669 lines
56 KiB
XML
669 lines
56 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.6280324" ID-GBIF-Dataset="647046fe-0ee5-401d-b4f8-aea0355fde7f" ID-GBIF-Taxon="124407472" ID-Zenodo-Dep="6280324" checkinTime="1471630429166" checkinUser="plazi" docAuthor="Piotr Naskrecki" docDate="2000" docId="634387D1A301FFE216D4FF0CFB443F01" docLanguage="en" docName="Naskrecki_KCR_2000.pdf.imf" docOrigin="Philadelphia, PA: The Orthopterists Society at the Academy of Natural Sciences of Philadelphia," docStyle="DocumentStyle{}" docTitle="Copiphora hastata Naskrecki, sp. n." docType="treatment" docVersion="8" lastPageId="44" lastPageNumber="41" masterDocId="9F7AFFA9A32AFFCE167AFFC3FFF13C6E" masterDocTitle="Katydids of Costa Rica / Vol. 1, Systematics and bioacoustics of the cone-head katydids (Orthoptera: Tettigoniidae: Conocephalinae sensu lato)." pageId="43" pageNumber="40" updateTime="1645797358202" updateUser="ExternalLinkService">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>Katydids of Costa Rica / Vol. 1, Systematics and bioacoustics of the cone-head katydids (Orthoptera: Tettigoniidae: Conocephalinae sensu lato).</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Piotr Naskrecki</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:originInfo>
|
||
<mods:dateIssued>2000</mods:dateIssued>
|
||
<mods:publisher>The Orthopterists Society at the Academy of Natural Sciences of Philadelphia,</mods:publisher>
|
||
<mods:place>
|
||
<mods:placeTerm>Philadelphia, PA</mods:placeTerm>
|
||
</mods:place>
|
||
</mods:originInfo>
|
||
<mods:classification>book</mods:classification>
|
||
<mods:identifier type="DOI">10.5281/zenodo.270035</mods:identifier>
|
||
<mods:identifier type="GBIF-Dataset">647046fe-0ee5-401d-b4f8-aea0355fde7f</mods:identifier>
|
||
<mods:identifier type="Zenodo-Dep">270035</mods:identifier>
|
||
</mods:mods>
|
||
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6280324" ID-GBIF-Taxon="124407472" ID-Zenodo-Dep="6280324" LSID="urn:lsid:plazi:treatment:634387D1A301FFE216D4FF0CFB443F01" httpUri="http://treatment.plazi.org/id/634387D1A301FFE216D4FF0CFB443F01" lastPageId="44" lastPageNumber="41" pageId="43" pageNumber="40">
|
||
<subSubSection box="[174,558,207,229]" pageId="43" pageNumber="40" type="nomenclature">
|
||
<paragraph blockId="43.[174,622,207,286]" box="[174,558,207,229]" pageId="43" pageNumber="40">
|
||
<taxonomicName authority="Naskrecki" authorityName="Naskrecki" box="[174,492,207,229]" class="Insecta" family="Tettigoniidae" genus="Copiphora" kingdom="Animalia" order="Orthoptera" pageId="43" pageNumber="40" phylum="Arthropoda" rank="species" species="hastata" status="sp. n.">
|
||
<emphasis bold="true" box="[174,377,207,229]" italics="true" pageId="43" pageNumber="40">Copiphora hastata</emphasis>
|
||
Naskrecki
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel box="[501,558,207,228]" pageId="43" pageNumber="40" rank="species">sp. n.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="43" pageNumber="40" type="description">
|
||
<paragraph blockId="43.[174,622,207,286]" box="[174,622,235,257]" pageId="43" pageNumber="40">Common name: Brown-faced spear bearer</paragraph>
|
||
<paragraph blockId="43.[174,622,207,286]" box="[174,543,264,286]" pageId="43" pageNumber="40">
|
||
<figureCitation box="[174,242,264,286]" captionStart="FIG. 7" captionStartId="38.[151,195,1684,1706]" captionTargetBox="[212,1301,204,1629]" captionTargetId="figure@38.[212,1302,204,1629]" captionTargetPageId="38" captionText="FIG. 7. Copiphora hastata. A. male - habitus, B. male face, C. male head and pronotum, dorsal view, D. titillators, posterior view (everted), E. male cerci, posterior view, F. male cerci, dorsal view." httpUri="https://zenodo.org/record/270042/files/figure.png" pageId="43" pageNumber="40">Figs. 7</figureCitation>
|
||
A-E, 36E, 42C, 55D-E,
|
||
<figureCitation box="[476,543,264,286]" captionStart="MAPS 1 - 6" captionStartId="157.[174,246,1655,1677]" captionTargetBox="[264,1285,204,1639]" captionTargetId="figure@157.[264,1285,204,1639]" captionTargetPageId="157" captionText="MAPS 1 - 6. Distribution of species of Conocephalinae s. l. in Costa Rica. 1. Conocephalus angustifrons, C. cinereus, 2. Conocephalus saltator, C. ictus, 3. Conocephalus magdalenae, Orchelimum fraternum, Lipotactomimus rowelli, 4. Lirometopum coronatum, Acantheremus colwelli, 5. Copiphora rhinoceros, C. cochleata, C. brevicauda costaricensis, 6. Copiphora hastata, C. cultricornis, C. ottei." httpUri="https://zenodo.org/record/270095/files/figure.png" pageId="43" pageNumber="40">Map 6</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="43" pageNumber="40" type="materials_examined">
|
||
<paragraph blockId="43.[174,736,321,400]" box="[174,692,321,343]" pageId="43" pageNumber="40">
|
||
<typeStatus box="[174,226,321,342]" pageId="43" pageNumber="40">Type</typeStatus>
|
||
locality:
|
||
<collectingCountry box="[322,437,321,343]" name="Costa Rica" pageId="43" pageNumber="40">Costa Rica</collectingCountry>
|
||
:
|
||
<collectingRegion box="[445,531,321,343]" country="Costa Rica" name="Heredia" pageId="43" pageNumber="40">Heredia</collectingRegion>
|
||
Prov., La Selva
|
||
</paragraph>
|
||
<paragraph blockId="43.[174,736,321,400]" pageId="43" pageNumber="40">
|
||
Biological Station;
|
||
<typeStatus box="[429,476,350,371]" pageId="43" pageNumber="40">type</typeStatus>
|
||
depository: Academy of Natural Sciences, Philadelphia –
|
||
<typeStatus box="[579,673,378,400]" pageId="43" pageNumber="40">holotype</typeStatus>
|
||
male
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="43" pageNumber="40" type="description">
|
||
<paragraph blockId="43.[174,763,435,1541]" pageId="43" pageNumber="40">
|
||
<emphasis box="[196,441,435,457]" italics="true" pageId="43" pageNumber="40">Diagnostic description.—</emphasis>
|
||
Body large, with wings in both sexes extending well beyond apex of abdomen (
|
||
<figureCitation box="[671,737,464,486]" captionStart="FIG. 7" captionStartId="38.[151,195,1684,1706]" captionTargetBox="[212,1301,204,1629]" captionTargetId="figure@38.[212,1302,204,1629]" captionTargetPageId="38" captionText="FIG. 7. Copiphora hastata. A. male - habitus, B. male face, C. male head and pronotum, dorsal view, D. titillators, posterior view (everted), E. male cerci, posterior view, F. male cerci, dorsal view." httpUri="https://zenodo.org/record/270042/files/figure.png" pageId="43" pageNumber="40">Figs. 7</figureCitation>
|
||
A, 36E). Fastigium of vertex forming large horn, somewhat flattened laterally, especially in its apical part, fastigium always slightly dilated subapically, about 3 times as long as eye diameter; dorsal part of fastigium strongly granulose (
|
||
<figureCitation box="[293,354,606,629]" captionStart="FIG. 7" captionStartId="38.[151,195,1684,1706]" captionTargetBox="[212,1301,204,1629]" captionTargetId="figure@38.[212,1302,204,1629]" captionTargetPageId="38" captionText="FIG. 7. Copiphora hastata. A. male - habitus, B. male face, C. male head and pronotum, dorsal view, D. titillators, posterior view (everted), E. male cerci, posterior view, F. male cerci, dorsal view." httpUri="https://zenodo.org/record/270042/files/figure.png" pageId="43" pageNumber="40">Fig. 7</figureCitation>
|
||
C). Face convex; genal carinae of head well developed, granulose. Pronotum smooth, flat dorsally; anterior dorsal margin straight, posterior margin weakly convex; lateral lobe of pronotum about 1.6 times longer than high. Male stridulatory file weakly curved,
|
||
<quantity box="[174,250,749,771]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="43" pageNumber="40" unit="mm" value="3.2">3.2 mm</quantity>
|
||
long, 275 μm wide, with 147 teeth; teeth in proximal half of file wider, thinner and more densely arranged, transition between two
|
||
<typeStatus box="[426,483,807,828]" pageId="43" pageNumber="40">types</typeStatus>
|
||
sudden (
|
||
<figureCitation box="[584,658,806,828]" captionStart="FIG. 42" captionStartId="140.[151,195,1684,1706]" captionTargetBox="[204,1305,204,1639]" captionTargetId="figure@140.[204,1306,204,1640]" captionTargetPageId="140" captionText="FIG. 42. Stridulatory files of species of Lirometopum and Copiphora. A. L. coronatum, B. C. cultricornis, C. C. hastata, D. C. brevicauda brevicauda, E. C. brevicauda costaricensis, F. C. ottei." httpUri="https://zenodo.org/record/270078/files/figure.png" pageId="43" pageNumber="40">Fig. 42</figureCitation>
|
||
C); mirror approximately oval; small, parallel vein next to AA1 present.
|
||
</paragraph>
|
||
<paragraph blockId="43.[174,763,435,1541]" pageId="43" pageNumber="40">
|
||
Ventral spines of
|
||
<taxonomicName box="[395,472,892,914]" class="Insecta" family="Tettigoniidae" genus="Sphyrometopa" kingdom="Animalia" order="Orthoptera" pageId="43" pageNumber="40" phylum="Arthropoda" rank="species" species="femora">femora</taxonomicName>
|
||
relatively short and thin, green; all genicular lobes armed with sharp spines, sometimes spine on anterior lobes of fore femur obtuse. Male cercus thick, blunt, and somewhat incurved, slightly thicker at midlength than at base; ventral, subapical spine of cercus long, thin and pointed; apex of cercus with sharply triangular lobe underneath (
|
||
<figureCitation box="[597,664,1063,1085]" captionStart="FIG. 7" captionStartId="38.[151,195,1684,1706]" captionTargetBox="[212,1301,204,1629]" captionTargetId="figure@38.[212,1302,204,1629]" captionTargetPageId="38" captionText="FIG. 7. Copiphora hastata. A. male - habitus, B. male face, C. male head and pronotum, dorsal view, D. titillators, posterior view (everted), E. male cerci, posterior view, F. male cerci, dorsal view." httpUri="https://zenodo.org/record/270042/files/figure.png" pageId="43" pageNumber="40">Fig. 7</figureCitation>
|
||
D). Male subgenital plate narrowed toward apex, apex straight; styli short, about 2.5 times as long as wide. Female subgenital plate approximately triangular, with distinct, triangular apical incision. Ovipositor extremely long, straight, more than 3 times as long as hind femur (ratio ovipositor/hind femur 3.3-3.5) (
|
||
<figureCitation box="[511,573,1234,1256]" captionStart="FIG. 7" captionStartId="38.[151,195,1684,1706]" captionTargetBox="[212,1301,204,1629]" captionTargetId="figure@38.[212,1302,204,1629]" captionTargetPageId="38" captionText="FIG. 7. Copiphora hastata. A. male - habitus, B. male face, C. male head and pronotum, dorsal view, D. titillators, posterior view (everted), E. male cerci, posterior view, F. male cerci, dorsal view." httpUri="https://zenodo.org/record/270042/files/figure.png" pageId="43" pageNumber="40">Fig. 7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="43.[174,763,435,1541]" pageId="43" pageNumber="40">
|
||
<emphasis box="[196,299,1263,1284]" italics="true" pageId="43" pageNumber="40">Coloration</emphasis>
|
||
.— General coloration light green (
|
||
<figureCitation box="[664,735,1263,1285]" captionStart="FIG. 36" captionStartId="134.[151,196,1655,1677]" captionTargetBox="[239,1270,204,1639]" captionTargetId="figure@134.[239,1270,204,1640]" captionTargetPageId="134" captionText="FIG. 36. Costa Rican species of Conocephalinae s. l. A. Conocephalus angustifrons - male, B. Caulopsis cuspidata - female, C. Acantheremus colwelli - male, D. Vestria punctata - male and female, in copula, E. Copiphora hastata - male, F. Podacanthophorus vargasi - female, G. Copiphora rhinoceros - male eating a snail, H. Subria sylvestris - male, brown form." httpUri="https://zenodo.org/record/270072/files/figure.png" pageId="43" pageNumber="40">Fig. 36</figureCitation>
|
||
E). Fastigium of vertex yellowish-brown; face with large, oval, brown spot covering most of the frons, upper portion of the clypeus, and mandibles (
|
||
<figureCitation box="[557,616,1348,1370]" captionStart="FIG. 7" captionStartId="38.[151,195,1684,1706]" captionTargetBox="[212,1301,204,1629]" captionTargetId="figure@38.[212,1302,204,1629]" captionTargetPageId="38" captionText="FIG. 7. Copiphora hastata. A. male - habitus, B. male face, C. male head and pronotum, dorsal view, D. titillators, posterior view (everted), E. male cerci, posterior view, F. male cerci, dorsal view." httpUri="https://zenodo.org/record/270042/files/figure.png" pageId="43" pageNumber="40">Fig. 7</figureCitation>
|
||
B); remainder of head yellow. Pronotum green, with two dark brown to dark olive green stripes along lateral dorsal edges of pronotum. Tegmina green, never mottled with dots; edges of tegmina, especially in their apical portions, red; male stridulatory file usually green; legs green, with green femoral spines. Abdomen green.
