185 lines
20 KiB
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185 lines
20 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.6280372" ID-GBIF-Dataset="647046fe-0ee5-401d-b4f8-aea0355fde7f" ID-Zenodo-Dep="6280372" checkinTime="1471630429166" checkinUser="plazi" docAuthor="Piotr Naskrecki" docDate="2000" docId="634387D1A361FF8315F9FB64FC053E8F" docLanguage="en" docName="Naskrecki_KCR_2000.pdf.imf" docOrigin="Philadelphia, PA: The Orthopterists Society at the Academy of Natural Sciences of Philadelphia," docStyle="DocumentStyle{}" docTitle="Metacaputus Naskrecki, sp. n." docType="treatment" docVersion="8" lastPageId="77" lastPageNumber="74" masterDocId="9F7AFFA9A32AFFCE167AFFC3FFF13C6E" masterDocTitle="Katydids of Costa Rica / Vol. 1, Systematics and bioacoustics of the cone-head katydids (Orthoptera: Tettigoniidae: Conocephalinae sensu lato)." pageId="75" pageNumber="72" updateTime="1645797358202" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Katydids of Costa Rica / Vol. 1, Systematics and bioacoustics of the cone-head katydids (Orthoptera: Tettigoniidae: Conocephalinae sensu lato).</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Piotr Naskrecki</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:originInfo>
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<mods:dateIssued>2000</mods:dateIssued>
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<mods:publisher>The Orthopterists Society at the Academy of Natural Sciences of Philadelphia,</mods:publisher>
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<mods:place>
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<mods:placeTerm>Philadelphia, PA</mods:placeTerm>
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</mods:place>
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</mods:originInfo>
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<mods:classification>book</mods:classification>
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<mods:identifier type="DOI">10.5281/zenodo.270035</mods:identifier>
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<mods:identifier type="GBIF-Dataset">647046fe-0ee5-401d-b4f8-aea0355fde7f</mods:identifier>
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<mods:identifier type="Zenodo-Dep">270035</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6280372" ID-Zenodo-Dep="6280372" LSID="urn:lsid:plazi:treatment:634387D1A361FF8315F9FB64FC053E8F" httpUri="http://treatment.plazi.org/id/634387D1A361FF8315F9FB64FC053E8F" lastPageId="77" lastPageNumber="74" pageId="75" pageNumber="72">
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<paragraph blockId="75.[899,1278,1191,1213]" box="[899,1278,1191,1213]" pageId="75" pageNumber="72">
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<heading box="[899,1278,1191,1213]" centered="true" fontSize="9" level="14" pageId="75" pageNumber="72" reason="4">
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<taxonomicName authority="Naskrecki" authorityName="Naskrecki" box="[899,1198,1191,1213]" class="Insecta" family="Tettigoniidae" genus="Metacaputus" kingdom="Animalia" order="Orthoptera" pageId="75" pageNumber="72" phylum="Arthropoda" rank="genus">
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<emphasis bold="true" box="[899,1082,1191,1212]" italics="true" pageId="75" pageNumber="72">METACAPUTUS</emphasis>
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Naskrecki
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</taxonomicName>
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, gen. n.
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</heading>
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</paragraph>
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<paragraph blockId="75.[794,1381,1248,1270]" box="[794,1381,1248,1270]" pageId="75" pageNumber="72">
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<typeStatus box="[794,846,1249,1270]" pageId="75" pageNumber="72">Type</typeStatus>
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species:
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<emphasis box="[939,1174,1248,1270]" italics="true" pageId="75" pageNumber="72">
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<taxonomicName box="[939,1140,1248,1270]" class="Insecta" family="Tettigoniidae" genus="Metacaputus" kingdom="Animalia" order="Orthoptera" pageId="75" pageNumber="72" phylum="Arthropoda" rank="species" species="brenesi" status="sp. n.">Metacaputus brenesi</taxonomicName>
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<taxonomicNameLabel box="[1146,1201,1249,1270]" pageId="75" pageNumber="72" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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, here designated
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</paragraph>
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<subSubSection pageId="75" pageNumber="72" type="diagnosis">
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<paragraph blockId="75.[794,1383,1306,1530]" box="[794,905,1306,1328]" pageId="75" pageNumber="72">
