treatments-xml/data/14/03/87/140387BF487AF01F0D88FAF7FA88F911.xml

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<mods:title id="D94A6486A6E93FBEF7F3E87F27B25A15">Glossing over cryptic species: Descriptions of four new species of Glossodoris and three new species of Doriprismatica (Nudibranchia: Chromodorididae)</mods:title>
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<mods:namePart id="316F5A486B940FEB9D2206659D72B7B8">Matsuda, Shayle B.</mods:namePart>
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<heading id="C75D81C5487AF01A0D88FAF7FD19FADE" bold="true" box="[151,733,1292,1318]" fontSize="11" level="1" pageId="2" pageNumber="503" reason="1">
<emphasis id="AEDEEABB487AF01A0D88FAF7FD19FADE" bold="true" box="[151,733,1292,1318]" pageId="2" pageNumber="503">
<taxonomicName id="5BAA4D2A487AF01A0D88FAF7FDBCFADE" ID-CoL="3GDJR" authority="Matsuda &amp; Gosliner" authorityName="Matsuda &amp; Gosliner" authorityYear="2018" box="[151,632,1292,1318]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="2" pageNumber="503" phylum="Mollusca" rank="species" species="buko" status="sp. nov.">
<emphasis id="AEDEEABB487AF01A0D88FAF7FEABFADE" bold="true" box="[151,367,1292,1318]" italics="true" pageId="2" pageNumber="503">Glossodoris buko,</emphasis>
Matsuda &amp; Gosliner
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<taxonomicNameLabel id="B5ED57C0487AF01A0F60FAF7FD19FADE" box="[639,733,1292,1318]" pageId="2" pageNumber="503" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
<paragraph id="9C1536A9487AF01A0D88FAD5FDDCFABF" blockId="2.[151,733,1292,1351]" box="[151,536,1326,1351]" pageId="2" pageNumber="503">Figures (1A–C, 2A, 3A–E, 4C–D)</paragraph>
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<subSubSection id="D4B06522487AF01A0D88FA8DFDA6FA10" pageId="2" pageNumber="503" type="reference_group">
<paragraph id="9C1536A9487AF01A0D88FA8DFC33FA53" blockId="2.[151,1436,1398,1512]" pageId="2" pageNumber="503">
<taxonomicName id="5BAA4D2A487AF01A0D88FA8DFD42FA74" ID-CoL="3GDLJ" authority="(Ruppell &amp; Leuckart 1830)" authorityName="Ruppell &amp; Leuckart" authorityYear="1830" baseAuthorityName="Ruppell &amp; Leuckart" baseAuthorityYear="1830" box="[151,646,1398,1420]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="2" pageNumber="503" phylum="Mollusca" rank="species" species="pallida">
<emphasis id="AEDEEABB487AF01A0D88FA8DFEA6FA74" box="[151,354,1398,1420]" italics="true" pageId="2" pageNumber="503">Glossodoris pallida</emphasis>
(Rüppell &amp; Leuckart 1830)
</taxonomicName>
, misidentification, 
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: figs. 9c, 10e–f; 
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Gosliner 
<emphasis id="AEDEEABB487AF01A09EEFA83FAE2FA74" box="[1265,1318,1398,1421]" italics="true" pageId="2" pageNumber="503">et al.</emphasis>
2008
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: 240, third photo; 
<bibRefCitation id="F83B4B58487AF01A0C5BFA6DFDE9FA53" box="[324,557,1429,1451]" pageId="2" pageNumber="503" refId="ref17286" refString="Turner, L. M. &amp; Wilson, N. G. (2008) Polyphyly across oceans: a molecular phylogeny of the Chromodorididae (Mollusca, Nudibranchia). Zoologica Scripta, 37 (1), 23 - 42. https: // doi. org / 10.1111 / j. 1463 - 6409.2007.00310. x" type="journal article">Turner &amp; Wilson 2008</bibRefCitation>
; 
<bibRefCitation id="F83B4B58487AF01A0F28FA6EFCC7FA53" box="[567,771,1429,1451]" pageId="2" pageNumber="503" refId="ref16064" refString="Gosliner, T. M., Valdes, A. &amp; Behrens, D. W. (2015) Nudibranch and Sea Slug Identification-Indo-Pacific. New World Publications, Jacksonville, Florida, 236 pp." type="book">
Gosliner 
<emphasis id="AEDEEABB487AF01A0F8BFA6DFD03FA53" box="[660,711,1429,1451]" italics="true" pageId="2" pageNumber="503">et al.</emphasis>
2015
</bibRefCitation>
: 237, upper right photo.
</paragraph>
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<taxonomicName id="5BAA4D2A487AF01A0D88FA4FFDA1FA32" ID-CoL="3GDMY" authority="Ehrenberg 1831: 92" authorityName="Ehrenberg" authorityPageNumber="92" authorityYear="1831" box="[151,613,1460,1482]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="2" pageNumber="503" phylum="Mollusca" rank="species" species="xantholeuca">
<emphasis id="AEDEEABB487AF01A0D88FA4FFE57FA32" box="[151,403,1460,1482]" italics="true" pageId="2" pageNumber="503">Glossodoris xantholeuca</emphasis>
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<bibRefCitation id="F83B4B58487AF01A0C85FA4FFD84FA32" box="[410,576,1460,1482]" pageId="2" pageNumber="503" refId="ref15719" refString="Ehrenberg, C. G. (1831) Symbolae physicae seu icones et descriptiones animalium evertebratorum sepositis insectis quae ex itinere per Africam Borealem et Asiam Occidentalem, novae aut illustrate redierunt. Decas I Mollusca. Vol. 2. Testo. Mittler, Officina Academica, Berlin. [page not numbered]" type="book">Ehrenberg 1831</bibRefCitation>
: 92
</treatmentCitation>
</taxonomicName>
; 
<bibRefCitation id="F83B4B58487AF01A0F70FA4FFCC5FA32" box="[623,769,1460,1482]" pageId="2" pageNumber="503" refId="ref17040" refString="Rudman, W. B. (1984) The Chrornodorididae (Opisthobranchia: Mollusca) of the Indo-West Pacific: a review of the genera. Zoological Journal of the Linnean Society, 81, 115 - 273. https: // doi. org / 10.1111 / j. 1096 - 3642.1984. tb 01174. x" type="journal article">Rudman 1984</bibRefCitation>
.
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</paragraph>
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<taxonomicName id="5BAA4D2A487AF01A0D88FA29FED6FA10" ID-CoL="7P3T9" authorityName="Ehrenberg" authorityYear="1831" box="[151,274,1490,1512]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="2" pageNumber="503" phylum="Mollusca" rank="genus">
<emphasis id="AEDEEABB487AF01A0D88FA29FED6FA10" box="[151,274,1490,1512]" italics="true" pageId="2" pageNumber="503">Glossodoris</emphasis>
</taxonomicName>
sp. A 
<bibRefCitation id="F83B4B58487AF01A0C4CFA29FD9AFA10" box="[339,606,1490,1512]" pageId="2" pageNumber="503" refId="ref16546" refString="Matsuda, S. B. &amp; Gosliner, T. M. (2017) Molecular phylogeny of Glossodoris (Ehrenberg, 1831) nudibranchs and related genera reveals cryptic and pseudocryptic species complexes. Cladistics, 34 (1), 41 - 56. https: // doi. org / 10.1111 / cla. 12194" type="journal article">Matsuda &amp; Gosliner 2017</bibRefCitation>
.
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Material
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. 
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<emphasis id="AEDEEABB487AF01A0C4BF9EEFE7FF9D5" box="[340,443,1557,1581]" italics="true" pageId="2" pageNumber="503">Holotype</emphasis>
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: CASIZ-223284 (ex CASIZ- 
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B) 
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preserved, 
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, 
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, 
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Island
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, coll: 
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, 
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<date id="E8141069487AF01A0F9FF9C1FCA0F9AA" box="[640,868,1593,1618]" pageId="2" pageNumber="503" value="2012-11-10">10 November 2012</date>
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, orig. fixative 95% EtOH, GenBank: 
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(
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)
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.
</paragraph>
<paragraph id="9C1536A9487AF01A0DD8F979FD72F86E" blockId="2.[151,1437,1557,2014]" pageId="2" pageNumber="503">
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<emphasis id="AEDEEABB487AF01A0DD8F979FEFDF961" box="[199,313,1666,1689]" italics="true" pageId="2" pageNumber="503">Paratypes</emphasis>
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: CASIZ- 
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A, 
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,1 dissected, 
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, same collection data as 
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. CASIZ- 0 86381, 
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, 1 dissected, 
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, 
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, 
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, North of 
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, approx. 
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<collectingMunicipality id="7C71ACD3487AF01A0D88F932FE7DF91A" box="[151,441,1737,1762]" pageId="2" pageNumber="503">South of Cape Croisilles</collectingMunicipality>
, 
<location id="99756072487AF01A0CD7F932FDCDF91A" LSID="urn:lsid:plazi:treatment:140387BF487AF01F0D88FAF7FA88F911:99756072487AF01A0CD7F932FDCDF91A" box="[456,521,1737,1762]" country="Papua New Guinea" county="North Coast" municipality="South of Cape Croisilles" name="South" pageId="2" pageNumber="503" stateProvince="Madang">South</location>
side of 
<collectorName id="315F537F487AF01A0F70F932FD32F91A" box="[623,758,1737,1762]" pageId="2" pageNumber="503">The Quarry</collectorName>
, coll: 
<collectorName id="315F537F487AF01A0E5CF931FC22F91A" box="[835,998,1737,1762]" pageId="2" pageNumber="503">T.M. Gosliner</collectorName>
, 
<collectingDate id="F850E981487AF01A0EEAF931FB52F91A" box="[1013,1174,1738,1762]" pageId="2" pageNumber="503" value="1992-06-13">
<date id="E8141069487AF01A0EEAF931FB52F91A" box="[1013,1174,1738,1762]" pageId="2" pageNumber="503" value="1992-06-13">13 June 1992</date>
</collectingDate>
, orig
</materialsCitation>
. 
<materialsCitation id="2CC23CF4487AF01A09FBF932FD76F86E" ID-GBIF-Occurrence="1913825175" collectingDate="2004-07-02" collectingDateMax="2008-03-16" collectingDateMin="2004-07-02" collectorName="Bouin's &amp; Panglao &amp; Pontog Lagoon I &amp; T. M. Gosliner &amp; Y. Camacho &amp; J. Templado &amp; M. Malaquias &amp; M. Poddubetskaia &amp; Layaglayag &amp; T. Gosliner &amp; A. Valdes &amp; M. Pola &amp; L. Witzel &amp; B. Moore &amp; A. Alejandrino" country="Philippines" county="Bohol Island" location="Caban Island" municipality="Luzon Island" pageId="2" pageNumber="503" specimenCount="1" stateProvince="Batangas Province" typeStatus="paratype">
fixative 
<collectorName id="315F537F487AF01A085AF932FA58F91A" box="[1349,1436,1737,1762]" pageId="2" pageNumber="503">Bouin’s</collectorName>
solution. CASIZ-181594, one specimen 
<quantity id="5B529B4C487AF01A0F46F916FD5DF8FD" box="[601,665,1773,1798]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="503" unit="mm" value="6.0">6 mm</quantity>
preserved, 
<collectingCountry id="E4BD7639487AF01A0E06F916FC5EF8FE" box="[793,922,1773,1798]" name="Philippines" pageId="2" pageNumber="503">Philippines</collectingCountry>
, 
<collectingCounty id="75744E25487AF01A0EBBF916FBF2F8FE" box="[932,1078,1773,1798]" pageId="2" pageNumber="503">Bohol Island</collectingCounty>
, 
<collectorName id="315F537F487AF01A095EF916FB5AF8FE" box="[1089,1182,1773,1798]" pageId="2" pageNumber="503">Panglao</collectorName>
, 
<collectorName id="315F537F487AF01A09B6F915FAA1F8FD" box="[1193,1381,1774,1798]" pageId="2" pageNumber="503">Pontog Lagoon I</collectorName>
, reef wall with small caves, coll: 
<collectorName id="315F537F487AF01A0CD2F8E9FDA8F8D2" box="[461,620,1809,1834]" pageId="2" pageNumber="503">T.M. Gosliner</collectorName>
, 
<collectorName id="315F537F487AF01A0F68F8E9FCC6F8D2" box="[631,770,1809,1834]" pageId="2" pageNumber="503">Y. Camacho</collectorName>
, 
<collectorName id="315F537F487AF01A0E13F8E9FC53F8D2" box="[780,919,1809,1834]" pageId="2" pageNumber="503">J. Templado</collectorName>
, 
<collectorName id="315F537F487AF01A0EBDF8E9FBFBF8D2" box="[930,1087,1809,1834]" pageId="2" pageNumber="503">M. Malaquias</collectorName>
, 
<collectorName id="315F537F487AF01A0955F8E9FAD3F8D2" box="[1098,1303,1809,1834]" pageId="2" pageNumber="503">M. Poddubetskaia</collectorName>
, 
<collectingDate id="F850E981487AF01A083DF8E9FA5CF8D2" box="[1314,1432,1809,1834]" pageId="2" pageNumber="503" value="2004-07-02">
<date id="E8141069487AF01A083DF8E9FA5CF8D2" box="[1314,1432,1809,1834]" pageId="2" pageNumber="503" value="2004-07-02">2 Jul 2004</date>
</collectingDate>
, Panglao Expedition 2004, 17–25 meters, orig. fixative 95% EtOH. CASIZ-177264, one specimen, 
<collectingCountry id="E4BD7639487AF01A0808F8CEFA5CF8B6" box="[1303,1432,1845,1870]" name="Philippines" pageId="2" pageNumber="503">Philippines</collectingCountry>
, 
<collectingMunicipality id="7C71ACD3487AF01A0D88F8A1FEF4F88A" box="[151,304,1881,1906]" pageId="2" pageNumber="503">Luzon Island</collectingMunicipality>
, 
<collectingRegion id="5E6EF84B487AF01A0C21F8A1FDD2F88A" box="[318,534,1881,1906]" country="Philippines" name="Batangas" pageId="2" pageNumber="503">Batangas Province</collectingRegion>
, Tingloy, 
<location id="99756072487AF01A0F93F8A2FCE1F88A" LSID="urn:lsid:plazi:treatment:140387BF487AF01F0D88FAF7FA88F911:99756072487AF01A0F93F8A2FCE1F88A" box="[652,805,1881,1906]" country="Philippines" county="Bohol Island" municipality="Luzon Island" name="Caban Island" pageId="2" pageNumber="503" stateProvince="Batangas Province">Caban Island</location>
, 
<collectorName id="315F537F487AF01A0E2BF8A2FC7DF88A" box="[820,953,1881,1906]" pageId="2" pageNumber="503">Layaglayag</collectorName>
, coll: 
<collectorName id="315F537F487AF01A091BF8A1FB42F88A" box="[1028,1158,1881,1906]" pageId="2" pageNumber="503">T. Gosliner</collectorName>
, 
<collectorName id="315F537F487AF01A098AF8A1FACCF88A" box="[1173,1288,1881,1906]" pageId="2" pageNumber="503">A. Valdés</collectorName>
, 
<collectorName id="315F537F487AF01A0808F8A1FAB1F88A" box="[1303,1397,1881,1906]" pageId="2" pageNumber="503">M. Pola</collectorName>
, 
<collectorName id="315F537F487AF01A089BF8A1FF20F86E" pageId="2" pageNumber="503">L. Witzel</collectorName>
, 
<collectorName id="315F537F487AF01A0DF1F885FE98F86D" box="[238,348,1918,1941]" pageId="2" pageNumber="503">B. Moore</collectorName>
, 
<collectorName id="315F537F487AF01A0C77F885FDD7F86E" box="[360,531,1917,1942]" pageId="2" pageNumber="503">A. Alejandrino</collectorName>
, 
<collectingDate id="F850E981487AF01A0F01F886FD76F86E" box="[542,690,1917,1942]" pageId="2" pageNumber="503" value="2008-03-16">
<date id="E8141069487AF01A0F01F886FD76F86E" box="[542,690,1917,1942]" pageId="2" pageNumber="503" value="2008-03-16">16 Mar 2008</date>
</collectingDate>
</materialsCitation>
.
</paragraph>
<paragraph id="9C1536A9487AF01B0DD8F859FC5EFDE0" blockId="2.[151,1437,1557,2014]" lastBlockId="3.[151,1437,151,536]" lastPageId="3" lastPageNumber="504" pageId="2" pageNumber="503">
<materialsCitation id="2CC23CF4487AF01B0DD8F859FE06FF2D" ID-GBIF-Occurrence="1913825177" collectingDate="2005-10-20" collectorName="Comparative &amp; Nosy Valiha &amp; T. M. Gosliner" country="Madagascar" elevation="13" lastPageId="3" lastPageNumber="504" location="Madagascar" pageId="2" pageNumber="503" specimenCount="1">
<emphasis id="AEDEEABB487AF01A0DD8F859FD01F841" box="[199,709,1953,1977]" italics="true" pageId="2" pageNumber="503">
<collectorName id="315F537F487AF01A0DD8F859FE9DF841" box="[199,345,1954,1977]" pageId="2" pageNumber="503">Comparative</collectorName>
material of 
<taxonomicName id="5BAA4D2A487AF01A0CF9F85AFD01F841" baseAuthorityName="Ruppell &amp; Leuckart" baseAuthorityYear="1830" box="[486,709,1953,1977]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="2" pageNumber="503" phylum="Mollusca" rank="species" species="pallida">Glossodoris pallida</taxonomicName>
</emphasis>
(
<figureCitation id="04912A2C487AF01A0FCAF85AFCADF842" box="[725,873,1953,1978]" captionStart="FIGURE 1" captionStartId="3.[151,250,1750,1772]" captionTargetBox="[248,1338,592,1723]" captionTargetId="figure@3.[246,1340,591,1726]" captionTargetPageId="3" captionText="FIGURE 1. A–C, Glossodoris buko sp. nov. A. CASIZ-191102 (i. large, ii. small) Papua New Guinea. B. CASIZ-181594 Philippines. C. CASIZ-086381 (A, B) Papua New Guinea. D–F, Glossodoris pallida: D. CASIZ-194338 Madagascar. E. CASIZ-175548 Madagascar. F. CASIZ-175554 Madagascar. Photos: T.M. Gosliner." httpUri="https://zenodo.org/record/1437226/files/figure.png" pageId="2" pageNumber="503">Figs. 1D–1E</figureCitation>
): CASIZ-173393, one specimen, dissected, 
<quantity id="5B529B4C487AF01A0845F859FA58F841" box="[1370,1436,1954,1978]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="503" unit="mm" value="9.0">9 mm</quantity>
preserved, 
<collectingCountry id="E4BD7639487AF01A0C0DF83EFE5FF826" box="[274,411,1989,2014]" name="Madagascar" pageId="2" pageNumber="503">Madagascar</collectingCountry>
, Iles Radama, Nosy Valiha, W of 
<collectorName id="315F537F487AF01A0E04F83DFC63F826" box="[795,935,1989,2014]" pageId="2" pageNumber="503">Nosy Valiha</collectorName>
, coll: 
<collectorName id="315F537F487AF01A0EF2F83DFB48F826" box="[1005,1164,1989,2014]" pageId="2" pageNumber="503">T.M. Gosliner</collectorName>
, 
<collectingDate id="F850E981487AF01A0987F83DFAE1F826" box="[1176,1317,1989,2014]" pageId="2" pageNumber="503" value="2005-10-20">
<date id="E8141069487AF01A0987F83DFAE1F826" box="[1176,1317,1989,2014]" pageId="2" pageNumber="503" value="2005-10-20">20 Oct 2005</date>
</collectingDate>
, 12– 
<elevation id="1787D19A487AF01A0841F83DFA53F825" box="[1374,1431,1990,2014]" metricMagnitude="1" metricUnit="m" metricValue="1.3" pageId="2" pageNumber="503" unit="m" value="13.0">
<quantity id="5B529B4C487AF01A0841F83DFA53F825" box="[1374,1431,1990,2014]" metricMagnitude="1" metricUnit="m" metricValue="1.3" pageId="2" pageNumber="503" unit="m" value="13.0">13 m</quantity>
</elevation>
, orig. fixative Bouin’s solution. CASIZ-173395, one specimen, dissected, 
<quantity id="5B529B4C487BF01B091DFF63FB8EFF57" box="[1026,1098,152,176]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="3" pageNumber="504" unit="mm" value="9.0">9 mm</quantity>
preserved, 
<collectingCountry id="E4BD7639487BF01B09C6FF6CFAA6FF48" box="[1241,1378,151,176]" name="Madagascar" pageId="3" pageNumber="504">Madagascar</collectingCountry>
, Iles Radama, Nosy Kalakajoro
</materialsCitation>
, 
<materialsCitation id="2CC23CF4487BF01B0CD3FF46FACBFF00" ID-GBIF-Occurrence="1913825172" collectingDate="2005-10-19" collectorName="T. M. Gosliner" country="Mozambique" elevation="15" location="West" pageId="3" pageNumber="504" specimenCount="1" stateProvince="West">
<collectingRegion id="5E6EF84B487BF01B0CD3FF46FDC7FF2C" box="[460,515,189,212]" country="Tanzania" name="Mjini Magharibi" pageId="3" pageNumber="504">West</collectingRegion>
of Nosy Kalakajoro and Nosy Beratia, coll: 
<collectorName id="315F537F487BF01B0EECFF46FB55FF2D" box="[1011,1169,188,213]" pageId="3" pageNumber="504">T.M. Gosliner</collectorName>
, 
<collectingDate id="F850E981487BF01B0983FF47FAECFF2D" box="[1180,1320,188,213]" pageId="3" pageNumber="504" value="2005-10-19">
<date id="E8141069487BF01B0983FF47FAECFF2D" box="[1180,1320,188,213]" pageId="3" pageNumber="504" value="2005-10-19">19 Oct 2005</date>
</collectingDate>
, 13– 
<elevation id="1787D19A487BF01B0841FF47FA53FF2C" box="[1374,1431,188,213]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="504" unit="m" value="15.0">
<quantity id="5B529B4C487BF01B0841FF47FA53FF2C" box="[1374,1431,188,213]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="504" unit="m" value="15.0">15 m</quantity>
</elevation>
, orig. fixative Bouin’s solution. CASIZ-176997, one specimen, 
<quantity id="5B529B4C487BF01B0E9EFF24FC2FFF0F" box="[897,1003,223,248]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="3" pageNumber="504" unit="mm" value="14.5">14.5 mm</quantity>
preserved, 
<collectingCountry id="E4BD7639487BF01B0967FF24FACBFF00" box="[1144,1295,223,248]" name="Mozambique" pageId="3" pageNumber="504">Mozambique</collectingCountry>
</materialsCitation>
, 
<materialsCitation id="2CC23CF4487BF01B083FFF24FA5CFE70" ID-GBIF-Occurrence="1913825176" collectingDate="2005-10-13" collectingDateMax="2008-02-06" collectingDateMin="2005-10-13" country="Madagascar" location="Radama Islands" pageId="3" pageNumber="504" specimenCount="1" stateProvince="Inhambane Province">
<collectingRegion id="5E6EF84B487BF01B083FFF24FF39FEE5" country="Mozambique" name="Inhambane" pageId="3" pageNumber="504">Inhambane Province</collectingRegion>
, Jangamo, Pandane Beach, coll: M. Pola and J. Reis, 
<collectingDate id="F850E981487BF01B0E79FEFFFC2EFEE4" box="[870,1002,260,285]" pageId="3" pageNumber="504" value="2008-02-06">
<date id="E8141069487BF01B0E79FEFFFC2EFEE4" box="[870,1002,260,285]" pageId="3" pageNumber="504" value="2008-02-06">6 Feb 2008</date>
</collectingDate>
, 1.5 meters, orig. fixative 95% EtOH. CASIZ-175548, one specimen 
<quantity id="5B529B4C487BF01B0F1BFED3FD8EFEC7" box="[516,586,296,320]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="504" unit="mm" value="4.0">4 mm</quantity>
preserved (large portion missing from tissue sample), 
<collectingCountry id="E4BD7639487BF01B09C5FEDCFAA7FEB8" box="[1242,1379,295,320]" name="Madagascar" pageId="3" pageNumber="504">Madagascar</collectingCountry>
, Iles Radama, Nosy Kalakajoro, coll: S. Fahey and T. M. Gosliner, 
<collectingDate id="F850E981487BF01B0E56FEB7FC10FE9D" box="[841,980,332,357]" pageId="3" pageNumber="504" value="2005-10-13">
<date id="E8141069487BF01B0E56FEB7FC10FE9D" box="[841,980,332,357]" pageId="3" pageNumber="504" value="2005-10-13">13 Oct 2005</date>
</collectingDate>
, CAS-WCS 
<location id="99756072487BF01B097CFEB7FAD0FE9D" LSID="urn:lsid:plazi:treatment:140387BF487AF01F0D88FAF7FA88F911:99756072487BF01B097CFEB7FAD0FE9D" box="[1123,1300,332,357]" country="Madagascar" name="Radama Islands" pageId="3" pageNumber="504" stateProvince="Inhambane Province">Radama Islands</location>
Expedition, 15-20 meters, orig. fixative 95% EtOH. CASIZ-194338, one specimen, dissected, 
<quantity id="5B529B4C487BF01B0959FE94FB4DFE7F" box="[1094,1161,367,392]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="504" unit="mm" value="6.0">6 mm</quantity>
preserved, 
<collectingCountry id="E4BD7639487BF01B0810FE94FA5CFE70" box="[1295,1432,367,392]" name="Madagascar" pageId="3" pageNumber="504">Madagascar</collectingCountry>
</materialsCitation>
, 
<materialsCitation id="2CC23CF4487BF01B0D88FE6FFBD3FE55" ID-GBIF-Occurrence="1913825170" box="[151,1047,404,429]" country="Madagascar" location="South" pageId="3" pageNumber="504" specimenCount="1" stateProvince="South">
<collectingRegion id="5E6EF84B487BF01B0D88FE6FFF1CFE55" box="[151,216,404,429]" country="Tanzania" name="Kusini Unguja" pageId="3" pageNumber="504">South</collectingRegion>
<collectingCountry id="E4BD7639487BF01B0DC0FE6FFEACFE55" box="[223,360,404,429]" name="Madagascar" pageId="3" pageNumber="504">Madagascar</collectingCountry>
, “Pointe Evatra, crique fond rocheux et gazon d’algues”, coll
</materialsCitation>
. 
<materialsCitation id="2CC23CF4487BF01B093EFE6FFEC2FE28" ID-GBIF-Occurrence="1913825174" collectingDate="2006-04-30" country="Madagascar" location="South" pageId="3" pageNumber="504" specimenCount="1" stateProvince="South">
<collectingRegion id="5E6EF84B487BF01B093EFE6FFBA6FE55" box="[1057,1122,404,429]" country="Tanzania" name="Kusini Unguja" pageId="3" pageNumber="504">South</collectingRegion>
<collectingCountry id="E4BD7639487BF01B0976FE6FFB34FE55" box="[1129,1264,404,429]" name="Madagascar" pageId="3" pageNumber="504">Madagascar</collectingCountry>
Expedition, 
<collectingDate id="F850E981487BF01B089FFE6FFEC2FE28" pageId="3" pageNumber="504" value="2006-04-30">
<date id="E8141069487BF01B089FFE6FFEC2FE28" pageId="3" pageNumber="504" value="2006-04-30">30 Apr 2006</date>
</collectingDate>
</materialsCitation>
– 
<materialsCitation id="2CC23CF4487BF01B0C0AFE4CFC5EFDE0" ID-GBIF-Occurrence="1913825173" collectingDate="2010-05-03" collectingDateMax="2010-05-08" collectingDateMin="2010-05-03" collectorName="S. Fahey &amp; T. M. Gosliner" country="Madagascar" location="Radama Islands" pageId="3" pageNumber="504" specimenCount="1">
<collectingDate id="F850E981487BF01B0C0AFE4CFE0EFE28" box="[277,458,439,464]" pageId="3" pageNumber="504" value="2010-05-03" valueMax="2010-05-08" valueMin="2010-05-03">
<date id="E8141069487BF01B0C0AFE4CFE0EFE28" box="[277,458,439,464]" pageId="3" pageNumber="504" value="2010-05-03" valueMax="2010-05-08" valueMin="2010-05-03">May 2010, 3–8</date>
</collectingDate>
meters, orig. fixative 95% EtOH. CASIZ-175554, one specimen, dissected, 
<quantity id="5B529B4C487BF01B0849FE43FA59FE37" box="[1366,1437,440,464]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="504" unit="mm" value="2.0">2 mm</quantity>
preserved, 
<collectingCountry id="E4BD7639487BF01B0C0CFE27FE58FE0D" box="[275,412,476,501]" name="Madagascar" pageId="3" pageNumber="504">Madagascar</collectingCountry>
, Iles Radama, Nosy Faly, NW of Nosy Faly, coll: 
<collectorName id="315F537F487BF01B0EC7FE27FBFFFE0D" box="[984,1083,476,501]" pageId="3" pageNumber="504">S. Fahey</collectorName>
and 
<collectorName id="315F537F487BF01B096CFE26FAD7FE0D" box="[1139,1299,476,501]" pageId="3" pageNumber="504">T.M. Gosliner</collectorName>
, CAS-WCS 
<location id="99756072487BF01B0D88FE04FE8DFDE0" LSID="urn:lsid:plazi:treatment:140387BF487AF01F0D88FAF7FA88F911:99756072487BF01B0D88FE04FE8DFDE0" box="[151,329,511,536]" country="Madagascar" name="Radama Islands" pageId="3" pageNumber="504">Radama Islands</location>
Expedition, 13–16 meters, orig. fixative 95% EtOH.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="C8D56621487BF01B0D88F92DFC2DF8D2" httpUri="https://zenodo.org/record/1437226/files/figure.png" pageId="3" pageNumber="504" startId="3.[151,250,1750,1772]" targetBox="[248,1338,592,1723]" targetPageId="3">
<paragraph id="9C1536A9487BF01B0D88F92DFC2DF8D2" blockId="3.[151,1436,1750,1834]" pageId="3" pageNumber="504">
<emphasis id="AEDEEABB487BF01B0D88F92DFED1F914" bold="true" box="[151,277,1750,1772]" pageId="3" pageNumber="504">FIGURE 1.</emphasis>
A–C, 
<taxonomicName id="5BAA4D2A487BF01B0C42F92DFDD7F914" authority="Matsuda &amp; Gosliner, 2018" authorityName="Matsuda &amp; Gosliner" authorityYear="2018" box="[349,531,1750,1772]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="3" pageNumber="504" phylum="Mollusca" rank="species" species="buko" status="sp. nov.">
<emphasis id="AEDEEABB487BF01B0C42F92DFDD7F914" box="[349,531,1750,1772]" italics="true" pageId="3" pageNumber="504">Glossodoris buko</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B5ED57C0487BF01B0F04F92CFDABF914" box="[539,623,1751,1772]" pageId="3" pageNumber="504" rank="species">
<emphasis id="AEDEEABB487BF01B0F04F92CFDABF914" bold="true" box="[539,623,1751,1772]" pageId="3" pageNumber="504">sp. nov.</emphasis>
</taxonomicNameLabel>
A. CASIZ-191102 (i. large, ii. small) Papua New Guinea. B. CASIZ-181594 Philippines. C. CASIZ-086381 (A, B) Papua New Guinea. D–F, 
<taxonomicName id="5BAA4D2A487BF01B0E4FF90EFBDAF8F3" baseAuthorityName="Ruppell &amp; Leuckart" baseAuthorityYear="1830" box="[848,1054,1781,1803]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="3" pageNumber="504" phylum="Mollusca" rank="species" species="pallida">
<emphasis id="AEDEEABB487BF01B0E4FF90EFBDAF8F3" box="[848,1054,1781,1803]" italics="true" pageId="3" pageNumber="504">Glossodoris pallida</emphasis>
</taxonomicName>
: D. CASIZ-194338 Madagascar. E. CASIZ-175548 Madagascar. F. CASIZ-175554 Madagascar. Photos: T.M. Gosliner.
</paragraph>
</caption>
<subSubSection id="D4B06522487BF01B0DD8F8ADFC8CF823" pageId="3" pageNumber="504" type="etymology">
<paragraph id="9C1536A9487BF01B0DD8F8ADFB67F86B" blockId="3.[151,1436,1878,2011]" pageId="3" pageNumber="504">
<emphasis id="AEDEEABB487BF01B0DD8F8ADFE82F897" bold="true" box="[199,326,1878,1903]" pageId="3" pageNumber="504">Etymology</emphasis>
. The name 
<taxonomicName id="5BAA4D2A487BF01B0CD2F8ADFD56F896" authorityName="Matsuda &amp; Gosliner" authorityYear="2018" box="[461,658,1878,1902]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="3" pageNumber="504" phylum="Mollusca" rank="species" species="buko">
<emphasis id="AEDEEABB487BF01B0CD2F8ADFD56F896" box="[461,658,1878,1902]" italics="true" pageId="3" pageNumber="504">Glossodoris buko</emphasis>
</taxonomicName>
comes from 
<taxonomicName id="5BAA4D2A487BF01B0E36F8ADFCA5F897" authorityName="Matsuda &amp; Gosliner" authorityYear="2018" box="[809,865,1878,1903]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="3" pageNumber="504" phylum="Mollusca" rank="species" species="buko">buko</taxonomicName>
(young coconut) owing to the resemblance of this species to the cream-colored coconut meat from the 
<collectingCountry id="E4BD7639487BF01B0FFEF881FCA6F86B" box="[737,866,1914,1939]" name="Philippines" pageId="3" pageNumber="504">Philippines</collectingCountry>
, where this species is found.
</paragraph>
<paragraph id="9C1536A9487BF01B0DD8F865FC8CF823" blockId="3.[151,1436,1878,2011]" pageId="3" pageNumber="504">
<emphasis id="AEDEEABB487BF01B0DD8F865FDC7F84F" bold="true" box="[199,515,1950,1975]" pageId="3" pageNumber="504">Geographical Distribution</emphasis>
. Specimens identified by 
<bibRefCitation id="F83B4B58487BF01B0E33F865FBA2F84F" box="[812,1126,1950,1975]" pageId="3" pageNumber="504" refId="ref16546" refString="Matsuda, S. B. &amp; Gosliner, T. M. (2017) Molecular phylogeny of Glossodoris (Ehrenberg, 1831) nudibranchs and related genera reveals cryptic and pseudocryptic species complexes. Cladistics, 34 (1), 41 - 56. https: // doi. org / 10.1111 / cla. 12194" type="journal article">Matsuda &amp; Gosliner (2017)</bibRefCitation>
range from the 
<collectingCountry id="E4BD7639487BF01B0801F865FA58F84F" box="[1310,1436,1950,1975]" name="Philippines" pageId="3" pageNumber="504">Philippines</collectingCountry>
to 
<collectingCountry id="E4BD7639487BF01B0DABF838FE48F823" box="[180,396,1986,2011]" name="Papua New Guinea" pageId="3" pageNumber="504">Papua New Guinea</collectingCountry>
and 
<collectingCountry id="E4BD7639487BF01B0CDDF839FDECF823" box="[450,552,1986,2011]" name="Australia" pageId="3" pageNumber="504">Australia</collectingCountry>
(
<bibRefCitation id="F83B4B58487BF01B0F28F838FCF8F823" box="[567,828,1986,2011]" pageId="3" pageNumber="504" refId="ref17286" refString="Turner, L. M. &amp; Wilson, N. G. (2008) Polyphyly across oceans: a molecular phylogeny of the Chromodorididae (Mollusca, Nudibranchia). Zoologica Scripta, 37 (1), 23 - 42. https: // doi. org / 10.1111 / j. 1463 - 6409.2007.00310. x" type="journal article">Turner &amp; Wilson 2008</bibRefCitation>
).
</paragraph>
</subSubSection>
<caption id="C8D56621487CF01C0D88F9D9FCDBF92A" httpUri="https://zenodo.org/record/1437228/files/figure.png" pageId="4" pageNumber="505" startId="4.[151,250,1570,1592]" targetBox="[334,1263,243,1490]" targetPageId="4">
<paragraph id="9C1536A9487CF01C0D88F9D9FCDBF92A" blockId="4.[151,1436,1570,1746]" pageId="4" pageNumber="505">
<emphasis id="AEDEEABB487CF01C0D88F9D9FED0F9CF" bold="true" box="[151,276,1570,1592]" pageId="4" pageNumber="505">FIGURE 2.</emphasis>
Mantle margins. A. 
<taxonomicName id="5BAA4D2A487CF01C0CF3F9D9FD64F9C0" authority="Matsuda &amp; Gosliner, 2018" authorityName="Matsuda &amp; Gosliner" authorityYear="2018" box="[492,672,1570,1592]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="4" pageNumber="505" phylum="Mollusca" rank="species" species="buko" status="sp. nov.">
<emphasis id="AEDEEABB487CF01C0CF3F9D9FD64F9C0" box="[492,672,1570,1592]" italics="true" pageId="4" pageNumber="505">Glossodoris buko</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B5ED57C0487CF01C0FB7F9D8FD3DF9C0" box="[680,761,1571,1592]" pageId="4" pageNumber="505" rank="species">
<emphasis id="AEDEEABB487CF01C0FB7F9D8FD3DF9C0" bold="true" box="[680,761,1571,1592]" pageId="4" pageNumber="505">sp. nov.</emphasis>
</taxonomicNameLabel>
, CASIZ-222107. B. 
<taxonomicName id="5BAA4D2A487CF01C0ECBF9D9FAFAF9C0" authority="(Cuvier, 1804)" authorityName="Cuvier" authorityYear="1804" baseAuthorityName="Cuvier" baseAuthorityYear="1804" box="[980,1342,1570,1592]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="4" pageNumber="505" phylum="Mollusca" rank="species" species="pallida">
<emphasis id="AEDEEABB487CF01C0ECBF9D9FB5AF9C0" box="[980,1182,1570,1592]" italics="true" pageId="4" pageNumber="505">Glossodoris pallida</emphasis>
(Cuvier, 1804)
</taxonomicName>
, CASIZ- 175554. C. 
<taxonomicName id="5BAA4D2A487CF01C0C0EF9BAFE34F9AF" authority="Matsuda &amp; Gosliner, 2018" authorityName="Matsuda &amp; Gosliner" authorityYear="2018" box="[273,496,1601,1623]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="4" pageNumber="505" phylum="Mollusca" rank="species" species="bowanga" status="sp. nov.">
<emphasis id="AEDEEABB487CF01C0C0EF9BAFE34F9AF" box="[273,496,1601,1623]" italics="true" pageId="4" pageNumber="505">Glossodoris bowanga</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B5ED57C0487CF01C0CE7F9BAFD8EF9AE" box="[504,586,1601,1622]" pageId="4" pageNumber="505" rank="species">
<emphasis id="AEDEEABB487CF01C0CE7F9BAFD8EF9AE" bold="true" box="[504,586,1601,1622]" pageId="4" pageNumber="505">sp. nov.</emphasis>
</taxonomicNameLabel>
, CASIZ-194204, photo by 
<taxonomicName id="5BAA4D2A487CF01C0E76F9BAFBB1F9AF" authority="Poddubetskaia. D." authorityName="Matsuda &amp; Gosliner" authorityYear="2018" box="[873,1141,1601,1623]" class="Magnoliopsida" family="Fabaceae" genus="Marina" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fabales" pageId="4" pageNumber="505" phylum="Tracheophyta" rank="genus" status="sp. nov.">Marina Poddubetskaia. D.</taxonomicName>
<taxonomicName id="5BAA4D2A487CF01C0962F9BAFAB1F9AF" authority="Matsuda &amp; Gosliner, 2018" authorityName="Matsuda &amp; Gosliner" authorityYear="2018" box="[1149,1397,1601,1623]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="4" pageNumber="505" phylum="Mollusca" rank="species" species="andersonae" status="sp. nov.">
<emphasis id="AEDEEABB487CF01C0962F9BAFAB1F9AF" box="[1149,1397,1601,1623]" italics="true" pageId="4" pageNumber="505">Glossodoris andersonae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B5ED57C0487CF01C0862F9BAFF07F98D" pageId="4" pageNumber="505" rank="species">
<emphasis id="AEDEEABB487CF01C0862F9BAFF07F98D" bold="true" pageId="4" pageNumber="505">sp. nov.</emphasis>
</taxonomicNameLabel>
, CASIZ-192288. E. 
<taxonomicName id="5BAA4D2A487CF01C0C89F99BFD90F98E" authority="Matsuda &amp; Gosliner, 2018" authorityName="Matsuda &amp; Gosliner" authorityYear="2018" box="[406,596,1632,1654]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="4" pageNumber="505" phylum="Mollusca" rank="species" species="acosti" status="sp. nov.">
<emphasis id="AEDEEABB487CF01C0C89F99BFD90F98E" box="[406,596,1632,1654]" italics="true" pageId="4" pageNumber="505">Glossodoris acosti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B5ED57C0487CF01C0F45F99BFD6EF98D" box="[602,682,1632,1653]" pageId="4" pageNumber="505" rank="species">
<emphasis id="AEDEEABB487CF01C0F45F99BFD6EF98D" bold="true" box="[602,682,1632,1653]" pageId="4" pageNumber="505">sp. nov.</emphasis>
</taxonomicNameLabel>
, CASIZ 191352, photo by Vanessa Knutson. F. 
<taxonomicName id="5BAA4D2A487CF01C098CF99BFACAF98E" authorityName="Ehrenberg" authorityYear="1831" box="[1171,1294,1632,1654]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="4" pageNumber="505" phylum="Mollusca" rank="genus">
<emphasis id="AEDEEABB487CF01C098CF99BFACAF98E" box="[1171,1294,1632,1654]" italics="true" pageId="4" pageNumber="505">Glossodoris</emphasis>
</taxonomicName>
sp. 
<taxonomicName id="5BAA4D2A487CF01C0828F99BFA52F98E" authorityName="Bergh" authorityYear="1888" box="[1335,1430,1632,1654]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="4" pageNumber="505" phylum="Mollusca" rank="species" species="cincta">
cf. 
<emphasis id="AEDEEABB487CF01C0847F99AFA52F98E" box="[1368,1430,1633,1654]" italics="true" pageId="4" pageNumber="505">cincta</emphasis>
</taxonomicName>
, CASIZ-199204. G. 
<taxonomicName id="5BAA4D2A487CF01C0C42F985FD65F96C" authority="Bergh, 1888" authorityName="Bergh" authorityYear="1888" box="[349,673,1662,1685]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="4" pageNumber="505" phylum="Mollusca" rank="species" species="cincta">
<emphasis id="AEDEEABB487CF01C0C42F985FDD8F96D" box="[349,540,1662,1685]" italics="true" pageId="4" pageNumber="505">Glossodoris cincta</emphasis>
Bergh, 1888
</taxonomicName>
, from Reunion, photo only, photo by Philibert Bidgrain. H. 
<taxonomicName id="5BAA4D2A487CF01C081FF97BFF0FF94B" authority="Matsuda &amp; Gosliner, 2018" authorityName="Matsuda &amp; Gosliner" authorityYear="2018" class="Gastropoda" family="Chromodorididae" genus="Doriprismatica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="4" pageNumber="505" phylum="Mollusca" rank="species" species="balut" status="sp. nov.">
<emphasis id="AEDEEABB487CF01C081FF97BFF0FF94B" italics="true" pageId="4" pageNumber="505">Doriprismatica balut</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B5ED57C0487CF01C0DCCF965FEE1F94B" box="[211,293,1694,1715]" pageId="4" pageNumber="505" rank="species">
<emphasis id="AEDEEABB487CF01C0DCCF965FEE1F94B" bold="true" box="[211,293,1694,1715]" pageId="4" pageNumber="505">sp. nov.</emphasis>
</taxonomicNameLabel>
, CASIZ-182787. I. 
<taxonomicName id="5BAA4D2A487CF01C0CEAF965FD0DF94B" authority="Matsuda &amp; Gosliner, 2018" authorityName="Matsuda &amp; Gosliner" authorityYear="2018" box="[501,713,1694,1715]" class="Gastropoda" family="Chromodorididae" genus="Doriprismatica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="4" pageNumber="505" phylum="Mollusca" rank="species" species="rossi" status="sp. nov.">
<emphasis id="AEDEEABB487CF01C0CEAF965FD0DF94B" box="[501,713,1694,1715]" italics="true" pageId="4" pageNumber="505">Doriprismatica rossi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B5ED57C0487CF01C0FCDF965FCE7F94B" box="[722,803,1694,1715]" pageId="4" pageNumber="505" rank="species">
<emphasis id="AEDEEABB487CF01C0FCDF965FCE7F94B" bold="true" box="[722,803,1694,1715]" pageId="4" pageNumber="505">sp. nov.</emphasis>
</taxonomicNameLabel>
CASIZ-192281. J. 
<taxonomicName id="5BAA4D2A487CF01C0EEFF965FB2CF94B" authority="Matsuda &amp; Gosliner, 2018" authorityName="Matsuda &amp; Gosliner" authorityYear="2018" box="[1008,1256,1694,1715]" class="Gastropoda" family="Chromodorididae" genus="Doriprismatica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="4" pageNumber="505" phylum="Mollusca" rank="species" species="marinae" status="sp. nov.">
<emphasis id="AEDEEABB487CF01C0EEFF965FB2CF94B" box="[1008,1256,1694,1715]" italics="true" pageId="4" pageNumber="505">Doriprismatica marinae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B5ED57C0487CF01C09EFF965FA86F94B" box="[1264,1346,1694,1715]" pageId="4" pageNumber="505" rank="species">
<emphasis id="AEDEEABB487CF01C09EFF965FA86F94B" bold="true" box="[1264,1346,1694,1715]" pageId="4" pageNumber="505">sp. nov.</emphasis>
</taxonomicNameLabel>
CASIZ- 194050, 
<taxonomicName id="5BAA4D2A487CF01C0DF0F947FE11F92A" authority="Poddubetskaia" authorityName="Poddubetskaia" box="[239,469,1724,1746]" class="Magnoliopsida" family="Fabaceae" genus="Marina" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fabales" pageId="4" pageNumber="505" phylum="Tracheophyta" rank="genus">Marina Poddubetskaia</taxonomicName>
, all other photos by. T. Gosliner.
</paragraph>
</caption>
<subSubSection id="D4B06522487CF01D0DD8F905FD95FBE8" lastPageId="5" lastPageNumber="506" pageId="4" pageNumber="505" type="description">
<paragraph id="9C1536A9487CF01D0DD8F905FB89FE55" blockId="4.[151,1437,1790,2031]" lastBlockId="5.[151,1437,151,2013]" lastPageId="5" lastPageNumber="506" pageId="4" pageNumber="505">
<emphasis id="AEDEEABB487CF01C0DD8F905FE15F8EF" bold="true" box="[199,465,1790,1815]" pageId="4" pageNumber="505">External Morphology.</emphasis>
<taxonomicName id="5BAA4D2A487CF01C0CC6F905FD5BF8EE" authorityName="Matsuda &amp; Gosliner" authorityYear="2018" box="[473,671,1790,1814]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="4" pageNumber="505" phylum="Mollusca" rank="species" species="buko">
<emphasis id="AEDEEABB487CF01C0CC6F905FD5BF8EE" box="[473,671,1790,1814]" italics="true" pageId="4" pageNumber="505">Glossodoris buko</emphasis>
</taxonomicName>
has a long and slender body that is transparent white in color (
<figureCitation id="04912A2C487CF01C0871F905FF1AF8C3" captionStart="FIGURE 1" captionStartId="3.[151,250,1750,1772]" captionTargetBox="[248,1338,592,1723]" captionTargetId="figure@3.[246,1340,591,1726]" captionTargetPageId="3" captionText="FIGURE 1. A–C, Glossodoris buko sp. nov. A. CASIZ-191102 (i. large, ii. small) Papua New Guinea. B. CASIZ-181594 Philippines. C. CASIZ-086381 (A, B) Papua New Guinea. D–F, Glossodoris pallida: D. CASIZ-194338 Madagascar. E. CASIZ-175548 Madagascar. F. CASIZ-175554 Madagascar. Photos: T.M. Gosliner." httpUri="https://zenodo.org/record/1437226/files/figure.png" pageId="4" pageNumber="505">Fig. 1A–C</figureCitation>
). There is an opaque white band that starts anteriorly on the mantle, narrows between the rhinophores and then widens again and narrows between the two major folds in the middle of the mantle, and ends circling the gills (
<figureCitation id="04912A2C487CF01C0D80F891FF39F87B" box="[159,253,1898,1923]" captionStart="FIGURE 2" captionStartId="4.[151,250,1570,1592]" captionTargetBox="[334,1263,243,1490]" captionTargetId="figure@4.[319,1268,225,1493]" captionTargetPageId="4" captionText="FIGURE 2. Mantle margins. A. Glossodoris buko sp. nov., CASIZ-222107. B. Glossodoris pallida (Cuvier, 1804), CASIZ- 175554. C. Glossodoris bowanga sp. nov., CASIZ-194204, photo by Marina Poddubetskaia. D. Glossodoris andersonae sp. nov., CASIZ-192288. E. Glossodoris acosti sp. nov., CASIZ 191352, photo by Vanessa Knutson. F. Glossodoris sp. cf. cincta, CASIZ-199204. G. Glossodoris cincta Bergh, 1888, from Reunion, photo only, photo by Philibert Bidgrain. H. Doriprismatica balut sp. nov., CASIZ-182787. I. Doriprismatica rossi sp. nov. CASIZ-192281. J. Doriprismatica marinae sp. nov. CASIZ- 194050, Marina Poddubetskaia, all other photos by. T. Gosliner." httpUri="https://zenodo.org/record/1437228/files/figure.png" pageId="4" pageNumber="505">Fig. 2A</figureCitation>
). In most specimens, the band is continuous, however there was a break in the band posterior to the rhinophores in some. A white opaque band runs the length of the foot on both sides and connects posteriorly. The mantle edge is rippled with the semi-permanent undulations that are characteristic of all 
<emphasis id="AEDEEABB487CF01C0967F848FAC3F833" box="[1144,1287,1971,1995]" italics="true" pageId="4" pageNumber="505">
<taxonomicName id="5BAA4D2A487CF01C0967F848FAC7F833" authorityName="Ehrenberg" authorityYear="1831" box="[1144,1283,1971,1995]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="4" pageNumber="505" phylum="Mollusca" rank="genus">Glossodoris</taxonomicName>
.
</emphasis>
One primary pair of undulations midway on the mantle is identifiable by an indentation of the white dorsal band. A thin, light yellow marginal band runs the length of the outer edge of the mantle and opaque white mantle dermal formations that appear as a thick white band that lies partly under the yellow band. The degree and number of the smaller semi-permanent undulations varies between individuals, but larger specimens have more pronounced undulations. A yellow band borders the foot, however no white band is visible due to the absence of mantle dermal formations. The rhinophores are elongate and conical with 11–12 lamellae. The bases of the rhinophores are white and the tips are yellow. The gill forms a semicircle surrounding the anus opening posteriorly, consisting of approximately 5–8 unipinnate branches. The lamellae are white with yellow tips and are shorter at the ends of the arc. The genital pore is located on the right side of the body below the mantle and behind the rhinophores.
</paragraph>
<paragraph id="9C1536A9487DF01D0DD8FE4CFC44FC78" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="506">
<emphasis id="AEDEEABB487DF01D0DD8FE4CFE5AFE28" bold="true" box="[199,414,439,464]" pageId="5" pageNumber="506">Internal Anatomy</emphasis>
. 
<emphasis id="AEDEEABB487DF01D0CB3FE43FD35FE28" box="[428,753,440,464]" italics="true" pageId="5" pageNumber="506">Radula and buccal armature</emphasis>
(
<figureCitation id="04912A2C487DF01D0E1EFE4CFCB9FE28" box="[769,893,439,464]" captionStart="FIGURE 3" captionStartId="6.[151,250,1919,1941]" captionTargetBox="[183,1406,193,1907]" captionTargetId="figure@6.[183,1407,193,1908]" captionTargetPageId="6" captionText="FIGURE 3. Scanning electron micrographs. Left column (A–E) Glossodoris buko sp. nov., CASIZ-086381A Papua New Guinea, A. Rachis B. Mid-laterals. CASIZ-191102A Papua New Guinea, C. Outer laterals. D. Jaws. E. Radular ribbon. Right column (F–J) Glossodoris pallida, CASIZ-173395 Madagascar: F. Rachis. G. Mid-laterals. H. Outer laterals. I. Jaws. J. Radular ribbon." httpUri="https://zenodo.org/record/1437230/files/figure.png" pageId="5" pageNumber="506">Fig. 3A–E</figureCitation>
). The radular ribbon (
<figureCitation id="04912A2C487DF01D096AFE4CFB0EFE28" box="[1141,1226,439,464]" captionStart="FIGURE 3" captionStartId="6.[151,250,1919,1941]" captionTargetBox="[183,1406,193,1907]" captionTargetId="figure@6.[183,1407,193,1908]" captionTargetPageId="6" captionText="FIGURE 3. Scanning electron micrographs. Left column (A–E) Glossodoris buko sp. nov., CASIZ-086381A Papua New Guinea, A. Rachis B. Mid-laterals. CASIZ-191102A Papua New Guinea, C. Outer laterals. D. Jaws. E. Radular ribbon. Right column (F–J) Glossodoris pallida, CASIZ-173395 Madagascar: F. Rachis. G. Mid-laterals. H. Outer laterals. I. Jaws. J. Radular ribbon." httpUri="https://zenodo.org/record/1437230/files/figure.png" pageId="5" pageNumber="506">Fig. 3E</figureCitation>
) is short and wide with a radula formula for a preserved specimen of 
<quantity id="5B529B4C487DF01D0FFBFE27FCE3FE0C" box="[740,807,476,500]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="506" unit="mm" value="3.0">3 mm</quantity>
of approximately 
<date id="E8141069487DF01D0EE3FE27FBEBFE0C" box="[1020,1071,476,500]" pageId="5" pageNumber="506" value="1928-10">28 x</date>
<date id="E8141069487DF01D0926FE27FB49FE0C" box="[1081,1165,476,500]" pageId="5" pageNumber="506" value="14-01-14">14.1.14</date>
(CASIZ- 
<date id="E8141069487DF01D09E5FE27FA8BFE0C" box="[1274,1359,476,500]" pageId="5" pageNumber="506" value="02-11-19">191102</date>
A) and 
<date id="E8141069487DF01D0D88FDFBFF2FFDE0" box="[151,235,512,536]" pageId="5" pageNumber="506" value="13-01-13">13.1.13</date>
for a 
<quantity id="5B529B4C487DF01D0C29FE04FE54FDEF" box="[310,400,511,536]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="5" pageNumber="506" unit="mm" value="6.5">6.5 mm</quantity>
preserved specimen (CASIZ-086381). The rachidian tooth (
<figureCitation id="04912A2C487DF01D0954FE04FB62FDE0" box="[1099,1190,511,536]" captionStart="FIGURE 3" captionStartId="6.[151,250,1919,1941]" captionTargetBox="[183,1406,193,1907]" captionTargetId="figure@6.[183,1407,193,1908]" captionTargetPageId="6" captionText="FIGURE 3. Scanning electron micrographs. Left column (A–E) Glossodoris buko sp. nov., CASIZ-086381A Papua New Guinea, A. Rachis B. Mid-laterals. CASIZ-191102A Papua New Guinea, C. Outer laterals. D. Jaws. E. Radular ribbon. Right column (F–J) Glossodoris pallida, CASIZ-173395 Madagascar: F. Rachis. G. Mid-laterals. H. Outer laterals. I. Jaws. J. Radular ribbon." httpUri="https://zenodo.org/record/1437230/files/figure.png" pageId="5" pageNumber="506">Fig. 3A</figureCitation>
) is very reduced and quasi-triangular. The first lateral tooth is almost bilaterally symmetrical. It is wide and has a relatively short triangular pointed central cusp. There are approximately five well-defined denticles, each about half the length of the central cusp, that point down and outward on the inner and outer sides of the central cusp. The mid-lateral teeth (
<figureCitation id="04912A2C487DF01D0D80FD74FF3DFD50" box="[159,249,655,680]" captionStart="FIGURE 3" captionStartId="6.[151,250,1919,1941]" captionTargetBox="[183,1406,193,1907]" captionTargetId="figure@6.[183,1407,193,1908]" captionTargetPageId="6" captionText="FIGURE 3. Scanning electron micrographs. Left column (A–E) Glossodoris buko sp. nov., CASIZ-086381A Papua New Guinea, A. Rachis B. Mid-laterals. CASIZ-191102A Papua New Guinea, C. Outer laterals. D. Jaws. E. Radular ribbon. Right column (F–J) Glossodoris pallida, CASIZ-173395 Madagascar: F. Rachis. G. Mid-laterals. H. Outer laterals. I. Jaws. J. Radular ribbon." httpUri="https://zenodo.org/record/1437230/files/figure.png" pageId="5" pageNumber="506">Fig. 3B</figureCitation>
) are longer and have a shorter central cusp with approximately 6–8 loosely packed and well defined denticles solely on the outer edge. Unlike the first lateral, the central cusp on the mid-laterals is almost indistinguishable from the denticles next to it. The denticles in the mid-laterals are almost indistinguishable in size and shape from the central cusp, and maintain this shape and their size integrity to the edge of the ribbon (
<figureCitation id="04912A2C487DF01D0828FD07FA4BFCEC" box="[1335,1423,764,789]" captionStart="FIGURE 3" captionStartId="6.[151,250,1919,1941]" captionTargetBox="[183,1406,193,1907]" captionTargetId="figure@6.[183,1407,193,1908]" captionTargetPageId="6" captionText="FIGURE 3. Scanning electron micrographs. Left column (A–E) Glossodoris buko sp. nov., CASIZ-086381A Papua New Guinea, A. Rachis B. Mid-laterals. CASIZ-191102A Papua New Guinea, C. Outer laterals. D. Jaws. E. Radular ribbon. Right column (F–J) Glossodoris pallida, CASIZ-173395 Madagascar: F. Rachis. G. Mid-laterals. H. Outer laterals. I. Jaws. J. Radular ribbon." httpUri="https://zenodo.org/record/1437230/files/figure.png" pageId="5" pageNumber="506">Fig. 3C</figureCitation>
). The jaw rodlets are short and well-spaced with distinct gaps between rodlets. They are predominantly bifid (
<figureCitation id="04912A2C487DF01D0870FCE4FF7EFCA4" captionStart="FIGURE 3" captionStartId="6.[151,250,1919,1941]" captionTargetBox="[183,1406,193,1907]" captionTargetId="figure@6.[183,1407,193,1908]" captionTargetPageId="6" captionText="FIGURE 3. Scanning electron micrographs. Left column (A–E) Glossodoris buko sp. nov., CASIZ-086381A Papua New Guinea, A. Rachis B. Mid-laterals. CASIZ-191102A Papua New Guinea, C. Outer laterals. D. Jaws. E. Radular ribbon. Right column (F–J) Glossodoris pallida, CASIZ-173395 Madagascar: F. Rachis. G. Mid-laterals. H. Outer laterals. I. Jaws. J. Radular ribbon." httpUri="https://zenodo.org/record/1437230/files/figure.png" pageId="5" pageNumber="506">Fig. 3D</figureCitation>
) with a few trifid rodlets. The ventral side of the buccal mass has a glandular sheath covering the oral tube that contains numerous densely packed white opaque glands (
<figureCitation id="04912A2C487DF01D0E06FC9CFCB0FC78" box="[793,884,871,896]" captionStart="FIGURE 4" captionStartId="7.[151,250,1117,1139]" captionTargetBox="[260,1353,211,1083]" captionTargetId="figure@7.[234,1353,193,1096]" captionTargetPageId="7" captionText="FIGURE 4. Glossodoris pallida, CASIZ-173395: (A) Reproductive system (B) Buccal mass, ventral side has no glands. Glossodoris buko sp. nov., CASIZ-086381A: (C) Reproductive system (D) Buccal mass, ventral side with glandular sheath. Abbreviations: p, penis; vd, vas deferens; pr, prostate; v, vaginal duct; rs, receptaculum seminis; bc, bursa copulatrix; am, ampulla; fgm, female gland mass; ot, oral tube, bb, buccal bulb." httpUri="https://zenodo.org/record/1437232/files/figure.png" pageId="5" pageNumber="506">Fig. 4D</figureCitation>
).
</paragraph>
<paragraph id="9C1536A9487DF01D0DD8FC77FD95FBE8" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="506">
<emphasis id="AEDEEABB487DF01D0DD8FC77FE6AFC5C" box="[199,430,908,932]" italics="true" pageId="5" pageNumber="506">Reproductive system</emphasis>
(
<figureCitation id="04912A2C487DF01D0CA2FC77FDD7FC5C" box="[445,531,908,933]" captionStart="FIGURE 4" captionStartId="7.[151,250,1117,1139]" captionTargetBox="[260,1353,211,1083]" captionTargetId="figure@7.[234,1353,193,1096]" captionTargetPageId="7" captionText="FIGURE 4. Glossodoris pallida, CASIZ-173395: (A) Reproductive system (B) Buccal mass, ventral side has no glands. Glossodoris buko sp. nov., CASIZ-086381A: (C) Reproductive system (D) Buccal mass, ventral side with glandular sheath. Abbreviations: p, penis; vd, vas deferens; pr, prostate; v, vaginal duct; rs, receptaculum seminis; bc, bursa copulatrix; am, ampulla; fgm, female gland mass; ot, oral tube, bb, buccal bulb." httpUri="https://zenodo.org/record/1437232/files/figure.png" pageId="5" pageNumber="506">Fig. 4C</figureCitation>
). The penial bulb is long and folded and leads to a coiled vas deferens followed by an approximately equal in length prostate gland. The receptaculum seminis duct and the vagina are relatively short. The receptaculum seminis is slightly smaller than the bursa copulatrix, and they are found adjacent to each other rather than being aligned linearly.
</paragraph>
</subSubSection>
<subSubSection id="D4B06522487DF01F0DD8FBE7FD3EFA79" lastPageId="7" lastPageNumber="508" pageId="5" pageNumber="506" type="discussion">
<paragraph id="9C1536A9487DF01D0DD8FBE7FAFAFA65" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="506">
<emphasis id="AEDEEABB487DF01D0DD8FBE7FEF0FBCD" bold="true" box="[199,308,1052,1077]" pageId="5" pageNumber="506">Remarks</emphasis>
. At first glance, 
<taxonomicName id="5BAA4D2A487DF01D0F13FBE7FDACFBCC" authorityName="Matsuda &amp; Gosliner" authorityYear="2018" box="[524,616,1052,1076]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="506" phylum="Mollusca" rank="species" species="buko">
<emphasis id="AEDEEABB487DF01D0F13FBE7FDACFBCC" box="[524,616,1052,1076]" italics="true" pageId="5" pageNumber="506">G. buko</emphasis>
</taxonomicName>
and 
<taxonomicName id="5BAA4D2A487DF01D0FB2FBE7FCE6FBCC" baseAuthorityName="Cuvier" baseAuthorityYear="1804" box="[685,802,1052,1076]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="506" phylum="Mollusca" rank="species" species="pallida">
<emphasis id="AEDEEABB487DF01D0FB2FBE7FCE6FBCC" box="[685,802,1052,1076]" italics="true" pageId="5" pageNumber="506">G. pallida</emphasis>
</taxonomicName>
could be easily confused, as there are few external morphological differences (
<figureCitation id="04912A2C487DF01D0CCEFBC4FDD2FBA0" box="[465,534,1087,1112]" captionStart="FIGURE 1" captionStartId="3.[151,250,1750,1772]" captionTargetBox="[248,1338,592,1723]" captionTargetId="figure@3.[246,1340,591,1726]" captionTargetPageId="3" captionText="FIGURE 1. A–C, Glossodoris buko sp. nov. A. CASIZ-191102 (i. large, ii. small) Papua New Guinea. B. CASIZ-181594 Philippines. C. CASIZ-086381 (A, B) Papua New Guinea. D–F, Glossodoris pallida: D. CASIZ-194338 Madagascar. E. CASIZ-175548 Madagascar. F. CASIZ-175554 Madagascar. Photos: T.M. Gosliner." httpUri="https://zenodo.org/record/1437226/files/figure.png" pageId="5" pageNumber="506">Fig. 1</figureCitation>
; 
<bibRefCitation id="F83B4B58487DF01D0F38FBC4FD0FFBA0" box="[551,715,1087,1112]" pageId="5" pageNumber="506" refId="ref17040" refString="Rudman, W. B. (1984) The Chrornodorididae (Opisthobranchia: Mollusca) of the Indo-West Pacific: a review of the genera. Zoological Journal of the Linnean Society, 81, 115 - 273. https: // doi. org / 10.1111 / j. 1096 - 3642.1984. tb 01174. x" type="journal article">Rudman 1984</bibRefCitation>
: fig 1b; 
<bibRefCitation id="F83B4B58487DF01D0E31FBC4FBD3FBA0" box="[814,1047,1087,1112]" pageId="5" pageNumber="506" refId="ref16064" refString="Gosliner, T. M., Valdes, A. &amp; Behrens, D. W. (2015) Nudibranch and Sea Slug Identification-Indo-Pacific. New World Publications, Jacksonville, Florida, 236 pp." type="book">
Gosliner 
<emphasis id="AEDEEABB487DF01D0E88FBBAFC0EFBA0" box="[919,970,1088,1112]" italics="true" pageId="5" pageNumber="506">et al</emphasis>
. 2015
</bibRefCitation>
: 237, upper right fig.; 
<bibRefCitation id="F83B4B58487DF01D0803FBC4FEFFFB84" pageId="5" pageNumber="506" refId="ref16546" refString="Matsuda, S. B. &amp; Gosliner, T. M. (2017) Molecular phylogeny of Glossodoris (Ehrenberg, 1831) nudibranchs and related genera reveals cryptic and pseudocryptic species complexes. Cladistics, 34 (1), 41 - 56. https: // doi. org / 10.1111 / cla. 12194" type="journal article">Matsuda &amp; Gosliner 2017</bibRefCitation>
: fig. 1). The 
<typeStatus id="4311880B487DF01D0CCFFB9FFDF6FB85" box="[464,562,1124,1149]" pageId="5" pageNumber="506" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="5BAA4D2A487DF01D0F46FB9FFD0CFB84" baseAuthorityName="Cuvier" baseAuthorityYear="1804" box="[601,712,1124,1148]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="506" phylum="Mollusca" rank="species" species="pallida">
<emphasis id="AEDEEABB487DF01D0F46FB9FFD0CFB84" box="[601,712,1124,1148]" italics="true" pageId="5" pageNumber="506">G. pallida</emphasis>
</taxonomicName>
was collected from the 
<collectingRegion id="5E6EF84B487DF01D0EC5FB9FFBFCFB84" box="[986,1080,1124,1149]" country="Sudan" name="Al Bahr al Ahmar" pageId="5" pageNumber="506">Red Sea</collectingRegion>
and subsequently examined by 
<bibRefCitation id="F83B4B58487DF01D0D88FB7CFE8EFB58" box="[151,330,1159,1184]" pageId="5" pageNumber="506" refId="ref17040" refString="Rudman, W. B. (1984) The Chrornodorididae (Opisthobranchia: Mollusca) of the Indo-West Pacific: a review of the genera. Zoological Journal of the Linnean Society, 81, 115 - 273. https: // doi. org / 10.1111 / j. 1096 - 3642.1984. tb 01174. x" type="journal article">Rudman (1984)</bibRefCitation>
together with a specimen from 
<collectingCountry id="E4BD7639487DF01D0FDEFB7CFCEDFB58" box="[705,809,1159,1184]" name="Tanzania" pageId="5" pageNumber="506">Tanzania</collectingCountry>
. Both specimens share the same color pattern, radular structure and reproductive system morphology as the five specimens of 
<taxonomicName id="5BAA4D2A487DF01D0EA8FB57FBE1FB3C" baseAuthorityName="Cuvier" baseAuthorityYear="1804" box="[951,1061,1196,1220]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="506" phylum="Mollusca" rank="species" species="pallida">
<emphasis id="AEDEEABB487DF01D0EA8FB57FBE1FB3C" box="[951,1061,1196,1220]" italics="true" pageId="5" pageNumber="506">G. pallida</emphasis>
</taxonomicName>
we comparatively examined here from 
<collectingCountry id="E4BD7639487DF01D0DC9FB34FE99FB10" box="[214,349,1231,1256]" name="Madagascar" pageId="5" pageNumber="506">Madagascar</collectingCountry>
and 
<collectingCountry id="E4BD7639487DF01D0C89FB34FDEEFB10" box="[406,554,1231,1256]" name="Mozambique" pageId="5" pageNumber="506">Mozambique</collectingCountry>
(
<figureCitation id="04912A2C487DF01D0F24FB34FD7FFB10" box="[571,699,1231,1256]" captionStart="FIGURE 1" captionStartId="3.[151,250,1750,1772]" captionTargetBox="[248,1338,592,1723]" captionTargetId="figure@3.[246,1340,591,1726]" captionTargetPageId="3" captionText="FIGURE 1. A–C, Glossodoris buko sp. nov. A. CASIZ-191102 (i. large, ii. small) Papua New Guinea. B. CASIZ-181594 Philippines. C. CASIZ-086381 (A, B) Papua New Guinea. D–F, Glossodoris pallida: D. CASIZ-194338 Madagascar. E. CASIZ-175548 Madagascar. F. CASIZ-175554 Madagascar. Photos: T.M. Gosliner." httpUri="https://zenodo.org/record/1437226/files/figure.png" pageId="5" pageNumber="506">Figs. 1D–F</figureCitation>
, 
<figureCitation id="04912A2C487DF01D0FD7FB2BFCC7FB10" box="[712,771,1232,1256]" captionStart="FIGURE 3" captionStartId="6.[151,250,1919,1941]" captionTargetBox="[183,1406,193,1907]" captionTargetId="figure@6.[183,1407,193,1908]" captionTargetPageId="6" captionText="FIGURE 3. Scanning electron micrographs. Left column (A–E) Glossodoris buko sp. nov., CASIZ-086381A Papua New Guinea, A. Rachis B. Mid-laterals. CASIZ-191102A Papua New Guinea, C. Outer laterals. D. Jaws. E. Radular ribbon. Right column (F–J) Glossodoris pallida, CASIZ-173395 Madagascar: F. Rachis. G. Mid-laterals. H. Outer laterals. I. Jaws. J. Radular ribbon." httpUri="https://zenodo.org/record/1437230/files/figure.png" pageId="5" pageNumber="506">3F–J</figureCitation>
). In 
<bibRefCitation id="F83B4B58487DF01D0E28FB34FC38FB10" box="[823,1020,1231,1256]" pageId="5" pageNumber="506" refId="ref17167" refString="Rudman, W. B. (1990) The Chromodorididae (Opisthobranchia: Mollusca) of the Indo-West Pacific: further species of Glossodoris, Thorunna and the Chromodoris aureomarginata colour group. Linnean Society of London, 100, 263 - 326. https: // doi. org / 10.1111 / j. 1096 - 3642.1990. tb 01864. x" type="journal article">Rudman’s (1990)</bibRefCitation>
<taxonomicName id="5BAA4D2A487DF01D091BFB2BFBB0FB10" baseAuthorityName="Cuvier" baseAuthorityYear="1804" box="[1028,1140,1232,1256]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="506" phylum="Mollusca" rank="species" species="pallida">
<emphasis id="AEDEEABB487DF01D091BFB2BFBB0FB10" box="[1028,1140,1232,1256]" italics="true" pageId="5" pageNumber="506">G. pallida</emphasis>
</taxonomicName>
description, he noted that specimens from East 
<collectingCountry id="E4BD7639487DF01D0C94FB0FFE36FAF5" box="[395,498,1268,1293]" name="Australia" pageId="5" pageNumber="506">Australia</collectingCountry>
have yellow gills and rhinophores, which are consistent with 
<taxonomicName id="5BAA4D2A487DF01D09ADFB0FFACCFAF4" authorityName="Matsuda &amp; Gosliner" authorityYear="2018" box="[1202,1288,1268,1292]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="506" phylum="Mollusca" rank="species" species="buko">
<emphasis id="AEDEEABB487DF01D09ADFB0FFACCFAF4" box="[1202,1288,1268,1292]" italics="true" pageId="5" pageNumber="506">G. buko</emphasis>
</taxonomicName>
, whereas his 
<collectingCountry id="E4BD7639487DF01D0D88FAECFF38FAC8" box="[151,252,1303,1328]" name="Tanzania" pageId="5" pageNumber="506">Tanzania</collectingCountry>
and 
<collectingCountry id="E4BD7639487DF01D0C2BFAECFEBEFAC8" box="[308,378,1303,1328]" name="Sudan" pageId="5" pageNumber="506">Sudan</collectingCountry>
specimens, the rhinophores and gills are white. However, he further remarks that yellow tips were reported from specimens in the 
<collectingRegion id="5E6EF84B487DF01D0F48FAC7FD7DFAAC" box="[599,697,1340,1365]" country="Sudan" name="Al Bahr al Ahmar" pageId="5" pageNumber="506">Red Sea</collectingRegion>
and 
<collectingCountry id="E4BD7639487DF01D0FE5FAC7FC9DFAAD" box="[762,857,1340,1365]" name="Réunion" pageId="5" pageNumber="506">Reunion</collectingCountry>
Island, indicating that yellow tips may not be a consistent identifier for 
<emphasis id="AEDEEABB487DF01D0CB0FA9BFDC9FA80" box="[431,525,1376,1400]" italics="true" pageId="5" pageNumber="506">
<taxonomicName id="5BAA4D2A487DF01D0CB0FA9BFDCDFA80" authorityName="Matsuda &amp; Gosliner" authorityYear="2018" box="[431,521,1376,1400]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="506" phylum="Mollusca" rank="species" species="buko">G. buko</taxonomicName>
.
</emphasis>
The rhinophores and gills of our 
<taxonomicName id="5BAA4D2A487DF01D0EBCFA9BFBD1FA80" baseAuthorityName="Cuvier" baseAuthorityYear="1804" box="[931,1045,1376,1400]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="506" phylum="Mollusca" rank="species" species="pallida">
<emphasis id="AEDEEABB487DF01D0EBCFA9BFBD1FA80" box="[931,1045,1376,1400]" italics="true" pageId="5" pageNumber="506">G. pallida</emphasis>
</taxonomicName>
specimens from 
<collectingCountry id="E4BD7639487DF01D09FEFAA4FAACFA80" box="[1249,1384,1375,1400]" name="Madagascar" pageId="5" pageNumber="506">Madagascar</collectingCountry>
and 
<collectingCountry id="E4BD7639487DF01D0D88FA7FFEEFFA65" box="[151,299,1412,1437]" name="Mozambique" pageId="5" pageNumber="506">Mozambique</collectingCountry>
all have frosted yellow tips, although the yellow is not as bright as in the 
<taxonomicName id="5BAA4D2A487DF01D097AFA7FFB7EFA64" authorityName="Matsuda &amp; Gosliner" authorityYear="2018" box="[1125,1210,1412,1436]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="506" phylum="Mollusca" rank="species" species="buko">
<emphasis id="AEDEEABB487DF01D097AFA7FFB7EFA64" box="[1125,1210,1412,1436]" italics="true" pageId="5" pageNumber="506">G. buko</emphasis>
</taxonomicName>
specimens.
</paragraph>
<paragraph id="9C1536A9487DF01D0DD8FA5CFBBEF824" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="506">
The most striking differences between the two species are found in the radula, jaws and buccal mass. The radular ribbon of 
<taxonomicName id="5BAA4D2A487DF01D0C43FA37FE02FA1D" baseAuthorityName="Cuvier" baseAuthorityYear="1804" box="[348,454,1484,1509]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="506" phylum="Mollusca" rank="species" species="pallida">G. pallida</taxonomicName>
(
<figureCitation id="04912A2C487DF01D0CC9FA37FDE0FA1C" box="[470,548,1484,1509]" captionStart="FIGURE 3" captionStartId="6.[151,250,1919,1941]" captionTargetBox="[183,1406,193,1907]" captionTargetId="figure@6.[183,1407,193,1908]" captionTargetPageId="6" captionText="FIGURE 3. Scanning electron micrographs. Left column (A–E) Glossodoris buko sp. nov., CASIZ-086381A Papua New Guinea, A. Rachis B. Mid-laterals. CASIZ-191102A Papua New Guinea, C. Outer laterals. D. Jaws. E. Radular ribbon. Right column (F–J) Glossodoris pallida, CASIZ-173395 Madagascar: F. Rachis. G. Mid-laterals. H. Outer laterals. I. Jaws. J. Radular ribbon." httpUri="https://zenodo.org/record/1437230/files/figure.png" pageId="5" pageNumber="506">Fig. 3J</figureCitation>
) is elongate (~ 
<date id="E8141069487DF01D0FD4FA37FD39FA1C" box="[715,765,1484,1509]" pageId="5" pageNumber="506" value="1985-10">85 x</date>
<date id="E8141069487DF01D0E1CFA37FC9CFA1C" box="[771,856,1484,1508]" pageId="5" pageNumber="506" value="23-01-23">23.1.23</date>
CASIZ-173395), whereas it is short and squat in 
<taxonomicName id="5BAA4D2A487DF01D089BFA37FF0BF9F0" authorityName="Matsuda &amp; Gosliner" authorityYear="2018" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="506" phylum="Mollusca" rank="species" species="buko">G. buko</taxonomicName>
(
<figureCitation id="04912A2C487DF01D0DFDFA14FEFEF9F0" box="[226,314,1519,1544]" captionStart="FIGURE 3" captionStartId="6.[151,250,1919,1941]" captionTargetBox="[183,1406,193,1907]" captionTargetId="figure@6.[183,1407,193,1908]" captionTargetPageId="6" captionText="FIGURE 3. Scanning electron micrographs. Left column (A–E) Glossodoris buko sp. nov., CASIZ-086381A Papua New Guinea, A. Rachis B. Mid-laterals. CASIZ-191102A Papua New Guinea, C. Outer laterals. D. Jaws. E. Radular ribbon. Right column (F–J) Glossodoris pallida, CASIZ-173395 Madagascar: F. Rachis. G. Mid-laterals. H. Outer laterals. I. Jaws. J. Radular ribbon." httpUri="https://zenodo.org/record/1437230/files/figure.png" pageId="5" pageNumber="506">Fig. 3E</figureCitation>
) (~ 
<date id="E8141069487DF01D0C6DFA0BFE63F9F0" box="[370,423,1520,1544]" pageId="5" pageNumber="506" value="1928-10">28 x</date>
<date id="E8141069487DF01D0CADFA0BFDCCF9F0" box="[434,520,1520,1544]" pageId="5" pageNumber="506" value="14-01-14">14.1.14</date>
) for specimens of comparable size. This is confirmed in Rudman’s 
<taxonomicName id="5BAA4D2A487DF01D0835FA0BFA5FF9F0" baseAuthorityName="Cuvier" baseAuthorityYear="1804" box="[1322,1435,1520,1544]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="506" phylum="Mollusca" rank="species" species="pallida">
<emphasis id="AEDEEABB487DF01D0835FA0BFA5FF9F0" box="[1322,1435,1520,1544]" italics="true" pageId="5" pageNumber="506">G. pallida</emphasis>
</taxonomicName>
specimen from the 
<collectingRegion id="5E6EF84B487DF01D0C6BF9EFFE17F9D4" box="[372,467,1556,1581]" country="Sudan" name="Al Bahr al Ahmar" pageId="5" pageNumber="506">Red Sea</collectingRegion>
(
<typeStatus id="4311880B487DF01D0CFBF9EFFDD2F9D4" box="[484,534,1556,1580]" pageId="5" pageNumber="506">type</typeStatus>
locality), which has a radular formula of 108(+4) x 
<date id="E8141069487DF01D096DF9EFFB02F9D4" box="[1138,1222,1556,1580]" pageId="5" pageNumber="506" value="39-01-39">39.1.39</date>
(
<quantity id="5B529B4C487DF01D09C7F9EFFAEEF9D4" box="[1240,1322,1556,1581]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="506" unit="mm" value="15.0">15 mm</quantity>
specimen alive) and 
<date id="E8141069487DF01D0C07F9C3FEA9F9A8" box="[280,365,1592,1616]" pageId="5" pageNumber="506" value="23-01-23">23.1.23</date>
for his Australian specimen (
<quantity id="5B529B4C487DF01D0FCAF9C3FCD8F9A8" box="[725,796,1592,1616]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="5" pageNumber="506" unit="mm" value="9.0">9 mm</quantity>
preserved) (
<bibRefCitation id="F83B4B58487DF01D0EADF9CCFB98F9A8" box="[946,1116,1591,1616]" pageId="5" pageNumber="506" refId="ref17040" refString="Rudman, W. B. (1984) The Chrornodorididae (Opisthobranchia: Mollusca) of the Indo-West Pacific: a review of the genera. Zoological Journal of the Linnean Society, 81, 115 - 273. https: // doi. org / 10.1111 / j. 1096 - 3642.1984. tb 01174. x" type="journal article">Rudman 1984</bibRefCitation>
), and while no length was reported, the number of lateral teeth and presence of a rachidian tooth suggest that this specimen is likely 
<emphasis id="AEDEEABB487DF01D0820F9A7FEBEF960" italics="true" pageId="5" pageNumber="506">
<taxonomicName id="5BAA4D2A487DF01D0820F9A7FA5CF98C" authorityName="Matsuda &amp; Gosliner" authorityYear="2018" box="[1343,1432,1628,1652]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="506" phylum="Mollusca" rank="species" species="buko">G. buko</taxonomicName>
. 
<taxonomicName id="5BAA4D2A487DF01D0D88F97BFEBEF960" baseAuthorityName="Cuvier" baseAuthorityYear="1804" box="[151,378,1664,1688]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="506" phylum="Mollusca" rank="species" species="pallida">Glossodoris pallida</taxonomicName>
</emphasis>
has a more prominent and pointed rachidian tooth (
<figureCitation id="04912A2C487DF01D0EF0F984FB8EF960" box="[1007,1098,1663,1688]" captionStart="FIGURE 3" captionStartId="6.[151,250,1919,1941]" captionTargetBox="[183,1406,193,1907]" captionTargetId="figure@6.[183,1407,193,1908]" captionTargetPageId="6" captionText="FIGURE 3. Scanning electron micrographs. Left column (A–E) Glossodoris buko sp. nov., CASIZ-086381A Papua New Guinea, A. Rachis B. Mid-laterals. CASIZ-191102A Papua New Guinea, C. Outer laterals. D. Jaws. E. Radular ribbon. Right column (F–J) Glossodoris pallida, CASIZ-173395 Madagascar: F. Rachis. G. Mid-laterals. H. Outer laterals. I. Jaws. J. Radular ribbon." httpUri="https://zenodo.org/record/1437230/files/figure.png" pageId="5" pageNumber="506">Fig. 3F</figureCitation>
), whereas the rachidians are significantly reduced and amorphous in 
<taxonomicName id="5BAA4D2A487DF01D0F7FF95FFD77F944" authorityName="Matsuda &amp; Gosliner" authorityYear="2018" box="[608,691,1700,1724]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="506" phylum="Mollusca" rank="species" species="buko">
<emphasis id="AEDEEABB487DF01D0F7FF95FFD77F944" box="[608,691,1700,1724]" italics="true" pageId="5" pageNumber="506">G. buko</emphasis>
</taxonomicName>
. There are also significant dissimilarities in the lateral teeth. The East African 
<taxonomicName id="5BAA4D2A487DF01D0C2CF933FE60F918" baseAuthorityName="Cuvier" baseAuthorityYear="1804" box="[307,420,1736,1760]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="506" phylum="Mollusca" rank="species" species="pallida">
<emphasis id="AEDEEABB487DF01D0C2CF933FE60F918" box="[307,420,1736,1760]" italics="true" pageId="5" pageNumber="506">G. pallida</emphasis>
</taxonomicName>
specimens examined here share the same lateral tooth structure as Rudman’s 
<collectingRegion id="5E6EF84B487DF01D0824F93CFA58F918" box="[1339,1436,1735,1760]" country="Sudan" name="Al Bahr al Ahmar" pageId="5" pageNumber="506">Red Sea</collectingRegion>
specimen (1984), with a long narrow hook-shaped central cusp with well-defined short denticles resting flat against the outer edge (
<figureCitation id="04912A2C487DF01D0C59F8F4FE04F8D0" box="[326,448,1807,1832]" captionStart="FIGURE 3" captionStartId="6.[151,250,1919,1941]" captionTargetBox="[183,1406,193,1907]" captionTargetId="figure@6.[183,1407,193,1908]" captionTargetPageId="6" captionText="FIGURE 3. Scanning electron micrographs. Left column (A–E) Glossodoris buko sp. nov., CASIZ-086381A Papua New Guinea, A. Rachis B. Mid-laterals. CASIZ-191102A Papua New Guinea, C. Outer laterals. D. Jaws. E. Radular ribbon. Right column (F–J) Glossodoris pallida, CASIZ-173395 Madagascar: F. Rachis. G. Mid-laterals. H. Outer laterals. I. Jaws. J. Radular ribbon." httpUri="https://zenodo.org/record/1437230/files/figure.png" pageId="5" pageNumber="506">Fig. 3G, H</figureCitation>
). 
<taxonomicName id="5BAA4D2A487DF01D0CCBF8EBFD5DF8D0" authorityName="Matsuda &amp; Gosliner" authorityYear="2018" box="[468,665,1808,1832]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="506" phylum="Mollusca" rank="species" species="buko">
<emphasis id="AEDEEABB487DF01D0CCBF8EBFD5DF8D0" box="[468,665,1808,1832]" italics="true" pageId="5" pageNumber="506">Glossodoris buko</emphasis>
</taxonomicName>
has a lateral tooth that is broad and concave with a central cusp that is almost indistinguishable from the denticles in size and shape that shares no similarities with 
<taxonomicName id="5BAA4D2A487DF01D09C6F8CFFA8DF8B4" baseAuthorityName="Cuvier" baseAuthorityYear="1804" box="[1241,1353,1844,1868]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="506" phylum="Mollusca" rank="species" species="pallida">
<emphasis id="AEDEEABB487DF01D09C6F8CFFA8DF8B4" box="[1241,1353,1844,1868]" italics="true" pageId="5" pageNumber="506">G. pallida</emphasis>
</taxonomicName>
. These differences are also visible in the jaw. 
<emphasis id="AEDEEABB487DF01D0F59F8A3FCE8F888" box="[582,812,1880,1904]" italics="true" pageId="5" pageNumber="506">
<taxonomicName id="5BAA4D2A487DF01D0F59F8A3FD09F888" authorityName="Ehrenberg" authorityYear="1831" box="[582,717,1880,1904]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="506" phylum="Mollusca" rank="genus">Glossodoris</taxonomicName>
pallida’
</emphasis>
s rodlets are long, curved, and tightly packed with bifid tips where one of the points protrudes from slightly below the adjacent rodlet (
<figureCitation id="04912A2C487DF01D0917F887FB93F86C" box="[1032,1111,1916,1941]" captionStart="FIGURE 3" captionStartId="6.[151,250,1919,1941]" captionTargetBox="[183,1406,193,1907]" captionTargetId="figure@6.[183,1407,193,1908]" captionTargetPageId="6" captionText="FIGURE 3. Scanning electron micrographs. Left column (A–E) Glossodoris buko sp. nov., CASIZ-086381A Papua New Guinea, A. Rachis B. Mid-laterals. CASIZ-191102A Papua New Guinea, C. Outer laterals. D. Jaws. E. Radular ribbon. Right column (F–J) Glossodoris pallida, CASIZ-173395 Madagascar: F. Rachis. G. Mid-laterals. H. Outer laterals. I. Jaws. J. Radular ribbon." httpUri="https://zenodo.org/record/1437230/files/figure.png" pageId="5" pageNumber="506">
Fig. 
<date id="E8141069487DF01D0922F887FB93F86C" box="[1085,1111,1916,1940]" pageId="5" pageNumber="506">3I</date>
</figureCitation>
). 
<taxonomicName id="5BAA4D2A487DF01D0973F887FAF4F86C" authorityName="Matsuda &amp; Gosliner" authorityYear="2018" box="[1132,1328,1916,1940]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="506" phylum="Mollusca" rank="species" species="buko">
<emphasis id="AEDEEABB487DF01D0973F887FAF4F86C" box="[1132,1328,1916,1940]" italics="true" pageId="5" pageNumber="506">Glossodoris buko</emphasis>
</taxonomicName>
has short and loosely packed rodlets that are less consistent in shape (
<figureCitation id="04912A2C487DF01D0E4EF85BFC6AF840" box="[849,942,1952,1977]" captionStart="FIGURE 3" captionStartId="6.[151,250,1919,1941]" captionTargetBox="[183,1406,193,1907]" captionTargetId="figure@6.[183,1407,193,1908]" captionTargetPageId="6" captionText="FIGURE 3. Scanning electron micrographs. Left column (A–E) Glossodoris buko sp. nov., CASIZ-086381A Papua New Guinea, A. Rachis B. Mid-laterals. CASIZ-191102A Papua New Guinea, C. Outer laterals. D. Jaws. E. Radular ribbon. Right column (F–J) Glossodoris pallida, CASIZ-173395 Madagascar: F. Rachis. G. Mid-laterals. H. Outer laterals. I. Jaws. J. Radular ribbon." httpUri="https://zenodo.org/record/1437230/files/figure.png" pageId="5" pageNumber="506">Fig. 3D</figureCitation>
). Finally, the large glandular sheath on the ventral side of the buccal mass in 
<taxonomicName id="5BAA4D2A487DF01D0F0CF83FFDACF824" authorityName="Matsuda &amp; Gosliner" authorityYear="2018" box="[531,616,1988,2012]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="506" phylum="Mollusca" rank="species" species="buko">
<emphasis id="AEDEEABB487DF01D0F0CF83FFDACF824" box="[531,616,1988,2012]" italics="true" pageId="5" pageNumber="506">G. buko</emphasis>
</taxonomicName>
(
<figureCitation id="04912A2C487DF01D0F68F83FFD0BF824" box="[631,719,1988,2013]" captionStart="FIGURE 4" captionStartId="7.[151,250,1117,1139]" captionTargetBox="[260,1353,211,1083]" captionTargetId="figure@7.[234,1353,193,1096]" captionTargetPageId="7" captionText="FIGURE 4. Glossodoris pallida, CASIZ-173395: (A) Reproductive system (B) Buccal mass, ventral side has no glands. Glossodoris buko sp. nov., CASIZ-086381A: (C) Reproductive system (D) Buccal mass, ventral side with glandular sheath. Abbreviations: p, penis; vd, vas deferens; pr, prostate; v, vaginal duct; rs, receptaculum seminis; bc, bursa copulatrix; am, ampulla; fgm, female gland mass; ot, oral tube, bb, buccal bulb." httpUri="https://zenodo.org/record/1437232/files/figure.png" pageId="5" pageNumber="506">Fig. 4D</figureCitation>
) is not present in 
<taxonomicName id="5BAA4D2A487DF01D0E88F83FFBC1F824" baseAuthorityName="Cuvier" baseAuthorityYear="1804" box="[919,1029,1988,2012]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="506" phylum="Mollusca" rank="species" species="pallida">
<emphasis id="AEDEEABB487DF01D0E88F83FFBC1F824" box="[919,1029,1988,2012]" italics="true" pageId="5" pageNumber="506">G. pallida</emphasis>
</taxonomicName>
(
<figureCitation id="04912A2C487DF01D090BF83FFBA9F824" box="[1044,1133,1988,2013]" captionStart="FIGURE 4" captionStartId="7.[151,250,1117,1139]" captionTargetBox="[260,1353,211,1083]" captionTargetId="figure@7.[234,1353,193,1096]" captionTargetPageId="7" captionText="FIGURE 4. Glossodoris pallida, CASIZ-173395: (A) Reproductive system (B) Buccal mass, ventral side has no glands. Glossodoris buko sp. nov., CASIZ-086381A: (C) Reproductive system (D) Buccal mass, ventral side with glandular sheath. Abbreviations: p, penis; vd, vas deferens; pr, prostate; v, vaginal duct; rs, receptaculum seminis; bc, bursa copulatrix; am, ampulla; fgm, female gland mass; ot, oral tube, bb, buccal bulb." httpUri="https://zenodo.org/record/1437232/files/figure.png" pageId="5" pageNumber="506">Fig. 4B</figureCitation>
).
</paragraph>
<caption id="C8D56621487EF01E0D88F884FF24F809" httpUri="https://zenodo.org/record/1437230/files/figure.png" pageId="6" pageNumber="507" startId="6.[151,250,1919,1941]" targetBox="[183,1406,193,1907]" targetPageId="6">
<paragraph id="9C1536A9487EF01E0D88F884FF24F809" blockId="6.[151,1436,1919,2033]" pageId="6" pageNumber="507">
<emphasis id="AEDEEABB487EF01E0D88F884FED2F86C" bold="true" box="[151,278,1919,1941]" pageId="6" pageNumber="507">FIGURE 3.</emphasis>
Scanning electron micrographs. Left column (A–E) 
<taxonomicName id="5BAA4D2A487EF01E0E59F884FC38F86D" authority="Matsuda &amp; Gosliner, 2018" authorityName="Matsuda &amp; Gosliner" authorityYear="2018" box="[838,1020,1919,1941]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="507" phylum="Mollusca" rank="species" species="buko" status="sp. nov.">
<emphasis id="AEDEEABB487EF01E0E59F884FC38F86D" box="[838,1020,1919,1941]" italics="true" pageId="6" pageNumber="507">Glossodoris buko</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B5ED57C0487EF01E091AF87BFB9DF86D" box="[1029,1113,1920,1941]" pageId="6" pageNumber="507" rank="species">
<emphasis id="AEDEEABB487EF01E091AF87BFB9DF86D" bold="true" box="[1029,1113,1920,1941]" pageId="6" pageNumber="507">sp. nov.</emphasis>
</taxonomicNameLabel>
, CASIZ-086381A Papua New Guinea, A. Rachis B. Mid-laterals. CASIZ-191102A Papua New Guinea, C. Outer laterals. D. Jaws. E. Radular ribbon. Right column (F–J) 
<taxonomicName id="5BAA4D2A487EF01E0C3BF847FE28F82A" baseAuthorityName="Cuvier" baseAuthorityYear="1804" box="[292,492,1980,2002]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="507" phylum="Mollusca" rank="species" species="pallida">
<emphasis id="AEDEEABB487EF01E0C3BF847FE28F82A" box="[292,492,1980,2002]" italics="true" pageId="6" pageNumber="507">Glossodoris pallida</emphasis>
</taxonomicName>
, CASIZ-173395 Madagascar: F. Rachis. G. Mid-laterals. H. Outer laterals. I. Jaws. J. Radular ribbon.
</paragraph>
</caption>
<caption id="C8D56621487FF01F0D88FBA6FCDAFB37" httpUri="https://zenodo.org/record/1437232/files/figure.png" pageId="7" pageNumber="508" startId="7.[151,250,1117,1139]" targetBox="[260,1353,211,1083]" targetPageId="7">
<paragraph id="9C1536A9487FF01F0D88FBA6FCDAFB37" blockId="7.[151,1437,1117,1231]" pageId="7" pageNumber="508">
<emphasis id="AEDEEABB487FF01F0D88FBA6FED3FB8B" bold="true" box="[151,279,1117,1139]" pageId="7" pageNumber="508">FIGURE 4.</emphasis>
<emphasis id="AEDEEABB487FF01F0C3EFBA6FE31FB8B" box="[289,501,1117,1139]" italics="true" pageId="7" pageNumber="508">
<taxonomicName id="5BAA4D2A487FF01F0C3EFBA6FE34FB8B" baseAuthorityName="Cuvier" baseAuthorityYear="1804" box="[289,496,1117,1139]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="508" phylum="Mollusca" rank="species" species="pallida">Glossodoris pallida</taxonomicName>
,
</emphasis>
CASIZ-173395: (A) Reproductive system (B) Buccal mass, ventral side has no glands. 
<taxonomicName id="5BAA4D2A487FF01F0D88FB87FE8EFB6A" authority="Matsuda &amp; Gosliner, 2018" authorityName="Matsuda &amp; Gosliner" authorityYear="2018" box="[151,330,1148,1170]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="508" phylum="Mollusca" rank="species" species="buko" status="sp. nov.">
<emphasis id="AEDEEABB487FF01F0D88FB87FE8EFB6A" box="[151,330,1148,1170]" italics="true" pageId="7" pageNumber="508">Glossodoris buko</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B5ED57C0487FF01F0C4CFB86FE61FB6A" box="[339,421,1149,1170]" pageId="7" pageNumber="508" rank="species">
<emphasis id="AEDEEABB487FF01F0C4CFB86FE61FB6A" bold="true" box="[339,421,1149,1170]" pageId="7" pageNumber="508">sp. nov.</emphasis>
</taxonomicNameLabel>
, CASIZ-086381A: (C) Reproductive system (D) Buccal mass, ventral side with glandular sheath. Abbreviations: p, penis; vd, vas deferens; pr, prostate; v, vaginal duct; rs, receptaculum seminis; bc, bursa copulatrix; am, ampulla; fgm, female gland mass; ot, oral tube, bb, buccal bulb.
</paragraph>
</caption>
<paragraph id="9C1536A9487FF01F0DD8FB07FD3EFA79" blockId="7.[151,1436,1276,1769]" pageId="7" pageNumber="508">
The reproductive system appears similar to the description by Rudman (1983) for 
<taxonomicName id="5BAA4D2A487FF01F0947FB06FB01FAEC" baseAuthorityName="Cuvier" baseAuthorityYear="1804" box="[1112,1221,1276,1300]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="508" phylum="Mollusca" rank="species" species="pallida">
<emphasis id="AEDEEABB487FF01F0947FB06FB01FAEC" box="[1112,1221,1276,1300]" italics="true" pageId="7" pageNumber="508">G. pallida</emphasis>
</taxonomicName>
from 
<collectingCountry id="E4BD7639487FF01F0817FB07FAA9FAED" box="[1288,1389,1276,1301]" name="Tanzania" pageId="7" pageNumber="508">Tanzania</collectingCountry>
and the 
<collectingRegion id="5E6EF84B487FF01F0DDDFADBFEDBFAC1" box="[194,287,1312,1337]" country="Sudan" name="Al Bahr al Ahmar" pageId="7" pageNumber="508">Red Sea</collectingRegion>
and examined here from 
<collectingCountry id="E4BD7639487FF01F0F58FADBFD0AFAC1" box="[583,718,1312,1337]" name="Madagascar" pageId="7" pageNumber="508">Madagascar</collectingCountry>
(
<figureCitation id="04912A2C487FF01F0FC0FADBFCFFFAC1" box="[735,827,1312,1337]" captionStart="FIGURE 4" captionStartId="7.[151,250,1117,1139]" captionTargetBox="[260,1353,211,1083]" captionTargetId="figure@7.[234,1353,193,1096]" captionTargetPageId="7" captionText="FIGURE 4. Glossodoris pallida, CASIZ-173395: (A) Reproductive system (B) Buccal mass, ventral side has no glands. Glossodoris buko sp. nov., CASIZ-086381A: (C) Reproductive system (D) Buccal mass, ventral side with glandular sheath. Abbreviations: p, penis; vd, vas deferens; pr, prostate; v, vaginal duct; rs, receptaculum seminis; bc, bursa copulatrix; am, ampulla; fgm, female gland mass; ot, oral tube, bb, buccal bulb." httpUri="https://zenodo.org/record/1437232/files/figure.png" pageId="7" pageNumber="508">Fig. 4A</figureCitation>
). The only noteworthy distinction in the reproductive systems of the two species is that the ejaculatory portion of the vas deferens of 
<taxonomicName id="5BAA4D2A487FF01F090CFABEFB46FAA4" baseAuthorityName="Cuvier" baseAuthorityYear="1804" box="[1043,1154,1348,1372]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="508" phylum="Mollusca" rank="species" species="pallida">
<emphasis id="AEDEEABB487FF01F090CFABEFB46FAA4" box="[1043,1154,1348,1372]" italics="true" pageId="7" pageNumber="508">G. pallida</emphasis>
</taxonomicName>
(
<figureCitation id="04912A2C487FF01F098EFABFFB2DFAA5" box="[1169,1257,1348,1373]" captionStart="FIGURE 4" captionStartId="7.[151,250,1117,1139]" captionTargetBox="[260,1353,211,1083]" captionTargetId="figure@7.[234,1353,193,1096]" captionTargetPageId="7" captionText="FIGURE 4. Glossodoris pallida, CASIZ-173395: (A) Reproductive system (B) Buccal mass, ventral side has no glands. Glossodoris buko sp. nov., CASIZ-086381A: (C) Reproductive system (D) Buccal mass, ventral side with glandular sheath. Abbreviations: p, penis; vd, vas deferens; pr, prostate; v, vaginal duct; rs, receptaculum seminis; bc, bursa copulatrix; am, ampulla; fgm, female gland mass; ot, oral tube, bb, buccal bulb." httpUri="https://zenodo.org/record/1437232/files/figure.png" pageId="7" pageNumber="508">Fig. 4A</figureCitation>
) contains many more convolutions than does that of 
<taxonomicName id="5BAA4D2A487FF01F0F2FFA92FD40FA78" authorityName="Matsuda &amp; Gosliner" authorityYear="2018" box="[560,644,1384,1408]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="508" phylum="Mollusca" rank="species" species="buko">
<emphasis id="AEDEEABB487FF01F0F2FFA92FD40FA78" box="[560,644,1384,1408]" italics="true" pageId="7" pageNumber="508">G. buko</emphasis>
</taxonomicName>
(
<figureCitation id="04912A2C487FF01F0F8BFA93FD29FA79" box="[660,749,1384,1409]" captionStart="FIGURE 4" captionStartId="7.[151,250,1117,1139]" captionTargetBox="[260,1353,211,1083]" captionTargetId="figure@7.[234,1353,193,1096]" captionTargetPageId="7" captionText="FIGURE 4. Glossodoris pallida, CASIZ-173395: (A) Reproductive system (B) Buccal mass, ventral side has no glands. Glossodoris buko sp. nov., CASIZ-086381A: (C) Reproductive system (D) Buccal mass, ventral side with glandular sheath. Abbreviations: p, penis; vd, vas deferens; pr, prostate; v, vaginal duct; rs, receptaculum seminis; bc, bursa copulatrix; am, ampulla; fgm, female gland mass; ot, oral tube, bb, buccal bulb." httpUri="https://zenodo.org/record/1437232/files/figure.png" pageId="7" pageNumber="508">Fig. 4C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="D4B06522487FF01F0DD8FA77FA88F911" pageId="7" pageNumber="508" type="reference_group">
<paragraph id="9C1536A9487FF01F0DD8FA77FA88F911" blockId="7.[151,1436,1276,1769]" pageId="7" pageNumber="508">
<taxonomicName id="5BAA4D2A487FF01F0DD8FA77FE56FA5C" ID-CoL="3GDJR" authorityName="Matsuda &amp; Gosliner" authorityYear="2018" box="[199,402,1420,1444]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="508" phylum="Mollusca" rank="species" species="buko">
<emphasis id="AEDEEABB487FF01F0DD8FA77FE56FA5C" box="[199,402,1420,1444]" italics="true" pageId="7" pageNumber="508">Glossodoris buko</emphasis>
</taxonomicName>
is distinct from 
<taxonomicName id="5BAA4D2A487FF01F0F7DFA76FD13FA5C" ID-CoL="3GDLJ" baseAuthorityName="Cuvier" baseAuthorityYear="1804" box="[610,727,1420,1444]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="508" phylum="Mollusca" rank="species" species="pallida">
<emphasis id="AEDEEABB487FF01F0F7DFA76FD13FA5C" box="[610,727,1420,1444]" italics="true" pageId="7" pageNumber="508">G. pallida</emphasis>
</taxonomicName>
in both internal morphology as shown here, and based on molecular analyses (
<bibRefCitation id="F83B4B58487FF01F0C9EFA4BFD70FA31" box="[385,692,1456,1481]" pageId="7" pageNumber="508" refId="ref16546" refString="Matsuda, S. B. &amp; Gosliner, T. M. (2017) Molecular phylogeny of Glossodoris (Ehrenberg, 1831) nudibranchs and related genera reveals cryptic and pseudocryptic species complexes. Cladistics, 34 (1), 41 - 56. https: // doi. org / 10.1111 / cla. 12194" type="journal article">Matsuda &amp; Gosliner 2017</bibRefCitation>
). This distinction is also maintained geographically. 
<taxonomicName id="5BAA4D2A487FF01F080BFA4BFF22FA14" ID-CoL="3GDLJ" baseAuthorityName="Cuvier" baseAuthorityYear="1804" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="508" phylum="Mollusca" rank="species" species="pallida">
<emphasis id="AEDEEABB487FF01F080BFA4BFF22FA14" italics="true" pageId="7" pageNumber="508">Glossodoris pallida</emphasis>
</taxonomicName>
has only been recorded off the coast of eastern Africa and the 
<collectingRegion id="5E6EF84B487FF01F0ED4FA2FFBE9FA15" box="[971,1069,1492,1517]" country="Sudan" name="Al Bahr al Ahmar" pageId="7" pageNumber="508">Red Sea</collectingRegion>
, and 
<taxonomicName id="5BAA4D2A487FF01F0972FA2EFB01FA14" ID-CoL="3GDJR" authorityName="Matsuda &amp; Gosliner" authorityYear="2018" box="[1133,1221,1492,1516]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="508" phylum="Mollusca" rank="species" species="buko">
<emphasis id="AEDEEABB487FF01F0972FA2EFB01FA14" box="[1133,1221,1492,1516]" italics="true" pageId="7" pageNumber="508">G. buko</emphasis>
</taxonomicName>
is solely from the western Pacific. 
<bibRefCitation id="F83B4B58487FF01F0C4BFA03FD67F9E9" box="[340,675,1528,1553]" pageId="7" pageNumber="508" refId="ref16546" refString="Matsuda, S. B. &amp; Gosliner, T. M. (2017) Molecular phylogeny of Glossodoris (Ehrenberg, 1831) nudibranchs and related genera reveals cryptic and pseudocryptic species complexes. Cladistics, 34 (1), 41 - 56. https: // doi. org / 10.1111 / cla. 12194" type="journal article">Matsuda &amp; Gosliner’s (2017)</bibRefCitation>
phylogeny of 
<taxonomicName id="5BAA4D2A487FF01F0E54FA03FC16F9E8" ID-CoL="7P3T9" authorityName="Ehrenberg" authorityYear="1831" box="[843,978,1528,1552]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="508" phylum="Mollusca" rank="genus">
<emphasis id="AEDEEABB487FF01F0E54FA03FC16F9E8" box="[843,978,1528,1552]" italics="true" pageId="7" pageNumber="508">Glossodoris</emphasis>
</taxonomicName>
provides support for the splitting of the previously hypothesized 
<taxonomicName id="5BAA4D2A487FF01F0CB0F9E7FD48F9CC" ID-CoL="3GDLJ" baseAuthorityName="Cuvier" baseAuthorityYear="1804" box="[431,652,1564,1588]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="508" phylum="Mollusca" rank="species" species="pallida">
<emphasis id="AEDEEABB487FF01F0CB0F9E7FD48F9CC" box="[431,652,1564,1588]" italics="true" pageId="7" pageNumber="508">Glossodoris pallida</emphasis>
</taxonomicName>
into two distinct species. This is further supported through p-distance values (
<bibRefCitation id="F83B4B58487FF01F0DEDF9BBFDE9F9A1" box="[242,557,1600,1625]" pageId="7" pageNumber="508" refId="ref16546" refString="Matsuda, S. B. &amp; Gosliner, T. M. (2017) Molecular phylogeny of Glossodoris (Ehrenberg, 1831) nudibranchs and related genera reveals cryptic and pseudocryptic species complexes. Cladistics, 34 (1), 41 - 56. https: // doi. org / 10.1111 / cla. 12194" type="journal article">Matsuda &amp; Gosliner 2017</bibRefCitation>
). Within 
<taxonomicName id="5BAA4D2A487FF01F0F81F9BAFD3CF9A0" ID-CoL="3GDJR" authorityName="Matsuda &amp; Gosliner" authorityYear="2018" box="[670,760,1600,1624]" class="Gastropoda" family="Chromodorididae" genus="Glossodoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="7" pageNumber="508" phylum="Mollusca" rank="species" species="buko">
<emphasis id="AEDEEABB487FF01F0F81F9BAFD3CF9A0" box="[670,760,1600,1624]" italics="true" pageId="7" pageNumber="508">G. buko</emphasis>
</taxonomicName>
, a grade and one clade are supported (a grade from the 
<collectingCountry id="E4BD7639487FF01F0D88F99FFED1F985" box="[151,277,1636,1661]" name="Philippines" pageId="7" pageNumber="508">Philippines</collectingCountry>
and a clade from 
<collectingCountry id="E4BD7639487FF01F0CFCF99FFD8EF985" box="[483,586,1636,1661]" name="Australia" pageId="7" pageNumber="508">Australia</collectingCountry>
and 
<collectingCountry id="E4BD7639487FF01F0F9DF99EFCA6F985" box="[642,866,1636,1661]" name="Papua New Guinea" pageId="7" pageNumber="508">Papua New Guinea</collectingCountry>
). However, there were no observed morphological differences and only a 5–6% p-distance between them and these were not recovered as distinct lineages in the ABGD analysis conducted by Matsuda and Gosliner (
<figureCitation id="04912A2C487FF01F0E0BF957FC99F93D" box="[788,861,1708,1733]" captionStart="FIGURE 5" captionStartId="9.[151,250,1981,2003]" captionTargetBox="[197,1432,287,1931]" captionTargetId="figure@9.[156,1433,263,1960]" captionTargetPageId="9" captionText="FIGURE 5. Phylogenetic tree (Matsuda &amp; Gosliner 2017) with ABGD partitions (colored bars) and p-distances. Values listed are Bayesian Inference posterior probabilities/Maximum Likelihood bootstraps/Maximum Parsimony bootstraps." httpUri="https://zenodo.org/record/1437234/files/figure.png" pageId="7" pageNumber="508">Fig. 5</figureCitation>
). This strongly supports that the western Pacific specimens represent a single species distinct from the Indian Ocean specimens (
<bibRefCitation id="F83B4B58487FF01F0909F92BFA84F911" box="[1046,1344,1744,1769]" pageId="7" pageNumber="508" refId="ref16546" refString="Matsuda, S. B. &amp; Gosliner, T. M. (2017) Molecular phylogeny of Glossodoris (Ehrenberg, 1831) nudibranchs and related genera reveals cryptic and pseudocryptic species complexes. Cladistics, 34 (1), 41 - 56. https: // doi. org / 10.1111 / cla. 12194" type="journal article">Matsuda &amp; Gosliner 2017</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>