|
||
</paragraph>
|
||
<paragraph blockId="43.[196,456,1577,1599]" box="[196,456,1577,1599]" pageId="43" pageNumber="40">
|
||
<emphasis box="[196,338,1577,1598]" italics="true" pageId="43" pageNumber="40">Measurements</emphasis>
|
||
.—
|
||
<tableCitation box="[375,452,1577,1599]" captionStart="TABLE 4" captionStartId="33.[174,254,207,228]" captionTargetBox="[174,1380,265,1093]" captionTargetPageId="33" captionText="TABLE 4. Body measurements of species of Copiphora (all measurements are lengths in mm: range, mean ± SD)" httpUri="http://table.plazi.org/id/BF95664FA30BFFEF16D4FF0CFAC63C8B" pageId="43" pageNumber="40" tableUuid="BF95664FA30BFFEF16D4FF0CFAC63C8B">Table 4</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="43.[174,763,1634,1741]" lastBlockId="43.[794,1383,207,800]" pageId="43" pageNumber="40">
|
||
<emphasis box="[196,518,1634,1655]" italics="true" pageId="43" pageNumber="40">
|
||
Bioacoustics.—
|
||
<taxonomicName box="[348,518,1634,1655]" class="Insecta" family="Tettigoniidae" genus="Copiphora" kingdom="Animalia" order="Orthoptera" pageId="43" pageNumber="40" phylum="Arthropoda" rank="species" species="hastata">Copiphora hastata</taxonomicName>
|
||
</emphasis>
|
||
has a call similar to that of
|
||
<emphasis box="[200,346,1662,1684]" italics="true" pageId="43" pageNumber="40">
|
||
<taxonomicName box="[200,341,1662,1684]" class="Insecta" family="Tettigoniidae" genus="Copiphora" kingdom="Animalia" order="Orthoptera" pageId="43" pageNumber="40" phylum="Arthropoda" rank="species" species="cultricornis">C. cultricornis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
although the series of chirps are shorter and the intervals between individual chirps are much shorter. Also, each chirp consists of a much higher number of pulses than that of
|
||
<emphasis box="[1056,1202,207,228]" italics="true" pageId="43" pageNumber="40">
|
||
<taxonomicName box="[1056,1199,207,228]" class="Insecta" family="Tettigoniidae" genus="Copiphora" kingdom="Animalia" order="Orthoptera" pageId="43" pageNumber="40" phylum="Arthropoda" rank="species" species="cultricornis">C. cultricornis</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Individual series of chirps last about
|
||
<geoCoordinate box="[1013,1061,235,257]" direction="south" orientation="latitude" pageId="43" pageNumber="40" precision="5555" value="-3.5">3.5 s</geoCoordinate>
|
||
and are separated by several seconds to several minutes of silence (
|
||
<figureCitation box="[1216,1291,264,286]" captionStart="FIG. 55" captionStartId="153.[174,218,1655,1677]" captionTargetBox="[249,1308,204,1636]" captionTargetId="figure@153.[249,1308,204,1636]" captionTargetPageId="153" captionText="FIG. 55. Oscillograms of male airborne calls and substrate tremulations of Costa Rican species of Conocephalinae s. l. A. Copiphora cultricornis, airborne call, B. ditto, 0.5 s fragment, C. ditto, substrate tremulations, D. Copiphora hastata, E. ditto, 0.5 s fragment, F. Copiphora brevicauda costaricensis, G. ditto, 0.5 s fragment." httpUri="https://zenodo.org/record/270091/files/figure.png" pageId="43" pageNumber="40">Fig. 55</figureCitation>
|
||
D). Each chirp lasts 153.2 - 241.5 ms and their length increases towards the end of the series. The chirps consist of 5 - 14 pulses, and the amplitude increases towards the end of each pulse (
|
||
<figureCitation box="[924,999,378,400]" captionStart="FIG. 55" captionStartId="153.[174,218,1655,1677]" captionTargetBox="[249,1308,204,1636]" captionTargetId="figure@153.[249,1308,204,1636]" captionTargetPageId="153" captionText="FIG. 55. Oscillograms of male airborne calls and substrate tremulations of Costa Rican species of Conocephalinae s. l. A. Copiphora cultricornis, airborne call, B. ditto, 0.5 s fragment, C. ditto, substrate tremulations, D. Copiphora hastata, E. ditto, 0.5 s fragment, F. Copiphora brevicauda costaricensis, G. ditto, 0.5 s fragment." httpUri="https://zenodo.org/record/270091/files/figure.png" pageId="43" pageNumber="40">Fig. 55</figureCitation>
|
||
E). Most of the energy of the call is allocated between 8.6 and 11.6 kHz.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="43" pageNumber="40" type="biology_ecology">
|
||
<paragraph blockId="43.[794,1383,207,800]" pageId="43" pageNumber="40">
|
||
Typical of the genus, the airborne signals of
|
||
<taxonomicName box="[1285,1382,435,457]" class="Insecta" family="Tettigoniidae" genus="Copiphora" kingdom="Animalia" order="Orthoptera" pageId="43" pageNumber="40" phylum="Arthropoda" rank="species" species="hastata">
|
||
<emphasis box="[1285,1382,435,457]" italics="true" pageId="43" pageNumber="40">C. hastata</emphasis>
|
||
</taxonomicName>
|
||
are accompanied by substrate tremulations. They are produced both in the absence of the female as well as after the female approaches the male. The substrate signals are similar to those observed in other species of
|
||
<taxonomicName box="[794,894,578,599]" class="Insecta" family="Tettigoniidae" genus="Copiphora" kingdom="Animalia" order="Orthoptera" pageId="43" pageNumber="40" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[794,894,578,599]" italics="true" pageId="43" pageNumber="40">Copiphora</emphasis>
|
||
</taxonomicName>
|
||
, and consist of rapid, shaking movements of the entire body, and unlike
|
||
<emphasis box="[1086,1333,607,628]" italics="true" pageId="43" pageNumber="40">
|
||
<taxonomicName box="[1086,1329,607,628]" class="Insecta" family="Tettigoniidae" genus="Lirometopum" kingdom="Animalia" order="Orthoptera" pageId="43" pageNumber="40" phylum="Arthropoda" rank="species" species="coronatum">Lirometopum coronatum</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
lack any percussive behavior. Tremulations are produced in series of bouts at 26 Hz. After the initial contact with the female, the male almost entirely ceases to produce airborne signals and switches to tremulations. Females respond with nearly identical tremulations but of much lower amplitude.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="43" pageNumber="40" type="distribution">
|
||
<paragraph blockId="43.[794,1384,835,1456]" pageId="43" pageNumber="40">
|
||
<emphasis box="[817,930,835,856]" italics="true" pageId="43" pageNumber="40">Remarks.—</emphasis>
|
||
This new species is widely distributed in
|
||
<collectingCountry box="[794,907,863,885]" name="Costa Rica" pageId="43" pageNumber="40">Costa Rica</collectingCountry>
|
||
, with the exception of the southern portion of
|
||
<collectingRegion box="[794,914,892,913]" country="Costa Rica" name="Puntarenas" pageId="43" pageNumber="40">Puntarenas</collectingRegion>
|
||
Prov., where it is replaced by the closely related
|
||
<taxonomicName box="[853,994,920,942]" class="Insecta" family="Tettigoniidae" genus="Copiphora" kingdom="Animalia" order="Orthoptera" pageId="43" pageNumber="40" phylum="Arthropoda" rank="species" species="cultricornis">
|
||
<emphasis box="[853,994,920,942]" italics="true" pageId="43" pageNumber="40">C. cultricornis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[1007,1077,920,942]" captionStart="MAPS 1 - 6" captionStartId="157.[174,246,1655,1677]" captionTargetBox="[264,1285,204,1639]" captionTargetId="figure@157.[264,1285,204,1639]" captionTargetPageId="157" captionText="MAPS 1 - 6. Distribution of species of Conocephalinae s. l. in Costa Rica. 1. Conocephalus angustifrons, C. cinereus, 2. Conocephalus saltator, C. ictus, 3. Conocephalus magdalenae, Orchelimum fraternum, Lipotactomimus rowelli, 4. Lirometopum coronatum, Acantheremus colwelli, 5. Copiphora rhinoceros, C. cochleata, C. brevicauda costaricensis, 6. Copiphora hastata, C. cultricornis, C. ottei." httpUri="https://zenodo.org/record/270095/files/figure.png" pageId="43" pageNumber="40">Map 6</figureCitation>
|
||
). It has also been collected in
|
||
<collectingCountry box="[794,908,949,971]" name="Nicaragua" pageId="43" pageNumber="40">Nicaragua</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="43" pageNumber="40" type="diagnosis">
|
||
<paragraph blockId="43.[794,1384,835,1456]" pageId="43" pageNumber="40">
|
||
<taxonomicName box="[817,915,977,999]" class="Insecta" family="Tettigoniidae" genus="Copiphora" kingdom="Animalia" order="Orthoptera" pageId="43" pageNumber="40" phylum="Arthropoda" rank="species" species="hastata">
|
||
<emphasis box="[817,915,977,999]" italics="true" pageId="43" pageNumber="40">C. hastata</emphasis>
|
||
</taxonomicName>
|
||
can be easily identified among Costa Rican members of the genus by the presence of a dark brown patch covering most of the face, and the shape of the fastigium of vertex. Females of this new species as well as those of
|
||
<taxonomicName box="[883,1024,1092,1113]" class="Insecta" family="Tettigoniidae" genus="Copiphora" kingdom="Animalia" order="Orthoptera" pageId="43" pageNumber="40" phylum="Arthropoda" rank="species" species="cultricornis">
|
||
<emphasis box="[883,1024,1092,1113]" italics="true" pageId="43" pageNumber="40">C. cultricornis</emphasis>
|
||
</taxonomicName>
|
||
have consistently the longest ovipositors among all
|
||
<taxonomicName box="[998,1098,1120,1141]" class="Insecta" family="Tettigoniidae" genus="Copiphora" kingdom="Animalia" order="Orthoptera" pageId="43" pageNumber="40" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[998,1098,1120,1141]" italics="true" pageId="43" pageNumber="40">Copiphora</emphasis>
|
||
</taxonomicName>
|
||
spp. in
|
||
<collectingCountry box="[1182,1295,1120,1142]" name="Costa Rica" pageId="43" pageNumber="40">Costa Rica</collectingCountry>
|
||
(and, in fact, all
|
||
<taxonomicName box="[891,1046,1149,1171]" class="Insecta" family="Tettigoniidae" kingdom="Animalia" order="Orthoptera" pageId="43" pageNumber="40" phylum="Arthropoda" rank="family">Tettigoniidae</taxonomicName>
|
||
). From the closely related
|
||
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Copiphora" kingdom="Animalia" order="Orthoptera" pageId="43" pageNumber="40" phylum="Arthropoda" rank="species" species="cultricornis">
|
||
<emphasis italics="true" pageId="43" pageNumber="40">C. cultricornis</emphasis>
|
||
</taxonomicName>
|
||
it differs in the coloration of the face (yellow in
|
||
<taxonomicName box="[824,968,1206,1227]" class="Insecta" family="Tettigoniidae" genus="Copiphora" kingdom="Animalia" order="Orthoptera" pageId="43" pageNumber="40" phylum="Arthropoda" rank="species" species="cultricornis">
|
||
<emphasis box="[824,968,1206,1227]" italics="true" pageId="43" pageNumber="40">C. cultricornis</emphasis>
|
||
</taxonomicName>
|
||
) and tegmina (posterior edges of tegmina reddish in
|
||
<taxonomicName box="[965,1061,1234,1256]" class="Insecta" family="Tettigoniidae" genus="Copiphora" kingdom="Animalia" order="Orthoptera" pageId="43" pageNumber="40" phylum="Arthropoda" rank="species" species="hastata">
|
||
<emphasis box="[965,1061,1234,1256]" italics="true" pageId="43" pageNumber="40">C. hastata</emphasis>
|
||
</taxonomicName>
|
||
, green in
|
||
<taxonomicName box="[1162,1302,1234,1256]" class="Insecta" family="Tettigoniidae" genus="Copiphora" kingdom="Animalia" order="Orthoptera" pageId="43" pageNumber="40" phylum="Arthropoda" rank="species" species="cultricornis">
|
||
<emphasis box="[1162,1302,1234,1256]" italics="true" pageId="43" pageNumber="40">C. cultricornis</emphasis>
|
||
</taxonomicName>
|
||
), shape of the fastigium of vertex (distinctly longer and more flattened laterally in
|
||
<taxonomicName box="[990,1135,1291,1313]" class="Insecta" family="Tettigoniidae" genus="Copiphora" kingdom="Animalia" order="Orthoptera" pageId="43" pageNumber="40" phylum="Arthropoda" rank="species" species="cultricornis">
|
||
<emphasis box="[990,1135,1291,1313]" italics="true" pageId="43" pageNumber="40">C. cultricornis</emphasis>
|
||
</taxonomicName>
|
||
), characteristics of the male’s stridulatory file (file with more teeth in
|
||
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Copiphora" kingdom="Animalia" order="Orthoptera" pageId="43" pageNumber="40" phylum="Arthropoda" rank="species" species="cultricornis">
|
||
<emphasis italics="true" pageId="43" pageNumber="40">C. cultricornis</emphasis>
|
||
</taxonomicName>
|
||
) as well as the characteristics of the male’s call (
|
||
<figureCitation box="[841,918,1377,1399]" captionStart="FIG. 55" captionStartId="153.[174,218,1655,1677]" captionTargetBox="[249,1308,204,1636]" captionTargetId="figure@153.[249,1308,204,1636]" captionTargetPageId="153" captionText="FIG. 55. Oscillograms of male airborne calls and substrate tremulations of Costa Rican species of Conocephalinae s. l. A. Copiphora cultricornis, airborne call, B. ditto, 0.5 s fragment, C. ditto, substrate tremulations, D. Copiphora hastata, E. ditto, 0.5 s fragment, F. Copiphora brevicauda costaricensis, G. ditto, 0.5 s fragment." httpUri="https://zenodo.org/record/270091/files/figure.png" pageId="43" pageNumber="40">Figs. 55</figureCitation>
|
||
D-E). Tremulations of these two species, however, are virtually identical, suggesting that they play little or no role in species recognition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="44" lastPageNumber="41" pageId="43" pageNumber="40" type="materials_examined">
|
||
<paragraph blockId="43.[794,1383,1490,1737]" lastBlockId="44.[151,741,206,1727]" lastPageId="44" lastPageNumber="41" pageId="43" pageNumber="40">
|
||
<emphasis box="[817,1030,1491,1512]" italics="true" pageId="43" pageNumber="40">Material examined.—</emphasis>
|
||
<emphasis bold="true" box="[1037,1357,1490,1513]" pageId="43" pageNumber="40">
|
||
<collectingCountry box="[1037,1192,1490,1513]" name="Costa Rica" pageId="43" pageNumber="40">COSTA RICA</collectingCountry>
|
||
:
|
||
<collectingRegion box="[1203,1296,1491,1513]" country="Costa Rica" name="Alajuela" pageId="43" pageNumber="40">Alajuela</collectingRegion>
|
||
Prov.
|
||
</emphasis>
|
||
,
|
||
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="43" pageNumber="40" unit="km" value="2.0">2 km</quantity>
|
||
S Pital,
|
||
<date box="[918,1154,1518,1541]" pageId="43" pageNumber="40" value="1988-09-05" valueMax="1988-09-28" valueMin="1988-09-05">5 - 28 September 1988</date>
|
||
(coll. F.D. Parker) -
|
||
<specimenCount pageId="43" pageNumber="40" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount box="[860,952,1547,1569]" pageId="43" pageNumber="40" type="female">1 female</specimenCount>
|
||
(
|
||
<typeStatus box="[968,1074,1547,1569]" pageId="43" pageNumber="40">paratypes</typeStatus>
|
||
) (
|
||
<collectionCode box="[1097,1170,1547,1569]" pageId="43" pageNumber="40">EMUS</collectionCode>
|
||
); Estac. San Ramón Oeste, elev.
|
||
<quantity box="[924,992,1575,1597]" metricMagnitude="2" metricUnit="m" metricValue="6.2" pageId="43" pageNumber="40" unit="m" value="620.0">620 m</quantity>
|
||
,
|
||
<date box="[1004,1149,1575,1597]" pageId="43" pageNumber="40" value="1994-04-19">19 April 1994</date>
|
||
(coll. F. Quesada) -
|
||
<specimenCount pageId="43" pageNumber="40" type="female">1 female</specimenCount>
|
||
(
|
||
<typeStatus box="[881,977,1603,1625]" pageId="43" pageNumber="40">paratype</typeStatus>
|
||
) (
|
||
<collectionCode box="[1001,1068,1603,1625]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="43" pageNumber="40">INBio</collectionCode>
|
||
); Zona Protectora de Arenal, Pocosol,
|
||
<date box="[891,1130,1630,1653]" pageId="43" pageNumber="40" value="1994-02-10" valueMax="1994-02-16" valueMin="1994-02-10">10 - 16 February 1994</date>
|
||
(coll. P. Naskrecki) - 1 nymph (
|
||
<collectionCode box="[892,929,1659,1680]" pageId="43" pageNumber="40">PN</collectionCode>
|
||
collection);
|
||
<emphasis bold="true" box="[1072,1228,1658,1680]" pageId="43" pageNumber="40">
|
||
<collectingRegion box="[1072,1162,1658,1680]" country="Costa Rica" name="Cartago" pageId="43" pageNumber="40">Cartago</collectingRegion>
|
||
Prov.
|
||
</emphasis>
|
||
, Monumento Nacional Guayabo, A.C.A.C. Amistad,
|
||
<date box="[1202,1326,1687,1709]" pageId="43" pageNumber="40" value="1994-07-15">15 July 1994</date>
|
||
(coll. G. Fonseca) -
|
||
<specimenCount box="[945,1037,1715,1737]" pageId="43" pageNumber="40" type="female">1 female</specimenCount>
|
||
(
|
||
<typeStatus box="[1053,1149,1715,1737]" pageId="43" pageNumber="40">paratype</typeStatus>
|
||
) (
|
||
<collectionCode box="[1172,1239,1715,1737]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="43" pageNumber="40">INBio</collectionCode>
|
||
); Turrialba,
|
||
<date lastPageId="44" lastPageNumber="41" pageId="43" pageNumber="40" value="1951-06-04">4 June 1951</date>
|
||
(coll. O.L. Cartwright) -
|
||
<specimenCount box="[528,620,207,229]" pageId="44" pageNumber="41" type="female">1 female</specimenCount>
|
||
(
|
||
<typeStatus box="[635,731,207,229]" pageId="44" pageNumber="41">paratype</typeStatus>
|
||
) (
|
||
<collectionCode box="[158,194,235,256]" pageId="44" pageNumber="41">PN</collectionCode>
|
||
collection); same locality,
|
||
<date box="[491,621,235,257]" pageId="44" pageNumber="41" value="1951-06-05">5 June 1951</date>
|
||
(coll. O.L. Cartwright) -
|
||
<specimenCount box="[317,417,263,285]" pageId="44" pageNumber="41" type="female">1 female</specimenCount>
|
||
(
|
||
<typeStatus box="[434,539,263,285]" pageId="44" pageNumber="41">paratype</typeStatus>
|
||
) (
|
||
<collectionCode box="[565,603,263,284]" pageId="44" pageNumber="41">PN</collectionCode>
|
||
collection);
|
||
<emphasis bold="true" box="[151,338,291,313]" pageId="44" pageNumber="41">
|
||
<collectingRegion box="[151,278,291,313]" country="Costa Rica" name="Guanacaste" pageId="44" pageNumber="41">Guanacaste</collectingRegion>
|
||
Prov.
|
||
</emphasis>
|
||
,
|
||
<quantity box="[351,404,291,313]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="44" pageNumber="41" unit="km" value="2.0">2 km</quantity>
|
||
N Colonia Blanca, P. N.
|
||
<collectingRegion box="[664,739,291,313]" country="Puerto Rico" name="Rincon" pageId="44" pageNumber="41">Rincón</collectingRegion>
|
||
de la Vieja, elev.
|
||
<quantity box="[345,416,320,341]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="44" pageNumber="41" unit="m" value="800.0">800 m</quantity>
|
||
,
|
||
<date box="[429,572,320,341]" pageId="44" pageNumber="41" value="1992-06-28">28 June 1992</date>
|
||
(coll. III curso Parataxon.) -
|
||
<specimenCount box="[293,363,348,370]" pageId="44" pageNumber="41" type="male">1 male</specimenCount>
|
||
(
|
||
<typeStatus box="[377,473,348,370]" pageId="44" pageNumber="41">paratype</typeStatus>
|
||
) (
|
||
<collectionCode box="[494,561,348,370]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="44" pageNumber="41">INBio</collectionCode>
|
||
);
|
||
<quantity box="[578,643,348,370]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="44" pageNumber="41" unit="km" value="14.0">14 km</quantity>
|
||
S Cañas,
|
||
<date box="[151,357,376,398]" pageId="44" pageNumber="41" value="1990-10-15" valueMax="1990-10-20" valueMin="1990-10-15">15 - 20 October 1990</date>
|
||
(coll. F.D. Parker) -
|
||
<specimenCount box="[557,624,376,398]" pageId="44" pageNumber="41" type="male">1 male</specimenCount>
|
||
(
|
||
<typeStatus box="[636,732,376,398]" pageId="44" pageNumber="41">paratype</typeStatus>
|
||
) (
|
||
<collectionCode box="[158,231,405,427]" pageId="44" pageNumber="41">EMUS</collectionCode>
|
||
); Estac. Cacao,
|
||
<quantity box="[397,452,404,426]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="44" pageNumber="41" unit="km" value="2.0">2 km</quantity>
|
||
SW del Cerro Cacao, elev.
|
||
<quantity box="[151,302,433,454]" metricMagnitude="3" metricUnit="m" metricValue="1.2" metricValueMax="1.4" metricValueMin="1.0" pageId="44" pageNumber="41" unit="m" value="1200.0" valueMax="1400.0" valueMin="1000.0">1000 - 1400 m</quantity>
|
||
,
|
||
<date box="[314,546,433,455]" pageId="44" pageNumber="41" value="1995-02-12" valueMax="1995-02-17" valueMin="1995-02-12">12 - 17 February 1995</date>
|
||
(coll. S. Avila) -
|
||
<specimenCount pageId="44" pageNumber="41" type="male">1 male</specimenCount>
|
||
(
|
||
<typeStatus box="[215,311,461,483]" pageId="44" pageNumber="41">paratype</typeStatus>
|
||
) (
|
||
<collectionCode box="[330,397,461,483]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="44" pageNumber="41">INBio</collectionCode>
|
||
); Estac. Cacao, Lado suroeste del Volcán Cacao, P.N.
|
||
<collectingRegion box="[361,486,489,510]" country="Costa Rica" name="Guanacaste" pageId="44" pageNumber="41">Guanacaste</collectingRegion>
|
||
, elev.
|
||
<quantity box="[553,703,489,511]" metricMagnitude="3" metricUnit="m" metricValue="1.2" metricValueMax="1.4" metricValueMin="1.0" pageId="44" pageNumber="41" unit="m" value="1200.0" valueMax="1400.0" valueMin="1000.0">1000 - 1400 m</quantity>
|
||
,
|
||
<date pageId="44" pageNumber="41" value="1990-06-15">15 June 1990</date>
|
||
(coll. II curso Parataxon) -
|
||
<specimenCount box="[534,623,518,540]" pageId="44" pageNumber="41" type="female">1 female</specimenCount>
|
||
(
|
||
<typeStatus box="[635,731,518,540]" pageId="44" pageNumber="41">paratype</typeStatus>
|
||
) (
|
||
<collectionCode box="[158,225,546,568]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="44" pageNumber="41">INBio</collectionCode>
|
||
); Estac. Maritza, lado O Vol. Orosí, elev.
|
||
<quantity box="[643,707,546,568]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="44" pageNumber="41" unit="m" value="600.0">600 m</quantity>
|
||
,
|
||
<date pageId="44" pageNumber="41" value="1992-03-10">10 March 1992</date>
|
||
(coll. K. Taylor) -
|
||
<specimenCount box="[450,536,574,596]" pageId="44" pageNumber="41" type="female">1 female</specimenCount>
|
||
(
|
||
<typeStatus box="[548,643,574,596]" pageId="44" pageNumber="41">paratype</typeStatus>
|
||
) (
|
||
<collectionCode box="[662,728,574,596]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="44" pageNumber="41">INBio</collectionCode>
|
||
); Estac. Pitilla,
|
||
<quantity box="[296,350,602,624]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="44" pageNumber="41" unit="km" value="9.0">9 km</quantity>
|
||
S. Santa Cecilia, elev.
|
||
<quantity box="[590,656,603,624]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="44" pageNumber="41" unit="m" value="700.0">700 m</quantity>
|
||
,
|
||
<date pageId="44" pageNumber="41" value="1993-07-18" valueMax="1993-07-23" valueMin="1993-07-18">18 - 23 July 1993</date>
|
||
(coll. C. Moraga) -
|
||
<specimenCount box="[445,533,631,653]" pageId="44" pageNumber="41" type="female">1 female</specimenCount>
|
||
(
|
||
<typeStatus box="[546,642,631,653]" pageId="44" pageNumber="41">paratype</typeStatus>
|
||
) (
|
||
<collectionCode box="[661,728,631,653]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="44" pageNumber="41">INBio</collectionCode>
|
||
); same locality,
|
||
<date box="[302,438,659,681]" pageId="44" pageNumber="41" value="1994-05-15">15 May 1994</date>
|
||
(coll. C. Moraga) -
|
||
<specimenCount box="[649,739,659,681]" pageId="44" pageNumber="41" type="female">1 female</specimenCount>
|
||
(
|
||
<typeStatus box="[158,254,687,709]" pageId="44" pageNumber="41">paratype</typeStatus>
|
||
) (
|
||
<collectionCode box="[277,344,687,709]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="44" pageNumber="41">INBio</collectionCode>
|
||
); Estac. Pitilla,
|
||
<quantity box="[509,564,687,709]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="44" pageNumber="41" unit="km" value="9.0">9 km</quantity>
|
||
St. Cecilia, elev.
|
||
<quantity box="[151,217,716,737]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="44" pageNumber="41" unit="m" value="700.0">700 m</quantity>
|
||
,
|
||
<date box="[229,358,715,737]" pageId="44" pageNumber="41" value="1988-07-15">15 July 1988</date>
|
||
(coll. GNP Biodiversity Survey) -
|
||
<specimenCount pageId="44" pageNumber="41" type="male">1 male</specimenCount>
|
||
(
|
||
<typeStatus box="[218,314,744,766]" pageId="44" pageNumber="41">paratype</typeStatus>
|
||
) (
|
||
<collectionCode box="[336,403,744,766]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="44" pageNumber="41">INBio</collectionCode>
|
||
); same locality,
|
||
<date box="[574,740,743,765]" pageId="44" pageNumber="41" value="1988-08-15">15 August 1988</date>
|
||
(coll. GNP Biodiversity Survey) - 1 nymph (
|
||
<collectionCode box="[602,669,772,794]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="44" pageNumber="41">INBio</collectionCode>
|
||
); same locality,
|
||
<date box="[238,370,800,822]" pageId="44" pageNumber="41" value="1989-06-15">15 June 1989</date>
|
||
(coll. GNP Biodiversity Survey) -
|
||
<specimenCount pageId="44" pageNumber="41" type="male">1 male</specimenCount>
|
||
(
|
||
<typeStatus box="[218,314,828,850]" pageId="44" pageNumber="41">paratype</typeStatus>
|
||
) (
|
||
<collectionCode box="[336,403,828,850]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="44" pageNumber="41">INBio</collectionCode>
|
||
); same locality,
|
||
<date box="[574,740,828,850]" pageId="44" pageNumber="41" value="1993-08-18">18 August 1993</date>
|
||
(coll. P. Rios) -
|
||
<specimenCount box="[301,389,857,879]" pageId="44" pageNumber="41" type="female">1 female</specimenCount>
|
||
(
|
||
<typeStatus box="[401,497,857,879]" pageId="44" pageNumber="41">paratype</typeStatus>
|
||
) (
|
||
<collectionCode box="[516,583,857,879]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="44" pageNumber="41">INBio</collectionCode>
|
||
); same locality,
|
||
<date box="[151,305,885,907]" pageId="44" pageNumber="41" value="1996-06-01" valueMax="1996-06-30" valueMin="1996-06-01">1 - 30 June 1996</date>
|
||
(coll. P. Rios) -
|
||
<specimenCount box="[454,540,885,907]" pageId="44" pageNumber="41" type="female">1 female</specimenCount>
|
||
(
|
||
<typeStatus box="[551,645,885,907]" pageId="44" pageNumber="41">paratype</typeStatus>
|
||
) (
|
||
<collectionCode box="[663,728,885,907]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="44" pageNumber="41">INBio</collectionCode>
|
||
); Estac Cacao, SW side Volcán Cacao, elev.
|
||
<quantity box="[578,719,913,935]" metricMagnitude="3" metricUnit="m" metricValue="1.2" metricValueMax="1.4" metricValueMin="1.0" pageId="44" pageNumber="41" unit="m" value="1200.0" valueMax="1400.0" valueMin="1000.0">1000 - 1400 m</quantity>
|
||
,
|
||
<date pageId="44" pageNumber="41" value="1989-12-01">1 December 1989</date>
|
||
(coll. R. Blanco and C. Chávez) -
|
||
<specimenCount pageId="44" pageNumber="41" type="female">1 female</specimenCount>
|
||
(
|
||
<typeStatus box="[217,313,970,992]" pageId="44" pageNumber="41">paratype</typeStatus>
|
||
) (
|
||
<collectionCode box="[334,401,970,992]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="44" pageNumber="41">INBio</collectionCode>
|
||
); Estac. Mengo, SW side Volcán Cacao, elev.
|
||
<quantity box="[301,388,998,1019]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="44" pageNumber="41" unit="m" value="1100.0">1100 m</quantity>
|
||
,
|
||
<date box="[402,606,998,1020]" pageId="44" pageNumber="41" value="1989-02-15">15 February 1989</date>
|
||
(coll. GNP Biodiversity Survey) -
|
||
<specimenCount box="[383,471,1026,1048]" pageId="44" pageNumber="41" type="female">1 female</specimenCount>
|
||
(
|
||
<typeStatus box="[482,578,1026,1048]" pageId="44" pageNumber="41">paratype</typeStatus>
|
||
) (
|
||
<collectionCode box="[597,663,1026,1048]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="44" pageNumber="41">INBio</collectionCode>
|
||
); Estac. Pitilla,
|
||
<quantity box="[226,280,1055,1077]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="44" pageNumber="41" unit="km" value="9.0">9 km</quantity>
|
||
S Santa Cecilia, elev.
|
||
<quantity box="[509,574,1055,1076]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="44" pageNumber="41" unit="m" value="700.0">700 m</quantity>
|
||
,
|
||
<date box="[585,739,1054,1077]" pageId="44" pageNumber="41" value="1989-03-15">15 March 1989</date>
|
||
(coll. GNP Biodiversity Survey) - 1 nymph (
|
||
<collectionCode box="[601,667,1083,1105]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="44" pageNumber="41">INBio</collectionCode>
|
||
); Finca Pasmompa Estación Pitilla,
|
||
<quantity box="[444,497,1111,1133]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="44" pageNumber="41" unit="km" value="5.0">5 km</quantity>
|
||
SW Santa Cecilia, elev.
|
||
<quantity box="[151,217,1139,1161]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="44" pageNumber="41" unit="m" value="400.0">400 m</quantity>
|
||
,
|
||
<date box="[228,362,1139,1161]" pageId="44" pageNumber="41" value="1989-05-15">15 May 1989</date>
|
||
(coll. GNP Biodiversity Survey) -
|
||
<specimenCount pageId="44" pageNumber="41" type="female">1 female</specimenCount>
|
||
(
|
||
<typeStatus box="[233,326,1167,1189]" pageId="44" pageNumber="41">paratype</typeStatus>
|
||
) (
|
||
<collectionCode box="[345,412,1167,1189]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="44" pageNumber="41">INBio</collectionCode>
|
||
);
|
||
<emphasis bold="true" box="[427,574,1167,1189]" pageId="44" pageNumber="41">
|
||
<collectingRegion box="[427,515,1167,1189]" country="Costa Rica" name="Heredia" pageId="44" pageNumber="41">Heredia</collectingRegion>
|
||
Prov.
|
||
</emphasis>
|
||
, Braulio Carillo N. P., Estac. El Ceibo, elev.
|
||
<quantity box="[451,580,1196,1218]" metricMagnitude="2" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="44" pageNumber="41" unit="m" value="500.0" valueMax="600.0" valueMin="400.0">400 - 600 m</quantity>
|
||
,
|
||
<date pageId="44" pageNumber="41" value="1989-11-15">15 November 1989</date>
|
||
(coll. R. Aguilar and M. Zumbado) -
|
||
<specimenCount box="[661,740,1224,1246]" pageId="44" pageNumber="41" type="male">1 male</specimenCount>
|
||
(
|
||
<typeStatus box="[158,253,1252,1274]" pageId="44" pageNumber="41">paratype</typeStatus>
|
||
) (
|
||
<collectionCode box="[272,338,1252,1274]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="44" pageNumber="41">INBio</collectionCode>
|
||
); Estac. Magsasay, P. N. Braulio Carillo, elev.
|
||
<quantity box="[203,268,1281,1302]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="44" pageNumber="41" unit="m" value="200.0">200 m</quantity>
|
||
,
|
||
<date box="[278,474,1280,1302]" pageId="44" pageNumber="41" value="1990-11-15">15 November 1990</date>
|
||
(coll. M.A. Zumbado) -
|
||
<specimenCount pageId="44" pageNumber="41" type="male">1 male</specimenCount>
|
||
(
|
||
<typeStatus box="[219,315,1309,1331]" pageId="44" pageNumber="41">paratype</typeStatus>
|
||
) (
|
||
<collectionCode box="[338,405,1309,1331]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="44" pageNumber="41">INBio</collectionCode>
|
||
); same locality, elev.
|
||
<quantity box="[635,702,1309,1330]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="44" pageNumber="41" unit="m" value="200.0">200 m</quantity>
|
||
,
|
||
<date pageId="44" pageNumber="41" value="1991-05-15">15 May 1991</date>
|
||
(coll. M.A. Zumbado) -
|
||
<specimenCount box="[527,619,1337,1359]" pageId="44" pageNumber="41" type="female">1 female</specimenCount>
|
||
(
|
||
<typeStatus box="[635,731,1337,1359]" pageId="44" pageNumber="41">paratype</typeStatus>
|
||
) (
|
||
<collectionCode box="[158,225,1365,1387]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="44" pageNumber="41">INBio</collectionCode>
|
||
); same locality, elev.
|
||
<quantity box="[446,511,1365,1387]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="44" pageNumber="41" unit="m" value="200.0">200 m</quantity>
|
||
,
|
||
<date box="[522,651,1365,1387]" pageId="44" pageNumber="41" value="1991-07-15">15 July 1991</date>
|
||
(coll. A. Fernández) -
|
||
<specimenCount box="[294,366,1394,1416]" pageId="44" pageNumber="41" type="male">1 male</specimenCount>
|
||
(
|
||
<typeStatus box="[380,476,1394,1416]" pageId="44" pageNumber="41">paratype</typeStatus>
|
||
) (
|
||
<collectionCode box="[499,566,1394,1416]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="44" pageNumber="41">INBio</collectionCode>
|
||
); Finca La Selva (on Río Puerto Viejo just above jct. with Río Sarapiquí),
|
||
<date pageId="44" pageNumber="41" value="1962-08-03">3 August 1962</date>
|
||
(coll. C.F. Walker) -
|
||
<specimenCount box="[523,617,1450,1472]" pageId="44" pageNumber="41" type="female">1 female</specimenCount>
|
||
(
|
||
<typeStatus box="[634,732,1450,1472]" pageId="44" pageNumber="41">paratype</typeStatus>
|
||
) (
|
||
<collectionCode box="[158,246,1479,1501]" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="44" pageNumber="41">UMMZ</collectionCode>
|
||
); La Selva,
|
||
<date box="[379,650,1478,1500]" pageId="44" pageNumber="41" value="1991-07-01" valueMax="1991-08-31" valueMin="1991-07-01">1 July - 31 August 1991</date>
|
||
(coll. J. Doubles) -
|
||
<specimenCount box="[267,338,1507,1529]" pageId="44" pageNumber="41" type="male">1 male</specimenCount>
|
||
(
|
||
<typeStatus box="[351,447,1507,1529]" pageId="44" pageNumber="41">paratype</typeStatus>
|
||
) (
|
||
<collectionCode box="[468,552,1507,1529]" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="44" pageNumber="41">UMMZ</collectionCode>
|
||
); Puerto Viejo, La Selva Biological Station, elev.
|
||
<quantity box="[461,569,1535,1557]" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="44" pageNumber="41" unit="m" value="100.0" valueMax="150.0" valueMin="50.0">50 - 150 m</quantity>
|
||
,
|
||
<geoCoordinate box="[578,674,1535,1557]" direction="north" orientation="latitude" pageId="44" pageNumber="41" precision="925" value="10.433333">10° 26' N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" pageId="44" pageNumber="41" precision="925" value="-84.01667">84° 1' W</geoCoordinate>
|
||
,
|
||
<date box="[183,317,1563,1585]" pageId="44" pageNumber="41" value="1992-03-01">1 March 1992</date>
|
||
(coll. S. Cunningham) -
|
||
<specimenCount box="[560,627,1563,1585]" pageId="44" pageNumber="41" type="male">1 male</specimenCount>
|
||
(
|
||
<typeStatus box="[638,732,1563,1585]" pageId="44" pageNumber="41">paratype</typeStatus>
|
||
) (
|
||
<collectionCode box="[158,194,1592,1613]" pageId="44" pageNumber="41">PN</collectionCode>
|
||
collection); same locality,
|
||
<date box="[488,650,1591,1613]" pageId="44" pageNumber="41" value="1994-03-15">15 March 1994</date>
|
||
(coll. P. Naskrecki) -
|
||
<specimenCount box="[285,383,1620,1642]" pageId="44" pageNumber="41" type="female">3 females</specimenCount>
|
||
(
|
||
<typeStatus box="[396,502,1620,1642]" pageId="44" pageNumber="41">paratypes</typeStatus>
|
||
) (
|
||
<collectionCode box="[523,558,1620,1641]" pageId="44" pageNumber="41">PN</collectionCode>
|
||
collection); same locality,
|
||
<date box="[239,404,1648,1670]" pageId="44" pageNumber="41" value="1994-04-01" valueMax="1994-04-04" valueMin="1994-04-01">1 - 4 April 1994</date>
|
||
(coll. P. Naskrecki) -
|
||
<specimenCount box="[633,716,1648,1670]" pageId="44" pageNumber="41" type="male">4 males</specimenCount>
|
||
,
|
||
<specimenCount pageId="44" pageNumber="41" type="female">1 female</specimenCount>
|
||
(
|
||
<typeStatus box="[244,345,1676,1698]" pageId="44" pageNumber="41">holotype</typeStatus>
|
||
male,
|
||
<typeStatus box="[426,518,1676,1698]" pageId="44" pageNumber="41">allotype</typeStatus>
|
||
female,
|
||
<typeStatus box="[620,733,1677,1698]" pageId="44" pageNumber="41">paratypes</typeStatus>
|
||
) (
|
||
<collectionCode box="[158,229,1705,1727]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="44" pageNumber="41">ANSP</collectionCode>
|
||
); same locality,
|
||
<date box="[408,649,1704,1726]" pageId="44" pageNumber="41" value="1995-12-07" valueMax="1995-12-10" valueMin="1995-12-07">7 - 10 December 1995</date>
|
||
(coll. P.
|
||
</paragraph>
|
||
<paragraph blockId="44.[771,1361,206,879]" pageId="44" pageNumber="41">
|
||
Naskrecki) -
|
||
<specimenCount box="[906,995,207,229]" pageId="44" pageNumber="41" type="female">1 female</specimenCount>
|
||
(
|
||
<typeStatus box="[1009,1105,207,229]" pageId="44" pageNumber="41">paratype</typeStatus>
|
||
) (
|
||
<collectionCode box="[1125,1160,207,228]" pageId="44" pageNumber="41">PN</collectionCode>
|
||
collection);
|
||
<emphasis bold="true" pageId="44" pageNumber="41">
|
||
<collectingRegion box="[1288,1360,206,228]" country="Costa Rica" name="Limon" pageId="44" pageNumber="41">Limón</collectingRegion>
|
||
Prov.
|
||
</emphasis>
|
||
, Cuatro Esquinas, P. N. Tortuguero, elev. 0 m,
|
||
<date pageId="44" pageNumber="41" value="1989-04-15">15 April 1989</date>
|
||
(coll. R. Aguilar and J. Solano) -
|
||
<specimenCount box="[1266,1360,263,285]" pageId="44" pageNumber="41" type="female">1 female</specimenCount>
|
||
(
|
||
<typeStatus box="[779,875,292,314]" pageId="44" pageNumber="41">paratype</typeStatus>
|
||
) (
|
||
<collectionCode box="[897,964,291,313]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="44" pageNumber="41">INBio</collectionCode>
|
||
); Cuatro Esquinas, Tortuguero N. P.,
|
||
<date box="[771,976,319,342]" pageId="44" pageNumber="41" value="1989-09-15">15 September 1989</date>
|
||
(coll. J. Solano) -
|
||
<specimenCount box="[1178,1253,320,342]" pageId="44" pageNumber="41" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount box="[1267,1360,320,342]" pageId="44" pageNumber="41" type="female">1 female</specimenCount>
|
||
(
|
||
<typeStatus box="[779,887,348,370]" pageId="44" pageNumber="41">paratypes</typeStatus>
|
||
) (
|
||
<collectionCode box="[911,979,348,370]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="44" pageNumber="41">INBio</collectionCode>
|
||
); same locality,
|
||
<date box="[1155,1361,348,370]" pageId="44" pageNumber="41" value="1989-11-15">15 November 1989</date>
|
||
(coll. J. Solano) -
|
||
<specimenCount box="[969,1065,376,398]" pageId="44" pageNumber="41" type="female">1 female</specimenCount>
|
||
(
|
||
<typeStatus box="[1082,1182,376,398]" pageId="44" pageNumber="41">paratype</typeStatus>
|
||
) (
|
||
<collectionCode box="[1206,1275,376,398]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="44" pageNumber="41">INBio</collectionCode>
|
||
); Estac. Cuatro Esquinas, Tortuguero N. P.,
|
||
<date box="[1145,1276,404,426]" pageId="44" pageNumber="41" value="1990-06-15">15 June 1990</date>
|
||
(coll. E. Quesada) -
|
||
<specimenCount box="[899,991,433,455]" pageId="44" pageNumber="41" type="female">1 female</specimenCount>
|
||
(
|
||
<typeStatus box="[1008,1105,433,455]" pageId="44" pageNumber="41">paratype</typeStatus>
|
||
) (
|
||
<collectionCode box="[1128,1195,433,455]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="44" pageNumber="41">INBio</collectionCode>
|
||
); same locality, elev.
|
||
<quantity box="[826,928,461,482]" metricMagnitude="1" metricUnit="m" metricValue="6.0" metricValueMax="12.0" metricValueMin="0.0" pageId="44" pageNumber="41" unit="m" value="60.0" valueMax="120.0" valueMin="0.0">0 - 120 m</quantity>
|
||
,
|
||
<date box="[939,1069,461,483]" pageId="44" pageNumber="41" value="1990-07-16">16 July 1990</date>
|
||
(coll. E. Quesada) -
|
||
<specimenCount box="[1289,1360,461,483]" pageId="44" pageNumber="41" type="male">1 male</specimenCount>
|
||
(
|
||
<typeStatus box="[779,877,489,511]" pageId="44" pageNumber="41">paratype</typeStatus>
|
||
), 2 nymphs (
|
||
<collectionCode box="[1016,1083,489,511]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="44" pageNumber="41">INBio</collectionCode>
|
||
); Estac. Hitoy-Cerere Res. Biol. Hitoy Cerere, Río Cerere, elev.
|
||
<quantity box="[1145,1209,518,539]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="44" pageNumber="41" unit="m" value="200.0">200 m</quantity>
|
||
,
|
||
<date pageId="44" pageNumber="41" value="1990-09-15">15 September 1990</date>
|
||
(coll. G. Carballo) -
|
||
<specimenCount box="[1026,1095,546,568]" pageId="44" pageNumber="41" type="male">1 male</specimenCount>
|
||
(
|
||
<typeStatus box="[1107,1203,546,568]" pageId="44" pageNumber="41">paratype</typeStatus>
|
||
) (
|
||
<collectionCode box="[1222,1289,546,568]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="44" pageNumber="41">INBio</collectionCode>
|
||
); same locality, elev.
|
||
<quantity box="[914,981,574,595]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="44" pageNumber="41" unit="m" value="100.0">100 m</quantity>
|
||
,
|
||
<date box="[993,1127,574,596]" pageId="44" pageNumber="41" value="1991-06-15">15 June 1991</date>
|
||
(coll. G. Carballo) - 1 nymph (
|
||
<collectionCode box="[862,929,602,624]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="44" pageNumber="41">INBio</collectionCode>
|
||
); same locality, elev.
|
||
<quantity box="[1150,1216,603,624]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="44" pageNumber="41" unit="m" value="100.0">100 m</quantity>
|
||
,
|
||
<date box="[1227,1360,602,624]" pageId="44" pageNumber="41" value="1992-06-15">15 June 1992</date>
|
||
(coll. G. Carballo) -
|
||
<specimenCount box="[980,1079,631,653]" pageId="44" pageNumber="41" type="female">2 females</specimenCount>
|
||
(
|
||
<typeStatus box="[1093,1199,631,653]" pageId="44" pageNumber="41">paratypes</typeStatus>
|
||
) (
|
||
<collectionCode box="[1221,1288,631,653]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="44" pageNumber="41">INBio</collectionCode>
|
||
); same locality, elev.
|
||
<quantity box="[915,982,659,680]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="44" pageNumber="41" unit="m" value="100.0">100 m</quantity>
|
||
,
|
||
<date box="[994,1125,659,681]" pageId="44" pageNumber="41" value="1992-07-15">15 July 1992</date>
|
||
(coll. G. Carballo) -
|
||
<specimenCount pageId="44" pageNumber="41" type="female">2 females</specimenCount>
|
||
(
|
||
<typeStatus box="[866,973,687,709]" pageId="44" pageNumber="41">paratypes</typeStatus>
|
||
) (
|
||
<collectionCode box="[992,1059,687,709]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="44" pageNumber="41">INBio</collectionCode>
|
||
); Hamburg Farm, (coll. C.W. Dodge) -
|
||
<specimenCount box="[866,952,715,737]" pageId="44" pageNumber="41" type="female">1 female</specimenCount>
|
||
(
|
||
<typeStatus box="[963,1058,716,738]" pageId="44" pageNumber="41">paratype</typeStatus>
|
||
) (
|
||
<collectionCode box="[1076,1152,716,738]" pageId="44" pageNumber="41">MCZC</collectionCode>
|
||
); same locality, (coll. C.W. Dodge) -
|
||
<specimenCount box="[933,1005,744,766]" pageId="44" pageNumber="41" type="male">1 male</specimenCount>
|
||
(
|
||
<typeStatus box="[1019,1115,744,766]" pageId="44" pageNumber="41">paratype</typeStatus>
|
||
) (
|
||
<collectionCode box="[1138,1215,744,766]" pageId="44" pageNumber="41">MCZC</collectionCode>
|
||
); Manzanillo,
|
||
<collectionCode box="[771,833,772,793]" pageId="44" pageNumber="41">RNFS</collectionCode>
|
||
Gandoca y Manzanillo, elev.
|
||
<quantity box="[1141,1236,772,794]" metricMagnitude="1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="44" pageNumber="41" unit="m" value="50.0" valueMax="100.0" valueMin="0.0">0 - 100 m</quantity>
|
||
,
|
||
<date pageId="44" pageNumber="41" value="1992-10-13">13 October 1992</date>
|
||
(coll. K. Taylor) -
|
||
<specimenCount box="[996,1082,800,822]" pageId="44" pageNumber="41" type="female">1 female</specimenCount>
|
||
(
|
||
<typeStatus box="[1094,1189,800,822]" pageId="44" pageNumber="41">paratype</typeStatus>
|
||
) (
|
||
<collectionCode box="[1207,1273,800,822]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="44" pageNumber="41">INBio</collectionCode>
|
||
);
|
||
<emphasis bold="true" pageId="44" pageNumber="41">
|
||
NICA-
|
||
<collectionCode box="[771,864,828,850]" pageId="44" pageNumber="41">RAGUA</collectionCode>
|
||
:
|
||
</emphasis>
|
||
Zelaya el Recreo,
|
||
<date box="[1050,1240,828,850]" pageId="44" pageNumber="41" value="1984-10-01" valueMax="1984-10-31" valueMin="1984-10-01">1 - 31 October 1984</date>
|
||
(coll. Foret) -
|
||
<specimenCount box="[785,857,857,879]" pageId="44" pageNumber="41" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount box="[867,966,857,879]" pageId="44" pageNumber="41" type="female">2 females</specimenCount>
|
||
(
|
||
<typeStatus box="[980,1087,857,879]" pageId="44" pageNumber="41">paratypes</typeStatus>
|
||
) (
|
||
<collectionCode box="[1107,1194,857,879]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="44" pageNumber="41">MNHN</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |