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<mods:title id="5FFF5904EB1C8D59448C4309C7CFE94B">Review of the mite family Pachylaelapidae (Acari: Mesostigmata)</mods:title>
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<subSubSection id="92BBA8FFFF81FF8AFF2368969EE4FD59" box="[151,752,692,718]" pageId="51" pageNumber="52" type="nomenclature">
<paragraph id="DA1EFB74FF81FF8AFF2368969EE4FD59" blockId="51.[151,752,692,718]" box="[151,752,692,718]" pageId="51" pageNumber="52">
<heading id="81564C18FF81FF8AFF2368969EE4FD59" box="[151,752,692,718]" pageId="51" pageNumber="52">
<emphasis id="E8D52766FF81FF8AFF2368969CF3FD59" bold="true" box="[151,231,692,718]" pageId="51" pageNumber="52">Genus</emphasis>
<taxonomicName id="1DA180F7FF81FF8AFF5A68969EE4FD59" authority="Berlese, 1916" authorityName="Berlese" authorityYear="1916" box="[238,752,692,718]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="51" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="E8D52766FF81FF8AFF5A68969E33FD59" bold="true" box="[238,551,692,718]" pageId="51" pageNumber="52">
<emphasis id="E8D52766FF81FF8AFF5A68969D68FD59" bold="true" box="[238,380,692,718]" italics="true" pageId="51" pageNumber="52">Pachylaella</emphasis>
<bibRefCitation id="BE308685FF81FF8AFE3768969E33FD59" author="Berlese, A." box="[387,551,692,718]" pageId="51" pageNumber="52" pagination="19 - 67" refId="ref39120" refString="Berlese, A. (1916 a) Centuria prima di Acari nuovi. Redia, 12, 19 - 67." type="journal article" year="1916">Berlese, 1916</bibRefCitation>
</emphasis>
(Macrochelidae)
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="92BBA8FFFF81FF8AFF2368D89910FC87" box="[151,1284,762,784]" pageId="51" pageNumber="52" type="reference_group">
<paragraph id="DA1EFB74FF81FF8AFF2368D89910FC87" blockId="51.[151,1284,762,784]" box="[151,1284,762,784]" pageId="51" pageNumber="52">
<taxonomicName id="1DA180F7FF81FF8AFF2368D89DF0FC87" authority="Berlese, 1916 b: 152" authorityName="Berlese" authorityPageNumber="152" authorityYear="1916" box="[151,484,762,784]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="51" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="E8D52766FF81FF8AFF2368D89D04FC87" box="[151,272,762,784]" italics="true" pageId="51" pageNumber="52">Pachylaella</emphasis>
<bibRefCitation id="BE308685FF81FF8AFEA268D89DF0FC87" author="Berlese" box="[278,484,762,784]" firstAuthor="Berlese" page="152" pageId="51" pageNumber="52" pagination="125 - 177" refId="ref39142" refString="Berlese, A. (1916 b) Centuria seconda di Acari nuovi. Redia, 12, 125 - 177." type="journal article" year="1916" yearSuffix="b">Berlese, 1916b: 152</bibRefCitation>
</taxonomicName>
.
<typeStatus id="051A45D6FF81FF8AFE5A68D99E34FC87" box="[494,544,763,784]" pageId="51" pageNumber="52">Type</typeStatus>
species
<taxonomicName id="1DA180F7FF81FF8AFDC268D8981DFC87" authority="Berlese, 1916" authorityName="Berlese" authorityYear="1916" box="[630,1033,762,784]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="robustissima">
<emphasis id="E8D52766FF81FF8AFDC268D89F62FC87" box="[630,886,762,784]" italics="true" pageId="51" pageNumber="52">Pachylaella robustissima</emphasis>
<bibRefCitation id="BE308685FF81FF8AFCC868D8981DFC87" author="Berlese, A." box="[892,1033,762,784]" pageId="51" pageNumber="52" pagination="19 - 67" refId="ref39120" refString="Berlese, A. (1916 a) Centuria prima di Acari nuovi. Redia, 12, 19 - 67." type="journal article" year="1916">Berlese, 1916</bibRefCitation>
</taxonomicName>
, by original designation.
