397 lines
38 KiB
XML
397 lines
38 KiB
XML
<document id="E730FC2794B47A2FF22771E291E9B0F5" ID-CLB-Dataset="29115" ID-DOI="10.11646/zootaxa.4455.1.3" ID-GBIF-Dataset="0d838ca1-52f1-4278-895d-68fc481b7558" ID-ISSN="1175-5326" ID-Zenodo-Dep="1456971" ID-ZooBank="2EA2EE4A-2BA4-438A-A1CB-D9C54F37DC78" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539260480029" checkinUser="plazi" docAuthor="Wüster, Wolfgang, Chirio, Laurent, Trape, Jean-François, Ineich, Ivan, Jackson, Kate, Greenbaum, Eli, Barron, Cesar, Kusamba, Chifundera, Nagy, Zoltán T., Storey, Richard, Hall, Cara, Wüster, Catharine E., Barlow, Axel & Broadley, Donald G." docDate="2018" docId="03ADDC33D479FA75FF59304CD7E1FD65" docLanguage="en" docName="zootaxa.4455.1.3.pdf" docOrigin="Zootaxa 4455 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Naja (Boulengerina) guineensis Wüster & Chirio & Trape & Ineich & Jackson & Greenbaum & Barron & Kusamba & Nagy & Storey & Hall & Wüster & Barlow & Broadley 2018, sp. nov." docType="treatment" docUuid="A7FFED1B-36FB-432A-ABEE-194346747D18" docUuidSource="ZooBank" docVersion="8" masterDocId="FF94A44BD472FA78FFCE3743D41AFF95" masterDocTitle="Integration of nuclear and mitochondrial gene sequences and morphology reveals unexpected diversity in the forest cobra (Naja melanoleuca) species complex in Central and West Africa (Serpentes: Elapidae)" updateTime="1698575068667" updateUser="ExternalLinkService">
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<mods:title id="8768F19367DDBE3341EA5C5E48FF114B">Integration of nuclear and mitochondrial gene sequences and morphology reveals unexpected diversity in the forest cobra (Naja melanoleuca) species complex in Central and West Africa (Serpentes: Elapidae)</mods:title>
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<mods:namePart id="804F5FF2BC3D4292D07B930FA9CCD3AE">Wüster, Wolfgang</mods:namePart>
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<mods:namePart id="1745BDC236F74EC4FE15990E230F84F8">Chirio, Laurent</mods:namePart>
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<mods:namePart id="BAB593E9275A1F2E54A277511A32C4D2">Trape, Jean-François</mods:namePart>
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<mods:namePart id="D32A7015265651B74EE1EB1F97E3D0FA">Ineich, Ivan</mods:namePart>
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<mods:namePart id="DCB8F2E3A6F205C25E6FF45226387258">Jackson, Kate</mods:namePart>
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<mods:namePart id="7D6496FF4BCDDD18BED6AC9AC3C1DB0A">Greenbaum, Eli</mods:namePart>
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<mods:namePart id="1891B14481915DDE9592E7BC6BA6F57C">Barron, Cesar</mods:namePart>
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<mods:namePart id="5F6829E0DE723587DF1C1CD7B2BFA477">Kusamba, Chifundera</mods:namePart>
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<mods:namePart id="5E436E77C6A2DCC3A8F818A5F9B27433">Nagy, Zoltán T.</mods:namePart>
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<mods:namePart id="87171D42D5D41C7718BAF8BCB4063C6A">Storey, Richard</mods:namePart>
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<mods:namePart id="38DDE9338DACF3A9C012EFD24445336A">Hall, Cara</mods:namePart>
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<mods:namePart id="43FE3717C9415E73BDD552FEED0FF225">Wüster, Catharine E.</mods:namePart>
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<mods:namePart id="EBBBF3B875937DF82C0B6FF85739E947">Barlow, Axel</mods:namePart>
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<mods:namePart id="E49622FE0F05A1565AF4AC5F2CE4C742">Broadley, Donald G.</mods:namePart>
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<mods:typeOfResource id="9E7432FAA1917C79BBE021EDBD180761">text</mods:typeOfResource>
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<mods:title id="CFC0A81263D9AF9259126DF749994F66">Zootaxa</mods:title>
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<mods:part id="21F78185DD806A01011761E723D1D9E2">
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<mods:date id="852BB83D155B0D05B8CF97DFE5F7DFBF">2018</mods:date>
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<mods:number id="B381C67F1BA8729248394BF0FF6221F0">2018-08-01</mods:number>
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<mods:detail id="6DCC7E366E55E1C85C57DD7BF3E9CFBE" type="volume">
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<mods:number id="8FB9BC80D4534FBED345CF7839861A82">4455</mods:number>
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<mods:number id="F09FDC5583DD4309FF920920F0E2F51D">1</mods:number>
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<mods:identifier id="89D0C73568AF304C33B1D617E789B532" type="DOI">10.11646/zootaxa.4455.1.3</mods:identifier>
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<mods:identifier id="FFD592F801820A895B3BD000490F0719" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="B6253E79F8694313EF8D09FEEB10B32A" type="Zenodo-Dep">1456971</mods:identifier>
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<treatment id="03ADDC33D479FA75FF59304CD7E1FD65" ID-DOI="http://doi.org/10.5281/zenodo.5950780" ID-GBIF-Taxon="148691101" ID-Zenodo-Dep="5950780" LSID="urn:lsid:zoobank.org:act:A7FFED1B-36FB-432A-ABEE-194346747D18" httpUri="http://treatment.plazi.org/id/03ADDC33D479FA75FF59304CD7E1FD65" lastPageId="13" pageId="11">
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<subSubSection id="C31E3EAED479FA73FF59304CD7AEF8F9" pageId="11" type="nomenclature">
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<paragraph id="8BBB6D25D479FA73FF59304CD095F8BF" blockId="11.[151,1167,1807,1834]" box="[151,1167,1807,1834]" pageId="11">
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<heading id="D0F3DA49D479FA73FF59304CD095F8BF" bold="true" box="[151,1167,1807,1834]" fontSize="11" level="1" pageId="11" reason="1">
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<emphasis id="B970B137D479FA73FF59304CD095F8BF" bold="true" box="[151,1167,1807,1834]" pageId="11">
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<taxonomicName id="4C0416A6D479FA73FF59304CD601F8BC" authority="Wüster & Chirio & Trape & Ineich & Jackson & Greenbaum & Barron & Kusamba & Nagy & Storey & Hall & Wüster & Barlow & Broadley, 2018" authorityName="Wüster & Chirio & Trape & Ineich & Jackson & Greenbaum & Barron & Kusamba & Nagy & Storey & Hall & Wüster & Barlow & Broadley" authorityYear="2018" box="[151,539,1807,1834]" class="Reptilia" family="Elapidae" genus="Naja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="11" phylum="Chordata" rank="species" species="guineensis" status="sp. nov." subGenus="Boulengerina">
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<emphasis id="B970B137D479FA73FF59304CD4C8F8BC" bold="true" box="[151,210,1807,1833]" italics="true" pageId="11">Naja</emphasis>
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(
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<emphasis id="B970B137D479FA73FF2D304CD593F8BC" bold="true" box="[227,393,1807,1833]" italics="true" pageId="11">Boulengerina</emphasis>
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)
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<emphasis id="B970B137D479FA73FE54304CD601F8BC" bold="true" box="[410,539,1807,1833]" italics="true" pageId="11">guineensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2430C4CD479FA73FDEC3053D665F8BF" box="[546,639,1808,1834]" pageId="11" rank="species">sp. nov.</taxonomicNameLabel>
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Broadley, Trape, Chirio, Ineich & Wüster
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BBB6D25D479FA73FF59301BD7AEF8F9" blockId="11.[151,1435,1878,2023]" box="[151,948,1878,1901]" pageId="11">
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<treatmentCitationGroup id="AB144A0BD479FA73FF59301BD7AEF8F9" box="[151,948,1878,1901]" pageId="11">
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<taxonomicName id="4C0416A6D479FA73FF59301BD555F8F9" authorityName="Hallowell" authorityYear="1857" box="[151,335,1878,1901]" class="Reptilia" family="Elapidae" genus="Naia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="11" phylum="Chordata" rank="species" species="melanoleuca">
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<emphasis id="B970B137D479FA73FF59301BD555F8F9" box="[151,335,1878,1901]" italics="true" pageId="11">Naia melanoleuca</emphasis>
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</taxonomicName>
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(not Hallowell)
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<treatmentCitation id="0AA54B34D479FA73FE363015D6C9F8F9" author="" box="[504,723,1878,1900]" page="376" pageId="11">
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<bibRefCitation id="EF9510D4D479FA73FE363015D6B9F8F9" box="[504,675,1878,1900]" pageId="11" refId="ref12241" refString="Boulenger, G. A. (1896) Catalogue of the Snakes in the British Museum (Natural History) III. British Museum (Natural History), London, 727 pp. https: // doi. org / 10.5962 / bhl. title. 54273" type="book">Boulenger, 1896</bibRefCitation>
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: 376
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</treatmentCitation>
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(part, var. B [c,d], C).
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31E3EAED479FA73FF593035D750F83F" pageId="11" type="reference_group">
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<paragraph id="8BBB6D25D479FA73FF593035D039F81E" blockId="11.[151,1435,1878,2023]" box="[151,1059,1909,1931]" pageId="11">
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<treatmentCitationGroup id="AB144A0BD479FA73FF593035D039F81E" box="[151,1059,1909,1931]" pageId="11">
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<taxonomicName id="4C0416A6D479FA73FF593035D4D3F81E" box="[151,201,1910,1931]" class="Reptilia" family="Elapidae" genus="Naja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="11" phylum="Chordata" rank="genus">
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<emphasis id="B970B137D479FA73FF593035D4D3F81E" box="[151,201,1910,1931]" italics="true" pageId="11">Naja</emphasis>
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</taxonomicName>
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sp. 2
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||
<taxonomicName id="4C0416A6D479FA73FECA3036D64AF81E" authority="Hallowell, 1857" authorityName="Hallowell" authorityYear="1857" box="[260,592,1909,1931]" class="Reptilia" family="Elapidae" genus="Naia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="11" phylum="Chordata" rank="species" species="melanoleuca">
|
||
cf.
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||
<emphasis id="B970B137D479FA73FEEB3036D5BDF81E" box="[293,423,1909,1931]" italics="true" pageId="11">melanoleuca</emphasis>
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<bibRefCitation id="EF9510D4D479FA73FE623036D64AF81E" box="[428,592,1909,1931]" pageId="11" refId="ref13793" refString="Hallowell, E. (1857) Notes of a collection of reptiles from the Gaboon country, West Africa, recently presented to the Academy of Natural Sciences of Philadelphia, by Dr. Henry A. Ford. Proceedings of Academy of Natural Sciences, Philadelphia, 9, 48 - 72." type="journal article">Hallowell, 1857</bibRefCitation>
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</taxonomicName>
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(blackish dorsum)
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||
<treatmentCitation id="0AA54B34D479FA73FCDA3035D005F81E" author="" box="[788,1055,1909,1931]" page="318" pageId="11">
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<bibRefCitation id="EF9510D4D479FA73FCDA3035D7F7F81E" box="[788,1005,1909,1931]" pageId="11" refId="ref16483" refString="Trape, J. - F. & Balde, C. (2014) A checklist of the snake fauna of Guinea, with taxonomic changes in the genera Philothamnus and Dipsadoboa (Colubridae) and a comparison with the snake fauna of some other West African countries. Zootaxa, 3900 (3), 301 - 338. https: // doi. org / 10.11646 / zootaxa. 3900.3.1" type="journal article">Trape & Baldé, 2014</bibRefCitation>
|
||
: 318
|
||
</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BBB6D25D479FA73FF5930D6D750F83F" blockId="11.[151,1435,1878,2023]" box="[151,842,1940,1962]" pageId="11">
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<treatmentCitationGroup id="AB144A0BD479FA73FF5930D6D750F83F" box="[151,842,1940,1962]" pageId="11">
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<taxonomicName id="4C0416A6D479FA73FF5930D6D4D3F83F" box="[151,201,1941,1962]" class="Reptilia" family="Elapidae" genus="Naja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="11" phylum="Chordata" rank="genus">
|
||
<emphasis id="B970B137D479FA73FF5930D6D4D3F83F" box="[151,201,1941,1962]" italics="true" pageId="11">Naja</emphasis>
|
||
</taxonomicName>
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||
sp. 2
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||
<taxonomicName id="4C0416A6D479FA73FECA30D7D5BDF83F" authorityName="Hallowell" authorityYear="1857" box="[260,423,1940,1962]" class="Reptilia" family="Elapidae" genus="Naia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="11" phylum="Chordata" rank="species" species="melanoleuca">
|
||
cf.
|
||
<emphasis id="B970B137D479FA73FEEB30D7D5BDF83F" box="[293,423,1940,1962]" italics="true" pageId="11">melanoleuca</emphasis>
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</taxonomicName>
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||
(forest form).
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||
<treatmentCitation id="0AA54B34D479FA73FDF530D6D75CF83F" author="" box="[571,838,1940,1962]" page="336" pageId="11">
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<bibRefCitation id="EF9510D4D479FA73FDF530D6D70EF83F" box="[571,788,1940,1962]" pageId="11" refId="ref16483" refString="Trape, J. - F. & Balde, C. (2014) A checklist of the snake fauna of Guinea, with taxonomic changes in the genera Philothamnus and Dipsadoboa (Colubridae) and a comparison with the snake fauna of some other West African countries. Zootaxa, 3900 (3), 301 - 338. https: // doi. org / 10.11646 / zootaxa. 3900.3.1" type="journal article">Trape & Baldé, 2014</bibRefCitation>
|
||
: 336
|
||
</treatmentCitation>
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.
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||
</treatmentCitationGroup>
|
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</paragraph>
|
||
</subSubSection>
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<subSubSection id="C31E3EAED479FA74FF5930F1D013FF3B" lastPageId="12" pageId="11" type="materials_examined">
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<paragraph id="8BBB6D25D479FA73FF5930F1D17DF872" blockId="11.[151,1435,1878,2023]" pageId="11">
|
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<typeStatus id="54BFD387D479FA73FF5930F1D4E8F85D" box="[151,242,1970,1992]" pageId="11" type="holotype">
|
||
<emphasis id="B970B137D479FA73FF5930F1D4E8F85D" box="[151,242,1970,1992]" italics="true" pageId="11">Holotype</emphasis>
|
||
</typeStatus>
|
||
:
|
||
<collectionCode id="ED15F5E0D479FA73FECE30F0D556F85D" box="[256,332,1971,1992]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="11">MNHN</collectionCode>
|
||
1921.0485, a male from N’Zébéla, Macenta Prefecture, Nzérékoré region of forested southeastern
|
||
<collectingCountry id="F3132DB5D479FA73FAE030F1D16FF85D" box="[1326,1397,1970,1992]" name="Guinea-Bissau" pageId="11">Guinea</collectingCountry>
|
||
(
|
||
<geoCoordinate id="EE300BE2D479FA73FA4830F1D4E6F872" direction="north" orientation="latitude" pageId="11" precision="925" value="8.083333">8° 05’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE300BE2D479FA73FEC83092D544F872" box="[262,350,2001,2023]" direction="west" orientation="longitude" pageId="11" precision="925" value="-9.083333">9° 05’W</geoCoordinate>
|
||
), elev.
|
||
<quantity id="4CFCC0C0D479FA73FE6D3092D5FEF872" box="[419,484,2001,2023]" metricMagnitude="2" metricUnit="m" metricValue="4.9" pageId="11" unit="m" value="490.0">490 m</quantity>
|
||
, Coll. Paul Chabanaud (
|
||
<date id="FFBA4BE5D479FA73FD1D3092D75FF872" box="[723,837,2001,2023]" pageId="11" value="1876" valueMax="1959">1876–1959</date>
|
||
) between
|
||
<date id="FFBA4BE5D479FA73FC653092D000F872" box="[939,1050,2001,2023]" pageId="11" value="1919" valueMax="1920">1919–1920</date>
|
||
(Chabanaud, 1921: 471) (
|
||
<figureCitation id="133F71A0D479FA73FAD03092D140F872" box="[1310,1370,2001,2023]" captionStart="FIGURE 5" captionStartId="12.[151,250,1409,1431]" captionTargetBox="[235,1352,511,1387]" captionTargetId="figure@12.[235,1352,511,1387]" captionTargetPageId="12" captionText="FIGURE 5. Naja (Boulengerina) guineensis sp. nov. Left and top right: holotype, MNHN 1921.0485, dorsal and ventral view and side view of head. Note extensive mottling of throat and anterior ventral side and limited posterior extent of lighter ventral markings. Bottom right: live adult specimen measuring approximately 200 cm total length, from Sekondi-Takoradi, Western Region, Ghana, displaying dark suffusion of throat and anterior venter (not preserved; photo L. Chirio)." httpUri="https://zenodo.org/record/1456981/files/figure.png" pageId="11">Fig. 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBB6D25D47EFA74FF5937DAD013FF3B" blockId="12.[151,1033,152,174]" box="[151,1033,152,174]" pageId="12">
|
||
<materialsCitation id="3B6C6778D47EFA74FF5937DAD013FF3B" ID-GBIF-Occurrence="1917075142" box="[151,1033,152,174]" collectionCode="BMNH" collectorName="Coll. C. T. & Pyne." country="Sierra Leone" location="Sierra Leone" pageId="12" specimenCode="BMNH 1960.1" specimenCount="1" typeStatus="paratype">
|
||
<typeStatus id="54BFD387D47EFA74FF5937DAD4EEFF3B" box="[151,244,153,174]" pageId="12" type="paratype">
|
||
<emphasis id="B970B137D47EFA74FF5937DAD4EEFF3B" box="[151,244,153,174]" italics="true" pageId="12">Paratype</emphasis>
|
||
</typeStatus>
|
||
:
|
||
<specimenCode id="DBA2C55ED47EFA74FECF37DAD581FF3B" box="[257,411,152,174]" collectionCode="BMNH" pageId="12">BMNH 1960.1</specimenCode>
|
||
.3.72, a male from Njala, Kori,
|
||
<collectingCountry id="F3132DB5D47EFA74FD1937DBD746FF3B" box="[727,860,152,174]" name="Sierra Leone" pageId="12">Sierra Leone</collectingCountry>
|
||
,
|
||
<collectorName id="26F108F3D47EFA74FCA837DBD013FF3B" box="[870,1033,152,174]" pageId="12">Coll. C.T. Pyne.</collectorName>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31E3EAED47EFA74FF593799D72CFE5E" pageId="12" type="diagnosis">
|
||
<paragraph id="8BBB6D25D47EFA74FF593799D72CFE5E" blockId="12.[151,1437,218,459]" pageId="12">
|
||
<emphasis id="B970B137D47EFA74FF593799D510FF66" bold="true" box="[151,266,218,243]" pageId="12">Diagnosis</emphasis>
|
||
.
|
||
<taxonomicName id="4C0416A6D47EFA74FED33798D5CFFF67" authorityName="Wüster & Chirio & Trape & Ineich & Jackson & Greenbaum & Barron & Kusamba & Nagy & Storey & Hall & Wüster & Barlow & Broadley" authorityYear="2018" box="[285,469,219,242]" class="Reptilia" family="Elapidae" genus="Naja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="12" phylum="Chordata" rank="species" species="guineensis">
|
||
<emphasis id="B970B137D47EFA74FED33798D5CFFF67" box="[285,469,219,242]" italics="true" pageId="12">Naja guineensis</emphasis>
|
||
</taxonomicName>
|
||
cAN bE distiNguishEd frOm thE pArtly sympAtric
|
||
<taxonomicName id="4C0416A6D47EFA74FBDB3798D0B4FF66" authority="Wüster & Chirio & Trape & Ineich & Jackson & Greenbaum & Barron & Kusamba & Nagy & Storey & Hall & Wüster & Barlow & Broadley, 2018" authorityName="Wüster & Chirio & Trape & Ineich & Jackson & Greenbaum & Barron & Kusamba & Nagy & Storey & Hall & Wüster & Barlow & Broadley" authorityYear="2018" box="[1045,1198,219,243]" class="Reptilia" family="Elapidae" genus="Naja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="12" phylum="Chordata" rank="species" species="savannula" status="sp. nov.">
|
||
<emphasis id="B970B137D47EFA74FBDB3798D0B4FF66" box="[1045,1198,219,243]" italics="true" pageId="12">N. savannula</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2430C4CD47EFA74FB743798D10DFF66" box="[1210,1303,219,243]" pageId="12" rank="species">sp. NOv.</taxonomicNameLabel>
|
||
by lAckiNg ExtENdEd dOrsAl bANdiNg, OftEN hAviNg 17 rAthEr thAN 19 dOrsAl scAlE rOws At midbOdy,
|
||
<collectingRegion id="49C0A3C7D47EFA74FBAD37BCD075FE82" box="[1123,1135,255,279]" country="Nigeria" name="Rivers" pageId="12">A</collectingRegion>
|
||
gENErAlly lOwEr subcAudAl scAlE cOuNt, fEwEr vENtrAl bANds,
|
||
<collectingRegion id="49C0A3C7D47EFA74FDD63660D63EFEAE" box="[536,548,291,315]" country="Nigeria" name="Rivers" pageId="12">A</collectingRegion>
|
||
lEssEr pOstEriOr ExtENt Of thE vENtrAl bANdiNg, ANd
|
||
<collectingRegion id="49C0A3C7D47EFA74FBBE3660D066FEAE" box="[1136,1148,291,315]" country="Nigeria" name="Rivers" pageId="12">A</collectingRegion>
|
||
strONg tENdENcy tOwArds mElANism iN Adults. SpEcimENs with 19 midbOdy dOrsAl scAlE rOws cAN bE distiNguishEd frOm
|
||
<taxonomicName id="4C0416A6D47EFA74FB263604D186FECB" authorityName="Hallowell" authorityYear="1857" box="[1256,1436,326,350]" class="Reptilia" family="Elapidae" genus="Naia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="12" phylum="Chordata" rank="species" species="melanoleuca">
|
||
<emphasis id="B970B137D47EFA74FB263604D186FECB" box="[1256,1436,326,350]" italics="true" pageId="12">N. melanoleuca</emphasis>
|
||
</taxonomicName>
|
||
thrOugh thE rEducEd NumbEr Of vENtrAl bANds, lEssEr pOstEriOr ExtENt Of bANdiNg ANd tENdENcy Of ONtOgENEtic mElANism frOm
|
||
<taxonomicName id="4C0416A6D47EFA74FE8236CCD5D4FE33" authorityName="LAurENt" authorityYear="1955" box="[332,462,398,422]" class="Reptilia" family="Elapidae" genus="Naja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="12" phylum="Chordata" rank="species" species="subfulva">
|
||
<emphasis id="B970B137D47EFA74FE8236CCD5D4FE33" box="[332,462,398,422]" italics="true" pageId="12">N. subfulva</emphasis>
|
||
</taxonomicName>
|
||
iN lAckiNg
|
||
<collectingRegion id="49C0A3C7D47EFA74FD9C36CCD644FE32" box="[594,606,399,423]" country="Nigeria" name="Rivers" pageId="12">A</collectingRegion>
|
||
lightEr ANtEriOr dOrsum ANd thrOugh ONtOgENEtic mElANism, ANd frOm
|
||
<taxonomicName id="4C0416A6D47EFA74FA4C36CCD530FE5E" class="Reptilia" family="Elapidae" genus="Naja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="12" phylum="Chordata" rank="species" species="peroescobari">
|
||
<emphasis id="B970B137D47EFA74FA4C36CCD530FE5E" italics="true" pageId="12">N. peroescobari</emphasis>
|
||
</taxonomicName>
|
||
iN hAviNg thE pOstEriOr
|
||
<collectingRegion id="49C0A3C7D47EFA74FDFA36F1D67EFE5E" box="[564,612,434,459]" country="Myanmar" name="Chin" pageId="12">chiN</collectingRegion>
|
||
shiElds iN cONtAct.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF7B3DADD47EFA74FF5932C2D0AAFA67" httpUri="https://zenodo.org/record/1456981/files/figure.png" pageId="12" startId="12.[151,250,1409,1431]" targetBox="[235,1352,511,1387]" targetPageId="12">
|
||
<paragraph id="8BBB6D25D47EFA74FF5932C2D0AAFA67" blockId="12.[151,1436,1409,1522]" pageId="12">
|
||
<emphasis id="B970B137D47EFA74FF5932C2D509FA02" bold="true" box="[151,275,1409,1431]" pageId="12">FIGURE 5.</emphasis>
|
||
<taxonomicName id="4C0416A6D47EFA74FED732C1D644FA02" authority="Wüster & Chirio & Trape & Ineich & Jackson & Greenbaum & Barron & Kusamba & Nagy & Storey & Hall & Wüster & Barlow & Broadley, 2018" authorityName="Wüster & Chirio & Trape & Ineich & Jackson & Greenbaum & Barron & Kusamba & Nagy & Storey & Hall & Wüster & Barlow & Broadley" authorityYear="2018" box="[281,606,1409,1431]" class="Reptilia" family="Elapidae" genus="Naja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="12" phylum="Chordata" rank="species" species="guineensis" status="sp. nov." subGenus="Boulengerina">
|
||
<emphasis id="B970B137D47EFA74FED732C1D550FA02" box="[281,330,1410,1431]" italics="true" pageId="12">Naja</emphasis>
|
||
(
|
||
<emphasis id="B970B137D47EFA74FE9432C2D5FEFA02" box="[346,484,1409,1431]" italics="true" pageId="12">Boulengerina</emphasis>
|
||
)
|
||
<emphasis id="B970B137D47EFA74FE3D32C1D644FA02" box="[499,606,1410,1431]" italics="true" pageId="12">guineensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2430C4CD47EFA74FDAA32C2D6AEFA02" box="[612,692,1409,1431]" pageId="12" rank="species">
|
||
<emphasis id="B970B137D47EFA74FDAA32C2D6AEFA02" bold="true" box="[612,692,1409,1431]" pageId="12">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
Left and top right: holotype, MNHN 1921.0485, dorsal and ventral view and side view of head. Note extensive mottling of throat and anterior ventral side and limited posterior extent of lighter ventral markings. Bottom right: live adult specimen measuring approximately 200 cm total length, from Sekondi-Takoradi, Western Region, Ghana, displaying dark suffusion of throat and anterior venter (not preserved; photo L. Chirio).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C31E3EAED47EFA75FF093159D72BFE60" lastPageId="13" pageId="12" type="description">
|
||
<paragraph id="8BBB6D25D47EFA74FF093159D7F8F9C2" blockId="12.[151,1437,1562,2022]" pageId="12">
|
||
<emphasis id="B970B137D47EFA74FF093159D5FFF9A6" bold="true" box="[199,485,1562,1587]" pageId="12">
|
||
Description of
|
||
<typeStatus id="54BFD387D47EFA74FEB93159D5FAF9A6" box="[375,480,1562,1587]" pageId="12" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
DimENsiONs:
|
||
<collectingRegion id="49C0A3C7D47EFA74FD4E3159D68EF9A7" box="[640,660,1562,1586]" country="Nigeria" name="Rivers" pageId="12">A</collectingRegion>
|
||
mAlE spEcimEN with dissEctEd tAil bAsE, sNOut–vENt lENgth
|
||
<quantity id="4CFCC0C0D47EFA74FAE53159D182F9A7" box="[1323,1432,1562,1587]" metricMagnitude="0" metricUnit="m" metricValue="1.85" pageId="12" unit="mm" value="1850.0">1850 mm</quantity>
|
||
, tAil lENgth
|
||
<quantity id="4CFCC0C0D47EFA74FEC0317CD574F9C3" box="[270,366,1599,1623]" metricMagnitude="-1" metricUnit="m" metricValue="3.7" pageId="12" unit="mm" value="370.0">370 mm</quantity>
|
||
, tOtAl lENgth
|
||
<quantity id="4CFCC0C0D47EFA74FE31317CD676F9C3" box="[511,620,1599,1623]" metricMagnitude="0" metricUnit="m" metricValue="2.2199999999999998" pageId="12" unit="mm" value="2220.0">2220 mm</quantity>
|
||
, rAtiO tOtAl lENgth: tAil lENgth 6:1.
|
||
</paragraph>
|
||
<paragraph id="8BBB6D25D47EFA74FF093121D7D6F90A" blockId="12.[151,1437,1562,2022]" pageId="12">BOdy scAlAtiON: 23 scAlE rOws ArOuNd hOOd, 17 ArOuNd midbOdy, 13 ONE hEAd lENgth AhEAd Of thE vENt, All smOOth ANd ObliquE; 208 vENtrAls, 62 subcAudAls, All dividEd, ANAl siNglE.</paragraph>
|
||
<paragraph id="8BBB6D25D47EFA74FF0931EFD624F8C3" blockId="12.[151,1437,1562,2022]" pageId="12">
|
||
HEAd scAlAtiON: 7/7 suprAlAbiAls, 3rd & 4th cONtAct Orbit, 6th lArgEst; 8/8 iNfrAlAbiAls, first 4 cONtAct ANtEriOr
|
||
<collectingRegion id="49C0A3C7D47EFA74FAA231EFD186F950" box="[1388,1436,1708,1733]" country="Myanmar" name="Chin" pageId="12">chiN</collectingRegion>
|
||
shiElds; ANtEriOr pAir Of
|
||
<collectingRegion id="49C0A3C7D47EFA74FE6E3192D5CAF97F" box="[416,464,1745,1770]" country="Myanmar" name="Chin" pageId="12">chiN</collectingRegion>
|
||
shiElds ANd iNfrAlAbiAls 3–4 with AN ANOmAly ON bOth sidEs, thEy ArE wArty ANd cOvErEd with circumvOlutiONs; 1/1 prEOculAr, twicE As lONg As high; 3/3 pOstOculArs; 1/1 ANtEriOr tEmpOrAl; pOstEriOr tEmpOrAls 3/3; rOstrAl brOAdEr thAN high ANd slightly dAmAgEd ANtEriOrly, clEArly visiblE frOm AbOvE; 7 tEmpOrAls ANd NuchAls cONtActiNg bOth pAriEtAls.
|
||
</paragraph>
|
||
<paragraph id="8BBB6D25D47EFA75FF093022D0B2FF6D" blockId="12.[151,1437,1562,2022]" lastBlockId="13.[151,1437,151,752]" lastPageId="13" pageId="12">
|
||
PAttErN: uppEr sidE Of hEAd ANd uppEr tEmpOrAl rEgiON dArk brOwN iNcludiNg pAriEtAls ANd tEmpOrAls but blAck bEhiNd thEm, lips ANd vENtrAl sidE Of hEAd crEAmy yEllOwish withOut dArk mOttliNg ON
|
||
<collectingRegion id="49C0A3C7D47EFA74FB9830C6D09CF80B" box="[1110,1158,1925,1950]" country="Myanmar" name="Chin" pageId="12">chiN</collectingRegion>
|
||
ANd ANtEriOr thrOAt; EAch pOstEriOr suprAlAbiAl EdgE brOAdly OutliNEd iN blAck frOm EyE lEvEl
|
||
<collectingRegion id="49C0A3C7D47EFA74FC4C30EAD782F854" box="[898,920,1961,1985]" country="South Sudan" name="Lakes" pageId="12">tO</collectingRegion>
|
||
mOuth cOrNEr; tEmpOrAls uNifOrmly brOwNish. DOrsum uNifOrm brOwNish-blAck thrOughOut. VENtEr: ANtEriOr first
|
||
<quantity id="4CFCC0C0D47EFA74FC51308ED7F3F870" box="[927,1001,1997,2021]" metricMagnitude="-1" metricUnit="m" metricValue="3.2" pageId="12" unit="cm" value="32.0">32 cm</quantity>
|
||
(14% Of tOtAl lENgth) Of vENtEr with AltErNAtiNg yEllOw ANd blAckish irrEgulAr bANds, thE pOstEriOr yEllOw bANds iNcrEAsiNgly suffusEd with dArk mOttliNg. ONly vENtrAl 1 is uNifOrm clEAr yEllOw, vENtrAls 2–10, 19, 28–29, 32–35 yEllOw but AlwAys iNcrEAsiNgly suffusEd with dArk mOttliNg, rEmAiNiNg vENtrAls ANd uNdErsidE Of tAil blAck (AftEr vENtrAl 35).
|
||
</paragraph>
|
||
<paragraph id="8BBB6D25D47FFA75FF093647D765FE1D" blockId="13.[151,1437,151,752]" pageId="13">
|
||
<emphasis id="B970B137D47FFA75FF093647D52FFE88" bold="true" box="[199,309,260,285]" pageId="13">Variation</emphasis>
|
||
. MidbOdy dOrsAl scAlE rOw cOuNts Of 17 ANd 19 ArE ApprOximAtEly EquAlly cOmmON iN this spEciEs. IN lArgE Adults, light pAttErN ElEmENts ON thE hEAd ANd thrOAt OftEN bEcOmE hEAvily suffusEd with blAck pigmENt, lEAdiNg
|
||
<collectingRegion id="49C0A3C7D47FFA75FF3C360FD512FEF1" box="[242,264,332,356]" country="South Sudan" name="Lakes" pageId="13">tO</collectingRegion>
|
||
<collectingRegion id="49C0A3C7D47FFA75FEDE360FD506FEF1" box="[272,284,332,356]" country="Nigeria" name="Rivers" pageId="13">A</collectingRegion>
|
||
virtuAlly ENtirEly mElANistic sNAkE. SOmE spEcimENs hAvE 1–4 gENErAlly fAiNt Or pOOrly dEfiNEd light bANds AcrOss thE NEck, ANd OccAsiONAlly AN OcEllAtE hOOd mArkiNg.
|
||
</paragraph>
|
||
<paragraph id="8BBB6D25D47FFA75FF0936D7D72BFE60" blockId="13.[151,1437,151,752]" pageId="13">
|
||
<emphasis id="B970B137D47FFA75FF0936D7D585FE38" bold="true" box="[199,415,404,429]" pageId="13">Largest recorded:</emphasis>
|
||
1818+437 =
|
||
<quantity id="4CFCC0C0D47FFA75FDF736D7D6BDFE39" box="[569,679,404,429]" metricMagnitude="0" metricUnit="m" metricValue="2.255" pageId="13" unit="mm" value="2255.0">2255 mm</quantity>
|
||
, frOm BAllAssOu,
|
||
<collectingCountry id="F3132DB5D47FFA75FCA136D7D7DAFE38" box="[879,960,404,429]" name="Guinea-Bissau" pageId="13">GuiNEA</collectingCountry>
|
||
(IRD 4213.G), but lArgEr spEcimENs hAvE bEEN ObsErvEd. BAsEd ON lOcAlity, MENziEs’ (1966) rEpOrt Of
|
||
<collectingRegion id="49C0A3C7D47FFA75FC8B36FBD74BFE45" box="[837,849,440,464]" country="Nigeria" name="Rivers" pageId="13">A</collectingRegion>
|
||
spEcimEN mEAsuriNg “Eight fEEt, Eight iNchEs” (
|
||
<quantity id="4CFCC0C0D47FFA75FABF36F4D4A1FE61" metricMagnitude="0" metricUnit="m" metricValue="2.6399999999999997" pageId="13" unit="cm" value="264.0">264 cm</quantity>
|
||
) frOm BO,
|
||
<collectingCountry id="F3132DB5D47FFA75FEFB369FD5D3FE61" box="[309,457,476,501]" name="Sierra Leone" pageId="13">SiErrA LEONE</collectingCountry>
|
||
, is likEly
|
||
<collectingRegion id="49C0A3C7D47FFA75FDFA369FD650FE61" box="[564,586,476,500]" country="South Sudan" name="Lakes" pageId="13">tO</collectingRegion>
|
||
rEfEr
|
||
<collectingRegion id="49C0A3C7D47FFA75FD42369FD6B8FE61" box="[652,674,476,500]" country="South Sudan" name="Lakes" pageId="13">tO</collectingRegion>
|
||
this spEciEs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31E3EAED47FFA75FF0936BCD6C2FD10" pageId="13" type="etymology">
|
||
<paragraph id="8BBB6D25D47FFA75FF0936BCD567FDF5" blockId="13.[151,1437,151,752]" pageId="13">
|
||
<emphasis id="B970B137D47FFA75FF0936BCD55CFD8D" bold="true" box="[199,326,511,536]" pageId="13">Etymology</emphasis>
|
||
. ThE spEcific EpithEt
|
||
<taxonomicName id="4C0416A6D47FFA75FDF03543D6AEFD82" authorityName="Wüster & Chirio & Trape & Ineich & Jackson & Greenbaum & Barron & Kusamba & Nagy & Storey & Hall & Wüster & Barlow & Broadley" authorityYear="2018" box="[574,692,512,535]" class="Reptilia" family="Elapidae" genus="Naja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="guineensis">
|
||
<emphasis id="B970B137D47FFA75FDF03543D6AEFD82" box="[574,692,512,535]" italics="true" pageId="13">guineensis</emphasis>
|
||
</taxonomicName>
|
||
mEANs “frOm
|
||
<collectingCountry id="F3132DB5D47FFA75FC9736BCD7B4FD8D" box="[857,942,511,536]" name="Guinea-Bissau" pageId="13">GuiNEA</collectingCountry>
|
||
” ANd is chOsEN
|
||
<collectingRegion id="49C0A3C7D47FFA75FBAC3543D062FD8D" box="[1122,1144,512,536]" country="South Sudan" name="Lakes" pageId="13">tO</collectingRegion>
|
||
rEflEct thE distributiON Of thE spEciEs iN thE UppEr
|
||
<collectingCountry id="F3132DB5D47FFA75FE0E3567D60BFDA8" box="[448,529,548,573]" name="Guinea-Bissau" pageId="13">GuiNEA</collectingCountry>
|
||
fOrEsts Of
|
||
<collectingRegion id="49C0A3C7D47FFA75FD5B3566D6D6FDA9" box="[661,716,549,572]" country="Tanzania" name="Mjini Magharibi" pageId="13">WEst</collectingRegion>
|
||
AfricA, pArt Of thE
|
||
<collectingRegion id="49C0A3C7D47FFA75FC773566D7EAFDA9" box="[953,1008,549,572]" country="Tanzania" name="Mjini Magharibi" pageId="13">WEst</collectingRegion>
|
||
AfricAN FOrEsts biOdivErsity hOtspOt (MyErs
|
||
<emphasis id="B970B137D47FFA75FF21350AD53DFDF5" box="[239,295,584,608]" italics="true" pageId="13">et al.</emphasis>
|
||
, 2000).
|
||
</paragraph>
|
||
<paragraph id="8BBB6D25D47FFA75FF09352FD6C2FD10" blockId="13.[151,1437,151,752]" box="[199,728,620,645]" pageId="13">
|
||
<emphasis id="B970B137D47FFA75FF09352FD5EEFD10" bold="true" box="[199,500,620,645]" pageId="13">Suggested common name</emphasis>
|
||
. BlAck fOrEst cObrA.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31E3EAED47FFA75FF0935CCD7E1FD65" pageId="13" type="distribution">
|
||
<paragraph id="8BBB6D25D47FFA75FF0935CCD7E1FD65" blockId="13.[151,1437,151,752]" pageId="13">
|
||
<emphasis id="B970B137D47FFA75FF0935CCD543FD3D" bold="true" box="[199,345,655,680]" pageId="13">Distribution</emphasis>
|
||
. ThE distributiON Of
|
||
<taxonomicName id="4C0416A6D47FFA75FD9E35D3D712FD32" authorityName="Wüster & Chirio & Trape & Ineich & Jackson & Greenbaum & Barron & Kusamba & Nagy & Storey & Hall & Wüster & Barlow & Broadley" authorityYear="2018" box="[592,776,656,679]" class="Reptilia" family="Elapidae" genus="Naja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="guineensis">
|
||
<emphasis id="B970B137D47FFA75FD9E35D3D712FD32" box="[592,776,656,679]" italics="true" pageId="13">Naja guineensis</emphasis>
|
||
</taxonomicName>
|
||
AppEArs
|
||
<collectingRegion id="49C0A3C7D47FFA75FCBA35D3D790FD3D" box="[884,906,656,680]" country="South Sudan" name="Lakes" pageId="13">tO</collectingRegion>
|
||
bE rEstrictEd
|
||
<collectingRegion id="49C0A3C7D47FFA75FBE335D3D059FD3D" box="[1069,1091,656,680]" country="South Sudan" name="Lakes" pageId="13">tO</collectingRegion>
|
||
thE UppEr
|
||
<collectingCountry id="F3132DB5D47FFA75FB0335CCD104FD3D" box="[1229,1310,655,680]" name="Guinea-Bissau" pageId="13">GuiNEA</collectingCountry>
|
||
FOrEsts Of
|
||
<collectingRegion id="49C0A3C7D47FFA75FF5935F6D4F4FD59" box="[151,238,693,716]" country="Sierra Leone" name="Western" pageId="13">wEstErN</collectingRegion>
|
||
AfricA, frOm
|
||
<collectingRegion id="49C0A3C7D47FFA75FE4435F6D5FBFD59" box="[394,481,693,716]" country="Sierra Leone" name="Western" pageId="13">wEstErN</collectingRegion>
|
||
<collectingCountry id="F3132DB5D47FFA75FE2635F7D63BFD59" box="[488,545,692,716]" name="Togo" pageId="13">TOgO</collectingCountry>
|
||
<collectingRegion id="49C0A3C7D47FFA75FDE735F7D625FD59" box="[553,575,692,716]" country="South Sudan" name="Lakes" pageId="13">tO</collectingRegion>
|
||
<collectingCountry id="F3132DB5D47FFA75FD8935F7D682FD58" box="[583,664,692,717]" name="Liberia" pageId="13">LibEriA</collectingCountry>
|
||
ANd
|
||
<collectingCountry id="F3132DB5D47FFA75FD0135F7D73AFD58" box="[719,800,692,717]" name="Guinea-Bissau" pageId="13">GuiNEA</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EF9510D4D47FFA75FCE035F7D03BFD59" box="[814,1057,692,717]" pageId="13" refId="ref16483" refString="Trape, J. - F. & Balde, C. (2014) A checklist of the snake fauna of Guinea, with taxonomic changes in the genera Philothamnus and Dipsadoboa (Colubridae) and a comparison with the snake fauna of some other West African countries. Zootaxa, 3900 (3), 301 - 338. https: // doi. org / 10.11646 / zootaxa. 3900.3.1" type="journal article">TrApE & BAldé, 2014</bibRefCitation>
|
||
) (
|
||
<figureCitation id="133F71A0D47FFA75FBF635F7D067FD58" box="[1080,1149,692,717]" captionStart="FIGURE 6" captionStartId="13.[151,250,1552,1574]" captionTargetBox="[157,1421,845,1502]" captionTargetId="figure@13.[151,1436,807,1530]" captionTargetPageId="13" captionText="FIGURE 6. Distribution of the five species of the N. melanoleuca complex. A full list of voucher specimens beyond those included in multivariate analyses and Appendix 2 is available on request from the first author." httpUri="https://zenodo.org/record/1456983/files/figure.png" pageId="13">Fig. 6</figureCitation>
|
||
). ThErE is
|
||
<collectingRegion id="49C0A3C7D47FFA75FB3835F7D118FD59" box="[1270,1282,692,716]" country="Nigeria" name="Rivers" pageId="13">A</collectingRegion>
|
||
siNglE rEcOrd frOm CONtubOEl,
|
||
<collectingCountry id="F3132DB5D47FFA75FE993594D5B2FD65" box="[343,424,727,752]" name="Guinea-Bissau" pageId="13">GuiNEA</collectingCountry>
|
||
<collectingRegion id="49C0A3C7D47FFA75FE613594D5E0FD65" box="[431,506,727,752]" country="Guinea-Bissau" name="Bissau" pageId="13">BissAu</collectingRegion>
|
||
(MBL 535). All OthEr rEcOrds 10°N Or lOwEr.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |