAE684" blockId="77.[508,1080,666,728]" box="[508,1080,666,693]" pageId="77" pageNumber="78">
<heading id="D0EB6139A6780667FE07FD33FBDAE684" box="[508,1080,666,693]" centered="true" fontSize="11" level="2" pageId="77" pageNumber="78" reason="2">
<taxonomicName id="4C1CADD6A6780667FE07FD33FC4DE684" authority="(Moser, 1913)" authorityName="Gao &amp; Coca-Abia" authorityYear="2021" baseAuthorityName="Moser" baseAuthorityYear="1913" box="[508,943,666,693]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="ciliatipennis" status="comb. nov.">
<emphasis id="B9680A47A6780667FE07FD33FCE3E684" bold="true" box="[508,769,666,693]" italics="true" pageId="77" pageNumber="78">Miridiba ciliatipennis</emphasis>
(Moser, 1913)
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<taxonomicNameLabel id="A25BB73CA6780667FC4CFD32FBDAE684" box="[951,1080,667,693]" pageId="77" pageNumber="78" rank="species">comb. nov.</taxonomicNameLabel>
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<subSubSection id="C30685DEA6780667FD23FD17FCB9E6E9" box="[728,859,702,728]" pageId="77" pageNumber="78" type="description">
<paragraph id="8BA3D655A6780667FD23FD17FCB9E6E9" blockId="77.[508,1080,666,728]" box="[728,859,702,728]" pageId="77" pageNumber="78">
<figureCitation id="1327CAD0A6780667FD23FD17FCD0E6E9" box="[728,818,702,728]" captionStart="Fig" captionStartId="15.[191,234,940,966]" captionTargetBox="[279,1310,277,894]" captionTargetId="figure-236@15.[264,1323,262,907]" captionTargetPageId="15" captionText="Fig. 23. Morphotype IX “Ciliatipennis”. Male genitalia of M. ciliatipennis (Moser, 1903), lectotype (MFNB). Parameres in dorsal (A), ventral (B) and lateral (C) views. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770326" httpUri="https://zenodo.org/record/4770326/files/figure.png" pageId="77" pageNumber="78">Figs 23</figureCitation>
,
<figureCitation id="1327CAD0A6780667FCC5FD17FCB9E6E9" box="[830,859,702,728]" captionStart="Figs 98–99. 98" captionStartId="77.[189,241,1953,1979]" captionTargetBox="[286,1303,1066,1896]" captionTargetId="figure-124@77.[264,1323,1047,1914]" captionTargetPageId="77" captionText="Figs 98–99. 98. Miridiba ciliatipennis (Moser, 1903), lectotype, ♂ (MFNB), habitus in dorsal view. 99. M. brunneipennis (Moser, 1916), lectotype, ♂ (MFNB), habitus in dorsal view. Scale bars = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770417" httpUri="https://zenodo.org/record/4770417/files/figure.png" pageId="77" pageNumber="78">98</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA6780667FF46FD56FBE4E74E" pageId="77" pageNumber="78" type="reference_group">
<paragraph id="8BA3D655A6780667FF46FD56FC5FE72B" blockId="77.[189,957,767,794]" box="[189,957,767,794]" pageId="77" pageNumber="78">
<treatmentCitationGroup id="AB0CF17BA6780667FF46FD56FC5FE72B" box="[189,957,767,794]" pageId="77" pageNumber="78">
<treatmentCitation id="0ABDF044A6780667FF46FD56FD21E72B" author="Moser J." box="[189,707,767,794]" page="344" pageId="77" pageNumber="78" year="1913">
<taxonomicName id="4C1CADD6A6780667FF46FD56FD21E72B" authority="Moser, 1913 b: 344" authorityName="Moser" authorityPageNumber="344" authorityYear="1913" box="[189,707,767,794]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="ciliatipennis">
<emphasis id="B9680A47A6780667FF46FD56FE3CE728" box="[189,478,767,793]" italics="true" pageId="77" pageNumber="78">Holotrichia ciliatipennis</emphasis>
<bibRefCitation id="EF8DABA4A6780667FE1DFCA9FD21E72B" author="Moser J." box="[486,707,767,794]" pageId="77" pageNumber="78" pagination="331 - 345" refId="ref45371" refString="Moser J. 1913 b. Beitrag zur Kenntnis der Melolonthiden (Col.) II. Annales de la Societe entomologique de Belgique 57: 331 - 345." type="journal article" year="1913">Moser, 1913b: 344</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(
<typeStatus id="54A768F7A6780667FD29FCA9FCE4E72B" box="[722,774,768,794]" pageId="77" pageNumber="78">type</typeStatus>
loc.:
<collectingCountry id="F30B96C5A6780667FCB3FD56FC53E72B" box="[840,945,767,794]" name="Vietnam" pageId="77" pageNumber="78">Vietnam</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA3D655A6780667FF46FCE8FBE4E74E" blockId="77.[189,1215,833,895]" pageId="77" pageNumber="78">
<treatmentCitationGroup id="AB0CF17BA6780667FF46FCE8FBE4E74E" pageId="77" pageNumber="78">
<taxonomicName id="4C1CADD6A6780667FF46FCE8FE3CE76A" authorityName="Moser" authorityYear="1913" box="[189,478,833,859]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="ciliatipennis">
<emphasis id="B9680A47A6780667FF46FCE8FE3CE76A" box="[189,478,833,859]" italics="true" pageId="77" pageNumber="78">Holotrichia ciliatipennis</emphasis>
</taxonomicName>
–
<treatmentCitation id="0ABDF044A6780667FE07FCE8FD2BE76A" author="Chang Y. - W." box="[508,713,833,859]" page="146" pageId="77" pageNumber="78" year="1964">
<bibRefCitation id="EF8DABA4A6780667FE07FCE8FD2BE76A" author="Chang Y. - W." box="[508,713,833,859]" pageId="77" pageNumber="78" pagination="139 - 152" refId="ref43461" refString="Chang Y. - W. 1964. Revision of Chinese may beetles of the genus Holotrichia Hope I. (Coleoptera, Scarabaeidae). Acta Zootaxonomica Sinica 1 (1): 139 - 152." type="journal article" year="1964">Chang 1964: 146</bibRefCitation>
</treatmentCitation>
, 149–150, figs 54–55 (species list; in key).
<taxonomicName id="4C1CADD6A6780667FF46FCCDFDF6E74F" box="[189,532,868,894]" class="Insecta" family="Melolonthidae" genus="Neodontocnema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="ciliatipennis">
<emphasis id="B9680A47A6780667FF46FCCDFDF6E74F" box="[189,532,868,894]" italics="true" pageId="77" pageNumber="78">Neodontocnema ciliatipennis</emphasis>
</taxonomicName>
–
<treatmentCitation id="0ABDF044A6780667FDCAFCCCFD0AE74E" author="Frey G." box="[561,744,868,895]" page="245" pageId="77" pageNumber="78" year="1970">
<bibRefCitation id="EF8DABA4A6780667FDCAFCCCFD0AE74E" author="Frey G." box="[561,744,868,895]" pageId="77" pageNumber="78" pagination="244 - 265" refId="ref43949" refString="Frey G. 1970. Bestimmungstabelle der Holotrichia - Arten Hinterindiens (Vietnam, Laos, Thailand, Kambodscha, Burma, Malaya) (Col., Scarab, Melolonthinae). Entomologische Arbeiten aus dem Museum G. Frey Tutzing bei Munchen 21: 244 - 265." type="journal article" year="1970">Frey 1970: 245</bibRefCitation>
</treatmentCitation>
, figs 1–2 (combination).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="DF6386DDA6780667FF46F808FA95E3EE" ID-DOI="http://doi.org/10.5281/zenodo.4770417" ID-Zenodo-Dep="4770417" httpUri="https://zenodo.org/record/4770417/files/figure.png" pageId="77" pageNumber="78" startId="77.[189,241,1953,1979]" targetBox="[286,1303,1066,1896]" targetPageId="77">
<paragraph id="8BA3D655A6780667FF46F808FA95E3EE" blockId="77.[189,1399,1953,2016]" pageId="77" pageNumber="78">
<emphasis id="B9680A47A6780667FF46F808FE96E38A" bold="true" box="[189,372,1953,1979]" pageId="77" pageNumber="78">Figs 98–99. 98</emphasis>
.
<taxonomicName id="4C1CADD6A6780667FE7DF808FCA2E38D" authority="(Moser, 1903)" authorityName="Gao &amp; Coca-Abia" authorityYear="2021" baseAuthorityName="Moser" baseAuthorityYear="1903" box="[390,832,1953,1980]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="ciliatipennis">
<emphasis id="B9680A47A6780667FE7DF808FD6BE38A" box="[390,649,1953,1979]" italics="true" pageId="77" pageNumber="78">Miridiba ciliatipennis</emphasis>
(Moser, 1903)
</taxonomicName>
, lectotype, ♂ (MFNB), habitus in dorsal view.
<emphasis id="B9680A47A6780667FF46F86DFF38E3EF" bold="true" box="[189,218,1988,2014]" pageId="77" pageNumber="78">99</emphasis>
.
<taxonomicName id="4C1CADD6A6780667FF13F86CFD86E3D1" authority="(Moser, 1916)" authorityName="Gao &amp; Coca-Abia" authorityYear="2021" baseAuthorityName="Moser" baseAuthorityYear="1916" box="[232,612,1989,2016]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="brunneipennis">
<emphasis id="B9680A47A6780667FF13F86CFE54E3EE" box="[232,438,1989,2015]" italics="true" pageId="77" pageNumber="78">M. brunneipennis</emphasis>
(
<bibRefCitation id="EF8DABA4A6780667FE3FF86CFDB9E3D1" author="Moser J." box="[452,603,1989,2016]" pageId="77" pageNumber="78" pagination="129 - 190" refId="ref45401" refString="Moser J. 1916. Beitrag zur Kenntnis der Melolonthiden. (Col.) V. Deutsche Entomologische Zeitschrift 1916: 129 - 190." type="journal article" year="1916">Moser, 1916</bibRefCitation>
)
</taxonomicName>
, lectotype, ♂ (MFNB), habitus in dorsal view. Scale bars = 5.0 mm.
</paragraph>
</caption>
<subSubSection id="C30685DEA67B0664FF46FEA1FAA9E750" pageId="78" pageNumber="79" type="diagnosis">
<paragraph id="8BA3D655A67B0664FF46FEA1FED4E513" blockId="78.[189,310,264,290]" box="[189,310,264,290]" pageId="78" pageNumber="79">
<heading id="D0EB6139A67B0664FF46FEA1FED4E513" bold="true" box="[189,310,264,290]" fontSize="11" level="3" pageId="78" pageNumber="79" reason="3">
<emphasis id="B9680A47A67B0664FF46FEA1FED4E513" bold="true" box="[189,310,264,290]" pageId="78" pageNumber="79">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A67B0664FF46FE9EFAA9E750" blockId="78.[186,1401,311,866]" pageId="78" pageNumber="79">
External morphology of adult (
<figureCitation id="1327CAD0A67B0664FDCFFE9EFD6CE560" box="[564,654,311,337]" captionStart="Figs 98–99. 98" captionStartId="77.[189,241,1953,1979]" captionTargetBox="[286,1303,1066,1896]" captionTargetId="figure-124@77.[264,1323,1047,1914]" captionTargetPageId="77" captionText="Figs 98–99. 98. Miridiba ciliatipennis (Moser, 1903), lectotype, ♂ (MFNB), habitus in dorsal view. 99. M. brunneipennis (Moser, 1916), lectotype, ♂ (MFNB), habitus in dorsal view. Scale bars = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770417" httpUri="https://zenodo.org/record/4770417/files/figure.png" pageId="78" pageNumber="79">Fig. 98</figureCitation>
). Dorsal surface glabrous and densely punctate. Clypeus with anterior edge widely emarginate at middle and oblique laterally. Frontal carina developed and sharp. Anterior pronotal margin flanged, with a concavity moderately developed at each lateral end; posterior margin glabrous; lateral margins smooth, elevated laterally at anterior third; anterior angle acute; posterior angle obtuse and rounded. Scutellum densely punctate. Elytral surface with punctures more scattered than those on pronotum; epipleuron with long yellow setae along all its length. Foretibia with dorsal carina moderately developed. Meso- and metatibia with transverse carina interrupted at middle; inner margin of metatibial dorsal surface with two strong spines. Pygidium dense punctate and conspicuously pubescent; apex with long pubescence. Ventrites punctate and conspicuously pubescent. Ventrite 2 with short pubescence decumbent. Ventrites 3 and 4 with scattered pubescence at middle. Ventrite 5 with long pubescence on posterior half, irregularly distributed. Ventrite 6 with long pubescence along posterior margin. Male genitalia: parameres (
<figureCitation id="1327CAD0A67B0664FD21FD13FCD3E6E5" box="[730,817,698,724]" captionStart="Fig" captionStartId="15.[191,234,940,966]" captionTargetBox="[279,1310,277,894]" captionTargetId="figure-236@15.[264,1323,262,907]" captionTargetPageId="15" captionText="Fig. 23. Morphotype IX “Ciliatipennis”. Male genitalia of M. ciliatipennis (Moser, 1903), lectotype (MFNB). Parameres in dorsal (A), ventral (B) and lateral (C) views. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770326" httpUri="https://zenodo.org/record/4770326/files/figure.png" pageId="78" pageNumber="79">Fig. 23</figureCitation>
) with dorsal branches of dorsal tubular complex sharp apically and with a tendency to converge (
<figureCitation id="1327CAD0A67B0664FD02FD74FC84E6C9" box="[761,870,733,760]" captionStart="Fig" captionStartId="15.[191,234,940,966]" captionTargetBox="[279,1310,277,894]" captionTargetId="figure-236@15.[264,1323,262,907]" captionTargetPageId="15" captionText="Fig. 23. Morphotype IX “Ciliatipennis”. Male genitalia of M. ciliatipennis (Moser, 1903), lectotype (MFNB). Parameres in dorsal (A), ventral (B) and lateral (C) views. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770326" httpUri="https://zenodo.org/record/4770326/files/figure.png" pageId="78" pageNumber="79">Fig. 23A</figureCitation>
), apices of ventral branches of dorsal tubular complex convergent (
<figureCitation id="1327CAD0A67B0664FE3FFCA8FDD2E72A" box="[452,560,769,795]" captionStart="Fig" captionStartId="15.[191,234,940,966]" captionTargetBox="[279,1310,277,894]" captionTargetId="figure-236@15.[264,1323,262,907]" captionTargetPageId="15" captionText="Fig. 23. Morphotype IX “Ciliatipennis”. Male genitalia of M. ciliatipennis (Moser, 1903), lectotype (MFNB). Parameres in dorsal (A), ventral (B) and lateral (C) views. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770326" httpUri="https://zenodo.org/record/4770326/files/figure.png" pageId="78" pageNumber="79">Fig. 23B</figureCitation>
) and curved ventrally (
<figureCitation id="1327CAD0A67B0664FCA9FCA8FC20E72A" box="[850,962,769,795]" captionStart="Fig" captionStartId="15.[191,234,940,966]" captionTargetBox="[279,1310,277,894]" captionTargetId="figure-236@15.[264,1323,262,907]" captionTargetPageId="15" captionText="Fig. 23. Morphotype IX “Ciliatipennis”. Male genitalia of M. ciliatipennis (Moser, 1903), lectotype (MFNB). Parameres in dorsal (A), ventral (B) and lateral (C) views. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770326" httpUri="https://zenodo.org/record/4770326/files/figure.png" pageId="78" pageNumber="79">Fig. 23C</figureCitation>
); in lateral view, dorsal and ventral branches of dorsal tubular complex forming a semi-circular concavity (
<figureCitation id="1327CAD0A67B0664FBDDFC8DFB74E70F" box="[1062,1174,804,830]" captionStart="Fig" captionStartId="15.[191,234,940,966]" captionTargetBox="[279,1310,277,894]" captionTargetId="figure-236@15.[264,1323,262,907]" captionTargetPageId="15" captionText="Fig. 23. Morphotype IX “Ciliatipennis”. Male genitalia of M. ciliatipennis (Moser, 1903), lectotype (MFNB). Parameres in dorsal (A), ventral (B) and lateral (C) views. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770326" httpUri="https://zenodo.org/record/4770326/files/figure.png" pageId="78" pageNumber="79">Fig. 23C</figureCitation>
). Ventral branches reduced. Phallobase longer than parameres. Endophallus, temones and female genitalia not studied.
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA67B0664FF46FC21FD44E0D0" pageId="78" pageNumber="79" type="materials_examined">
<paragraph id="8BA3D655A67B0664FF46FC21FE48E793" blockId="78.[189,426,904,930]" box="[189,426,904,930]" pageId="78" pageNumber="79">
<heading id="D0EB6139A67B0664FF46FC21FE48E793" bold="true" box="[189,426,904,930]" fontSize="11" level="3" pageId="78" pageNumber="79" reason="3">
<emphasis id="B9680A47A67B0664FF46FC21FE48E793" bold="true" box="[189,426,904,930]" pageId="78" pageNumber="79">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A67B0664FF11FC1EFDDAE7E3" blockId="78.[189,1399,951,1048]" box="[234,568,951,978]" pageId="78" pageNumber="79">
<emphasis id="B9680A47A67B0664FF11FC1EFE84E7E0" bold="true" box="[234,358,951,977]" pageId="78" pageNumber="79">
<typeStatus id="54A768F7A67B0664FF11FC1EFE84E7E0" box="[234,358,951,977]" pageId="78" pageNumber="79" type="lectotype">Lectotype</typeStatus>
</emphasis>
(here designated)
</paragraph>
<paragraph id="8BA3D655A67B0664FF46FC73FE8DE029" blockId="78.[189,1399,951,1048]" pageId="78" pageNumber="79">
<materialsCitation id="3B74DC08A67B0664FF46FC73FE8DE029" ID-GBIF-Occurrence="3114121328" collectionCode="MFNB" country="Vietnam" determinerName="CH. Bu Gao" location="VIETNAM" pageId="78" pageNumber="79" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
<collectingCountry id="F30B96C5A67B0664FF46FC73FEA6E7C5" box="[189,324,986,1012]" name="Vietnam" pageId="78" pageNumber="79">VIETNAM</collectingCountry>
•
<specimenCount id="9D1A1DDCA67B0664FEA7FC72FE97E7C4" box="[348,373,987,1013]" pageId="78" pageNumber="79" type="male">♂</specimenCount>
; “
<collectingCountry id="F30B96C5A67B0664FE77FC73FE0AE7C5" box="[396,488,986,1012]" name="Vietnam" pageId="78" pageNumber="79">Annam</collectingCountry>
” [Centre of Vietnam]; “
<taxonomicName id="4C1CADD6A67B0664FD04FC73FAA7E7C5" authority="(Moser, 1913) CH. Bu Gao" authorityName="CH. Bu Gao" baseAuthorityName="Moser" baseAuthorityYear="1913" box="[767,1349,986,1013]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="ciliatipennis">
<emphasis id="B9680A47A67B0664FD04FC73FC1DE7C5" box="[767,1023,986,1012]" italics="true" pageId="78" pageNumber="79">Miridiba ciliatipennis</emphasis>
(Moser, 1913)
<determinerName id="7E7F3A91A67B0664FB48FC73FAA7E7C5" box="[1203,1349,986,1013]" pageId="78" pageNumber="79">CH. Bu Gao</determinerName>
</taxonomicName>
det. 2017”;
<collectionCode id="ED0D4E90A67B0664FEEFFC57FE89E029" box="[276,363,1022,1048]" collectionName="Italy, Udine, Museo Friulano di Storia Naturale" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="78" pageNumber="79">MFNB</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BA3D655A67B0664FF11FB96FDB5E06B" blockId="78.[189,986,1087,1149]" box="[234,599,1087,1114]" pageId="78" pageNumber="79">
<emphasis id="B9680A47A67B0664FF11FB96FE75E068" bold="true" box="[234,407,1087,1113]" pageId="78" pageNumber="79">
<typeStatus id="54A768F7A67B0664FF11FB96FE75E068" box="[234,407,1087,1113]" pageId="78" pageNumber="79" type="paralectotype">Paralectotype</typeStatus>
</emphasis>
(here indicated)
</paragraph>
<paragraph id="8BA3D655A67B0664FF46FBCBFC38E04C" blockId="78.[189,986,1087,1149]" box="[189,986,1122,1149]" pageId="78" pageNumber="79">
<materialsCitation id="3B74DC08A67B0664FF46FBCBFC34E04C" ID-GBIF-Occurrence="3114121303" box="[189,982,1122,1149]" collectingDate="1887-07" collectionCode="MFNB" collectorName="A. E. Pratt &amp; Coll. Leon Fairmaire" country="Vietnam" determinerName="D. Keith &amp; CH. Bu Gao &amp; M. Coca-Abia" location="VIETNAM" pageId="78" pageNumber="79" specimenCount="1" specimenCount-female="1" typeStatus="paralectotype">
<collectingCountry id="F30B96C5A67B0664FF46FBCBFEA6E04D" box="[189,324,1122,1148]" name="Vietnam" pageId="78" pageNumber="79">VIETNAM</collectingCountry>
•
<specimenCount id="9D1A1DDCA67B0664FEA6FBCBFE68E04C" box="[349,394,1122,1149]" pageId="78" pageNumber="79" type="female">1 ♀</specimenCount>
; same collection data as for the lectotype;
<collectionCode id="ED0D4E90A67B0664FC84FBCAFC34E04C" box="[895,982,1123,1149]" collectionName="Italy, Udine, Museo Friulano di Storia Naturale" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="78" pageNumber="79">MFNB</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BA3D655A67B0664FF11FB0AFE44E08C" blockId="78.[189,678,1187,1249]" box="[234,422,1187,1213]" pageId="78" pageNumber="79">
<emphasis id="B9680A47A67B0664FF11FB0AFE44E08C" bold="true" box="[234,422,1187,1213]" pageId="78" pageNumber="79">Other material</emphasis>
</paragraph>
<paragraph id="8BA3D655A67B0664FF46FB6EFD44E0D0" blockId="78.[189,678,1187,1249]" box="[189,678,1223,1249]" pageId="78" pageNumber="79">
<materialsCitation id="3B74DC08A67B0664FF46FB6EFD40E0D0" ID-GBIF-Occurrence="3114121389" box="[189,674,1223,1249]" collectionCode="MFNB" country="Vietnam" determinerName="Frey" location="VIETNAM" pageId="78" pageNumber="79" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<collectingCountry id="F30B96C5A67B0664FF46FB6EFEA6E0D0" box="[189,324,1223,1249]" name="Vietnam" pageId="78" pageNumber="79">VIETNAM</collectingCountry>
•
<specimenCount id="9D1A1DDCA67B0664FEA6FB6EFE69E0D0" box="[349,395,1223,1249]" pageId="78" pageNumber="79" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D1A1DDCA67B0664FE63FB6EFE27E0D0" box="[408,453,1223,1249]" pageId="78" pageNumber="79" type="female">1 ♀</specimenCount>
;
<determinerName id="7E7F3A91A67B0664FE28FB6EFDEBE0D0" box="[467,521,1223,1249]" pageId="78" pageNumber="79">Frey</determinerName>
det.;
<collectionCode id="ED0D4E90A67B0664FDB0FB6EFD40E0D0" box="[587,674,1223,1249]" collectionName="Italy, Udine, Museo Friulano di Storia Naturale" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="78" pageNumber="79">MFNB</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA67B0664FF46FAA1FB29E215" pageId="78" pageNumber="79" type="discussion">
<paragraph id="8BA3D655A67B0664FF46FAA1FECDE113" blockId="78.[189,303,1288,1314]" box="[189,303,1288,1314]" pageId="78" pageNumber="79">
<heading id="D0EB6139A67B0664FF46FAA1FECDE113" bold="true" box="[189,303,1288,1314]" fontSize="11" level="3" pageId="78" pageNumber="79" reason="3">
<emphasis id="B9680A47A67B0664FF46FAA1FECDE113" bold="true" box="[189,303,1288,1314]" pageId="78" pageNumber="79">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A67B0664FF46FA9EFB29E215" blockId="78.[186,1399,1335,1572]" pageId="78" pageNumber="79">
<bibRefCitation id="EF8DABA4A67B0664FF46FA9EFE8BE163" author="Moser J." box="[189,361,1335,1362]" pageId="78" pageNumber="79" pagination="331 - 345" refId="ref45371" refString="Moser J. 1913 b. Beitrag zur Kenntnis der Melolonthiden (Col.) II. Annales de la Societe entomologique de Belgique 57: 331 - 345." type="journal article" year="1913">Moser (1913b)</bibRefCitation>
described
<taxonomicName id="4C1CADD6A67B0664FE1CFA9EFCE0E160" authorityName="Moser" authorityYear="1913" box="[487,770,1335,1361]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="ciliatipennis">
<emphasis id="B9680A47A67B0664FE1CFA9EFCE0E160" box="[487,770,1335,1361]" italics="true" pageId="78" pageNumber="79">Holotrichia ciliatipennis</emphasis>
</taxonomicName>
without indicating how many species he studied.
<bibRefCitation id="EF8DABA4A67B0664FAB9FA9EFEE8E145" author="Frey G." pageId="78" pageNumber="79" pagination="244 - 265" refId="ref43949" refString="Frey G. 1970. Bestimmungstabelle der Holotrichia - Arten Hinterindiens (Vietnam, Laos, Thailand, Kambodscha, Burma, Malaya) (Col., Scarab, Melolonthinae). Entomologische Arbeiten aus dem Museum G. Frey Tutzing bei Munchen 21: 244 - 265." type="journal article" year="1970">Frey (1970)</bibRefCitation>
transferred this species to
<taxonomicName id="4C1CADD6A67B0664FDCDFAF3FC65E145" authority="Arrow, 1948" authorityName="Arrow" authorityYear="1948" box="[566,903,1370,1396]" class="Insecta" family="Melolonthidae" genus="Neodontocnema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A67B0664FDCDFAF3FD13E145" box="[566,753,1370,1396]" italics="true" pageId="78" pageNumber="79">Neodontocnema</emphasis>
<bibRefCitation id="EF8DABA4A67B0664FD0DFAF3FC65E145" author="Arrow G. J." box="[758,903,1370,1396]" pageId="78" pageNumber="79" pagination="49 - 54" refId="ref43061" refString="Arrow G. J. 1948. Further notes on the beetle genus Lachnosterna (Col.: Melolonthinae), with descriptions of three new genera. The Proceedings of the Royal Entomological Society of London. Series B, Taxonomy 17: 49 - 54. https: // doi. org / 10.1111 / j. 1365 - 3113.1948. tb 00885. x" type="journal article" year="1948">Arrow, 1948</bibRefCitation>
</taxonomicName>
and illustrated the parameres only in dorsal view.The first author of the present paper has examined
<specimenCount id="9D1A1DDCA67B0664FCD6FAD4FC3BE1A6" box="[813,985,1405,1431]" pageId="78" pageNumber="79" type="generic">four specimens</specimenCount>
of
<taxonomicName id="4C1CADD6A67B0664FC01FAD7FAF0E1A6" authority="(MFNB)" baseAuthorityName="MFNB" box="[1018,1298,1405,1431]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="ciliatipennis">
<emphasis id="B9680A47A67B0664FC01FAD7FB4BE1A6" box="[1018,1193,1405,1431]" italics="true" pageId="78" pageNumber="79">H. ciliatipennis</emphasis>
(MFNB)
</taxonomicName>
(
<specimenCount id="9D1A1DDCA67B0664FADBFAD4FA95E1A6" box="[1312,1399,1405,1431]" pageId="78" pageNumber="79" type="male">3 males</specimenCount>
and
<specimenCount id="9D1A1DDCA67B0664FF17FA09FEB6E18B" box="[236,340,1440,1466]" pageId="78" pageNumber="79" type="female">1 female</specimenCount>
, including
<specimenCount id="9D1A1DDCA67B0664FE2EFA08FDABE18A" box="[469,585,1441,1467]" pageId="78" pageNumber="79" type="generic">two types</specimenCount>
), which show external and genital features that characterize
<taxonomicName id="4C1CADD6A67B0664FAF1FA09FA8DE18B" authorityName="Pseudosinensis" authorityYear="2021" baseAuthorityName="Matsumoto" baseAuthorityYear="2011" box="[1290,1391,1440,1466]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A67B0664FAF1FA09FA8DE18B" box="[1290,1391,1440,1466]" italics="true" pageId="78" pageNumber="79">Miridiba</emphasis>
</taxonomicName>
. However, male genitalia show features not shared with species belonging to other genital morphotypes of
<taxonomicName id="4C1CADD6A67B0664FF46FA4EFEC1E230" authorityName="Pseudosinensis" authorityYear="2021" baseAuthorityName="Matsumoto" baseAuthorityYear="2011" box="[189,291,1511,1537]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A67B0664FF46FA4EFEC1E230" box="[189,291,1511,1537]" italics="true" pageId="78" pageNumber="79">Miridiba</emphasis>
</taxonomicName>
. Hence, we transfer
<taxonomicName id="4C1CADD6A67B0664FDE8FA4EFD27E230" box="[531,709,1511,1537]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="ciliatipennis">
<emphasis id="B9680A47A67B0664FDE8FA4EFD27E230" box="[531,709,1511,1537]" italics="true" pageId="78" pageNumber="79">H. ciliatipennis</emphasis>
</taxonomicName>
to
<taxonomicName id="4C1CADD6A67B0664FD11FA4EFCB2E230" authorityName="Pseudosinensis" authorityYear="2021" baseAuthorityName="Matsumoto" baseAuthorityYear="2011" box="[746,848,1511,1537]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A67B0664FD11FA4EFCB2E230" box="[746,848,1511,1537]" italics="true" pageId="78" pageNumber="79">Miridiba</emphasis>
</taxonomicName>
and describe the male genitalia morphotype IX “
<emphasis id="B9680A47A67B0664FF31F9A3FEBCE215" box="[202,350,1546,1572]" italics="true" pageId="78" pageNumber="79">Ciliatipennis</emphasis>
” based on this species. Moreover, we designate
<typeStatus id="54A768F7A67B0664FC73F9A3FC13E215" box="[904,1009,1546,1572]" pageId="78" pageNumber="79" type="lectotype">lectotype</typeStatus>
and
<typeStatus id="54A768F7A67B0664FBD2F9A3FB25E215" box="[1065,1223,1546,1572]" pageId="78" pageNumber="79" type="paralectotype">paralectotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA67B0664FF46F9E2FE7DE2A5" box="[189,415,1611,1684]" pageId="78" pageNumber="79" type="distribution">
<paragraph id="8BA3D655A67B0664FF46F9E2FEB4E254" blockId="78.[189,342,1611,1637]" box="[189,342,1611,1637]" pageId="78" pageNumber="79">
<heading id="D0EB6139A67B0664FF46F9E2FEB4E254" bold="true" box="[189,342,1611,1637]" fontSize="11" level="3" pageId="78" pageNumber="79" reason="3">
<emphasis id="B9680A47A67B0664FF46F9E2FEB4E254" bold="true" box="[189,342,1611,1637]" pageId="78" pageNumber="79">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A67B0664FF46F9D3FE7DE2A5" blockId="78.[189,415,1658,1684]" box="[189,415,1658,1684]" pageId="78" pageNumber="79">
<collectingCountry id="F30B96C5A67B0664FF46F9D3FEC1E2A5" box="[189,291,1658,1684]" name="Vietnam" pageId="78" pageNumber="79">Vietnam</collectingCountry>
,
<collectingCountry id="F30B96C5A67B0664FED5F9D3FE78E2A5" box="[302,410,1658,1684]" name="Thailand" pageId="78" pageNumber="79">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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