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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.530.6047" ID-GBIF-Dataset="ee8a20f5-fcd1-46eb-ab80-99d4f61ac046" ID-PMC="PMC4668906" ID-Pensoft-Pub="1313-2970-530-37" ID-PubMed="26692799" ID-ZBK="6B52F31401BE48EE9C400A467D24040E" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-530-37" ModsDocOrigin="ZooKeys 530" ModsDocTitle="Review of New Caledonian species of Oxyethira Eaton, with description of 17 new species, and new records for Hydroptila Dalman and Hellyethira Neboiss (Trichoptera, Hydroptilidae)" checkinTime="1451243854172" checkinUser="pensoft" docAuthor="Wells, Alice &amp; Johanson, Kjell Arne" docDate="2015" docId="1BF30168D63627799E79AB4E25665823" docLanguage="en" docName="ZooKeys 530: 37-90" docOrigin="ZooKeys 530" docSource="http://dx.doi.org/10.3897/zookeys.530.6047" docTitle="Oxyethira (Pacificotrichia) melasma Kelley" docType="treatment" docVersion="3" lastPageNumber="62" masterDocId="FFDCFFA8FFE22A19DE777A04FFC50560" masterDocTitle="Review of New Caledonian species of Oxyethira Eaton, with description of 17 new species, and new records for Hydroptila Dalman and Hellyethira Neboiss (Trichoptera, Hydroptilidae)" masterLastPageNumber="90" masterPageNumber="37" pageNumber="61" updateTime="1668160879621" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Review of New Caledonian species of Oxyethira Eaton, with description of 17 new species, and new records for Hydroptila Dalman and Hellyethira Neboiss (Trichoptera, Hydroptilidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wells, Alice</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Johanson, Kjell Arne</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2015</mods:date>
<mods:detail type="volume">
<mods:number>530</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>37</mods:start>
<mods:end>90</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.530.6047</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.530.6047</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-530-37</mods:identifier>
<mods:identifier type="ZBK">6B52F31401BE48EE9C400A467D24040E</mods:identifier>
<mods:identifier type="ZooBank">6B52F31401BE48EE9C400A467D24040E</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152062918" LSID="urn:lsid:plazi:treatment:1BF30168D63627799E79AB4E25665823" httpUri="http://treatment.plazi.org/id/1BF30168D63627799E79AB4E25665823" lastPageId="25" lastPageNumber="62" pageId="24" pageNumber="61">
<subSubSection pageId="24" pageNumber="61" type="multiple">
<paragraph pageId="24" pageNumber="61">Taxon classification Animalia Trichoptera Hydroptilidae</paragraph>
</subSubSection>
<subSubSection pageId="24" pageNumber="61" type="nomenclature">
<paragraph pageId="24" pageNumber="61">
<taxonomicName authority="Kelley" class="Insecta" family="Hydroptilidae" genus="Oxyethira" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oxyethira (Pacificotrichia) melasma" order="Trichoptera" pageId="24" pageNumber="61" phylum="Arthropoda" rank="species" species="melasma" subGenus="Pacificotrichia">Oxyethira (Pacificotrichia) melasma Kelley</taxonomicName>
Figs 55-57, 105
</paragraph>
</subSubSection>
<subSubSection pageId="25" pageNumber="62" type="reference_group">
<paragraph pageId="25" pageNumber="62">
<taxonomicName class="Insecta" family="Hydroptilidae" genus="Oxyethira" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oxyethira melasma" order="Trichoptera" pageId="25" pageNumber="62" phylum="Arthropoda" rank="species" species="melasma">
<pageBreakToken pageId="25" pageNumber="62" start="start">Oxyethira</pageBreakToken>
melasma
</taxonomicName>
Kelley, 1989: 200, figs 37, 45, 59.
</paragraph>
</subSubSection>
<subSubSection pageId="25" pageNumber="62" type="note">
<paragraph pageId="25" pageNumber="62">Note.</paragraph>
<paragraph pageId="25" pageNumber="62">
A mismatch between the published description and
<bibRefCitation author="Kelley, RW" journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="41" pageNumber="78" pagination="190 - 202" title="New species of micro-caddisflies (Trichoptera: Hydroptilidae) from New Caledonia, Vanuatu and Fiji." volume="91" year="1989">
<normalizedToken originalValue="Kelleys">Kelley's</normalizedToken>
(1989)
</bibRefCitation>
figures and the holotype specimen in the BPBM labelled &quot;
<taxonomicName class="Insecta" family="Hydroptilidae" genus="Oxyethira" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oxyethira melasma" order="Trichoptera" pageId="25" pageNumber="62" phylum="Arthropoda" rank="species" species="melasma">Oxyethira melasma</taxonomicName>
&quot; is a puzzle. The genital features of the holotype (examined by AW) are as in Fig. 56, and disagree with
<normalizedToken originalValue="Kelleys">Kelley's</normalizedToken>
text description and illustrations (his figs 37, 45, 59) of &quot;
<taxonomicName class="Insecta" family="Hydroptilidae" genus="Oxyethira" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oxyethira melasma" order="Trichoptera" pageId="25" pageNumber="62" phylum="Arthropoda" rank="species" species="melasma">Oxyethira melasma</taxonomicName>
&quot;. However, the tibial spur count of the holotype is 0,2,4, as given by Kelley, and as in the type species.
<normalizedToken originalValue="Kelleys">Kelley's</normalizedToken>
three figures of genital structures agree with the features of a series of specimens, described here as
<taxonomicName class="Insecta" family="Hydroptilidae" genus="Oxyethira" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oxyethira nehoue" order="Trichoptera" pageId="25" pageNumber="62" phylum="Arthropoda" rank="species" species="nehoue">Oxyethira nehoue</taxonomicName>
and illustrated in Figs 58-60; in contrast to
<normalizedToken originalValue="Kelleys">Kelley's</normalizedToken>
<normalizedToken originalValue="“holotype”">&quot;holotype&quot;</normalizedToken>
, these specimens all have a tibial spur count of 0,3,4.
</paragraph>
<paragraph pageId="25" pageNumber="62">
<taxonomicName class="Insecta" family="Hydroptilidae" genus="Oxyethira" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oxyethira melasma" order="Trichoptera" pageId="25" pageNumber="62" phylum="Arthropoda" rank="species" species="melasma">Oxyethira melasma</taxonomicName>
Kelley is here redescribed and figured from the holotype specimen. We can only suggest that Kelley had several specimens at hand, macerated one and labelled it
<normalizedToken originalValue="“holotype”">&quot;holotype&quot;</normalizedToken>
, but described and illustrated a specimen other than that labelled
<normalizedToken originalValue="“Type”">&quot;Type&quot;</normalizedToken>
. The type has to be the name-bearing specimen and is redescribed here.
</paragraph>
</subSubSection>
<subSubSection pageId="25" pageNumber="62" type="revised diagnosis">
<paragraph pageId="25" pageNumber="62">Revised diagnosis.</paragraph>
<paragraph pageId="25" pageNumber="62">
Males resemble
<taxonomicName class="Insecta" family="Hydroptilidae" genus="Oxyethira" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oxyethira nehoue" order="Trichoptera" pageId="25" pageNumber="62" phylum="Arthropoda" rank="species" species="nehoue">Oxyethira nehoue</taxonomicName>
sp. n.,
<taxonomicName class="Insecta" family="Hydroptilidae" genus="Oxyethira" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oxyethira spicula" order="Trichoptera" pageId="25" pageNumber="62" phylum="Arthropoda" rank="species" species="spicula">Oxyethira spicula</taxonomicName>
sp. n., and
<taxonomicName class="Insecta" family="Hydroptilidae" genus="Oxyethira" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oxyethira ouenghi" order="Trichoptera" pageId="25" pageNumber="62" phylum="Arthropoda" rank="species" species="ouenghi">Oxyethira ouenghi</taxonomicName>
in having have a more or less triangular median ventral processes in the male genitalia. However, they most closely resemble
<taxonomicName class="Insecta" family="Hydroptilidae" genus="Oxyethira" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oxyethira nehoue" order="Trichoptera" pageId="25" pageNumber="62" phylum="Arthropoda" rank="species" species="nehoue">Oxyethira nehoue</taxonomicName>
sp. n. from which they are distinguished by spur count 0,2,4, gonopods fused, in ventral view sharply tapered proximally, narrowly parallel-sided distally, rounded apically, apex of phallic apparatus rounded without apical spine, in contrast to
<taxonomicName class="Insecta" family="Hydroptilidae" genus="Oxyethira" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oxyethira nehoue" order="Trichoptera" pageId="25" pageNumber="62" phylum="Arthropoda" rank="species" species="nehoue">Oxyethira nehoue</taxonomicName>
sp. n. in which the fused gonopods appear triangular in ventral view, and acuminate apically, and ventral process in lateral view sclerotised and arched ventrally.
</paragraph>
</subSubSection>
<subSubSection pageId="25" pageNumber="62" type="revised description">
<paragraph pageId="25" pageNumber="62">Revised description</paragraph>
<paragraph pageId="25" pageNumber="62">male. Antennae with 19-26 flagellomeres, apical 3 flagellomeres pale, next 5 dark, then 13 pale and basal flagellomeres dark; anterior wing length 1.4-1.9 mm (n=10); tibial spurs 0,2,4; abdominal sternite VII with small sharp median spine, offset from margin.</paragraph>
<paragraph pageId="25" pageNumber="62">Male, genitalia (Figs 55-57). Abdominal segment VIII more or less conical, a deep broadly V-shaped excision apico-ventrally. Abdominal segment IX in lateral view broadly bell-shaped; in ventral view rounded proximally, tapered distally, a pair of small short processes apically, each bearing a single seta. Gonopods sclerotised, fused, tapered to narrowly truncate apex, basal setose processes widely separated, slender, elongate, but shorter than fused gonopods, a slender basal apodeme midventrally; subgenital plate broad based, tapered to rounded apex, mostly membranous, but with a small ventrally curved prominence subapically. Phallic apparatus swollen in basal third, narrow in distal 2/3, without apical spine, ejaculatory tube medial.</paragraph>
</subSubSection>
<subSubSection pageId="25" pageNumber="62" type="materials_examined">
<paragraph pageId="25" pageNumber="62">Material examined.</paragraph>
<paragraph pageId="25" pageNumber="62">
Holotype. Male, Mountain stream up Boulari River (BPBM). Other material. 18 males, Province Sud, W part of Plaine des lacs, 150 m downstream bridge at La Capture,
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-22.266117">22°15.967'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="166.82489">166°49.493'E</geoCoordinate>
, 261 m, 04-22.xi.2003, Malaise trap, loc#007, leg. K.A. Johanson (NHRS); 31males, Province Sud, Mt Dzumac, source stream of Ouinne River, near crosspoint to mountain track,
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-22.03455">22°02.073'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="166.47433">166°28.460'E</geoCoordinate>
, 810 m, 18.
<normalizedToken originalValue="xi">xi-</normalizedToken>
4.xii.2003, Malaise trap, loc#030, leg. K.A. Johanson (NHRS); 1 male, Province Sud, Mt Dzumac, source stream of Ouinne River, downstream crosspoint to mountain track,
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-22.033283">22°01.997'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="166.47476">166°28.486'E</geoCoordinate>
, 795 m, over about 30 m waterfall, 18.
<normalizedToken originalValue="xi">xi-</normalizedToken>
4.xii.2003, Malaise trap, loc#031, leg. K.A. Johanson (NHRS); 1 male (on slide), Province Sud, W slope Mt Ningua,
<normalizedToken originalValue="Kwé">Kwe</normalizedToken>
<normalizedToken originalValue="Néco">Neco</normalizedToken>
, Stream, at Camp Jacob, 3.7 km WNW summit of Mt Ningua, on
<normalizedToken originalValue="BoulouparisThio">Boulouparis-Thio</normalizedToken>
Road, about 50 m upstream road,
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-21.726883">21°43.613'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="166.10945">166°06.567'E</geoCoordinate>
, 150 m, 29.
<normalizedToken originalValue="xi">xi-</normalizedToken>
12.xii.2003, Malaise trap, loc#054, leg. K.A. Johanson (NHRS); 4 males, Province Sud, Co Rigule Stream, 2.1 km N bridge over Baie de
<normalizedToken originalValue="Yaté">Yate</normalizedToken>
, 4.3 km S
<normalizedToken originalValue="Wé">We</normalizedToken>
<normalizedToken originalValue="Ngéré">Ngere</normalizedToken>
,
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-22.135784">22°08.147'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="166.93454">166° 56.072'E</geoCoordinate>
, 14 m, 18.i.2004, light trap, loc#122, leg. K.A. Johanson (NHRS).
</paragraph>
</subSubSection>
<subSubSection pageId="25" pageNumber="62" type="remarks">
<paragraph pageId="25" pageNumber="62">Remarks.</paragraph>
<paragraph pageId="25" pageNumber="62">The species was taken at a number of sites in the southern province of the island (Fig. 105).</paragraph>
</subSubSection>
</treatment>
</document>