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<heading bold="true" box="[794,905,1306,1328]" fontSize="9" level="13" pageId="75" pageNumber="72" reason="6">
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<emphasis bold="true" box="[794,905,1306,1328]" pageId="75" pageNumber="72">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="75.[794,1383,1306,1530]" pageId="75" pageNumber="72">
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Body slender; both sexes macropterous (
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<figureCitation box="[1273,1349,1335,1357]" captionStart="FIG. 17" captionStartId="76.[151,195,1684,1706]" captionTargetBox="[151,1352,204,1593]" captionTargetId="figure@76.[151,1352,204,1593]" captionTargetPageId="76" captionText="FIG. 17. Metacaputus brenesi. A. female - habitus, B. female face, C. female head and pronotum, dorsal view, D. female subgenital plate, ventral view, E. male cerci, dorsal view." httpUri="https://zenodo.org/record/270053/files/figure.png" pageId="75" pageNumber="72">Fig. 17</figureCitation>
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A); tegumen moderately rugose. Fastigium of vertex 3 times as broad as scapus, long, triangular; genal carinae weakly indicated; frons weakly concave; eyes small. All legs very long and slender. Male cercus armed apically with 2 strongly incurved spines; ovipositor short, distinctly bent upwards at base; rounded apically.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="77" lastPageNumber="74" pageId="75" pageNumber="72" type="description">
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<paragraph blockId="75.[794,1382,1566,1736]" box="[794,1245,1566,1588]" pageId="75" pageNumber="72">
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<emphasis bold="true" box="[794,923,1566,1588]" pageId="75" pageNumber="72">Description</emphasis>
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(male except where specified)
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</paragraph>
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<paragraph blockId="75.[794,1382,1566,1736]" lastBlockId="77.[174,763,207,1726]" lastPageId="77" lastPageNumber="74" pageId="75" pageNumber="72">
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<emphasis box="[817,897,1595,1616]" italics="true" pageId="75" pageNumber="72">Head.—</emphasis>
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Fastigium of vertex 3 times as wide as scapus, triangular, strongly projecting in front of eyes, 3 times as long as eye diameter; fastigium of vertex separated from fastigium of frons by small gap (
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<figureCitation box="[1153,1227,1684,1706]" captionStart="FIG. 17" captionStartId="76.[151,195,1684,1706]" captionTargetBox="[151,1352,204,1593]" captionTargetId="figure@76.[151,1352,204,1593]" captionTargetPageId="76" captionText="FIG. 17. Metacaputus brenesi. A. female - habitus, B. female face, C. female head and pronotum, dorsal view, D. female subgenital plate, ventral view, E. male cerci, dorsal view." httpUri="https://zenodo.org/record/270053/files/figure.png" pageId="75" pageNumber="72">Fig. 17</figureCitation>
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A). Eyes small relative to size of head, weakly protruding. Frons weakly concave; tegumen of head weakly rugose, with traces of genal carinae; face narrowly triangular; mandibles and clypeus weakly asymmetrical (right mandible smaller) (
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<figureCitation box="[181,253,291,314]" captionStart="FIG. 17" captionStartId="76.[151,195,1684,1706]" captionTargetBox="[151,1352,204,1593]" captionTargetId="figure@76.[151,1352,204,1593]" captionTargetPageId="76" captionText="FIG. 17. Metacaputus brenesi. A. female - habitus, B. female face, C. female head and pronotum, dorsal view, D. female subgenital plate, ventral view, E. male cerci, dorsal view." httpUri="https://zenodo.org/record/270053/files/figure.png" pageId="77" pageNumber="74">Fig. 17</figureCitation>
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B).
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</paragraph>
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<caption httpUri="https://zenodo.org/record/270053/files/figure.png" pageId="76" pageNumber="73" targetBox="[151,1352,204,1593]" targetPageId="76">
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<paragraph blockId="76.[151,1359,1684,1735]" pageId="76" pageNumber="73">
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<emphasis bold="true" box="[151,238,1684,1706]" pageId="76" pageNumber="73">FIG. 17.</emphasis>
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<taxonomicName box="[246,449,1684,1705]" class="Insecta" family="Tettigoniidae" genus="Metacaputus" kingdom="Animalia" order="Orthoptera" pageId="76" pageNumber="73" phylum="Arthropoda" rank="species" species="brenesi">
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<emphasis box="[246,449,1684,1705]" italics="true" pageId="76" pageNumber="73">Metacaputus brenesi</emphasis>
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</taxonomicName>
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.
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<emphasis bold="true" box="[464,488,1684,1705]" pageId="76" pageNumber="73">A.</emphasis>
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female - habitus,
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<emphasis bold="true" box="[685,707,1684,1705]" pageId="76" pageNumber="73">B.</emphasis>
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female face,
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<emphasis bold="true" box="[851,874,1684,1706]" pageId="76" pageNumber="73">C.</emphasis>
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female head and pronotum, dorsal view,
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<emphasis bold="true" box="[1334,1359,1684,1705]" pageId="76" pageNumber="73">D.</emphasis>
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female subgenital plate, ventral view,
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<emphasis bold="true" box="[552,572,1713,1734]" pageId="76" pageNumber="73">E.</emphasis>
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male cerci, dorsal view.
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</paragraph>
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</caption>
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<paragraph blockId="77.[174,763,207,1726]" pageId="77" pageNumber="74">
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<emphasis box="[196,396,320,341]" italics="true" pageId="77" pageNumber="74">Thorax and wings.—</emphasis>
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Dorsal surface of pronotum weakly rugose, flat; both anterior and posterior dorsal margins straight (
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<figureCitation box="[275,352,376,398]" captionStart="FIG. 17" captionStartId="76.[151,195,1684,1706]" captionTargetBox="[151,1352,204,1593]" captionTargetId="figure@76.[151,1352,204,1593]" captionTargetPageId="76" captionText="FIG. 17. Metacaputus brenesi. A. female - habitus, B. female face, C. female head and pronotum, dorsal view, D. female subgenital plate, ventral view, E. male cerci, dorsal view." httpUri="https://zenodo.org/record/270053/files/figure.png" pageId="77" pageNumber="74">Fig. 17</figureCitation>
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C); lateral lobes with posterior angle broadly rounded; humeral sinus well developed. Thoracic auditory spiracle large, elliptical, completely hidden under lateral lobe of pronotum. Prosternum armed with two thin, widely separated spines (modified basisternum); meso- and metasternum with lateral lobes of basisterna triangular, their inner margins touching.
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</paragraph>
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<paragraph blockId="77.[174,763,207,1726]" pageId="77" pageNumber="74">
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Wings in both sexes fully developed, well surpassing apices of hind
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<taxonomicName box="[337,415,602,624]" class="Insecta" family="Tettigoniidae" genus="Sphyrometopa" kingdom="Animalia" order="Orthoptera" pageId="77" pageNumber="74" phylum="Arthropoda" rank="species" species="femora">femora</taxonomicName>
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. Stridulatory apparatus of male well developed; stridulatory area of left wing thickened, with dense network of secondary veinlets; stridulatory file (vein AA1) very short, consisting of only 17 (n=1) wide, thick teeth (
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<figureCitation box="[366,439,715,738]" captionStart="FIG. 43" captionStartId="141.[174,218,1655,1677]" captionTargetBox="[242,1315,204,1640]" captionTargetId="figure@141.[242,1316,204,1640]" captionTargetPageId="141" captionText="FIG. 43. Stridulatory files of Costa Rican species of Conocephalinae s. l. A. Copiphora cochleata, B. Copiphora rhinoceros, C. ditto, enlarged fragment, D. Acantheremus colwelli, enlarged fragment, E. ditto, entire file, F. Metacaputus brenesi." httpUri="https://zenodo.org/record/270079/files/figure.png" pageId="77" pageNumber="74">Fig. 43</figureCitation>
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F); mirror of right wing nearly circular. Posterior margin of tegmen weakly concave; apex of tegmen obliquely truncated.
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</paragraph>
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<paragraph blockId="77.[174,763,207,1726]" pageId="77" pageNumber="74">
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<emphasis box="[196,270,801,822]" italics="true" pageId="77" pageNumber="74">Legs.—</emphasis>
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All legs exceptionally thin and long, fore coxa with elongate, forward projecting spine dorsally. Fore and mid
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<taxonomicName box="[226,301,857,879]" class="Insecta" family="Tettigoniidae" genus="Sphyrometopa" kingdom="Animalia" order="Orthoptera" pageId="77" pageNumber="74" phylum="Arthropoda" rank="species" species="femora">femora</taxonomicName>
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completely unarmed; hind
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<taxonomicName box="[609,684,857,879]" class="Insecta" family="Tettigoniidae" genus="Sphyrometopa" kingdom="Animalia" order="Orthoptera" pageId="77" pageNumber="74" phylum="Arthropoda" rank="species" species="femora">femora</taxonomicName>
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armed with minute spines on both ventral margins in apical half; all genicular lobes of
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<taxonomicName box="[409,484,913,935]" class="Insecta" family="Tettigoniidae" genus="Sphyrometopa" kingdom="Animalia" order="Orthoptera" pageId="77" pageNumber="74" phylum="Arthropoda" rank="species" species="femora">femora</taxonomicName>
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armed with short spines. Fore and mid tibiae unarmed dorsally, both ventral margins with immovable spines as long as 1/4 diameter of tibia; hind tibia armed on all four dorsal and ventral margins; apex of tibia with two pairs of ventral and one pair of dorsal movable spurs. Tympanum on fore tibia bilaterally closed, tympanal slits facing forward, tympanal area only weakly swollen, with pair of small, elongated pits below tympanal slits.
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</paragraph>
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<paragraph blockId="77.[174,763,207,1726]" pageId="77" pageNumber="74">
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<emphasis box="[196,317,1168,1189]" italics="true" pageId="77" pageNumber="74">Abdomen.—</emphasis>
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Dorsal surface of abdominal terga smooth, unmodified. Male 10th tergite with shallow apical emargination; supraanal plate small, triangular; female 10th tergite with narrow, triangular apical incision; female supraanal plate narrowly triangular. Male cercus with 2 apical spines, both strongly bent inwards under acute angle; upper spine 1/4 as long as lower spine; lower spine indented apically; female cercus, simple, slender, straight (
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<figureCitation box="[181,251,1394,1416]" captionStart="FIG. 17" captionStartId="76.[151,195,1684,1706]" captionTargetBox="[151,1352,204,1593]" captionTargetId="figure@76.[151,1352,204,1593]" captionTargetPageId="76" captionText="FIG. 17. Metacaputus brenesi. A. female - habitus, B. female face, C. female head and pronotum, dorsal view, D. female subgenital plate, ventral view, E. male cerci, dorsal view." httpUri="https://zenodo.org/record/270053/files/figure.png" pageId="77" pageNumber="74">Fig. 17</figureCitation>
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D). Subgenital plate of male with pair of styli and shallow, triangular apical emargination; female subgenital plate longer than wide, with shallow apical emargination (
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<figureCitation box="[330,401,1479,1501]" captionStart="FIG. 17" captionStartId="76.[151,195,1684,1706]" captionTargetBox="[151,1352,204,1593]" captionTargetId="figure@76.[151,1352,204,1593]" captionTargetPageId="76" captionText="FIG. 17. Metacaputus brenesi. A. female - habitus, B. female face, C. female head and pronotum, dorsal view, D. female subgenital plate, ventral view, E. male cerci, dorsal view." httpUri="https://zenodo.org/record/270053/files/figure.png" pageId="77" pageNumber="74">Fig. 17</figureCitation>
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E).
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</paragraph>
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<paragraph blockId="77.[174,763,207,1726]" pageId="77" pageNumber="74">Male concealed genitalia with part of phallic membrane partially sclerotized and covered by large chitinous callosities. Ovipositor distinctly bent upwards at base, about as long as 1/2 of hind femur, gradually narrowing towards apex; apex rounded.</paragraph>
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<paragraph blockId="77.[174,763,207,1726]" lastBlockId="77.[794,1384,207,737]" pageId="77" pageNumber="74">
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<emphasis box="[196,328,1648,1669]" italics="true" pageId="77" pageNumber="74">Coloration.—</emphasis>
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Coloration probably green, the only available specimens have yellowish-brown coloration, typical of specimens that had been stored in alcohol before being dried.Apex of fastigium of vertex with dark edges.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="77" pageNumber="74" type="discussion">
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<paragraph blockId="77.[794,1384,207,737]" pageId="77" pageNumber="74">
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<emphasis box="[817,1065,235,256]" italics="true" pageId="77" pageNumber="74">
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Remarks.—
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<taxonomicName box="[938,1065,235,256]" class="Insecta" family="Tettigoniidae" genus="Metacaputus" kingdom="Animalia" order="Orthoptera" pageId="77" pageNumber="74" phylum="Arthropoda" rank="genus">Metacaputus</taxonomicName>
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</emphasis>
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seems to be closely related to the genera
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<taxonomicName authority="Karny" authorityName="Karny" box="[907,1141,263,285]" class="Insecta" family="Tettigoniidae" genus="Neoconocephalus" kingdom="Animalia" order="Orthoptera" pageId="77" pageNumber="74" phylum="Arthropoda" rank="genus">
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<emphasis box="[907,1071,263,284]" italics="true" pageId="77" pageNumber="74">Neoconocephalus</emphasis>
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Karny
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</taxonomicName>
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and
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<taxonomicName authority="Karny" authorityName="Karny" box="[1190,1379,264,285]" class="Insecta" family="Tettigoniidae" genus="Paroxyprora" kingdom="Animalia" order="Orthoptera" pageId="77" pageNumber="74" phylum="Arthropoda" rank="genus">
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<emphasis box="[1190,1309,264,285]" italics="true" pageId="77" pageNumber="74">Paroxyprora</emphasis>
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Karny
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</taxonomicName>
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, sharing similar general body form, shape of male cerci, and weakly asymmetrical mouthparts. From
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<taxonomicName box="[794,960,348,369]" class="Insecta" family="Tettigoniidae" genus="Neoconocephalus" kingdom="Animalia" order="Orthoptera" pageId="77" pageNumber="74" phylum="Arthropoda" rank="genus">
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<emphasis box="[794,960,348,369]" italics="true" pageId="77" pageNumber="74">Neoconocephalus</emphasis>
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</taxonomicName>
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it differs in having the ovipositor relatively very short and distinctly bent (long and straight or weakly downcurved in
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<taxonomicName box="[1084,1250,404,425]" class="Insecta" family="Tettigoniidae" genus="Neoconocephalus" kingdom="Animalia" order="Orthoptera" pageId="77" pageNumber="74" phylum="Arthropoda" rank="genus">
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<emphasis box="[1084,1250,404,425]" italics="true" pageId="77" pageNumber="74">Neoconocephalus</emphasis>
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</taxonomicName>
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), male cerci with apical teeth bent under acute angle (right angle in
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<taxonomicName box="[794,960,461,482]" class="Insecta" family="Tettigoniidae" genus="Neoconocephalus" kingdom="Animalia" order="Orthoptera" pageId="77" pageNumber="74" phylum="Arthropoda" rank="genus">
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<emphasis box="[794,960,461,482]" italics="true" pageId="77" pageNumber="74">Neoconocephalus</emphasis>
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</taxonomicName>
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), and very thin, unarmed fore and mid
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<taxonomicName box="[794,872,489,511]" class="Insecta" family="Tettigoniidae" genus="Sphyrometopa" kingdom="Animalia" order="Orthoptera" pageId="77" pageNumber="74" phylum="Arthropoda" rank="species" species="femora">femora</taxonomicName>
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. From
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<taxonomicName box="[947,1068,490,511]" class="Insecta" family="Tettigoniidae" genus="Paroxyprora" kingdom="Animalia" order="Orthoptera" pageId="77" pageNumber="74" phylum="Arthropoda" rank="genus">
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<emphasis box="[947,1068,490,511]" italics="true" pageId="77" pageNumber="74">Paroxyprora</emphasis>
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</taxonomicName>
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this new genus differs in the shape of the ovipositor (weakly curved, dilated midlength in
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<taxonomicName box="[937,1058,546,567]" class="Insecta" family="Tettigoniidae" genus="Paroxyprora" kingdom="Animalia" order="Orthoptera" pageId="77" pageNumber="74" phylum="Arthropoda" rank="genus">
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<emphasis box="[937,1058,546,567]" italics="true" pageId="77" pageNumber="74">Paroxyprora</emphasis>
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</taxonomicName>
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and characteristics of the male stridulatory file (short, reduced in
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<taxonomicName box="[1183,1313,575,596]" class="Insecta" family="Tettigoniidae" genus="Metacaputus" kingdom="Animalia" order="Orthoptera" pageId="77" pageNumber="74" phylum="Arthropoda" rank="genus">
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<emphasis box="[1183,1313,575,596]" italics="true" pageId="77" pageNumber="74">Metacaputus</emphasis>
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</taxonomicName>
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, long, multidentate in
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<taxonomicName box="[964,1085,603,624]" class="Insecta" family="Tettigoniidae" genus="Paroxyprora" kingdom="Animalia" order="Orthoptera" pageId="77" pageNumber="74" phylum="Arthropoda" rank="genus">
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<emphasis box="[964,1085,603,624]" italics="true" pageId="77" pageNumber="74">Paroxyprora</emphasis>
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</taxonomicName>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="77" pageNumber="74" type="etymology">
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<paragraph blockId="77.[794,1384,207,737]" pageId="77" pageNumber="74">
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<emphasis box="[817,993,631,652]" italics="true" pageId="77" pageNumber="74">Etymology.— The</emphasis>
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generic epithet is derived from Latin words
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<emphasis box="[867,955,659,680]" italics="true" pageId="77" pageNumber="74">“Meta- =</emphasis>
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pyramidal” and
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<emphasis box="[1136,1202,659,680]" italics="true" pageId="77" pageNumber="74">“caput</emphasis>
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= head,” indicating the strongly triangular shape of the head of the only species of the genus.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |