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<document id="6F7F2D4E49F680276D34C6814D8B1342" ID-DOI="10.5852/ejt.2023.916.2381" ID-ISSN="2118-9773" ID-Zenodo-Dep="10453460" ID-ZooBank="0DC587F6-9DAA-4F6E-BA2A-AD528990BA24" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1704271610563" checkinUser="plazi" docAuthor="Wood, Thomas James" docDate="2023" docId="03ADDB11FFA7FF870A42FA69FDA4FCC7" docLanguage="en" docName="EJT.2023.916.1-85.pdf" docOrigin="European Journal of Taxonomy 916 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2381/10463" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Andrena (Micrandrena) anammas Wood 2023, sp. nov." docType="treatment" docUuid="F44C6808-4589-49ED-BD9B-D3FC6E1B91C5" docUuidSource="ZooBank" docVersion="2" lastPageNumber="50" masterDocId="FF94A369FF88FFB6085AFFA3FE1EFFB2" masterDocTitle="Revisions to the Andrena fauna of north-western Africa with a focus on Morocco (Hymenoptera: Andrenidae)" masterLastPageNumber="85" masterPageNumber="1" pageNumber="48" updateTime="1704272290945" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="FBD9AB4498B19A228DAB0023806C981F">Revisions to the Andrena fauna of north-western Africa with a focus on Morocco (Hymenoptera: Andrenidae)</mods:title>
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<mods:namePart id="02D4B9069E30C62B712A19664450FCFD">Wood, Thomas James</mods:namePart>
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<mods:title id="6A7F5228DD678632561CEF1EAECE8DC6">European Journal of Taxonomy</mods:title>
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<mods:date id="8CDA93B2368FF3D0AA91814DDE727472">2023</mods:date>
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<mods:identifier id="4E88E5580FA84B55D6AF2DEA753A8C8D" type="DOI">10.5852/ejt.2023.916.2381</mods:identifier>
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<treatment id="03ADDB11FFA7FF870A42FA69FDA4FCC7" LSID="urn:lsid:zoobank.org:act:F44C6808-4589-49ED-BD9B-D3FC6E1B91C5" httpUri="http://treatment.plazi.org/id/03ADDB11FFA7FF870A42FA69FDA4FCC7" lastPageId="49" lastPageNumber="50" pageId="47" pageNumber="48">
<subSubSection id="C31E398CFFA7FF990A42FA69FAD6F9BA" pageId="47" pageNumber="48" type="nomenclature">
<paragraph id="8BBB6A07FFA7FF990A42FA69FA02FA57" blockId="47.[363,1224,1482,1579]" box="[536,1052,1482,1509]" pageId="47" pageNumber="48">
<heading id="D0F3DD6BFFA7FF990A42FA69FA02FA57" box="[536,1052,1482,1509]" centered="true" fontSize="11" level="2" pageId="47" pageNumber="48" reason="2">
<taxonomicName id="4C041184FFA7FF990A42FA69FDA7FA57" authority="Wood, 2023" authorityName="Wood" authorityYear="2023" box="[536,953,1482,1509]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="anammas" status="sp. nov." subGenus="Micrandrena">
<emphasis id="B970B615FFA7FF990A42FA69FDA7FA57" bold="true" box="[536,953,1482,1509]" pageId="47" pageNumber="48">
<emphasis id="B970B615FFA7FF990A42FA69FC9CFA57" bold="true" box="[536,642,1482,1509]" italics="true" pageId="47" pageNumber="48">Andrena</emphasis>
(
<emphasis id="B970B615FFA7FF990AC9FA69FD2DFA57" bold="true" box="[659,819,1482,1509]" italics="true" pageId="47" pageNumber="48">Micrandrena</emphasis>
)
<emphasis id="B970B615FFA7FF990B1EFA68FDA7FA57" bold="true" box="[836,953,1483,1509]" italics="true" pageId="47" pageNumber="48">anammas</emphasis>
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2430B6EFFA7FF990B9BFA68FA02FA57" box="[961,1052,1483,1509]" pageId="47" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<uri id="FF956605FFA7FF990931FA4EFAD6F9BA" box="[363,1224,1517,1544]" pageId="47" pageNumber="48">
urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
</subSubSection>
<subSubSection id="C31E398CFFA7FF990AAAF9B2FD5DF999" box="[752,835,1553,1579]" pageId="47" pageNumber="48" type="description">
<paragraph id="8BBB6A07FFA7FF990AAAF9B2FD5DF999" blockId="47.[363,1224,1482,1579]" box="[752,835,1553,1579]" pageId="47" pageNumber="48">
<figureCitation id="133F7682FFA7FF990AAAF9B2FD5DF999" box="[752,835,1553,1579]" captionStart="Fig" captionStartId="49.[189,232,1882,1908]" captionTargetBox="[189,1399,947,1848]" captionTargetId="figure-175@49.[189,1399,947,1848]" captionTargetPageId="49" captionText="Fig. 28. Andrena (Micrandrena) anammas sp. nov., holotype, ♀ (OÖLM). A. Lateral habitus. B. Face, frontal view. C. Scutum, dorsal view. D. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10453558" httpUri="https://zenodo.org/record/10453558/files/figure.png" pageId="47" pageNumber="48">Fig. 28</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31E398CFFA7FF8608E7F9F1FAA8FDE9" lastPageId="48" lastPageNumber="49" pageId="47" pageNumber="48" type="diagnosis">
<paragraph id="8BBB6A07FFA7FF9908E7F9F1FF28F9DE" blockId="47.[189,310,1618,1644]" box="[189,310,1618,1644]" pageId="47" pageNumber="48">
<heading id="D0F3DD6BFFA7FF9908E7F9F1FF28F9DE" bold="true" box="[189,310,1618,1644]" fontSize="11" level="3" pageId="47" pageNumber="48" reason="3">
<emphasis id="B970B615FFA7FF9908E7F9F1FF28F9DE" bold="true" box="[189,310,1618,1644]" pageId="47" pageNumber="48">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BBB6A07FFA7FF8608E7F922FF5BFED8" blockId="47.[188,1399,1665,2008]" lastBlockId="48.[189,1398,265,362]" lastPageId="48" lastPageNumber="49" pageId="47" pageNumber="48">
<taxonomicName id="4C041184FFA7FF9908E7F922FF85F928" authority="Wood, 2023" authorityName="Wood" authorityYear="2023" box="[189,411,1665,1691]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="anammas" status="sp. nov.">
<emphasis id="B970B615FFA7FF9908E7F922FF85F928" box="[189,411,1665,1691]" italics="true" pageId="47" pageNumber="48">Andrena anammas</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2430B6EFFA7FF9909FCF922FC1DF929" box="[422,515,1665,1691]" pageId="47" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
can quickly be recognised as a member of
<taxonomicName id="4C041184FFA7FF990C42F922FAACF929" authorityName="Ashmead" authorityYear="1899" box="[1048,1202,1665,1691]" pageId="47" pageNumber="7" rank="subGenus" subGenus="Micrandrena">
<emphasis id="B970B615FFA7FF990C42F922FAACF929" box="[1048,1202,1665,1691]" italics="true" pageId="47" pageNumber="48">Micrandrena</emphasis>
</taxonomicName>
due to its small body size (
<figureCitation id="133F7682FFA7FF990961F907FFB9F90C" box="[315,423,1700,1726]" captionStart="Fig" captionStartId="49.[189,232,1882,1908]" captionTargetBox="[189,1399,947,1848]" captionTargetId="figure-175@49.[189,1399,947,1848]" captionTargetPageId="49" captionText="Fig. 28. Andrena (Micrandrena) anammas sp. nov., holotype, ♀ (OÖLM). A. Lateral habitus. B. Face, frontal view. C. Scutum, dorsal view. D. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10453558" httpUri="https://zenodo.org/record/10453558/files/figure.png" pageId="47" pageNumber="48">Fig. 28A</figureCitation>
), dark integument, and entirely rugose propodeal triangle. Due to the comparatively (for a
<taxonomicName id="4C041184FFA7FF990951F964FFBBF953" authorityName="Ashmead" authorityYear="1899" box="[267,421,1735,1761]" pageId="47" pageNumber="7" rank="subGenus" subGenus="Micrandrena">
<emphasis id="B970B615FFA7FF990951F964FFBBF953" box="[267,421,1735,1761]" italics="true" pageId="47" pageNumber="48">Micrandrena</emphasis>
</taxonomicName>
) long face and clypeus (
<figureCitation id="133F7682FFA7FF990A8FF964FD5DF953" box="[725,835,1735,1761]" captionStart="Fig" captionStartId="49.[189,232,1882,1908]" captionTargetBox="[189,1399,947,1848]" captionTargetId="figure-175@49.[189,1399,947,1848]" captionTargetPageId="49" captionText="Fig. 28. Andrena (Micrandrena) anammas sp. nov., holotype, ♀ (OÖLM). A. Lateral habitus. B. Face, frontal view. C. Scutum, dorsal view. D. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10453558" httpUri="https://zenodo.org/record/10453558/files/figure.png" pageId="47" pageNumber="48">Fig. 28B</figureCitation>
; head overall only 1.1 times as wide as long), narrow facial fovea (dorsally occupying ¼ of space between the lateral ocellus and the compound eye, consistently wide along its length), and densely punctate scutum (
<figureCitation id="133F7682FFA7FF990B9CF8AEFA2CF89A" box="[966,1074,1805,1832]" captionStart="Fig" captionStartId="49.[189,232,1882,1908]" captionTargetBox="[189,1399,947,1848]" captionTargetId="figure-175@49.[189,1399,947,1848]" captionTargetPageId="49" captionText="Fig. 28. Andrena (Micrandrena) anammas sp. nov., holotype, ♀ (OÖLM). A. Lateral habitus. B. Face, frontal view. C. Scutum, dorsal view. D. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10453558" httpUri="https://zenodo.org/record/10453558/files/figure.png" pageId="47" pageNumber="48">Fig. 28C</figureCitation>
; punctures separated by &lt;1 puncture diameter) they are comparable to
<taxonomicName id="4C041184FFA7FF990AE0F892FDCCF8F9" authority="Wood, 2021" authorityName="Wood" authorityYear="2021" box="[698,978,1841,1867]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="atlantea">
<emphasis id="B970B615FFA7FF990AE0F892FD25F8F9" box="[698,827,1841,1867]" italics="true" pageId="47" pageNumber="48">A. atlantea</emphasis>
Wood, 2021
</taxonomicName>
(High and Middle Atlas Mountains in
<collectingCountry id="F3132A97FFA7FF990886F8F7FF56F8DC" box="[220,328,1876,1902]" name="Morocco" pageId="47" pageNumber="48">Morocco</collectingCountry>
) and
<taxonomicName id="4C041184FFA7FF9909D2F8F7FFF9F8DC" authorityName="Wood" authorityYear="2023" box="[392,487,1876,1902]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="ortizi">
<emphasis id="B970B615FFA7FF9909D2F8F7FFF9F8DC" box="[392,487,1876,1902]" italics="true" pageId="47" pageNumber="48">A. ortizi</emphasis>
</taxonomicName>
(
<collectingCountry id="F3132A97FFA7FF9909ACF8F7FC23F8DC" box="[502,573,1876,1902]" name="Spain" pageId="47" pageNumber="48">Spain</collectingCountry>
, Sierra Nevada).
<taxonomicName id="4C041184FFA7FF990B53F8F7FDFAF8DC" authorityName="Wood" authorityYear="2023" box="[777,996,1876,1902]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="anammas">
<emphasis id="B970B615FFA7FF990B53F8F7FDFAF8DC" box="[777,996,1876,1902]" italics="true" pageId="47" pageNumber="48">Andrena anammas</emphasis>
</taxonomicName>
can be separated from
<taxonomicName id="4C041184FFA7FF990CACF8F7FB68F8DC" authorityName="Wood" authorityYear="2021" box="[1270,1398,1876,1902]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="atlantea">
<emphasis id="B970B615FFA7FF990CACF8F7FB68F8DC" box="[1270,1398,1876,1902]" italics="true" pageId="47" pageNumber="48">A. atlantea</emphasis>
</taxonomicName>
(with which it can be found in direct sympatry) by the stronger sculpture of the scutum and terga, these areas with clear microreticulation and at most weakly shining (in
<taxonomicName id="4C041184FFA7FF990C47F838FABFF806" authorityName="Wood" authorityYear="2021" box="[1053,1185,1946,1973]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="atlantea">
<emphasis id="B970B615FFA7FF990C47F838FABFF806" box="[1053,1185,1946,1973]" italics="true" pageId="47" pageNumber="48">A. atlantea</emphasis>
</taxonomicName>
with sculpture of the scutum and terga weaker, more strongly shining). Due to this stronger sculpture, the punctation of the tergal discs is weak and obscure (
<figureCitation id="133F7682FFB8FF860AF1FEAAFD05FE91" box="[683,795,265,291]" captionStart="Fig" captionStartId="49.[189,232,1882,1908]" captionTargetBox="[189,1399,947,1848]" captionTargetId="figure-175@49.[189,1399,947,1848]" captionTargetPageId="49" captionText="Fig. 28. Andrena (Micrandrena) anammas sp. nov., holotype, ♀ (OÖLM). A. Lateral habitus. B. Face, frontal view. C. Scutum, dorsal view. D. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10453558" httpUri="https://zenodo.org/record/10453558/files/figure.png" pageId="48" pageNumber="49">Fig. 28D</figureCitation>
), whereas in
<taxonomicName id="4C041184FFB8FF860BE5FEAAFA5DFE91" authorityName="Wood" authorityYear="2021" box="[959,1091,265,291]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="atlantea">
<emphasis id="B970B615FFB8FF860BE5FEAAFA5DFE91" box="[959,1091,265,291]" italics="true" pageId="48" pageNumber="49">A. atlantea</emphasis>
</taxonomicName>
it is more clearly visible against the underlying sculpture. Specimens of
<taxonomicName id="4C041184FFB8FF860A85FE8FFDA7FEF4" authorityName="Wood" authorityYear="2023" box="[735,953,300,326]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="anammas">
<emphasis id="B970B615FFB8FF860A85FE8FFDA7FEF4" box="[735,953,300,326]" italics="true" pageId="48" pageNumber="49">Andrena anammas</emphasis>
</taxonomicName>
are also slightly smaller (
<quantity id="4CFCC7E2FFB8FF860CB9FE8FFB38FEF5" box="[1251,1318,300,327]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="48" pageNumber="49" unit="mm" value="6.0">6 mm</quantity>
versus
<quantity id="4CFCC7E2FFB8FF8608E7FEECFF27FED8" box="[189,313,335,362]" metricMagnitude="-3" metricUnit="m" metricValue="7.75" metricValueMax="8.0" metricValueMin="7.5" pageId="48" pageNumber="49" unit="mm" value="7.75" valueMax="8.0" valueMin="7.5">7.5–8 mm</quantity>
).
</paragraph>
<paragraph id="8BBB6A07FFB8FF8608E7FE32FAA8FDE9" blockId="48.[188,1399,401,603]" pageId="48" pageNumber="49">
Finally,
<taxonomicName id="4C041184FFB8FF86097AFE32FFAAFE19" authority="Wood, 2023" authorityName="Wood" authorityYear="2023" box="[288,436,401,427]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="anammas" status="sp. nov.">
<emphasis id="B970B615FFB8FF86097AFE32FFAAFE19" box="[288,436,401,427]" italics="true" pageId="48" pageNumber="49">A. anammas</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2430B6EFFB8FF8609E5FE32FC03FE19" box="[447,541,401,427]" pageId="48" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
is most similar to
<emphasis id="B970B615FFB8FF860B5DFE32FD6EFE19" box="[775,880,401,427]" italics="true" pageId="48" pageNumber="49">
<taxonomicName id="4C041184FFB8FF860B5DFE32FD75FE19" authorityName="Wood" authorityYear="2023" box="[775,875,401,427]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="ortizi">A. ortizi</taxonomicName>
,
</emphasis>
which measures
<quantity id="4CFCC7E2FFB8FF860C1EFE32FA92FE19" box="[1092,1164,401,427]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="48" pageNumber="49" unit="mm" value="7.0">7 mm</quantity>
in length. The two species can be separated by the width of the tergal margins; in
<taxonomicName id="4C041184FFB8FF860BFFFE17FA28FE7C" authorityName="Wood" authorityYear="2023" box="[933,1078,436,462]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="anammas">
<emphasis id="B970B615FFB8FF860BFFFE17FA28FE7C" box="[933,1078,436,462]" italics="true" pageId="48" pageNumber="49">A. anammas</emphasis>
</taxonomicName>
, the tergal margins occupy at most ⅓ of the disc of T2 and ½ of the discs of T3–4, whereas in
<taxonomicName id="4C041184FFB8FF860B9FFE7BFA3DFE43" authorityName="Wood" authorityYear="2023" box="[965,1059,472,498]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="ortizi">
<emphasis id="B970B615FFB8FF860B9FFE7BFA3DFE43" box="[965,1059,472,498]" italics="true" pageId="48" pageNumber="49">A. ortizi</emphasis>
</taxonomicName>
the marginal areas occupy at least ½ of the disc of T2 and over ½ of the discs of T3–4. Additionally, in
<taxonomicName id="4C041184FFB8FF860C4DFE58FAB9FDA6" authorityName="Wood" authorityYear="2023" box="[1047,1191,507,533]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="anammas">
<emphasis id="B970B615FFB8FF860C4DFE58FAB9FDA6" box="[1047,1191,507,533]" italics="true" pageId="48" pageNumber="49">A. anammas</emphasis>
</taxonomicName>
the discs of T2–3 are obscurely punctate, with punctures disappearing into the background microreticulation (
<figureCitation id="133F7682FFB8FF860CA0FDBDFB77FD8A" box="[1274,1385,542,568]" captionStart="Fig" captionStartId="49.[189,232,1882,1908]" captionTargetBox="[189,1399,947,1848]" captionTargetId="figure-175@49.[189,1399,947,1848]" captionTargetPageId="49" captionText="Fig. 28. Andrena (Micrandrena) anammas sp. nov., holotype, ♀ (OÖLM). A. Lateral habitus. B. Face, frontal view. C. Scutum, dorsal view. D. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10453558" httpUri="https://zenodo.org/record/10453558/files/figure.png" pageId="48" pageNumber="49">Fig. 28D</figureCitation>
), whereas in
<taxonomicName id="4C041184FFB8FF860919FDE2FFBFFDE9" authorityName="Wood" authorityYear="2023" box="[323,417,577,603]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="ortizi">
<emphasis id="B970B615FFB8FF860919FDE2FFBFFDE9" box="[323,417,577,603]" italics="true" pageId="48" pageNumber="49">A. ortizi</emphasis>
</taxonomicName>
they are strongly and densely punctate (see figures in
<bibRefCitation id="EF9517F6FFB8FF860C44FDE2FAB4FDE9" author="Wood T. J." box="[1054,1194,577,603]" pageId="48" pageNumber="49" pagination="241 - 484" refId="ref41222" refString="Wood T. J. 2023. The genus Andrena in the Iberian Peninsula: revisionary taxonomy, ecology, new species, and a key to their identification (Hymenoptera: Andrenidae). Journal of Hymenoptera Research 96: 241 - 484. https: // doi. org / 10.3897 / jhr. 96.101873" type="journal article" year="2023">Wood 2023</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31E398CFFB8FF8608E7FD21FB69FD79" box="[189,1399,642,715]" pageId="48" pageNumber="49" type="etymology">
<paragraph id="8BBB6A07FFB8FF8608E7FD21FF5AFD2E" blockId="48.[189,324,642,668]" box="[189,324,642,668]" pageId="48" pageNumber="49">
<heading id="D0F3DD6BFFB8FF8608E7FD21FF5AFD2E" bold="true" box="[189,324,642,668]" fontSize="11" level="3" pageId="48" pageNumber="49" reason="3">
<emphasis id="B970B615FFB8FF8608E7FD21FF5AFD2E" bold="true" box="[189,324,642,668]" pageId="48" pageNumber="49">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BBB6A07FFB8FF8608E7FD12FB69FD79" blockId="48.[189,1399,689,715]" box="[189,1399,689,715]" pageId="48" pageNumber="49">Named after the Amazigh (Berber) name for the Middle Atlas (Atlas Anammas). It is a noun in apposition.</paragraph>
</subSubSection>
<subSubSection id="C31E398CFFB8FF8608E7FD51FAC6FBDC" pageId="48" pageNumber="49" type="materials_examined">
<paragraph id="8BBB6A07FFB8FF8608E7FD51FFB4FCBE" blockId="48.[189,426,754,780]" box="[189,426,754,780]" pageId="48" pageNumber="49">
<heading id="D0F3DD6BFFB8FF8608E7FD51FFB4FCBE" bold="true" box="[189,426,754,780]" fontSize="11" level="3" pageId="48" pageNumber="49" reason="3">
<emphasis id="B970B615FFB8FF8608E7FD51FFB4FCBE" bold="true" box="[189,426,754,780]" pageId="48" pageNumber="49">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BBB6A07FFB8FF8608B0FC82FF42FC89" blockId="48.[189,1399,801,900]" box="[234,348,801,827]" pageId="48" pageNumber="49">
<emphasis id="B970B615FFB8FF8608B0FC82FF42FC89" bold="true" box="[234,348,801,827]" pageId="48" pageNumber="49">
<typeStatus id="54BFD4A5FFB8FF8608B0FC82FF42FC89" box="[234,348,801,827]" pageId="48" pageNumber="49" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BBB6A07FFB8FF8608E7FCE7FA28FC36" blockId="48.[189,1399,801,900]" pageId="48" pageNumber="49">
<materialsCitation id="3B6C605AFFB8FF8608E7FCE7FA28FC36" collectingDate="2022-05-24" collectionCode="OOLM" collectorName="Ifrane, P &amp; T. J. Wood" country="Morocco" elevation="1900" latitude="33.4323" location="Fes-Meknes" longLatPrecision="6" longitude="-5.0629" pageId="48" pageNumber="49" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<collectingCountry id="F3132A97FFB8FF8608E7FCE7FF4FFCEC" box="[189,337,836,862]" name="Morocco" pageId="48" pageNumber="49">MOROCCO</collectingCountry>
•
<specimenCount id="9D02A18EFFB8FF86092FFCE6FF92FCED" box="[373,396,837,863]" count="1" pageId="48" pageNumber="49" type="female">♀</specimenCount>
;
<location id="8EDB3CDCFFB8FF8609FAFCE7FC2CFCEC" LSID="urn:lsid:plazi:treatment:03ADDB11FFA7FF870A42FA69FDA4FCC7:8EDB3CDCFFB8FF8609FAFCE7FC2CFCEC" box="[416,562,836,862]" country="Morocco" latitude="33.4323" longLatPrecision="6" longitude="-5.0629" name="Fes-Meknes" pageId="48" pageNumber="49">Fès-Meknès</location>
,
<collectorName id="26F10FD1FFB8FF860A19FCE7FCB0FCEC" box="[579,686,836,862]" pageId="48" pageNumber="49">Ifrane, P</collectorName>
7231,
<locationDeviation id="9A097325FFB8FF860AA7FCE7FA05FCEC" box="[765,1051,836,863]" location="Michlifen" pageId="48" pageNumber="49">
<quantity id="4CFCC7E2FFB8FF860AA7FCE7FD23FCEC" box="[765,829,836,862]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="48" pageNumber="49" unit="km" value="2.0">2 km</quantity>
NE of Michlifen
</locationDeviation>
;
<elevation id="00298D34FFB8FF860C76FCE7FAD1FCEC" box="[1068,1231,836,863]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="48" pageNumber="49" unit="m" value="1900.0">
<quantity id="4CFCC7E2FFB8FF860C76FCE7FA94FCED" box="[1068,1162,836,863]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="48" pageNumber="49" unit="m" value="1900.0">1900 m</quantity>
a.s.l.
</elevation>
;
<geoCoordinate id="EE300CC0FFB8FF860CB8FCE7FB6CFCED" box="[1250,1394,836,863]" degrees="33.4323" direction="north" orientation="latitude" pageId="48" pageNumber="49" precision="5" value="33.4323">33.4323° N</geoCoordinate>
,
<geoCoordinate id="EE300CC0FFB8FF8608E7FCC4FF20FC30" box="[189,318,871,898]" degrees="5.0629" direction="west" orientation="longitude" pageId="48" pageNumber="49" precision="5" value="-5.0629">5.0629° W</geoCoordinate>
;
<date id="FFBA4CC7FFB8FF860910FCC4FFF4FC33" box="[330,490,871,898]" pageId="48" pageNumber="49" value="2022-05-24">
<collectingDate id="EFFEB52FFFB8FF860910FCC4FFF4FC33" box="[330,490,871,898]" pageId="48" pageNumber="49" value="2022-05-24">24 May 2022</collectingDate>
</date>
;
<collectorName id="26F10FD1FFB8FF8609ACFCCBFC75FC33" box="[502,619,871,898]" pageId="48" pageNumber="49">T.J. Wood</collectorName>
leg.; BOLD: WPATW685-22;
<collectionCode id="ED15F2C2FFB8FF860B82FCCBFA2CFC36" box="[984,1074,872,900]" pageId="48" pageNumber="49">OÖLM</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BBB6A07FFB8FF8608B0FC0AFF42FC71" blockId="48.[189,1398,937,1033]" box="[234,348,937,963]" pageId="48" pageNumber="49">
<emphasis id="B970B615FFB8FF8608B0FC0AFF42FC71" bold="true" box="[234,348,937,963]" pageId="48" pageNumber="49">
<typeStatus id="54BFD4A5FFB8FF8608B0FC0AFF42FC71" box="[234,348,937,963]" pageId="48" pageNumber="49" type="paratype">Paratype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BBB6A07FFB8FF8608E7FC6FFC68FBBB" blockId="48.[189,1398,937,1033]" pageId="48" pageNumber="49">
<materialsCitation id="3B6C605AFFB8FF8608E7FC6FFC6FFBBB" collectingDate="2022-05-18" collectionCode="TJWC" collectorName="Azrou, P &amp; T. J. Wood" country="Morocco" elevation="1800" location="Fes-Meknes" municipality="Azrou" pageId="48" pageNumber="49" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<collectingCountry id="F3132A97FFB8FF8608E7FC6FFF4FFC54" box="[189,337,972,998]" name="Morocco" pageId="48" pageNumber="49">MOROCCO</collectingCountry>
•
<specimenCount id="9D02A18EFFB8FF860933FC6FFF9EFC54" box="[361,384,972,998]" count="1" pageId="48" pageNumber="49" type="female">♀</specimenCount>
;
<location id="8EDB3CDCFFB8FF8609D7FC6FFC01FC54" LSID="urn:lsid:plazi:treatment:03ADDB11FFA7FF870A42FA69FDA4FCC7:8EDB3CDCFFB8FF8609D7FC6FFC01FC54" box="[397,543,972,998]" country="Morocco" municipality="Azrou" name="Fes-Meknes" pageId="48" pageNumber="49">Fès-Meknès</location>
,
<collectorName id="26F10FD1FFB8FF860A73FC6FFC8FFC54" box="[553,657,972,998]" pageId="48" pageNumber="49">
<collectingMunicipality id="6BDFF07DFFB8FF860A73FC6FFC6BFC54" box="[553,629,972,998]" pageId="48" pageNumber="49">Azrou</collectingMunicipality>
, P
</collectorName>
7217,
<locationDeviation id="9A097325FFB8FF860A83FC6FFDB4FC54" box="[729,938,972,998]" location="Azrou" pageId="48" pageNumber="49">
<quantity id="4CFCC7E2FFB8FF860A83FC6FFD3CFC54" box="[729,802,972,998]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="48" pageNumber="49" unit="km" value="10.0">10 km</quantity>
S of Azrou
</locationDeviation>
;
<elevation id="00298D34FFB8FF860BEFFC6FFA52FC54" box="[949,1100,972,998]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="48" pageNumber="49" unit="m" value="1800.0">
<quantity id="4CFCC7E2FFB8FF860BEFFC6FFA13FC54" box="[949,1037,972,998]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="48" pageNumber="49" unit="m" value="1800.0">1800 m</quantity>
a.s.l.
</elevation>
;
<date id="FFBA4CC7FFB8FF860C02FC6FFAE9FC54" box="[1112,1271,972,998]" pageId="48" pageNumber="49" value="2022-05-18">
<collectingDate id="EFFEB52FFFB8FF860C02FC6FFAE9FC54" box="[1112,1271,972,998]" pageId="48" pageNumber="49" value="2022-05-18">18 May 2022</collectingDate>
</date>
;
<collectorName id="26F10FD1FFB8FF860D5BFC6FFB68FC54" box="[1281,1398,972,998]" pageId="48" pageNumber="49">T.J. Wood</collectorName>
leg.; BOLD: WPATW686-22;
<collectionCode id="ED15F2C2FFB8FF860A7BFC4CFC6FFBBB" box="[545,625,1007,1033]" pageId="48" pageNumber="49">TJWC</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BBB6A07FFB8FF8608B0FB93FA02FBF9" blockId="48.[189,1240,1072,1134]" box="[234,1052,1072,1099]" pageId="48" pageNumber="49">
<emphasis id="B970B615FFB8FF8608B0FB93FFB8FBF8" bold="true" box="[234,422,1072,1098]" pageId="48" pageNumber="49">Other material</emphasis>
(identified as
<taxonomicName id="4C041184FFB8FF860A17FB93FD06FBF8" authorityName="Wood" authorityYear="2021" box="[589,792,1072,1098]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="atlantea">
<emphasis id="B970B615FFB8FF860A17FB93FD06FBF8" box="[589,792,1072,1098]" italics="true" pageId="48" pageNumber="49">Andrena atlantea</emphasis>
</taxonomicName>
; see also Wood 2021)
</paragraph>
<paragraph id="8BBB6A07FFB8FF8608E7FBF7FAC6FBDC" blockId="48.[189,1240,1072,1134]" box="[189,1240,1107,1134]" pageId="48" pageNumber="49">
<materialsCitation id="3B6C605AFFB8FF8608E7FBF7FACAFBDC" box="[189,1236,1107,1134]" collectingDate="2022-05-18" collectionCode="TJWC" collectorName="Azrou, P &amp; T. J. Wood" country="Morocco" elevation="1800" location="Fes-Meknes" municipality="Azrou" pageId="48" pageNumber="49" specimenCount="1" specimenCount-female="1">
<collectingCountry id="F3132A97FFB8FF8608E7FBF7FF4FFBDC" box="[189,337,1108,1134]" name="Morocco" pageId="48" pageNumber="49">MOROCCO</collectingCountry>
•
<specimenCount id="9D02A18EFFB8FF860930FBF7FF89FBDC" box="[362,407,1108,1134]" count="1" pageId="48" pageNumber="49" type="female">1 ♀</specimenCount>
; same collection data as for paratype; BOLD: WPATW551-22;
<collectionCode id="ED15F2C2FFB8FF860CDEFBF7FACAFBDC" box="[1156,1236,1108,1134]" pageId="48" pageNumber="49">TJWC</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31E398CFFB8FF8708E7FB36FF27FD37" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49" type="description">
<paragraph id="8BBB6A07FFB8FF8608E7FB36FF50FB1D" blockId="48.[189,334,1173,1199]" box="[189,334,1173,1199]" pageId="48" pageNumber="49">
<heading id="D0F3DD6BFFB8FF8608E7FB36FF50FB1D" bold="true" box="[189,334,1173,1199]" fontSize="11" level="3" pageId="48" pageNumber="49" reason="3">
<emphasis id="B970B615FFB8FF8608E7FB36FF50FB1D" bold="true" box="[189,334,1173,1199]" pageId="48" pageNumber="49">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BBB6A07FFB8FF8608B0FB67FF5BFB6C" blockId="48.[189,745,1220,1281]" box="[234,325,1220,1246]" pageId="48" pageNumber="49">
<emphasis id="B970B615FFB8FF8608B0FB67FF5BFB6C" bold="true" box="[234,325,1220,1246]" pageId="48" pageNumber="49">Female</emphasis>
</paragraph>
<paragraph id="8BBB6A07FFB8FF8608E7FB44FCF7FAB3" blockId="48.[189,745,1220,1281]" box="[189,745,1255,1281]" pageId="48" pageNumber="49">
MEASUREMENTS. Body length
<quantity id="4CFCC7E2FFB8FF860A41FB44FC41FAB3" box="[539,607,1255,1281]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="48" pageNumber="49" unit="mm" value="6.0">6 mm</quantity>
(
<figureCitation id="133F7682FFB8FF860A35FB44FCC3FAB3" box="[623,733,1255,1281]" captionStart="Fig" captionStartId="49.[189,232,1882,1908]" captionTargetBox="[189,1399,947,1848]" captionTargetId="figure-175@49.[189,1399,947,1848]" captionTargetPageId="49" captionText="Fig. 28. Andrena (Micrandrena) anammas sp. nov., holotype, ♀ (OÖLM). A. Lateral habitus. B. Face, frontal view. C. Scutum, dorsal view. D. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10453558" httpUri="https://zenodo.org/record/10453558/files/figure.png" pageId="48" pageNumber="49">Fig. 28A</figureCitation>
).
</paragraph>
<paragraph id="8BBB6A07FFB8FF8608E7FA8BFF14F9EF" blockId="48.[189,1399,1320,1629]" pageId="48" pageNumber="49">
HEAD. Dark, 1.1 times as wide as long (
<figureCitation id="133F7682FFB8FF860AEAFA8BFD01FAF0" box="[688,799,1320,1346]" captionStart="Fig" captionStartId="49.[189,232,1882,1908]" captionTargetBox="[189,1399,947,1848]" captionTargetId="figure-175@49.[189,1399,947,1848]" captionTargetPageId="49" captionText="Fig. 28. Andrena (Micrandrena) anammas sp. nov., holotype, ♀ (OÖLM). A. Lateral habitus. B. Face, frontal view. C. Scutum, dorsal view. D. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10453558" httpUri="https://zenodo.org/record/10453558/files/figure.png" pageId="48" pageNumber="49">Fig. 28B</figureCitation>
). Clypeus elongate, not much broader than long, apically distinctly exceeding lower level of compound eyes, evenly domed, with large punctures, punctures separated by 1–2 puncture diameters; underlying surface medially and basally with granular microreticulation, weakly shining, apically more strongly shining. Process of labrum narrow, rounded, as long as broad. Gena equalling width of compound eye; ocelloccipital distance 0.5 times diameter of lateral ocellus. Foveae narrow, occupying ⅓ of space between lateral ocellus and compound eye, very slightly narrowing ventrally at level of antennal insertions; foveae filled with whitish hairs. Face, gena, vertex, and scape with white hairs. Antennae dark, A3 equalling or very slightly exceeding length of A4+5.
</paragraph>
<paragraph id="8BBB6A07FFB8FF8608E7F920FAACF868" blockId="48.[188,1401,1667,2011]" pageId="48" pageNumber="49">
MESoSoMA. Scutum and scutellum with granular microreticulation, weakly shining, regularly punctate, punctures separated by 0.5–1 puncture diameter (
<figureCitation id="133F7682FFB8FF860B55F905FD60F973" box="[783,894,1702,1729]" captionStart="Fig" captionStartId="49.[189,232,1882,1908]" captionTargetBox="[189,1399,947,1848]" captionTargetId="figure-175@49.[189,1399,947,1848]" captionTargetPageId="49" captionText="Fig. 28. Andrena (Micrandrena) anammas sp. nov., holotype, ♀ (OÖLM). A. Lateral habitus. B. Face, frontal view. C. Scutum, dorsal view. D. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10453558" httpUri="https://zenodo.org/record/10453558/files/figure.png" pageId="48" pageNumber="49">Fig. 28C</figureCitation>
). Pronotum rounded. Mesepisternum with strong granular microreticulation, dull. Dorsolateral parts of propodeum with granular microreticulation overlain by additional network of raised reticulation. Propodeal triangle laterally delineated by raised carinae, internal surface with pattern of strongly raised rugae, interspaces shining. Mesepisternum laterally with whitish hairs, scutum and scutellum with shorter whitish hairs. Propodeal corbicula incomplete, dorsal fringe composed of whitish plumose hairs, internal surface with long, white simple hairs. Legs dark, pubescence brownish. Flocculus incomplete, composed of plumose hairs; flocculus, femoral scopae, and tibial scopae composed of white simple hairs. Hind tarsal claws with inner tooth. Wings hyaline, stigma dark brown to black, venation dark brown, nervulus interstitial.
</paragraph>
<paragraph id="8BBB6A07FFB9FF8708E7FEAAFD86FDA8" blockId="49.[187,1399,265,538]" pageId="49" pageNumber="50">
METASoMA. Terga dark, apical margins at most with apical rims narrowly and obscurely lightened dark brown (
<figureCitation id="133F7682FFB9FF870901FE8FFFD2FEF4" box="[347,460,300,326]" captionStart="Fig" captionStartId="49.[189,232,1882,1908]" captionTargetBox="[189,1399,947,1848]" captionTargetId="figure-175@49.[189,1399,947,1848]" captionTargetPageId="49" captionText="Fig. 28. Andrena (Micrandrena) anammas sp. nov., holotype, ♀ (OÖLM). A. Lateral habitus. B. Face, frontal view. C. Scutum, dorsal view. D. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10453558" httpUri="https://zenodo.org/record/10453558/files/figure.png" pageId="49" pageNumber="50">Fig. 28D</figureCitation>
). Tergal margins relatively broad, on T2 occupying ⅓ of visible segment, on T3–4 occupying slightly more than ½ of visible segment. Tergal discs microreticulate, weakly shining, obscurely punctate, punctures disappearing into underlying sculpture, separated by 1–2 puncture diameters. T2–4 laterally with narrow apical hair fringes on apex of tergal margins, obscuring underlying surface, broadly interrupted medially. Apical fringe of T5 and hairs flanking pygidial plate dark brown, hairs of T5 laterally overlain by long white hairs. Pygidial plate rounded triangular, medially with elevated longitudinal midline, surface with obscure punctures.
</paragraph>
<paragraph id="8BBB6A07FFB9FF8708B0FDEBFF34FDD0" blockId="49.[189,313,584,645]" box="[234,298,584,610]" pageId="49" pageNumber="50">
<emphasis id="B970B615FFB9FF8708B0FDEBFF34FDD0" bold="true" box="[234,298,584,610]" pageId="49" pageNumber="50">Male</emphasis>
</paragraph>
<paragraph id="8BBB6A07FFB9FF8708E7FDC8FF27FD37" blockId="49.[189,313,584,645]" box="[189,313,619,645]" pageId="49" pageNumber="50">Unknown.</paragraph>
</subSubSection>
<subSubSection id="C31E398CFFB9FF8708E7FD16FAF0FD4F" box="[189,1262,693,766]" pageId="49" pageNumber="50" type="discussion">
<paragraph id="8BBB6A07FFB9FF8708E7FD16FF31FD7D" blockId="49.[189,303,693,719]" box="[189,303,693,719]" pageId="49" pageNumber="50">
<heading id="D0F3DD6BFFB9FF8708E7FD16FF31FD7D" bold="true" box="[189,303,693,719]" fontSize="11" level="3" pageId="49" pageNumber="50" reason="3">
<emphasis id="B970B615FFB9FF8708E7FD16FF31FD7D" bold="true" box="[189,303,693,719]" pageId="49" pageNumber="50">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BBB6A07FFB9FF8708E7FD40FAF0FD4F" blockId="49.[189,1262,739,766]" box="[189,1262,739,766]" pageId="49" pageNumber="50">
The species was collected from small
<taxonomicName id="4C041184FFB9FF870A21FD40FCC3FD4F" box="[635,733,739,765]" class="Magnoliopsida" family="Brassicaceae" genus="Alyssum" kingdom="Plantae" order="Brassicales" pageId="49" pageNumber="50" phylum="Tracheophyta" rank="genus">
<emphasis id="B970B615FFB9FF870A21FD40FCC3FD4F" box="[635,733,739,765]" italics="true" pageId="49" pageNumber="50">Alyssum</emphasis>
</taxonomicName>
(
<taxonomicName id="4C041184FFB9FF870AB6FD40FD99FD4F" baseAuthorityName="Wood" baseAuthorityYear="2023" box="[748,903,739,765]" class="Magnoliopsida" family="Brassicaceae" kingdom="Plantae" order="Brassicales" pageId="49" pageNumber="50" phylum="Tracheophyta" rank="family">Brassicaceae</taxonomicName>
) flowers in subalpine habitats.
</paragraph>
</subSubSection>
<subSubSection id="C31E398CFFB9FF8708E7FC8FFDA4FCC7" box="[189,954,812,885]" pageId="49" pageNumber="50" type="distribution">
<paragraph id="8BBB6A07FFB9FF8708E7FC8FFF48FCF4" blockId="49.[189,342,812,838]" box="[189,342,812,838]" pageId="49" pageNumber="50">
<heading id="D0F3DD6BFFB9FF8708E7FC8FFF48FCF4" bold="true" box="[189,342,812,838]" fontSize="11" level="3" pageId="49" pageNumber="50" reason="3">
<emphasis id="B970B615FFB9FF8708E7FC8FFF48FCF4" bold="true" box="[189,342,812,838]" pageId="49" pageNumber="50">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BBB6A07FFB9FF8708E7FCF8FDA4FCC7" blockId="49.[189,954,859,885]" box="[189,954,859,885]" pageId="49" pageNumber="50">High altitude sites in the Moroccan Middle Atlas (Azrou, Ifrane).</paragraph>
</subSubSection>
</treatment>
</document>
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