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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.985.51622" ID-GBIF-Dataset="8e4b4855-0ead-4bfa-9444-338191711d0c" ID-PMC="PMC7661952" ID-Pensoft-Pub="1313-2970-985-71" ID-Pensoft-UUID="673D74424EF656EF8EC4AC8BFB4D00FA" ID-PubMed="33223876" ID-ZooBank="7A38B594F29D43F18CB18B108AC18A1C" ModsDocID="1313-2970-985-71" checkinTime="1604647783870" checkinUser="pensoft" docAuthor="Martinez, Jose I." docDate="2020" docId="2E45AF379CF45CA0BABBFA0B0513F8B6" docLanguage="en" docName="ZooKeys 985: 71-126" docOrigin="ZooKeys 985" docSource="http://dx.doi.org/10.3897/zookeys.985.51622" docTitle="Oculicattus raizae Martinez 2020, sp. nov." docType="treatment" docUuid="BD31472B-72CC-4B58-B011-ED8E5339F5D4" docUuidSource="ZooBank" docVersion="4" id="673D74424EF656EF8EC4AC8BFB4D00FA" lastPageNumber="71" masterDocId="673D74424EF656EF8EC4AC8BFB4D00FA" masterDocTitle="Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species" masterLastPageNumber="126" masterPageNumber="71" pageNumber="71" updateTime="1668169762986" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of the South American genus Gaujonia Dognin (Noctuidae, Pantheinae) with descriptions of five new genera and twenty-one new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Martinez, Jose I.</mods:namePart>
<mods:affiliation>Florida Museum of Natural History, McGuire Center for Lepidoptera and Biodiversity, University of Florida, Gainesville, FL, 32611, USA, &amp; Entomology and Nematology Department, University of Florida, Gainesville, FL, 326011, USA</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0003-4368-2729</mods:nameIdentifier>
<mods:nameIdentifier type="email">joemartinez@ufl.edu</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>985</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>71</mods:start>
<mods:end>126</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.985.51622</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.985.51622</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-985-71</mods:identifier>
<mods:identifier type="ZooBank">7A38B594F29D43F18CB18B108AC18A1C</mods:identifier>
<mods:identifier type="Pensoft-UUID">673D74424EF656EF8EC4AC8BFB4D00FA</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="168815811" LSID="urn:lsid:zoobank.org:act:BD31472B-72CC-4B58-B011-ED8E5339F5D4" httpUri="http://treatment.plazi.org/id/2E45AF379CF45CA0BABBFA0B0513F8B6" lastPageNumber="71" pageId="0" pageNumber="71">
<subSubSection pageId="0" pageNumber="71" type="nomenclature">
<paragraph pageId="0" pageNumber="71">
<taxonomicName LSID="2E45AF37-9CF4-5CA0-BABB-FA0B0513F8B6" authority="Martinez, 2020" authorityName="Martinez" authorityYear="2020" class="Insecta" family="Noctuidae" genus="Oculicattus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Oculicattus raizae" order="Lepidoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="raizae" status="sp. nov.">Oculicattus raizae</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="71">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figures 43–58" captionStartId="F6" captionText="Figures 43 - 58. Adult habitus of Oculicattus species 43 Oculicattus renifera, ♂, MGCL, Cusco, Peru 44 O. renifera, ♂, MGCL, Junin, Peru 45 O. schmidti, ♂, holotype, MGCL, Pasco, Peru 46 O. schmidti, ♂, paratype, MGCL, Junin, Peru 47 O. inca, ♂, holotype, MGCL, Sierra Siberia, Bolivia 48 O. boliviana, ♂, holotype, MGCL, Sierra Siberia, Bolivia 49 O. uturunku, ♂, holotype, MGCL, Morona, Ecuador 50 O. raizae, ♂, holotype, MGCL, Tolima, Colombia 51 O. raizae, ♂, paratype, ZSM, Pichincha, Ecuador 52 O. brehmi, ♂, holotype, FSU, Loja, Ecuador 53 O. brehmi, ♂, paratype, MGCL, Napo, Ecuador 54 O. brehmi, ♀, paratype, MGCL, Napo, Ecuador 55 O. brehmi, ♀, paratype, MGCL, Napo, Ecuador 56 O. renifera, ♀, MGCL, Cusco, Peru 57 O. renifera, ♀, MGCL, Cusco, Peru 58 O. raizae, ♀, paratype, MGCL, La Paz, Bolivia." figureDoi="10.3897/zookeys.985.51622.figures43-58" httpUri="https://binary.pensoft.net/fig/471606" pageId="0" pageNumber="71">Figs 50</figureCitation>
<figureCitation captionStart="Figures 43–58" captionStartId="F6" captionText="Figures 43 - 58. Adult habitus of Oculicattus species 43 Oculicattus renifera, ♂, MGCL, Cusco, Peru 44 O. renifera, ♂, MGCL, Junin, Peru 45 O. schmidti, ♂, holotype, MGCL, Pasco, Peru 46 O. schmidti, ♂, paratype, MGCL, Junin, Peru 47 O. inca, ♂, holotype, MGCL, Sierra Siberia, Bolivia 48 O. boliviana, ♂, holotype, MGCL, Sierra Siberia, Bolivia 49 O. uturunku, ♂, holotype, MGCL, Morona, Ecuador 50 O. raizae, ♂, holotype, MGCL, Tolima, Colombia 51 O. raizae, ♂, paratype, ZSM, Pichincha, Ecuador 52 O. brehmi, ♂, holotype, FSU, Loja, Ecuador 53 O. brehmi, ♂, paratype, MGCL, Napo, Ecuador 54 O. brehmi, ♀, paratype, MGCL, Napo, Ecuador 55 O. brehmi, ♀, paratype, MGCL, Napo, Ecuador 56 O. renifera, ♀, MGCL, Cusco, Peru 57 O. renifera, ♀, MGCL, Cusco, Peru 58 O. raizae, ♀, paratype, MGCL, La Paz, Bolivia." figureDoi="10.3897/zookeys.985.51622.figures43-58" httpUri="https://binary.pensoft.net/fig/471606" pageId="0" pageNumber="71">, 51</figureCitation>
<figureCitation captionStart="Figures 43–58" captionStartId="F6" captionText="Figures 43 - 58. Adult habitus of Oculicattus species 43 Oculicattus renifera, ♂, MGCL, Cusco, Peru 44 O. renifera, ♂, MGCL, Junin, Peru 45 O. schmidti, ♂, holotype, MGCL, Pasco, Peru 46 O. schmidti, ♂, paratype, MGCL, Junin, Peru 47 O. inca, ♂, holotype, MGCL, Sierra Siberia, Bolivia 48 O. boliviana, ♂, holotype, MGCL, Sierra Siberia, Bolivia 49 O. uturunku, ♂, holotype, MGCL, Morona, Ecuador 50 O. raizae, ♂, holotype, MGCL, Tolima, Colombia 51 O. raizae, ♂, paratype, ZSM, Pichincha, Ecuador 52 O. brehmi, ♂, holotype, FSU, Loja, Ecuador 53 O. brehmi, ♂, paratype, MGCL, Napo, Ecuador 54 O. brehmi, ♀, paratype, MGCL, Napo, Ecuador 55 O. brehmi, ♀, paratype, MGCL, Napo, Ecuador 56 O. renifera, ♀, MGCL, Cusco, Peru 57 O. renifera, ♀, MGCL, Cusco, Peru 58 O. raizae, ♀, paratype, MGCL, La Paz, Bolivia." figureDoi="10.3897/zookeys.985.51622.figures43-58" httpUri="https://binary.pensoft.net/fig/471606" pageId="0" pageNumber="71">, 58</figureCitation>
<figureCitation captionStart="Figures 74–79" captionStartId="F10" captionText="Figures 74 - 79. Male genitalia of Oculicattus species 74 Oculicattus inca, holotype, MGCL, Sierra Siberia, Bolivia 75 O. boliviana, holotype, MGCL, Sierra Siberia, Bolivia 76 O. uturunku, holotype, MGCL, Morona, Ecuador 77 O. brehmi, holotype, FSU, Loja, Ecuador 78 O. raizae, holotype, MGCL, Tolima, Colombia 79 O. schmidti, holotype, MGCL, Pasco, Peru." figureDoi="10.3897/zookeys.985.51622.figures74-79" httpUri="https://binary.pensoft.net/fig/471610" pageId="0" pageNumber="71">, 78</figureCitation>
<figureCitation captionStart="Figures 85–90" captionStartId="F12" captionText="Figures 85 - 90. Female genitalia of Gaujonia and Oculicattus species 85 Gaujonia arbosi, MGCL, Napo, Ecuador 86 G. chiqyaq, holotype, MGCL, Marinio, Colombia 87 G. kanakusika, paratype, MGCL, Cundinamarca, Colombia 88 Oculicattus renifera, MGCL, Cusco, Peru 89 O. brehmi, paratype, MGCL, Napo, Ecuador 90 O. raizae, paratype, MGCL, La Paz, Bolivia." figureDoi="10.3897/zookeys.985.51622.figures85-90" httpUri="https://binary.pensoft.net/fig/471612" pageId="0" pageNumber="71">, 90</figureCitation>
<figureCitation captionStart="Figure 95" captionStartId="F17" captionText="Figure 95. Distribution of examined specimens of the genus Oculicattus." figureDoi="10.3897/zookeys.985.51622.figure95" httpUri="https://binary.pensoft.net/fig/471617" pageId="0" pageNumber="71">, 95</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="type material">
<paragraph pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Holotype</emphasis>
♂,
<emphasis bold="true" pageId="0" pageNumber="71">Colombia</emphasis>
: Colombia, Tolima, Nevado del Tolima,
<geoCoordinate degrees="4" direction="north" minutes="36" orientation="latitude" precision="15" seconds="02" value="4.6005554">
4°36'02
<normalizedToken originalValue="’’">''</normalizedToken>
N
</geoCoordinate>
,
<geoCoordinate degrees="75" direction="west" minutes="19" orientation="longitude" precision="15" seconds="51" value="-75.33083">
75°19'51
<normalizedToken originalValue="’’">''</normalizedToken>
W
</geoCoordinate>
, 2600 m, 05-07.12.2013, legit Victor Sinyaev &amp; Mildred
<normalizedToken originalValue="Márquez">Marquez</normalizedToken>
Martinez / UF, FLMNH, MGCL 1049101. [DNA voucher MGCL-NOC- 65284] deposited in MGCL.
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Paratypes</emphasis>
(6 ♂, 1 ♀, MGCL):
<emphasis bold="true" pageId="0" pageNumber="71">Ecuador</emphasis>
: Same collecting data as holotype (2 ♂); Ecuador, Carchi, El Angel Ecological Reserve,
<geoCoordinate degrees="0" direction="north" minutes="45" orientation="latitude" precision="15" seconds="31" value="0.7586111">0°45'31&quot;N</geoCoordinate>
,
<geoCoordinate degrees="78" direction="west" minutes="01" orientation="longitude" precision="15" seconds="40" value="-78.02778">78°01'40&quot;W</geoCoordinate>
, 7-8. XI 2012, 3320 m, leg. Sinjaev &amp; Romanov (1 ♂); Ecuador, Carchi prov., El Moran,
<geoCoordinate degrees="0" direction="north" minutes="45" orientation="latitude" precision="15" seconds="50" value="0.7638889">0°45'50&quot;N</geoCoordinate>
,
<geoCoordinate degrees="78" direction="west" minutes="02" orientation="longitude" precision="15" seconds="38" value="-78.04389">78°02'38&quot;W</geoCoordinate>
, 1-3.05.2012, H = 2940 m, Exped. Ron Brechlin &amp; Victor Sinyaev (1 ♂); Ecuador, Pichincha, Camping Bella Vista, 2230 m,
<geoCoordinate degrees="0" direction="south" minutes="00" orientation="latitude" precision="15" seconds="41" value="-0.011388889">0°00'41&quot;S</geoCoordinate>
,
<geoCoordinate degrees="78" direction="west" minutes="41" orientation="longitude" precision="15" seconds="17" value="-78.68806">78°41'17&quot;W</geoCoordinate>
, 19.XII.2012-7.I.2013, leg. Sinjaev &amp; Romanov &amp; [coll.] Dr. R. Brechlin (1 ♂); Ecuador, Pichincha, Quito/Chiriboga, K40, 2480 m, 22 Mar. 1982, coll. N. Venedictoff (1 ♀);
<emphasis bold="true" pageId="0" pageNumber="71">Peru</emphasis>
: Peru-Junin Near Calabaza vill.,
<geoCoordinate degrees="11" direction="south" minutes="30.4" orientation="latitude" precision="92" value="-11.506666">11°30.4'S</geoCoordinate>
,
<geoCoordinate degrees="74" direction="west" minutes="51.7" orientation="longitude" precision="92" value="-74.861664">74°51.7'W</geoCoordinate>
, 20.12.2010, H = 2722 m, leg/coll. Viktor &amp; Svetlana Sinyaev + Vladimir Izerskiy (1 ♂);
<emphasis bold="true" pageId="0" pageNumber="71">Bolivia</emphasis>
: BOLIVIA, La Paz, Death road Coroico,
<geoCoordinate degrees="16" direction="south" minutes="18.3" orientation="latitude" precision="92" value="-16.305">16°18.3'S</geoCoordinate>
,
<geoCoordinate degrees="67" direction="west" minutes="48.8" orientation="longitude" precision="92" value="-67.81333">67°48.8'W</geoCoordinate>
, 28-30.10.2010, H = 3060 m, leg. Viktor Sinyaev &amp; Oleg Romanov (1 ♀).
<emphasis bold="true" pageId="0" pageNumber="71">Additional examined specimens</emphasis>
(1 ♂, ZSM):
<emphasis bold="true" pageId="0" pageNumber="71">Ecuador</emphasis>
: Ecuador, Anden Oskordillere, Prov. Pichincha, Tandayapa, Km. 3 S (Bellavista, Lodge), 2310 m, 00°03'694&quot;N, 78°40'929&quot;W 1-20.ix.2012, Dietl Monika + Stefan &amp;, R. Beck leg. / BC ZSM Lep 65183 / coll. G. Behounek, grafing bei
<normalizedToken originalValue="Müchen">Muechen</normalizedToken>
/ &quot;
<emphasis italics="true" pageId="0" pageNumber="71">Gaujonia renifera</emphasis>
♂&quot; (1 ♂).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="etymology">
<paragraph pageId="0" pageNumber="71">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="71">The species is dedicated to my wife, Raiza Castillo, for her love and support since the beginning of my career.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="diagnosis">
<paragraph pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="71">
<taxonomicName authorityName="Martinez" authorityYear="2020" class="Insecta" family="Noctuidae" genus="Oculicattus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Oculicattus raizae" order="Lepidoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="raizae">
<emphasis italics="true" pageId="0" pageNumber="71">Oculicattus raizae</emphasis>
</taxonomicName>
is closely related to
<taxonomicName lsidName="O. uturunku" pageId="0" pageNumber="71" rank="species" species="uturunku">
<emphasis italics="true" pageId="0" pageNumber="71">O. uturunku</emphasis>
</taxonomicName>
; they differ from the other
<taxonomicName authorityName="Martinez" authorityYear="2020" class="Insecta" family="Noctuidae" genus="Oculicattus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Oculicattus" order="Lepidoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Oculicattus</emphasis>
</taxonomicName>
species by the darker color, and the unusual reniform spot that is completely brown or black.
<taxonomicName authorityName="Martinez" authorityYear="2020" class="Insecta" family="Noctuidae" genus="Oculicattus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Oculicattus raizae" order="Lepidoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="raizae">
<emphasis italics="true" pageId="0" pageNumber="71">Oculicattus raizae</emphasis>
</taxonomicName>
can be identified by its brownish yellow coloration, and the brown forewing pattern. In the male genitalia the cucullus is longer and wider than in
<taxonomicName lsidName="O. uturunku" pageId="0" pageNumber="71" rank="species" species="uturunku">
<emphasis italics="true" pageId="0" pageNumber="71">O. uturunku</emphasis>
</taxonomicName>
. Additionally, DNA barcodes are ca. 4% different.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="description">
<paragraph pageId="0" pageNumber="71">Description.</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Head</emphasis>
.
</emphasis>
Palp reduced in size; dorsal surface covered by brown scales, ventral surface by dark sulfur-yellow scales; frons covered by dark sulfur-yellow scales, mixed with black; female similar to male externally.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
.
</emphasis>
Dark sulfur-yellow with some brown patches.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Wing</emphasis>
.
</emphasis>
Forewing with dark sulfur-yellow and brown scales covering venation and margins; forewing length: male: 22-24 mm; female: 27-29 mm; antemedial, postmedial, and subterminal lines not defined; orbicular spot small, poorly-defined oblong brown spot; reniform spot brown, narrow, and barely defined; base of CuA2 with Y-shaped mark large, brown; a brown line across fold on cell at CuA2; hindwing hyaline with some dark sulfur-yellow scales on venation, but more notable along margins; a Y-shape mark at base of CuA2; some small brown lines scattered through M3+CuA1+CuA2.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Leg</emphasis>
.
</emphasis>
Dark sulfur-yellow with brown patches on anterior area of prothoracic legs, which is lighter on mesothoracic and metathoracic legs.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Abdomen</emphasis>
.
</emphasis>
Clothed in grayish yellow hair-like scales with white scales at terminus and with a dark sulfur-yellow tuft on A8.
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
.
</emphasis>
Cucullus wide, parallel sided with the apex rounded; costal margin swollen basally; sacculus triangular with saccular process entirely coated with setae; tegumen wide; saccus broadly V-shaped; juxta small with parallel sides and a V-shaped notch posteriorly; aedeagus 3
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide, with a wide opening to vesica ca.
<normalizedToken originalValue="¼">1/4</normalizedToken>
<normalizedToken originalValue="×">x</normalizedToken>
total length of aedeagus; vesica with basal area ca. same width that of aedeagus, and it is ventrally curved; basal area with a large patch of spines; vesica with two patches of spines on each side, one small slightly beyond middle of dorsal wall of the vesica, and truncated anteriorly, whereas other larger apically narrow, and broader towards the aedeagus.
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
. Small and truncated apically; anal papilla with posterior apophysis
<normalizedToken originalValue="¼">1/4</normalizedToken>
<normalizedToken originalValue="×">x</normalizedToken>
shorter than anal papilla; anterior apophysis short; rectangular sterigma that is fused above ostium; ductus bursae wide and long; appendix bursae ⅔
<normalizedToken originalValue="×">x</normalizedToken>
shorter than corpus bursae which is partially sclerotized at base.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="immature stages">
<paragraph pageId="0" pageNumber="71">Immature stages.</paragraph>
<paragraph pageId="0" pageNumber="71">Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="distribution">
<paragraph pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="71">
Recorded from cloud forests in Colombia, Ecuador, Peru, and Bolivia in middle and high elevations from 2000 m to above 3000 m (Fig.
<figureCitation captionStart="Figure 95" captionStartId="F17" captionText="Figure 95. Distribution of examined specimens of the genus Oculicattus." figureDoi="10.3897/zookeys.985.51622.figure95" httpUri="https://binary.pensoft.net/fig/471617" pageId="0" pageNumber="71">95</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="biology_ecology">
<paragraph pageId="0" pageNumber="71">Biology.</paragraph>
<paragraph pageId="0" pageNumber="71">Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="remarks">
<paragraph pageId="0" pageNumber="71">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="71">
A specimen of
<taxonomicName authorityName="Martinez" authorityYear="2020" class="Insecta" family="Noctuidae" genus="Oculicattus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Oculicattus raizae" order="Lepidoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="raizae">
<emphasis italics="true" pageId="0" pageNumber="71">Oculicattus raizae</emphasis>
</taxonomicName>
was misidentified as
<taxonomicName lsidName="G. renifera" pageId="0" pageNumber="71" rank="species" species="renifera">
<emphasis italics="true" pageId="0" pageNumber="71">G. renifera</emphasis>
</taxonomicName>
by G. Behounek and posted on Barcode Of Life Data System v4 (www.barcodinglife.org). Holotype (Fig.
<figureCitation captionStart="Figures 43–58" captionStartId="F6" captionText="Figures 43 - 58. Adult habitus of Oculicattus species 43 Oculicattus renifera, ♂, MGCL, Cusco, Peru 44 O. renifera, ♂, MGCL, Junin, Peru 45 O. schmidti, ♂, holotype, MGCL, Pasco, Peru 46 O. schmidti, ♂, paratype, MGCL, Junin, Peru 47 O. inca, ♂, holotype, MGCL, Sierra Siberia, Bolivia 48 O. boliviana, ♂, holotype, MGCL, Sierra Siberia, Bolivia 49 O. uturunku, ♂, holotype, MGCL, Morona, Ecuador 50 O. raizae, ♂, holotype, MGCL, Tolima, Colombia 51 O. raizae, ♂, paratype, ZSM, Pichincha, Ecuador 52 O. brehmi, ♂, holotype, FSU, Loja, Ecuador 53 O. brehmi, ♂, paratype, MGCL, Napo, Ecuador 54 O. brehmi, ♀, paratype, MGCL, Napo, Ecuador 55 O. brehmi, ♀, paratype, MGCL, Napo, Ecuador 56 O. renifera, ♀, MGCL, Cusco, Peru 57 O. renifera, ♀, MGCL, Cusco, Peru 58 O. raizae, ♀, paratype, MGCL, La Paz, Bolivia." figureDoi="10.3897/zookeys.985.51622.figures43-58" httpUri="https://binary.pensoft.net/fig/471606" pageId="0" pageNumber="71">50</figureCitation>
) and paratypes are well preserved.
</paragraph>
</subSubSection>
</treatment>
</document>
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