</paragraph>
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<subSubSection id="92BBA8FFFF81FF8AFF23691F9EA2FA91" pageId="51" pageNumber="52" type="discussion">
<paragraph id="DA1EFB74FF81FF8AFF23691F9EA2FA91" blockId="51.[151,1436,829,1286]" pageId="51" pageNumber="52">
<emphasis id="E8D52766FF81FF8AFF23691F9CF5FCC1" bold="true" box="[151,225,829,854]" pageId="51" pageNumber="52">Notes.</emphasis>
The genus
<taxonomicName id="1DA180F7FF81FF8AFED5691F9DF3FCC1" authorityName="Berlese" authorityYear="1916" box="[353,487,829,854]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="51" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="E8D52766FF81FF8AFED5691F9DF3FCC1" box="[353,487,829,854]" italics="true" pageId="51" pageNumber="52">Pachylaella</emphasis>
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was proposed by
<bibRefCitation id="BE308685FF81FF8AFD1A691F9F76FCC1" author="Berlese" box="[686,866,829,854]" firstAuthor="Berlese" pageId="51" pageNumber="52" pagination="125 - 177" refId="ref39142" refString="Berlese, A. (1916 b) Centuria seconda di Acari nuovi. Redia, 12, 125 - 177." type="journal article" year="1916" yearSuffix="b">Berlese (1916b)</bibRefCitation>
with
<taxonomicName id="1DA180F7FF81FF8AFC14691F98AEFCC1" authorityName="Berlese" authorityYear="1916" box="[928,1210,829,854]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="robustissima">
<emphasis id="E8D52766FF81FF8AFC14691F98AEFCC1" box="[928,1210,829,854]" italics="true" pageId="51" pageNumber="52">Pachylaella robustissima</emphasis>
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as
<typeStatus id="051A45D6FF81FF8AFB6A691C991AFCC1" box="[1246,1294,830,854]" pageId="51" pageNumber="52">type</typeStatus>
species, and placed in the family
<taxonomicName id="1DA180F7FF81FF8AFE3569439E2FFCED" authorityName="Berlese" authorityYear="1913" box="[385,571,865,890]" class="Arachnida" family="Pachylaelapidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="51" pageNumber="52" phylum="Arthropoda" rank="family">Pachylaelapidae</taxonomicName>
. There are four slides and
<specimenCount id="CCA730FDFF81FF8AFCD369409FF3FCED" box="[871,999,865,890]" pageId="51" pageNumber="52" type="adult">seven adult</specimenCount>
specimens (
<specimenCount id="CCA730FDFF81FF8AFBC76943991BFCED" box="[1139,1295,865,890]" pageId="51" pageNumber="52" type="female">three females</specimenCount>
,
<specimenCount id="CCA730FDFF81FF8AFAAE69439980FCED" box="[1306,1428,865,890]" pageId="51" pageNumber="52" type="male">four males</specimenCount>
) of this species in the Berlese Acaroteca, viz. 182/9 (
<typeStatus id="051A45D6FF81FF8AFD6969A79F04FC09" box="[733,784,901,926]" pageId="51" pageNumber="52">type</typeStatus>
) and 182/68, all from the dung beetle
<emphasis id="E8D52766FF81FF8AFB7869A79927FC09" box="[1228,1331,901,926]" italics="true" pageId="51" pageNumber="52">Atheucus</emphasis>
sp. in La Plata,
<collectingCountry id="A2B6BBE4FF81FF8AFF50698B9D42FC55" box="[228,342,937,962]" name="Argentina" pageId="51" pageNumber="52">Argentina</collectingCountry>
, but only
<specimenCount id="CCA730FDFF81FF8AFE6969889E74FC55" box="[477,608,937,962]" pageId="51" pageNumber="52" type="female">one female</specimenCount>
specimen is in suitable condition for examination (on slide 182/6).
<taxonomicName id="1DA180F7FF81FF8AFF2369EF9DA0FC71" authorityName="Berlese" authorityYear="1916" box="[151,436,973,998]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="robustissima">
<emphasis id="E8D52766FF81FF8AFF2369EF9DA0FC71" box="[151,436,973,998]" italics="true" pageId="51" pageNumber="52">Pachylaella robustissima</emphasis>
</taxonomicName>
does not appear to be a member of the family
<taxonomicName id="1DA180F7FF81FF8AFC6E69EF9880FC71" authorityName="Berlese" authorityYear="1913" box="[986,1172,973,998]" class="Arachnida" family="Pachylaelapidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="51" pageNumber="52" phylum="Arthropoda" rank="family">Pachylaelapidae</taxonomicName>
. The sternal shield has three pairs of setae; there is a pair of small circular metasternal platelets separate from the sternal shield; the epigynal shield is tongue-shaped, not enlarged posteriorly, and bears only one pair of genital setae; the anal shield is subcircular, enlarged anteriorly, almost adjacent to the epigynal shield, and appears to have some ventral setae on its surface.
<taxonomicName id="1DA180F7FF81FF8AFE956E7F9DB3FBE1" authorityName="Berlese" authorityYear="1916" box="[289,423,1117,1142]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="51" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="E8D52766FF81FF8AFE956E7F9DB3FBE1" box="[289,423,1117,1142]" italics="true" pageId="51" pageNumber="52">Pachylaella</emphasis>
</taxonomicName>
could be classified in the family
<taxonomicName id="1DA180F7FF81FF8AFC8A6E7F9FF0FBE1" baseAuthorityName="Krantz" baseAuthorityYear="1965" box="[830,996,1117,1142]" class="Arachnida" family="Macrochelidae" kingdom="Animalia" order="Mesostigmata" pageId="51" pageNumber="52" phylum="Arthropoda" rank="family">Macrochelidae</taxonomicName>
rather than the
<taxonomicName id="1DA180F7FF81FF8AFB136E7F9975FBE1" authorityName="Berlese" authorityYear="1913" box="[1191,1377,1117,1142]" class="Arachnida" family="Pachylaelapidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="51" pageNumber="52" phylum="Arthropoda" rank="family">Pachylaelapidae</taxonomicName>
. The ventral idiosoma of
<taxonomicName id="1DA180F7FF81FF8AFE3C6EA09E2CFB0D" authorityName="Berlese" authorityYear="1916" box="[392,568,1153,1178]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="robustissima">
<emphasis id="E8D52766FF81FF8AFE3C6EA09D82FB0E" box="[392,406,1154,1177]" italics="true" pageId="51" pageNumber="52">P</emphasis>
.
<emphasis id="E8D52766FF81FF8AFE1E6EA39E2CFB0D" box="[426,568,1153,1178]" italics="true" pageId="51" pageNumber="52">robustissima</emphasis>
</taxonomicName>
resembles some species of
<taxonomicName id="1DA180F7FF81FF8AFC3F6EA398C2FB0D" authority="Berlese, 1903" authorityName="Berlese" authorityYear="1903" box="[907,1238,1153,1178]" class="Arachnida" family="Macrochelidae" genus="Holostaspella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="51" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="E8D52766FF81FF8AFC3F6EA39833FB0D" box="[907,1063,1153,1178]" italics="true" pageId="51" pageNumber="52">Holostaspella</emphasis>
<bibRefCitation id="BE308685FF81FF8AFB876EA398C2FB0D" author="Berlese, A." box="[1075,1238,1153,1178]" pageId="51" pageNumber="52" pagination="235 - 252" refId="ref38861" refString="Berlese, A. (1903 a) Acari nuovi. Manipulus I. Redia, 1, 235 - 252." type="journal article" year="1903">Berlese, 1903</bibRefCitation>
</taxonomicName>
. In addition, the peritremes of
<taxonomicName id="1DA180F7FF81FF8AFE816E849DF4FB29" authorityName="Berlese" authorityYear="1916" box="[309,480,1189,1214]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="robustissima">
<emphasis id="E8D52766FF81FF8AFE816E849DF4FB29" box="[309,480,1189,1214]" italics="true" pageId="51" pageNumber="52">P. robustissima</emphasis>
</taxonomicName>
are looped posteriorly, and join the stigma posteriorly, which is a specific character typical of most of the known macrochelid genera. Unfortunately, some important characters on the chelicerae were obscure and not visible in the available material.
</paragraph>
</subSubSection>
</treatment>
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