172 lines
19 KiB
XML
172 lines
19 KiB
XML
<document id="3809AB56233C931B4317D7DD6FC81099" ID-DOI="10.5281/zenodo.186952" ID-GBIF-Dataset="4239119c-102b-417f-b91e-619957e061c1" ID-ISSN="1175-5326" ID-Zenodo-Dep="186952" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461059822508" checkinUser="plazi" docAuthor="Levy, Gershom" docDate="2009" docId="0E2987C979370C253DF6FA5AFC80FE28" docLanguage="en" docName="zt02066p049.pdf" docOrigin="Zootaxa 2066" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Leptopilos Levy, 2009, n. gen." docType="treatment" docVersion="11" lastPageNumber="8" masterDocId="F210FFB179300C2D3D61FFB2FFA7FFE8" masterDocTitle="New ground-spider genera and species with annexed checklist of the Gnaphosidae (Araneae) of Israel" masterLastPageNumber="49" masterPageNumber="1" pageNumber="8" updateTime="1698581975113" updateUser="plazi">
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<mods:title id="5E9457A4D23753CE405EA895FA86FC0D">New ground-spider genera and species with annexed checklist of the Gnaphosidae (Araneae) of Israel</mods:title>
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<mods:namePart id="F66BDDA6CAB8F25C81CC07FA8BEF67ED">Levy, Gershom</mods:namePart>
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<mods:title id="F3D6738827944B94B204FEBB83D657D6">Zootaxa</mods:title>
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<mods:part id="69F3648CAD12D210DAE46643DD7B55C4">
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<mods:date id="54829A597B326263D751682927BDC505">2009</mods:date>
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<mods:number id="7FF5DCD3D58D1BDB7BC018BCCEF7662E">2066</mods:number>
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<mods:identifier id="7E85A14F2AF800166886C6901D93B5E8" type="DOI">10.5281/zenodo.186952</mods:identifier>
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<treatment id="0E2987C979370C253DF6FA5AFC80FE28" ID-DOI="http://doi.org/10.5281/zenodo.6221977" ID-GBIF-Taxon="119613850" ID-Zenodo-Dep="6221977" LSID="urn:lsid:plazi:treatment:0E2987C979370C253DF6FA5AFC80FE28" httpUri="http://treatment.plazi.org/id/0E2987C979370C253DF6FA5AFC80FE28" lastPageId="8" lastPageNumber="8" pageId="7" pageNumber="8">
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<subSubSection id="CE9A655479370C2A3DF6FA5AFD5BF9A0" pageId="7" pageNumber="8" type="nomenclature">
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<paragraph id="863F36DF79370C2A3DF6FA5AFECAF9EA" blockId="7.[151,365,1512,1538]" box="[151,365,1512,1538]" pageId="7" pageNumber="8">
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<heading id="DD7781B379370C2A3DF6FA5AFECAF9EA" bold="true" box="[151,365,1512,1538]" fontSize="11" level="1" pageId="7" pageNumber="8" reason="1">
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<emphasis id="B4F4EACD79370C2A3DF6FA5AFECAF9EA" bold="true" box="[151,365,1512,1538]" pageId="7" pageNumber="8">
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<taxonomicName id="41804D5C79370C2A3DF6FA5AFEB3F9EA" ID-CoL="5D9V" box="[151,276,1512,1538]" class="Arachnida" family="Gnaphosidae" genus="Leptopilos" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B4F4EACD79370C2A3DF6FA5AFEB3F9EA" bold="true" box="[151,276,1512,1538]" italics="true" pageId="7" pageNumber="8">Leptopilos</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="AFC757B679370C2A3C7DFA5AFECAF9EA" box="[284,365,1512,1538]" pageId="7" pageNumber="8" rank="genus">n. gen.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="863F36DF79370C2A3DF6F983FD5BF9A0" blockId="7.[151,764,1585,1609]" box="[151,764,1585,1609]" pageId="7" pageNumber="8">
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<emphasis id="B4F4EACD79370C2A3DF6F983FE88F9A1" bold="true" box="[151,303,1585,1609]" pageId="7" pageNumber="8">Type-species:</emphasis>
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<taxonomicName id="41804D5C79370C2A3C54F980FDA0F9A1" box="[309,519,1586,1609]" class="Arachnida" family="Gnaphosidae" genus="Drassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="tenerrimus">
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<emphasis id="B4F4EACD79370C2A3C54F980FDA0F9A1" box="[309,519,1586,1609]" italics="true" pageId="7" pageNumber="8">Drassus tenerrimus</emphasis>
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</taxonomicName>
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O.P.-
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<bibRefCitation id="E2114B2E79370C2A3F5FF983FD5BF9A0" author="Cambridge" box="[574,764,1585,1608]" pageId="7" pageNumber="48" refString="Cambridge, O. P. -. (1872) General list of the spiders of Palestine and Syria. Proceedings of the Zoological Society of London, 1872, 212 - 354." type="proceedings paper" year="1872">Cambridge, 1872</bibRefCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="CE9A655479370C2A3DF6F9CAFBA8F951" pageId="7" pageNumber="8" type="etymology">
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<paragraph id="863F36DF79370C2A3DF6F9CAFBA8F951" blockId="7.[151,1437,1656,2030]" pageId="7" pageNumber="8">
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<emphasis id="B4F4EACD79370C2A3DF6F9CAFEBBF97A" bold="true" box="[151,284,1656,1682]" pageId="7" pageNumber="8">Etymology</emphasis>
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.The generic name is a combination derived from
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<taxonomicName id="41804D5C79370C2A3E01F9CBFC5BF97A" box="[864,1020,1657,1682]" class="Arachnida" family="Gnaphosidae" genus="Leptodrassus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
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<emphasis id="B4F4EACD79370C2A3E01F9CBFC5BF97A" box="[864,1020,1657,1682]" italics="true" pageId="7" pageNumber="8">Leptodrassus</emphasis>
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</taxonomicName>
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and
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<emphasis id="B4F4EACD79370C2A3955F9CBFBCAF97A" box="[1076,1133,1657,1682]" italics="true" pageId="7" pageNumber="8">pilos</emphasis>
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, a cap in Greek, a marked diagnostic sclerite on the female
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<taxonomicName id="41804D5C79370C2A3F78F92DFD36F951" box="[537,657,1695,1721]" class="Magnoliopsida" family="Apocynaceae" genus="Epigynum" kingdom="Plantae" order="Gentianales" pageId="7" pageNumber="41" phylum="Tracheophyta" rank="genus">epigynum</taxonomicName>
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, characteristic of the new genus.
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</paragraph>
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</subSubSection>
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<subSubSection id="CE9A655479370C2A3DA7F977FDB9F849" pageId="7" pageNumber="8" type="diagnosis">
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<paragraph id="863F36DF79370C2A3DA7F977FDB9F849" blockId="7.[151,1437,1656,2030]" pageId="7" pageNumber="8">
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<emphasis id="B4F4EACD79370C2A3DA7F977FE98F937" bold="true" box="[198,319,1733,1759]" pageId="7" pageNumber="8">Diagnosis</emphasis>
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. Small gnaphosids with colouration, body shape and general eye configuration as in
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<taxonomicName id="41804D5C79370C2A3DF6F95EFE9DF8ED" box="[151,314,1772,1797]" class="Arachnida" family="Gnaphosidae" genus="Leptodrassus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
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<emphasis id="B4F4EACD79370C2A3DF6F95EFE9DF8ED" box="[151,314,1772,1797]" italics="true" pageId="7" pageNumber="8">Leptodrassus</emphasis>
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</taxonomicName>
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. AME, the largest eyes, are borne on a common black patch. Male without dorsal scutum. Chelicerae with small conventional teeth, unlike the peculiar, angular teeth of
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<taxonomicName id="41804D5C79370C2A3925F8A1FB44F8C4" box="[1092,1251,1811,1836]" class="Arachnida" family="Gnaphosidae" genus="Leptodrassus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
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<emphasis id="B4F4EACD79370C2A3925F8A1FB44F8C4" box="[1092,1251,1811,1836]" italics="true" pageId="7" pageNumber="8">Leptodrassus</emphasis>
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</taxonomicName>
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. Labium wider than long. Palpus of male apically armed with closely grouped, often pointed laminae and a single-lobed tibial retrolateral apophysis. Female
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<taxonomicName id="41804D5C79370C2A3F63F8D2FDD0F892" box="[514,631,1888,1914]" class="Magnoliopsida" family="Apocynaceae" genus="Epigynum" kingdom="Plantae" order="Gentianales" pageId="7" pageNumber="41" phylum="Tracheophyta" rank="genus">epigynum</taxonomicName>
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with a median cavity and a broad, prominent cap-like anterior hood unlike the scape of
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<taxonomicName id="41804D5C79370C2A3C1AF835FDB0F848" box="[379,535,1927,1952]" class="Arachnida" family="Gnaphosidae" genus="Leptodrassus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
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<emphasis id="B4F4EACD79370C2A3C1AF835FDB0F848" box="[379,535,1927,1952]" italics="true" pageId="7" pageNumber="8">Leptodrassus</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="CE9A655479370C253DA7F81FFC80FE28" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="discussion">
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<paragraph id="863F36DF79370C253DA7F81FFC80FE28" blockId="7.[151,1437,1656,2030]" lastBlockId="8.[151,1436,152,448]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
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<emphasis id="B4F4EACD79370C2A3DA7F81FFEEAF82F" bold="true" box="[198,333,1965,1991]" pageId="7" pageNumber="8">Comments</emphasis>
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. The former genus
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<taxonomicName id="41804D5C79370C2A3F56F81CFD74F82F" box="[567,723,1966,1991]" class="Arachnida" family="Gnaphosidae" genus="Leptodrassus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
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<emphasis id="B4F4EACD79370C2A3F56F81CFD74F82F" box="[567,723,1966,1991]" italics="true" pageId="7" pageNumber="8">Leptodrassus</emphasis>
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</taxonomicName>
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has been recently divided by having the cheliceral dentition into two groups (
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<bibRefCitation id="E2114B2E79370C2A3C3FF866FDABF806" author="Murphy" box="[350,524,2004,2030]" pageId="7" pageNumber="49" refString="Murphy, J. A. (2007) Gnaphosid genera of the world. British Arachnological Society, St Neots, Cambs, 2 Volumes, 1 - 605." type="journal article" year="2007">Murphy, 2007</bibRefCitation>
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).
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<taxonomicName id="41804D5C79370C2A3F7EF866FD1CF805" box="[543,699,2004,2029]" class="Arachnida" family="Gnaphosidae" genus="Leptodrassus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
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<emphasis id="B4F4EACD79370C2A3F7EF866FD1CF805" box="[543,699,2004,2029]" italics="true" pageId="7" pageNumber="8">Leptodrassus</emphasis>
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</taxonomicName>
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includes those species which like the type-species
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<taxonomicName id="41804D5C79370C2A3875F866FA3BF805" box="[1300,1436,2004,2029]" class="Arachnida" family="Gnaphosidae" genus="Leptodrassus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="femineus">
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<emphasis id="B4F4EACD79370C2A3875F866FA3BF805" box="[1300,1436,2004,2029]" italics="true" pageId="7" pageNumber="8">L. femineus</emphasis>
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</taxonomicName>
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have two large angular retromarginal teeth, while the recently described genus
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<taxonomicName id="41804D5C79380C253958FF2AFA9CFF5A" authority="Murphy" authorityName="Murphy" box="[1081,1339,152,178]" class="Arachnida" family="Gnaphosidae" genus="Leptodrassex" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
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<emphasis id="B4F4EACD79380C253958FF2AFB72FF59" box="[1081,1237,152,177]" italics="true" pageId="8" pageNumber="9">Leptodrassex</emphasis>
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Murphy
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</taxonomicName>
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, with
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<taxonomicName id="41804D5C79380C2538E5FF2AFF4FFF30" class="Arachnida" family="Gnaphosidae" genus="Leptodrassex" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="simoni">
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<emphasis id="B4F4EACD79380C2538E5FF2AFF4FFF30" italics="true" pageId="8" pageNumber="9">L. simoni</emphasis>
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</taxonomicName>
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as type-species, possesses small, discrete conventional teeth. However, among the latter, a definite group of species is distinguished by their genital characters, namely a peculiar cap-like sclerite on the female
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<taxonomicName id="41804D5C79380C253DF6FEBEFEA8FECE" box="[151,271,268,294]" class="Magnoliopsida" family="Apocynaceae" genus="Epigynum" kingdom="Plantae" order="Gentianales" pageId="8" pageNumber="41" phylum="Tracheophyta" rank="genus">epigynum</taxonomicName>
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, and closely grouped laminae on the male palpus, unlike
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<taxonomicName id="41804D5C79380C253ED4FEBEFB84FECD" box="[949,1059,268,293]" class="Arachnida" family="Gnaphosidae" genus="Leptodrassex" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="simoni">
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<emphasis id="B4F4EACD79380C253ED4FEBEFB84FECD" box="[949,1059,268,293]" italics="true" pageId="8" pageNumber="9">L. simoni</emphasis>
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</taxonomicName>
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(
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<bibRefCitation id="E2114B2E79380C253955FEBEFAFDFECE" author="Levy" box="[1076,1370,268,294]" pageId="8" pageNumber="49" refString="Levy, G. (1999 b) Spiders of six uncommon drassodine genera (Araneae: Gnaphosidae) from Israel. Israel Journal of Zoology, 45, 427 - 452." type="journal article" year="1999" yearSuffix="b">Levy, 1999b, figs 35–37</bibRefCitation>
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). The females within
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<taxonomicName id="41804D5C79380C253C2FFE80FE6DFEA3" box="[334,458,306,331]" class="Arachnida" family="Gnaphosidae" genus="Leptopilos" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
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<emphasis id="B4F4EACD79380C253C2FFE80FE6DFEA3" box="[334,458,306,331]" italics="true" pageId="8" pageNumber="9">Leptopilos</emphasis>
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</taxonomicName>
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are easily separated from each other by the different shapes of the epigynal hood. Apart from the type-species
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<taxonomicName id="41804D5C79380C253C9DFEEBFD03FE9A" box="[508,676,345,370]" class="Arachnida" family="Gnaphosidae" genus="Drassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="tenerrimus">
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<emphasis id="B4F4EACD79380C253C9DFEEBFD03FE9A" box="[508,676,345,370]" italics="true" pageId="8" pageNumber="9">L. tenerrimus</emphasis>
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</taxonomicName>
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the following species are here placed in
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<taxonomicName id="41804D5C79380C2539C9FEEBFA8CFE9A" box="[1192,1323,345,370]" class="Arachnida" family="Gnaphosidae" genus="Leptopilos" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
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<emphasis id="B4F4EACD79380C2539C9FEEBFA8CFE9A" box="[1192,1323,345,370]" italics="true" pageId="8" pageNumber="9">Leptopilos</emphasis>
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</taxonomicName>
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:
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<taxonomicName id="41804D5C79380C25385CFEEBFE9EFE72" authority="Dalmas, 1919" authorityName="Dalmas" authorityYear="1919" class="Arachnida" family="Gnaphosidae" genus="Leptopilos" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="pupa">
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<emphasis id="B4F4EACD79380C25385CFEEBFA3BFE9A" box="[1341,1436,345,370]" italics="true" pageId="8" pageNumber="9">L. pupa</emphasis>
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<bibRefCitation id="E2114B2E79380C253DF6FE32FE9EFE72" author="Dalmas" box="[151,313,384,410]" pageId="8" pageNumber="48" refString="Dalmas, R. de. (1919) Catalogue des araignees du genre Leptodrassus (Gnaphosidae), d'apres les materiaux de la collection E. Simon au Museum d'histoire naturelle. Bulletin du Museum d'histoire naturelle, Paris, (4), 1919, 243 - 250." type="journal article" year="1919">Dalmas, 1919</bibRefCitation>
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</taxonomicName>
|
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(not
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<bibRefCitation id="E2114B2E79380C253C17FE32FDD1FE72" author="Chatzaki" box="[374,630,384,410]" pageId="8" pageNumber="48" refString="Chatzaki, M., Thaler, K. & Mylonas, M. (2002 a) Ground spiders (Gnaphosidae; Araneae) of Crete (Greece). Taxonomy and distribution I. Revue Suisse Zoology, 109 (3), 559 - 601." type="journal article" year="2002" yearSuffix="a">Chatzaki et al., 2002a</bibRefCitation>
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||
,
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<bibRefCitation id="E2114B2E79380C253F1BFE32FD2EFE72" author="Chatzaki" box="[634,649,384,410]" pageId="8" pageNumber="48" refString="Chatzaki, M., Thaler, K. & Mylonas, M. (2002 b) Ground spiders (Gnaphosidae; Araneae) of Crete and adjacent areas of Greece. Taxonomy and distribution II. Revue Suisse de Zoology, 109 (3), 603 - 633." type="journal article" year="2002" yearSuffix="b">b</bibRefCitation>
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=
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<taxonomicName id="41804D5C79380C253FC7FE32FC99FE71" box="[678,830,384,409]" class="Arachnida" family="Gnaphosidae" genus="Leptopilos" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="levantinus" status="sp. nov.">
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<emphasis id="B4F4EACD79380C253FC7FE32FC99FE71" box="[678,830,384,409]" italics="true" pageId="8" pageNumber="9">L. levantinus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="AFC757B679380C253E25FE32FC24FE72" box="[836,899,384,410]" pageId="8" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
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),
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<taxonomicName id="41804D5C79380C253EFBFE32FAA9FE72" authority="Chatzaki, 2002" authorityName="Chatzaki" authorityYear="2002" box="[922,1294,384,410]" class="Arachnida" family="Gnaphosidae" genus="Leptopilos" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="hadjissaranti">
|
||
<emphasis id="B4F4EACD79380C253EFBFE32FBF5FE71" box="[922,1106,384,409]" italics="true" pageId="8" pageNumber="9">L. hadjissaranti</emphasis>
|
||
Chatzaki, 2002
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="41804D5C79380C253878FE32FEECFE28" authority="Chatzaki, 2002" authorityName="Chatzaki" authorityYear="2002" class="Arachnida" family="Gnaphosidae" genus="Leptopilos" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="manolisi">
|
||
<emphasis id="B4F4EACD79380C253878FE32FA3BFE71" box="[1305,1436,384,409]" italics="true" pageId="8" pageNumber="9">L. manolisi</emphasis>
|
||
Chatzaki, 2002
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="41804D5C79380C253C36FE15FE49FE28" box="[343,494,423,448]" class="Arachnida" family="Gnaphosidae" genus="Leptopilos" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="levantinus" status="sp. nov.">
|
||
<emphasis id="B4F4EACD79380C253C36FE15FE49FE28" box="[343,494,423,448]" italics="true" pageId="8" pageNumber="9">L. levantinus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="AFC757B679380C253C94FE14FD93FE28" box="[501,564,422,448]" pageId="8" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="41804D5C79380C253F0CFE15FD46FE28" box="[621,737,423,448]" class="Arachnida" family="Gnaphosidae" genus="Leptopilos" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="lakhish" status="sp. nov.">
|
||
<emphasis id="B4F4EACD79380C253F0CFE15FD46FE28" box="[621,737,423,448]" italics="true" pageId="8" pageNumber="9">L. lakhish</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="AFC757B679380C253F89FE14FC80FE28" box="[744,807,422,448]" pageId="8" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |