284 lines
34 KiB
XML
284 lines
34 KiB
XML
<document id="F87CFAA540D305C6A482E30596FD42D9" ID-DOI="10.5281/zenodo.173690" ID-GBIF-Dataset="cc2aee04-de27-4429-a003-6f5549e36675" ID-ISSN="11755326" ID-Zenodo-Dep="173690" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459782831777" checkinUser="plazi" docAuthor="Framenau, Volker W." docDate="2006" docId="D1708793FFE1D32B0A6CB47B0A6FFAF0" docLanguage="en" docName="zt01304p020.pdf" docOrigin="Zootaxa 1304" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Anomalosa Roewer 1960" docType="treatment" docVersion="9" lastPageNumber="5" masterDocId="2D49FFEBFFE2D32F0B64B7570959FF9E" masterDocTitle="Revision of the Australian wolf spider genus Anomalosa Roewer, 1960 (Araneae: Lycosidae)" masterLastPageNumber="20" masterPageNumber="1" pageNumber="4" updateTime="1698218029850" updateUser="plazi">
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<mods:title id="BD4C74DD13385E0345F4B6B8F93775E6">Revision of the Australian wolf spider genus Anomalosa Roewer, 1960 (Araneae: Lycosidae)</mods:title>
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<mods:name id="1599CE9D20C7344A9750BF184072F9D1" type="personal">
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<mods:roleTerm id="9B198A5225451B4EB38B942589DDA83A">Author</mods:roleTerm>
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<mods:namePart id="B34E9CF5065F0B93DE051D47C9F23FF0">Framenau, Volker W.</mods:namePart>
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<mods:title id="680A161FEA33879CA74C6A57E637D502">Zootaxa</mods:title>
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<mods:part id="84B9A3ACF45279C9C4BDEACCFFDE4517">
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<mods:date id="1AC816472E750B9C8CF9B14176144AF9">2006</mods:date>
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<mods:detail id="F02D02718B7FD36CA5F660EEBC401C57" type="volume">
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<mods:number id="CC316DA264DCC9AFAF2075B79A079118">1304</mods:number>
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<mods:identifier id="84251F56D4AB054E692A2180B7DD5A44" type="DOI">10.5281/zenodo.173690</mods:identifier>
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<treatment id="D1708793FFE1D32B0A6CB47B0A6FFAF0" ID-DOI="http://doi.org/10.5281/zenodo.6262735" ID-GBIF-Taxon="119348431" ID-Zenodo-Dep="6262735" LSID="urn:lsid:plazi:treatment:D1708793FFE1D32B0A6CB47B0A6FFAF0" httpUri="http://treatment.plazi.org/id/D1708793FFE1D32B0A6CB47B0A6FFAF0" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
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<subSubSection id="11C3650EFFE1D32C0A6CB47B0BCFFCD8" box="[264,662,812,838]" pageId="3" pageNumber="4" type="nomenclature">
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<paragraph id="59663685FFE1D32C0A6CB47B0BCFFCD8" blockId="3.[264,662,812,838]" box="[264,662,812,838]" pageId="3" pageNumber="4">
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<heading id="022E81E9FFE1D32C0A6CB47B0BCFFCD8" bold="true" box="[264,662,812,838]" fontSize="11" level="1" pageId="3" pageNumber="4" reason="1">
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<emphasis id="6BADEA97FFE1D32C0A6CB47B0BCFFCD8" bold="true" box="[264,662,812,838]" pageId="3" pageNumber="4">
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Genus
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<taxonomicName id="9ED94D06FFE1D32C0A3BB47B0BCFFCD8" ID-CoL="WSY" ID-ENA="332768" authority="Roewer, 1960" authorityName="Roewer" authorityYear="1960" box="[351,662,812,838]" class="Arachnida" family="Lycosidae" genus="Anomalosa" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="6BADEA97FFE1D32C0A3BB47B08B1FCD8" bold="true" box="[351,488,812,838]" italics="true" pageId="3" pageNumber="4">Anomalosa</emphasis>
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<bibRefCitation id="3D484B74FFE1D32C0A8BB47B0BCFFCD8" author="Roewer" box="[495,662,812,838]" pageId="3" pageNumber="19" refString="Roewer, C. F. (1960) [imprint date 1959]. Araneae Lycosiformae II (Lycosidae) (Fortsetzung und Schluss). Exploration du Parc National de l'Upemba - Mission GF de Witte, 55, 519 - 1040." type="journal article" year="1960">Roewer, 1960</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="11C3650EFFE1D32C0A6CB4200A20FC13" box="[264,889,886,910]" pageId="3" pageNumber="4" type="reference_group">
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||
<paragraph id="59663685FFE1D32C0A6CB4200A20FC13" blockId="3.[264,889,886,910]" box="[264,889,886,910]" pageId="3" pageNumber="4">
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<treatmentCitationGroup id="79C911ABFFE1D32C0A6CB4200A20FC13" box="[264,889,886,910]" pageId="3" pageNumber="4">
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<taxonomicName id="9ED94D06FFE1D32C0A6CB4200B16FC13" ID-CoL="WSY" authority="Roewer 1960: 977" authorityName="Roewer" authorityPageNumber="977" authorityYear="1960" box="[264,591,886,910]" class="Arachnida" family="Lycosidae" genus="Anomalosa" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="6BADEA97FFE1D32C0A6CB42008D9FC10" box="[264,384,887,910]" italics="true" pageId="3" pageNumber="4">Anomalosa</emphasis>
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<treatmentCitation id="D8781094FFE1D32C0AE3B4210B16FC13" author="Roewer" box="[391,591,886,909]" page="977" pageId="3" pageNumber="4" year="1960">
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<bibRefCitation id="3D484B74FFE1D32C0AE3B4210B40FC13" author="Roewer" box="[391,537,886,909]" pageId="3" pageNumber="19" refString="Roewer, C. F. (1960) [imprint date 1959]. Araneae Lycosiformae II (Lycosidae) (Fortsetzung und Schluss). Exploration du Parc National de l'Upemba - Mission GF de Witte, 55, 519 - 1040." type="journal article" year="1960">Roewer 1960</bibRefCitation>
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: 977
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</treatmentCitation>
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</taxonomicName>
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.
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<treatmentCitation id="D8781094FFE1D32C093DB4210A2CFC13" author="Lehtinen" box="[601,885,886,909]" page="7" pageId="3" pageNumber="4" year="1979">
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<bibRefCitation id="3D484B74FFE1D32C093DB4210A03FC13" author="Lehtinen" box="[601,858,886,909]" pageId="3" pageNumber="19" refString="Lehtinen, P. T. & Hippa, H. (1979) Spiders of the Oriental-Australian region. I. Lycosidae: Venoniinae and Zoicinae. Annales Zoologici Fennici, 16, 1 - 22." type="journal article" year="1979">Lehtinen & Hippa 1979</bibRefCitation>
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: 7
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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||
</paragraph>
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</subSubSection>
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<subSubSection id="11C3650EFFE1D32C0A6CB4E90D3BFC47" box="[264,1122,958,985]" pageId="3" pageNumber="4" type="materials_examined">
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||
<paragraph id="59663685FFE1D32C0A6CB4E90D3BFC47" blockId="3.[264,1323,958,1905]" box="[264,1122,958,985]" pageId="3" pageNumber="4">
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||
<emphasis id="6BADEA97FFE1D32C0A6CB4E908F3FC46" bold="true" box="[264,426,958,984]" pageId="3" pageNumber="4">
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<typeStatus id="86628827FFE1D32C0A6CB4E9081CFC46" box="[264,325,958,984]" pageId="3" pageNumber="4">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="9ED94D06FFE1D32C0AD6B4E80A65FC47" authority="Simon, 1898" authorityName="Simon" authorityYear="1898" box="[434,828,959,985]" class="Arachnida" family="Lycosidae" genus="Anomalomma" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="kochi">
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<emphasis id="6BADEA97FFE1D32C0AD6B4E80BC2FC46" box="[434,667,959,984]" italics="true" pageId="3" pageNumber="4">Anomalomma kochi</emphasis>
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Simon, 1898
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</taxonomicName>
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. By original designation.
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</paragraph>
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</subSubSection>
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<subSubSection id="11C3650EFFE1D32C0A5CB4B108FAFA7F" pageId="3" pageNumber="4" type="diagnosis">
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<paragraph id="59663685FFE1D32C0A5CB4B108FAFA7F" blockId="3.[264,1323,958,1905]" pageId="3" pageNumber="4">
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<emphasis id="6BADEA97FFE1D32C0A5CB4B108EEFB9E" bold="true" box="[312,439,998,1024]" pageId="3" pageNumber="4">Diagnosis.</emphasis>
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Within the Venoniinae
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<emphasis id="6BADEA97FFE1D32C09B9B4B00A71FB9E" box="[733,808,999,1024]" italics="true" pageId="3" pageNumber="4">(sensu</emphasis>
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<bibRefCitation id="3D484B74FFE1D32C0850B4B00D02FB9F" author="Lehtinen" box="[820,1115,999,1025]" pageId="3" pageNumber="19" refString="Lehtinen, P. T. & Hippa, H. (1979) Spiders of the Oriental-Australian region. I. Lycosidae: Venoniinae and Zoicinae. Annales Zoologici Fennici, 16, 1 - 22." type="journal article" year="1979">Lehtinen & Hippa 1979</bibRefCitation>
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),
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<taxonomicName id="9ED94D06FFE1D32C0F10B4B00DA1FB9E" box="[1140,1272,999,1024]" class="Arachnida" family="Lycosidae" genus="Anomalosa" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="6BADEA97FFE1D32C0F10B4B00DA1FB9E" box="[1140,1272,999,1024]" italics="true" pageId="3" pageNumber="4">Anomalosa</emphasis>
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</taxonomicName>
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has somatic and genitalic characters similar to
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<taxonomicName id="9ED94D06FFE1D32C0853B3580ACCFBB6" box="[823,917,1039,1064]" class="Arachnida" family="Lycosidae" genus="Venonia" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="6BADEA97FFE1D32C0853B3580ACCFBB6" box="[823,917,1039,1064]" italics="true" pageId="3" pageNumber="4">Venonia</emphasis>
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</taxonomicName>
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. It differs from
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<taxonomicName id="9ED94D06FFE1D32C0F1CB3580D8EFBB6" box="[1144,1239,1039,1064]" class="Arachnida" family="Lycosidae" genus="Venonia" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="6BADEA97FFE1D32C0F1CB3580D8EFBB6" box="[1144,1239,1039,1064]" italics="true" pageId="3" pageNumber="4">Venonia</emphasis>
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</taxonomicName>
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in its colouration, as the dorsal shield of the prosoma and the opisthosoma have light median bands, in particular distinct in males, whereas the dorsal shield of the prosoma and the opisthosoma of
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<taxonomicName id="9ED94D06FFE1D32C0AA1B3D00B7AFB3E" box="[453,547,1159,1184]" class="Arachnida" family="Lycosidae" genus="Venonia" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="6BADEA97FFE1D32C0AA1B3D00B7AFB3E" box="[453,547,1159,1184]" italics="true" pageId="3" pageNumber="4">Venonia</emphasis>
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</taxonomicName>
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are of uniform colouration with a posterodorsal white spot above the spinnerets. Pedipalp morphology shows distinct differences between
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<taxonomicName id="9ED94D06FFE1D32C0F16B3F80DAFFB56" box="[1138,1270,1199,1224]" class="Arachnida" family="Lycosidae" genus="Anomalosa" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="6BADEA97FFE1D32C0F16B3F80DAFFB56" box="[1138,1270,1199,1224]" italics="true" pageId="3" pageNumber="4">Anomalosa</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="9ED94D06FFE1D32C0A6CB380083FFB6E" box="[264,358,1239,1264]" class="Arachnida" family="Lycosidae" genus="Venonia" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="6BADEA97FFE1D32C0A6CB380083FFB6E" box="[264,358,1239,1264]" italics="true" pageId="3" pageNumber="4">Venonia</emphasis>
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</taxonomicName>
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. The cymbium is not truncated apicoretrolaterally as in
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<taxonomicName id="9ED94D06FFE1D32C0F38B3800DE3FB6E" box="[1116,1210,1239,1264]" class="Arachnida" family="Lycosidae" genus="Venonia" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="6BADEA97FFE1D32C0F38B3800DE3FB6E" box="[1116,1210,1239,1264]" italics="true" pageId="3" pageNumber="4">Venonia</emphasis>
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</taxonomicName>
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and the prolateral tegular lobe is much larger than the tegular apophysis in
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<taxonomicName id="9ED94D06FFE1D32C0F58B3A80D99FA86" box="[1084,1216,1279,1304]" class="Arachnida" family="Lycosidae" genus="Anomalosa" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="6BADEA97FFE1D32C0F58B3A80D99FA86" box="[1084,1216,1279,1304]" italics="true" pageId="3" pageNumber="4">Anomalosa</emphasis>
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</taxonomicName>
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(smaller than the tegular apophysis in
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<taxonomicName id="9ED94D06FFE1D32C091EB2700B81FADE" box="[634,728,1319,1344]" class="Arachnida" family="Lycosidae" genus="Venonia" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="6BADEA97FFE1D32C091EB2700B81FADE" box="[634,728,1319,1344]" italics="true" pageId="3" pageNumber="4">Venonia</emphasis>
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</taxonomicName>
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).
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<taxonomicName id="9ED94D06FFE1D32C0992B2700A23FADE" box="[758,890,1319,1344]" class="Arachnida" family="Lycosidae" genus="Anomalosa" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="6BADEA97FFE1D32C0992B2700A23FADE" box="[758,890,1319,1344]" italics="true" pageId="3" pageNumber="4">Anomalosa</emphasis>
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</taxonomicName>
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clearly differs from
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<taxonomicName id="9ED94D06FFE1D32C0FE4B2700C7AFADE" box="[1152,1315,1319,1344]" class="Arachnida" family="Lycosidae" genus="Anomalomma" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="6BADEA97FFE1D32C0FE4B2700C7AFADE" box="[1152,1315,1319,1344]" italics="true" pageId="3" pageNumber="4">Anomalomma</emphasis>
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</taxonomicName>
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, where
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<taxonomicName id="9ED94D06FFE1D32C0A39B218089DFAF6" box="[349,452,1359,1384]" class="Arachnida" family="Lycosidae" genus="Anomalomma" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="kochi">
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<emphasis id="6BADEA97FFE1D32C0A39B218089DFAF6" box="[349,452,1359,1384]" italics="true" pageId="3" pageNumber="4">A. kochi</emphasis>
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</taxonomicName>
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was initially placed, as
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<taxonomicName id="9ED94D06FFE1D32C0999B2180AF9FAF6" box="[765,928,1359,1384]" class="Arachnida" family="Lycosidae" genus="Anomalomma" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="6BADEA97FFE1D32C0999B2180AF9FAF6" box="[765,928,1359,1384]" italics="true" pageId="3" pageNumber="4">Anomalomma</emphasis>
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</taxonomicName>
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has short spinnerets (Simon in
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<bibRefCitation id="3D484B74FFE1D32C0A6CB22008FCFA0F" author="Hasselt" box="[264,421,1399,1425]" pageId="3" pageNumber="18" refString="Hasselt, A. W. M. van (1890) Araneae ex Archipelago Malayano. In: Weber, M., Zoologische Ergebnisse einer Reise in Niederlandisch Ost-Indien. Leiden, 1, pp. 193 - 210." type="journal article" year="1890">Hasselt 1890</bibRefCitation>
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: 199: “Mammillae brevissimae, ut in
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<emphasis id="6BADEA97FFE1D32C0839B2200AF1FA0E" box="[861,936,1399,1424]" italics="true" pageId="3" pageNumber="4">Cybeo</emphasis>
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||
L. Koch.”). The
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||
<typeStatus id="86628827FFE1D32C0F15B2200DFDFA0F" box="[1137,1188,1399,1425]" pageId="3" pageNumber="4">type</typeStatus>
|
||
material of the
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<typeStatus id="86628827FFE1D32C0A51B2C80831FA27" box="[309,360,1439,1465]" pageId="3" pageNumber="4">type</typeStatus>
|
||
species of the genus,
|
||
<taxonomicName id="9ED94D06FFE1D32C0916B2C80AF6FA27" authority="Simon, 1890" authorityName="Simon" authorityYear="1890" box="[626,943,1439,1465]" class="Arachnida" family="Lycosidae" genus="Anomalomma" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="lycosinum">
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||
<emphasis id="6BADEA97FFE1D32C0916B2C80A55FA26" box="[626,780,1439,1464]" italics="true" pageId="3" pageNumber="4">A. lycosinum</emphasis>
|
||
Simon, 1890
|
||
</taxonomicName>
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, appears to be lost (
|
||
<bibRefCitation id="3D484B74FFE1D32C0FC4B2C808CFFA7F" author="Lehtinen" lastPageId="3" lastPageNumber="19" pageId="3" pageNumber="18" refString="Hippa, H. & Lehtinen, P. T. (1983) The zantheres group of Zoicinae (Araneae, Lycosidae) and a relimitation of the subfamily. Annales Zologici Fennici, 20, 151 - 156." type="journal article" year="1983">Lehtinen & Hippa 1983</bibRefCitation>
|
||
).
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</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="11C3650EFFE1D32B0A5CB2B90885FD00" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="description">
|
||
<paragraph id="59663685FFE1D32B0A5CB2B90B1DFE18" blockId="3.[264,1323,958,1905]" lastBlockId="4.[264,1324,284,1390]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
|
||
<emphasis id="6BADEA97FFE1D32C0A5CB2B90896F996" bold="true" box="[312,463,1518,1544]" pageId="3" pageNumber="4">Description.</emphasis>
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Small wolf spiders (TL ca.
|
||
<quantity id="9E219B60FFE1D32C0871B2B80AFDF997" box="[789,932,1519,1545]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" metricValueMax="6.8" metricValueMin="3.8" pageId="3" pageNumber="4" unit="mm" value="5.3" valueMax="6.8" valueMin="3.8">3.8–6.8 mm</quantity>
|
||
) with narrow, elongated prosoma and opisthosoma. Dorsal profile of prosoma straight in lateral view (
|
||
<figureCitation id="C1E22A00FFE1D32C0F5DB1400DDAF9AF" box="[1081,1155,1559,1585]" captionStart="FIGURE 4" captionStartId="7.[264,367,1143,1167]" captionTargetBox="[531,1056,328,1112]" captionTargetId="figure@7.[531,1056,326,1119]" captionTargetPageId="7" captionText="FIGURE 4. Male pedipalp (SEM) of Anomalosa kochi (Simon) from Boggomoss, Queensland, Australia (QM S 36893), ventral view." httpUri="https://zenodo.org/record/173693/files/figure.png" pageId="3" pageNumber="4">Fig. 4</figureCitation>
|
||
). Head flanks steep in frontal view (
|
||
<figureCitation id="C1E22A00FFE1D32C096DB1680B08F9C7" box="[521,593,1599,1625]" captionStart="FIGURES 5 – 11" captionStartId="8.[264,382,1670,1694]" captionTargetBox="[342,1258,335,1649]" captionTargetId="figure@8.[335,1258,326,1649]" captionTargetPageId="8" captionText="FIGURES 5 – 11. Anomalosa kochi (Simon), male (QM S 36714) from Nathan Gorge, Queensland, Australia, female (QM S 36727) from Boggomoss, Queensland, Australia. 5 Male prosoma, lateral view; 6 male eye pattern, frontal view; 7 – 8 Left male pedipalp (7 ventral, 8 retrolateral view); 9 Apical part of bulbus, ventral view; 10 – 11 Female epigyne (10 ventral, 11 dorsal view). Scale bar: 5, 2.30 mm; 6, 0.90 mm; 7 – 8, 0.45 mm; 9, 0.30 mm; 10 – 11, 0.35 mm. The tip of the resting embolus in Fig. 9 is dislodged from its normal resting position behind, and in a grove of, the terminal apophysis, similar to A. oz sp. nov. (see Fig. 18)." httpUri="https://zenodo.org/record/173694/files/figure.png" pageId="3" pageNumber="4">Fig. 5</figureCitation>
|
||
). Uniform brown to black colouration, but centrally lighter or with a distinct light median band. Indistinct darker radial pattern. Anterior median eyes slightly larger than anterior lateral eyes, row of AE narrower than row of PME; row of AE procurved (
|
||
<figureCitation id="C1E22A00FFE1D32C0AF2B1E008BCF94F" box="[406,485,1719,1745]" captionStart="FIGURES 5 – 11" captionStartId="8.[264,382,1670,1694]" captionTargetBox="[342,1258,335,1649]" captionTargetId="figure@8.[335,1258,326,1649]" captionTargetPageId="8" captionText="FIGURES 5 – 11. Anomalosa kochi (Simon), male (QM S 36714) from Nathan Gorge, Queensland, Australia, female (QM S 36727) from Boggomoss, Queensland, Australia. 5 Male prosoma, lateral view; 6 male eye pattern, frontal view; 7 – 8 Left male pedipalp (7 ventral, 8 retrolateral view); 9 Apical part of bulbus, ventral view; 10 – 11 Female epigyne (10 ventral, 11 dorsal view). Scale bar: 5, 2.30 mm; 6, 0.90 mm; 7 – 8, 0.45 mm; 9, 0.30 mm; 10 – 11, 0.35 mm. The tip of the resting embolus in Fig. 9 is dislodged from its normal resting position behind, and in a grove of, the terminal apophysis, similar to A. oz sp. nov. (see Fig. 18)." httpUri="https://zenodo.org/record/173694/files/figure.png" pageId="3" pageNumber="4">Fig. 5</figureCitation>
|
||
). Chelicerae with three promarginal and three retromarginal teeth. Labium as wide or slightly wider than long. Opisthosoma uniformly olivegrey to black, with a light median line, in particularly distinct in males. Posterior spinnerets distinctly longer than anterior spinnerets (
|
||
<figureCitation id="C1E22A00FFE1D32C091AB0780B9FF8D7" box="[638,710,1839,1865]" captionStart="FIGURE 3" captionStartId="6.[264,367,1777,1801]" captionTargetBox="[401,1184,1146,1752]" captionTargetId="figure@6.[401,1186,1146,1753]" captionTargetPageId="6" captionText="FIGURE 3. Spinnerets of male Anomalosa kochi (Simon) from Laidley Creek, Queensland, Australia (QM S 61256), ventral view. Scale bar 0.80 mm." httpUri="https://zenodo.org/record/173692/files/figure.png" pageId="3" pageNumber="4">Fig. 3</figureCitation>
|
||
). Coxae light yellowbrown, femora II–IV with one basal and one central light annulation. Leg formula generally VI>I>II>III, but variable as legs I–III are of very similar length. Spination of legs: Femur: 2 or 3 dorsal, 0–1 (rarely 2) apicoprolateral; patella: 1 apicodorsal; tibia: 2 dorsal; 2 ventral pairs, 1 prolateral; metatarsus: 3 ventral pairs.
|
||
</paragraph>
|
||
<paragraph id="59663685FFE6D32B0A5CB6C30885FD00" blockId="4.[264,1324,284,1390]" pageId="4" pageNumber="5">
|
||
Tegulum with a prolateral, bipartite (i.e. partly sclerotised and partly membranous) lobe and a retrolateral extension in which the spermduct is visible. Tegular apophysis membranous (heavily reduced in
|
||
<taxonomicName id="9ED94D06FFE6D32B09C0B6B30A50FE63" box="[676,777,484,509]" class="Arachnida" family="Lycosidae" genus="Anomalomma" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="kochi">
|
||
<emphasis id="6BADEA97FFE6D32B09C0B6B30A50FE63" box="[676,777,484,509]" italics="true" pageId="4" pageNumber="5">A. kochi</emphasis>
|
||
</taxonomicName>
|
||
). Embolus originating prolaterally on palea, long and slender and with a curled tip that rests in a cavity of the terminal apophysis. Terminal apophysis originates centrally on the apical part of the bulb. Epigyne a simple sclerotised plate with a posterior incision. Large, oval spermathecae and reduced fertilisation ducts.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="11C3650EFFE6D32B0A5CB5FB0D62FD58" box="[312,1083,684,710]" pageId="4" pageNumber="5" type="distribution">
|
||
<paragraph id="59663685FFE6D32B0A5CB5FB0D62FD58" blockId="4.[264,1324,284,1390]" box="[312,1083,684,710]" pageId="4" pageNumber="5">
|
||
<emphasis id="6BADEA97FFE6D32B0A5CB5FB0881FD58" bold="true" box="[312,472,684,710]" pageId="4" pageNumber="5">Distribution.</emphasis>
|
||
Currently only known from
|
||
<collectingCountry id="21CE7615FFE6D32B084EB5FB0ACFFD58" box="[810,918,684,710]" name="Australia" pageId="4" pageNumber="5">Australia</collectingCountry>
|
||
(
|
||
<figureCitation id="C1E22A00FFE6D32B08C1B5FB0D59FD58" box="[933,1024,684,710]" captionStart="FIGURE 12" captionStartId="9.[264,367,1059,1083]" captionTargetBox="[348,1245,321,1040]" captionTargetId="figure@9.[345,1251,309,1052]" captionTargetPageId="9" captionText="FIGURE 12. Records of Anomalosa kochi (Simon) in Australia." httpUri="https://zenodo.org/record/173695/files/figure.png" pageId="4" pageNumber="5">Figs 12</figureCitation>
|
||
,
|
||
<figureCitation id="C1E22A00FFE6D32B0F68B5FB0D74FD58" box="[1036,1069,684,710]" captionStart="FIGURE 21" captionStartId="15.[264,367,1091,1115]" captionTargetBox="[329,1262,326,1072]" captionTargetId="figure@15.[326,1267,310,1083]" captionTargetPageId="15" captionText="FIGURE 21. Records of Anomalosa oz sp. nov. in Australia." httpUri="https://zenodo.org/record/173699/files/figure.png" pageId="4" pageNumber="5">21</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="11C3650EFFE6D32B0A5CB5830A6FFAF0" pageId="4" pageNumber="5" type="discussion">
|
||
<paragraph id="59663685FFE6D32B0A5CB58308CFFBC8" blockId="4.[264,1324,284,1390]" pageId="4" pageNumber="5">
|
||
<emphasis id="6BADEA97FFE6D32B0A5CB58308E8FD70" bold="true" box="[312,433,724,750]" pageId="4" pageNumber="5">Remarks.</emphasis>
|
||
A recent molecular phylogeny inferred from 12S rRNA and NADH1 including seventy worldwide lycosid species with the majority from the Australasian region, included
|
||
<emphasis id="6BADEA97FFE6D32B0AB5B4730B54FCA3" box="[465,525,804,829]" italics="true" pageId="4" pageNumber="5">A. oz</emphasis>
|
||
<emphasis id="6BADEA97FFE6D32B0972B4730B2CFCA0" bold="true" box="[534,629,804,830]" pageId="4" pageNumber="5">
|
||
<taxonomicNameLabel id="709E57ECFFE6D32B0972B4730B2CFCA0" box="[534,629,804,830]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(as
|
||
<taxonomicName id="9ED94D06FFE6D32B09CDB4730A2CFCA3" box="[681,885,804,829]" class="Arachnida" family="Lycosidae" genus="Anomalosa" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="kochi">
|
||
<emphasis id="6BADEA97FFE6D32B09CDB4730A2CFCA3" box="[681,885,804,829]" italics="true" pageId="4" pageNumber="5">Anomalosa kochi</emphasis>
|
||
</taxonomicName>
|
||
, WAM T56076) as representative of
|
||
<taxonomicName id="9ED94D06FFE6D32B0A6CB41B0BE2FCF8" authority="Murphy et al. 2006" authorityName="Murphy et al." authorityYear="2006" box="[264,699,844,870]" class="Arachnida" family="Lycosidae" genus="Anomalosa" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6BADEA97FFE6D32B0A6CB41B08D5FCFB" box="[264,396,844,869]" italics="true" pageId="4" pageNumber="5">Anomalosa</emphasis>
|
||
(
|
||
<bibRefCitation id="3D484B74FFE6D32B0AC3B41B0BEAFCF8" author="Murphy" box="[423,691,844,870]" pageId="4" pageNumber="19" refString="Murphy, N. P., Framenau, V. W., Donellan, S. C., Harvey, M. S., Park, Y. - C. & Austin, A. D. (2006) Phylogenetic reconstruction of the wolf spiders (Araneae: Lycosidae) using sequences from the 12 S rRNA, 28 rRNA and NADH 1 genes: implications for classification, biogeography and the evolution of web building behavior. Molecular Phylogenetics and Evolution, 38, 583 - 602." type="journal article" year="2006">
|
||
Murphy
|
||
<emphasis id="6BADEA97FFE6D32B097EB41B0B38FCFB" box="[538,609,844,869]" italics="true" pageId="4" pageNumber="5">et al.</emphasis>
|
||
2006
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Independent of the phylogenetic analysis used (parsimony, Bayesian),
|
||
<emphasis id="6BADEA97FFE6D32B0945B4230B05FC13" box="[545,604,884,909]" italics="true" pageId="4" pageNumber="5">A. oz</emphasis>
|
||
<emphasis id="6BADEA97FFE6D32B0901B4230B9DFC10" bold="true" box="[613,708,884,910]" pageId="4" pageNumber="5">
|
||
<taxonomicNameLabel id="709E57ECFFE6D32B0901B4230B9DFC10" box="[613,708,884,910]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
represented the sister taxon of
|
||
<taxonomicName id="9ED94D06FFE6D32B0F5DB423089CFC28" authority="L. Koch, 1877" authorityName="L. Koch" authorityYear="1877" class="Arachnida" family="Lycosidae" genus="Venonia" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="micarioides">
|
||
<emphasis id="6BADEA97FFE6D32B0F5DB4230C72FC13" box="[1081,1323,884,909]" italics="true" pageId="4" pageNumber="5">Venonia micarioides</emphasis>
|
||
(
|
||
<bibRefCitation id="3D484B74FFE6D32B0A75B4CB08E4FC28" author="Koch" box="[273,445,924,950]" pageId="4" pageNumber="19" refString="Koch, L. (1877) Die Arachniden Australiens, nach der Natur beschrieben und abgebildet. Bauer and Raspe, Nurnberg, pp. 889 - 968." type="book chapter" year="1877">L. Koch, 1877</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
in a clade that also included both species of
|
||
<taxonomicName id="9ED94D06FFE6D32B08B7B4CB08C6FC40" authority="Roewer, 1960 (Vink 2001)" authorityName="Roewer, 1960 (Vink" authorityYear="2001" class="Arachnida" family="Lycosidae" genus="Allotrochosina" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6BADEA97FFE6D32B08B7B4CB0DD9FC2B" box="[979,1152,924,949]" italics="true" pageId="4" pageNumber="5">Allotrochosina</emphasis>
|
||
<bibRefCitation id="3D484B74FFE6D32B0FE3B4CB0C72FC28" author="Roewer" box="[1159,1323,924,950]" pageId="4" pageNumber="19" refString="Roewer, C. F. (1960) [imprint date 1959]. Araneae Lycosiformae II (Lycosidae) (Fortsetzung und Schluss). Exploration du Parc National de l'Upemba - Mission GF de Witte, 55, 519 - 1040." type="journal article" year="1960">Roewer, 1960</bibRefCitation>
|
||
(
|
||
<bibRefCitation id="3D484B74FFE6D32B0A74B49308CFFC40" author="Vink" box="[272,406,964,990]" pageId="4" pageNumber="20" refString="Vink, C. J. (2001) A revision of the genus Allotrochosina Roewer (Lycosidae: Araneae). Invertebrate Taxonomy, 15, 461 - 466." type="journal article" year="2001">Vink 2001</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. This confirmed the close relationship of
|
||
<taxonomicName id="9ED94D06FFE6D32B08C8B4930D69FC43" box="[940,1072,964,989]" class="Arachnida" family="Lycosidae" genus="Anomalosa" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6BADEA97FFE6D32B08C8B4930D69FC43" box="[940,1072,964,989]" italics="true" pageId="4" pageNumber="5">Anomalosa</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="9ED94D06FFE6D32B0F17B4930D88FC43" box="[1139,1233,964,989]" class="Arachnida" family="Lycosidae" genus="Venonia" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6BADEA97FFE6D32B0F17B4930D88FC43" box="[1139,1233,964,989]" italics="true" pageId="4" pageNumber="5">Venonia</emphasis>
|
||
</taxonomicName>
|
||
that is evident by similar somatic and genital morphology. This clade (Venoniinae
|
||
<emphasis id="6BADEA97FFE6D32B0FE8B4BB0C78FB9B" box="[1164,1313,1004,1029]" italics="true" pageId="4" pageNumber="5">sensu stricto</emphasis>
|
||
) is sister to a clade including all species of
|
||
<taxonomicName id="9ED94D06FFE6D32B0999B3430A11FBB3" box="[765,840,1044,1069]" class="Arachnida" family="Lycosidae" genus="Pirata" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6BADEA97FFE6D32B0999B3430A11FBB3" box="[765,840,1044,1069]" italics="true" pageId="4" pageNumber="5">Pirata</emphasis>
|
||
</taxonomicName>
|
||
(Piratinae
|
||
<emphasis id="6BADEA97FFE6D32B08AEB3430D52FBB3" box="[970,1035,1044,1069]" italics="true" pageId="4" pageNumber="5">sensu</emphasis>
|
||
<bibRefCitation id="3D484B74FFE6D32B0F70B3430DEBFBB0" author="Zyuzin" box="[1044,1202,1044,1070]" pageId="4" pageNumber="20" refString="Zyuzin, A. A. (1993) Studies on the wolf spiders (Araneae: Lycosidae). I. A new genus and species from Kazakhstan, with comments on the Lycosinae. Memoirs of the Queensland Museum, 33, 693 - 700." type="journal article" year="1993">Zyuzin, 1993</bibRefCitation>
|
||
) (
|
||
<bibRefCitation id="3D484B74FFE6D32B0FAEB34308D3FBC8" author="Murphy" pageId="4" pageNumber="19" refString="Murphy, N. P., Framenau, V. W., Donellan, S. C., Harvey, M. S., Park, Y. - C. & Austin, A. D. (2006) Phylogenetic reconstruction of the wolf spiders (Araneae: Lycosidae) using sequences from the 12 S rRNA, 28 rRNA and NADH 1 genes: implications for classification, biogeography and the evolution of web building behavior. Molecular Phylogenetics and Evolution, 38, 583 - 602." type="journal article" year="2006">
|
||
Murphy
|
||
<emphasis id="6BADEA97FFE6D32B0A6CB36B081AFBCB" box="[264,323,1084,1109]" italics="true" pageId="4" pageNumber="5">et al.</emphasis>
|
||
2006
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="59663685FFE6D32B0A5CB3330A6FFAF0" blockId="4.[264,1324,284,1390]" pageId="4" pageNumber="5">
|
||
The male pedipalp of
|
||
<taxonomicName id="9ED94D06FFE6D32B0953B3330BE2FBE3" box="[567,699,1124,1149]" class="Arachnida" family="Lycosidae" genus="Anomalosa" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6BADEA97FFE6D32B0953B3330BE2FBE3" box="[567,699,1124,1149]" italics="true" pageId="4" pageNumber="5">Anomalosa</emphasis>
|
||
</taxonomicName>
|
||
is characterised by a large prolateral tegular lobe and membranous tegular apophysis (heavily reduced in
|
||
<taxonomicName id="9ED94D06FFE6D32B08C4B3DB0D52FB3B" box="[928,1035,1164,1189]" class="Arachnida" family="Lycosidae" genus="Anomalosa" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="kochi">
|
||
<emphasis id="6BADEA97FFE6D32B08C4B3DB0D52FB3B" box="[928,1035,1164,1189]" italics="true" pageId="4" pageNumber="5">A. kochi</emphasis>
|
||
</taxonomicName>
|
||
). These structures are equivalent to structures in
|
||
<taxonomicName id="9ED94D06FFE6D32B0921B3E30BFDFB53" box="[581,676,1204,1229]" class="Arachnida" family="Lycosidae" genus="Venonia" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6BADEA97FFE6D32B0921B3E30BFDFB53" box="[581,676,1204,1229]" italics="true" pageId="4" pageNumber="5">Venonia</emphasis>
|
||
</taxonomicName>
|
||
in which the prolateral tegular lobe is generally much smaller and the tegular apophysis much larger (Yoo & Framenau in press). Lethinen and
|
||
<bibRefCitation id="3D484B74FFE6D32B0A6CB25308FCFA80" box="[264,421,1284,1310]" pageId="4" pageNumber="19" refString="Lehtinen, P. T. & Hippa, H. (1979) Spiders of the Oriental-Australian region. I. Lycosidae: Venoniinae and Zoicinae. Annales Zoologici Fennici, 16, 1 - 22." type="journal article">Hippa (1979)</bibRefCitation>
|
||
did not know the second species of
|
||
<taxonomicName id="9ED94D06FFE6D32B0830B2530A81FA83" box="[852,984,1284,1309]" class="Arachnida" family="Lycosidae" genus="Anomalosa" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6BADEA97FFE6D32B0830B2530A81FA83" box="[852,984,1284,1309]" italics="true" pageId="4" pageNumber="5">Anomalosa</emphasis>
|
||
</taxonomicName>
|
||
, which bears both, a distinct prolateral tegular lobe and a tegular apophysis (e.g.
|
||
<figureCitation id="C1E22A00FFE6D32B0811B27B0A95FAD8" box="[885,972,1324,1350]" captionStart="FIGURES 16 – 20" captionStartId="14.[264,382,1692,1716]" captionTargetBox="[294,1303,728,1658]" captionTargetId="figure@14.[287,1307,726,1677]" captionTargetPageId="14" captionText="FIGURES 16 – 20. Anomalosa oz sp. nov., male holotype (WAM T 65677) from Barmah State Forest, Victoria, Australia, and female paratype (WAM T 70050) from Lake Eppalock, Victoria, Australia. 16 – 17 Left male pedipalp (16 ventral, 17 retrolateral view); 18 Apical part of bulbus, ventral view. 19 – 20 Female epigyne (19 ventral, 20 dorsal view). Scale bar: 16 – 17, 1.30 mm; 18, 0.75 mm; 19 – 20, 0.30 mm." httpUri="https://zenodo.org/record/173698/files/figure.png" pageId="4" pageNumber="5">Fig. 16</figureCitation>
|
||
), and erroneously termed the prolateral lobe as median (= tegular) apophysis.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |